identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
203D141B5228FFC7558852380529F87D.text	203D141B5228FFC7558852380529F87D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ambrysus bifidus La Rivers & Nieser	<div><p>Ambrysus bifidus La Rivers &amp; Nieser</p><p>(Figs. 1–2)</p><p>Ambrysus bifidus La Rivers &amp; Nieser 1972: Biol. Soc. Nevada Occas. Pap. 4–8.</p><p>Ambrysus bifidus: Nieser 1975, Stud. Fauna Suriname Guyanas 56–57 (suppl. descript.).</p><p>Discussion. This species is known from Suriname and northern Brazil (La Rivers &amp; Nieser 1972). Reported here is the first record from Guyana. This species was distinguished from A. scolius by a "slight bifurcation" of the apical median lobe of the female subgenital plate (Fig. 2 F) (La Rivers &amp; Nieser 1972), and from A. tricuspis by the same bifurcation of the subgenital plate and also the male accessory genitalic process less massive or elongated laterally (Fig. 2 B) (La Rivers 1974). Nieser (1975) provided a supplemental description in his treatment of the water bugs of the Guyana region, in which he documented the variation in the shape of the accessory genitalic process of tergum VI. A. bifidus can be distinguished from the other species in this complex by the bifurcated median lobe of the female subgenital plate.</p><p>The habitat of the type series was given in the original description as "floating plant-debris upstream of a barrier of tree branches, etc., velocity of current 5 m /min." In Suriname, A. bifidus, A. stali, and A. usingeri La Rivers were considered to be common in small streams in savannah woodlands with dark water low in electrolytes (La Rivers &amp; Nieser 1972). A. bifidus was collected with A. stali in Surinam mostly in barriers of branches and dead leaves or at the bases of trees standing in water where the current is strong.</p><p>Type locality. SURINAME, Zanderijsavanne. Holotype is housed at CAS.</p><p>Material examined. GUYANA: Pirara Ranch, 3º 22.1' N, 59º 40.5' W, Pirara River, 25 Apr 1995, colln #19, PJ Spangler &amp; SA Perry (1♀, UMC). SURINAME: Zanderijsavanna Carolina Cr. / (reverse side of locality label) 1969 Aug 18, N. Nieser / Ambrysus bifidus L&amp;N 1972 Paratype / Ira La Rivers Collection Bequeathed to the California Academy of Sciences (1♂, 1♀, CAS).</p><p>Published records. (La Rivers &amp; Nieser 1972): BRAZIL, Amazonas, Pará; SURINAM, Zanderijsavanne. (Pereira &amp; Melo 2007): BRAZIL, Amazonas, Waimiri-Atroari Biological Reserve.</p></div>	https://treatment.plazi.org/id/203D141B5228FFC7558852380529F87D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sites, Robert W.;Reynoso-Velasco, Daniel	Sites, Robert W., Reynoso-Velasco, Daniel (2015): Review of the Ambrysus stali La Rivers species complex (Heteroptera: Nepomorpha: Naucoridae) with the description of a new species from Mesoamerica. Zootaxa 4018 (2): 279-291, DOI: 10.11646/zootaxa.4018.2.7
203D141B522AFFC2558853A202ADFE2C.text	203D141B522AFFC2558853A202ADFE2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ambrysus maya Sites and Reynoso	<div><p>Ambrysus maya Sites and Reynoso, NEW SPECIES</p><p>(Figs. 1, 3–4)</p><p>Description. Macropterous male. HOLOTYPE, length 7.84; maximum width 4.40. Paratypes (n = 2), length 7.84–7.92 (mean = 7.88); maximum width 4.32–4.52 (mean = 4.42). General shape fusiform; widest across abdominal segment III (Fig. 3 A). Overall dorsal coloration anteriorly pale yellow with dark brown pattern on forelegs, head, and pronotum (Fig. 3 A); anterior pale yellow has greenish tone in live specimens (Fig. 4 B); posteriorly with dark brown scutellum and hemelytra. Exposed connexiva with alternating creamy white and dark checkered appearance. Dorsal surface coarsely punctate. Ventrally, mostly brown, lighter centrally.</p><p>Head. Head length 1.36; maximum width 1.98. Eyes convergent, synthlipsis 0.72; thin band of cuticle posterior to eye; eyes flat and not raised above level of vertex, slightly raised above level of pronotum. Anterior margin between eyes convex, extending anteriorly in front of eyes 7% of head length; posterior margin between eyes strongly convex, extending posteriorly 30% of head length. Labrum width 1.8 x length, evenly rounded. Labium brown, with three visible segments, extending 0.48 beyond labrum. Antennal segment proportions 2:9:11:6, length 0.54, elongate hairs on segment 4 and distal 2/3 of 3.</p><p>Thorax. Pronotum mostly pale yellowish brown with dark brown punctures, anteromedial brown maculation, irregular longitudinal brown stripes, some longitudinal stripes enhanced with dark punctures; slightly rugose; lateral margins convex; sparse erect setae; explanate, strongly convergent anteriorly; posterior margin nearly straight; anterior margin deeply concave receiving convex posterior margin of head; transverse line of punctures defining anterior margin of band in posterior 1/4; posterolateral corners obtuse, rounded; width 3.2 x length; length at midline 1.22; maximum width at posterolateral corners 3.92. Prothorax ventrally brown and pruinose medially, with yellow glabrous lateral band; apices of propleura meeting broadly at midline, separated from level of prosternellum. Probasisternum brown, with sharp medial carina and generally appearing pruinose, row of elongate pale hairs lateral to carina. Prosternellum light brown, extending beneath apices of propleura. Elongate setae along posterior margin of propleuron. Scutellum mostly brown with yellow anterior corners and apex, triangular, densely punctate, width 1.9 x length, width 2.60, length 1.36. Hemelytra densely punctate. Claval commissure length 0.72. Embolium length 2.30, greatest width 0.62; lateral margin slightly convex in proximal half, more pronounced in distal half. Hind wings well developed. Mesobasisternum with midventral longitudinal tumescence with sulcus on midline; tumescence terminating posteriorly as acute mesosternellum. Metasternellum (=metaxyphus) transverse, subtriangular, with apex acute.</p><p>Legs. All leg segments pale yellowish brown, darker distally on fore and hind legs. Profemur pale yellowish brown with dark brown along anterior margin continuous to apex; dark brown posterior marking extending anteriorly, becoming discontinuous where it approaches or meets anterior margin dark marking; row of 10–12 combs of short brown spines in basal half of posterior margin; anterior margin with dense pad of setae without associated spines. Protibia and tarsus with inner surface flattened and with spatulate setae; tarsus immovable, onesegmented; pretarsal claw single, minute, triangular. Procoxa with dark brown basal marking on lateral to anterior surfaces, cluster of stout brown anteromedial spines. Mesocoxa partially recessed into thorax. Metacoxa with longitudinal sulcus that can accommodate flexed metafemur. Meso- and metafemora with row of short, brown spines on anterior margin; spines restricted to basal 1/3 and single on mesofemur, combs of 1–4 spines irregularly spaced along full length of metafemur. Mesotibia with ventrolateral, ventromedial, dorsolateral, and dorsomedial rows of stout reddish-brown spines, dorsolateral and dorsomedial rows include combs of up to 4 spines; stout spines at apical rim dorsally, two transverse rows of stout spines at apex of ventral margin. Metatibia with ventrolateral, ventromedial, dorsolateral, and dorsomedial rows of stout reddish-brown spines, transverse rows of stout spines at apex of ventral margin. Meso- and metatibiae and -tarsi with long, pale swimming hairs; hairs sparse on mesotibia and -tarsus, profuse on metatibia and -tarsus. Meso- and metapretarsi with paired claws slender, curved in distal half, with basal tooth. Leg measurements as follows: foreleg, femur 1.86, tibia 1.60, tarsus 0.32; middle leg, femur 1.88, tibia 1.64, tarsomeres 1–3, 0.10, 0.28, 0.36; hind leg, femur 2.32, tibia 2.72, tarsomeres 1– 3, 0.16, 0.52, 0.47.</p><p>Abdomen. Abdomen dorsally with connexiva of III–VII exposed and with checkered appearance (Fig. 4 B), posterior half of III–VII dark brown, anterior half of III–V creamy white and of VI–VII brown; VIII mostly brown; lateral margin with row of pale setae, group of trichobothria on dorsal surface near posterolateral corners. Posterolateral corner of III slightly acuminate, IV–VII spinose. Accessory genitalic process of VI broad, apical margin flattened with corners broadly rounded, broadly prolonged laterally (Fig. 3 B). Medial lobes of VIII (pseudoparameres) with apical corners rounded, distal margin flat (Fig. 3 D). Ventrally mostly brown laterally, brownish yellow medially, pruinose, with pile of fine hairs. Laterosternites III–VI with yellow near middle of lateral margin. Lateral margin with thin glabrous band. Glabrous patches on laterosternites II–VII. Mediosternite V with posterior margin slightly asymmetrical. Phallosoma with left ventral lobe reflexed dorsad, unarmed with teeth; right ventral lobe appressed to body of phallosoma, unarmed (Fig. 3 C). Parameres subquadrate, pedunculate, symmetrical, mesal margin slightly convex with corners broadly rounded (Fig. 3 E). Pygophore with anterior margin between parameres slightly produced at midline, posterior apex with fine setae.</p><p>Macropterous female. Paratypes (n = 11), length 7.68–8.96 (mean = 8.45); maximum width 4.36–4.72 (mean = 4.55). Similar to male in general structure and coloration except as follows: Mediosternite V symmetrical. Mediosternite VII (subgenital plate) with medial convexity and distolateral production (Fig. 3 F). Laterosternite VI with posteromedial corner on left side produced into well-developed spine directed posteroventrad, posteromedial corner on right side produced, tuberculate, deflexed ventrad.</p><p>Diagnosis and comparative notes. This species can be distinguished from other members of the species complex by the significantly narrower fore femora (t=3.803, df=15, p=0.002) and its noncoincident geographic range. It can be distinguished from all other naucorids in Mesoamerica by its small size, fusiform body shape, and dark longitudinal markings of the pronotum and dark markings on the dorsal surface of the profemora.</p><p>Habitat description. At the type locality, specimens were collected from marginal vegetation including dead leaves, grasses, and other vegetation (Fig. 4 A). The river was shallow (ca. 1 m) with cobble substrate and swift current. In the riffles were the naucorids Ambrysus circumcinctus Montandon and Limnocoris insularis Champion, and the byrrhoid beetles Helichus suturalis LeConte (Dryopidae), Phanocerus clavicornis Sharp, Xenelmis bufo (Sharp), Heterelmis sp., Hexacylloepus sp., and Microcylloepus sp. ( Elmidae). Specimens of Ambrysus maya from the Sibun River in Belize were collected mostly from vegetational debris caught on branches or along the margins. Other insect species in the Sibun River included Argia sp. ( Coenagrionidae); Laccophilus oscillator laevipennis Sharp, L. proximus Say (Dytiscidae); Anopheles albimanus Wiedemann, and Chagasia bathana (Dyar) (Culicidae) (R. Vogtsberger, pers. com.).</p><p>Discussion. Although the Central American countries south of Guatemala have been well-collected and the fauna studied by many scientists over the past century beginning with Champion (1901), A. maya has not been reported from there. Thus, this species appears to be widely separated from A. tricuspis in Colombia, the nearest known member of the A. stali complex (Fig. 1). The majority of insects in Central America are suggested to have originated from taxonomic elements of northern South America (Kimsey 1992). Ambrysus maya follows this pattern since most of the diversity of this species complex is distributed throughout the Guyana Shield region with a single incursion into Mesoamerica. Alternatively, the ancestor of this complex might have moved southward from the Mexican geographic center of Ambrysus diversity, and subsequently became discontinuous with the current species through vicariance. Substantially more geological time for either scenario to have occurred than was previously considered was shown recently. Montes et al. (2015) provided evidence that the Central American land bridge formed in the Miocene epoch 10–12 million years earlier than the 3 million year Pliocene model that had been widely accepted. Moreover, a phylogenetic analysis of New World naucorid groups with attention to divergence times may be able to evaluate invasion patterns. The diversity of Ambrysus is greatest in North America and of Cryphocricos and Limnocoris in South America.</p><p>Etymology. The specific epithet is a noun in apposition in reference to the Maya civilization, which occupied this region of Mesoamerica until the 17th century.</p><p>Repositories. The holotype and some paratypes are deposited in the Enns Entomology Museum (University of Missouri); additional paratypes are deposited in Midwestern State University (Wichita Falls), Universidad del Valle de Guatemala (Guatemala City), Universidad Nacional Autónoma de México (Mexico City), and the United States National Museum of Natural History (Washington D.C.).</p><p>Type material examined. HOLOTYPE ♂: BELIZE: Orange Walk District: Río Bravo ca. 10 mi. N. of Gallon Jug at Cedar Crossing, 19 April 2014, 44 m, 17°41.415´N, 89°01.615´W, R. Sites &amp; W. Shepard, vegetated margins, L-1740; PARATYPES: same data as holotype (1♀); Belize District: Sibun River, 17º24.690'N, 088º25.288'W, 1 VII 2009, bamboo &amp; branches, coll J. H. Adams (1♀); same data except 17º24.281'N, 088º26.012'W (1♀); same data except 17º24.702'N, 088º25.297'W (1♂); same data except 17º16.112'N, 088º34.276'W, 29 VI 2009, submerged stones (1♀); Cayo District: Sibun River, 0338582N, 1916372W, detritus patch, 26–30 VI 2009, coll J. H. Adams (1♀); same data except 0333330N, 1912078W (1♀); same data except 0348708N, 1925658W (1♀); same data except 0338622N, 1916554W (1♀); same data except 033283N, 1911970W (1♀). GUATEMALA: Alta Verapaz: Lag[una]. Lachua, 19 VII 1995, J. C. Schuster (1♀). MEXICO: Chiapas: Chajal Arroyo Manzana, 9-XII-2008, G. Mendez / Colección del Instituto de Biología, UNAM México, D.F. (1♀); Tabasco: Río Carrizal, Est. No. 7, 10-VI-1996, 17º57.23'N, 93º07.17'W, J. Bueno, A. Contreras, R. Barba / Colección del Instituto de Biología, UNAM México, D.F. (1♂).</p></div>	https://treatment.plazi.org/id/203D141B522AFFC2558853A202ADFE2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sites, Robert W.;Reynoso-Velasco, Daniel	Sites, Robert W., Reynoso-Velasco, Daniel (2015): Review of the Ambrysus stali La Rivers species complex (Heteroptera: Nepomorpha: Naucoridae) with the description of a new species from Mesoamerica. Zootaxa 4018 (2): 279-291, DOI: 10.11646/zootaxa.4018.2.7
203D141B522EFFCD5588524102CBFB30.text	203D141B522EFFCD5588524102CBFB30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ambrysus scolius La Rivers	<div><p>Ambrysus scolius La Rivers</p><p>(Figs. 1, 5)</p><p>Ambrysus scolius La Rivers 1970: Great Basin Natur. 1–4.</p><p>Discussion. This species is known by only three females from the Guyana Shield countries of Guyana, Trinidad, and Venezuela. Males are not known. Although we have been unable to locate the holotype, the paratypes in the CAS (Fig. 5) that we examined do not resemble the habitus figure of the holotype in the original description. La Rivers (1970) gave the sigmoid posterior margin of laterosternite VI (La Rivers' segment V) as the diagnostic feature of this species. We have examined the two paratypes housed at the CAS and considered this feature in both anterior/posterior and dorsal/ventral planes, but are uncertain how to interpret the sigmoid nature of the margin and how it differs interspecifically. In addition, the left laterosternite VI has a well-developed posteromedial spine directed posteroventrad, whereas the right laterosternite is developed differently in the two paratypes: In one specimen the posterior margin is more strongly developed medially and becomes deflexed ventrad at the corner, whereas the other specimen has the posteromedial corner tuberculate and deflexed ventrad.</p><p>La Rivers (1970) described A. scolius without knowledge of the related species he would describe in the next few years; thus, he did not know if the sigmoid posterior margin of laterosternite VI would be a unique feature. However, he made no mention of its condition in the original descriptions of A. bifidus and A. tricuspis . Because A. scolius and A. tricuspis have the same subgenital plate shape and our examination reveals nearly the same condition for the posterior margin of laterosternite VI, we are not able to satisfactorily distinguish between them.</p><p>We provisionally determined eight specimens including four females from northwestern Pará, Brazil as A. scolius . However, there is substantial variation in the condition of the female subgenital plate and laterosternite VI. Specifically, the subgenital plate posterior margin varies from straight to strongly convex, but is not bifurcated. The right laterosternite VI varies from a flat, unmodified posterior margin to bearing a short posteromedian spine. This suggests that considerable intraspecific variation in these features might also exist in other species in the complex. As such, species determinations can be equivocal for most of these species and series of specimens should be collected when possible to establish the limits of expression of these features. For these specimens from Brazil, geographic proximity suggests conspecificity with A. scolius rather than with A. tricuspis .</p><p>Type locality. VENEZUELA, Monagas, 42 km SE Maturin. Repository of holotype unknown; two female paratypes are housed at CAS.</p><p>Material examined. BRAZIL: Pará state, 378, Rio Mapuera, Igarapé do Aracu, 19 June 1986, Py-Daniel &amp; Barbosa (2♂, 2♀, INPA; 2♂, 2♀, UMC). BRITISH GUIANA: Honey Camp Cr., Oct. 24 1937, S. Harris / A. scolius ♀ Paratype / Ira La Rivers Collection Bequeathed to the California Academy of Sciences (♀, CAS). TRINIDAD: B.W.I., Port of Spain, Nov. 5 1931, W. E. Broadway / A. scolius ♀ Paratype / Ira La Rivers Collection Bequeathed to the California Academy of Sciences (♀, CAS).</p><p>Published records. (La Rivers 1970): VENEZUELA, Monagas state.</p></div>	https://treatment.plazi.org/id/203D141B522EFFCD5588524102CBFB30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sites, Robert W.;Reynoso-Velasco, Daniel	Sites, Robert W., Reynoso-Velasco, Daniel (2015): Review of the Ambrysus stali La Rivers species complex (Heteroptera: Nepomorpha: Naucoridae) with the description of a new species from Mesoamerica. Zootaxa 4018 (2): 279-291, DOI: 10.11646/zootaxa.4018.2.7
203D141B5221FFCF55885026040AFD8F.text	203D141B5221FFCF55885026040AFD8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ambrysus stali La Rivers	<div><p>Ambrysus stali La Rivers</p><p>(Figs. 1, 6)</p><p>Ambrysus stali La Rivers 1962: Bull. S. Cal. Acad. Sci. Bull. 185–187. Ambrysus bourquini De Carlo 1968: Rev. Soc. Entomol. Arg. 100.</p><p>Discussion. La Rivers (1962) considered this species to be closely related to A. oblongulus and presented characteristics to distinguish them. The male accessory genitalic process of tergum VI (Fig. 6 B) is slightly narrower than in other members of the A. stali complex, although intraspecific variation is evident. Also distinctive is that the longitudinal stripes of the pronotum are less linear and more coalescent than in other members of the complex, and the hemelytra are uniformly brown or dark brown without lighter mottling (Fig. 6 A).</p><p>Reported here are the first records of A. stali from Guyana. Nieser (1975) indicated that this species is common in Suriname and occurs in small, shaded streams where it occurs in "barriers of branches and trees in moderate to quite strong currents (3– over 10 m /min.)", and less commonly at the margins on vegetation and debris.</p><p>In papers concerning the fauna of Argentina, López Ruf (1987, 2007) and Mazzucconi et al. (2009) indicated the presence of A. stali in Misiones Province, which is in the northeastern part of the country. López Ruf (2007) also provided a description of egg structure of the Argentina population. We have been unable to confirm their identifications because voucher specimens are not present at Museo de La Plata or Museo Argentino Ciencias Naturales. An undescribed species of Ambrysus occurs nearby in Paraguay that bears superficial resemblance to the A. stali complex; however, male genitalic features indicate that it is clearly does not belong to this complex.</p><p>Because the Argentina records are widely separated from all other records of the species and a similar species occurs in that region, we suspect that the Argentina material is probably not conspecific with A. stali . Type locality. FRENCH GUIANA, St. Laurent. Holotype is housed at CAS.</p><p>Material examined., FRENCH GUIANA: St. Laurent [du Maroni], October 1937, H. E. Hinton, Cal. Acad. Sci. Coll. / Ira La Rivers Collection, Bequeathed to the California Academy of Sciences—1978 / California Academy of Sciences Type No. 13412 / Ambrysus stäli La Rivers HOLOTYPE (♂, CAS). GUYANA: REGION 9, Parabara, elev 274 m, 2º 6' 29.52", -59 º13' 39.183", 3 November 2013, Short, Isaacs, Salisbury / GY–1103–02A, Small flowing creek with sand-detritus margins &amp; leaf packs-stick jams (2♂, 2♀, UMC); Kusad Mts., Basecamp, 2.811917, - 59.866833, 163, 23 October 2013, A. E. Short, small sandy stream with detritus, GY 13–1023–01 A (1♂, 1–5th instar, UMC); Region 9, Kusad Mts., Basecamp, 2.811917, -59.866833, 163 m, 23 October 2013, A. E. Short, pool in mostly dried streambed GY 13–1023–02 A (1♂, UMC); N. Parabara, creek by basecamp (Bototo wau creek), 2.181800, - 59.338432, 289 m, 31 October 2013, Short, Isaacs, Salisbury, stream margins, GY 13-1031-01 A (1♂, 1♀, UMC). SURINAME: SIPALIWINI, Camp 1, Kutari river, 2°10.973'N, 56°47.235'W, 238 m, 22 August 2010, Short &amp; Kadosoe, forest swamp, SR10-0822-02A (1♂, 2♀, UMC); Camp 1, Kutari river, 2.175350, -56.787399, 228 m, 20 August 2010, Short &amp; Kadosoe, forest stream, SR10-0820-01A (4♂, UMC); Camp 4 (low): Kasikasima, 2.977310, - 55.384998, 200 m, 20 March 2012, A. E. Short, sandy stream, SR12-0320-02A (2♂, UMC); Central Suriname Nature Reserve: Tafelberg Summit, near Augustus Creek Camp, 3.926667, -56.188332, 600 m, 17 August 2013, Short &amp; Bloom, forested detrital pools and creeks on trail into Arrowhead basin, SR13-0817-01A (1♀, UMC); Central Suriname Nature Reserve, Tafelberg Summit, near Augustus Creek Camp, 3.926667, -56.188332, 600 m, 17 August 2013, SR13–0817–03A (4♂, 5♀, UMC); Central Suriname Nature Reserve, Tafelberg Summit, near Caiman Creek Camp, 3.899033, -56.180817, 733 m, 19 August 2013, Short &amp; Bloom, Caimen Creek, pools and creek margins, SR13-0819-04B (1♂). TRINIDAD: bet. Four Roads &amp; Carmichael, 5 VII 2005, 220 ft, unnamed stream, 10º30'N, 61º13'W / (underside of same label) WDS-A-1647 / William D. Shepard, leg. (1♂, 1♀, UMC).</p><p>Published records. La Rivers (1962): BRAZIL, Pará; FRENCH GUIANA, St. Laurent [du Maroni]; [SURINAME]; TRINIDAD. La Rivers (1972), Nieser (1975): BRAZIL, Amazonas and Pará states; SURINAME, Brokopondo, Marowijne, and Saramacca Districts. Araujo &amp; Beserra (2007): VENEZUELA, Amazonas.</p></div>	https://treatment.plazi.org/id/203D141B5221FFCF55885026040AFD8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sites, Robert W.;Reynoso-Velasco, Daniel	Sites, Robert W., Reynoso-Velasco, Daniel (2015): Review of the Ambrysus stali La Rivers species complex (Heteroptera: Nepomorpha: Naucoridae) with the description of a new species from Mesoamerica. Zootaxa 4018 (2): 279-291, DOI: 10.11646/zootaxa.4018.2.7
203D141B5222FFCE5588558D05BBFC63.text	203D141B5222FFCE5588558D05BBFC63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ambrysus tricuspis La Rivers	<div><p>Ambrysus tricuspis La Rivers</p><p>(Figs. 1, 7)</p><p>Ambrysus tricuspis La Rivers 1974: Biol. Soc. Nevada Occas. Pap. 2–3.</p><p>Discussion. This species is known only from the holotype and allotype, which were collected together in northern Colombia. No information regarding habitat or biology was given in the original description. It differs from A. bifidus by the unbifurcated median lobe of the subgenital plate (Fig. 7 E). Although La Rivers (1974) suggested the male accessory genitalic process is more massive and elongated laterally (Fig. 7 B) than in A. bifidus, this feature exhibits intraspecific variation in other species of Ambrysus, including within this species complex. Specifically, Nieser (1975) documented sufficient intraspecific variation in the shape of the accessory genitalic process among paratypes of A. bifidus from Suriname that its precise shape should not be considered a reliable diagnostic attribute. La Rivers made no attempt to distinguish this species specifically from A. scolius, but thought the "three-pointed subgenital plate" and "exaggerated dog's head outline" of the male accessory genitalic process would satisfactorily distinguish A. tricuspis from all congeners. Although we are unable to find reliable characters to distinguish A. tricuspis from A. scolius, the slight difference in shape of the male accessory genitalic process, the geographic distance between the known ranges, and the paucity of specimens are enough that we chose not to synonymize these species at this time. More specimens of both species are needed to determine the extent of intraspecific variation in these features, which will then enable us to more effectively address the validity of this species.</p><p>Type locality. COLOMBIA, Bolivar Department. Type specimen is housed at CAS.</p><p>Material examined. COLOMBIA: Bolivar, San Marcos / (reverse side of locality label) 1964 Aug 8, O. Pinada / Ambrysus tricuspis Holotype / California Academy of Sciences Type No. 13406 / Ira La Rivers Collection Bequeathed to the California Academy of Sciences (♀, CAS); same data, Allotype (♂, CAS).</p></div>	https://treatment.plazi.org/id/203D141B5222FFCE5588558D05BBFC63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sites, Robert W.;Reynoso-Velasco, Daniel	Sites, Robert W., Reynoso-Velasco, Daniel (2015): Review of the Ambrysus stali La Rivers species complex (Heteroptera: Nepomorpha: Naucoridae) with the description of a new species from Mesoamerica. Zootaxa 4018 (2): 279-291, DOI: 10.11646/zootaxa.4018.2.7
