identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
216587E00D2EFF96EFC4F7DE3317FF07.text	216587E00D2EFF96EFC4F7DE3317FF07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abantiades cephalocorvus Moore & Beaver & Velasco-Castrillón & Stevens 2020	<div><p>Abantiades cephalocorvus Moore &amp; Beaver, sp. nov.</p><p>Figs 2a, 3 a–d.</p><p>Material Examined:</p><p>Holotype ♂. In SAMA collection. 1♂, South Australia, Eyre Pen [insula]., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=135.98097&amp;materialsCitation.latitude=-31.579084" title="Search Plazi for locations around (long 135.98097/lat -31.579084)">Goog’s Lake</a>, 31 o 34’44.7”S, 135 o 58’51.5”E, 28 th May 2016. J. Forrest. Dissected M.D. Moore. Leg taken for DNA analysis (Spec. No. 1611, sample designation M 018). SAMA No. 31-019757.</p><p>Paratypes ♂. SAMA . 1♂, South Australia, Eyre Pen., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=135.98097&amp;materialsCitation.latitude=-31.579084" title="Search Plazi for locations around (long 135.98097/lat -31.579084)">Goog’s Lake</a>, 31 o 34’44.7”S, 135 o 58’51.5”E, 28 th May 2016. J. Forrest. SAMA No. 31-019758. SAMA , 1♂, West. Aust. Skirmish Hill. Dissected M.D. Moore. SAMA . 1♂, South Australia, Eyre Pen., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=135.98097&amp;materialsCitation.latitude=-31.579084" title="Search Plazi for locations around (long 135.98097/lat -31.579084)">Goog’s Lake</a>, 31 o 34’44.7” S, 135 o 58’51.5”E, 28 th May 2016. J. Forrest. Leg taken for DNA analysis. (Sample designation M 019). SAMA No. 31-019759. ANIC , 1♂, Napperby Ck., NT, 26 May 1968, K.J. McKie. Spec. Ref. No. 18197. Dissected M.D. Moore.</p></div>	https://treatment.plazi.org/id/216587E00D2EFF96EFC4F7DE3317FF07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moore, Michael D.;Beaver, Ethan P.;Velasco-Castrillón, Alejandro;Stevens, Mark I.	Moore, Michael D., Beaver, Ethan P., Velasco-Castrillón, Alejandro, Stevens, Mark I. (2020): Description of two new Australian species of Abantiades Herrich-Schäffer (Lepidoptera: Hepialidae) and females of two further species with notes on their biogeography. Zootaxa 4822 (1): 71-93, DOI: 10.11646/zootaxa.4822.1.3
216587E00D2EFF97EFC4F1EA3577FB83.text	216587E00D2EFF97EFC4F1EA3577FB83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abantiades Herrich-Schaffer 1855	<div><p>Systematics of Abantiades</p><p>Molecular analyses</p><p>(see Fig. 1)</p><p>The Bayesian analysis for each species (Fig. 1) reveals a well supported (PP = 1.0) monophyletic short-beaked Abantiades clade compared to our six outgroup taxa. Although this is only based on the single mtDNA (COI) gene, it does corroborate the morphological distinction of the two new species from the six other Abantiades species in our analysis, which are each also defined by unique morphological characters (see Taxonomy section below). Supplementary Table S1 details all specimens used, including GenBank accessions and locality information. For instances where we obtained multiple individuals within species we found close intraspecific sequence divergences (Supplementary Table S2). For example, the greatest sequence divergence observed within any of our species varied between 0.6–3.3%, indicating variation that we would expect within species included here (Supplementary Table S2). These distances also correspond to those reported elsewhere for Lepidoptera that generally indicate below 3% intraspecific while above 3% interspecific comparisons using the same COI gene (e.g. Grund et al. 2019; Beaver et al. 2020). However, sequence divergences have been found to vary outside these ranges for some Abantiades species (Simonsen et al. 2019; Moore et al. 2020).</p><p>Sequence divergence comparisons between all Abantiades species compared to our outgroup taxa were between 8.6–12.5% (Supplementary Table S2). Examining between Abantiades species reveal interspecific sequence divergences of up to 10.2% (between A. cephalocorvus sp. nov. and A. labyrinthicus (Donovan)) and up to 6.8% between the ‘triforked’ clade, which includes all Abantiades species in Fig. 1 except A. labyrinthicus . We also observed several interspecific sequence divergences that are below 3%, for example A. pallida, A. kristenseni, and A. tembyi sp. nov. and A. inexpecta (Supplementary Table S2), which has been observed in other ghost moths (e.g. Moore et al. 2020). Although sequence divergence can be low between a number of Abantiades species, Fig. 1 indicates that these are each monophyletic species (PP&gt; 75%) that correspond to morphological differences (see Taxonomy section below). Comparisons between these trees and the sequence divergences of our new species described (see below) reveal that A. cephalocorvus sp. nov. is 3.9–5.0% divergent from its closest neighbouring species ( A. kristenseni), while Abantiades tembyi sp. nov. differs from its closest neighbour ( A. pallida) by 2.3–4.4%.</p></div>	https://treatment.plazi.org/id/216587E00D2EFF97EFC4F1EA3577FB83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moore, Michael D.;Beaver, Ethan P.;Velasco-Castrillón, Alejandro;Stevens, Mark I.	Moore, Michael D., Beaver, Ethan P., Velasco-Castrillón, Alejandro, Stevens, Mark I. (2020): Description of two new Australian species of Abantiades Herrich-Schäffer (Lepidoptera: Hepialidae) and females of two further species with notes on their biogeography. Zootaxa 4822 (1): 71-93, DOI: 10.11646/zootaxa.4822.1.3
216587E00D2EFF97EFC4F422355DF9AF.text	216587E00D2EFF97EFC4F422355DF9AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abantiades Herrich-Schaffer 1855	<div><p>Genus Abantiades Herrich-Schäffer, [1855]</p><p>Diagnosis:</p><p>Simonsen (2018) produced a new diagnosis for the genus: “A patch of elongate scales at the forewing base and the ‘sensory tubercles’ on S2 are both unique and diagnostic for Abantiades .” He also noted that they have hepialine venation (Dumbleton 1966; Dugdale 1994), and monopectinate antennae that lack scales on the flagellum.</p></div>	https://treatment.plazi.org/id/216587E00D2EFF97EFC4F422355DF9AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moore, Michael D.;Beaver, Ethan P.;Velasco-Castrillón, Alejandro;Stevens, Mark I.	Moore, Michael D., Beaver, Ethan P., Velasco-Castrillón, Alejandro, Stevens, Mark I. (2020): Description of two new Australian species of Abantiades Herrich-Schäffer (Lepidoptera: Hepialidae) and females of two further species with notes on their biogeography. Zootaxa 4822 (1): 71-93, DOI: 10.11646/zootaxa.4822.1.3
216587E00D25FF99EFC4F1EA35FEF8B3.text	216587E00D25FF99EFC4F1EA35FEF8B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abantiades tembyi Moore & Beaver & Velasco-Castrillón & Stevens 2020	<div><p>Abantiades tembyi Moore &amp; Beaver, sp. nov.</p><p>Figs 2b, 4 a–d.</p><p>Material Examined:</p><p>Holotype ♂. In SAMA collection. South Australia, Eyre Pen., Ceduna, 32 o 07’27.5” S, 133 o 49’27.3” E, 28 th April 2014, N. Temby. Holotype label attached.</p><p>Paratypes ♂. SAMA, 1♂, South Australia, Eyre Pen., Ceduna, 32 o 07’27.5” S, 133 o 49’27.3” E, 28 th April 2014, N. Temby. Dissected M.D. Moore. Leg taken for DNA analysis Spec. No. S 043, DNA sample designation M032. SAMA, 1♂, South Australia, Eyre Pen., Ceduna, 32 o 07’27.5” S, 133 o 49’27.3” E, 28 th April 2014, N. Temby. Spec. No. 17016. SAMA, 1♂, 23 rd April 2017, 30 km E by N of Swan Reach, S. Aust., 34 o 31.478’ S, 139 o 54.879’ E, Coll: M. Hura. SAMA, 2 ♂, M. H. Hopgood, Yurgo. SAMA Nos 31-019502, 31-019793 ANIC, 1♂, Australia: South Australia, Ceduna area during 13.4 mm rain, 28 April 2014 Leg. N. Temby, Abantiades sp. indet. et nov., det. J.E. Nielsen. ANIC, 1♂, 23 rd May 2017, 30 km E by N of Swan Reach, S. Aust., 34 o 31.478’ S, 139 o 54.879’ E, Coll: M. Hura. John Nielsen Private Collection. 1♂, Australia: South Australia, Ceduna area during 13.4 mm rain, 28 April 2014 Leg. N. Temby, Abantiades sp. indet. et nov., det. J.E. Nielsen. Spec. No. 17270, DNA Sample designation H063. Dissected M.D. Moore. Mark Hura private collection, 1♂, 10 th May 2015, 30 km E by N of Swan Reach, S. Aust., 34 o 31.478’ S, 139 o 54.879’ E, Coll: M. Hura. Spec. no. 17261. DNA Sample designation H061. Mark Hura private collection, 1♂, 19 th May 2019, 30 km E by N of Swan Reach, S. Aust., 34 o 31.478’ S, 139 o 54.879’ E, Coll: M. Hura. E. Beaver private collection, 1♂, 23 rd April 2017, 30 km E by N of Swan Reach, S. Aust., 34 o 31.478’ S, 139 o 54.879’ E, Coll: E. P. Beaver + M. Hura. Spec. No. 19019, DNA Sample designation G047, coll. E. P. Beaver and M. Hura, 1♂, 25 APR 2020, 30 km E by N of Swan Reach, S. Aust., 34 o 29.604’ S, 139 o 54.926’ E, Coll: M. Hura. MusVic, 1♂, Timberoo Station, Walpeup, Vic., 10-5-1920, (G. Lyell Coll. – presented 21-7-32.) Registration No. HET – 44837.</p><p>Diagnosis:</p><p>A medium sized species found east of the Nullarbor in mallee habitats, that has both the fore and hind wings dark brown, but with the forewing covered in fine white or pale grey scrolling lines over most of its surface. Males of Abantiades tembyi sp. nov. differ from all other species of Abantiades by: i) the shape of the sternite VIII (Fig. 4d), ii) the narrow set, relatively short, narrowly inwardly curved vincular arms (Fig. 4c), and iii) the heavily sclerotized, erect, pseudoteguminal lobes with prominent triangular processes and long concave disto-posterior margins (Fig. 4a). Analysis of the mtDNA (COI) will also allow positive identification. For a complete comparison of the species within this clade, and other triforked Abantiades species likely to be found within the range, see the Remarks section below.</p><p>Description:</p><p>Male</p><p>Head: Ventrally clothed in elongate red-straw coloured scales, fronto-clypeal region red-straw coloured scales tipped grey. Two large compound eyes, each only slightly smaller than head capsule. Labial palps three segmented, second palpomere longest, not straight and slightly angled laterally near the terminal end, third smallest, sub-spherical, clothed in reddish, straw coloured scales. Antennae total 72 segments, monopectinate but dividing almost immediately into triforked rami, red-brown. In resting position central fork projects downwards, others laterally to anterior and posterior, then vertically downwards until tips level with tip of central fork.</p><p>Thorax: Clothed in elongate piliform grey scales, dorsal darker. Forewing length, 38–55 mm, veins red-brown. Dorsal surface; area between costa and Rs thickly covered by short scales, grey near base, browner towards apex, and interspersed with scattered whitish scales. Discoidal and submarginal bands broad, white, linear; discoidal band with irregular margin and comprising densely packed shining white scales, margin highlighted with dark brown scales. In anterior of discoidal cell grey or brown scroll lines. Posterior to white line, thick area of dark brown scaled highlighting particularly towards apical end. Submarginal mark variable in shape, length, and amount of highlighting. Sides sub parallel swollen in middle, apex to CuA2, filled with shining white scales. In some specimens the area between the two major white marks can be brown and largely empty of markings, or have irregular white spots, or have fine white scrolling, or be bridged by white scales linking the discoidal and sub-marginal marks. A sub-terminal line suggested, edged white, but this can be series of diamond shapes. Grey/white scrolling lines extensive. Anterior portion of wing, two or three scales thick, large shapes, posterior and termen scrolling finer, only one scale thick, often parallel lines though larger shapes can be present. Ventral surface; forward Sc sparingly covered, longish brown piliform scales, rest sparsely covered, long straw-brown scales. Hindwing; 30–40 mm. Wing membranes only slightly opaque. Dorsal surface; Veins straw-brown. Forward of Sc dense covering longish brown scales. Rest sparsely covered, longish straw-coloured scales. Ventral surface; Forward of Sc densely covered smaller light brown scales. Rest sparsely covered in long light brown piliform scales. Legs; Dorsal coloured grey, lateral and ventral covered in longer lighter scales. Arolium and epiphysis present.</p><p>Abdomen: All sternites and tergites densely covered in dark grey, long, piliform scales. Sometimes dorsally, there can be a light coloured scale band at the meeting of abdomen and thorax, with lighter bases to the hind wings and light coloured tergites I and II. Sternite VIII (S8) (Fig. 4d) Anterior margin continuously rounded with deep central concavity. Posterior margin shallow convex with flattened central portion. Posterior corners sharply pointed.</p><p>Genitalia: (Figs 4 a–c). Arms of vinculum concave laterally. Meet producing a broad base. Saccus deep, slightly concave sides, bell shaped, posterior margin of ventral face, flat with central raised mesa like lobe. Posterior margin of dorsal face broad shallow “V” shape. Distal twin tergal processes short and stubby, very broad, roundly obtuse ends. Pseudoteguminal lobes, with disto-posterior process set forward, extended flattish posterior. Dorso-posterior margin almost straight. Posterior corner short, broad based, pointed. Disto-posterior margin strongly concave ending steeply in the disto-posterior process, the latter acuminate, rear margin steep, anterior margin slightly convex, tip roundly obtuse. Ventro-posterior margin convex, but at pseudoteguminal arms distinctly concave. Arms beak like, approximately parallel for most of length, tip pointed. Ventro-dorso margin concave. Valva, basal sacculus relatively large (2.5 mm by 1 mm), distal processes (harpe) relatively thin (1.2 mm by 0.6 mm.), setose.</p><p>Female. Unknown.</p><p>Distribution and Biology:</p><p>Recorded Localities: (Figs 11, 12 c–d) About 12 kms east of Ceduna; 30 kms ENE of Swan Reach; and Yurgo, SA, as well as a specimen from near Walpeup, Victoria. These four locations are along a line 850 km in length. Whether we are looking at two or more isolated populations, brought about by development and land clearance or a single population not fully sampled is uncertain.</p><p>Flight times: Most probably highly dependent on rainfall. These moths were collected 23 rd April to 19 th May.</p><p>Larval biology: Unknown.</p><p>Etymology:</p><p>An eponym honouring the extensive collecting work done by, and for the support and friendship given by Nick Temby to my work in this Family, and for his collection of the first specimens.</p><p>Remarks:</p><p>See above for intra-cladal differences and for comparison to A. cephalocorvus sp. nov. In Fig. 1, A. tembyi sp. nov. is most similar to A. pallida, which is not surprising given their morphological similarities (see below).</p><p>In appearance A. tembyi sp. nov. differs from A. kristenseni by the former having a dark brown forewing broadly covered in very fine grey scaled scrolling lines and dark brown/grey hindwing whereas A. kristenseni has a mid-brown forewing with large white and more distinctive scrolling lines and shapes in the apical and terminal areas of the wing. The hindwing is a light, yellow-brown giving the moth a distinct two-tone colouration. The genitalia differ in many ways but most particularly in the shape of the pseudoteguminal lobe. Abantiades tembyi sp. nov. has a rather irregularly linear dorso-posterior margin, an angled dorso-posterior corner, and a long deeply concave disto-posterior margin that produces an acuminate disto-posterior process. Abantiades kristenseni on the other hand has virtually no dorso-posterior margin, a large well rounded dorso-posterior corner and a gentler concave distoposterior margin that produces a broad smooth rounded disto-posterior process. The twin processes differ markedly too, A. kristenseni having cylindrical twin processes with slightly clubbed ends, where A. tembyi sp. nov. has broad, flat, processes with blunt ends.</p><p>To differentiate A. tembyi sp. nov. (Fig. 2b) from A. pallida (Fig. 8a), sternite VIII and genitalia must be examined (see Figs 4 a–d, 5a–d). Some differences can be seen without dissection. The disto-posterior margin on the pseudoteguminal lobe of A. tembyi sp. nov. is a long deep concavity producing a steep verticality towards the apex, resulting in the disto-posterior process being triangular with a long flat posterior margin. The same margin in A. pallida is much less concave resulting in a less steep posterior margin, producing a disto-posterior process that is rounded and near hemispherical in A. pallida and a short curved posterior margin. The twin processes are also quite different. In A. tembyi sp. nov., they are short, failing to make the dorso-posterior corner, broad and round topped, in A. pallida whilst they are broad based, they are longer, easily make the dorso-posterior corner and are conical and pointed. The apodemal aspect of the vincular arms differs in the two species. In A. tembyi sp. nov. they are slender and more erect resulting in a narrow genital structure and a bell shaped saccus but in A. pallida the arms are more massive and more concave, producing a much broader genital structure and a different shaped saccus (Figs 4c and 5c). Sternite VIII is different in the two species too. Abantiades tembyi sp. nov. has inwardly curved sides and an anterior margin with a deep central concavity, in A. pallida the structure has stepped sides and flattish anterior and posterior margins.</p><p>The first specimen of A. tembyi sp. nov. to have its DNA sequenced was collected at Ceduna in SA, and although suspected to be of the A. pallida group, Simonsen et al. (2019) discovered that its DNA was notably different from A. pallida specimens from Western Australia and they did not list the specimen as an example of that species.</p></div>	https://treatment.plazi.org/id/216587E00D25FF99EFC4F1EA35FEF8B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moore, Michael D.;Beaver, Ethan P.;Velasco-Castrillón, Alejandro;Stevens, Mark I.	Moore, Michael D., Beaver, Ethan P., Velasco-Castrillón, Alejandro, Stevens, Mark I. (2020): Description of two new Australian species of Abantiades Herrich-Schäffer (Lepidoptera: Hepialidae) and females of two further species with notes on their biogeography. Zootaxa 4822 (1): 71-93, DOI: 10.11646/zootaxa.4822.1.3
216587E00D21FF9AEFC4F1A231BEFBEF.text	216587E00D21FF9AEFC4F1A231BEFBEF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abantiades macropusinsulariae Simonsen 2018	<div><p>Abantiades macropusinsulariae Simonsen, 2018 (female)</p><p>Figs 6 a–b, 7a–d.</p><p>Material Examined:</p><p>SAMA. 1♀, SA, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=137.59464&amp;materialsCitation.latitude=-35.59658" title="Search Plazi for locations around (long 137.59464/lat -35.59658)">Kangaroo Island</a>, North Cape Road, Bay of Shoals, M+ M Moore, 35 o 35’47.7”S 137 o 35’40.7”E, 2 May 2019. Specimen No 19169. Leg removed for DNA analysis; Designation G 116. Dissected M.D. Moore 12 th Feb. 2020 . SAMA No. 31-021005.</p><p>Diagnosis:</p><p>Endemic to Kangaroo Island, the males of A. macropusinsulariae have patterned forewings with many scattered, irregular shaped, white coloured marks, and lighter coloured hindwings giving a two-tone effect to the specimens and they also have a red-brown antennal flagellum. Specifically, Simonsen (2018) states, “second segment of the labial palp very long, more than 3x the length of the terminal segment, S8 short and pentagonal with a medially slightly projecting posterior margin, distal pseudoteguminal lobes smoothly triangularly rounded without a concave curvature on the dorso-posterior margin, disto-posterior arm of vinculum with a small, lateral ridge; saccus in posteroventral view narrowly V-shaped”. The female of this species is black/grey and white, has patterned forewings and a red-brown antennal flagellum with distinctly triforked pectinations. Specifically, it varies from all other triforked Abantiades females by; i) the heavily sclerotized bands bordering the medial lobe of the antevaginal lamella, and ii) the long, conical end projection on the corpus bursae. The wings in both sexes of this species are shorter and broader than in the other species in this clade.</p><p>Description:</p><p>Female</p><p>Head: Head, thickly covered in medium grey piliform scales, fronto-clypeal scales darker. Two large compound eyes, each only slightly smaller than head capsule. Labial palps approximately straight, three segmented, second palpomere longest, third smallest, sub-spherical, all densely covered in brown coloured scales. Antennae: (Figs 7 c–d). total 48 segments, length 11–12 mm, 20% of forewing length; flagellum dark red-brown, monopectinate, basal pectinations tear-drop shaped, then transitional to triforked about 1/5 from base. Lateral projections red-brown approximately parallel sides with blunt, rounded end, central fork white, long triangular with narrowly acute tip.</p><p>Thorax: Densely clothed in medium-grey piliform scales. Forewing: length 48 mm, membrane whitish opaque, veins white, thinly covered with short, dark grey scales. Dorsal surface: space between costa and Sc vein thickly covered in small piliform grey scales particularly near base. Two large white areas; discoidal, filled with dull white scales, and in this specimen divided medially into two irregular patches, patch of dark brown/grey scales along anterior edge in basal half. Sub-terminal band dull white, sublinear between M3 and near apex. Also, a series of intervein, irregular white shapes along termen. White, strongly marked scrolling lines in apical area, less apparent elsewhere. Circular white mark present on posterior wing half way along the dorsum (Fig. 6b – arrow). White markings edged in blackish-brown scales with lighter, grey scales filling the inter-scrolled spaces. Ventral surface unmarked but with dorsal markings faintly visible, sparsely covered by reddish-brown digitiform scales that are longer near dorsum particularly near the base. Hindwing: 39 mm, dorsal surface with whitish veins, and elongate reddish-brown piliform scales. Ventral surface, as for forewing. Legs: Dorsal and lateral surfaces densely covered in grey piliform scales, longer laterally, ventrally covered in sandy red scales. Epiphysis and arolium present.</p><p>Abdomen: Covered in long dark grey piliform scales.</p><p>Genitalia: (Figs 7 a–b). Dorsal plate lightly sclerotized, bilobed, with deep “V” shaped central invagination of plate. Outer margin curved with triangular prominences in central region of each lobe (Fig. 7a – arrow).Antevaginal lamella trilobate with broad areas of connection between lateral and medial plates. Medial plate more heavily sclerotized, dorsal margin centrally raised with minute bilobed prominence and heavily sclerotized convex sides. Long, thick setae on prominence and generally over central surface of medial plate. Also, on inner margins of lateral lobes and on lobes of dorsal plate. Sub-anal plates sclerotized, pentagonal. Corpus bursae broad at junction with narrow ductus bursae and then tapering as a long conical terminal. No diverticulum observed.</p><p>Distribution and Biology:</p><p>Recorded Localities: (Figs 10, 12 e–f) Coastal, previously only from Vivonne Bay (south-central of Island), but now also on North Cape (NE of Island).</p><p>Flight times: April 19 th to May 3 rd (Simonsen 2018). The female and an associated male were collected on the 2 nd of May.</p><p>Larval biology: Unknown.</p><p>Remarks:</p><p>The female specimen (MT052730) in Fig. 1 was initially identified as A. macropusinsulariae by sequencing of the mtDNA (COI) gene, and groups with the four other male A. macropusinsulariae specimens. All five A. macropusinsulariae specimens are closely related and form a sister group to A. pallida and A. tembyi sp. nov.</p><p>Comparison with the females of A. inexpecta, A. cephalocorvus sp. nov., and A. tembyi sp. nov. is not possible because none have been located in collections. We can compare the females of the three remaining species if we use Simonsen (2018) for the features of A. kristenseni . In A. macropusinsulariae the structure of the genitalia is particularly interesting as the heavily sclerotized bands bordering the medial lobe of antevaginal lamella are not present in any other as yet dissected Abantiades female, triforked or otherwise. These bands presumably provide rigidity. In our opinion this provides a three dimensionality to the structure not seen in any other triforked Abantiades . The only other female Abantiades that shows any marked three dimensionality in its structure is A. equipalpus Moore, 2014 . The corpus bursae is also unique in having a long taper at its distal end, that was particularly notable upon first opening the abdomen, the taper being bent forward over the whole structure.</p><p>Abantiades macropusinsulariae is a dark grey/black coloured moth with distinct discoidal and submarginal white marks whereas both A. kristenseni and A. pallida are light grey in colour. In the specimens available it seems that A. kristenseni will usually have two large marks on the forewings, discoidal and submarginal, whereas A. pallida will only have a submarginal mark. Upon dissection A. macropusinsulariae stands out having a narrow medial lobe to the antevaginal lamella edged in thick sclerotization, more regularly shaped, more rounded and more widely separated lobes on the dorsal plate and sub-anal plates that are double the area of the other two species. Abantiades pallida and A. kristenseni both have shallow rounded lobes on the dorsal plate and no thick sclerotization on the medial lobe. Also, the medial lobe of the antevaginal lamella is over triple the width in A. kristenseni and the lateral processes on the inner margin are triangular and pointed whereas in A. pallida where the medial lobe is a little narrower than A. kristenseni (but still triple the width of A. macropusinsulariae) the lateral processes are ten percent longer, trapezoidal in form and blunt ended. The setose nipple like eruptions (Fig. 9 – arrow)) on the lateral lobes of the antevaginal lamella seen in A. pallida has not been seen on either of the two other females.</p><p>Endemic to Kangaroo Island, A. macropusinsulariae can be separated from the other two triforked species present on the island by its antennae. Abantiades macropusinsulariae has a dark red flagellum, distinctly three lobed pectinations with a long, pointed pale central fork. Abantiades atripalpis (Walker, 1856) antennae are tan/russet in colour, pectinations are more rhomboidal and with a shorter central fork, and scrolling lines that are fine and indistinct. Abantiades argentata has dark brown, black rami similar in shape to A. atripalpis, and brown forewings with more obvious scrolling lines.</p></div>	https://treatment.plazi.org/id/216587E00D21FF9AEFC4F1A231BEFBEF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moore, Michael D.;Beaver, Ethan P.;Velasco-Castrillón, Alejandro;Stevens, Mark I.	Moore, Michael D., Beaver, Ethan P., Velasco-Castrillón, Alejandro, Stevens, Mark I. (2020): Description of two new Australian species of Abantiades Herrich-Schäffer (Lepidoptera: Hepialidae) and females of two further species with notes on their biogeography. Zootaxa 4822 (1): 71-93, DOI: 10.11646/zootaxa.4822.1.3
216587E00D23FF86EFC4F51F3472FF06.text	216587E00D23FF86EFC4F51F3472FF06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abantiades pallida Simonsen 2018	<div><p>Abantiades pallida Simonsen, 2018 (female)</p><p>Figs 8 a–b, 9a–b.</p><p>Material Examined:</p><p>SAMA; 1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.486664&amp;materialsCitation.latitude=-30.821667" title="Search Plazi for locations around (long 117.486664/lat -30.821667)">West. Aust. Wheatbelt</a>, Koorda Caravan Park, Koorda, 30 o 49’18”S, 117 o 29’12”E, M+ M Moore 14 May 17. Spec. No., 17185, leg removed, for tissue, storage. M.D. Moore. Sample No H 043 . SAMA No. 31-020314 . SAMA; 1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=129.00194&amp;materialsCitation.latitude=-31.63861" title="Search Plazi for locations around (long 129.00194/lat -31.63861)">West. Aust.</a>; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=129.00194&amp;materialsCitation.latitude=-31.63861" title="Search Plazi for locations around (long 129.00194/lat -31.63861)">Nullarbor</a>, Quarantine Station, Border Village, 31 o 38’19”S, 129 o 00’07”E , R. Penning, 7 th May 2018. Spec. No. 18122, leg removed, for tissue, storage. M.D. Moore. Sample No H 122. Dissected E. Beaver, 16 th March 2020 . SAMA No. 31-020414 .</p><p>Diagnosis:</p><p>A large grey moth with a distinct white submarginal line but only an indistinct brown/grey coloured discoidal mark. Specifically, the presence of setose covered nipple shaped projections on the lateral lobes of the antevaginal lamella is unique in all described triforked females (Fig. 9 – arrow).</p><p>Description:</p><p>Female</p><p>Head: Dorsally covered in thick, elongate light grey scales, fronto-clypeal region dark grey. Each eye only slightly smaller than head capsule. Labial palps approximately straight, three segmented; basal palpomere thickly covered in long brown/grey scales, some whitish at base; second palpomere longest, thickly covered in light brown scales, darker ventrally; third palpomere, smallest, sub-spherical, densely covered in brown scales. Antennae: total 60 segments, 12–13 mm long, less than 20% forewing length; monopectinate, basal rami lamellar transitioning distally to spatulate then with lateral forks; rami cruciform on basal half then fully triple branched before spade shaped near the tip. Scattered robust dark setae and more numerous long colourless setae on central fork. Colourless setae present on upper surface of lateral fork and on faces. Pectinations lean slightly towards distal end and central fork a little more so.</p><p>Thorax: Dorsal surface densely clothed by medium grey, elongate, robust scales, slightly paler than head. Ventral surface light white grey. Forewing length 62–68 mm, membrane transparent, veins whitish, browner towards base, covered in small dark grey scales; dorsal costa covered in small grey/brown scales; costa to Rs and apically Rs1, sparsely covered in dark grey scales. Two large marks on forewing present, an indistinct discoidal mark, medium brown basally, whiter distally and a distinct dull white submarginal band that extends between Rs2 to CuA1, linear but wider centrally and highlighted by narrow border of dark grey scales. The anterior portion of the discoidal mark has three or four light grey scroll lines present then out to R and Rs1+2 a thin band of dark brown highlighting scales. Irregular obscure, white scaled scroll lines present over forewing, apically scroll marks elliptical, along termen, linear, with pale grey scales between lines. Dorsally and basally wing scales become longer and more red-brown, sparse throughout. Ventral surface: dorsal markings visible through wing; costa covered in small yellow-grey scales with longer grey-brown scales towards Sc. Apex with a few, thinly spread dark grey scales, rest of wing sparsely covered in white-grey piliform scales that are longer and more golden towards base. Hindwing length 45–53 mm, veins white-yellow, membrane transparent. Dorsal surface, sparsely covered in long light grey to reddish-brown scales, basally piliform golden scales. Ventral surface, as for ventral forewing. Legs: femur dorsal surface covered by short yellow sandy-brown scales, lateral scales light grey; inner lateral surface light grey; ventrally whitish grey. Tibia dorsally sandy-grey, ventrally sandy yellow. Tarsus, dorsally sandy-grey, ventrally sandy-yellow, increasingly golden towards distal tarsus. Arolium and epiphysis present.</p><p>Abdomen: Dorsal surface thickly covered in long, dark grey piliform scales. Ventrally thickly covered in piliform scales, sternites I–III light white grey, IV–V yellow-grey, VI–VIII yellowish.</p><p>Genitalia: (Fig. 9). Dorsal plate lightly sclerotized, bilobed, each hemispherical, central “V’ shaped invagination, inner margin setose. Antevaginal lamella trilobed; medial lobe more heavily sclerotized, setose, inner margin continuously curved with wide, blunt ended lateral projections and small concave depression centrally. Inner margin lightly sclerotized and membranous in some places. Lateral lobes with setose nipple like projection which is quite distinct in the more lateral view (see Fig. 9b –arrow). Subanal plates with heavily sclerotized distal margin. No bursa copulatrix was recovered.</p><p>Distribution and Biology:</p><p>Recorded Localities: (Fig. 11) Previously known only from Border Village, SA, and Eucla, WA (Simonsen 2018). Now also from Coolgardie; Bullabulling; Hyden; Koorda; and Kellerberrin all in Western Australia.</p><p>Flight times: Simonsen (2018) lists the flight period as between 28 th of April to the 11 th of May. The females examined here were collected on the 04 th and 14 th of May.</p><p>Larval biology: Unknown.</p><p>Remarks:</p><p>See above for intra-cladal differences and comparison to A. macropusinsulariae and A. kristenseni .</p><p>The females (MT052733 and MT052735) were identified as belonging to A. pallida by sequencing of the mtDNA (COI) gene, and are clustered within a clade that includes three male specimens (Fig. 1). Even though separated by hundreds of kilometres, neither of the two females had an obvious discoidal mark on their forewings, which is rare in Abantiades .</p><p>Abantiades pallida has a wide distribution that significantly overlaps with both A. kristenseni and A. zonatriticum Moore &amp; Beaver, 2020 (Moore et al. 2020). The females of the triforked Abantiades species are inherently difficult to identify without dissection. However, A. pallida has one obvious white submarginal line and only an almost obscure discoidal one, while the other two species have two obvious marks (either white or brown). Antennal structure is useful; A. zonatriticum has triangular shaped lateral processes on the antenna whereas in A. pallida they are linear. Upon dissection of the genitalia, A. zonatriticum has a process which seems like an extension to the medial lobe (rather than part of it) of the antevaginal lamella and projects more into the central space (has triangular shaped lateral processes), whereas with A. pallida the projection is more part of the medial lobe, has a smaller projection into the central space with blunt ended lateral processes trapezoidal in shape. The lateral lobes of the antevaginal lamella have a nipple like protrusion in A. pallida but not so in A. zonatriticum .</p></div>	https://treatment.plazi.org/id/216587E00D23FF86EFC4F51F3472FF06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moore, Michael D.;Beaver, Ethan P.;Velasco-Castrillón, Alejandro;Stevens, Mark I.	Moore, Michael D., Beaver, Ethan P., Velasco-Castrillón, Alejandro, Stevens, Mark I. (2020): Description of two new Australian species of Abantiades Herrich-Schäffer (Lepidoptera: Hepialidae) and females of two further species with notes on their biogeography. Zootaxa 4822 (1): 71-93, DOI: 10.11646/zootaxa.4822.1.3
