taxonID	type	description	language	source
244E8D140B1C214F508EC0E5F526FCE0.taxon	description	Description. Macropterous female. HOLOTYPE, length 11.50; maximum width 7.55. Paratypes (n = 10), length 10.72 – 11.44 (mean = 11.17); maximum width 6.72 – 7.52 (mean = 7.14). General shape elongate, parallel-sided; widest across embolia (Fig. 1 A). Overall dorsal coloration of hemelytra brown to dark brown with yellowish brown marks; head and pronotum yellowish with brown marks; legs yellowish brown. Dorsal surface coarsely punctate. Ventral coloration of head and thorax yellowish, abdomen with yellowish pubescence. Head. Head length 1.68; maximum width 3.16. Mostly yellowish with brown marks along medial line and Thorax. Pronotum coarsely punctate, ground color yellowish, brown marking posteromedially, brown longitudinal line behind eye; transverse sulcus marking anterior border of transverse band in posterior 1 / 4; transverse band yellowish anteriorly, brown posteriorly; lateral margins yellowish, convergent, evenly convex, explanate; posterior margin almost straight; anterior margin deeply concave between eyes to embrace convex posterior margin of head; posterolateral corners rounded; width 3.1 × length; length at midline 2.08; maximum width at posterolateral corners 6.44. Prothorax ventrally pruinose throughout, except laterally; apices of propleura meeting at midline, not appressed to prosternellum; propleuron yellowish, light brown posteriorly, medial 2 / 3 of posterior margin with golden setae. Probasisternum with sharp median carina and row of setae lateral to carina. Prosternellum not covered by apices of propleura. Scutellum coarsely punctate, triangular, yellowish brown, two reniform to ovate brown marks medially, width 2.2 × length, width 4.32, length 1.92. Hemelytra densely punctate, brown to dark brown, length 8.0 (chord measurement); corium dark brown, yellowish irregularly shaped and brown-spotted yellowish checkmarkshaped markings at lateral and posterior margins, respectively (Fig. 1 A). Clavus dark brown in lateral half, brown in mesal half with dark brown close to base and apex, length 3.8 × width, length 4.56, width 1.20; claval commissure dark brown, length 1.12. Embolium length 3.48, greatest width 1.11; lateral margin convex, yellowish in anterior 2 / 3, brown posteriorly. Membrane dark brown (Fig. 1 A). Hindwings extending to posterior margin of tergum VI. Mesobasisternum tumescence negligible, with sulcus on midline continuing through triangular mesosternellum. Metasternellum (= metaxyphus) transverse, subtriangular, with median carina. Legs. All leg segments yellowish brown. Profemur yellowish brown; posterior margin with row of tightly arranged setae in basal 2 / 3, row of short brown spines along middle third; anterior margin with dense pad of setae without associated spines. Protibia and - tarsus with occlusal inner surface flattened and with spatulate setae; tarsus immovable, one-segmented; pretarsal claw single, minute, triangular. Procoxa with cluster of stout, brown anteromedial spines. Meso- and metacoxae partially recessed into thorax. Meso- and metafemora with row of short, light brown spines on anterior margin; spines restricted to basal half on mesofemur, nearly full length of metafemur. Mesotibia with ventrolateral and dorsolateral rows of stout reddish-brown spines; mesolateral rows intermixed with combs of two to five spines. Metatibia with ventrolateral and dorsolateral rows of stout reddish-brown spines. Meso- and metatibiae with semi-circlet of spines at apex of dorsal and mesal margins; two comb rows of stout spines near apex of ventral margin. Meso- and metatibiae and metatarsus with long, yellow swimming hairs; hairs profuse on metatibia and - tarsus. Meso- and metapretarsi with paired claws slender, gently curved, with small basal tooth. Leg measurements as follows: foreleg, femur 2.52, tibia 2.00, tarsus 0.48; middle leg, femur 2.60, tibia 2.30, tarsomeres 1 – 3, 0.22, 0.45, 0.51; hind leg, femur 3.48, tibia 4.00, tarsomeres 1 – 3, 0.25, 0.82, 0.72. Abdomen. Dorsally with lateral margins of III – VI exposed, each brown in anterior 1 / 3 and light brown posteriorly; lateral margins of II – basal half of III nearly smooth and distal half of III – VI finely serrate, marginal row of profuse and short yellow setae, group of trichobothria near posterolateral corners. Posterolateral corner of II (visible ventrally) narrowly rounded, right angled, III – VII acute. Ventrally yellowish, with dense pile of fine hairs. Lateral margin with thin, glabrous band. Glabrous rounded to oval patches near spiracles on laterosternites II – VI. Mediosternite VII (subgenital plate) asymmetrical, width 1.2 × length; length at midline 1.17; maximum width 1.42; low lateral lobe at base of left margin; posterolateral corners pointed; wide central lobe notched giving the appearance of two pointed lobes, lobes produced posteriorly further than posterolateral corners (Fig. 1 B). Macropterous male. Paratypes (n = 10), length 9.92 – 10.48 (mean = 10.21); maximum width 6.32 – 6.88 (mean = 6.66). Coloration, setation, pronotum proportions, posterolateral corners of pronotum, scutellum proportions, embolium length generally same as for holotype. Otherwise hindwings extending to posterior margin of tergum V. Accessory genitalic process of tergum VI slender, extending nearly straight posteriorly in basal 2 / 3, abruptly curved to right at approximately 70 degree angle in apical 1 / 3, apex narrowly rounded (Fig. 1 C). Medial lobes of tergum VIII (pseudoparameres) symmetrical, laterally produced, posterolateral and posteromesal corners broadly rounded (Fig. 1 D). Phallosoma elongate, linear, slender, slightly constricted basally; left endosomal sclerite subtriangular (lateral view), with transverse row of small denticles apically; right endosomal sclerite small, rounded, with a few denticles anteriorly (Fig. 1 E). Parameres symmetrical, almost as long as maximum width, mesal margin rounded, setae emanating from distal 1 / 3 of dorsal surface. Proctiger as long as width at base. Pygophore with elongate setae sparsely distributed over most of surface, with thick brush of elongate setae on posterior margin (Fig. 1 F).	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B1C214F508EC0E5F526FCE0.taxon	discussion	Discussion. Based on the generalized shape of the female SGP and the male phallosoma with the right endosomal sclerite small and rounded we consider this species to be closely related to A. cayo Sites & Shepard, A. colimanus Polhemus & Polhemus, A. inflatus La Rivers, and A. quadracies La Rivers. Diagnostic features and comparative notes. Females of this species exhibit a characteristic SGP with a low and rounded lateral lobe at the base of the left margin. The central lobe on the posterior margin is deeply notched and appears as two pointed lobes that are produced further than the posterolateral corners. A similar morphology of the posterior margin of the SGP is present in the species A. cayo, A. colimanus, A. inflatus, and A. quadracies, although in A. cayo, A. colimanus, and A. quadracies the posterolateral corners and central notched lobe are almost at the same level. In A. inflatus, the posterolateral corners are more pointed, the notched central lobe is narrower, and the lateral lobe is at mid-length of the left margin. We have noticed some variation on the SGP of A. brailovskyi that includes a low and rounded lateral lobe at the base of the right margin (symmetrical SGP) and the central lobe on the posterior margin appearing as two rounded lobes.	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B1C214F508EC0E5F526FCE0.taxon	description	Habitat description. The type locality at San José Veracruz (L- 1808) is in the northwestern area of the Mexican state of Chiapas, at an elevation of 862 meters. At this locality the stream is shallow, 3 – 10 meters wide, and with limestone boulders and cobble in the channel (Fig. 2). The microhabitat where most specimens of A. brailovskyi were found was the protected area behind rocks in the channel.	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B1C214F508EC0E5F526FCE0.taxon	distribution	Distribution. This species is only known from the Mexican state of Chiapas (Fig. 3). Ambrysus brailovskyi is distributed in the highlands (569 – 1010 meters) of the Chiapas biogeographic province. The species was collected syntopically with several species of Ambrysus, including with A. pulchellus Montandon and A. spicatus La Rivers at Río Matasano (L- 1348), with A. circumcinctus Montandon at Río Prusia (L- 1345), and with A. sitesi n. sp. at Finca Prusia (12 - V- 1985), Río Prusia (L- 1345), Río Matasano (L- 1348), and the Tenejapa municipality (14 - VII- 1964). Ambrysus brailovskyi also was collected syntopically with the rock-clinging saucer bug Cataractocoris macrocephalus (Montandon) at Río Prusia (L- 1345).	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B1C214F508EC0E5F526FCE0.taxon	etymology	Etymology. This species is named in honor of our colleague Professor Harry Brailovsky, of the Universidad Nacional Autónoma de México, to recognize his remarkable career on the study of Heteroptera, which heretofore includes the descriptions of 153 genera and 987 species. Repositories. The holotype and some paratypes will be deposited in the Colección Entomológica del Instituto de Ecología A. C. (Xalapa); additional paratypes will be deposited in the Colección Nacional de Insectos (Mexico City), Enns Entomology Museum (Columbia), and the United States National Museum of Natural History (Washington D. C.).	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B1C214F508EC0E5F526FCE0.taxon	materials_examined	Type material examined. HOLOTYPE ♀. MÉXICO: CHIAPAS: Mpio. Chilón, San José Veracruz, 17 ° 04 ’ 23.9 ’’ N, 91 ° 58 ’ 05.7 ’’ W, 862 m, 02 - Jun- 2014, L- 1808, DRV & PRH colls. PARATYPES: same data as holotype (7 ♂, 13 ♀ UMC); [Mpio. Ángel Albino Corzo], Finca Prusia, 12 - V- 85, M Vértiz / Colección del Instituto de Biología, UNAM. México, D. F (1 ♂ CNIN); Mpio. Ángel Albino Corzo, 1 km S of Finca Prusia, Río Prusia, 15 º 43 ’ 23.3 ’’ N, 92 º 47 ’ 46.5 ’’ W, 1010 m, 18 - May- 2012, L- 1345, DRV & PRH colls. (3 ♂, 2 ♀ UMC); Mpio. Chilón, Guadalupe Paxila, Río Paxila, 17 ° 07 ’ 01.6 ’’ N, 92 ° 01 ’ 43.9 ’’ W, 569 m, 02 - Jun- 2014, L- 1809, DRV & PRH colls. (4 ♂, 4 ♀ IEXA); Mpio. Chilón, Jet-Mesil, Río Bolontina, 17 ° 07 ’ 11.1 ’’ N, 92 ° 05 ’ 18.8 ’’ W, 584 m, 02 - Jun- 2014, L- 1810, DRV & PRH colls. (1 ♂, 2 ♀ IEXA); [Mpio. Ixhuatán], 4 mi N of Tapiula [Tapilula], CL 1097, 5 May 1964, JT & MS Polhemus / C. J. Drake Accession (1 ♂ USNM); [Mpio. Ixtacomitán], Ixtacomitán, Río Blanco, 10 - XII- 85, cols. F Arias, L Cervantes y R Barba / Colección del Instituto de Biología, UNAM. México, D. F. (3 ♂, 2 ♀ CNIN); Mpio. Ocosingo, Las Lajas, Río Las Lajas, 16 ° 57 ’ 13.6 ’’ N, 92 ° 06 ’ 51.1 ’’ W, 922 m, 02 - Jun- 2014, L- 1811, DRV & PRH colls. (1 ♂, 1 ♀ IEXA); Mpio. Siltepec, Matasano, Río Matasano, 15 º 36 ’ 13.7 ’’ N, 92 º 33 ’ 37.1 ’’ W, 950 m, 19 - May- 2012, L- 1348, DRV & PRH colls. (3 ♂, 3 ♀ UMC); Mpio. Tenejapa, ponds in stream above Habenal / 14. VII. 1964, coll. D Breedlove / C. J. Drake Accession / CAS (1 ♀ USNM); km 88 carr. Palenque-Ocosingo, 8 - V- 85, F Arias / Colección del Instituto de Biología, UNAM. México, D. F (1 ♀ CNIN).	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B182143508EC09EF204FC76.taxon	description	(Figs. 3 – 5) Description. Macropterous female. HOLOTYPE, length 12.12; maximum width 8.12. Paratypes (n = 10), length 10.64 – 12.48 (mean = 11.73); maximum width 6.88 – 8.16 (mean = 7.60). General shape ovate; widest across embolia (Fig. 4 A). Overall dorsal coloration of hemelytra dark brown to black with yellowish brown marks; head and pronotum yellowish brown with brown marks; legs yellowish brown. Dorsal surface coarsely punctate. Ventral coloration of head and thorax yellowish, abdomen with yellowish pubescence. Head. Head length 1.76; maximum width 3.48. Mostly yellowish brown with brown marks along midline and posteriorly; coarsely punctate. Eyes convergent anteriorly, synthlipsis 1.72; thin band of cuticle along posterolateral margin of eye; eyes not raised above level of vertex or pronotum. Anterior margin between eyes slightly convex, extending anteriorly in front of eyes 8.5 % of head length; posterior margin between eyes strongly convex, extending posteriorly 30.6 % of head length. Labrum width 1.9 × length, evenly rounded. Labium with three visible yellowish brown segments, darkening distally, extending 0.37 beyond labrum not including extruded stylets. Antennal proportions 3: 19: 21: 11, length 0.81, extending to lateral margin of eye, elongate hairs on segment 4 and distal half of 3. Thorax. Pronotum coarsely punctate, ground color yellowish brown, brown marking posteromedially, black longitudinal line behind eye; transverse sulcus marking anterior border of transverse band in posterior 1 / 3; transverse band yellowish anteriorly, brown posteriorly; lateral margins yellowish, convergent, evenly convex, explanate; posterior margin almost straight; anterior margin deeply concave between eyes to embrace convex posterior margin of head; posterolateral corners rounded; width 2.8 × length; length at midline 2.52; maximum width at posterolateral corners 7.06. Prothorax ventrally pruinose throughout, except laterally; apices of propleura meeting at midline, not appressed to prosternellum; propleuron yellowish, light brown posteriorly, medial 2 / 3 of posterior margin with golden setae. Probasisternum with sharp median carina and row of setae lateral to carina. Prosternellum not covered by apices of propleura. Scutellum coarsely punctate, triangular, brown, two reniform to ovate dark brown marks medially, width 2.3 × length, width 4.76, length 2.04. Hemelytra densely punctate, dark brown to black, length 8.96 (chord measurement); corium dark brown, brown suboval and yellowish brown checkmark-shaped markings at lateral and posterior margins, respectively. Clavus dark brown to black, yellow at each end, faint yellowish spot near middle, length 3.8 × width, length 4.80, width 1.24; claval commissure black, length 1.20. Embolium length 3.84, greatest width 1.17; lateral margin convex, yellowish brown in anterior 2 / 3, dark brown posteriorly. Membrane black (Fig. 4 A). Hindwings extending to posterior margin of tergum VII. Mesobasisternum tumescence negligible, with sulcus on midline continuing through triangular mesosternellum. Metasternellum (= metaxyphus) transverse, subtriangular, with median carina. Legs. All leg segments yellowish brown. Profemur yellowish; posterior margin with row of tightly arranged setae in basal 2 / 3, row of short brown spines along middle third; anterior margin with dense pad of setae without associated spines. Protibia and - tarsus with occlusal inner surface flattened and with spatulate setae; tarsus immovable, one-segmented; pretarsal claw single, minute, triangular. Procoxa with cluster of stout, brown anteromedial spines. Meso- and metacoxae partially recessed into thorax. Meso- and metafemora with row of short, light brown spines on anterior margin; spines restricted to basal half on mesofemur, nearly full length of metafemur. Mesotibia with ventrolateral and dorsolateral rows of stout reddish-brown spines; mesolateral rows intermixed with combs of two to six spines. Metatibia with ventrolateral and dorsolateral rows of stout reddish-brown spines. Meso- and metatibiae with semi-circlet of spines at apices on dorsal and mesal margins; two comb rows of stout spines near apices on ventral margin. Meso- and metatibiae and metatarsus with long, yellow swimming hairs; hairs profuse on metatibia and - tarsus. Meso- and metapretarsi with paired claws slender, gently curved, with small basal tooth. Leg measurements as follows: foreleg, femur 2.65, tibia 1.90, tarsus 0.52; middle leg, femur 2.75, tibia 2.37, tarsomeres 1 – 3, 0.18, 0.37, 0.52; hind leg, femur 3.72, tibia 4.04, tarsomeres 1 – 3, 0.25, 0.84, 0.73. Abdomen. Dorsally with lateral margins of III – VI exposed, each brown in anterior 1 / 3 and light brown posteriorly; lateral margins of II smooth and III – VI finely serrate, marginal row of profuse and short yellow setae, group of trichobothria near posterolateral corners. Posterolateral corner of II (visible ventrally) narrowly rounded, right angled, III – VII acute. Ventrally yellowish, with dense pile of fine hairs. Lateral margin with thin, glabrous band. Glabrous rounded to oval patches near spiracles on laterosternites II – VII. Mediosternite VII (subgenital plate) symmetrical, subtriangular, width 1.05 × length; length at midline 1.18; maximum width 1.24; inconspicuous posterolateral corners; apex truncate (Fig. 4 B). Macropterous male. Paratypes (n = 10), length 10.32 – 11.68 (mean = 10.87); maximum width 6.56 – 7.52 (mean = 7.06). Coloration, setation, pronotum proportions, posterolateral corners of pronotum, scutellum proportions, embolium length generally same as for holotype. Otherwise hindwings extending to posterior margin of tergum VI. Accessory genitalic process of tergum VI robust, abruptly curved to right close to the base at approximately 90 degree angle, apex narrowly rounded (Fig. 4 C). Medial lobes of tergum VIII (pseudoparameres) symmetrical, posterolaterally produced, posterolateral corner broadly rounded, extending posteriorly further than posteromesal corner (Fig. 4 D). Phallosoma elongate, linear, slender, slightly constricted basally; left endosomal sclerite subtriangular (lateral view), with group of small denticles apically; right endosomal sclerite knob-like, with conspicuous denticles (Fig.	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B182143508EC09EF204FC76.taxon	discussion	Discussion. This species superficially resembles A. hydor La Rivers, although A. henryi is narrower and the posterolateral corners of abdominal segments III – V are less produced. The morphology of the endosomal sclerites of the males suggests that A. henryi is related to A. dyticus La Rivers, A. hydor, A. magniceps La Rivers, A. occidentalis La Rivers, A. scalenus La Rivers, and A. sonorensis in the A. signoreti species complex. Diagnostic features and comparative notes. Females of this species exhibit a unique subtriangular and truncate SGP. Another species from this complex that presents the SGP with similar inconspicuous posterolateral corners is A. sonorensis; however, this species has clear lateral lobes and the SGP is almost as wide as long (quadrate). The AGP exhibited by males of A. henryi is the widest among species of this complex. Males of A. inflatus present a similar AGP, although it is narrower and less strongly curved.	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B182143508EC09EF204FC76.taxon	description	Habitat description. The type locality at Río San Juan (L- 1904) is in the southern area of the Mexican state of Oaxaca, on the Sierra Madre del Sur Mountain Range, at an elevation of 400 meters. At this locality the river is shallow, 3 – 6 meters wide, and the substratum is mainly composed of gravel and cobble (Fig. 5). The microhabitat where we collected this species was among rocks, close to the river margins.	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B182143508EC09EF204FC76.taxon	distribution	Distribution. This species occurs in the southern Mexican states of Chiapas, Guerrero, and Oaxaca (Fig. 3). Ambrysus henryi is distributed in the Chiapas (Chiapas), Mexican Pacific Coast (Chiapas, Oaxaca), and Sierra Madre del Sur (Guerrero, Oaxaca) biogeographic provinces. In the state of Oaxaca, this species was collected syntopically with various congeners, including A. angularis La Rivers, A. baeus Polhemus & Polhemus, A. pudicus Stål, and A. spicatus at Río San Juan (L- 1904, the type locality of A. henryi); with A. baeus and A. spicatus at Río San Juan (L- 1375, L- 1376, L- 1905); with A. baeus, A. pulchellus, and A. schuhi n. sp. at Río Rana (L- 1380); with A. baeus, A. schuhi n. sp., A. sitesi n. sp., and A. spicatus at Río Rana (L- 1906); and with A. angularis, A. parviceps Montandon, A. pudicus, and A. pulchellus at Pte. Yutaniñe (L- 1385) and Mártires de Tacubaya (L- 1390). In the state of Chiapas, at the Velo de Novia Waterfall (L- 1359, L- 1903), this species was found syntopically with A. pudicus, A. pulchellus, and the three known species in genus Cataractocoris Usinger. This waterfall is the type locality of Cataractocoris shepardi Sites, Reynoso & Novelo.	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B182143508EC09EF204FC76.taxon	etymology	Etymology. The specific epithet henryi is in honor of our colleague Professor Thomas J. Henry, of the United States National Museum of Natural History, to commemorate a lifetime dedicated to the study of Heteroptera. Among his numerous accomplishments are the descriptions of approximately 300 insect taxa. Repositories. The holotype and some paratypes will be deposited in the Colección Entomológica del Instituto de Ecología A. C. (Xalapa); additional paratypes will be deposited in the Colección Nacional de Insectos (Mexico City), Enns Entomology Museum (Columbia), and the United States National Museum of Natural History (Washington D. C.).	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B182143508EC09EF204FC76.taxon	materials_examined	Type material examined. HOLOTYPE ♀. MÉXICO: OAXACA: Mpio. Candelaria Loxicha, km 207 carr. Oaxaca-San Pedro Pochutla, Candelaria Loxicha, Río San Juan, 15 º 55 ’ 32.5 ’’ N, 96 º 29 ’ 12.6 ’’ W, 400 m, L- 1904, 02 - Apr- 2015, DRV, Sites, Shepard, Barr & PRH colls. PARATYPES: same data as holotype (4 ♂, 1 ♀ IEXA); same but 25 - May- 2012, L- 1375, DRV & PRH colls. (1 ♂, 1 ♀ UMC); Mpio. Candelaria Loxicha, Candelaria Loxicha, Río San Juan at Pte. San Juan, 15 º 55 ’ 32.9 ’’ N, 96 º 29 ’ 25 ’’ W, 403 m, 26 - May- 2012, L- 1376, DRV & PRH colls. (2 ♀ UMC); same but 02 - Apr- 2015, L- 1905, DRV, Sites, Shepard, Barr & PRH colls. (1 ♂ IEXA); Mpio. Mártires de Tacubaya, Mártires de Tacubaya, 16 º 34 ’ 41.1 ’’ N, 98 º 13 ’ 08.7 ’’ W, 206 m, L- 1390, 29 - May- 2012, DRV & PRH colls. (1 ♂, 1 ♀ IEXA); Mpio. San Gabriel Mixtepec, km 194 carr. Puerto Escondido-Oaxaca, San Gabriel Mixtepec, Río Rana, 16 º 06 ’ 06.3 ’’ N, 97 º 03 ’ 52.9 ’’ W, 712 m, L- 1380, DRV & PRH colls. (1 ♂, 1 ♀ UMC); same but 03 - Apr- 2015, L- 1906, DRV, Sites, Shepard, Barr & PRH colls. (1 ♂, 1 ♀ IEXA, 3 ♂, 7 ♀ UMC); Mpio. San Miguel Tlacamama, San Miguel Tlacamama, Pte. Yutaniñe, 16 º 25 ’ 32.6 ’’ N, 98 º 03 ’ 29.2 ’’ W, 260 m, L- 1385, 28 - May- 2012, DRV & PRH colls. (1 ♂ UMC); 29 mi W [Magdalena] Tequisistlán, CL 1064, 30 Apr 1964, JT & MS Polhemus / C. J. Drake Accession (1 ♂ USNM). CHIAPAS: Mpio. La Concordia, Km 17 carr. Jaltenango-Rev. Mexicana, 15 º 58 ’ 12.6 ’’ N 92 º 48 ’ 18.7 ’’ W, 562 m, L- 1316, 07 - Jan- 2012, submerged veg. DRV coll. (1 ♀ UMC); Mpio. Tonalá, Velo de Novia Waterfall, 15 º 55 ’ 11.3 ’’ N, 93 º 29 ’ 45.9 ’’ W, 144 m, nr. Pte. Las Hermanas, 21 - May- 2012, L- 1359, DRV & PRH colls. (1 ♀ UMC); same but 01 - Apr- 2015, L- 1903, DRV, Sites, Shepard, Barr & PRH colls. (3 ♂, 3 ♀ CNIN; 6 ♂, 4 ♀ IEXA; 5 ♂, 7 ♀ UMC). GUERRERO: [Mpio. Chilpancingo de Los Bravo], Agua de Obispo, 8 - 37, P Roveglia / Colección del Instituto de Biología, UNAM. México D. F. (1 ♀ CNIN).	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B142146508EC5B8F390FC3E.taxon	description	(Figs. 3, 6 – 7) Description. Macropterous female. HOLOTYPE, length 13.87; maximum width 8.30. Paratypes (n = 9), length 12.48 – 14.04 (mean = 13.12); maximum width 7.84 – 8.48 (mean = 8.10). General shape ovate; widest across embolia (Fig. 6 A). Overall dorsal coloration of hemelytra brown to dark brown with light brown marks; head light brown with dark brown marks; pronotum brown with dark brown marks; legs brown. Dorsal surface coarsely punctate. Ventral coloration of head and thorax brown, abdomen with yellowish brown pubescence. Head. Head length 2.04; maximum width 3.92. Mostly brown with dark brown marks along midline and posteriorly; coarsely punctate. Eyes convergent anteriorly, synthlipsis 1.80; thin band of cuticle along posterolateral margin of eye; eyes not raised above level of vertex or pronotum. Anterior margin between eyes slightly convex, extending anteriorly in front of eyes 10.7 % of head length; posterior margin between eyes strongly convex, extending posteriorly 30.6 % of head length. Labrum width 2.1 × length, evenly rounded. Labium with three visible yellowish brown segments, darkening distally, extending 0.75 beyond labrum not including extruded stylets. Antennal proportions 5: 17: 22: 11, length 1.37, extending to lateral margin of eye, elongate hairs on segments 3 and 4. Thorax. Pronotum coarsely punctate, ground color brown, dark brown marking posteromedially, dark brown longitudinal line behind eye; transverse sulcus marking anterior border of transverse band in posterior 1 / 3; transverse band light brown anteriorly, brown posteriorly; lateral margins brown, convergent, evenly convex, explanate; posterior margin almost straight; anterior margin deeply concave between eyes to embrace convex posterior margin of head; posterolateral corners rounded; width 2.4 × length; length at midline 2.96; maximum width at posterolateral corners 7.37. Prothorax ventrally pruinose throughout, except laterally; apices of propleura meeting at midline, not appressed to prosternellum; propleuron yellowish brown, brown posteriorly, medial 2 / 3 of posterior margin with golden setae. Probasisternum with sharp median carina and row of setae lateral to carina. Prosternellum not covered by apices of propleura. Scutellum coarsely punctate, triangular, dark brown, brown coloration on anterior and posterior corners and along midline, width 2.2 × length, width 5.00, length 2.24. Hemelytra densely punctate, brown to dark brown, length 10.0 (chord measurement); corium dark brown, faint brown ovoid and yellowish brown comma-shaped markings at lateral and posterior margins, respectively. Clavus dark brown, indistinct light brown marking at mid-length in mesal half, length 4.0 × width, length 5.45, width 1.36; claval commissure dark brown, length 1.27. Embolium length 4.20, greatest width 1.17; lateral margin convex, light brown in anterior 2 / 3, brown posteriorly. Membrane dark brown (Fig. 6 A). Hindwings extending to posterior margin of tergum VI. Mesobasisternum tumescence negligible, with sulcus on midline continuing through triangular mesosternellum. Metasternellum (= metaxyphus) transverse, subtriangular, with median carina. Legs. All leg segments yellowish brown to brown. Profemur yellowish brown; posterior margin with row of tightly arranged setae in basal 2 / 3, row of short brown spines along middle third; anterior margin with dense pad of setae without associated spines. Protibia and - tarsus with occlusal inner surface flattened and with spatulate setae; tarsus immovable, one-segmented; pretarsal claw single, minute, triangular. Procoxa with cluster of stout, brown anteromedial spines. Meso- and metacoxae partially recessed into thorax. Meso- and metafemora with row of short, brown spines on anterior margin; spines restricted to basal half on mesofemur, nearly full length of metafemur. Mesotibia with ventrolateral and dorsolateral rows of stout reddish-brown spines; mesolateral rows intermixed with combs of two to six spines. Metatibia with ventrolateral and dorsolateral rows of stout reddish-brown spines. Meso- and metatibiae with semi-circlet of spines at apices on dorsal and mesal margins; two comb rows of stout spines near apex on ventral margin. Meso- and metatibiae and metatarsus with long, yellow swimming hairs; hairs profuse on metatibia and - tarsus. Meso- and metapretarsi with paired claws slender, gently curved, with small basal tooth. Leg measurements as follows: foreleg, femur 3.25, tibia 2.54, tarsus 0.52; middle leg, femur 2.91, tibia 2.75, tarsomeres 1 – 3, 0.27, 0.60, 0.58; hind leg, femur 4.00, tibia 4.48, tarsomeres 1 – 3, 0.34, 1.05, 0.75. Abdomen. Dorsally with lateral margins of III – VI exposed, each dark brown in anterior 1 / 3 and light brown posteriorly; lateral margins of II – basal half of III smooth and distal half of III – VI finely serrate, marginal row of profuse and short yellow setae, group of trichobothria near posterolateral corners. Posterolateral corner of II (visible ventrally) narrowly rounded, right angled, III – VII acute. Ventrally yellowish brown, with dense pile of fine hairs. Lateral margin with thin, glabrous band. Glabrous rounded to oval patches near spiracles on laterosternites II – VII. Mediosternite VII (subgenital plate) asymmetrical, width 1.1 × length; length at midline 1.45; maximum width 1.69; with pointed lateral lobe at apex of left margin; posterolateral corners pointed, asymmetrical, right posterolateral corner wider, both corners posteriorly produced further than central lobe; wide central lobe notched giving the appearance of two rounded lobes (Fig. 6 B). Macropterous male. Paratypes (n = 10), length 11.68 – 12.64 (mean = 12.19); maximum width 7.36 – 7.84 (mean = 7.62). Coloration, setation, pronotum proportions, posterolateral corners of pronotum, scutellum proportions, embolium length generally same as for holotype. Otherwise hindwings extending to posterior margin of tergum V. Accessory genitalic process of tergum VI slender, gently curved to right in basal 2 / 3, curved to right at approximately 90 degree angle in apical 1 / 3, apex pointed (Fig. 6 C). Medial lobes of tergum VIII (pseudoparameres) symmetrical, posterolaterally produced, posterolateral corner narrowly rounded, extending posteriorly further than posteromesal corner (Fig. 6 D). Phallosoma elongate, linear, slender, slightly constricted basally; endosomal sclerites subtriangular, knob-like apically, with conspicuous denticles, right sclerite basally produced further than left sclerite (Fig. 6 E). Parameres symmetrical, almost as long as maximum width, mesal margin rounded, setae emanating from distal 1 / 3 of dorsal surface. Proctiger almost as long as width at base. Pygophore with elongate setae sparsely distributed over most of surface, with thick brush of elongate setae on posterior margin (Fig. 6 F).	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B142146508EC5B8F390FC3E.taxon	discussion	Discussion. The general morphology of the female SGP, the male AGP, and the endosomal sclerites exhibited by A. schuhi suggest a close relationship with A. cosmius La Rivers. Diagnostic features and comparative notes. Females have a diagnostic SGP that presents a pointed lateral lobe at the apex of the left margin, just basal to the left posterolateral corner. Also, the pointed posterolateral corners are asymmetrical with that on the right generally wider. A similar SGP is also present in A. cosmius, although the lateral lobe on the left margin is rounded and at mid-length and the posterolateral corners are symmetrical. Males of A. schuhi exhibit a characteristic AGP that is particularly long and pointed in the apical third, whereas A. cosmius presents a comparable AGP, but is longer and the apex is anterolaterally directed (laterally directed in A. schuhi).	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B142146508EC5B8F390FC3E.taxon	description	Habitat description. The type locality at El Rosario (L- 1394) is in the western area of the Mexican state of Oaxaca, at an elevation of 361 meters. The stream at the type locality is shallow and approximately 5 meters wide. The river bottom is mainly composed of gravel (Fig. 7). The microhabitat where we collected specimens of this species was among rocks and algae along the stream margins.	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B142146508EC5B8F390FC3E.taxon	distribution	Distribution. This species is present in the southern area of the Mexican states of Guerrero and Oaxaca (Fig. 3). Ambrysus schuhi is distributed in the Mexican Pacific Coast (Guerrero) and Sierra Madre del Sur (Oaxaca) biogeographic provinces. At its type locality (El Rosario, Oaxaca) this species was collected syntopically with A. baeus and A. spicatus. At Río Rana, in May 2012 (L- 1380), A. schuhi was found syntopically with A. baeus, A. pulchellus, and the new species A. henryi. Also, at Río Rana, but in April 2015 (L- 1906), it was collected with A. baeus, A. spicatus, and the new species A. henryi and A. sitesi. At the locality northeast of San Pedro Amuzgos (L- 1391) we found this species with A. baeus, A. pudicus, A. pulchellus, and A. spicatus. Additionally, at the locality south of El Tapanco (L- 1393) A. schuhi was collected with the congeners A. angularis and A. pulchellus, as well as with Cataractocoris marginiventris Usinger.	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B142146508EC5B8F390FC3E.taxon	etymology	Etymology. It is our pleasure to name this species in honor of our dear colleague Professor Randal T. Schuh, of the American Museum of Natural History, to commemorate the astounding work he has conducted on the group of “ True Bugs. ” Recently, during the VI Quadrennial Meeting of the International Heteropterists’ Society (La Plata, Argentina), Dr. Schuh was awarded with the newly institutionalized Carl Stål Medal, to recognize his accomplishments on the study of Heteroptera, which include the description of approximately 100 genera and 600 species of insects. Repositories. The holotype and some paratypes will be deposited in the Colección Entomológica del Instituto de Ecología A. C. (Xalapa); additional paratypes will be deposited in the Colección Nacional de Insectos (Mexico City), Enns Entomology Museum (Columbia), University of Michigan Museum of Zoology (Ann Arbor), and the United States National Museum of Natural History (Washington D. C.).	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B142146508EC5B8F390FC3E.taxon	materials_examined	Type material examined. HOLOTYPE ♀. MÉXICO: OAXACA: Mpio. Sta. Maria Zacatepec, km 179 carr. Sta. Maria Zacatepec-Putla, El Rosario, Pte. El Rosario, 29 - May- 2012, L- 1394 / 361 m, 16 º 51 ’ 34.6 ’’ N, 97 º 58 ’ 44.7 ’’ W, DRV & PRH colls. PARATYPES: same data as holotype (3 ♂ IEXA; 6 ♂, 6 ♀ UMC); Mpio. Mesones Hidalgo, km 173 carr. Sta. Maria Zacatepec-Putla, Pte. La Junta, 29 - May- 2012, L- 1395 / 396 m, 16 º 53 ’ 59.7 ’’ N 97 º 58 ’ 02 ’’ W, DRV & PRH (1 ♂ UMC); Mpio. San Gabriel Mixtepec, Río Rana, km 194 carr. Puerto Escondido- Oaxaca, San Gabriel Mixtepec, L- 1380 / 27 - May- 2012, 712 m, 16 º 06 ’ 06.3 ’’ N 97 º 03 ’ 52.9 ’’ W, DRV & PRH colls. (1 ♂, 1 ♀ UMC); same but 3 - Apr- 2015, L- 1906, DRV, Sites, Shepard, Barr & PRH colls., rocks & gravel (5 ♂, 1 ♀ IEXA; 3 ♂, 1 ♀ UMC); [Mpio. San Gabriel Mixtepec], Sn. Gabriel Mixtepec, 20 - V- 84, M García / Colección del Instituto de Biología, UNAM. México D. F. (1 ♂ CNIN; 1 ♂, 1 ♀ USNM); Mpio. Sta. Maria Zacatepec, km 189 carr. Sta. Maria Zacatepec-Putla, S of El Tapanco, L- 1393 / 29 - May- 2012, 341 m, 16 º 47 ’ 28.3 ’’ N 97 º 59 ’ 20.4 ’’ W, DRV & PRH (2 ♂ UMC); Mpio. Sta Maria Zacatepec, Km 201 carr. Mártires de Tacubaya-Santa Maria Zacatepec ca. 13 km NE of San Pedro Amuzgos, L- 1391 / 29 - May- 2012, 276 m, 16 º 42 ’ 18.9 ’’ N 98 º 00 ’ 52.9 ’’ W, DRV & PRH (1 ♂ UMC). GUERRERO: stream tributary to Laguna Tres Palos, about 22 miles from Acapulco on highway to Mexico City, III: 22: 1951 RR & M Miller (2 ♂, 1 ♀ UMMZ).	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B10215B508EC6D9F56DFE26.taxon	description	(Figs. 3, 8 – 9) Description. Macropterous female. HOLOTYPE, length 11.87; maximum width 7.25. Paratypes (n = 10), length 10.70 – 11.60 (mean = 11.31); maximum width 6.80 – 7.50 (mean = 7.17). General shape ovate; widest across embolia (Fig. 8 A). Overall dorsal coloration of hemelytra brown to dark brown with yellowish brown marks; head and pronotum light brown with dark brown marks; legs yellowish brown. Dorsal surface coarsely punctate. Ventral coloration of head and thorax yellowish, abdomen with yellowish pubescence. Head. Head length 1.60; maximum width 3.20. Mostly light brown with dark brown marks along midline and posteriorly; coarsely punctate. Eyes convergent anteriorly, synthlipsis 1.48; thin band of cuticle along posterolateral margin of eye; eyes not raised above level of vertex or pronotum. Anterior margin between eyes slightly convex, extending anteriorly in front of eyes 7.8 % of head length; posterior margin between eyes strongly convex, extending posteriorly 29.7 % of head length. Labrum width 1.7 × length, evenly rounded. Labium with three visible yellowish brown segments, darkening distally, extending 0.49 beyond labrum not including extruded stylets. Antennal proportions 3: 14: 22: 9, length 0.72, extending to lateral margin of eye, elongate hairs on segment 4 and distal half of 3. Thorax. Pronotum coarsely punctate, ground color medium brown, profuse dark brown markings, dark brown longitudinal line behind eye; transverse sulcus marking anterior border of transverse band in posterior 1 / 3; transverse band light brown anteriorly, brown posteriorly; lateral margins yellowish brown, convergent, evenly convex, explanate; posterior margin almost straight; anterior margin deeply concave between eyes to embrace convex posterior margin of head; posterolateral corners rounded; width 2.7 × length; length at midline 2.28; maximum width at posterolateral corners 6.26. Prothorax ventrally pruinose throughout, except laterally; apices of propleura meeting at midline, not appressed to prosternellum; propleuron yellowish, light brown posteriorly, medial 2 / 3 of posterior margin with golden setae. Probasisternum with sharp median carina and row of setae lateral to carina. Prosternellum not covered by apices of propleura. Scutellum coarsely punctate, triangular, dark brown, light brown coloration at anterior corners and along midline, width 2.1 × length, width 4.32, length 2.00. Hemelytra densely punctate, brown to dark brown, length 8.12 (chord measurement); corium dark brown, yellowish brown ovate and yellowish commashaped marks at lateral and posterior margins, respectively. Clavus dark brown in lateral half, brown in mesal half with dark brown close to base and apex, length 3.8 × width, length 4.56, width 1.20; claval commissure dark brown, length 1.30. Embolium length 3.32, greatest width 1.05; lateral margin convex, yellowish in anterior 2 / 3, brown posteriorly. Membrane dark brown (Fig. 8 A). Hindwings extending to middle of tergum VII. Mesobasisternum tumescence negligible, with sulcus on midline continuing through triangular mesosternellum. Metasternellum (= metaxyphus) transverse, subtriangular, with median carina. Legs. All leg segments yellowish brown. Profemur yellowish brown; posterior margin with row of tightly arranged setae in basal 2 / 3, row of short brown spines along middle third; anterior margin with dense pad of setae without associated spines. Protibia and - tarsus with occlusal inner surface flattened and with spatulate setae; tarsus immovable, one-segmented; pretarsal claw single, minute, triangular. Procoxa with cluster of stout, brown anteromedial spines. Meso- and metacoxae partially recessed into thorax. Meso- and metafemora with row of short, light brown spines on anterior margin; spines restricted to basal half on mesofemur, nearly full length of metafemur. Mesotibia with ventrolateral and dorsolateral rows of stout reddish-brown spines; mesolateral rows intermixed with combs of two to six spines. Metatibia with ventrolateral and dorsolateral rows of stout reddish-brown spines. Meso- and metatibiae with semi-circlet of spines at apices on dorsal and mesal margins; two comb rows of stout spines near apex of ventral margin. Meso- and metatibiae and metatarsus with long, yellow swimming hairs; hairs profuse on metatibia and - tarsus. Meso- and metapretarsi with paired claws slender, gently curved, with small basal tooth. Leg measurements as follows: foreleg, femur 2.60, tibia 2.02, tarsus 0.47; middle leg, femur 2.55, tibia 2.30, tarsomeres 1 – 3, 0.22, 0.49, 0.52; hind leg, femur 3.34, tibia 3.72, tarsomeres 1 – 3, 0.28, 0.88, 0.67. Abdomen. Dorsally with lateral margins of III – VI exposed, each brown in anterior 1 / 3 and light brown posteriorly; lateral margins of II to basal half of III smooth and distal half of III to VI finely serrate, marginal row of profuse and short yellow setae, group of trichobothria near posterolateral corners. Posterolateral corner of II (visible ventrally) narrowly rounded, right angled, III – VII acute. Ventrally yellowish, with dense pile of fine hairs. Lateral margin with thin, glabrous band. Glabrous rounded to oval patches near spiracles on laterosternites II – VI. Mediosternite VII (subgenital plate) asymmetrical, width 1.1 × length; length at midline 1.24; maximum width 1.38; lateral lobe at mid-length of left margin broadly rounded; posterolateral corners pointed; wide central lobe produced posteriorly slightly further than posterolateral corners (Fig. 8 B). Macropterous male. Paratypes (n = 10), length 9.80 – 11.00 (mean = 10.52); maximum width 6.20 – 7.00 (mean = 6.62). Coloration, setation, pronotum proportions, posterolateral corners of pronotum, scutellum proportions, embolium length generally same as for holotype. Otherwise hindwings extending to posterior margin of tergum V. Accessory genitalic process of tergum VI slender, gently curved to right close to the base, widening distally, apex pointed (Fig. 8 C). Medial lobes of tergum VIII (pseudoparameres) symmetrical, posterolaterally produced, posterolateral corner narrowly rounded, extending posteriorly further than posteromesal corner (Fig. 8 D). Phallosoma elongate, linear, slender, slightly constricted basally; left endosomal sclerite reduced, with small and inconspicuous denticles; right endosomal sclerite absent (Fig. 8 E). Parameres symmetrical, almost as long as maximum width, mesal margin rounded, setae emanating from distal 1 / 3 of dorsal surface. Proctiger slightly shorter than width at base. Pygophore with elongate setae sparsely distributed over most of surface, with thick brush of elongate setae on posterior margin (Fig. 8 F).	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B10215B508EC6D9F56DFE26.taxon	discussion	Discussion. The conspicuous dorsal pigmentation of specimens of A. sitesi is similar to that of A. lunatus Usinger and A. signoreti Stål. The morphology of the endosomal sclerites of the males suggests that this species could be closely related to A. lundbladi Usinger. In his work on the Ambrysus of Mexico, La Rivers (1953) described the subspecies A. lunatus menoides from the central state of Puebla. He cited paratype specimens from the state of Chiapas (Ixtapa) that were deposited in the UMMZ. We examined a female paratype that was deposited in the CAS and found that the morphology of the SGP was different to that reported for the subspecies. Instead, the morphology corresponded to the herein described A. sitesi. Based on this and our extensive collecting in the area, we consider the subspecies A. l. menoides not to be distributed in the state of Chiapas. Diagnostic features and comparative notes. Females of this species exhibit a characteristic SGP with a broadly rounded lateral lobe at mid-length of the left margin. Other species in the A. signoreti complex present a lateral lobe on the left margin of the SGP (sometimes also on the right margin), but in general the lobe is low and pointed (e. g., A. colimanus, A. lunatus). The species A. scalenus presents a similar left-lobed margin of the SGP (examined on the allotype in the SEMC); however, the lobe extends further laterally and appears truncate. Males of A. sitesi present the phallosoma with no right endosomal sclerite and a reduced left endosomal sclerite with inconspicuous rounded denticles along the mesal margin. To our knowledge, the rare A. lundbladi is the only other species from this species complex that exhibits the phallosoma with an absent right endosomal sclerite and the left sclerite reduced.	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B10215B508EC6D9F56DFE26.taxon	description	Habitat description. The type locality at Río Rana (L- 1906), close to the town of San Gabriel Mixtepec, is in the south central area of the Mexican state of Oaxaca, on the Sierra Madre del Sur mountain range, at an elevation of 712 meters. At the type locality the stream is 5 – 10 meters wide, with fast current, and boulders and cobble (Fig. 9). Specimens were collected by disturbing the substratum with the net along the stream margin.	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B10215B508EC6D9F56DFE26.taxon	distribution	Distribution. Ambrysus sitesi is distributed in the highlands of the Mexican states of Chiapas and Oaxaca and the western area of Guatemala, in the Huehuetenango Department (Fig. 3). The records from Oaxaca are in the area of the Sierra Madre del Sur biogeographic province, while those from Chiapas and Huehuetenango are in the Chiapas biogeographic province. One record from Chiapas (L- 1899) is in the Mexican Pacific Coast biogeographic province. This species was collected syntopically with various congeners, including A. circumcinctus (L- 1339, L- 1341, L- 1350, L 1884, L 1897), A. circumcinctus and A. pudicus (L- 1881, L- 1898), A. circumcinctus and A. spicatus (L- 1349), A. circumcinctus, A. parviceps, and A. pudicus (L- 1899); A. chiapanecus Reynoso, Sites & Novelo (L- 1340, L- 1886); A. chiapanecus and A. spicatus (L- 1885); A. mexicanus Montandon, A. pygmaeus La Rivers, and A. spicatus (L- 1397). At its type locality (L- 1906) this species was found syntopically with A. baeus, A. henryi n. sp., A. schuhi n. sp., and A. spicatus. Additionally, A. sitesi was collected with A. brailovskyi n. sp. at Finca Prusia (12 - V- 1985), Río Matasano (L- 1348), Río Prusia (L- 1345), and the Tenejapa municipality (14 - VII- 1964).	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B10215B508EC6D9F56DFE26.taxon	etymology	Etymology. It is my (DRV) pleasure to name this species in honor of Professor Robert. W. Sites, my former doctoral advisor at the University of Missouri and coauthor of this manuscript. I am grateful for his guidance, help, and friendship throughout these years. Repositories. The holotype and some paratypes will be deposited in the Colección Entomológica del Instituto de Ecología A. C. (Xalapa); additional paratypes will be deposited in the California Academy of Sciences (San Francisco), Colección Nacional de Insectos (Mexico City), Enns Entomology Museum (Columbia), Snow Entomological Museum Collection (Lawrence), and the United States National Museum of Natural History (Washington D. C.)	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
244E8D140B10215B508EC6D9F56DFE26.taxon	materials_examined	Type material examined. HOLOTYPE ♀. MÉXICO: OAXACA: Mpio. San Gabriel Mixtepec, km 194 carr. Puerto Escondido-Oaxaca, San Gabriel Mixtepec, Río Rana, 16 º 06 ’ 06.3 ’’ N, 97 º 03 ’ 52.9 ’’ W, 712 m, 03 - Apr- 2015, L- 1906, DRV, Sites, Shepard, Barr & PRH colls. PARATYPES: same data as holotype (2 ♂, 2 ♀ CNIN; 10 ♂, 7 ♀ IEXA; 5 ♂, 4 ♀ UMC); Mpio. Jalapa de Díaz, Sto. Domingo, Río Sto. Domingo, 18 ° 02 ’ 15.9 ’’ N, 96 ° 32 ’ 57.2 ’’ W, 100 m, L- 1881, 25 - Mar- 2015, DRV, Sites, Shepard, Barr & PRH colls. (2 ♂, 6 ♀ IEXA; 1 ♂, 1 ♀ UMC); Mpio. Putla Villa de Guerrero, Pluta de Guerrero, 17 º 02 ’ 21.5 ’’ N, 97 º 54 ’ 42.9 ’’ W, 767 m, 30 - May- 2012, L- 1397, DRV & PRH colls. (1 ♂ UMC); Mpio. Sta. Maria Tecomavaca, carr. Tehuacán-San Francisco Telixtlahuaca, 16 km S of San Juan de los Cués, Río Salado, 17 º 55 ’ 50.5 ’’ N, 97 º 00 ’ 04.2 ’’ W, 544 m, 01 - Jun- 2012, L- 1407, DRV & PRH colls. (1 ♂ UMC). CHIAPAS: Mpio. Altamirano, Cuauhtemoc, Balneario Tzaconejá, Río Tzaconejá, 16 ° 41 ’ 17.5 ’’ N, 92 ° 01 ’ 03.0 ’’ W, 1172 m, 28 - Mar- 2015, L- 1886, DRV, Sites, Shepard, Barr & PRH colls. (6 ♂, 3 ♀ IEXA; 4 ♂ UMC); Mpio. Am- atengo de la Frontera, Barrio Sabinalito, Río Agua Caliente, 15 ° 25 ’ 0.1 ’’ N, 92 ° 08 ’ 33.7 ’’ W, 942 m, L- 1898, 31 - Mar- 2015, DRV, Sites, Shepard, Barr & PRH colls. (4 ♂ IEXA); [Mpio. Ángel Albino Corzo], Finca Prusia, 10 - V- 85, M. Vértiz / Colección del Instituto de Biología, UNAM. México, D. F (1 ♂ CNIN); [Mpio. Ángel Albino Corzo], Finca Prusia, 12 - V- 85, F Arias / Colección del Instituto de Biología, UNAM. México, D. F (1 ♂, 2 ♀ CNIN); [Mpio. Ángel Albino Corzo], Finca Prusia, 12 - V- 85, H Velasco / Colección del Instituto de Biología, UNAM. México, D. F (3 ♂ CNIN); [Mpio. Ángel Albino Corzo], Finca Prusia, 12 - V- 85, M Vértiz / Colección del Instituto de Biología, UNAM. México, D. F (1 ♀ CNIN); Mpio. Ángel Albino Corzo, 1 km S of Finca Prusia, Río Prusia, 15 º 43 ’ 23.3 ’’ N, 92 º 47 ’ 46.5 ’’ W, 1010 m, 18 - May- 2012, L- 1345, DRV & PRH colls. (1 ♂, 4 ♀ UMC); Mpio. Frontera Comalapa, El Caracol, 8 km S of Frontera Comalapa, Río Grijalva, 15 ° 36 ’ 07.0 ’’ N, 92 ° 09 ’ 14.5 ’’ W, 677 m, 31 - Mar- 2015, L- 1897, DRV, Sites, Shepard, Barr & PRH colls. (1 ♂ IEXA); Mpio. Huixtla, Huixtla, Río Huixtla, 15 ° 08 ’ 59.9 ’’ N, 92 ° 27 ’ 04.6 ’’ W, 67 m, 31 - Mar- 2015, L- 1899, DRV, Sites, Shepard, Barr & PRH colls. (1 ♂ IEXA); Mpio. Ixtapa, Ixtapa, 16 º 48 ’ 43.7 ’’ N, 92 º 54 ’ 30 ’’ W, 1030 m, 17 - May- 2012, L- 1341, DRV & PRH colls. (6 ♂, 3 ♀ IEXA); [Mpio. Ixtapa], Ixtapa, # 132, IV- 27 - 1941, IJ Cantrall / Ambrysus lunatus menoides La Rivers PARATYPE / Ira La Rivers Collection Bequeathed to the California Academy of Sciences- 1978 (1 ♀ CAS); Mpio. Ixtapa, Río Lajas at Pte. Río Lajas, 16 º 50 ’ 42.6 ’’ N, 92 º 54 ’ 01.7 ’’ W, 1020 m, 17 - May- 2012, L- 1340, DRV & PRH colls. (2 ♂, 1 ♀ UMC); same but 28 - Mar- 2015, L- 1885, DRV, Sites, Shepard, Barr & PRH colls. (10 ♂, 10 ♀ CNIN; 20 ♂, 21 ♀ IEXA; 6 ♂, 15 ♀ UMC); [Mpio. Ixhuatán], 4 mi N of Tapiula [Tapilula], CL 1097, 5 May 1964, JT & MS Polhemus (1 ♂, 1 ♀ USNM); [Mpio. Ocosingo], Ocosingo, June 25 1950, C & M Goodnight & L Stannard, mountain stream / Chiapas VI- 25 - 57, Mex. Stannard (1 ♂ SEMC); Mpio. Siltepec, km 72 carr. Jaltenango-Siltepec, nr. Nuevo Galeana, 15 º 35 ’ 54.9 ’’ N, 92 º 32 ’ 28.9 ’’ W, 933 m, L- 1349, 19 - May- 2012, DRV & PRH colls. (2 ♂, 1 ♀ UMC); Mpio. Siltepec, km 92 carr. Jaltenango-Siltepec, Campo Aéreo, Río Bravo, 15 º 35 ’ 17.9 ’’ N, 92 º 28 ’ 02.3 ’’ W, 1068 m, 19 - May- 2012, L- 1350, DRV & PRH colls. (1 ♂, 1 ♀ UMC); Mpio. Siltepec, Matasano, Río Matasano, 15 º 36 ’ 13.7 ’’ N, 92 º 33 ’ 37.1 ’’ W, 950 m, 19 - May- 2012, L- 1348, DRV & PRH colls. (6 ♂, 6 ♀ IEXA; 16 ♂, 19 ♀ UMC); Mpio. Solosuchiapa, El Beneficio, Río La Sierra, 17 ° 21 ’ 01.1 ’’ N, 93 ° 00 ’ 21.1 ’’ W, 303 m, 27 - Mar- 2015, L- 1884, DRV, Sites, Shepard, Barr & PRH colls. (1 ♂ UMC); Mpio. Solosuchiapa, Solosuchiapa, Río La Sierra, 17 º 25 ’ 40.5 ’’ N, 93 º 01 ’ 35.7 ’’ W, 122 m, 17 - May- 2012, L- 1339, DRV & PRH colls. (1 ♂ UMC); [Mpio. Soyaló], Soyalo, Rio Blanco, CL 1086, 4 May 1964, JT & MS Polhemus / C. J. Drake Accession (16 ♂, 10 ♀ USNM); Mpio. Tenejapa, ponds in stream above Habenal / 14. VII. 1964, coll. D Breedlove / C. J. Drake Accession / CAS (2 ♂ USNM); Rio Laja [s] nr Bochil, CL 668, I- 6 - 74, JT Polhemus (9 ♂, 5 ♀ USNM); Soyalo, Rio Lajas, CL 1087, 4 May 1964, JT & MS Polhemus / A. guttatipennis close, No! / C. J. Drake Accession (1 ♀ USNM). GUATEMALA: HUEHUETENANGO: Rio Nentón, ca. 8 km NE of La Laguna, 10 - May- 2013, 15.79915 º N, 91.88951 ºW, 613 m, RS Zack coll. (1 ♀ WSUC).	en	Reynoso-Velasco, Daniel, Sites, Robert W. (2019): Descriptions of four new species of the Ambrysus signoreti Stål species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa 4615 (2): 285-302, DOI: 10.11646/zootaxa.4615.2.3
