taxonID	type	description	language	source
90E568FB8FB5581081B30227A31044BC.taxon	materials_examined	Material examined. In addition to the type series, we examined one male (15 - 313) collected on 17 August 2015 by X. Freilich, J. Reyes-Velasco and S. Boissinot southeast of Debre Markos (10.2745 ° N, 37.8564 ° E, 2261 m a. s. l.), one female (15 - 367) collected on 19 August 2015 by X. Freilich J. Reyes-Velasco and S. Boissinot southwest of Debarq (13.0359 ° N, 37.799 ° E, 2583 m a. s. l.), one female (16 - 175) collected on 15 July 2016 by X. Freilich, J. Reyes-Velasco and S. Boissinot in Debre Markos (10.35195 ° N, 37.7414 ° E, 2803 m a. s. l.), one female (SB 580) collected on 8 July 2018 by S. Goutte and Y. Bourgeois north-west of Debre Markos (10.3566 ° N, 37.6571 ° E, 2589 m a. s. l.), one female (SB 584) collected on 8 July 2018 by S. Goutte and Y. Bourgeois in Debre Markos (10.35195 ° N, 37.7414 ° E, 2388 m a. s. l.) and five males (SB 591, SB 592, SB 593, SB 597 and SB 606) collected on 9 July 2018 by S. Goutte and Y. Bourgeois in Debre Markos (10.3520 ° N, 37.7414 ° E, 2388 m a. s. l.).	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
90E568FB8FB5581081B30227A31044BC.taxon	diagnosis	Diagnosis. Medium-sized species (male (6) SVL 36.9 + / - 2.7 mm, female (7) SVL 42.5 + / - 2.0 mm) of the Ptychadena cooperi species group (Fig. 9). It differs from other members of the Ptychadena neumanni species complex by the following combination of characters: (1) tibia half of the snout-vent length, (2) eye close to one another (male IOD / HW 0.16 + / - 0.02, female IOD / HW 0.17 + / - 0.02), (3) vertical light stripe on the tympanum, (4) vocal sacs are light grey to cream, sometimes mottled with light grey, (5) adult males' bodies covered in small warts.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
90E568FB8FB5581081B30227A31044BC.taxon	description	Description of the holotype. Juvenile, in poor condition (Fig. 10). Right hind limb missing. No visible ridges or coloration on the back due to the poor condition of the specimen. Finger formula: 1 <2 <4 <3, hand free of webbing. Toe formula: 1 <2 <5 <3 <4. Toe webbing indistinguishable. Tongue bifurcated and free for half is length. Maxillary and premaxillary teeth present, vomerine teeth not visible. Throat cream. Measurements of the holotype (mm): SVL 21.4, HW 8.8, HL 9.3, SL 3.7, NS 2.1, IND 2.1, EN 1.8, IOD 1.6, ETD 0.9, TD 1.6, ED 2, UEW 1.1, FLL 3.9, HAL 4.9, F 4 DW 0.3, THL 9.6, TL 9.7, FL 10.1, T 4 DW 0.3, MTL 0.9.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
BE21FD2A9D415C909D56E7446862297C.taxon	diagnosis	Diagnosis. A medium-sized species (male (9) SVL 37.9 + / - 2.6 mm, female (8) SVL 43.2 + / - 2.9 mm) of the Ptychadena neumanni species group (Fig. 18), distinguished by the following combination of characters: (1) relatively short hind limbs (male TL / SVL 0.51 + / - 0.03, female TL / SVL 0.57 + / - 0.03), (2) tympanum with a light vertical bar, (3) medium or wide vertebral stripe always present, (4) vocal sacs of most males are bicolored, from light to dark grey posteriorly and from yellow to cream anteriorly, rarely, they are light grey, (5) forearms of adult males not thickened.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
BE21FD2A9D415C909D56E7446862297C.taxon	description	Description of the holotype. A medium-sized (SVL 36.3 mm) adult male (Fig. 18). Head slightly wider than long. Snout projecting beyond the lower jaw. Interorbital distance 0.62 x the eye diameter. Internarial distance 1.12 x interorbital distance. Tympanum 0.67 x eye diameter. Finger tips not expanded but rounded, with very small subarticular tubercles. Finger formula: I <II <IV <III. Hand free of webbing, palmar tubercle absent. Nuptial pads light grey. Hindlimbs moderately elongated, with tibia length 0.51 x snout-vent length. Foot as long as thigh and shorter than tibia (FL / TL 0.97). Toe tips rounded. Subarticular tubercles extremely small. Inner metatarsal tubercle present, external absent. Toe formula: I <II <V <III <IV. Foot webbing formula: Ie (1), IIi / e (1 - 1), IIIi / e (1 - 2), IVi / e (2 - 2), Vi (2). Two light brown, continuous lateral ridges, two continuous and four interrupted dorsal ridges. No vertebral nor sacral ridges. No warts on the body. Small and round light brown warts on tibias and thighs.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
BE21FD2A9D415C909D56E7446862297C.taxon	etymology	Etymology. The specific name corresponds to the translation of the word " enough " or " that's it " in Amharic (በቃ), in reference to the controversial taxonomic history of the group that we hope has come to an end. It is an invariable noun used in apposition.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
C980B0B63C5C503AA82FC846A6ECF097.taxon	diagnosis	Diagnosis. Large member (male (6) SVL 40.1 + / - 2.9 mm, female (6) SVL 47.2 + / - 2.3 mm) of the Ptychadena neumanni species group (Fig. 19) distinguished by the following combination of characters: (1) moderately long hind limbs (male TL / SVL 0.53 + / - 0.02, female TL / SVL 0.55 + / - 0.03), (2) long forearms (FLL / SVL 0.20 + / - 0.01), (3) eye close to one another (male IOD / HW 0.17 + / - 0.02, female IOD / HW 0.20 + / - 0.03), (4) light vertical stripe on the tympanum, (5) vocal sacs are dark grey or dark grey posteriorly and lighter anteriorly.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
C980B0B63C5C503AA82FC846A6ECF097.taxon	description	Description of the holotype. Relatively large (SVL 44.2 mm) male (Fig. 19). Head wider than long (HL / HW 0.87). Snout projecting beyond the lower jaw. Interorbital distance 0.54 x eye diameter. Internarial distance 1.68 x interorbital distance. Tympanum 0.71 x eye diameter. Finger tips not expanded but rounded, with small subarticular tubercles. Finger formula: I <II <IV <III. Hand free of webbing. Hindlimbs elongated, with tibia length 0.55 x snout-vent length. Foot longer than thigh (FL / THL 1.27) and slightly longer than tibia (FL / TL 1.02). Toe tips rounded. Subarticular tubercles small and round. Inner metatarsal tubercle present, external absent. Fourth toe on the left foot amputated. Toe formula: I <II <V <III <IV. Foot webbing formula: Ie (minimal), IIi / e (minimal- 1), IIIi / e (2 - 2), IVi / e (2 - 2), Vi (2). Two light lateral ridges, continuous on the right side and discontinuous on the left side. Two continuous dorsal ridges and two interrupted dorsolateral ridges. No vertebral nor sacral ridges. Body and eyelids covered with minute transparent warts. Nuptial pad developed along finger I. Tongue longer than wide, free for half of its length, bifurcated at the end. Vomerine, maxillary and premaxillary teeth present.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
C980B0B63C5C503AA82FC846A6ECF097.taxon	etymology	Etymology. The specific name originates from the Latin delphinus, dolphin, in reference to the advertisement call of the species resembling a dolphin's clicking sound. We have Latinized delphinus into the adjective Ptychadena delphina to be in accordance with the gender of genus Ptychadena. The advertisement call best distinguishes P. delphina sp. nov. from P. doro sp. nov.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
0705404D7B4056A188565655B54204C3.taxon	diagnosis	Diagnosis. Moderately large (male (13) SVL 35.8 + / - 2.2 mm, female (6) SVL 42.4 + / - 5.7 mm) member of the Ptychadena neumanni species group (Fig. 20) distinguished by the following combination of characters: (1) long hind limbs (male TL / SVL 0.58 + / - 0.03, female TL / SVL 0.64 + / - 0.11), (2) long hands (male HAL / SVL 0.24 + / - 0.02, female HAL / SVL 0.25 + / - 0.05), (3) long snout (male SL / SVL 0.16 + / - 0.01, female SL / SVL 0.16 + / - 0.02), (4) relatively small tympanum (male TD / ED 0.64 + / - 0.09, female TD / ED 0.63 + / - 0.1), (5) light vertical stripe on the tympanum, (6) vocal sacs are grey or dark grey posteriorly and lighter anteriorly,	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
0705404D7B4056A188565655B54204C3.taxon	description	Description of the holotype. A medium-sized (SVL 36.4 mm) adult male with long hind limbs (TL / SVL 0.58; Fig. 20, Suppl. material 3: Table S 1). Head as long as wide. Snout projecting beyond the lower jaw. Interorbital distance 0.60 x the eye diameter. Internarial distance 1.08 x interorbital distance. Tympanum 0.63 x eye diameter. Finger tips not expanded but rounded, with very small subarticular tubercles. Finger formula: I <II <IV <III. Hand free of webbing, palmar tubercle absent. Nuptial pads light grey to cream, extending dorsally to the second finger. Hindlimbs elongated, with tibia length 0.57 x snout-vent length. Foot longer than thigh and slightly longer than tibia (FL / THL 1.20, FL / TL 1.01). Toe tips rounded. Subarticular tubercles extremely small. Inner metatarsal tubercle present, external absent. Toe formula: I <II <V <III <IV. Foot webbing formula: Ie (1), IIi / e (1 - 1), IIIi / e (1 - 2), IVi / e (2 - 2), Vi (2). Two light brown, continuous lateral ridges, two continuous and four interrupted dorsal ridges. No vertebral nor sacral ridges. Very small, translucent warts on the body and small round warts on tibias and feet. Tongue free for half of its length, bifurcated at the end. Vomerine, maxillary and premaxillary teeth present.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
0705404D7B4056A188565655B54204C3.taxon	etymology	Etymology. The species name is the Amharic translation of chicken (ዶሮ), in reference to the advertisement call of the species resembling a chicken's song. The advertisement call distinguishes best Ptychadena doro sp. nov. from the syntopic species P. delphina sp. nov.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
87B773BC7FF35B5B818DA5DB58E809FC.taxon	materials_examined	Material examined. Except for the type specimen and the type series of P. largeni, all the material examined is deposited at ZNHM. One female (15 - 47) collected by X. Freilich, J. Reyes-Velasco and S. Boissinot south of Assela (7.9068 ° N, 39.1238 ° E, 2520 m a. s. l.), one male (15 - 400) collected by X. Freilich, J. Reyes-Velasco and S. Boissinot on 22 August 2015 north of Fitche (9.7877 ° N, 38.6974 ° E, 2821 m a. s. l.), two males (15 - 417 and 15 - 420) collected by X. Freilich, J. Reyes-Velasco and S. Boissinot on 26 August 2015 north (6.3844 ° N, 38.5927 ° E, 2655 m a. s. l.) and south (6.3256 ° N, 38.6645 ° E, 2684 m a. s. l.) of Bore, respectively, one male (16 - 6) collected by J. Reyes-Velasco and S. Boissinot on 4 July 2016 between Addis Ababa and Ambo, eight males (16 - 6, 16 - 10, 16 - 11, 16 - 12, 16 - 14, 16 - 16, 16 - 17 and 16 - 24) collected by J. Reyes-Velasco and S. Boissinot on 5 July 2016 south of Assela (7.7776 - 7.8431 ° N, 39.1384 - 39.1529 ° E, 2553 - 2637 m a. s. l.), one female (16 - 131) and six males (16 - 99, 16 - 106, 16 - 112, 16 - 113, 16 - 114 and 16 - 118) collected by J. Reyes-Velasco and S. Boissinot on 12 July 2016 southeast of Mehal Meda (9.9894 - 10.3316 ° N, 39.7452 - 39.8092 ° E, 3017 - 3394 m a. s. l.), one female (16 - 142) and one male (16 - 155) collected by J. Reyes-Velasco and S. Boissinot on 13 July 2016 north of Debre Birhan (9.6822 - 9.6979 ° N, 39.5505 - 39.5628 ° E, 2833 - 2837 m a. s. l.), one male (16 - 166) collected by J. Reyes-Velasco and S. Boissinot on 14 July 2016 south of Fitche (9.7502 ° N, 38.7445 ° E, 2726 m a. s. l.), one female (SB 231) and one male (SB 232) collected by S. Goutte and J. Reyes-Velasco on 23 April 2018 south of Gumer (7.9125 ° N, 38.0644 ° E, 2831 m a. s. l.), two males (SB 552 and SB 553) collected by S. Goutte and Y. Bourgeois on 3 July 2018 east of Mehal Meda (10.3247 ° N, 39.8092 ° E, 2795 m a. s. l.), one male (SB 562) collected by S. Goutte and Y. Bourgeois on 4 July 2018 east of Mehal Meda (10.3247 ° N, 39.8092 ° E, 2795 m a. s. l.), three males (SB 570, SB 571 and SB 577) collected by S. Goutte and Y. Bourgeois on 7 July 2018 north of Gebre Guracha (9.8818 ° N, 38.3660 ° E, 2558 m a. s. l.), one male (SB 615) collected by S. Goutte and Y. Bourgeois on 14 July 2018 near Holeta (9.0692 ° N, 38.5214 ° E, 2397 m a. s. l.), P. largeni male holotype (MHNG- 2513.31), P. largeni male paratypes (MHNG- 2513.38, MHNG- 2513.42, MHNG- 2513.44, MHNG- 2513.45, MHNG- 2513.49, MHNG- 2513.52), P. largeni female paratypes (MHNG- 2513.56, MHNG- 2513.57, MHNG- 2513.58, MHNG- 2513.59 and MHNG- 2513.60) collected by M. Largen on 12 June 1977 in Addis Ababa, Shewa (Shoa) (9.03 ° N, 38.75 ° E, 2500 m a. s. l.).	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
87B773BC7FF35B5B818DA5DB58E809FC.taxon	diagnosis	Diagnosis. A medium-sized (male (35) SVL 34.1 + / - 2.5 mm, female (10) SVL 34.6 + / - 1.7 mm) member of the Ptychadena erlangeri species group (Fig. 11). Ptychadena erlangeri differs from other members of the P. neumanni complex by the following combination of characters: (1) relatively short hind limbs (male TL / SVL 0.53 + / - 0.04, female TL / SVL 0.54 + / - 0.05), (2) tympanum translucent, without any light bar or blotch, (3) vocal sacs light in color, from cream to light grey, very rarely with a bit of grey, (4) adult males with robust forelimbs, (5) adult males often covered in small warts.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
87B773BC7FF35B5B818DA5DB58E809FC.taxon	description	Description of the holotype. Medium sized (SVL 37.6 mm), slender, gravid female with long hind limbs (TL / SVL 0.63, Fig. 12, Suppl. material 3: Table S 1). Head longer than wide (HW / HL 0.94). Snout pointed, projecting beyond the lower jaw. Interorbital distance 0.58 x eye diameter. Nostril half-way between the tip of the snout and the eye. Internarial distance 1.2 x interorbital distance. Tympanum 0.61 x eye diameter. Finger tips not expanded but rounded, with moderate subarticular tubercles. Finger formula: I <II <IV <III. Hand free of webbing. Hindlimbs elongated, with tibia length 0.63 x snout-vent length. Rounded white warts irregularly spread on the dorsal side of tibia. Foot as long as thigh and almost as long as tibia (FL / TL 0.98). Toe tips rounded. Subarticular tubercles small and round. Inner metatarsal tubercle present, external absent. Toe formula: I <II <III <V <IV (right foot) and I <II <VIII <IV (left foot). Foot webbing formula (toe internal / external sides, number of phalanges webbed): Ie (1), IIi / e (1 - 2), IIIi / e (2 - 2), IVi / e (2 - 2), Vi (2). Two light, continuous lateral ridges, four interrupted dorsal ridges. No vertebral nor sacral ridges. Note that we did not see the nasal ridges that Perret (1980) noted as the single major diagnostic character for the species.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
D81C7126345D57D4A71B845C742CC2E5.taxon	materials_examined	Material examined. In addition to the holotype, we examined one male (TJC 218) collected by T. J. Colston, one female (15 - 85) and two males (15 - 103, 15 - 105) collected on 7 August 2015 by X. Freilich, J. Reyes-Velasco and S. Boissinot in the Harenna forest (6.5866 ° N, 39.7417 ° E, 1778 m a. s. l.), one male (15 - 121) collected on 7 August 2015 by X. Freilich, J. Reyes-Velasco and S. Boissinot in the Harenna forest (6.6634 ° N, 39.7302 ° E, 2002 m a. s. l.), one female (15 - 425) and two males (15 - 426, 15 - 427) collected on 27 September 2015 by X. Freilich, J. Reyes-Velasco and S. Boissinot northwest of Kibre Mengist (5.9055 - 6.04546 ° N, 38.837 - 38.9334 ° E, 1745 - 2238 m a. s. l.), two males (15 - 448, 15 - 449) collected on 28 September 2015 by X. Freilich, J. Reyes-Velasco and S. Boissinot in Harenna forest (6.71925 N, 39.7202 E), two males (SB 99 and SB 100) collected on 10 April 2018 by S. Goutte and J. Reyes-Velasco in the Harenna forest (6.6634 ° N, 39.7302 ° E, 2440 m a. s. l.), two females (SB 158, SB 159) and two males (SB 160, SB 161) collected on 17 April 2018 by S. Goutte and J. Reyes-Velasco northwest of Kibre Mengist (6.0093 ° N, 38.8576 ° E, 2105 m a. s. l.), one female (SB 807) collected on 7 May 2019 by S. Goutte in the Harenna forest (6.6640 ° N, 39.7301 ° E, 1992 m a. s. l.), one female (SB 808) collected on 10 May 2019 by S. Goutte in the Harenna forest (6.7164 ° N, 39.7257 ° E, 2375 m a. s. l.).	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
D81C7126345D57D4A71B845C742CC2E5.taxon	diagnosis	Diagnosis. Large species (male (15) SVL 42.4 + / - 2.7 mm, female (6) SVL 51.2 + / - 4.2 mm) from the Ptychadena neumanni species group (Fig. 21) distinguished by the following combination of characters: (1) long hind limbs (male TL / SVL 0.60 + / - 0.05, female TL / SVL 0.59 + / - 0.02), (2) long feet (male FL / SVL 0.61 + / - 0.05, female FL / SVL 0.58 + / - 0.02), (3) long hands (male HAL / SVL 0.24 + / - 0.03, female HAL / SVL 0.23 + / - 0.01), (4) long head (male HL / SVL 0.38 + / - 0.04, female HL / SVL 0.36 + / - 0.04), (5) long snout (male SL / SVL 0.16 + / - 0.02, female SL / SVL 0.15 + / - 0.01), (6) vocal sacs light grey, cream or yellow, sometimes mottled with light grey, (7) male skin smooth.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
D81C7126345D57D4A71B845C742CC2E5.taxon	description	Description of the holotype. Relatively large (SVL 41.9 mm) male (Fig. 22) with long hind limbs (TL / SVL 0.58, Suppl. material 3: Table S 3). Head as long as wide. Snout projecting beyond the lower jaw. Interorbital distance almost equal to the eye diameter. Internarial distance 0.88 x interorbital distance. Tympanum 0.85 x eye diameter. Finger tips not expanded but rounded, with moderate subarticular tubercles. Finger formula: I <II <IV <III. Hand free of webbing. Hindlimbs elongated, with tibia length 0.58 x snout-vent length. Foot slightly longer than thigh and tibia (FL / THL 1.14, FL / TL 1.05). Toe tips rounded. Subarticular tubercles small and round. Inner metatarsal tubercle present, external absent. Toe formula: I <II <V <III <IV. Foot webbing formula: Ie (1), IIi / e (1 - 1.5), IIIi / e (1.5 - 2), IVi / e (2 - 2), Vi (2). Two continuous lateral ridges, brown anteriorly and light brown posteriorly, barely visible, six interrupted dorsal ridges. No vertebral, nor sacral ridges. No wart on body or limbs. Tongue free for less than a third of its length, divided in two lobes. Vomerine, maxillary and premaxillary teeth present.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
18DEDED7345456FAA395B2D2793E692D.taxon	materials_examined	Material examined. Beside the male holotype (TJC 219) we examined one female (15 - 155) and one male (15 - 158) collected by X. Freilich, J. Reyes-Velasco, and S. Boissinot on 9 August 2015 east of Dinsho (7.1061 ° N, 39.8181 ° E, 3058 m a. s. l.), one female (15 - 482) collected by X. Freilich, J. Reyes-Velasco and S. Boissinot on 3 October 2015 south of Dodola (6.8465 ° N, 39.1933 ° E, 3404 m a. s. l.), one female (16 - 89) and one male (16 - 90) collected by J. Reyes-Velasco and S. Boissinot on 10 July 2016 east of Dinsho (7.1065 ° N, 39.8184 ° E, 3065 m a. s. l.), one male (SB 43) collected by S. Goutte and J. Reyes-Velasco on 6 April 2018 south of Dodola (6.8632 ° N, 39.1948 ° E, 3260 m a. s. l.), two males (SB 66 and SB 67) collected by S. Goutte and J. Reyes-Velasco on 8 April 2018 south of Dinsho, one female (SB 85) and two males (SB 90 and SB 94) collected by S. Goutte and J. Reyes-Velasco on 9 April 2018 east of Dinsho.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
18DEDED7345456FAA395B2D2793E692D.taxon	diagnosis	Diagnosis. A small species (male (8) SVL 34.3 + / - 2.3 mm, female (4) SVL 37.0 + / - 2.5 mm) of the Ptychadena erlangeri species group (Fig. 13) with variable coloration, distinguished by the following combination of characters: (1) short hindlimbs (male TL / SVL 0.49 + / - 0.06, female TL / SVL 0.51 + / - 0.04), (2) head as long as wide, (3) snout short (male SN / SVL 0.13 + / - 0.02, female SN / SVL 0.13 + / - 0.01), (4) vocal sacs cream, light grey, or bicolored cream and grey.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
18DEDED7345456FAA395B2D2793E692D.taxon	description	Description of the holotype. Small sized (SVL 33.3 mm) male (Fig. 14). Head longer than wide (HW / HL 0.93). Snout slightly rounded, projecting beyond the lower jaw. Interorbital distance shorter than eye diameter (IOD / ED 0.78) and internarial distance (IOD / IND 0.93). Nostril closer to the eye than the tip of the snout. Tympanum 0.72 x eye diameter. Finger tips not expanded but rounded. Finger formula: I <II <IV <III. Hand free of webbing. Hindlimbs moderately long, with tibia length 0.54 x snout-vent length. Thigh shorter than tibia. Foot 1.3 x tibia length. Toe tips rounded. Inner metatarsal tubercle present, outer absent. Toe formula: I <II <V <III <IV. Foot webbing formula (toe internal / external sides, number of phalanges webbed): Ie (1), IIi / e (1 - 1.5), IIIi / e (1.5 - 2), IVi / e (2 - 2), Vi (2). Four discontinuous dorsal ridges and two discontinuous cream lateral ridges.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
A4613A6C39D755EF81344DEF810341F6.taxon	materials_examined	Material examined. Except for the type series, all the material examined is deposited at ZNHM. Beside the female holotype (ZMB- 26878 H), we examined four female paratypes (ZMB- 57189, ZMB- 57190, ZMB- 57193, ZMB- 57199) and four male paratypes (ZMB- 26877, ZMB- 57191, ZMB- 57192, ZMB- 57195) collected by C. von Erlanger and O. Neumann in 1900 likely between June and mid-August in Didda, East Arussi plateau, one female (SB 488) and two males (SB 486, SB 487) collected by S. Goutte and Y. Bourgeois on 26 June 2018 west of Ch'ange (8.1303 ° N, 39.3985 ° E, 2357 m a. s. l.), one female (SB 493) and two males (SB 494, SB 495) collected by S. Goutte and Y. Bourgeois on 27 June 2018 between Robe and Sedika (7.7307 ° N, 39.7133 ° E, 2377 m a. s. l.), five females (SB 508, SB 510, SB 512, SB 515, SB 516) and four males (SB 523, SB 524, SB 525, SB 526) collected by S. Goutte and Y. Bourgeois on 29 June 2018 southeast of Ch'ange (8.1086 ° N, 39.4486 ° E, 2573 m a. s. l.).	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
A4613A6C39D755EF81344DEF810341F6.taxon	diagnosis	Diagnosis. The smallest species (male (12) SVL 28.3 + / - 3.3 mm, female (12) SVL 29.6 + / - 3.2 mm) of the P. neumanni complex (Fig. 15), distinguished by the following combination of characters: (1) short hind limbs (male TL / SVL 0.45 + / - 0.03, female TL / SVL 0.44 + / - 0.03), (2) short hands (HAL / SVL 0.20 + / - 0.02), (3) short feet (male FL / SVL 0.49 + / - 0.04, female FL / SVL 0.48 + / - 0.04), (4) reduced foot webbing, (5) no stripe or blotch on the tympanum, (6) adult males almost always covered in small warts during the breeding season.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
A4613A6C39D755EF81344DEF810341F6.taxon	description	Description of the holotype. Small sized (SVL 26.9 mm) gravid female with short hind limbs (TL / SVL 0.43, Fig. 16, Suppl. material 3: Table S 1). Head longer than wide (HW / HL 0.95). Snout rounded, slightly projecting beyond the lower jaw. Interorbital distance 0.58 x eye diameter. Nostril half-way between the tip of the snout and the eye. Internarial distance 1.4 x interorbital distance. Tympanum 0.65 x eye diameter. Finger tips not expanded but rounded. Finger formula: III <IV <III. Hand free of webbing. Hindlimbs short, with tibia length 0.42 x snout-vent length. Foot 1.2 x tibia length and 1.3 thigh length. Toe tips rounded. Outer metatarsal tubercle absent. Toe formula: I <II <V <III <IV. Foot webbing formula (toe internal / external sides, number of phalanges webbed): Ie (1), IIi / e (1 - 1), IIIi / e (1 - 2), IVi / e (2 - 2), Vi (2). Ridges hardly distinguishable due to the state of preservation of the specimen.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
F94186280C5155C0B4CA4F978B36CC5E.taxon	materials_examined	Material examined. Except for the type series, all examined specimens are deposited at ZNHM. In addition to the type series, we examined one male (15 - 173) collected by X. Freilich, J. Reyes-Velasco and S. Boissinot on 9 August 2015 in Wondo Genet (7.0833 ° N, 38.6381 ° E, 1896 m a. s. l.), one female (15 - 181) and four males (15 - 183, 15 - 191, 15 - 208, 15 - 209) collected by X. Freilich, J. Reyes-Velasco and S. Boissinot on 12 August 2015 northwest of Bonga (7.3076 ° N, 36.1226 ° E, 1861 m a. s. l.), one female (16 - 203) collected on 18 July 2016 by J. Reyes-Velasco and S. Boissinot southwest of Bonga (7.2542 ° N, 36.2628 ° E, 1963 m a. s. l.), one female (16 - 302) and two males (16 - 303, 16 - 305) collected on 19 July 2016 by J. Reyes-Velasco and S. Boissinot north of Maji (6.2365 ° N, 35.5712 ° E, 1936 m a. s. l.), one female (16 - 326) and one male (16 - 313) collected on 20 July 2016 by J. Reyes-Velasco and S. Boissinot northeast of Machi (6.3780 ° N, 35.6659 ° E, 2063 m a. s. l.), two males (16 - 327, 16 - 329) collected on 20 July 2016 by J. Reyes-Velasco and S. Boissinot northeast of Mizan Teferi (7.0203 ° N, 35.7545 ° E, 2449 m a. s. l.), two females (SB 333, SB 334) collected on 9 June 2018 by S. Goutte and J. Reyes-Velasco northwest of Gore (8.2014 ° N, 35.3772 ° E, 1666 m a. s. l.), one male (SB 388) collected on 11 June 2018 by S. Goutte and J. Reyes-Velasco south of Gech'a (7.5544 ° N, 35.4148 ° E, 1936 m a. s. l.), one female (SB 405) collected on 13 June 2018 by S. Goutte and J. Reyes-Velasco south of Gech'a (7.5185 ° N, 35.4163 ° E, 1917 m a. s. l.), one female (SB 462) collected on 20 June 2018 by S. Goutte and J. Reyes-Velasco northeast of Shebe (7.5423 ° N, 36.5732 ° E, 2240 m a. s. l.).	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
F94186280C5155C0B4CA4F978B36CC5E.taxon	diagnosis	Diagnosis. Medium-sized species (male (20) SVL 35.7 + / - 2 mm, female (8) SVL 45.6 + / - 1.4 mm) of the Ptychadena neumanni species group (Fig. 23) distinguished by the following combination of characters: (1) long hind limbs (male TL / SVL 0.58 + / - 0.02, female TL / SVL 0.57 + / - 0.01), (2) vertical cream bar on the tympanum, (3) vocal sacs uniformly dark grey.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
F94186280C5155C0B4CA4F978B36CC5E.taxon	description	Description of the lectotype. The lectotype is very desiccated (Fig. 24 A) and the description of some characters is hindered by the state of conservation of the specimen. Medium sized (SVL 32.2 mm), slender adult male. Snout pointed, projecting beyond the lower jaw. Interorbital distance 0.83 x eye diameter. Head longer than wide (HW / HL 0.80). Nostril half-way between the tip of the snout and the eye. Internarial distance 1.1 x interorbital distance. Tympanum 0.64 x eye diameter. Finger tips not expanded but rounded, with moderate subarticular tubercles. Finger formula: I <II <IV <III. Hand free of webbing. Hindlimbs elongated, with tibia length 0.57 x snout-vent length. Foot longer than thigh (FL / THL 1.1) and as long as tibia. Toe tips rounded. Subarticular tubercles small and round. Inner metatarsal tubercle present, external absent. Toe formula: I <II <III <V <IV. Foot webbing formula: Ie (1), IIi / e (1 - 2), IIIi / e (2 - 2), IVi / e (2 - 2), Vi (2). Two light, continuous lateral ridges, six dorsal ridges difficult to see due to preservation of the specimen. No vertebral nor sacral ridges. Small warts on the body. No sacral, femoral, or crural folds.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
45EEF9BCD0515803A1CF764921588C18.taxon	diagnosis	Diagnosis. Small species (male (13) SVL 30.3 + / - 0.7 mm, female (4) SVL 31.6 + / - 2.9 mm) of the Ptychadena erlangeri species group (Fig. 17), distinguishable from other species of the P. neumanni complex by the following combination of characters: (1) short hind limbs (male TL / SVL 0.45 + / - 0.0.03, female TL / SVL 0.44 + / - 0.02), (2) short forearms (male FLL / SVL 0.18 + / - 0.02, female FLL / SVL 0.18 + / - 0.00), (3) short feet (male FL / SVL 0.5 + / - 0.03, female FL / SVL 0.49 + / - 0.05), (4) reduced foot webbing, (5) a vertical cream or golden stripe or blotch on the tympanum (sometimes faint), (6) warts absent in adult males during the breeding season.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
45EEF9BCD0515803A1CF764921588C18.taxon	description	Description of the holotype. Small sized (SVL 29.3 mm) male with short hind limbs (TL / SVL 0.45, Fig. 17, Suppl. material 3: Table S 1). Head longer than wide (HW / HL 0.96). Snout slightly rounded, projecting beyond the lower jaw. Interorbital distance equal to the eye diameter and to the internarial distance. Nostril half-way between the tip of the snout and the eye. Tympanum 0.77 x eye diameter. Finger tips not expanded but rounded. Finger formula: I <II <IV <III. Hand free of webbing. Hindlimbs short, with tibia length 0.45 x snout-vent length. Tibia and thigh lengths equal. Foot 1.3 x tibia length. Toe tips rounded. Inner metatarsal tubercle present, outer absent. Toe formula: I <II <V <III <IV. Foot webbing formula (toe internal / external sides, number of phalanges webbed): Ie (1), IIi / e (1 - 1), IIIi / e (1 - 1), IVi / e (1 - 1), Vi (1). Four continuous dorsal ridges and one faint cream lateral ridge, interrupted on one side and continuous on the other.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
45EEF9BCD0515803A1CF764921588C18.taxon	etymology	Etymology. The specific name refers to the town of Robe, near the type locality.	en	Goutte, Sandra, Reyes-Velasco *, Jacobo, Freilich, Xenia, Kassie, Abeje, Boissinot, Stephane (2021): Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands. ZooKeys 1016: 77-141, DOI: http://dx.doi.org/10.3897/zookeys.1016.59699, URL: http://dx.doi.org/10.3897/zookeys.1016.59699
