taxonID	type	description	language	source
270B87F2C6231356FF0DFC65FBDFFC95.taxon	diagnosis	Diagnosis. Peritrematal shields truncate behind fourth pair of coxae, and separated by triangular slit from body margin behind posterior peritrematal setae. Two pairs of peritrematal setae present, anterior peritrematal setae short, smooth or very finely pilose, posterior peritrematal setae considerably longer (3 – 10 times as long as anterior setae), finely pilose or smooth, apically tapering. Both pairs inserted dorsally on peritrematal shield. Peritremes short, reaching only posterior edge of third pair of coxae. Adgenital gland openings present, situated on conspicuous platelets. One pair of post-genital sclerites often present. Ventri-anal shield caudally fused to opisthonotal shield. Anterior margin of ventri-anal shield with two pairs of setae. Ventri-anal pores situated posterolaterally to adanal setae. Epistome with two pairs of longer central points and several shorter spines. Anterior margin of podonotum sloping gradually, setae j 1 inserted dorsally. Setae z 1 absent. Margin of opisthonotum with 7 – 8 pairs of setae. Notes on the genus. On the basis of a detailed observation of Zerconella leitnerae specimens collected in Slovenia and Croatia (Ujvári, 2009, 2010), I raise the subgenus Zercon (Zerconella) Willmann, 1953 (Figs 1, 2) to full generic rank because of the following features: posterior peritrematal setae are very long, smooth or very finely pilose, apically tapering and inserted dorsally in Zerconella; but shorter, usually apically pilose or feathered, often with hyaline ending and inserted ventrally in Zercon; peritremes are short (neotenic) (15 – 20 μm), reaching only the posterior edge of the third pair of coxae in Zerconella; peritremes are longer (60 – 80 μm), reaching the anterior edge of the third pair of coxae in Zercon; post-genital sclerites are usually present in Zerconella, absent in Zercon; the epistome has four long, central points and several shorter spines in Zerconella; usually three long, central points of which the medial one is often apically bifurcate or trifurcate in Zercon. The genus Zercon is heterogeneous in important generic characters, such as the number of setae on the anterior margin of the ventri-anal shield and the distribution of dorsal glands, therefore it can not be objectively compared with the genus Zerconella in its current form on the basis of these features. Regarding the genus Metazercon, Błaszak (1975 a) the author described similar generic characters of peritrematal shields, peritrematal setae, peritremes, number of ventri-anal setae, adgenital and post-genital attributes to that of Zerconella leitnerae. However Błaszak omitted to compare Metazercon with the former subgenus Zerconella. In addition Błaszak (1975 a) mentioned two characters which would distinguish the type species of Metazercon from Zerconella leitnerae. The number of marginal setae is eight in Metazercon, seven in Zerconella leitnerae, and the former has a deep incision on posterolateral side of peritrematal shields which is absent in the latter. Later, examining other specimens Halašková (1979) recognised that Metazercon species can have seven or eight pairs of marginal setae on the opisthonotum, and the incision of in the peritrematal shields is often absent. Dorsal gland openings are minute in most of the species, some of them often not visible, but it seems that the second pair of opisthonotal pores (gdJ 2) is associated with setae J 2 (on a line connecting setae J 2 and Z 2), while the third pair (gdJ 4) is associated with caudodorsal lobes in the Asian species. It is interesting that some of the species lack one pair of J-setae. Unfortunately no larvae or protonymphs are known from the redefined group, while ontogenetic studies of different Zerconidae genera show that setae J 2 – 5 are present in larval stage and J 1 appears in protonymphs (Lindquist & Moraza 1998). Presumably the four pairs of larval J-setae are present in each Zerconella species, while in species having only four pairs of J-setae the protonymphal J 1 setae are lacking. However the absence of setae J 1 may be an important derivative character. The zoogeographic distribution of Zerconella species suggests that the former character appeared independently in the European and some Asian species, as an evolutionary convergence. Besides the above-mentioned characters it is important to note that each species of the redefined genus Zerconella has an oblong shape, a relatively small body-size (230 – 350 μm) and two or four pairs of strongly sclerotised dorsal cavities. Setae of each species are smooth or very finely pilose, setiform, and often elongate. The genital opening of known males is situated below the line of sternal setae st 3.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6221351FF0DF8CEFDA1FF3B.taxon	diagnosis	Diagnosis. Caudodorsal region without elevations; surface of dorsal shields smooth, some dorsal setae and all marginal setae short.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6221351FF0DF8CEFDA1FF3B.taxon	distribution	Distribution. Central Europe.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6251351FF0DF98DFDCCF835.taxon	diagnosis	Diagnosis. Dorsal setae needle-like, elongated, except setae J 5 which are short. Setae of J-series diverging posteriorly. Glands gdJ 5 (Po 3) situated laterally to setae J 5. Margin of opisthonotum with 7 – 8 pairs of setae. A single pair of large, flat dorsal cavities present, with undulate margins and axes converging posteriorly. Three small, lobe-like structures present above level of dorsal cavities, lateral lobes with well-sclerotised anterolateral margins. Anterior surface of opisthonotum with reticulate pattern, posterior surface covered by small pits.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6251351FF0DF98DFDCCF835.taxon	distribution	Distribution. Korean Peninsula.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6251351FF0DFC78FD82FA0B.taxon	diagnosis	Diagnosis. Caudodorsal region with characteristic elevations. Gland openings gdJ 4 (Po 3) associated with former caudodorsal lobes. Surface of dorsal shields ornamented, all dorsal setae elongated.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6251351FF0DFC78FD82FA0B.taxon	discussion	Remarks. It is important to note that Zercon armiger Aoki, 1966 probably belongs to the subgenus Zerconella (Metazercon) as well, as was noted by Halašková (1979). Ventral characters of the latter were not described or figured in the original description, but the body size, shape of the idiosoma, the special caudodorsal structures, appearance of dorsal setae, small and inconspicuous gland openings and the geographic distribution (currently known from Japan) suggest that the species is closely related to the Eastern Asian Zerconella (Metazercon) species.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6251351FF0DFC78FD82FA0B.taxon	distribution	Distribution. Eastern Asia.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6251351FF0DFEE8FBEEFC45.taxon	diagnosis	Diagnosis. Setae z 3, s 5, r 3, I 5, Z 4, Z 5 and S 3 – 5 elongated, other dorsal setae significantly shorter. Setae J 1 absent, J 5 adjacent to each other. Margin of opisthonotum with seven pairs of setae. Two pairs of dorsal cavities present, each strongly sclerotised, rounded, with undulate margins, lateral pair slightly larger than medial pair. Both podonotum and opisthonotum smooth.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6251351FF0DFEE8FBEEFC45.taxon	distribution	Distribution. Central Europe.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6251351FF0DFEE8FBEEFC45.taxon	discussion	Remarks. The situation of the gland pores has not been described so far, because these pores are quite small and difficult to recognise in spite of the absence of dorsal ornamentation. The positions of the newly located poroid structures are the following: glands gdj 2 (po 1) situated anteriorly to setae j 3, gdj 4 seem to open anterolaterally to setae j 4, gds 4 posteromedially to setae s 4, gdz 6 on a line connecting z 6 and J 1, gdJ 2 not conspicuous, gdJ 4 on a line connecting J 3 and Z 4, gdS 5 lying anterolaterally to S 5.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6271353FF0DFF38FC44FD30.taxon	diagnosis	Diagnosis. Dorsal setae elongated, needle-like. Six pairs of J-setae present, which consist of larval setae J 2 – 5 and presumably two pairs of supplementary setae. Margin of opisthonotum with eight pairs of setae. Two pairs of dorsal cavities present, outer pair large, with strikingly undulate margins, inner pair smaller, with slightly undulate margins, both pairs with axes parallel to that of body. Two posterodorsal lobes situated anterolaterally to dorsal cavities, emerging towards body axis. Gland openings gdJ 4 situated on posteromedial edge of caudodorsal lobes, shifted beyond line of dorsal cavities. Whole opisthonotum covered by small pits.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6271353FF0DFF38FC44FD30.taxon	distribution	Distribution. Korean Peninsula.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6271353FF0DFF38FC44FD30.taxon	discussion	Remarks. This is the only known neotrichous Zerconella species. In the present study two pairs of central J-setae are considered as supplementary setae while setae J 1 is considered as absent. However the origin of the J-setae needs to be clarified through ontogenetic studies. Regarding the original description, some opisthonotal lyrifissures were misidentified as gland openings. Unfortunately, gdj 2 (po 1) and opisthonotal gland openings are not conspicuous on the three examined specimens.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6271353FF0DFCEDFDCCFB00.taxon	diagnosis	Diagnosis. Dorsal setae elongated, needle-like. Setae J 5 situated laterally to J 4. Margin of opisthonotum with eight pairs of setae. A single pair of large, rounded dorsal cavities present, with undulate margins and axes converging posteriorly. Two large, anteriorly sclerotised lateral and one smaller central lobe can be found above the level of the dorsal cavities. Glands gdJ 4 situated on anterior edge of lateral lobes, on a line connecting J 5 and Z 4. Whole opisthonotum covered by small pits.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6271353FF0DFCEDFDCCFB00.taxon	distribution	Distribution. Korean Peninsula.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6271358FF0DFABDFACDFDC5.taxon	materials_examined	Type material. Holotype female (AS- 876), Taiwan, Nantou county, Kunyang, logging road, primary broadleaved forest, from leaf-litter, 24 o 06.705 ' N, 121 o 11.977 ' E, 2200 m a. s. l., 22 May 2008., leg. L. Dányi, Z. Korsós & E. Lazányi. (in the Collection of Soil Zoology of the Hungarian Natural History Museum, Budapest). Paratypes: 2 females (AS- 905), Taiwan, Miaoli County, Taian, township, Shei-Pa National Park, Kuanwu, Yulun Trail, 24 ° 30.002 ' N, 121 ° 6.434 ' E, 2200 m a. s. l., primary broad-leaved forest, leaf-litter, 20 October 2009, leg. L. Dányi & E. Lazányi; 1 female (AS- 917), Taiwan, Nantou County, Ren-ai, township, Taroko N. P., Dayuling, Western slope of Mt. Maomu, Mt. Bilu Trail, 24 ° 11.095 ' N, 121 ° 18.615 ' E, 2580 m a. s. l., primary broad-leaved forest with stream, moss from trunks, 11 October 2009, leg. L. Dányi & E. Lazányi; 35 females, 2 males, 1 deutonymph (AS- 932), Taiwan, Hualien County, Xiulin, township, Taroko N. P., Dayuling, Eastern slope of Mt. Maomu, 24 ° 10.881 ' N, 121 ° 19.408 ' E, 2498 m a. s. l., disturbed secondary mixed forest, leaf-litter, 11 October 2009, leg. L. Dányi & E. Lazányi.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6271358FF0DFABDFACDFDC5.taxon	diagnosis	Diagnosis. Dorsal setae rarely, very finely pilose or smooth. Setae z 3, r 3, s 5 and S 3 – 5 very long, extending beyond margin of idiosoma by more than half of their length. Opisthonotal setae relatively long, thickened, apically tapering. Setae J 5 situated medially to J 4. Margin of opisthonotum with seven pairs of setae. Central surface of opisthonotum covered by alveolar pits. Single pair of well-sclerotised dorsal cavities present, covered by large, posterocentral lobes which have strongly sclerotised, continuous posterior margin running arcuately between setae S 5.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6271358FF0DFABDFACDFDC5.taxon	description	Description. Female. Length of idiosoma: 265 – 280 μm; width 175 – 195 μm (n = 39). Dorsum (Figs 11, 15). Podonotum with 20 pairs of finely pilose setae (Fig. 16). Setae z 3 and s 5 very long (48 – 65 μm), apically tapering, reaching beyond margin of idiosoma by more than half of their length. Other podonotal setae similar in shape, setiform (16 – 36 μm). Setae j 6 reaching beyond bases of J 2, z 6 reaching beyond insertions of Z 1. Gland openings gdj 2 (po 1) situated below line connecting insertions of j 3 and s 1, remaining podonotal gland pores not conspicuous. Sculpturing pattern of podonotum squamous, posteromedial surface with small, alveolar pits. Opisthonotum with 20 pairs of elongate, apically tapering, finely pilose setae. Setae J 1 absent. Setae J 3 reaching beyond insertions of J 4 and J 5, 1.5 – 2 times as long as J 2. Setae J 5 situated medially to J 4, shorter than J 3 – 4. Setae Z 1 shorter than any other opisthonotal setae. Setae Z 2 – 5 similar in shape and length to central J-setae. S 2 similar in shape and length to J 2. Setae S 3 – 5 (Fig. 20) very long, reaching beyond margin of idiosoma, apically tapering. Marginal setae uniform, similar to that of podonotal margin. Size of opisthonotal setae and distance between their bases according to Table 1. Only exterior pair of dorsal cavities present, which large, strongly sclerotised, with smooth margins and axes converging posteriorly. Two large, lobe-like structures (Figs 17 – 18) emerging on posterior part of opisthonotum towards the caudal margin of idiosoma, covering dorsal cavities and bases of setae JV 5. These lobe-like structures have a continuous, strongly sclerotised posterior margin expanding between setae S 5. Small mound can be observed posteromedially to dorsal cavities. Gland openings gdJ 4 (Po 3) situated on a line connecting Z 4 and Z 5, near caudolateral margin of characteristic lobes, others not conspicuous. Marginal serration of dorsal idiosoma obtuse. Central and lateral surface of opisthonotum covered by alveolar pits. Ve n te r (Fig. 12). Peritrematal shields truncated at level of fourth pair of coxae, between level of S 1 - R 1, separated by triangular slit from podonotal shield, and ornamented by longitudinal fissures. Peritremes extremely short. Both peritrematal setae very finely pilose, r 3 six times as long as r 1. Sternal shield 46 – 50 μm long, 37 – 40 μm wide at level of setae st 2, posterior margin slightly concave. Adgenital platelets small, bearing 2 gland-openings of gv 2. One pair of post-genital sclerites often present. Ventri-anal shield with 7 pairs of preanal setae, posteriorly fused to opisthonotal shield. All ventral setae smooth or finely pilose. Ventri-anal pores situated near adanal setae, in posterolateral direction. Anal valves with vestigial euanal setae. Sternal shield with reticulate ornamentation, ventri-anal shield covered by tile-like pattern. Male (Figs 13 – 14). Length of idiosoma: 210 – 215 μm; width 151 – 154 μm (n = 2). Dorsal chaetotaxy, adenotaxy, poroidotaxy and sculpturing pattern similar to that of female. Length of opisthonotal setae and distance between their bases according to Table 1. Sternogenital shield with five pairs of setae (st 1 – 5). Genital opening situated behind level of setae st 3. Posterior ends of peritrematal shields free, truncated at level of S 1. Ƥ 3 DN Ƥ 3 DN Ƥ 3 DN Z 1 16 22 28 S 1 30 22 23 Z 1 - Z 2 33 21 30 S 1 - S 2 24 17 20 J 2 32 22 25 Z 2 45 29 36 S 2 33 22 31 J 2 - J 3 30 20 25 Z 2 - Z 3 30 17 18 S 2 - S 3 20 17 20 J 3 53 40 45 Z 3 55 49 67 S 3 63 39 30 J 3 - J 4 25 15 18 Z 3 - Z 4 15 15 16 S 3 - S 4 33 24 28 J 4 45 28 39 Z 4 47 38 48 S 4 69 52 72 J 4 - J 5 23 17 17 Z 4 - Z 5 48 41 38 S 4 - S 5 33 23 25 J 5 38 24 41 Z 5 50 41 46 S 5 69 60 77 Deutonymph (Fig. 19). Length of idiosoma: 247 μm, width 164 μm (n = 1). Dorsal chaetotaxy, adenotaxy and poroidotaxy similar to that of adults, sculpturing pattern on podonotum and lateral surface of opisthonotum weakly developed. Shape of posterodorsal lobes as in Fig. 19. Length of opisthonotal setae and distance between their bases according to Table 1.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6271358FF0DFABDFACDFDC5.taxon	etymology	Etymology. The name of the new species refers to the characteristic caudodorsal lobes.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
270B87F2C6271358FF0DFABDFACDFDC5.taxon	discussion	Remarks. The new species is closely related to Zerconella (Metazercon) athiasae (Błaszak, 1975) and Zerconella (Metazercon) rafalskii (Błaszak et al., 1997) by having only two dorsal cavities. Females of the three species can be distinguished by the following features. In Zerconella (Metazercon) lobata, setae J 5 are situated medially to J 4, setae J 1 are absent, the sculpturing pattern on the lateral parts of the opisthonotum is weakly developed, the surface between the J-series setae is covered by large, alveolar depressions, and a well sclerotised arcuate margin runs between setae S 5, laterally forming two caudodorsal lobes; in Zerconella (Metazercon) athiasae, setae J 5 are situated posterolaterally to J 4, setae J 1 are present, the anterior and lateral surface of the opisthonotum is covered by a tile-like pattern, the anteromedian surface between the J-series setae has reticulate ornamentation, spot-like depression can only be found between setae J 4 and J 5, on the caudodorsal part three separate, well sclerotised mounds can be observed; in Zerconella (Metazercon) rafalskii setae J 5 are situated posteromedially to J 4, setae J 1 are present, the sculpturing pattern on lateral parts of opisthonotum is weakly developed, the surface between the J-series setae is covered by small, spotlike depressions, and on the caudodorsal part three separate, well sclerotised mounds can be observed.	en	Ujvári, Zsolt (2010): Zerconella Willmann, 1953, a forgotten group of Zerconidae (Acari: Mesostigmata). Zootaxa 2558: 33-47, DOI: 10.5281/zenodo.196960
