taxonID	type	description	language	source
207887EF2B09B74627F7FD15146C33DA.taxon	description	Description. Body size small (length 4.3 – 7.3 mm), shape oval, convex. General color shining black; antennomeres VII and VIII white, sometimes extending to IX and X. Head. Labrum transverse, with rounded anterior angles, anterior margin concave, dorsally with few setigerous punctures. Frontal tubercles subtriangular, with anterior angles divergent, basally separated from frons by impressed line. Interocular space wide, 3.2 – 3.3 times as wide as transverse diameter of eye. Interantennal space 1.2 – 1.3 times as wide as transverse diameter of antennal socket. Eyes small. Vertex wide, moderately convex and glabrous. Antennae with 11 antennomeres, as long as or slightly longer than body. Pronotum transverse, about 1.9 – 2.0 x as wide as long. Anterior margins broadly concave, posterior margin truncate or slightly concave, lateral margins posteriorly narrowed. Only lateral margins widely bordered; all margins without setae. Each angle bearing one setigerous puncture and a long pale seta. Surface glabrous, indistinctly finely punctate, with a pair of distinct depressions laterally. Scutellum large, subtriangular. Elytra convex, posteriorly widened, widest near apex, with punctation sparse and randomly arranged; without setae except several minute setae along longitudinal ridge. One distinct ridge arising from humerus, longitudinal, reduced before apical 1 / 3. Epipleura slender, gradually narrowed apically, and disappearing before apices. Brachelytrous. Hind wings absent. Legs slender, apices of all tibiae lacking spines. First metatarsomere elongate, 1.2 x as long as following two tarsomeres combined. All claws normal and appendiculate. Procoxal cavities closed, intercoxal prosternal process wide, not elevated, visible between procoxae. Posterior margin of last ventrite with short distinct incisions in male, central lobe evenly rounded and flat, disc largely and triangularly depressed. Aedeagus slender, parallel-sided, apically narrowed, moderately curved in lateral view; with one longitudinal endophallic sclerite, with one cluster of setae at basal 1 / 3, and basally bifurcate. Females. Abdomen with last ventrite truncate or slightly convex or notched (in S. tsoui sp. nov.). Spermatheca slender, as wide as pump, hardly separated from pump; pump wide and strongly curved, apex widely rounded, apex with transverse, short sclerite, size variable; proximal spermathecal duct hardly separated from receptacle, apically narrowed. Gonocoxae longitudinal, conjoined from base to middle, slightly narrowed at apical 1 / 3, apices rounded, each gonocoxa with seven to nine elongate setae. Ventrite VIII with only apical area and spiculum sclerotized; with dense elongate setae inside apical sclerotized area, apical margin rounded; spiculum elongate and wide.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B09B74627F7FD15146C33DA.taxon	diagnosis	Diagnosis. The genus Shairella is among the genera classified by Wilcox (1971 – 1975) in the section Adoxiites, characterized by the wide pronotum in combination with the unmargined basal border. It is similar to Shaira Maulik, 1936 but apart from the characters mentioned before, all species in Shairella have the humerus raised, whereas it is flat in Shaira. Also, the procoxal cavities in Shaira are open whereas they are closed in Shairella. The genus Khasia Jacoby, 1899 (type species Khasia kraatzi Jacoy, 1899) is characterized by a square pronotum which seems elongate due to the strong constriction in the basal half. Moreover, Khasia species are small species with complete elytra that bear relatively long bristles.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B09B74627F7FD15146C33DA.taxon	biology_ecology	Biology. Shairella species appear to be univoltine based on field observations (unpublished data TCRT). Larvae are diurnal and found on the host plant’s leaves during late February. They are easily discovered when feeding on the tops of leaves during the day (Fig. 1 E). Larval development takes about one month, based on laboratory rearing. Mature larvae leave the host plant and burrow into the soil where they build underground chambers for pupation. Pupal stage (Fig. 1 F) duration is about 20 days, and adults begin to emerge after early April. Adults are nocturnal and have been observed in the field from May to August.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B09B74627F7FD15146C33DA.taxon	distribution	Distribution. Endemic to Taiwan.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B08B74027F7FE6716ED3479.taxon	materials_examined	Type locality. Chiayi County, Chushan (ṞƜ), 23 ° 30 ’ 44 ” N, 120 ° 49 ’ 22 ” E, 2400 m. Types. Shairella aeneipennis. Holotype ♂ (TARI): “ Iwai-yama (= Chushan, ṞƜ) / Mt. Arisan (ȐHƜ) / Tainan-syu / FORMOSA / 6. viii. 1940 / Col. K. ENDO [p, w] // Shairella / aeneipennis / CHÛJÔ [h] / DET. M. CHUJO [p, w] // HoLo / Type [p, w] (circle label, red letters but faded) // 2315 [p, w] ”.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B08B74027F7FE6716ED3479.taxon	description	Khasia itorum. Holotype ♀ (OMNH): “ Mt. ALI (ȐHƜ) / TAIWAN / 17. V. 1981 / T. ITO [p, y] // Khasia / itoi / Kimoto, n. sp. [h, w] // HOLOTYPE [p, r] ”. Paratypes: 2 ♂ (KMNH, OMNH): “ Mt. ALI / TAIWAN / 17. V. 1981 / T. ITO [p, y] // Khasia / itoi / Kimoto, n. sp. [h, w] // PARATOPOTYPE [p, b] // PHOTO [p, r] ”; 1 ♂ (KMNH): “ Mt. ALI / TAIWAN / 18. V. 1981 / N. ITO [p, y] // Khasia / itoi / Kimoto, n. sp. [h, w] // PARATYPE [p, b] // (blank) [w] ”.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B08B74027F7FE6716ED3479.taxon	materials_examined	Additional specimens examined (n = 34). Chiayi: 6 ♂, 3 ♀ (TARI, RBCN), Alishan (ȐHƜ), 17. V. 2010, leg. M. - H. Tsou; 1 ♂, 1 ♀ (TARI), Erhwanping (ΞΑ ‼, near Alishan), 9. V. 2011, leg. T. - H. Lee & M. - H. Tsou; Nantou: 1 ♂ (TARI), Hsitou (Ā Ḇ), 14. VI. 2011, leg. T. - H. Lee; 1 ♂ (TARI), 24 - 31. VII. 2008, leg. C. - S. Tung; Tatachia (Ṙ Ṙffi), 1 ♂, 5 ♀ (TARI), same locality, 8. VI. 2009, leg. C. - F. Lee; 5 ♂, 4 ♀ (TARI), same locality, 20. VII. 2009, leg. C. - F. Lee, H. Lee & S. - F. Yu; 1 ♀ (TARI), same locality, 21. IX. 2009, leg. C. - F. Lee; 4 ♂, 1 ♀ (TARI), same locality, 16. V. 2010, leg. M. - H. Tsou; 1 ♂ (TARI), same locality, 9. VII. 2014, leg. C. - F. Lee. Males. Length 4.7 – 5.2 mm; width 2.9 – 3.1 mm. Antenna (Fig. 2 A) long, filiform, about 1.2 x longer than body, length ratios of antennomeres I – XI 1.0: 0.3: 0.7: 1.0: 0.9: 1.0: 1.0: 1.0: 1.0: 0.9: 0.9, length to width ratios of antennomeres I – XI 3.1: 1.5: 2.7: 3.4: 3.4: 4.2: 4.4: 4.2: 4.5: 4.3: 4.7. Elytra strongly widened apically; about 1.1 x longer than wide; surface with sparse, fine punctures. Apical margin of last abdominal ventrite with two well developed incisions. Penis (Figs 2 C – 2 D) elongate, about 5.8 x longer than wide, parallel-sided, apex narrowly rounded, strongly curved in lateral view; endophallic sclerite short, about 0.5 x as long as penis, slender and longitudinal, apex rounded, with dense, short hairs along lateral margin at apical 1 / 4, bifurcate at basal 1 / 4.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B08B74027F7FE6716ED3479.taxon	description	Females. Length 5.0 – 5.2 mm, width 3.0 – 3.1 mm. Antenna (Fig. 2 B) long, filiform, about 1.1 x longer than body, length ratios of antennomeres I – XI 1.0: 0.3: 0.7: 0.9: 0.8: 0.8: 0.8: 0.8: 0.9: 0.8: 0.9, length to width ratios of antennomeres I – XI 3.6: 1.4: 3.1: 4.1: 3.9: 4.2: 3.9: 4.3: 5.0: 4.7: 5.8. Elytra strongly widened apically; about 1.1 x longer than wide; surface with sparse, fine punctures. Apical margin of last abdominal ventrite truncate. Gonocoxae (Fig. 2 E) longitudinal and wide, 3.1 x longer than wide, conjoined from base to middle, slightly narrowed at apical 1 / 3, apices rounded, each gonocoxa with eight or nine elongate setae. Ventrite VIII (Fig. 2 F) with only apical area and spiculum sclerotized; with dense, elongate setae inside apical sclerotized area, apical margin rounded; spiculum elongate, wide. Receptacle of spermatheca (Fig. 2 G) slender, as wide as pump, hardly separated from pump; pump wide, strongly curved, apex broadly rounded, apex with transverse and short appendix, size variable (Fig. 2 H); proximal spermathecal duct hardly separated from receptacle, apically narrowed.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B08B74027F7FE6716ED3479.taxon	diagnosis	Differential diagnosis. Shairella aeneipennis is similar to S. guoi sp. nov. and S. motienensis sp. nov. by possessing fine punctures on the elytra; but this species differs others by the shorter endophallic sclerite of the penis (Figs 2 C – 2 D) (longer endophallic sclerite in S. motiensis (Figs 9 C – 9 D )), rounded apex of the endophallic sclerite (depressed apex of the endophallic slcerite in S. guoi sp. nov. (Figs 8 C – 8 D) and pointed apex in S. motiensis sp. nov.), and the narrowed apex of the penis (tapering apex from apical 1 / 6 in S. guoi sp. nov. and lanceolate apex in S. motienensis sp. nov.).	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B08B74027F7FE6716ED3479.taxon	discussion	Remarks. The allotype of S. aenepennis is prooved to belong to another species (see S. tsoui sp. nov.). Host plant. Clinopodium laxiflorum var. taiwanianum Hsieh & Huang (Labiatae). Distribution. Chiayi county: Alishan (ȐHƜ); Nantou county: Hsitou (Ā Ḇ) and Tatachia (ṘṘffi) (Fig. 3 A).	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B0EB74227F7F9C2154631F6.taxon	materials_examined	Type locality. Taiwan: Taitung County, Lichia trail (ffl ḃŔả), 22 ° 48 ’ 09 ” N, 120 ° 59 ’ 15 ” E, 1100 m. Types (n = 12). Holotype ♂ (TARI), Taitung: Lichia trail (ffl ḃŔả), 15. VII. 2014, leg. B. - X. Guo. Paratypes: 2 ♂ (TARI), same data as holotype; 1 ♀ (TARI), same locality, 25. VII. 2015, leg. Y. - T. Chung, P. - H. Kuo & S. - P. Wu; 7 ♂, 1 ♀ (TARI, RBCN), same locality, 1. VII. 2016, leg. B. - X. Guo.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B0EB74227F7F9C2154631F6.taxon	description	Males. Length 5.7 – 5.9 mm; width 3.6 – 3.7 mm. Antenna (Fig. 4 A) long, filiform, about 1.1 x longer than body, length ratios of antennomeres I – XI 1.0: 0.3: 0.7: 0.9: 0.8: 0.9: 0.9: 0.8: 0.8: 0.8: 0.8, length to width ratios of antennomeres I – XI 3.4: 1.6: 3.1: 4.2: 4.1: 4.6: 4.6: 5.1: 5.5: 5.0: 6.0. Elytra apically and weakly widened; about 1.1 x longer than wide; surface with sparse and coarse punctures, and with several indistinct longitudinal ridges. Apical margin of last abdominal ventrite with two well developed incisions. Penis (Figs 4 C – 4 D) elongate, about 5.6 x longer than wide, gradually narrowed towards basal 1 / 5, apically tapering from apical 1 / 9, strongly curved in lateral view; endophallic sclerite long, about 0.7 x as long as penis, slender and longitudinal, apical margin rounded, with dense short hairs along lateral margin at middle, bifurcate at basal 1 / 6. Females. Length 6.1 – 6.2 mm, width 4.1 – 4.2 mm. Antenna (Fig. 4 B) extremely long, filiform, as long as body, length ratios of antennomeres I – XI 1.0: 0.4: 0.7: 0.9: 0.8: 0.9: 0.8: 0.8: 0.8: 0.8: 0.9, length to width ratios of antennomeres I – XI 3.7: 1.7: 3.1: 4.3: 4.5: 4.8: 4.5: 5.1: 5.4: 5.4: 7.0. Elytra strongly widened apically; as long as wide; surface with sparse, fine punctures and with several indistinct longitudinal ridges. Apical margin of last abdominal ventrite subtruncate. Gonocoxae (Fig. 4 E) longitudinal and narrow, 2.9 X longer than wide, conjoined from base to middle, slightly narrowed at apical 1 / 3, apices rounded, each gonocoxa with eight or nine elongate setae. Ventrite VIII (Fig. 4 F) with only apical area and spiculum sclerotized; with dense elongated inside apical sclerotized area, apical margin rounded; spiculum elongate and wide. Receptacle of spermatheca (Fig. 4 G) slender, as wide as pump, hardly separated from pump; pump wide and strongly curved, apex broadly rounded, apex with transverse, broad sclerite (Fig. 4 H), size variable; proximal spermathecal duct hardly separated from receptacle, apically narrowed.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B0EB74227F7F9C2154631F6.taxon	diagnosis	Differential diagnosis. Shairella cheni sp. nov. is similar to S. chungi sp. nov. based on the large body size and coarse punctures on the elytra. This species differs from S. chungi sp. nov. by the tapering of the penis from the apical 1 / 4 to apex (Figs 4 C – 4 D) (lanceolate apex in S. chungi sp. nov. (Figs 6 C – 6 D )).	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B0EB74227F7F9C2154631F6.taxon	biology_ecology	Host plant. Strobilanthes flexicaulis Hayata (Acanthaceae).	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B0EB74227F7F9C2154631F6.taxon	etymology	Etymology. This new species is dedicated to Mr. Chang Chin Chen for supporting the TCRT in various ways.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B0EB74227F7F9C2154631F6.taxon	distribution	Distribution. Only known from the type locality (Fig. 3 A).	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B0CB74D27F7FC5015463312.taxon	materials_examined	Type locality. Pingtung County, Tahanshan (XÄƜ), 22 ° 24 ’ 27 ” N, 120 ° 45 ’ 23 ” E, 1400 m. Types (n = 34). Holotype ♂ (TARI), Pingtung: Tahanshan (XÄƜ), 29. VI. 2013, leg. B. - X. Guo. Paratypes: 6 ♂, 3 ♀ (TARI), same data as holotype; 5 ♂, 3 ♀ (TARI, RBCN), same locality, 11. VII. 2013, leg. B. - X. Guo; 2 ♂ (TARI), same locality, 12. VII. 2013, leg. Y. - T. Chung; 1 ♀ (TARI), same locality, 21. VII. 2013, leg. Y. - T. Chung; 1 ♀ (TARI), same locality, 9. VIII. 2013, leg. B. - X. Guo; 1 ♂ (TARI), same locality, 28. VIII. 2014, leg. Y. - T. Chung; 3 ♂, 1 ♀ (TARI), same locality, 19. V. 2015, leg. Y. - T. Chung; 3 ♂, 1 ♀ (TARI), same locality, 27. V. 2015, leg. Y. - T. Chung; 1 ♂, 3 ♀ (TARI), same locality, 6. VI. 2015, leg. Y. - T. Chung.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B0CB74D27F7FC5015463312.taxon	description	Males (Figs 5 A – 5 C). Length 6.2 – 6.7 mm; width 3.7 – 3.9 mm. Antenna (Fig. 6 A) long, filiform, about 1.1 x longer than body, length ratios of antennomeres I – XI 1.0: 0.3: 0.7: 1.0: 0.9: 0.8: 0.8: 0.8: 0.8: 0.8: 0.9, length to width ratios of antennomeres I – XI 3.3: 1.8: 3.1: 4.6: 4.4: 4.4: 4.5: 5.5: 6.1: 5.4: 7.0. Elytra apically weakly broadened; about 1.2 - 1.3 x longer than wide; surface with sparse, coarse punctures and several indistinct longitudinal ridges. Apical margin of last abdominal ventrite with two well developed incisions. Penis (Figs 6 C – 6 D) elongate, about 5.1 x longer than wide, gradually narrowed towards middle, apex lanceolate, strongly curved in lateral view; endophallic sclerite long, about 0.8 x as long as penis, slender and longitudinal, apical margin rounded, with dense short hairs along lateral margin at middle, bifurcate at basal 1 / 6. Females (Figs 5 D – 5 F). Length 6.9 – 7.3 mm, width 4.5 – 4.7 mm. Antenna (Fig. 6 B) long, filiform, as long as body, length ratios of antennomeres I – XI 1.0: 0.3: 0.7: 0.9: 0.9: 0.8: 0.8: 0.8: 0.8: 0.8: 0.8, length to width ratios of antennomeres I – XI 4.0: 1.5: 3.1: 4.6: 4.9: 5.5: 5.3: 5.6: 6.5: 5.8: 7.1. Elytra strongly broadened apically; about 1.0 – 1.1 x longer than wide; surface with sparse, fine punctures and several indistinct longitudinal ridges. Apical margin of last abdominal ventrite subtruncate. Gonocoxae (Fig. 6 E) longitudinal and narrow, 3.2 x longer than wide, conjoined from base to middle, slightly narrowed in apical 1 / 3, apices rounded, each gonocoxa with eight elongate setae. Ventrite VIII (Fig. 6 F) with only apical area and spiculum sclerotized; with dense elongate setae inside apical sclerotized area, apical margin rounded; spiculum elongate and broad. Receptacle of spermatheca (Fig. 6 G) slender, as wide as pump, hardly separated from pump; pump broad and strongly curved, apex broadly rounded, apex with transverse, broad sclerite (Fig. 6 H), size variable; proximal spermathecal duct hardly separated from receptacle, apically narrowed.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B0CB74D27F7FC5015463312.taxon	diagnosis	Differential diagnosis. See diagnosis of Shairella cheni sp. nov.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B0CB74D27F7FC5015463312.taxon	biology_ecology	Host plants. Strobilanthes flexicaulis Hayata (Acanthaceae) and Hemiboea bicornuta (Hayata) Ohwi (Gesneriaceae).	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B0CB74D27F7FC5015463312.taxon	etymology	Etymology. The new species is dedicated to Mr. Yi-Ting Chung, who collected the type specimens.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B0CB74D27F7FC5015463312.taxon	distribution	Distribution. Only known from the type locality (Fig. 3 A).	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B01B74927F7FF2B132A319E.taxon	materials_examined	Type locality. Taiwan: Kaoshiung County, Tienchih (Ŧè), 23 ° 16 ’ 37 ” N, 120 ° 55 ’ 01 ” E, 2200 m. Types (n = 27). Holotype ♂ (TARI), Kaohsiung: Tienchih (Ŧè), 17. V. 2015, leg. B. - X. Guo. Paratypes: 3 ♂, 2 ♀ (TARI), same data as holotype; 10 ♂, 9 ♀ (TARI, RBCN), Kueiku (Ñê), 11. VI. 2015, leg. C. - F. Lee; 2 ♀ (TARI), Kuanshanyakou (DƜ ḄŊ), 30. VII. 2015, leg. C. - F. Lee.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B01B74927F7FF2B132A319E.taxon	description	Males (Figs 7 A – 7 C). Length 4.3 – 4.6 mm; width 2.5 – 2.7 mm. Antenna (Fig. 8 A) long, filiform, about 1.2 x longer than body, length ratios of antennomeres I – XI 1.0: 0.4: 0.8: 0.9: 0.8: 0.8: 0.8: 0.8: 0.8: 0.7: 0.8, length to width ratios of antennomeres I – XI 3.2: 1.9: 3.6: 4.2: 3.6: 4.1: 4.0: 4.1: 4.5: 4.1: 4.8. Elytra strongly broadened apically; about 1.1 x longer than wide; surface with sparse, fine punctures. Apical margin of last abdominal ventrite with two well developed incisions. Penis (Figs 8 C – 8 D) elongate, about 6.9 x longer than wide, parallel-sided, apically tapering from apical 1 / 6, strongly curved in lateral view; endophallic sclerite short, about 0.5 x as long as penis, slender and longitudinal, apical margin medially depressed, with dense short hairs along lateral margin at apical 1 / 3, bifurcate at basal 1 / 4. Females (Figs 7 D – 7 F). Length 5.0 – 5.3 mm, width 3.2 – 3.3 mm. Antenna extremely long, filiform, as long as body, length ratios of antennomeres I – XI 1.0: 0.3: 0.7: 0.7: 0.7: 0.7: 0.8: 0.7: 0.7: 0.7: 0.7, length to width ratios of antennomeres I – XI 3.5: 1.5: 2.9: 3.3: 4.1: 4.0: 4.4: 3.9: 4.1: 3.9: 4.3. Elytra strongly widened apically; about 1.0 – 1.1 x longer than wide; surface with sparse, fine punctures. Apical margin of last abdominal ventrite truncate. Gonocoxae (Fig. 8 E) longitudinal and narrow, 3.8 x longer than wide, conjoined from base to middle, slightly narrowed at apical 1 / 3, apices rounded, each gonocoxa with eight elongate setae. Ventrite VIII (Fig. 8 F) with only apical area and spiculum sclerotized; with dense elongate setae inside apical sclerotized area, apical margin rounded; spiculum elongate and broad. Receptacle of spermatheca (Fig. 8 G) slender, as wide as pump, hardly separated from pump; pump broad and strongly curved, apex broadly rounded, apex with transverse, short sclerite, size variable (Figs 8 H – 8 I); proximal spermathecal duct hardly separated from receptacle, apically narrowed.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B01B74927F7FF2B132A319E.taxon	diagnosis	Differential diagnosis. See diagnosis of Shairella aeneipennis Chûjô.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B01B74927F7FF2B132A319E.taxon	biology_ecology	Host plant. Clinopodium laxiflorum var. taiwanianum Hsieh & Huang (Labiatae).	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B01B74927F7FF2B132A319E.taxon	etymology	Etymology. The new species is dedicated to Mr. Bo-Xin Guo, who collected the type specimens.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B01B74927F7FF2B132A319E.taxon	distribution	Distribution. Vicinity along western half of South Cross-Island Highway (Ħ ḂḦR) (Fig. 3 A).	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B07B74B27F7FBA3132232AE.taxon	materials_examined	Type locality. Taiwan: Taitung County, Motien (= Ŧ), 23 ° 11 ’ 35 ” N, 121 ° 01 ’ 24 ” E, 1500 m. Types (n = 54). Holotype ♂ (TARI), Taitung: Motien (= Ŧ), 20. VI. 2011, leg. C. - F. Lee. Paratypes: 24 ♂, 8 ♀ (TARI, RBCN), same data as holotype; 3 ♂ (TARI), Liyuan (M ḋ), 23. VI. 2010, leg. M. - H. Tsou; 1 ♂ (TARI), same locality, 3. X. 2010, leg. M. - H. Tsou; 2 ♂, 3 ♀ (TARI), same locality, 19. VI. 2013, leg. C. - F. Lee; 8 ♂, 4 ♀ (TARI), same locality, 24. VII. 2013, leg. C. - F. Lee.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B07B74B27F7FBA3132232AE.taxon	description	Males. Length 4.6 – 5.1 mm; width 2.7 – 3.1 mm. Antenna (Fig. 9 A) long, filiform, about 1.3 x longer than body, length ratios of antennomeres I – XI 1.0: 0.4: 0.8: 0.9: 0.9: 0.9: 0.9: 0.8: 0.9: 0.8: 0.9, length to width ratios of antennomeres I – XI 3.3: 1.8: 3.2: 3.7: 4.0: 4.2: 4.2: 4.5: 5.0: 4.8: 5.2. Elytra apically, moderately broadened; about 1.2 x longer than wide; surface with sparse, fine punctures. Apical margin of last abdominal ventrite with two well developed incisions. Penis (Figs 9 C – 9 D) elongate, about 5.3 x longer than wide, parallel-sided, apex lanceolate, moderately curved in lateral view; endophallic sclerite long, about 0.7 x as long as penis, slender and longitudinal, apically tapering, with dense short hairs along lateral margin in apical 1 / 4, bifurcate at basal 1 / 4. Females. Length 5.4 – 6.4 mm, width 3.5 – 3.9 mm. Antenna (Fig. 9 B) long, filiform, 1.2 x longer than body, length ratios of antennomeres I – XI 1.0: 0.4: 0.7: 0.9: 0.9: 0.8: 0.9: 0.8: 0.8: 0.7: 0.9, length to width ratios of antennomeres I – XI 3.6: 1.2: 2.9: 4.1: 4.4: 4.7: 4.8: 5.8: 5.7: 5.2: 6.1. Elytra strongly broadened apically; about 1.1 x longer than wide; surface with sparse, fine punctures. Apical margin of last abdominal ventrite slightly convex at middle (Fig. 9 G). Gonocoxae (Fig. 9 E) longitudinal and broad, 3.0 x longer than wide, conjoined from base to middle, slightly narrowed at apical 1 / 3, apices rounded, each gonocoxa with seven or eight elongate setae. Ventrite VIII (Fig. 9 F) with only apical area and spiculum sclerotized; with dense elongate setae inside apical sclerotized area, apical margin rounded; spiculum elongate and broad. Receptacle of spermatheca (Fig. 9 K) slender, as wide as pump, hardly separated from pump; pump broad, strongly curved, apex broadly rounded, apex with transverse, short sclerite, size extremely variable (Figs 9 H – 9 J); proximal spermathecal duct hardly separated from receptacle, apically narrowed.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B07B74B27F7FBA3132232AE.taxon	diagnosis	Differential diagnosis. See diagnosis of Shairella aeneipennis Chûjô.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B07B74B27F7FBA3132232AE.taxon	biology_ecology	Host plant. Clinopodium laxiflorum var. taiwanianum Hsieh & Huang (Labiatae). Etymology. This new species is named after the type locality.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B07B74B27F7FBA3132232AE.taxon	distribution	Distribution. Vicinity along eastern half of South Cross-Island Highway (Ħ ḂḦR) (Fig. 3 A).	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B05B74B27F7FEF31327356A.taxon	materials_examined	Type locality. Taiwan: Hualien County, Kuanyuan (Dë), 24 ° 11 ’ 03 ” N, 121 ° 20 ’ 32 ” E, 2300 m. Types (n = 79). Holotype ♂ (TARI), Hualien: Kuanyuan (Dë), 2. VI. 2016, leg. C. - F. Lee. Paratypes: 11 ♂, 15 ♀ (TARI, RBCN), same data as holotype; 5 ♂, 6 ♀ (TARI), same locality, 2. VI. 2016, leg. Y. - T. Chung; 10 ♂, 10 ♀, same locality, 2. VI. 2016, leg. B. - X. Guo; Taichung: 3 ♂, 2 ♀ (TARI), Pilu (Bõ), 8. VIII. 2014, leg. S. - F. Yu; 5 ♀ (TARI), same locality, 8. VIII. 2014, leg. M. - H. Tsou; 2 ♂, 5 ♀ (TARI), same locality, 7. VII. 2015, leg. C. - F. Lee; 4 ♂ (TARI), same locality, 3. VI. 2016, leg. B. - X. Guo; 1 ♀ (TARI), labeled “ TAIWAN / HASSENZAN (= Pahsienshan, Ã ḾƜ) [p] / 4. VI. 1942 [h] / A. MUTUURA [p, w] // ữaeȁ (Chiapaotai) — Ë Ṛ (Liming) [h, on the back of the same label] // Shairella / aeneipennis / CHÛJÔ [h] / DET. M. CHUJO [p, w] // ALLo / Type [p, w, circle label with gray letters] ”.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B05B74B27F7FEF31327356A.taxon	description	Males (Figs 10 A – 10 C). Length 4.5 – 5.0 mm; width 2.6 – 2.7 mm. Antenna (Fig. 11 A) long, filiform, about 1.3 x longer than body, length ratios of antennomeres I – XI 1.0: 0.4: 0.8: 1.0: 0.9: 0.9: 1.0: 0.8: 0.8: 0.8: 0.9, length to width ratios of antennomeres I – XI 2.8: 1.6: 2.8: 3.6: 3.3: 3.8: 3.9: 3.3: 3.9: 4.0: 4.4. Elytra apically, moderately broadened; about 1.1 – 1.3 x longer than wide; surface with sparse, coarse punctures and with several longitudinal indistinct ridges. Apical margin of last abdominal ventrite with two well developed incisions. Penis (Figs 11 C – 11 D) extremely slender, about 8.0 x longer than wide, apically tapering from basal 1 / 3, moderately curved at basal 1 / 3 in lateral view; endophallic sclerite short, about 0.4 x as long as penis, slender and longitudinal, apex pointed, with dense, short hairs along lateral margin in apical 1 / 4, bifurcate at basal 1 / 4. Females (Figs 10 D – 10 F). Length 5.2 – 5.4 mm, width 3.5 – 3.6 mm. Antenna (Fig. 11 B) long, filiform, 1.2 x longer than body, length ratios of antennomeres I – XI 1.0: 0.4: 0.7: 0.9: 0.8: 0.8: 0.8: 0.8: 0.7: 0.7: 0.8, length to width ratios of antennomeres I – XI 3.2: 1.5: 2.6: 3.8: 3.7: 3.8: 3.8: 3.9: 3.7: 3.7: 5.1. Elytra strongly broadened apically; about 1.0 – 1.1 x longer than wide; surface with sparse and coarse punctures. Apical margin of last abdominal ventrite with median notch (Fig. 11 G). Gonocoxae (Fig. 11 E) longitudinal, broad, 2.9 X longer than wide, conjoined from base to middle, slightly narrowed at apical 1 / 3, apices rounded, each gonocoxa with seven or nine elongate setae. Ventrite VIII (Fig. 11 F) without sclerotized areas but spiculum sclerotized; with dense elongate setae along apical margin, apical margin irregular; spiculum elongate and broad. Receptacle of spermatheca (Fig. 11 H) slender, as wide as pump, hardly separated from pump; pump broad, strongly curved, apex broadly rounded, apex with transverse, tiny sclerite (Fig. 11 I); proximal spermathecal duct hardly separated from receptacle, apically narrowed.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B05B74B27F7FEF31327356A.taxon	discussion	Remarks. The allotype of Shairella aeneipennis Chûjô is misidentified and belongs to S. tsoui sp. nov .. It is a member to this species.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B05B74B27F7FEF31327356A.taxon	diagnosis	Differential diagnosis. Shairella tsoui sp. nov. appears similar to S. chungi sp. nov. and S. cheni sp. nov. based on the coarse punctures of the elytra; but this species can be separated from them with their smaller sizes, the slender and apically tapering penis, the long endophallic sclerite, and presence of the median notch on the apical margin of the last abdominal ventrite in females.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B05B74B27F7FEF31327356A.taxon	biology_ecology	Host plant. Clinopodium laxiflorum var. taiwanianum Hsieh & Huang (Labiatae).	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B05B74B27F7FEF31327356A.taxon	etymology	Etymology. The new species is dedicated to Mr. Mei-Hua Tsou, who collected the type specimens.	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
207887EF2B05B74B27F7FEF31327356A.taxon	distribution	Distribution. Kuanyuan (Dë), Hualien County and Pilu (Bõ), Taichung County (Fig. 3 A).	en	Lee, Chi-Feng, Beenen, Ron (2017): Revision of the genus Shairella Chûjô, 1962 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species. Zootaxa 4268 (4): 489-507, DOI: 10.11646/zootaxa.4268.4.2
