taxonID	type	description	language	source
32452E2A6708582BFE79586C28587CE7.taxon	description	(Figs. 1 – 5)	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A6708582BFE79586C28587CE7.taxon	materials_examined	Type locality. Yemen, Socotra Island, Aloove area, Hassan village. Type material. HOLOTYPE: J (NMPC), labelled: ‘ YEMEN, SOCOTRA Island / Aloove area, HASSAN vill. env. / N 12 ° 30 ′ 58.2 ″, E 54 ° 06 ′ 39.2 ″ / 270 - 350 m, 3. - 4. ii. 2010 / L. Purchart lgt. ’. PARATYPES: 18 JJ 3 ♀♀, same data as holotype (1 ♀ NMPC, 1 J BMNH, 1 J HNHM, 1 J SMNS, 1 J ZSMC, 7 JJ 1 ♀ LPCB, 7 JJ 1 ♀ VNCP); 4 JJ, same data as holotype, but ‘ 300 m, at light / L. Purchart & J. Vybíral lgt. ’ (2 JJ LPCB, 2 JJ VNCP); 1 J: ‘ YEMEN, SOCOTRA Island / Kesa env., 220 - 300 m / 12 ° 39 ′ 37 ″ N, 53 ° 26 ′ 42 ″ E / 28. - 29. i. 2010, L. Purchart lgt. ’ (LPCB); 1 J, ‘ Yemen: Socotra Isl. / Ayhaft, 15.3.2000 / V. Bejček & K. Šťastný lgt. ’ (VNCP); 2 JJ: ‘ Yemen, Socotra Isl., Calanthia, / 29. - 30. iii. 2001, / leg. V. Bejček & K. Šťastný’ (VNCP); 1 J: ‘ SOCOTRA: Homhill / 23 - 24. II. 2009 - leg. P. / Lo Cascio & F. Grita’ (PLFG); 1 J: ‘ YEMEN, SOCOTRA Island / Aloove area, ALOOVE vill. env. / Jatropha unicostata shrubland with / Boswellia elongata trees / 19. - 20. vi. 2012 / 12 ° 31.2 ’ N, 54 ° 07.4 ’ E, 221 m // SOCOTRA expedition 2012, J. Bezděk, J. Hájek, V. Hula, P. Kment, I. Malenovský, J. Niedobová & L. Purchart leg. ’ (LPCB).	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A6708582BFE79586C28587CE7.taxon	description	Description. Male (holotype). Habitus as in Fig. 1. Body small, elongate, brown, with pale brown setation, BL 6.1 mm. Widest near elytral midlength; BL / EW 2.69. Head (Fig. 2) brown, with long, pale brown setation and microgranulation, slightly shiny, punctation distinct, punctures relatively large. HW 1.1 mm; HW / PW 0.69; HL 0.5 mm. Eyes dark, very large, transverse, deeply excised, space between eyes distinctly shorter than length of antennomere 2; OI equal to 13.42. Antennae (Fig. 3). Long, relatively narrow, unicoloured pale brown with microgranulation, punctation and pale brown setation, slightly shiny; AL 3.8 mm; AL / BL 0.62. Antennomere 2 shortest, antennomere 3 – 10 distinctly broadest at apex. RLA (1 – 11): 0.82: 0.47: 1.00: 1.11: 1.07: 1.02: 1.05: 1.13: 1.05: 0.93: 1.14. RL / WA (1 – 11): 2.30: 1.77: 3.39: 3.13: 2.66: 2.79: 2.46: 2.81: 2.67: 2.46: 3.19. Maxillary palpus. Pale brown with pale brown setation and microgranulation. Palpomeres 2 – 4 distinctly broadened from base to apex, slightly shiny. Ultimate palpomere broadly triangular. Pronotum (Fig. 2) unicoloured brown, elongate, distinctly longer than semicircular, bellshaped, with long, dense pale brown setation, setae directed backwards, closely covering body, with microgranulation and punctation. Punctures conspicuous, relatively large, coarse. PL 1.3 mm; PW 1.6 mm; PI equal to 81.25. Borders complete, pronotal base bisinuate. Posterior angles of pronotum rectangular, anterior angles indistinct. Elytra. Unicoloured brown, slightly oval, covered with dense, long, backward directed, closely adhering to body, pale brown setation. Broadest near midlength. EL 4.3 mm; EW 2.3 mm; EL / EW 1.87. Elytra with microgranulation and punctation; punctures relatively large. Rows of punctures in striae not clearly conspicuous, punctures in elytral interspaces smaller than in striae. Scutellum triangular, brown, with darker sides, covered with long, pale brown setae. Epipleura well developed, brown, with large punctures and pale brown setation, regularly narrowing from pronotal base to first abdominal ventrite, then continuing as parallel. Ventral part. Reddish-brown, with pale brown setation, punctation and microgranulation. Punctures large. Abdomen with shallow, medium-sized punctures. Legs. Unicoloured pale brown, covered with dense, short, pale brown setation. Tibiae and tarsi narrow, tibiae slightly dilated anteriorly. Penultimate tarsomere of each tarsus very slightly broadened, distinctly lobed. RLT: protarsus: 1.00: 0.49: 0.46: 0.55: 1.12; mesotarsus: 1.00: 0.54: 0.37: 0.35: 0.93; metatarsus: 1.00: 0.34: 0.29: 0.50. Both anterior tarsal claws with 10 visible teeth. Aedeagus (Figs. 4, 5). Pale brown, with microgranulation, slightly shiny. Basal part slightly rounded dorsally in lateral view, in dorsal view regularly narrowing laterally. Apical part (apex) short, in lateral view in form of long narrow triangle and dorsal view regularly narrowing laterally, with sharp tip. AEB / AEA 4.53. Female. Antennae slightly shorter than in male. Anterior tarsal claws with seven visible teeth. RLA (1 – 11): 0.76: 0.54: 1.00: 1.30: 1.11: 1.22: 1.24: 1.41: 1.19: 1.16: 1.30. RL / WA (1 – 11): 1.40: 1.54: 2.31: 2.67: 2.16: 2.50: 2.88: 2.89: 2.32: 2.39: 3.20. RLT: protarsus: 1.00: 0.56: 0.59: 0.72: 1.48; mesotarsus: 1.00: 0.63: 0.45: 0.51: 0.89; metatarsus: 1.00: 0.32: 0.35: 0.61. BL 6.1 mm; HL 0.5 mm; HW 1.0 mm; OI 17.05; PL 1.2 mm; PW 1.5 mm; PI 80.00; EL 4.4 mm; EW 2.4 mm; HW / PW 0.67; BL / EW 2.54; EL / EW 1.83; AL 3.6 mm; AL / BL 0.59. Variability. Measurements: mean (minimum – maximum). Males (n = 28). BL 5.2 mm (4.5 – 6.1 mm); HL 0.5 mm (0.4 – 0.6 mm); HW 1.0 mm (0.8 – 1.0 mm); OI 18.05 (13.42 – 24.95), PL 1.2 mm (1.0 – 1.3 mm) PW 1.4 mm (1.2 – 1.7 mm); PI 82.39 (76.53 – 88.48); EL 3.5 mm (3.0 – 4.4 mm); EW 1.9 mm (1.7 – 2.4 mm). Females (n = 3). BL 5.7 mm (5.2 – 6.0 mm); HL 0.5 mm (0.4 – 0.7 mm); HW 1.0 mm (0.9 – 1.0 mm); OI 19.27 (17.05 – 23.18), PL 1.1 mm (0.9 – 1.3 mm) PW 1.5 mm (1.4 – 1.6 mm); PI 75.25 (65.49 – 81.28); EL 4.1 mm (3.8 – 4.4 mm); EW 2.1 mm (1.9 – 2.4 mm).	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A6708582BFE79586C28587CE7.taxon	diagnosis	Differential diagnosis. Socotralia intermedia sp. nov. differs, besides characters stated in the key, from all similar species of Socotralia mainly in having space between eyes very narrow, distinctly narrower than length of antennomere 2, while other Socotralia species have space between eyes distinctly broader than length of antennomere 2.	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A6708582BFE79586C28587CE7.taxon	etymology	Etymology. Latin adjective intermedius, - a, - um (= intermediate), expressing the species body size, which is halfway within the size range of the genus Socotralia. Collection circumstances. Members of this species were collected during the day on herb layer – flowering plants of the family Fabaceae. They were also attracted to the light.	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A6708582BFE79586C28587CE7.taxon	distribution	Distribution. Yemen: Socotra Island.	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A6709582BFE105C6428967E17.taxon	materials_examined	Material examined. 8 JJ 5 ♀♀, ‘ YEMEN, SOCOTRA Island E / Homhil area, 400 - 510 m / N 12 ° 34 ′ 25 ″, E 54 ° 18 ′ 53 ″ / 9. - 10. ii. 2010 / L. Purchart & J. Vybíral lgt. ’ (1 ♀ NMPC, 4 JJ 2 ♀♀ LPCB, 4 JJ 2 ♀♀ VNCP); 2 JJ, ‘ YEMEN, SOCOTRA Island / Diksam plateau, Bidehor, Digeila / Cave env., 920 m, 8. ii. 2010 / N 12 ° 30 ′ 31 ″, E 53 ° 56 ′ 18 ″ / L. Purchart & J. Vybíral lgt. ’ (1 J LPCB, 1 J VNCP); 2 JJ 2 ♀♀, ‘ SOCOTRA: W. Ayheft / 28. II- 1. III. 2009 – leg. P. / Lo Cascio & F. Grita’ (PLFG); 1 J, ‘ Coll. I. R. Sc. N. B. / N. SOKOTRA isld., / Top of Ayheft valley / 17 - I- 2010 / Leg Saldaitis / Achat Saldaitis. I. G 31.512 ’ (IRSNB).	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A670E582CFE7B59A52E607B10.taxon	materials_examined	Material examined. 4 JJ 5 ♀♀, ‘ YEMEN, SOCOTRA Island E / Homhil area, 400 - 510 m / 12 ° 34 ′ 25 ″ N, 54 ° 18 ′ 53 ″ E / 9. - 10. ii. 2010 / L. Purchart lgt. ’ (1 ♀ NMPC, 4 JJ 4 ♀♀ LPCB); 2 JJ, ‘ YEMEN, SOCOTRA Island E / Homhil area, 400 - 510 m / N 12 ° 34 ′ 25 ″, E 54 ° 18 ′ 53 ″ / 9. - 10. ii. 2010 / L. Purchart & J. Vybíral lgt. ’ (LPCB); 1 J, ‘ YEMEN, Socotra Isl., / Deiqub cave env., / 10. vi. 2010, / V. Hula & J. Niedobová leg. ’ (LPCB); 1 J, ‘ YEMEN, SOCOTRA Island / Zemhon, 260 - 320 m / N 12 ° 32 ′ 17.5 ″, E 54 ° 4 ′ 12.7 ″ / 20. vi. 2009 / L. Purchart lgt. ’ (LPCB); 1 J, ‘ YEMEN, SOCOTRA Island / Aloove area, HASSAN vill. env. / N 12 ° 30 ′ 58.2 ″, E 54 ° 06 ′ 39.2 ″ / 270 - 350 m, 3. - 4. ii. 2010 / L. Purchart lgt. ’ (LPCB); 1 J, ‘ Republic of Yemen / Socotra Isl. 16. - 17.6.2010 / Aloove area, 270 - 300 m a. s. l. / N 12 ° 30.58 ′, E 054 ° 06.39 ′ / lgt. V. Hula & J. Niedobová’ (LPCB); 2 JJ 1 ♀, ‘ YEMEN, SOCOTRA Island, S / Noged plain, Deiqyub Cave / 16. vi. 2009 / L. Purchart & J. Vybíral lgt. ’ (1 J 1 ♀ LPCB, 1 J VNCP); 2 JJ, ‘ YEMEN, SOCOTRA Island / Aloove area, HASSAN vill. env. / N 12 ° 30 ′ 58.2 ″, E 54 ° 06 ′ 39.2 ″ / 300 m, 9. - 10. ii. 2010, at light / L. Purchart & J. Vybíral lgt. ’ (1 J LPCB, 1 J VNCP); 27 JJ 36 ♀♀, ‘ YEMEN, SOCOTRA Island / Aloove area, 270 - 350 m / N 12 ° 30 ′ 58 ″, E 54 ° 06 ′ 39 ″ / 16. - 17. vi. 2010, V. Hula lgt. // window trap / Installed 4. ii. 2010 / by L. Purchart / medium – ethylenglycol 98 % ’ (1 J 1 ♀ BMNH, 1 J 1 ♀ HNHM, 1 J 1 ♀ NMPC, 1 J 1 ♀ SMNS, 1 J 1 ♀ ZSMC, 12 JJ 16 ♀♀ LPCB, 10 JJ 15 ♀♀ VNCP); 3 JJ, ‘ YEMEN, SOCOTRA Island / Firmihin, 400 - 500 m / N 12 ° 28 ′ 27.9 ″, E 54 ° 0 ′ 54.2 ″ / 22. - 25. vi. 2009, L. Purchart lgt. ’ (2 JJ LPCB, 1 J VNCP); 1 J, ‘ Yemen, Soqotra Is., 2003 / 2 - 3 / xii., Dixam plateau, / WADI ESGEGO, 300 m / N 12 ° 28 ′ 09 ″ E 54 ° 00 ′ 36 ″ / [GPS], David Král lgt. // YEMEN – SOQOTRA 2003 / Expedition; Jan Farkač, Petr Kabátek & David Král’ (NMPC); 4 JJ, ‘ YEMEN, SOCOTRA Island / Dixam plateau / Firmihin (Dracaena forest) / 12 ° 28.6 ′ N, 54 ° 01.1 ′ E, 490 m / P. Hlaváč leg., 15 - 16. xi. 2010 ’ (NMPC).	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A670E5820FE475A5428737A07.taxon	description	(Figs. 6 – 10)	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A670E5820FE475A5428737A07.taxon	materials_examined	Type locality. Yemen, Socotra Island, Al Haghier Mts., Scand Mt. env. Type material. HOLOTYPE: J (NMPC), labelled: ‘ YEMEN, SOCOTRA Island / Al Haghier Mts. / Scant Mt. env. / 12 ° 34.6 ′ N, 54 ° 01.5 ′ E, 1450 m / 12. - 13. xi. 2010, J. Bezděk leg. ’. PARATYPES: 1 J, same data as holotype, but J. Hájek leg. (NMPC); 1 J, same data as holotype, but P. Hlaváč lgt. (VNCP); 4 JJ, same data as holotype, but J. Batelka leg. (3 J J JBCP, 1 J LPCB); 3 ♀♀, ‘ YEMEN, SOCOTRA Island / Al Haghier Mts. / wadi Madar, 1180 - 1230 m / 12 ° 33.2 ′ N, 54 ° 00.4 ′ E / 13. - 14. xi. 2010, P. Hlaváč lgt. ’ (1 ♀ VNCP, 1 ♀ LPCB, 1 ♀ JBCP).	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A670E5820FE475A5428737A07.taxon	description	Description. Male (holotype). Habitus as in Fig. 6. Body relatively large, elongate, narrow, from pale reddish-brown to dark blackish-brown, with pale brown setation, slightly shiny, BL 7.4 mm. Widest near midlength of elytra; BL / EW 3.08. Head (Fig. 7) relatively small, reddish-brown, with sparse, pale brown setation and microgranulation, dull. HW 1.2 mm; HW / PW 0.71; HL 0.9 mm. Eyes dark, large, transverse, deeply excised, space between eyes approximately as broad as length of antennomere 1, distinctly broader than length of antennomere 2; OI equal to 30.09. Antennae (Fig. 8). Long, relatively narrow with pale reddish-brown setation; antennomeres 1 and 2 reddish-brown, antennomere 3 bicolored, antennomeres 4 – 11 dark blackish-brown with distinctly paler apex. AL 4.5 mm, AL / BL 0.61. Antennomere 2 shortest, antennomere 3 distinctly longer than antennomere 2. RLA (1 – 11): 0.55: 0.33: 1.00: 1.16: 1.01: 0.99: 1.02: 1.03: 0.94: 0.85: 0.88. RL / WA (1 – 11): 1.85: 1.59: 3.57: 4.13: 3.60: 3.65: 3.88: 3.92: 3.48: 3.16: 3.24. Maxillary palpus. Pale brown with sparse, short, pale brown setation and a few long pale brown setae. Palpomeres 2 – 4 with microgranulation, distinctly broadened from base to apex. Ultimate palpomere broadly triangular, with darker posterior half. Pronotum (Fig. 7). Narrower than base of elytra, blackish-brown with sides narrowly reddish-brown, covered with pale brown setation, microgranulation and punctation, punctures small. PL 1.2 mm; PW 1.7 mm; PI equal to 70.59. Borders complete but not clearly conspicuous. Base of pronotum finely bisinuate, on antescutellar area straight. Posterior angles finely obtuse, anterior angles indistinct. Elytra. Narrow, parallel, distinctly shiny, blackish-brown, covered with dense pale brown setation, microgranulation and punctation. Broadest near midlength. EL 5.4 mm; EW 2.4 mm; EL / EW 2.25. Elytral striae indistinct. Elytral surface with rows of medium-sized punctures. Scutellum small, triangular, reddish-brown with sides dark blackish-brown, covered with pale brown setae. Elytral epipleura well-developed, blackish-brown, covered with pale brown setation, regularly narrowing from elytral base to first abdominal ventrite, then continues as parallel. Ventral aspect. Pale reddish-brown, with pale brown setation and punctation. Metaventrite and abdominal ventrites 1 – 3 partly dark brown. Abdomen reddish-brown with pale brown setation and punctation, slightly shiny. Legs. Narrow, brown, covered with dense pale brown setation, anterior part of femora pale brown, posterior part dark brown. Tibiae and tarsi narrow, tibiae slightly dilated anteriorly. Penultimate tarsomere of each tarsus very slightly broadened and distinctly lobed. RLT: protarsus: 1.00: 0.80: 0.86: 1.00: 1.57; mesotarsus: 1.00: 0.47: 0.40: 0.61: 0.84; metatarsus: 1.00: 0.45: 0.42: 0.60. Both anterior tarsal claws with 15 visible teeth. Aedeagus (Figs. 9, 10). Pale brown, slightly shiny. In dorsal view basal part long, shortly rounded, then parallel laterally. In lateral view basal half of basal part rounded dorsally and apical half slightly narrowing dorsally. Apical part (apex) short, covered with short spines, in lateral view in form of long narrow triangle and in dorsal view broadly cross-shaped, respectively. AEB / AEA 3.47. Female. Anterior tarsal claws with nine visible teeth. RLA (1 – 11): 0.55: 0.36: 1.00: 1.23: 1.10: 1.07: 1.05: 1.11: 1.05: 0.97: 1.02. RL / WA (1 – 11): 1.79: 1.47: 3.65: 3.62: 3.24: 3.00: 2.71: 3.29: 3.10: 3.00: 3.32. RLT: protarsus: 1.00: 0.83: 1.00: 0.96: 1.39; mesotarsus: 1.00: 0.57: 0.53: 0.64: 0.97; metatarsus: 1.00: 0.41: 0.35: 0.59. BL 8.2 mm; HL 1.0 mm; HW 1.2 mm; OI 31.62; PL 1.3 mm; PW 2.0 mm; PI 65.00; EL 5.9 mm; EW 2.8 mm; HW / PW 0.60; BL / EW 2.93; EL / EW 2.11; AL 4.4 mm; AL / BL 0.54. Variability. Measurements: mean (minimum – maximum). Males (n = 2). BL 7.35 mm (7.3 – 7.4 mm); HL 0.85 mm (0.8 – 0.9 mm); HW 1.15 mm (1.1 – 1.2 mm); OI 34.06 (30.09 – 38.02), PL 1.15 mm (1.1 – 1.2 mm) PW 1.65 mm (1.6 – 1.7 mm); PI 69.67 (68.75 – 70.59); EL 5.55 mm (5.2 – 5.9 mm); EW 2.35 mm (2.3 – 2.4 mm). Females (n = 3). BL 8.10 mm (8.00 – 8.2 mm); HL 0.95 mm (0.9 – 1.00 mm); HW 1.19 mm (1.1 – 1.2 mm); OI 34.64 (31.62 – 37.07), PL 1.27 mm (1.2 – 1.3 mm) PW 1.93 mm (1.9 – 2.00 mm); PI 65.88 (63.16 – 68.54); EL 5.85 mm (5.7 – 5.9 mm); EW 2.72 mm (2.7 – 2.8 mm).	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A670E5820FE475A5428737A07.taxon	diagnosis	Differential diagnosis (for more details see the key below). Socotralia montana sp. nov. differs from the similar species S. vybirali sp. nov. mainly in having antennomeres 3 – 10 relatively narrow while in S. vybirali sp. nov. antennomeres 3 – 10 are slightly serrate and distinctly broadest at apex. From S. intermedia sp. nov., S. reitteri sp. nov., S. major and S. minor it differs mainly in having antennomere 11 distinctly shorter than antennomere 3, while in the previous species the antennomere 11 is distinctly longer than antennomere 3. Socotralia montana sp. nov. can be distinguished from S. brunnea by its narrow, elongate body and parallel pronotum, while in S. brunnea the body is slightly oval and pronotum more transverse and rounded.	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A670E5820FE475A5428737A07.taxon	etymology	Etymology. Latin adjective montanus, - a, - um (= montane), expressing the species affinity to high (mountain) altitudes. Collection circumstances. The adults were collected at night by shaking off the shrub vegetation. They were also attracted to light.	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A670E5820FE475A5428737A07.taxon	distribution	Distribution. Yemen: Socotra Island.	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A67025821FE535B4428587E20.taxon	description	(Figs. 11 – 15)	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A67025821FE535B4428587E20.taxon	materials_examined	Type locality. Yemen, Socotra Island, Firmihin. Type material. HOLOTYPE: J (NMPC), labelled: ‘ YEMEN, SOCOTRA Island / Firmihin, 400 - 500 m / N 12 ° 28 ′ 27 ″, E 54 ° 0 ′ 54 ″ / 6. - 7. ii. 2010 at light / L. Purchart & J. Vybíral lgt. ’ PARATYPES: 3 JJ 5 ♀♀, same data as holotype (1 ♀ NMPC, 1 J 2 ♀♀ LPCB, 2 JJ 2 ♀♀ VNCP).	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A67025821FE535B4428587E20.taxon	description	Description. Male (holotype). Habitus as in Fig. 11. Body small, elongate, from pale reddish-brown to dark blackish-brown, with pale brown setation, BL 4.6 mm. Widest near elytral midlength; BL / EW 2.56. Head (Fig. 12). Small, reddish-brown, with long, pale brown setation and microgranulation, slightly shiny. Anterior aspect and clypeus distinctly paler, punctation conspicuous, punctures relatively large. HW 0.9 mm; HW / PW 0.64; HL 0.5 mm. Eyes dark, large, transverse, deeply excised, space between eyes distinctly broader than antennomere 2 is long; OI equal to 22.28. Antennae (Fig. 13). Long, with microgranulation, punctation and pale brown setation, bicoloured, with antennomeres 1 – 2 and base of antennomere 3 pale reddish-brown, slightly shiny, anterior half of antennomere 3 and antennomeres 4 – 11 black, duller. AL 3.2 mm, AL / BL 0.70. Antennomere 2 shortest, antennomere 3 – 10 distinctly broader at apex. RLA (1 – 11): 0.77: 0.51: 1.00: 1.22: 1.20: 1.24: 1.20: 1.33: 1.31: 1.31: 1.45. RL / WA (1 – 11): 1.93: 1.56: 2.58: 2.46: 2.38: 2.38: 2.61: 2.51: 2.69: 2.64: 3.50. Maxillary palpus. Pale reddish-brown with pale brown setation and microgranulation. Palpomeres 2 – 4 distinctly broadened from base to apex, slightly shiny. Ultimate palpomere broadly triangular. Pronotum (Fig. 12) unicoloured reddish-brown, slightly longer than semicircular, bellshaped, with long, pale brown setation, setae directed backwards, with microgranulation and punctation. Punctures conspicuous, relatively large. PL 1.0 mm; PW 1.4 mm; PI equal to 70.43. Borders complete, pronotal base bisinuate, on antescutellar area straight. Posterior angles rectangular, anterior angles indistinct. Ventral part. Reddish-brown, with pale brown setation, microgranulation and punctation, punctures large and coarse. Elytra unicoloured dark blackish-brown, elongate, covered by dense, long, backward directed, pale brown setation. EL 3.2 mm. Broadest near elytral two thirds, EW 1.8 mm; EL / EW 1.78. Elytra with microgranulation and punctation; punctures relatively large. Rows of punctures in elytral striae not clearly conspicuous. Scutellum triangular, pale reddish-brown with pale brown setae and sparse microgranulation, shiny. Elytral epipleura well developed, pale reddish-brown, with large punctures and pale brown setation, regularly narrowing from elytral base to abdominal ventrite 1, then continues as parallel. Legs. Unicoloured pale reddish-brown, with dense, short, pale brown setation. Tibiae, tarsi narrow, tibiae slightly dilated anteriorly. Penultimate tarsomere of each tarsus very slightly broadened, distinctly lobed. RLT: protarsus: 1.00: 0.72: 0.72: 0.80: 1.66; mesotarsus: 1.00: 0.44: 0.36: 0.48: 0.91; metatarsus: 1.00: 0.33: 0.30: 0.54. Both anterior tarsal claws with eight visible teeth. Aedeagus (Figs 14, 15). Pale brown, slightly shiny. Basal part rounded in lateral view, in dorsal view regularly narrowing anteriorly. Apical part (apex) short, in lateral view narrowly triangular, in dorsal view in form of a long triangle. AEB / AEA 3.54. Female. More oval, antennae distinctly shorter in than male. Anterior tarsal claws with five visible teeth. RLA (1 – 11): 0.71: 0.40: 1.00: 1.11: 1.09: 1.22: 1.31: 1.36: 1.27: 1.22: 1.29. RL / WA (1 – 11): 1.95: 1.00: 2.29: 2.26: 1.94: 2.03: 2.06: 2.21: 1.76: 1.91: 2.09. RLT: protarsus: 1.00: 0.62: 0.76: 0.97: 1.76; mesotarsus: 1.00: 0.49: 0.44: 0.52: 0.88; metatarsus: 1.00: 0.35: 0.32: 0.48. BL 4.4 mm; HL 0.5 mm; HW 0.8 mm; OI 29.86; PL 0.8 mm; PW 1.2 mm; PI 66.67; EL 3.2 mm; EW 1.7 mm; HW / PW 0.67; BL / EW 2.59; EL / EW 1.88; AL 2.6 mm; AL / BL 0.59. Variability. Measurements: mean (minimum – maximum). Males (n = 4). BL 4.5 mm (4.1 – 4.9 mm); HL 0.5 mm (0.4 – 0.6 mm); HW 0.8 mm (0.7 – 0.9 mm); OI 25.44 (22.28 – 28.85), PL 0.9 mm (0.8 – 1.0 mm) PW 1.2 mm (1.1 – 1.4 mm); PI 70.39 (68.61 – 71.59); EL 3.1 mm (2.8 – 3.5 mm); EW 1.7 mm (1.4 – 1.9 mm). Females (n = 6). BL 4.3 mm (4.00 – 4.5 mm); HL 0.4 mm (0.4 – 0.5 mm); HW 0.75 mm (0.7 – 0.8 mm); OI 33.10 (29.86 – 36.25), PL 0.8 mm (0.7 – 0.8 mm) PW 1.2 mm (1.1 – 1.3 mm); PI 66.89 (64.31 – 69.36); EL 3.1 mm (2.9 – 3.2 mm); EW 1.7 mm (1.5 – 1.9 mm).	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A67025821FE535B4428587E20.taxon	diagnosis	Differential diagnosis (for more details see the key below). Socotralia reitteri sp. nov. differs from similar species S. montana sp. nov., S. vybirali sp. nov. and S. brunnea mainly in having antennomere 11 distinctly longer than antennomere 3, while S. montana sp. nov., S. vybirali sp. nov. and S. brunnea have antennomere 11 shorter than antennomere 3. S. reitteri sp. nov. differs from the similar species S. intermedia sp. nov. mainly in having space between eyes broader than length of antennomere 2, while S. intermedia sp. nov. has space between eyes distinctly shorter than length of antennomere 2. S. reitteri sp. nov. differs from the similar species S. major in having antennomere 3 – 10 distinctly serrate, while S. major has antennomeres 3 – 10 narrower. S. reitteri sp. nov. differs from the similar species S. minor in having sides of pronotum conically narrowing, while S. minor has rounded sides of pronotum.	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A67025821FE535B4428587E20.taxon	etymology	Etymology. Named in honour of Edmund Reitter (1845 – 1920), world famous expert in Coleoptera. Collecting circumstances. Members of this species were collected only at night when they were attracted to light. They were not found during the day on herb or shrub vegetation.	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A67025821FE535B4428587E20.taxon	distribution	Distribution. Yemen: Socotra Island.	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A67005823FE4F59A528597CA0.taxon	description	(Figs. 16 – 20)	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A67005823FE4F59A528597CA0.taxon	materials_examined	Type locality. Yemen, Socotra Island, Homhil area. Type material. HOLOTYPE: J (NMPC), labelled: ‘ YEMEN, SOCOTRA Island / Homhil area, 400 - 510 m / 12 ° 34 ′ 25 ″ N 54 ° 18 ′ 53 ″ E / 9. - 10. i. 2010, at light / L. Purchart & J. Vybíral lgt. ’. PARATYPES: 5 JJ 4 ♀♀, same data as holotype (1 ♀ NMPC, 2 JJ 2 ♀♀ LPCB, 3 JJ 1 ♀ VNCP); 2 ♀♀, ‘ YEMEN, SOCOTRA Island / Shibhon plateau / ESERHE, 13. vi. 2012 / Croton socotranus shrubland / 12 ° 25.2 ’ N, 53 ° 56.6 ’ E, 547 m // SOCOTRA expedition 2012, J. Bezděk, J. Hájek, V. Hula, P. Kment, I. Malenovský, J. Niedobová & L. Purchart leg. ’ (1 ♀ NMPC, 1 ♀ LPCB).	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A67005823FE4F59A528597CA0.taxon	description	Description. Male (holotype). Habitus as in Fig. 16. Body small, elongate, from pale brown to dark blackish-brown, with pale brown setation, BL 4.5 mm. Widest near two thirds of elytral length; BL / EW 2.65. Head (Fig. 17). Relatively narrow, small, brown, with long, pale brown setation and microgranulation, slightly shiny, punctation conspicuous, punctures relatively large. HW 0.9 mm; HW / PW 0.56; HL 0.4 mm. Eyes dark, very large, transverse, deeply excised, space between eyes approximately as long as antennomere 2; OI equal to 20.05. Antennae (Fig. 18). Shorter, relatively narrow; pale brown with microgranulation, punctation and pale brown setation, slightly shiny; AL 2.4 mm, AL / BL 0.53. Antennomere 2 shortest, antennomere 3 - 10 distinctly broadest at apex. Antennomere 3 and 4 distinctly longer than each of antennomeres 5 – 11. RLA (1 – 11): 0.61: 0.57: 1.00: 1.06: 0.98: 1.04: 0.97: 1.06: 0.98: 0.95: 0.96. RL / WA (1 – 11): 1.61: 1.97: 3.45: 2.87: 2.58: 2.48: 2.26: 2.26: 2.09: 1.86: 2.26. Maxillary palpus. Pale brown with pale brown setation and microgranulation. Palpomeres 2 – 4 distinctly broadened from base to apex, slightly shiny. Ultimate palpomere broadly triangular. Pronotum (Fig. 17) unicoloured dark reddish-brown, transverse, with long, dense, pale brown setation, setae directed backwards, closely covering to body, with punctation, microgranulation not clearly conspicuous. Punctures conspicuous, relatively large and coarse. PL 0.9 mm; PW 1.6 mm; PI equal to 56.25. Borders complete, base bisinuate, on antescutellar area straight. Posterior angles roundly rectangular, anterior angles indistinct, sides slightly rounded, in posterior half parallel. Ventral side of body. Brown, with pale brown setation. Abdomen dark brown with pale brown setation, microgranulation and punctation, punctures relatively large, sparse. Elytra unicoloured dark blackish-brown, slightly oval, shiny, covered by dense, long, backward directed, body closely covering, pale brown setation. EL 3.1 mm. Elytra broadest near midlength, EW 1.7 mm. EL / EW 1.82. Elytra with very fine microgranulation and punctation; punctures in elytral striae and elytral interspaces of approximately same diameter, relatively large. Rows of punctures in striae not clearly conspicuous. Scutellum triangular, pale brown, paler than elytra, with sides darker, with microgranulation and long, pale brown setae. Elytral epipleura well-developed, at base pale brown, then brown, with large punctures, pale brown setation and distinct microgranulation, regularly narrowing from pronotal base to abdominal ventrite 1, then continues as parallel. Legs. Unicoloured pale reddish-brown, with dense, pale brown setation. Tibiae, tarsi narrow, tibiae slightly dilated anteriorly. Penultimate tarsomere of each tarsus very slightly broadened and distinctly lobed. RLT: protarsus: 1.00: 0.90: 0.68: 0.84: 2.00; mesotarsus: 1.00: 0.61: 0.34: 0.42: 0.87; metatarsus: 1.00: 0.28: 0.22: 0.38. Anterior tarsal claws with nine visible teeth. Aedeagus (Figs. 19, 20). Pale brown, with fine microgranulation, slightly shiny. Basal part rounded in lateral view and in dorsal view regularly narrowing. Apex short, in dorsal view slightly broadening from base towards end of 1 / 3 then narrowing anteriorly, without sharp tip; in lateral view slightly bent upwards. AEB / AEA 4.82. Female. More oval. Anterior tarsal claws with 5 – 7 teeth. RLA (1 – 11): 0.64: 0.61: 1.00: 1.14: 1.04: 1.07: 1.05: 1.05: 1.00: 0.89: 1.07. RL / WA (1 – 11): 1.44: 1.42: 1.81: 2.00: 1.71: 1.77: 1.74: 1.79: 1.81: 1.43: 1.82. BL 4.4 mm; HL 0.4 mm; HW 0.9 mm; OI 24.43; PL 0.8 mm; PW 1.3 mm; PI 61.54; EL 3.2 mm; EW 1.7 mm; HW / PW 0.69; BL / EW 2.59; EL / EW 1.88; AL 2.1 mm; AL / BL 0.48. Variability. Measurements: mean (minimum – maximum). Males (n = 6). BL 4.5 mm (4.2 – 4.6 mm); HL 0.4 mm (0.3 – 0.5 mm); HW 0.8 mm (0.8 – 0.9 mm); OI 21.01 (17.34 – 26.35), PL 0.9 mm (0.8 – 0.9 mm) PW 1.3 mm (1.2 – 1.6 mm); PI 66.85 (62.45 – 70.16); EL 3.1 mm (3.0 – 3.2 mm); EW 1.7 mm (1.6 – 1.8 mm). Females (n = 4). BL 4.4 mm (4.3 – 4.7 mm); HL 0.5 mm (0.4 – 0.6 mm); HW 0.8 mm (0.8 – 0.9 mm); OI 29.94 (24.43 – 34.53), PL 0.8 mm (0.8 – 0.9 mm) PW 1.3 mm (1.2 – 1.5 mm); PI 62.42 (61.27 – 64.86); EL 3.1 mm (3.0 – 3.3 mm); EW 1.8 mm (1.7 – 1.9 mm).	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A67005823FE4F59A528597CA0.taxon	diagnosis	Differential diagnosis (for more details see the key below). Socotralia vybirali sp. nov. differs from similar species S. intermedia sp. nov., S. reitteri sp. nov., S. major and S. minor mainly in having antennomere 11 shorter than antennomere 3, while S. intermedia sp. nov., S. reitteri sp. nov., S. major and S. minor have antennomere 11 distinctly longer than antennomere 3. S. vybirali sp. nov. differs from similar species S. montana sp. nov. and S. brunnea mainly in smaller body length and distinctly serrate antennomeres 3 – 10, while in S. montana sp. nov. and S. brunnea the body is large and antennomeres 3 – 10 are narrower.	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A67005823FE4F59A528597CA0.taxon	etymology	Etymology. Named in honour of Jan Vybíral (Židlochovice, Czech Republic), one of the collectors of the type series. Collection circumstances. Specimens of the new species were collected at night when they were attracted to light. Two specimens (Eserhe) were found also during the day on herb and shrub vegetation.	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
32452E2A67005823FE4F59A528597CA0.taxon	distribution	Distribution. Yemen, Socotra Island.	en	Novák, Vladimír, Purchart, Luboš (2012): New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae). Acta Entomologica Musei Nationalis Pragae 52: 323-336, DOI: 10.5281/zenodo.5338404
