taxonID	type	description	language	source
E4A8D7CD19877C832522EF86590B1D67.taxon	description	Male, Figs 1, 7 - 9, 22, 34 - 39, 40 - 45; Female, Figs 29, 31, 46 - 50	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
E4A8D7CD19877C832522EF86590B1D67.taxon	diagnosis	Diagnosis. The male of this species shares with that of E. oliveri and E. simillima the subapical projection of the anterior margin of the male metatibia, which is formed by the medial portion of the ridge that borders the depressed area and thus located above the inner spur, and the dorsal lobe of the gonostylus, which is apically about as broad as its base and largely bare on its outer surface. It can be separated from E. simillima by the length of the glossa (reaching mesotrochanter in E. coerulescens vs. reaching metatrochanter in E. simillima), presence of a longitudinal median depression on clypeus (absent in E. simillima), outer surface of mesotibia with area between the medial margin of the posterior felty patch and the anterior margin of tibia pubescent throughout except for small area apically (half bare apically in E. simillima), and by the body color (largely metallic green in the male of E. coerulescens vs. dark blue with violet hues in both sexes of E. simillima). In addition, both species are geographically separated: E. simillima is restricted to the western slope of the Cordillera of Chihuahua whereas E. coerulescens is more widely distributed, occurring along the Sierra Madre Oriental and eastern Mexico. From E. oliveri, which shares the same body coloration, it can be separated by the length of the glossa (reaching S 2 in E. oliveri), punctation of T 2 (punctures on disc separated by at least half a puncture width in E. coerulescens, closer in E. oliveri), and posterior felty patch of mesotibia, which is broader medially than apically (about the same width across its length in E. oliveri). The female can be recognized by the following combination of features: glossa extending to metatrochanter; dorsum of mesosoma with pubescence not obscuring integument; metasoma with apical terga and sterna with white setae at least laterally; T 2 with sparse, coarse punctures; mesoscutellum with fine, dense punctures; and metabasitarsus short, 1.7 - 1.8 times longer than broad. In E. simillima the pubescence of metasoma is black, T 2 is more finely punctate, and metabasitarsus is longer (2.2 times longer than broad). In E. micheneri the pubescence on the dorsum of mesosoma is denser, partially obscuring the integument, the mesoscutellum is more coarsely and sparsely punctate, and the metabasitarsus is about twice as long as broad.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
E4A8D7CD19877C832522EF86590B1D67.taxon	materials_examined	Material examined. (n = 46 ♀, 3 ♂,) 1 ♂, USA: Texas, Culberson County: Guadalupe Mountains National Park, Pine Springs, N 31.8955 W 104.8271, 20 Jul 2010, J. D. Herndon, A. Druk, H. Ikerd, pantrap, GUMO 27853 (BBSL); 1 ♂, New Mexico, Eddy County: Longview Spring, 0.7 km E, N 32.1007 W 104.6317, 1551 m, 22 Jul 2010, J. D. Herndon, Cirsium sp., CAVE 20415 (BBSL); Mexico: 2 ♀, Chihuahua, General Trias, August 20, 1991, J. Rozen / N. Pember, SM 0307820, - 21, KUNHM-ENT (SEMC); 1 ♀, Coahuila, Cuesta la Muralla, ix- 12 - 76 / J. A. Chemsak, J. Powell, A. & M Michelbacher collectors / EMEC 1069129 (EMEC); 8 ♀, Durango, Dgo., Mex., 6200 ft. Aug. 14, 1947 / D. Rockefeller, Exp. Gertsch / SM 0504556, - 59 - 65 KUNHM-ENT (SEMC); 1 ♂, idem (FSCA); 1 ♀, idem / BBSL 849648 (BBSL); 1 ♀, idem / D. Rockefeller, Exp. Spieth / SM 0504557, KUNHM-ENT (SEMC); 1 ♀, idem / D. Rockefeller, Exp. Cazier / SM 0504558, KUNHM-ENT (SEMC); 1 ♀, ECO-TAP-E- 105054, Mex., Hgo. [Hidalgo], La Mision, Puerto de Piedra, 1678 m, 21,05076 N, 99,11475 W, 19 / 8 / 2013, 09: 10, Col. Jorge Merida; 3 ♀, Mexico: Hidalgo, 24 km NW Cardonal (Barranco del Tulanlango), 1590 m / 13 July 1990, R. L. Minckley, ex., Cassia / SM 0504551 - 53, KUNHM-ENT (SEMC); 1 ♀, Mexico: Hidalgo, 3.5 km W Canyon of Tolantongo / NE of Ixmiquilipan, 1590 m, 13 July 1990, I. Yarom, on Cassia? / SM 0504576, KUNHM-ENT (SEMC); 3 ♀, Mexico: Hidalgo, Grutas Tolantongo, 30 km NW Cardonal, 1230 m, 13 July 1990. W. J. Bell # 2 / SM 0504521 - 22, - 75, KUNHM-ENT (SEMC); 1 ♀, Mexico: Hidalgo, 31 km S Jacala, 1050 m, 12 July 1990, W. J. Bell # 2 / SM 0504579, KUNHM-ENT (SEMC); 1 ♀, Mexico: Hidalgo, Santorun, Atotonlico el Grande, 13 Sept. 1997, L. Godinez, # 736, ex. Senecio salignus / SM 0504580, KUNHM-ENT (SEMC); 1 ♂, ECO-TAP-E- 104040, NL [Nuevo Leon], Aranberri, P. de Anteojitos, 2041 m, 24,21185 N, 99,88922 W, 8 / 8 / 2013, 09: 05, Col. Jorge Merida (ECOSUR); 2 ♀, ECO-TAP-E- 104736, - 53, Mex., Qro. [Queretaro], La Loma, 1830 m, 21,36714 N, 99,76277 W, 16 / 8 / 2013, 10: 35, Col. Jorge Merida, Erika Esquivel; 1 ♀, Mexico: Queretaro, 27 km W San Luis Potosi, 1270 m, 8 July 1990, W. J. Bell # 1 / SM 0504577, KUNHM-ENT (SEMC); 2 ♀, idem, 26.5 km W San Luis Potosi / borde ron Hwy. 120, 1270 m, 9 July 1990, R. L. Minckley / SM 0504523, - 70, KUNHM-ENT (SEMC); 1 ♀, ECO-TAP-E- 104700, Mex., SLP [San Luis Potosi], Alaquines, Las Huertas, 1384 m, 22,04671 N, 99,53239 W, 15 / 8 / 2013, 11: 30, Col. Erika Esquivel; 4 ♀, idem, ECO-TAP-E- 104661, S. J. de Corito, 1353 m, 22,00349 N, 99,49312 W, 8: 25, Col. Jorge Merida; 2 ♀, ECO-TAP-E- 104939, Mex., SLP, Xilitla, La Soledad, 1356 m, 21,33348 N, 99,08237 W, 18 / 8 / 2013, 10: 40, Col. Jorge Merida, Erika Esquivel; 1 ♀, idem, ECO-TAP-E- 104965, Buenavista, 1648 m, 21,33774 N, 99,10539 W; 1 ♀, 17 mi. W, Xilitla, S. L. P., Mex., vii- 22 - 54,4700 ft / Univ. Kans. Mex. Expedition / SM 05045, KUNHM-ENT (SEMC); 1 ♀, Mexico, San Luis Potosi, 17 mi E Cd. Maiz, 3200 ', 23 July 196, U. Kans. Mex. Exped. / SM 0504569, KUNHM-ENT (SEMC); 1 ♀, idem, 15 mi. E Cd. Maiz, 3700 ' / SM 0504528, KUNHM-ENT (SEMC); 2 ♀, idem, 5 mi. E Ciudad del Maiz, S. L. P., Mex. 4700 feet, viii- 22, 23 - 1954 / SM 0504554, - 55, KUNHM-ENT (SEMC); 1 ♀, Mex: San Luis Potosi, 15 mi. W, Xilitla, 1350 m, viii- 15 - 1977, E. I. Schlinger / on plant # 8, Cassia sp. / EMEC 1069131 (EMEC); 1 ♀, Villagran, Tamaulipas, Mex, ix- 8 - 1966, D. H. Janzen, Thevetia / AMNH _ BEE 00164041 (AMNH); 1 ♀, Mex: Tamaulipas, 15 mi. SW Cd. Victoria, 5000 ', ix- 19 - 76 / J. A. Chemsak, J. Powell, A. & M Michelbacher collectors / EMEC 1069128, - 30 (EMEC).	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
E4A8D7CD19877C832522EF86590B1D67.taxon	distribution	Distribution. (Fig. 128) Mexico: Chihuahua, Coahuila, Durango, Hidalgo, Nuevo Leon, Queretaro, San Luis Potosi, Tamaulipas. USA (accidental?, see results and discussion): Texas, New Mexico.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
2642EBBB8655B7B9E50F7C95A91B1BB6.taxon	description	Male, Figs 2, 10, 11, 21, 23 - 25, 51, 53, 55, 56, 58 - 62; Female, Figs 28, 33, 52, 54, 57, 63	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
2642EBBB8655B7B9E50F7C95A91B1BB6.taxon	diagnosis	Diagnosis. Both sexes of this species are most similar to E. coerulescens, from which it can be separated easily by the longer glossa and more densely punctate disc of T 2. The female of E. oliveri can be recognized from that of E. barthelli by the mesoscutellum coarsely and non-contiguously punctate submedially, with a weak medial groove lacking a distinct row of setae (finely, contiguously punctate with stronger medial groove having a distinct row of setae in E. barthelli), and by the T 2, which is more densely punctate on disc, with punctures separate by at most a puncture width (punctures separated by 1.0 - 2.0 times a puncture width in E. barthelli).	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
2642EBBB8655B7B9E50F7C95A91B1BB6.taxon	description	Description. Holotype, ♂: As described for E. coerulescens except as follows: Head width 5.7 mm; intertegular distance 4.6 mm; body length 15.4 mm; forewing length 13.5 mm. Glossa in repose reaching S 2; metabasitarsus with posterodistal margin more rounded, not as acute as in E. coerulescens. Hidden sterna and genitalia as in Figs 58 - 62. Vertex and gena green with weak blue and purple hues. Mesosoma (excluding legs) predominantly green except mesoscutellum green with weak blue and golden hues; metatibia basally mostly green with golden hues; wing membrane infuscate, lighter than in E. coerulescens. Metasoma green with distal margins of terga and sterna (excluding S 2) weakly blue to purple. Head with whitish setae, with scattered gray to black setae on vertex. Mesotibia with area between medial margin of posterior patch and anterior margin of tibia setose on basal half, distal half asetose; posterior patch 5.7 times longer than broad, subapically narrower than distance between its medial margin and anterior margin of tibia. T 2 with punctures on disc separated by at most half a puncture width, impunctate distal margin very narrow, at most as wide as a puncture width. ♀: Head width 6.3 mm; intertegular distance 4.9 mm; body length 18.4 mm; forewing length 13.5 mm. Metatibia with emargination on distal margin about half width of posterodistal angle; metabasitarsus twice as long as broad. Blue with purple hues on mandible basally, labrum, frons, vertex, gena, dorsum of mesosoma, mesepisternum, propodeum, legs, discs of sterna, and marginal zones of terga. Pubescence black, except off-white on sides of T 2, T 3 - T 6, sides of S 2 and S 3, entire S 4 and S 5, and S 2 basally. Mesoscutellum without row of dense setae on weak medial longitudinal groove. Clypeus with median longitudinal ridge continuing onto supraclypeal area; labrum with distinct median and sublateral carinae. Mesoscutellum with coarse, non-contiguous punctures submedially. T 2 with punctures sparser than in the male, separated by a puncture width or less.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
2642EBBB8655B7B9E50F7C95A91B1BB6.taxon	materials_examined	Additional material. (n = 5 ♂, 12 ♀, not designated as paratypes) 3 ♀, Mexico: Morelos, 4 mi SW Yautepec, 2 July 1961, 3800 ', C. D. Michener / on flowers of Cassia sp. / KUNHM-ENT 504573, 504526, 504529 (KU); 1 ♀, Canon de Lobos, Mor. [Morelos], 16 - xi- 83, R y G. Ayala / # 10447 (UNAM); 1 ♂, Ocuilan, @ Stanhopea hernandezii, 2001, M. A. Soto (FSCA); 3 ♀, Mex [Mexico], Oaxaca, Tehuantepec, 56 mi N. W., vii- 27 - 63 / W. A. Foster collectors / AMNH _ BEE 00265723 (AMNH); 2 ♀, Mexico, Puebla, 12 mi NW Tehuitzingo, 4050 ft, 29 June 1961, U. Kans. Mex. Exped. / KUNHM-ENT 504524, 504572 (KU); Mexico, Jalisco, Estacion de Biologia Chamela, 5 - viii- 86 / M. Sanchez / / KUNHM-ENT 730197; 1 ♀, idem, 10 / 14 Jul 1989, T. Griswold / BBSL 861169 (BBSL); 1 ♀, idem, 4 - vii- 1984 / R. Ayala / # 10448 (UNAM); 1 ♂, Tzararacua, near Uruapan, Mich. [Michoacan], Mexico, 28 June 68 [1968], N. H. Williams, Eugenol, 276 (FSCA); 1 ♂, Mexico, Mich. [Michoacan], Tzararacua, nr. Uruapan, 29 VI 1968, N. H. Williams, 276, Eugenol (FSCA); 1 ♂, Mexico, Mich. [Michoacan], Tzararacua, nr. Uruapan, 6 VII 1968, N. H. Williams, 276, Eugenol (FSCA); C. [Clowesia] glaucoglossa, La Huacana, Mich. [Michoacan], Mex [Mexico], Ernesto Aguirre, 3 - 22 - 1977 (FSCA).	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
2642EBBB8655B7B9E50F7C95A91B1BB6.taxon	etymology	Etymology. This species is named after Oliver Mitchell Betancourt, son of the first author (March 11, 2015), who daily brings love and joy.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
2642EBBB8655B7B9E50F7C95A91B1BB6.taxon	distribution	Distribution. (Fig. 128) Mexico: Guerrero, Jalisco, Michoacan, Morelos, Oaxaca, Puebla.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
69CAEBD29A3885E93147F4645C77678A.taxon	description	Male, Figs 3, 12, 13, 67 - 72, 73 - 78; Female, Figs 64, 79 - 83	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
69CAEBD29A3885E93147F4645C77678A.taxon	diagnosis	Diagnosis. Both sexes of this species can be easily recognized by their body color, which is predominantly dark purple, thus resembling E. micheneri. However, they can be easily separated from that species by the shorter glossa (not surpassing metatrochanter in E. simillima and reaching S 1 in E. micheneri), shape of the subapical projection on the anterior surface of the male metatibia, male gonostylus, and metasoma with dark brown to black pubescence (apical terga and sterna with white setae at least laterally in E. micheneri). (See comparative diagnosis for E. micheneri)	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
69CAEBD29A3885E93147F4645C77678A.taxon	distribution	Distribution. (Fig. 128) This species is known only from the western slope of the Cordillera of Chihuahua, Mexico.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
79CE22354F6F05F56D219D46F4FE5855.taxon	description	Male, Figs 4, 14, 15, 84 - 89, 90 - 95; Female, Figs 27, 65, 96 - 99	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
79CE22354F6F05F56D219D46F4FE5855.taxon	diagnosis	Diagnosis. This species shares with E. micheneri and E. engeli the subapical projection of the anterior margin of the male metatibia, which is formed by the upper ridge that borders the depressed area, and the dorsal lobe of the gonostylus, which is apically broad and covered by setae on its outer surface. It is most similar to E. engeli from southern Mexico. It can be separated from that species by the posterior felty patch of the mesotibia, which is subapically about as wide as the distance between its medial margin and the anterior margin of the tibia (broader in E. engeli), the finer and slightly sparser punctures on disc of T 2, and its geographical distribution (E. barthelli occurs in central Mexico). The female of E. barthelli is similar to that of E. oliveri in the long glossa, reaching at least to S 1. However, they can be separated primarily by the punctation of the mesoscutellum. In E. barthelli it is finely, contiguously punctate throughout, with a distinct medial longitudinal groove that bears a row of dense setae. In E. oliveri, the mesoscutellum is coarsely, non-contiguously punctate, and with a weak medial longitudinal groove lacking a distinct row of dense pubescence.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
79CE22354F6F05F56D219D46F4FE5855.taxon	description	Description. Holotype, ♂: Head width 6.0 mm; intertegular distance 5.0 mm; body length 18.9 mm; forewing length 15.0 mm. Glossa in repose reaching S 2. Anterior margin of metatibia in inner view with elevated ridge bordering depressed, smooth and hairless area above tibial spurs, upper margin of ridge medially projected, appearing as a spine in profile; metabasitarsus about twice as long as broad, inner surface near base weakly protuberant in frontal view, posterior margin gently convex, posterodistal margin broadly rounded. Hidden sterna and genitalia as in Figs 90 - 95; dorsal lobe of gonostylus broader apically than basally, distinctly setose on outer surface. Mandible black on apical two-thirds, basally blue; labrum blue with weak green hues; remainder areas of head green with weak golden hues on face and bluish hues around ocelli and gena; antenna black. Mesosoma excluding legs predominantly green, with weak golden hues on mesoscutum anteriorly, disc of tegula and mesepisternum dorsally; weak bluish hues on axilla laterally, mesoscutellum, and propodeum basal and laterally. Legs mostly blue-purple except green on pro- and mesotibiae anteriorly and most of metatibia. Wing membrane infuscate, veins dark brown to black. T 1 - T 4 green except distal margins blue to purple; T 5 - T 7 blue-purple; sterna mainly green with weak blue-purple hues except S 2 dark brown, with blue-purple hues barely visible basolaterally. Head mainly with off-white setae, with gray to black setae on vertex. Mesosoma with gray to black setae except: whitish setae on posterior margin of meso- and metatibiae, outer surfaces of mesobasitarsus and metatibia; off-white on anterior half of mesoscutum, and lateral face of mesepisternum. Mesotibia with area between medial margin of posterior patch and anterior margin of tibia setose except on apical one-fourth; posterior patch 4.5 times longer than broad, subapically about as wide as distance between its medial margin and anterior margin of tibia. Metasoma with off-white to light brown setae, longer and denser on apical terga and sterna. Clypeus without longitudinal medial depression. T 2 with fine punctures on disc separated by at most a puncture width, distal margin narrow, about twice a puncture width. ♀: Head width 6.3 - 6.5 mm; body length 16.5 - 17.8 mm; forewing length 15.0 mm. Metatibia with emargination on distal margin about half width of posterodistal angle; metabasitarsus twice as long as broad. Blue with purple hues on mandible basally, labrum, dorsum of mesosoma, mesepisternum ventrally, propodeum, legs, discs of sterna, and marginal zones of terga. Pubescence black, except off-white on sides of T 2, T 3 - T 6, S 1 - S 5, and S 2 basally. Pubescence denser on mesosoma, not obscuring integument. Clypeus with median longitudinal ridge continuing onto supraclypeal area; labrum with basal, longitudinally elongate tubercle, sublateral carinae sometimes weak. Mesoscutellum with fine, contiguous punctures submedially, median longitudinal groove well defined, with distinct row of dense pubescence. T 2 with punctures on disc sparser than in the male, separated by at least a puncture width, impunctate distal margin wide, at least three or four times a puncture width.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
79CE22354F6F05F56D219D46F4FE5855.taxon	materials_examined	Additional material. (n = 6 ♀, not designated as paratypes) 1 ♀, ECO-TAP-E- 95712, Mex., Jal., [Jalisco], Tamazula, Agua Zarca, 1829 m, 19.82484 N, 103.27449 W, 31 / 10 / 2012, 10: 00, Col. Jorge Merida (ECOSUR); 1 ♀, Mexico: Jalisco, 13 km SE La Manzanilla, La Loma Alla, 31 - x- 1990, C. M. Estrada # 69 / SM 0730193, KUNHM-ENT (SEMC); 1 ♀, Museum Paris, Mexique, Etat de Jalisco, L. Diguet, 1900 (MNHN); 1 ♀, Ahuacatlan, Nay. [Nayarit], Mex., vii- 18 - 22 - 51 / H. E. Evans, collector / SM 0504525, KUNHM-ENT (SEMC); 1 ♀, 3 mi W. Jacona, Mich [Michoacan], vii- 18 - 53, 5300 ft / Univ. Kans. Mex. Expedition / SM 0504527, KUNHM-ENT (SEMC); 1 ♀, Mexico, Morelos, 4 mi. SW Yautepec, 2 July 1961, 3800 ', C. D. Michener, on flowers of Cassia sp. (FSCA). Variation. The blue-purple coloration is strong in both males from Morelos and one of the males from Jalisco. In particular, one of the males from Morelos (UNAM, # 02290) has extensive blue hues on all tagmata.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
79CE22354F6F05F56D219D46F4FE5855.taxon	etymology	Etymology. This species is dedicated to our friend and colleague Dr. John Barthell (University of Central Oklahoma) for his contributions to bee ecology and efforts to promote undergraduate research on bees.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
79CE22354F6F05F56D219D46F4FE5855.taxon	distribution	Distribution. Central Mexico: Jalisco, Michoacan, Morelos, Nayarit, Oaxaca.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
91E8AC0EB5E4AE58B30BBCC263D4741F.taxon	description	Male, Figs 5, 16, 17, 26, 100 - 105, 106 - 111	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
91E8AC0EB5E4AE58B30BBCC263D4741F.taxon	diagnosis	Diagnosis. This species is known only from the male sex. It along with E. micheneri and E. barthelli belong to a group of species that differs from other concolorous metallic blue to dark green Mexican Eufriesea by the anterior margin of metatibia in inner view without a distinct spine-like subapical projection above inner spur, but upper margin of distinctly depressed area projected medially, thus often appearing spine-like in profile (Figs 14 - 19) and the genitalia with dorsal lobe of gonostylus apically broad, distinctly setose on outer surface (Fig. 26). It can be separated from E. micheneri by the metabasitarsus broadly rounded posterodistally (angled in E. micheneri), T 2 with punctures separated by at most a puncture width on disc (punctures separated by 1 - 2 times a puncture width in E. micheneri), and by the body color (largely metallic green in the male of E. engeli vs. dark blue with violet hues in both sexes of E. micheneri). From E. barthelli it can be separated by the posterior felty patch of mesotibia, which is subapically broader than the distance between its medial margin and anterior margin of tibia (narrower in E. barthelli), punctation of T 2 (punctures coarser and nearly contiguous in E. engeli, finer and slightly sparser in E. barthelli), and its geographical distribution (E. engeli occurs in southern Mexico whereas E. barthelli in central Mexico).	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
91E8AC0EB5E4AE58B30BBCC263D4741F.taxon	description	Description. Holotype, ♂: As described for E. barthelli except as follows: Head width 5.9 mm; body length 16.7 mm; intertegular distance 4.6 mm; forewing length 14.4 mm. Glossa in repose reaching S 2. Hidden sterna and genitalia as in Figs 106 - 111. Gena mostly green. Mesosoma excluding legs predominantly green, with weak golden hues on mesoscutum anteriorly, disc of tegula and mesepisternum dorsally; weak bluish hues on axilla laterally, mesepisternum ventrally, and propodeum basal and laterally. Legs mostly blue-purple except green on pro- and mesotibiae anteriorly and metatibia basally. T 1 - T 4 green except distal margins blue to purple; T 5 - T 7 blue with purple marginal zones; sterna mainly green except S 2 dark brown, with blue to purple hues on apical sterna. Mesosoma with mostly off-white setae except gray to black setae on pronotum, posterior half of mesoscutum, mesoscutellum, mesepisternum ventrally, metepisternum, and propodeum. Legs with off-white setae, except gray to dark brown setae on inner surfaces of tibiae, basitarsi, and tarsi of all legs. Mesotibia with posterior patch subapically broader than distance between its medial margin and anterior margin of tibia. Disc of T 2 with coarser and denser punctures on disc than in E. barthelli, punctures separated by less than a puncture width to nearly contiguous. Female. Unknown.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
91E8AC0EB5E4AE58B30BBCC263D4741F.taxon	materials_examined	Additional material. (n = 7 ♂, not designated as paratypes) 1 ♂, Mexico: Chiapas, L. [Lago] Montebello, 20 v 1970, 1410, R. L. Dressler (FSCA); 5 ♂, Tuxtla Gutierrez, Chiapas, Mexico, C. Dodson, 7 - 21 - 1968, No. 310, Cineole (FSCA); 1 ♂, Mexico: Chis [Chiapas], Tuxtla Gutierrez, Cineole, C. H. Dodson, 310, 19 - 23 VII 1968 (FSCA).	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
91E8AC0EB5E4AE58B30BBCC263D4741F.taxon	etymology	Etymology. This species is dedicated to our friend and colleague Dr. Michael S. Engel (University of Kansas), in recognition of his significant contributions to systematic melittology.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
91E8AC0EB5E4AE58B30BBCC263D4741F.taxon	distribution	Distribution. (Fig. 128) This species is known only from Chiapas, southern Mexico, but can be expected to be found in Guatemala since Lago Montebello is on the border with Guatemala.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
CBA58212A16F7419578001F64212937C.taxon	description	Male, Figs 6, 18, 19, 20, 112 - 117, 118 - 123; Female, Figs 30, 32, 66, 124 - 127	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
CBA58212A16F7419578001F64212937C.taxon	diagnosis	Diagnosis. Superficially this species resembles E. simillima in that both sexes are primarily blue with purple hues. In addition to their geographical separation (E. simillima occurs in the Sierra Madre de Occidental whereas E. micheneri occupies western parts of the Transverse Volcanic Belt), it can be separated by the length of the tongue (reaching S 2 in E. micheneri and only the metatrochanter in E. simillima), the subapical projection of the anterior margin of the male metatibia (formed by the upper portion of the ridge that borders the depressed area in E. micheneri and by the medial portion of the ridge, and thus located above the inner spur, in E. simillima), and by the dorsal lobe of gonostylus (apically about as broad as its base and largely bare on its outer surface in E. simillima and apically broad and setose on outer surface in E. micheneri). From E. barthelli and E. engeli it can be separated by T 2 with sparser punctures on disc (contiguous or separated by at most a puncture width in E. barthelli and E. engeli), metabasitarsus with posterodistal margin angled (broadly rounded in E. barthelli and E. engeli) and pubescence of mesotibia, between the medial margin of posterior felty patch and anterior margin of tibia (presence on basal half in E. micheneri and basal two-thirds in E. barthelli and E. engeli).	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
CBA58212A16F7419578001F64212937C.taxon	distribution	Distribution. (Fig. 128) Mexico: Durango, Jalisco, Michoacan, Nayarit. This species appears to be restricted to mid elevation oak and pine forests in the western parts of the Transverse Volcanic Belt.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
CBA58212A16F7419578001F64212937C.taxon	materials_examined	Material examined. In addition to the paratypes deposited in SEMC (five males and one female from Mascota, Jaslico, and one male from Santa Tereresa, Nayarit), we also examined the following two females deposited in ECOSUR: 1 ♀, ECO-TAP-E- 118160, Mex., Dgo. [Durango], San Dimas, 1760 m, 24,53210 N, 105,81339 W, 2 / 10 / 2013, 10: 00, Col. Liliana Tlapaya; 1 ♀, ECO-TAP-E- 102831, Mex., Mich. [Michoacan], Morelia, J. del Monte, 2172 m, 19,63395 N, 101,14644 W, 4 / 7 / 2013, 10: 05, Col. Oscar Martinez Lopez.	en	Gonzalez, Victor H., Griswold, Terry, Simoes, Marianna (2017): On the identity of the adventive species of Eufriesea Cockerell in the USA: systematics and potential distribution of the coerulescens species group (Hymenoptera, Apidae). Journal of Hymenoptera Research 55: 55-102, DOI: http://dx.doi.org/10.3897/jhr.55.12209, URL: http://dx.doi.org/10.3897/jhr.55.12209
