taxonID	type	description	language	source
AAAAF214F32639A1ACC40E9553063910.taxon	description	Description. Holotype male. Form elongate, slender; metathoracic wings present and fully developed. TL = 11.9 mm. Color: Head, pronotum and thorax piceous; elytra slightly lighter; antennae and mouthparts brunneous; legs testaceous; abdomen brunneous mesally, becoming pale testaceous laterally. Elytra with a pattern of testaceous markings and maculae as follows: the anterior 1 / 2 with a roughly X-shaped marking; each elytron at posterior 1 / 4 with a large, irregularly bordered, ovate macula narrowly separated at the elytral suture (Fig. 1). Head: HL = 2.15 mm, HW = 1.95 mm. Measured across eyes wider than pronotum; surface smooth, moderately shiny, moderately, finely punctate; frons bi-impressed; sparsely clothed with short, very fine, recumbent setae, more profusely vested behind the eyes with longer setae; eyes moderately small, subsinuate, taller than wide, moderately emarginate in front, moderately protuberant laterally, separated by approximately 2.5 eye-widths. Antennae slender; loosely composed; extending beyond posterior margin of elytra; third antennomere about two times the length of second antennomere, fourth antennomere slightly longer than third antennomere; antennomeres 5 - 10 subequal in length; antennomeres 4 - 10 weakly serrate, serration very gradually increasing distally; last antennomere flattened apically, as long as tenth antennomere (Fig. 6). Thorax: PL = 2.75 mm, PW = 1.9 mm. Pronotum elongate; widest at middle; middle slightly broader than anterior margin; sides constricted laterally; more strongly constricted behind middle; disc flat; moderately impressed in front of middle; subbasal tumescence feebly pronounced; surface rugulose; moderately, finely punctate; vested with short, fine, pale recumbent setae intermingled with less numerous, long erect setae, the latter more numerous laterally. Prosternum wider than long, rugulose, moderately puncticulate, scarcely vested with fine, pale, semirecumbent setae. Mesosternum smooth, shiny, feebly, coarsely, deeply punctate. Metasternum convex; rugose; moderately, shallowly punctate; moderately clothed with pale, semirecumbent setae, vestiture more abundant medially. Legs: Femora clothed with short, recumbent setae interspersed with a few erect and semierect setae; tibiae moderately vested with some short and long erect and semierect setae; femora and tibiae transversely rugose. Elytra: EL = 8.8 mm, EW = 2.6 mm. Anterior margin arcuately emarginate; wider than widest portion of pronotum; humeri pronounced; sides subparallel; widest on posterior fourth; disc convex; apex moderately dehiscent; elytral sculpturing as follows: anterior third set with regular, rather coarse and deep striae that abruptly diminish after anterior third and disappear entirely on posterior half; punctations at elytral base coarse and deep; surface moderately clothed with short, very fine, pale, recumbent setae intermixed with very few fine, long, erect setae. Abdomen: Ventrites 1 - 5 rugulose, strongly convex; shallowly, moderately punctate; subequal in length; each ventrite with a pair of large, pale, shallow impressions near sides; surface clothed with fine, pale, moderately long, recumbent setae. Fifth ventrite (Fig. 11) moderately smooth; sides moderately oblique and arcuate; posterior margin broadly, deeply emarginate, emargination reaches medial portion of segment, posterolateral angles acuminate; sixth ventrite (Fig. 11) longer than wide, surface rugulose, with a pair of oblique, V-shaped, longitudinal carinae that initiate on anterior 1 / 4 and end slightly beyond segmental mid-length; highly modified distally, posterior margin deeply emarginate, emargination U-shaped, with the posterolateral angles each produced as a dorsally recurved and conspicuously elongate extension, the last third of each posterolateral extension bearing an internal spine or acuminate protuberance. Fifth tergite (Fig. 20) moderately convex; rugulose; lateral margins subparallel; posterior margin bisinuate, broadly, moderately deeply, triangularly emarginate at middle. Sixth tergite (Fig. 20) conspicuously longer than wide; subtriangular; surface strongly convex; rugulose; moderately, shallowly punctate; lateral margin oblique; posterior margin narrowly, moderately rounded, much surpassed by posterolateral extensions of the sixth ventrite. Aedeagus 2.8 mm long; moderately robust; ratio of length of paramere to whole tegmen 0.45: 1; tegmen fully covering phallus; parameres robust throughout their length; lateral margins feebly obtuse, subparallel, pointed distally; phallobase wide; phallic plate armed with a row of denticles along dorsal margin; phallobasic apodeme robust distally, moderately short; phallobasic struts rather slender throughout their length (Fig. 23). Female: The female paratype is somewhat longer than the male, having a total body length of 13.8 mm. The female also differs from the male holotype by having the sixth ventrite strongly, deeply, V-shaped emarginate, lacking the pair of elongate, posterolateral extensions observed in the male (Figs 11, 20). Additionally, the female has a slightly paler integument than the male, with a less distinct elytral pattern anteriorly by comparison.	en	Burke, Alan F., Rifkind, Jacques, Zolnerowich, Gregory (2015): Four new species of Cymatodera Gray from central and southern Mexico (Coleoptera, Cleridae, Tillinae). ZooKeys 513: 105-121, DOI: http://dx.doi.org/10.3897/zookeys.513.9935, URL: http://dx.doi.org/10.3897/zookeys.513.9935
AAAAF214F32639A1ACC40E9553063910.taxon	distribution	Distribution. Presently known only from the northern portion of Hidalgo and the southeastern part of Tamaulipas, Mexico. The vegetation at the collecting localities is predominantly pine-oak forest (Fig. 29).	en	Burke, Alan F., Rifkind, Jacques, Zolnerowich, Gregory (2015): Four new species of Cymatodera Gray from central and southern Mexico (Coleoptera, Cleridae, Tillinae). ZooKeys 513: 105-121, DOI: http://dx.doi.org/10.3897/zookeys.513.9935, URL: http://dx.doi.org/10.3897/zookeys.513.9935
AAAAF214F32639A1ACC40E9553063910.taxon	etymology	Etymology. The specific epithet tortuosa (from the Latin tortilis or tortuosus, meaning twisted or winding), is a reference to the intricate and elaborate structure of the male pygidium of this species.	en	Burke, Alan F., Rifkind, Jacques, Zolnerowich, Gregory (2015): Four new species of Cymatodera Gray from central and southern Mexico (Coleoptera, Cleridae, Tillinae). ZooKeys 513: 105-121, DOI: http://dx.doi.org/10.3897/zookeys.513.9935, URL: http://dx.doi.org/10.3897/zookeys.513.9935
F0CAAF3B8A4B7FBAD81D0A28F7C09029.taxon	description	Description. Holotype male. Medium sized, moderately slender anteriorly, rather robust posteriorly; brachypterous, TL = 13.1 mm. Color: Head, pronotum, thorax, femora and anterior portion of tibiae brunneous; posterior portion of femora and tarsomeres testaceous; antennae and mouthparts fuscous; abdomen fuscous, slightly darker than thorax, distal portion of abdominal segments with a depressed testaceous mark; elytral ground light testaceous, with a slight greenish tinge. Each elytron bearing a pair of irregular, sinuate, darkened fasciae: the first fascia located on anterior third, extending from the elytral suture to the epipleural fold, conspicuously slender proximal to elytral suture then abruptly widening before epipleural fold; the second fascia located at the elytral mid-length, moderately wide, extending from the elytral suture to before the epipleural fold. Punctation on elytral ground infuscate (Fig. 2). Head: HL = 2.9 mm, HW = 2.5 mm. Large, measured across eyes wider than pronotum; surface rugose; frons bi-impressed; surface moderately punctate; clothed with short, fine, recumbent setae intermixed with long, semierect and erect setae; eyes mod erately small, form subsinuate, longer than wide, moderately emarginate in front, very feebly bulging laterally, separated by approximately 3 eye-widths. Antennae slender; loosely composed; extending beyond posterior margin of elytra; antennomeres 2 - 3 subequal in length; fourth antennomere slightly longer than third; fifth antennomere very slightly longer than fourth, antennomeres 5 - 10 subequal in length, antennomeres 4 - 10 weakly serrate; last antennomere flattened apically, somewhat acuminate distally, approximately the same length as ninth antennomere (Fig. 7). Thorax: PL = 4.05 mm, PW = 2.1 mm. Pronotum elongate; widest at middle; middle slightly broader than anterior margin; sides constricted subapically; more constricted behind middle; moderately impressed in front of middle; subbasal tumescence pronounced; surface rugose, moderately punctate, punctation rather coarse; clothed with short, pale, semirecumbent setae, intermingled with long, stiff, semierect pale setae. Prosternum convex; wider than long; surface rugose, shining, very feebly punctate. Mesosternum feebly convex; surface rugulose, moderately, deeply punctate, vested with fine, pale, recumbent setae. Metasternum convex; shortened longitudinally; surface rugulose, devoid of tubercles or carinae, moderately, shallowly punctate. Scutellum conspicuously wider than long; moderately setose. Legs: Femora clothed with short, recumbent setae intermingled with less numerous erect and semierect setae; tibiae vested with short and long erect and semierect setae; femora and tibiae transversely rugose; metathoracic legs with tarsomeres longer than those of pro- and mesothoracic legs. Elytra: EL = 7.7 mm, EW = 3.2 mm. Form: subovate (brachypterous type). Anterior margin arcuately emarginate; narrower than widest portion of pronotum; humeri very feebly indicated; sides widest on posterior fourth; disc convex; apex rounded, broadly dehiscent, not covering sixth ventrite; surface smooth, moderately clothed with short, fine, pale, recumbent setae intermixed with long, pale, fine, erect setae; sculpturing consisting of small, coarse punctures and larger punctation irregularly arranged from base to apex, punctures becoming less numerous behind anterior third, interstices about 3 x the diameter of punctures at elytral base. Abdomen: Ventrites 1 - 5 rugulose; shallowly, moderately punctate; each segment with a pair of large, shallow impressions near sides; surface clothed with short, recumbent setae intermixed with less numerous, long, semi-erect setae. Fifth ventrite (Fig. 12) moderately convex; sides oblique, moderately arcuate; posterior margin broadly, very deeply emarginate. Sixth ventrite (Fig. 12) subquadrate; protruding laterally (visible in dorsal view); rugose; surface moderately concave; somewhat punctate; lateral margins oblique, feebly arcuate; posterior margin broadly, very deeply emarginate; posterolateral angles somewhat blunt, slightly procurved inwardly. Fifth tergite (Fig. 21) convex, rugulose; lateral margins slightly oblique; posterior margin narrowly, shallowly emarginate. Sixth tergite (Fig. 21) subquadrate; moderately convex; surface rugulose; lateral margins moderately oblique; posterolateral angles rounded; posterior margin broadly, shallowly, triangularly emarginate. Aedeagus 2.3 mm long; ratio of length of parameres to whole tegmen 0.4: 1; tegmen fully covering phallus; parameres subparallel, pointed at apex, lateral margins fee bly oblique; phallobase moderately broad; phallus with copulatory piece feebly tapered distally; phallic plate armed with a row of moderately long denticles along the dorsal margin, these denticles increasing in size toward distal end; phallobasic apodeme short, robust, dilated distally; phallobasic struts slender throughout their length, each as long as phallobasic apodeme (Fig. 24). Females of the type series differ from males by having the sixth ventrite (Fig. 17) with lateral margins moderately oblique, feebly arcuate, and posterior margin very feebly, shallowly, narrowly emarginate, rather than broadly, deeply, semicircularly emarginate, as observed in males (Fig. 12); additionally, females have the sixth tergite (Fig. 22) subquadrate, with the lateral margins oblique and the posterior margin very feebly, shallowly, narrowly emarginate. Females closely resemble males in other respects.	en	Burke, Alan F., Rifkind, Jacques, Zolnerowich, Gregory (2015): Four new species of Cymatodera Gray from central and southern Mexico (Coleoptera, Cleridae, Tillinae). ZooKeys 513: 105-121, DOI: http://dx.doi.org/10.3897/zookeys.513.9935, URL: http://dx.doi.org/10.3897/zookeys.513.9935
F0CAAF3B8A4B7FBAD81D0A28F7C09029.taxon	distribution	Distribution. Available specimens were collected in the central-occidental part of Mexico, in the states of Colima, Jalisco and Michoacan. Distribution appears limited to the southern tip of the Sierra Madre Occidental (Fig. 29).	en	Burke, Alan F., Rifkind, Jacques, Zolnerowich, Gregory (2015): Four new species of Cymatodera Gray from central and southern Mexico (Coleoptera, Cleridae, Tillinae). ZooKeys 513: 105-121, DOI: http://dx.doi.org/10.3897/zookeys.513.9935, URL: http://dx.doi.org/10.3897/zookeys.513.9935
F0CAAF3B8A4B7FBAD81D0A28F7C09029.taxon	etymology	Etymology. The specific epithet is a patronymic honoring Cristina Ortega, a friend of the first author.	en	Burke, Alan F., Rifkind, Jacques, Zolnerowich, Gregory (2015): Four new species of Cymatodera Gray from central and southern Mexico (Coleoptera, Cleridae, Tillinae). ZooKeys 513: 105-121, DOI: http://dx.doi.org/10.3897/zookeys.513.9935, URL: http://dx.doi.org/10.3897/zookeys.513.9935
C32FCECDC3A1F6D76983688676F145E5.taxon	description	Description. Holotype male. Moderately small, rather robust, metathoracic wings complete, TL = 10.2 mm. Color: head, pronotum, prosternum, mesosternum ferrugineous-brown, the rest of the body uniformly testaceous brown. Each elytron with a fuscous, irregular fascia near the mid-length, extending from the elytral suture to the eighth row of striae, but not reaching the epipleural fold (Fig. 3). Head: HL = 1.2 mm, HW = 1.85 mm. Measured across eyes wider than pronotum; surface feebly rugose, shiny; frons bi-impressed; moderately, coarsely punctate; vested with short, recumbent setae interspersed with some erect, stiff setae on and behind eyes; eyes rather rounded, moderately large, slightly longer than wide, feebly emarginate in front, bulging laterally. Antennae reaching posterior margin of pronotum; second antennomere 0.75 x longer than third antennomere, antennomeres 3 - 10 subequal in length; antennomeres 2 - 3 subcylindrical; fourth antennomere feebly serrate; antennomeres 5 - 10 moderately serrate; last antennomere flattened apically, posterior margin acuminate, about 2 x the length of tenth antennomere (Fig. 8). Thorax: PL = 2.9 mm, PW = 1.5 mm. Pronotum rugose; moderately punctate; anterior margin as wide as middle; sides feebly constricted subapically; slightly more constricted behind middle; disc flat, very feebly impressed in front of middle; subbasal tumescence absent; surface clothed with moderately long, semirecumbent setae interspersed with some long, erect setae. Prosternum sparsely vested, feebly punctate. Mesosternum convex; surface shiny, moderately, shallowly punctate. Metasternum with surface rugulose, shiny, moderately, shallowly puncticulate. Legs: Moderately vested with semierect and some recumbent setae; femora puncticulate, rugulose; tibia moderately punctate, rugulose. Elytra: EL = 6.9 mm, EW = 4.4 mm. Broader than pronotum; humeri pronounced, rounded; sides subparallel, widest portion behind posterior third; disc flattened above; surface moderately rugulose; apices rounded, feebly dehiscent; elytral declivity gradual; integument clothed with short, semierect setae intermixed with fewer long, erect setae; surface bearing coarse punctation arranged in regular striae that gradually become smaller and shallower on posterior half, punctation not reaching elytral apex; interstices at elytral base about 1.2 x the width of punctation. Abdomen: Ventrites 1 - 5 rugulose, moderately, finely punctate, clothed with short, pale, fine, recumbent setae. First ventrite rather convex, subquadrate, conspicuously elevated at posterior 1 / 4 with a transverse, arcuate carina which does not attain posterolateral angles (Fig. 27). Fifth ventrite (Fig. 13) feebly convex; subquadrate in shape; surface shiny, moderately, shallowly, finely punctate; lateral margins oblique, somewhat arcuate; posterolateral angles rounded; posterior margin feebly, broadly emarginate. Sixth ventrite (Fig. 13) reduced; broader than long; subtriangular; lateral margins oblique, arcuate; hind margin moderately acuminate; surface feebly convex, shiny, moderately, finely, shallowly puncticulate. Fifth tergite subtriangular in shape, rugulose; lateral margin moderately oblique, posterior margin truncate. Sixth tergite subtriangular, broader than long; lateral margins strongly oblique, feebly arcuate; converging posteriorly; extending beyond apical margin of sixth ventrite. Aedeagus 1.2 mm long; feebly sclerotized; wide; ratio of length of paramere to whole tegmen 0.45: 1; tegmen partially covering phallus; parameres subtriangular; lateral margins obtuse, strongly oblique, pointed distally; phallobase wide; phallic plate armed with two long rows of moderately large denticles along dorsal and distal margins; phallobasic apodeme slender, somewhat shorter than endophallic struts; phallobasic struts slender throughout their length, 1.5 x the length of phallobasic apodeme (Fig. 25). Females can be distinguished from males based on the structure of the pygidium. The sixth ventrite and the sixth tergite are broadly rounded posteriorly in females (Fig. 18). Other characters are constant in both sexes.	en	Burke, Alan F., Rifkind, Jacques, Zolnerowich, Gregory (2015): Four new species of Cymatodera Gray from central and southern Mexico (Coleoptera, Cleridae, Tillinae). ZooKeys 513: 105-121, DOI: http://dx.doi.org/10.3897/zookeys.513.9935, URL: http://dx.doi.org/10.3897/zookeys.513.9935
C32FCECDC3A1F6D76983688676F145E5.taxon	distribution	Distribution. All specimens in the type series were collected in the locality of El Aguacero, approximately 10 miles northwest of Ocozocuautla, Chiapas, Mexico. The elevation at the type locality is approximately 650 m and the vegetation is predominantly tropical deciduous forest (Fig. 29).	en	Burke, Alan F., Rifkind, Jacques, Zolnerowich, Gregory (2015): Four new species of Cymatodera Gray from central and southern Mexico (Coleoptera, Cleridae, Tillinae). ZooKeys 513: 105-121, DOI: http://dx.doi.org/10.3897/zookeys.513.9935, URL: http://dx.doi.org/10.3897/zookeys.513.9935
C32FCECDC3A1F6D76983688676F145E5.taxon	etymology	Etymology. We name this beetle for Prof. Dr. Roland Gerstmeier (Technische Universitaet Muenchen, Germany), in recognition of his many contributions to the study of Cleridae.	en	Burke, Alan F., Rifkind, Jacques, Zolnerowich, Gregory (2015): Four new species of Cymatodera Gray from central and southern Mexico (Coleoptera, Cleridae, Tillinae). ZooKeys 513: 105-121, DOI: http://dx.doi.org/10.3897/zookeys.513.9935, URL: http://dx.doi.org/10.3897/zookeys.513.9935
F8A4CC0FC7B1A648A23F18B375CF1240.taxon	description	Description. Holotype male. Small, moderately slender, metathoracic wings complete. TL = 9.3 mm. Color: head, pronotum, prosternum, mesosternum and mouthparts testaceous; remainder of body pale testaceous (Fig. 4). Head. HL = 1.1 mm, HW = 1.6 mm. Measured across eyes wider than pronotum; surface feebly rugose, shiny; frons not bi-impressed; moderately, finely punctate; vested with pale, short, recumbent, fine setae interspersed with some erect, fine, long and less numerous setae; eyes moderately rounded, large, slightly longer than wide, feebly emarginate in front, conspicuously bulging laterally. Antennae long, extending to posterior half of elytral length; second and third antennomere small, slender, about the same length; fourth antennomere about 3 x longer than third antennomere, antennomeres 4 - 10 robust, moderately elongate, subequal in length; antennomeres 2 - 3 subcylindrical; antennomeres 4 - 10 moderately serrate; last antennomere acuminate posteriorly, flattened apically, about the same length of tenth antennomere (Fig. 9). Thorax: PL = 1.7 mm, PW = 0.9 mm. Pronotum moderately rugose, feebly, finely punctate; anterior margin as wide as middle and posterior margin; sides feebly constricted subapically; more constricted behind middle; disc flat, feebly impressed in front of middle; anterior pronotal impression present, subbasal tumescence obvious; surface moderately clothed with pale, stiff, short and long, semirecumbent setae. Prosternum sparsely vested, feebly, finely punctate. Mesosternum convex; surface shiny, smooth, moderately, shallowly punctate. Metasternum with surface feebly rugose, moderately, shallowly puncticulate. Scutellum ovoid, wider than long, posteriorly emarginate. Legs: Moderately vested with pale, fine, recumbent setae intermixed with some scattered, very long, pale, stiff setae; femora transversally rugulose; tibia feebly punctate, longitudinally, finely rugulose. Elytra: EL = 4.9 mm, EW = 2.1 mm. Broader than pronotum; humeri pronounced, rounded; sides slightly ovoid; widest portion at posterior fourth; disc moderately flat tened above, slightly depressed medially, smooth; apices subtriangular, feebly dehiscent; elytral declivity steep; integument clothed with short, pale, fine, recumbent setae intermixed with long, erect, pale, stiff setae; sculpture consisting of moderately coarse punctation arranged in regular striae that gradually become smaller and shallower on toward apex, punctation disappear before elytral apex; interstices at elytral base smooth, about 2.0 x the width of punctation. Abdomen: Ventrites 1 - 4 shiny, smooth; feebly, finely punctate; clothed with few short, pale, fine, recumbent setae; posterior margins truncate; lateral margins not depressed. Fifth ventrite (Fig. 14) conspicuously wider than long; surface smooth, shiny, moderately concave; lateral margins oblique, finely arcuate; posterior margin broadly, shallowly emarginate. Sixth ventrite (Fig. 14) small; broader than long; subtriangular; surface shiny, smooth, very finely rugulose, medially convex; lateral margins strongly oblique, arcuate, hind margin broadly, very shallowly emarginate; posterolateral angles rounded. Fifth tergite subquadrate; rugulose; lateral margin moderately oblique, posterior margin truncate. Sixth tergite subquadrate, broader than long; surface concave; lateral margins oblique, feebly arcuate; posterior margin truncate; posterolateral angles broadly rounded. Sixth tergite extending slightly beyond the apical margin of sixth ventrite. Aedeagus 0.9 mm long; feebly sclerotized; moderately wide; ratio of length of paramere to whole tegmen 0.3: 1; tegmen partially covering phallus; parameres ovoid; lateral margins obtuse, oblique, pointed distally; phallobase wide; phallic plate devoid of denticles, distal portion of phallic plate spinous, spines reduced; phallus rounded at apex, conspicuously wide at middle; phallobasic apodeme robust, swollen distally, longer than phallobasic struts; phallobasic struts moderately robust, swollen distally, approximately 1.2 x the length of phallobasic apodeme (Fig. 26). Females of the type series can be differentiated from males by the shape of the sixth ventrite. This segment is broadly rounded posteriorly (Fig. 19), rather than shallowly emarginate, as in males (Fig. 14). Remaining characters are constant in both sexes.	en	Burke, Alan F., Rifkind, Jacques, Zolnerowich, Gregory (2015): Four new species of Cymatodera Gray from central and southern Mexico (Coleoptera, Cleridae, Tillinae). ZooKeys 513: 105-121, DOI: http://dx.doi.org/10.3897/zookeys.513.9935, URL: http://dx.doi.org/10.3897/zookeys.513.9935
F8A4CC0FC7B1A648A23F18B375CF1240.taxon	distribution	Distribution. The type series was collected from various localities in the Sierra Mixteca of Mexico, specifically in the south and southwestern portion of the state of Puebla, and in central Guerrero state (Fig. 29). This region is characterized by tropical deciduous to thorny forest habitats.	en	Burke, Alan F., Rifkind, Jacques, Zolnerowich, Gregory (2015): Four new species of Cymatodera Gray from central and southern Mexico (Coleoptera, Cleridae, Tillinae). ZooKeys 513: 105-121, DOI: http://dx.doi.org/10.3897/zookeys.513.9935, URL: http://dx.doi.org/10.3897/zookeys.513.9935
F8A4CC0FC7B1A648A23F18B375CF1240.taxon	etymology	Etymology. The specific epithet makes allusion to the regional home of the Mixteca people, and of this new species.	en	Burke, Alan F., Rifkind, Jacques, Zolnerowich, Gregory (2015): Four new species of Cymatodera Gray from central and southern Mexico (Coleoptera, Cleridae, Tillinae). ZooKeys 513: 105-121, DOI: http://dx.doi.org/10.3897/zookeys.513.9935, URL: http://dx.doi.org/10.3897/zookeys.513.9935
