taxonID	type	description	language	source
32553F1BFF91FFA4FF3E73FA28D5DACE.taxon	discussion	Remarks. The species of the genus were revised in detail by Ahrens 2000, with subsequent additions of Ahrens (2002), Ahrens & Pacholátko (2003, 2007), Liu et al. (2011, 2014), and Kobayashi (2015). Based on previous evidence from a molecular phylogeny (Eberle et al. 2017) and this new comprehensive morphological and taxonomical revision, a part of the species so far assigned to the genus Microserica Brenske, 1894, have to be classified as Gastroserica, although key characters of this genus are not present (see below).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9EFFA5FF3E759629EAD996.taxon	type_taxon	Type species of the new subgenus: Microserica varians Moser, 1915 (by current designation).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9EFFA5FF3E759629EAD996.taxon	diagnosis	Diagnosis. The new subgenus differs from its sister clade (Fig. 1), Gastroserica sensu stricto, by the well-produced and distinctly angled anterior angles of pronotum, as well as by the distinctly emarginated anterior margin of labroclypeus. Furthermore, the apical abdomen is not as robustly developed as in most Gastroserica sensu stricto species, with the pygidium barely protruding under the apex of elytra.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9EFFA5FF3E759629EAD996.taxon	discussion	Remarks. Given, of being the sister lineage of Gastroserica (Fig. 1; see also Eberle et al. 2017), and thus representing a monophyletic group with the species of Gastroserica sensu stricto, we nevertheless decided to erect a new and separate subgenus for these species, since key diagnostic features of Gastroserica sensu stricto are not present among these species, namely the obsolete anterior angles of pronotum and the straight, non-emarginate anterior margin of labroclypeus. All species so far classified as Gastroserica (Ahrens & Bezděk 2016) are subsequently assigned to the subgenus Gastroserica Brenske, 1897.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9EFFA5FF3E759629EAD996.taxon	etymology	Etymology. The name of the new subgenus is composed of the Greek word “ helios ” (sun) and the word root Serica (type genus of the tribe Sericini), with reference to the diurnal activity of the species of this subgenus.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9FFFA7FF3E77202D0FDFCF.taxon	description	Figs. 2 A – D, 26	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9FFFA7FF3E77202D0FDFCF.taxon	materials_examined	Type material examined. Syntypes: 1 Ô “ China Yunnan / Microserica varians Type Ô Mos. ” (ZMHB), 1 ♀ “ China Yunnan / Microserica varians Type ♀ Mos. ” (ZMHB), 5 ÔÔ, 2 ♀♀ “ China Yunnan ” (ZMHB), 1 Ô “ Yünnan China ” (ZMHB). Additional material examined. China: 2 ex. [China] “ Yun-nan sen ” (ZMHB), 2 ex. “ Cine Yunnan Recu de Lou-Nan 1931 ” (MNHN), 7 ex. [China] “ Yunnan 2000 - 2800 m 25.11 N 100.24 E Weibaoshan mts. W slope 25 - 28 / 6.92 Vit Kubáň leg. ” (CN), 11 ex. “ China Yunnan 1800 m Lijiang 23. VI. - 21. VII. 26 ° 53 ’ N 100 ° 18 ’ E E. Jendek leg. 1992 ” (NHMW), 43 ex. “ China Yunnan 22.5. - 2.6. 100 km W Kunming Diaolin Nat. Res. 1993 E. Jendek & O. Sausa leg. [sic] ” (NHMW), 1 ex. “ CH-Yunnan 13 - 14. VI. 1995 100 km W of Kunming Dialin Nat. Reservation E. Jendek & O. Sausa leg. ” (CP), 1 Ô “ CH-Yunnan 1995 Diaolin 13 - 14. VI. E. Jendek leg. ” (CP), 12 ex. “ China, Yunnan, 19. VII. 1993 Dali, Hua Dian Ba 50 km N of Dali C. Holzschuh leg. / CS 4 ” (CP), 7 ex. “ China Yunnan, 1800 m Lijiang 23.6. - 21.7. 26.53 N 100.18 E E. Jendek leg. 1992 / CS 16 ” (CP), 3 ex. “ Coll. R. I. Sc. N. B. Chine / Sud Yunnan Tche-Ping-Tcheou / Coll. P. Guerry Le Moult vendit ” (ISNB), 1 ex. “ China: N-Yunnan Baiyungshan (Bai Railing Mts.) 2400 m Yong Ren, VII- 2003 leg. Ying et al. ” (ZFMK), 7 ex. “ China, Yunnan, 19. VII. 1993 Dali, Hua Dian Ba 50 km N of Dali C. Holzschuh leg. ” (CP), 3 ex. “ CH-Yunnan, 13 - 14. VI. 1995 100 km W of Kunming Diaolin Nat. Reservation E. Jendek & O. Sausa leg. ” (CP), 10 ex. “ China, W. Yunnan, Diaolin, 18. VI. 1993 E. Jendek & O. Sausa leg. ” (CP), 2 ex. [China] “ Yunnan 2500 - 2700 m, 25.58 N 100.21 E Juzu Shan 6 - 10.7. Vit Kuban leg. 1994 ” (CP), 23 ex. [China] “ Yunnan 1800 - 2000 m 25.04 N 101.55 E Vipinglang 8 - 10.6. Vit Kuban leg. 1994 ” (CP), 21 ex. [China] “ Yunnan 1800 - 2000 m 25.04 N 101.55 E Vipinglang 17 - 20.6. Vit Kuban leg. 1994 ” (CP), 7 ex. [China] “ Yunnan, 1700 m Yipinglang 28 - 29. Jul 1995 BOLM lgt. ” (CP), 1 ex. “ China Yunnan Lijiang 2100 m 13. - 27. vi. 1995 leg. Pekarovic ” (CP), 1 ex. “ CH-Yunnan 1995 Diaolin 13 - 14. VI. E. Jendek leg. ” (CP), 1 ex. “ Kunmin China 61 lgt. S. Skopek / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 7 ex. “ China (Yunnan) Dali Bai Aut. Pref. Wuliang Shan, 9 km SW Weishan, 2450 - 2500 m, 25 ° 10 ’ 14 ’’ N / 100 ° 14 ’ 22 ’’ E (sec. Oak / pine for., beaten from trees and bushes) 13. VI. 2007 D. W. Wrase [350] ” (ZFMK), 3 ÔÔ [China] “ Heilinpu, Kunming, Yunnan, 23. Ⅴ. 1941 ” (CAU), 1 Ô [China] “ Kunming, Yunnan, 16. V. 1956, 1900, leg. Kryschanovsknja ” (IZAS), 1 Ô [China] “ Defu, Napo, Guangxi, 18. VI. 2000, 1350 m, leg. Li Wenzhu ” (IZAS), 2 ÔÔ [China] “ Mengsong, Xishuangbanna, Yunnan, 26. IV. 1958, 1600 m, leg. Zhang Yiran / LW- 359 ” (IZAS), 1 Ô “ China: S-Yunnan (Xishuangbanna) ca. 30 km NW Jinghong vic. Bameng, 17 - 2000 m / Hua Zhuliangzi Mts. N 22 ° 08.01 / E 100 ° 31.54 ’ 1700 - 2000 m, 30. V. 2008 leg. A. Weigel sec. forest ” (NME), 2 ÔÔ “ CHINA: S- Yunnan (Xishuangbanna) 45 km SW Jinghong vic. Bangzhang vill. / N 21 ° 44 ’ 37 ’’, E 100 ° 27 ’ 02 ’’, 16 - 1700 m, 03. - 05. V. 2009 leg. A. Weigel, blüh. Cast ” (NME). Laos: 1 ex. “ LAOS, Xieng Khouang prov., 19 ° 37 – 8 ’ N, 103 ° 20 ’ E, Phonsavan (30 km NE) Phou Sane Mt., 1420 m, 10 - 30. v. 2009, Z. Kraus leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 1 ex. “ LAOS, Xieng Khouang prov., 19 ° 37 – 8 ’ N, 103 ° 20 ’ E, Phonsavan (30 km NE) Phou Sane Mt., 1420 m, 10 - 30. v. 2009, M. Brancucci leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 18 ex. “ LAOS-NE, Houa Phan prov., Ban Saluei-> Phou Pane Mt., 20 ° 12 - 13.5 ’ N 103 ° 59.5 ’ - 104 ° 01 ’ E, 1340 - 1870 m, 15. iv. - 15. v. 2008, Lao collector leg. ” (NMPC), 3 ex. “ LAONE, Hua Phan prov., 20 ° 12 ’ N 104 ° 01 ’ E, Phu Phan Mt., 1500 - 1900 m, 17. v. - 3. vi. 2007, M. Brancucci leg. / NHMB Basel expedition to Laos, 2007 ” (NHMB), 7 ex. “ LAOS, Houa Phan prov., 20 ° 12 - 13 ’ N, 103 ° 59.5 ’ - 104 ° 01 ’ E, Ban Saluei-> Phou Pane Mt., 1340 - 1870 m, 10. v. - 16. vi. 2009, M. Brancucci & local coll. leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 4 ex. “ LAOS, Hua Phan prov., 20 ° 13 ’ N, 104 ° 00 ’ E, Phou Pane Mt., 1 - 16. vi. 2009, 1350 - 1500 m, M. Brancucci leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 2 ex. “ LAOS, Houa Phan prov., 20 ° 13 ’ 09 - 19 ’’ N, 103 ° 59 ’ 54 ’’ - 104 ° 00 ’ 03 ’’ E, 1480 - 1550 m, Phou Pane Mt., 9 - 16. vi. 2009, David Hauck leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 991 ex. “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ’ N, 104 ° 01 ’ E; 11. iv. - 15. v. 2012; 1300 - 1900 m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012 / 13 ” (ZFMK), 679 ex. “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ’ N, 104 ° 01 ’ E; 11. iv. - 15. v. 2012; 1300 - 1900 m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012 ” (ZFMK), 24 ex. “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) ~ 20 ° 12 ’ N, 104 ° 01 ’ E; 1300 - 1900 m; 01. - 31. V. 2011; leg. C. Holzschuh Ankauf ZFMK Bonn 2011 ” (ZFMK), 189 ex. “ Laos-NE, Houa Phan prov., 20 ° 13 ’ 09 - 19 ’’ N 103 ° 59 ’ 54 ’’ - 104 ° 00 ’ 03 ’’ E, 1480 - 1510 m Phou Pane Mt., 22. IV. - 14. V. 2008 Vit Kuban leg. ” (NMPC, ZFMK), 43 ex. “ Laos-NE Hua Phan prov., 20 ° 12 ’ N, 104 ° 01 ’ E, Phu Phan Mt., 1500 - 1900 m, 17.5. - 3.6.2007, leg. Vit Kuban ” (ZFMK), 28 ex. “ Laos-NE Hua Phan prov., 20 ° 12 ’ N, 104 ° 01 ’ E, Phu Phan Mt., 1500 - 1900 m, 17.5. - 3.6.2007, leg. C. Holzschuh ” (ZFMK), 7 ex. “ Laos (west): Oudomxao province, Beng, 20.20 ° N 101.80 ° E, May 1 - 14.2010 local collectors of Li Jingke ” (ZFMK), 1 Ô “ X-DA 4734 / X- DA 4734 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 15. - 30. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica varians ” (ZFMK), 1 Ô “ X-DA 4732 / X- DA 4732 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 15. - 30. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica varians ” (ZFMK), 1 Ô “ X-DA 4673 / X- DA 4673 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 15. - 30. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica varians ” (ZFMK), 1 Ô “ X-DA 4765 / X- DA 4765 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 6. - 14. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica varians ” (ZFMK), 1 Ô “ X-DA 4766 / X- DA 4766 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 6. - 14. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica varians ” (ZFMK), 1 Ô “ XDA 5115 / X- DA 5115 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 6. - 15. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica spLAOS Phupane 11 ” (ZFMK). Vietnam: 2 ÔÔ “ N-Vietnam, Cao Bang Pr. vic. Tinh Tuc, Son Dong Nu Pa Oac Nature Res. 09. - 15. V. 2014, 850 - 1300 m 22 ° 37 ’ 55 ’’ N, 105 ° 52 ’ 98 ’’ E leg. A. Weigel by light ” (NME), 1 Ô “ N-Vietnam, Cao Bang Pr. vic. Tinh Tuc, Son Dong Nu Pa Oac Nature Res. 22 ° 36 ’ 50 ’’ N, 105 ° 53 ’ 20 ’’ E 13. V. 2014, 1400 - 1800 m leg. A. Weigel ” (NME).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9FFFA7FF3E77202D0FDFCF.taxon	description	Redescription. Length: 5.3 mm, length of elytra: 3.6 mm, width: 3.1 mm. Body oval, body black, legs, center of elytra, margins of pronotum, labroclypeus, and antennal funiculus yellowish, dorsal surface iridescent shiny and nearly glabrous. Labroclypeus elliptical, slightly wider than long, widest shortly before base, lateral margins strongly convex and convergent to strongly rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, medially almost flat, finely and coarsely, densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons dull, with sparse, coarse punctures, beside eyes and behind frontoclypeal suture with a few single short setae, otherwise with minute setae in punctures. Smooth area in front of eyes twice as wide as long. Ocular canthus sparsely finely punctate, narrow and moderately long (1 / 3 of ocular width), with a terminal seta. Eyes moderately large, ratio of diameter / interocular width: 0.66. Antenna with ten antennomeres; club composed of four antennomeres, slightly reflexed at apex, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest shortly before base, lateral margins evenly convex and convergent to blunt but distinctly produced anterior angles, posterior angles strongly rounded. Anterior margin of pronotum weakly convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, on midline narrowly impunctate. Elytra short, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd intervals with a few short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.76. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, with mixed finer and larger punctures, without smooth midline, with numerous short setae and a few longer in the larger punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin finely serrate at apex, only weakly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 3.1; dorsal margin sharply carinate, with two groups of spines, basal group at middle, apical one at three quarter of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere little shorter than following two tarsomeres combined, twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 2 A – C. Habitus: Fig. 2 D. Variation. The coloration of this species is quite variable, ranging from entirely yellow (females) to entirely black (some males).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9DFFA1FF3E70972D77D953.taxon	description	Figs. 2 E – H, 24	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9DFFA1FF3E70972D77D953.taxon	materials_examined	Type material examined. Holotype (fukiensis): Ô [China] “ Kuatun (2300 m) 27, 40 n. Br. 117, 40 ö. L. J. Klapperich 20.5.1938 (Fukien) / Type Gastroserica fukiensis G. Frey 1971 ” (CF 2). Paratypes: 1 Ô, 1 ♀ [China] “ Kuatun (2300 m) 27, 40 n. Br. 117, 40 ö. L. J. Klapperich 20.5.1938 (Fukien) / Paratype Gastroserica fukiensis G. Frey 1971 ” (CF). Additional material examined. (see also Ahrens 2002). China. 2 ex. “ China: SW Zhejiang, 5. VI. Fangyangshan, Huangmao Jian 27 ° 53 ’ N 119 ° 11 ’ E, 1500 - 1850 m Jaroslav Turna leg., 2008 ” (ZFMK), 6 ex. “ C-China, Shaanxi, Qinling Shan, 6 km E of Xunyangba 1000 - 1300 m, 23. V. - 13. VI. leg. C. Holzschuh 2000 ” (CP), 4 ex. “ China, W Hubei, 8 + 15. - 17. VI. Muyuping S env. - 1100 m 31.45 N 110.4 E Jaroslav Turna leg., 2002 ” (CP), 2 ex. “ China, 1000 - 1300 m, Shaanxi, Qinling Mts. Xunyangba (6 km E), 23. v. - 13. vi. 1998, J. H. Marshal leg. ” (CP), 3 ex. [China] “ Kuatun, Fukien China 25.5.46 (Tschung Sen.) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 2 ex. [China] “ Kuatun (2300 m) 27,40 n. Br. 117,40 o. L. J. Klapperich 11.5. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 3 ex. [China] “ Kuatun (2300 m) 27,40 n. Br. 117,40 o. 2 Frey (1972) did not specify in which museum the type specimens of this taxon are preserved, however, we found only type specimens in the collection G. Frey (NHMB). Since he noticed ZFMK for type depository in all other species that originated from the ZFMK Bonn (leg. Klapperich) treated in that paper (Frey 1972), the holotype of G. fukiensis should also belong to ZFMK. L. J. Klapperich 13.5. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. [China] “ Kuatun (2300 m) 27,40 n. Br. 117,40 o. L. J. Klapperich 4.6. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. [China] “ Kuatun (2300 m) 27,40 n. Br. 117,40 o. L. J. Klapperich 30.5. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. [China] “ Kuatun (2300 m) 27,40 n. Br. 117,40 o. L. J. Klapperich 25.5. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. [China] “ Kuatun (2300 m) 27,40 n. Br. 117,40 o. L. J. Klapperich 23.5. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. [China] “ Kuatun (2300 m) 27,40 n. Br. 117,40 o. L. J. Klapperich 11.4. 1938 (Fukien) / ex. Coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ex. “ China-Guangdong, Daqiao env., 1000 - 1200 m, 24 ° 54 ’ N- 113 ° 01 ’ E, 1. - 3. v. 2002, Dr. R. Fencl lgt. ” (CP), 1 Ô [China] “ Tienmushan, 12. VI. 1936, 1350 m, leg. O. Piel, Musee Heude ” (IZAS), 1 Ô [China] “ Kuling, 7. VII. 1935, leg. O. Piel, Musee Heude ” (IZAS), 2 ÔÔ [China] “ Mt. Dayaoshan, Jinxiu, Guangxi, 12. VI. 1982, leg. Li Fasheng ” (CAU), 1 Ô [China] “ Mt. Jianfengling, Ledong, Hainan, 17 - 21. V. 2007, leg. Ba Yibin, Lang Juntong ” (HBUM), 1 Ô [China] “ San’gang, Mt. Wuyishan, Fujian, 16 - 28. Ⅴ. 2004, leg. Yuan Caixia, Li Jing ” (HBUM), 1 Ô “ China: Zhejiang, Lin’an City, Daming Shan, 30 ° 02 ’ 04 ’’ N, 118 ° 39 ’ 14 ’’ E, light trap, 860 m, 7. VII. 2014, Jia-Yao-Hu leg. ” (SNUC), 1 Ô “ CHINA: Zhejiang Prov. West Tiammu Shan (Mts.) reserve forest, esp. on flowering Castanea sequinii, 5. - 6. vii. 2017; 1500 m 30 ° 20 ’ 58.5 ’’ N, 119 ° 25 ’ 26.5 ’’ E J. Hájek & J. Růžička leg. / 1060 Asia Sericini spec. ” (NMPC), 1 Ô, 1 ♀ “ CHINA: Zhejiang Prov. West Tiammu Shan (Mts.) reserve forest, esp. on flowering Castanea sequinii, 5. - 6. vii. 2017; 1500 m 30 ° 20 ’ 58.5 ’’ N, 119 ° 25 ’ 26.5 ’’ E J. Hájek & J. Růžička leg. ” (NMPC).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9DFFA1FF3E70972D77D953.taxon	description	Redescription. Length: 5.8 mm, length of elytra: 3.9 mm, width: 3.4 mm. Body oval, body yellowish brown, head and ventral surface including abdomen and most parts of pygidium blackish, dorsal surface dull and nearly glabrous. Labroclypeus elliptical, slightly wider than long, widest shortly before base, lateral margins strongly convex and convergent to strongly rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, medially almost flat, finely and coarsely, densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons dull, with sparse, coarse punctures, beside eyes and behind frontoclypeal suture with a few single short setae, otherwise with minute setae in punctures. Smooth area in front of eyes twice as wide as long. Ocular canthus sparsely finely punctate, narrow and moderately long (1 / 3 of ocular width), with a terminal seta. Eyes moderately large, ratio of diameter / interocular width: 0.68. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest shortly before base, lateral margins evenly convex and convergent to blunt but moderately produced anterior angles, posterior angles strongly rounded. Anterior margin of pronotum weakly convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, on midline narrowly impunctate. Elytra short, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd intervals with a few short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.72. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, with mixed finer and larger punctures, without smooth midline, with numerous short setae and a few longer in the larger punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin finely serrate at apex, only weakly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 3.1; dorsal margin sharply carinate, with two groups of spines, basal group at middle, apical one at three quarter of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and very sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined, twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 2 E – G. Habitus: Fig. 2 H. Variation. The coloration of this species is also quite variable, ranging from almost entirely yellow (females, some males – except black head and ventral surface) to bicolored with margins and disc as well as frons dark brown (some males). In addition, we found a color variant with blackish elytra and red pronotum, very similar to the next species; this color form from Zheijang (1060 Asia Sericini spec.) is in shape of parameres clearly identical with G. fukiensis.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9BFFA2FF3E76E02DC6DF7F.taxon	description	Figs. 3 A – D, 25	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9BFFA2FF3E76E02DC6DF7F.taxon	materials_examined	Type material examined. Holotype Ô [China] “ Simao, Yunnan, No. 49 / LW- 360 ” (IZAS). Paratypes: Laos: 11 ÔÔ “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ’ N, 104 ° 01 ’ E; 11. iv. - 15. v. 2012; 1300 - 1900 m; leg. C. Holzschuh - ZFMK Ankauf Bonn 2012 ” (ZFMK), 5 ÔÔ “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ’ N, 104 ° 01 ’ E; 11. iv. - 15. v. 2012; 1300 - 1900 m, leg. C. Holzschuh - ZFMK Ankauf Bonn 2012 / 2013 ” (ZFMK), 1 Ô “ 836956 / 836956 Laos Xieng Khouang, Phonsavan (30 km NE), Phou Sane Mt. 1400 - 1600 m, 10. - 30.5.2009 V. Kuban Microserica spLa 1 - cfvarians / 1112 Asia Sericini spec. ” (ZFMK), 2 ÔÔ “ LAOS, Xieng Khouang prov., 19 ° 37 – 8 ’ N, 103 ° 20 ’ E, Phonsavan (30 km NE) Phou Sane Mt., 1420 m, 10 - 30. v. 2009, Z. Kraus leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 1 Ô “ XDA 4729 / X- DA 4729 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 15. - 30. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica spLAOS Phupane 13 ” (ZFMK), 1 Ô “ X-DA 4731 / X- DA 4731 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 15. - 30. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica spLAOS Phupane 13 ” (ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9BFFA2FF3E76E02DC6DF7F.taxon	description	Description of holotype. Length: 5.1 mm, length of elytra: 3.8 mm, width: 3.2 mm. Body oval, body including legs reddish brown, antenna yellow, club slightly darker, head, elytra and abdomen blackish dark, pronotum and scutellum red, dorsal surface iridescent shiny, partly dull, nearly glabrous. Labroclypeus elliptical, slightly wider than long, widest shortly before base, lateral margins strongly convex and convergent to strongly rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface weakly shiny, medially convexly elevated, coarsely and densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons dull, with sparse, coarse punctures, beside eyes and behind frontoclypeal suture with a few single short setae, otherwise with minute setae in punctures. Smooth area in front of eyes twice as wide as long. Ocular canthus sparsely finely punctate, narrow and moderately long (1 / 3 of ocular width), with a terminal seta. Eyes moderately large, ratio of diameter / interocular width: 0.65. Antenna with ten antennomeres; club composed of four antennomeres, slightly reflexed at apex, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest shortly before base, lateral margins evenly convex and convergent to blunt but distinctly produced anterior angles, posterior angles moderately rounded. Anterior margin of pronotum weakly convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, on midline narrowly impunctate. Elytra short, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd intervals with a few short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.9. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, with mixed finer and larger punctures, without smooth midline, with numerous short setae and a few longer in the larger punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin finely serrate at apex, only weakly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 2.4; dorsal margin sharply carinate, with two groups of spines, basal group at middle, apical one at three quarter of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere little as long as following two tarsomeres combined, dorsal tibial spur missing in holotype. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 3 A – C. Habitus: Fig. 3 D. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9BFFA2FF3E76E02DC6DF7F.taxon	diagnosis	Diagnosis. Gastroserica simaoensis Ahrens, Lukic & Liu, new species is in external appearance and shape of aedeagus most similar to G. fukiensis Frey, 1972. The new species differs by the longer and more elongate basal lobe of left paramere, and the color.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9BFFA2FF3E76E02DC6DF7F.taxon	etymology	Etymology. The name of the new species is derived from the type locality, Simao (adjective in singular nominative case).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF9BFFA2FF3E76E02DC6DF7F.taxon	description	Variation. Length: 5.1 – 5.9 mm, length of elytra: 3.8 – 4.0 mm, width: 3.2 – 3.4 mm. The coloration of this species is quite variable, ranging from yellowish brown having laterally and basally darkened elytra (some males), to the red form of the holotype.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF98FFA3FF3E70072E2DDEF3.taxon	description	Figs. 3 E – H, 24	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF98FFA3FF3E70072E2DDEF3.taxon	materials_examined	Type material examined. Lectotype (here designated): Ô “ N China Donkier V. / Microserica hiulca Typ. Brsk. / coll. Brenske ” (ZMHB). Additional material examined. China: 1 ex. “ Museum Paris Kouy-Tcheou Reg. de Pin-Fa, Pere Cavalerie 1908 ” (MNHN), 3 ex. “ China: Kiangsi Hong Shan, 840 m 30. VI. 1936 / L. & M. Gressitt Collectors Bishop Mus. ” (BPBM), 1 ex. “ So. Kiangsi Prov., S. China SE Corner Hong shan, Alt. 950 meters. June 30, 1936 J. Linsley Gressitt ” (BPBM), 1 ex. “ Kiangsi, SE. China Hong Shan. 1000 m VI- 25 - 36. Gressitt ” (BPBM), 8 ex. “ China Jiangxi W Jinggang Shan Ciping env. 2. - 14. VI. 1994 ” (NHMW), 2 ex. “ China: Hunan; Mupu Mt. 1600 m, Pingjiang VII- 2003, leg. Li et al. ” (ZFMK), 12 ex. “ China: S- Hubei Mupushan, Tongshan vi. 2004 leg. Wen ” (ZFMK), 4 ex. “ China: Hunan prov., Zhang Jia Jie 15. vi. 1992; C. Holzschuh leg. ” (CP), 3 ex. “ CH, Guizhou prov., Leigong Shan (E slope), 14 - 24. vi. 2001, Leishan-> Fangxiang (pass), 26 ° 22.74 ’ N 108 ° 12.99 ’ E, ~ 1700 m, C. Holzschuh leg. ” (CP), 3 ex. “ China, W Hubei, 12. - 13. VII. Muyuping S env. 31,45 N 110,4 E, ~ 1300 m Jaroslav Turna leg., 2003 ” (ZFMK), 1 Ô “ China, W Jiangxi Jinggang Shan-Ciping 2 - 14. VI. 1994 E. Jendek & O. Sausa leg. / CS 78 ” (CP), 2 ÔÔ, 1 ♀ [China] “ Longmenhe, Xingshan, Hubei, 14, 16. VII. 1993, 1350 m, leg. Yang Xingke, Sun Baowen / LW- 387 ” (IZAS), 1 Ô, 1 ♀ [China] “ Mts. Leigongshan, Guizhou, 13. VII. 1983, 1300 - 1400 m, leg. Zou ” (NKU), 1 Ô [China] “ Baiyan, Longsheng, Guangxi, 23. VI. 1963, 1150 m, leg. Wang Shuyong ” (IZAS). Vietnam: 2 ex. (Ô) “ Tonkin Montes Mauson April, Mai 2 - 3000 ’ [feet] H. Fruhstorfer ” (ZMHB, MNHN).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF98FFA3FF3E70072E2DDEF3.taxon	description	Redescription. Length: 5.4 mm, length of elytra: 3.6 mm, width: 3.4 mm. Body oval, body yellowish brown, head, center of pronotum, and ventral surface including abdomen and most parts of pygidium darker brown, dorsal surface iridescent shiny or sometimes dull and nearly glabrous. Labroclypeus elliptical, slightly wider than long, widest shortly before base, lateral margins strongly convex and convergent to strongly rounded anterior angles; anterior margin emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, medially almost flat, finely and coarsely, densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons dull, with sparse, coarse punctures, beside eyes and behind frontoclypeal suture with a few single short setae, otherwise with minute setae in punctures. Smooth area in front of eyes twice as wide as long. Ocular canthus sparsely finely punctate, narrow and moderately long (1 / 3 of ocular width), with a terminal seta. Eyes small, ratio of diameter / interocular width: 0.47. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum widest at base, lateral margins evenly convex and convergent to blunt but distinctly produced anterior angles, posterior angles strongly rounded. Anterior margin of pronotum weakly convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, on midline narrowly impunctate. Elytra short, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd intervals with a few short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.76. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, with mixed finer and larger punctures, without smooth midline, with numerous short setae and a few longer in the larger punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin finely serrate at apex, only weakly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 2.9; dorsal margin weakly carinate, with two groups of spines, basal group at middle, apical one at three quarter of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and very sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined, almost twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 3 E – G. Habitus: Fig. 3 H.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF99FFA3FF3E718B2DECDCBC.taxon	description	Figs. 4 A – D, 24	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF99FFA3FF3E718B2DECDCBC.taxon	materials_examined	Material examined. 1 Ô “ Mt. An-ma-shan Taichung, Taiwan 12 th vii 1992 C. Yu leg. ” (ZFMK). Gastroserica nitidipyga (Nomura, 1974) new combination Figs. 4 E – H, 25	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF99FFA3FF3E718B2DECDCBC.taxon	materials_examined	Material examined. 1 Ô [China] “ Cujiang, Huaping, Guangxi, 7. VI. 1963, leg. Yang Jikun / LW- 356 ” (CAU), 1 Ô “ China: Fujian Province; Mt. Liang-shan-ding, Wuping County, July 2 - 13, 2009 local collectors Ankauf via Li Jingke 2010 ” (ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF86FFBCFF3E74F32E6ADCB3.taxon	description	Figs. 4 I – L, 25	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF86FFBCFF3E74F32E6ADCB3.taxon	materials_examined	Type material examined. Holotype Ô [China] “ Shaanxi, Zhouzhi, Yueba, 2012 - VI- 30. N: 33.585 - 33.572, E: 107.765 - 107.788, 1139 - 1573 m, daytime / LW 1321 ” (ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF86FFBCFF3E74F32E6ADCB3.taxon	description	Description of holotype. Length: 6.0 mm, length of elytra: 4.4 mm, width: 3.5 mm. Body oval, dark brown, legs, one longitudinal stripe at middle of the elytra and its lateral margin, margins of pronotum, labroclypeus, apex of scutellum, and antennal funiculus yellowish, antennal club brown, dorsal surface dull and nearly glabrous. Labroclypeus transverse-elliptical, distinctly wider than long, widest at middle, lateral margins strongly convex and convergent to strongly rounded anterior angles; anterior margin widely but not shallowly emarginate medially, anterior margin strongly reflexed, lateral margin weakly reflexed; labrum strongly enlarged; lateral margins and ocular canthus produce a blunt angle; surface shiny, weakly convex, finely, densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons dull, with sparse, fine punctures, beside eyes and behind frontoclypeal suture with a few single short setae in larger punctures, otherwise with minute setae in punctures. Smooth area in front of eyes narrow, as wide as long. Ocular canthus sparsely finely punctate, narrow and moderately long (1 / 3 of ocular width), with a terminal seta. Eyes small, ratio of diameter / interocular width: 0.49. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins evenly convex and convergent to blunt but weakly produced anterior angles, posterior angles moderately rounded. Anterior margin of pronotum straight, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and evenly densely punctate. Elytra elongate, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd intervals with a few short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.7. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, with large, dense punctures, without smooth midline, with numerous short setae at apex. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin finely serrate at apex, only weakly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 3.0; dorsal margin sharply carinate, with two groups of spines, basal group at middle, apical one at three quarter of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere little shorter than following two tarsomeres combined, a quarter of its length longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 4 I – K. Habitus: Fig. 4 L. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF86FFBCFF3E74F32E6ADCB3.taxon	diagnosis	Diagnosis. This new species is very similar to Gastroserica nitidipyga (Nomura, 1974). Gastroserica yuebaensis Ahrens, Lukic & Liu, new species differs from G. nitidipyga by the shorter and narrower left paramere, the narrower right paramere (lateral view).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF86FFBCFF3E74F32E6ADCB3.taxon	etymology	Etymology. The name of the new species is derived from the name of the type locality, Yueba (adjective in nominative singular case). Genbank accessions. Table 1.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF87FFBDFF3E74F32EF8DAC6.taxon	description	Fig. 24	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF87FFBDFF3E74F32EF8DAC6.taxon	materials_examined	Material examined. See Ahrens & Fabrizi (2009 b: 268). Aedeagus. See Ahrens & Fabrizi (2009 b: p. 282, fig. 6 G – I).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF87FFBDFF3E74F32EF8DAC6.taxon	distribution	Distribution. India (Assam / Patkai Mts).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF87FFBDFF3E759E2EE0D864.taxon	description	Fig. 25	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF87FFBDFF3E759E2EE0D864.taxon	materials_examined	Material examined. See Ahrens & Fabrizi (2016: 133) Aedeagus. See Ahrens & Fabrizi (2016: p. 298, fig. 10 X – Z)	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF87FFBDFF3E759E2EE0D864.taxon	distribution	Distribution. India (Arunachal Pradesh).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF87FFBEFF3E77382E69DE17.taxon	description	Figs. 5 A – D, 24	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF87FFBEFF3E77382E69DE17.taxon	materials_examined	Type material examined. Holotype: Ô “ Laos: Vientiane Prov. Ban Van Eue 31. VII. 1965 / Native collector Bishop Mus. / 17 Sericini Asia spec. ” (BPBM). Paratypes: Laos: 1 Ô “ Laos: Vientiane Prov. Ban Van Eue 14,15. IV. 1965 / Native collector Bishop Mus. ” (BPBM), 2 ÔÔ “ LAOS centr., Bolikhamsai prov., Ban Non env., 9. - 14.5.1998 Route No. 8, alt. 220 + 50 m, N 18 ° 08.7, E 104 ° 28.1 (GPS), E. Jendek & O. Šauša leg. ” (CP), 1 Ô “ 836950 / 836950 Laos Xieng Khouang, Phonsavan (30 km NE), Phou Sane Mt. 1400 - 1600 m 10. - 30.5.2009 V. Kuban Microserica spLA 1 / 540 Sericini Asia spec. ” (ZFMK), 1 Ô “ 836958 / 836958 Laos Xieng Khouang, Phonsavan (30 km NE), Phou Pane Mt. 1400 - 1600 m 10. - 30.5.2009 V. Kuban Microserica spLA 3 ” (ZFMK), 8 ÔÔ “ LAOS, Xieng Khouang prov., 19 ° 37 – 8 ’ N, 103 ° 20 ’ E, Phonsavan (30 km NE) Phou Sane Mt., 1420 m, 10 - 30. v. 2009, Z. Kraus leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň / 1095 Sericini Asia spec. ” (NHMB), 2 ÔÔ, 1 ♀ “ LAOS, Xieng Khouang prov., 19 ° 37 – 8 ’ N, 103 ° 20 ’ E, 30 km NE Phonsavan: Ban Na Lam -> Phou Sane Mt., 1300 - 1700 m, 10 - 30. v. 2009, M. Geiser leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB). China: 1 Ô “ Kunluo Road 591 km, Simao County, Yunnan, 11. V. 1957, 1350 m, leg. Pu Fuji ” (IZAS), 1 Ô “ Menghun, Xishuangbanna, Yunnan, 22. V. 1958, 1200 - 1400 m, leg. Zhang Yiran ” (IZAS).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF87FFBEFF3E77382E69DE17.taxon	description	Description of holotype. Length: 5.0 mm, length of elytra: 3.6 mm, width: 3.4 mm. Body oval, body including legs dark reddish brown, antennal funiculus little lighter, dorsal surface iridescent shiny to moderately simply shiny, partly dull, nearly glabrous. Labroclypeus subrectangular, slightly wider than long, widest shortly at base, lateral margins subparallel, strongly convex and convergent to strongly rounded anterior angles; anterior margin emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, medially weakly convexly elevated, coarsely and densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons dull, with sparse, coarse punctures, impunctate along midline, beside eyes and behind frontoclypeal suture with a few single short setae, otherwise with minute setae in punctures. Smooth area in front of eyes twice as wide as long. Ocular canthus sparsely finely punctate, narrow and moderately long (1 / 3 of ocular width), with a terminal seta. Eyes moderately large, ratio of diameter / interocular width: 0.7. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest shortly before base, lateral margins evenly convex and convergent to sharp but weakly produced anterior angles, posterior angles strongly rounded. Anterior margin of pronotum straight, with robust and complete marginal line; basal marginal line absent; surface finely and moderately densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, on midline narrowly impunctate. Elytra short, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd intervals with a few short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.72. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium weakly convex and dull, with mixed finer and larger punctures, without smooth midline, with numerous short setae and a few longer in the larger punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin finely serrate at apex, only weakly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 2.7; dorsal margin sharply carinate, with two groups of spines, basal group at middle, apical one at three quarter of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined, and twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 5 A – C. Habitus: Fig. 5 D.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF87FFBEFF3E77382E69DE17.taxon	diagnosis	Diagnosis. Gastroserica fumaria Ahrens, Lukic & Liu, new species is in external appearance and shape of aedeagus rather similar to G. varians. Gastroserica fumaria differs by the longer and dorsally evenly arched phallobase, the raspulose field at the right side of phallobasal apex; G. fumaria appears more unicolored, with dark elytra.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF87FFBEFF3E77382E69DE17.taxon	etymology	Etymology. The name of the new species is derived from the Latin adjective, “ fumarius ” (smoky-brown).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF87FFBEFF3E77382E69DE17.taxon	description	Variation. Length: 4.8 – 5.2 mm, length of elytra: 3.4 – 3.6 mm, width: 3.0 – 3.4 mm. In coloration rather uniform, some specimens have a deep large concavity at the middle of pronotum, which we think are simply deformed. Female: Antennal club little shorter than remaining antennomeres combined, composed of three antennomeres; eyes little smaller than in male, ratio of diameter / interocular width: 0.54; body lighter, pygidium almost flat. Genbank accessions. Table 1.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF84FFBFFF3E71AF2D6ADD3F.taxon	description	Figs. 5 E – H, 25	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF84FFBFFF3E71AF2D6ADD3F.taxon	materials_examined	Type material examined. Holotype: Ô “ THAI-NE, Loei prov. Phu Kradung N. P. 16 ° 52 ’ N 101 ° 49 ’ E, 16. - 18. v. 1999, 1000 m, D. Hauck leg. / 13 Sericini Asia spec. ” (CP). Paratypes: 4 ÔÔ, 1 ♀ “ Phu Rua NP (12 - 1300 m) Loei P., NE Thai. 27 - 30. IV. 2006 Takakuwa, M. leg. ” (ZFMK), 1 Ô “ NE Thailand, Phu Rua, Loei Prov., Phu Rua National Park 22. V. 2004 leg. S. Ohmomo ” (ZFMK), 7 ÔÔ “ Mt. Phu Rua Summit, Loei Prov., NE Thailand, 24. V. 2004 leg. Y. Komiya ” (ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF84FFBFFF3E71AF2D6ADD3F.taxon	description	Description of holotype. Length: 5.6 mm, length of elytra: 3.5 mm, width: 3.1 mm. Body oval, body including legs and antenna yellowish brown, abdomen, middle of pronotum, margins of elytra and frons blackish, antennal club little darker, dorsal surface iridescent shiny to weakly dull, nearly glabrous. Labroclypeus semicircular, slightly wider than long, widest at base, lateral margins strongly convex and convergent to strongly rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, medially weakly convexly elevated, finely and coarsely, densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons dull, with sparse, fine punctures, beside eyes and behind frontoclypeal suture with a few single short setae, otherwise with minute setae in punctures. Smooth area in front of eyes twice as wide as long. Ocular canthus sparsely finely punctate, narrow and moderately long (1 / 3 of ocular width), with a terminal seta. Eyes moderately large, ratio of diameter / interocular width: 0.63. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins evenly convex and convergent to sharp but weakly produced anterior angles, posterior angles strongly rounded. Anterior margin of pronotum straight, with robust and complete marginal line; basal marginal line absent; surface finely and moderately densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, on midline narrowly impunctate. Elytra short, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd intervals with a few short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.63. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium moderately convex and dull, with mixed finer and larger punctures, without smooth midline, with numerous short setae and a few longer in the larger punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin finely serrate at apex, only weakly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 2.94; dorsal margin sharply carinate, with two groups of spines, basal group at middle, apical one at three quarter of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth, with a few fine punctures; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined, and more than twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 5 E – G. Habitus: Fig. 5 H. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF84FFBFFF3E71AF2D6ADD3F.taxon	diagnosis	Diagnosis. Gastroserica phukradung Ahrens, Lukic & Liu, new species is in external appearance and shape of aedeagus rather similar to Gastroserica fumaria Ahrens, Lukic & Liu, new species. Gastroserica phukradung differs most of all by the shape of the left paramere which is basally more narrow (lateral view), the right one is before apex strongly bent externally, while it is almost straight in G. fumaria.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF84FFBFFF3E71AF2D6ADD3F.taxon	etymology	Etymology. The name of the new species is named after its occurrence in the Phu Kradun province (noun in apposition).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF85FFB9FF3E72C72D4ADBBB.taxon	description	Figs. 6 A – D, 24	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF85FFB9FF3E72C72D4ADBBB.taxon	materials_examined	Type material examined. Holotype: Ô “ THAI-N, 1. - 19. v. 1998 Chiang Mai prov., Ban San Pakia, 1400 m, Ivo Martinů leg. / Coll. Dirk Ahrens / 15 Sericini Asia spec. ” (ZFMK). Paratypes: Thailand: 1 Ô “ NW Thailand 9. - 16. V. Mae Hong Son 1991 Ban Huai Po 1600 m leg. P. Pacholátko / TS 72 ” (CP), 21 ÔÔ “ NE Thailand 1. - 15.5 [?]. 1991 Mae Hong Son Ban Huai Po 800 - 1600 m S. Bily leg. ” (NHMW, ZFMK), 25 ÔÔ “ NW Thailand, 19.19 N, 97.59 E Mae Hong Son, 1991 Ban Huai Po 1600 - 2000 m 17. - 23.5., L. Dembicky leg. ” (NHMW, ZFMK), 4 ÔÔ “ NE Thailand 28.4.1991 Mae Hong Son S. Bily leg. ” (NHMW), 5 ÔÔ, 2 ♀♀ “ N. Thailand Ban Huai Po 9. - 16.5. 1991 I. Horák leg. / Coll. Milan Nikodym Praha ” (ZFMK), 1 Ô “ NW Thailand, Chieng Dao, Ban San Pakia, 5. - 10. v. 2004, 1200 m, Sv. Bily ” (NME). China: 1 Ô “ China: S-Yunnan (Xishuangbanna) 45 km SW Jinghong vic. Bangzhang vill. / N 21 ° 44 ’ 37 ’’, E 100 ° 27 ’ 02 ’’, 16 - 1700 m, 03. - 05. V. 2009 leg. A. Weigel, blüh Cast ” (NME).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF85FFB9FF3E72C72D4ADBBB.taxon	description	Description of holotype. Length: 5.1 mm, length of elytra: 3.1 mm, width: 2.9 mm. Body oval, body including legs and antenna yellow, two spots on pronotum, margins of elytra and frons blackish, dorsal surface shiny, nearly glabrous. Labroclypeus narrow and semicircular, as wide as long, widest shortly before base, lateral margins strongly convex and convergent to strongly rounded anterior angles; anterior margin weakly incised medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface medially weakly convexly elevated, finely and coarsely, densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons shiny, with dense, fine punctures, beside eyes and behind frontoclypeal suture with a few single short setae, otherwise with minute setae in punctures. Smooth area in front of eyes twice as wide as long. Ocular canthus sparsely finely punctate, narrow and moderately long (1 / 3 of ocular width), terminal seta abraded. Eyes moderately large, ratio of diameter / interocular width: 0.63. Antenna with ten antennomeres; club composed of four antennomeres, strongly reflexed, three times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins evenly convex and convergent to sharp but moderately produced anterior angles, posterior angles blunt and moderately rounded. Anterior margin of pronotum convex, with robust and complete marginal line; basal marginal line absent; surface coarsely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, coarsely and densely punctate, on median base impunctate. Elytra short, widest at middle, striae finely impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd intervals with a few short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface shiny, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.5. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and shiny, with mixed finer and larger punctures, without smooth midline, with numerous short setae and a few longer in the larger punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin finely serrate at apex, only weakly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 3.33; dorsal margin sharply carinate, with two groups of spines, basal group at middle, apical one at three quarter of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally sparsely finely punctate; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined and twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 6 A – C. Habitus: Fig. 6 D.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF85FFB9FF3E72C72D4ADBBB.taxon	diagnosis	Diagnosis. Gastroserica ivoi Ahrens, Lukic & Liu, new species is in external appearance and shape of aedeagus rather similar to Gastroserica lucens (Ahrens & Fabrizi, 2009). Gastroserica ivoi differs by long antennal club in male, the light yellowish body color, and by the shape of aedeagus: the phallobase is distinctly shorter compared to the longer parameres, and strongly curved (lateral view); the basal lobe of the left paramere is distally distinctly produced.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF85FFB9FF3E72C72D4ADBBB.taxon	etymology	Etymology. The name of the new species is named after one of its collectors Ivo Martinů (noun in singular genitive).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF85FFB9FF3E72C72D4ADBBB.taxon	description	Variation. Length: 5.0 – 5.1 mm, length of elytra: 3.0 – 3.1 mm, width: 2.8 – 2.9 mm. Paratypes without brown dots on the pronotum. Female: completely yellow, antennal club short, composed of three antennomeres; pygidium flat.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF83FFBAFF3E75432E6ADD8F.taxon	description	Figs. 6 E – I, 25	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF83FFBAFF3E75432E6ADD8F.taxon	materials_examined	Type material examined. Holotype Ô “ 836957 / 836957 Laos Xieng Khouang, Phonsavan (30 km NE), Phou Sane Mt. 1400 - 1600 m 10. - 30.2009 V. Kuban Microserica spLA 2 / 43 Asia Sericini spec. ” (ZFMK). Paratypes: Laos: 1 Ô “ LAOS north 24. - 30. V. 1997 20 km NW Lounag Namtha, N 21 ° 09.2 E 101 ° 18.7, alt. 900 + 100 m, E. Jendek & O. Šauša leg. ” (CP), 6 ÔÔ “ LAOS, Xieng Khouang prov., 19 ° 37 – 8 ’ N, 103 ° 20 ’ E, Phonsavan (30 km NE) Phou Sane Mt., 1420 m, 10 - 30. v. 2009, Z. Kraus leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 2 ÔÔ “ LAOS, Xieng Khouang prov., 19 ° 37 – 8 ’ N, 103 ° 20 ’ E, Phonsavan (30 km NE) Phou Sane Mt., 1300 - 1500 m 10 - 30. v. 2009, M. Brancucci leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 5 ÔÔ “ LAOS, Xieng Khouang prov., 19 ° 37 – 8 ’ N, 103 ° 20 ’ E, 30 km NE Phonsavan: Ban Na Lam -> Phou Sane Mt., 1300 - 1700 m, 10 - 30. v. 2009, M. Geiser leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 1 Ô “ LAOS, Hua Phan prov., 20 ° 13 ’ N, 104 ° 00 ’ E, Phou Pane Mt., 1 - 16. vi. 2009, 1350 - 1500 m, M. Brancucci leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 18 ÔÔ “ LAOS, Xieng Khoang prov., 19 ° 37 - 8 ’ N 103 ° 20 - 1 ’ E, Phonsavan (30 km NE) Phou Pane Mt., 1400 - 1700 m, 10. - 30. v. 2009, D. Hauck leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 1 Ô “ Laos: Luang Namtha Distr. Lak Kham Mai, Muang Sing, 5 - 9.5.2007, M. Pejcha ” (NME), 11 ÔÔ “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ’ N, 104 ° 01 ’ E; 14. iv. - 15. v. 2012; 1300 - 1900 m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012 / 13 ” (ZFMK), 2 ÔÔ “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ’ N, 104 ° 01 ’ E; 1300 - 1900 m; 01. - 31. v. 2011; leg. C. Holzschuh Ankauf ZFMK Bonn 2011 ” (ZFMK), 16 ÔÔ “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ’ N, 104 ° 01 ’ E; 11. iv. - 15. v. 2012; 1300 - 1900 m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012 ” (ZFMK), 1 Ô “ Laos-NE Hua Phan prov. 20 ° 12 ’ N, 104 ° 01 ’ E Phou Phan Mt., 1900 m 17.5. - 3.6.2007, leg. C. Holzschuh ” (ZFMK), 1 Ô “ LAOS, Attapeau prov. Annam Highlands Mts Dong Amphan; NBCA ca 1160 m Nong Fa (crater lake) env.; 15 ° 05.9 ’ N, 107 ° 25.6 ’ E St. Jakl lgt. 30.4. - 6.5.2010 ” (ZFMK). China: 1 Ô “ China: SYunnan (Xishuangbanna) 23 km NW Jinghong vic. Na Ban (NNNR) / N 21 ° 09.49 ’, E 100 ° 39.92 20. V. 2008, leg. A. Weigel, 730 m Waldrand KF ” (NME), 1 ♀ “ China: S-Yunnan (Xishuangbanna) 20 km NW Jinghong vic. Man Dian (NNNR) / N 22 ° 07.80 ’, E 100 ° 40.05 730 m 28. VI. 2008, GS leg. A. Weigel, forest ” (NME), 1 Ô [China] “ Mandian (Forest), Nabanhe Nature Reserve, Jinghong, Xishuangbannna, Yunnan, 16. V. 2009, 753 m, leg. Meng LZ / LW- 401 ” (IZAS), 1 Ô [China] “ Mt. Pinglongshan, Shangsi, Guangxi, 6. IV. 2002, leg. Shi Aimin / LW- 397 ” (HBUM), 1 Ô, 2 ♀♀ [China] “ Mengla, Xishuangbanna, Yunnan, 10. VI. 1959, 620 - 650 m, leg. Pu Fuji, Zhang Facai ” (IZAS), 1 ♀ [China] “ Shuiman, Hainan, Guangdong, 26. V. 1960, 1879 m, leg. Li Suofu ” (IZAS). Vietnam: 3 ÔÔ “ N-Vietnam Cao Bang Prov., vic. Vin Den, Nui Pia Oac Nat. Res. 06. - 10. V. 2013, 22 ° 33 ’ 53 ’’ N, 105 ° 52 ’ 53 ’’ E 900 - 1300 m A. Skale ” (ZFMK, NME), 1 Ô “ N-Vietnam Cao Bang Prov., vic. Vin Den, Nui Pia Oac Nature Res., 6. - 10. V. 2013 22 ° 33 ’ 53 ’’ N, 105 ° 52 ’ 53 ’’ E 900 - 1300 m leg. A. Weigel ” (NME), 2 ÔÔ, 1 ♀ “ N-Vietnam Cao Bang Prov., vic. Vin Den, Nui Pia Oac Nat. Res. 10. - 15. V. 2014, 22 ° 33 ’ 53 ’’ N, 105 ° 52 ’ 53 ’’ E 900 - 1300 m A. Skale ” (ZFMK, NME), 1 Ô “ N-Vietnam Cao Bang Prov., vic. Tinh Tuc, Son Dong, Nui Piac Oac Nature Reserve, 9. - 15. V. 2014, 22 ° 37 ’ 55 ’’ N 105 ° 52 ’ 98 ’’ E 850 - 1300 m leg. A. Skale ” (NME).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF83FFBAFF3E75432E6ADD8F.taxon	description	Description of holotype. Length: 4.9 mm, length of elytra: 3.4 mm, width: 3.1 mm. Body oval, body dark brown to blackish, pronotum, center of elytra and pygidium, legs and antenna yellow, margins of elytra and frons dark, dorsal surface shiny, nearly glabrous. Labroclypeus narrow and almost semicircular, as wide as long, widest at base, lateral margins strongly convex and convergent to strongly rounded, almost obsolete anterior angles; anterior margin weakly incised medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface medially weakly convexly elevated, finely and coarsely, densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons shiny, with moderately dense, fine punctures, beside eyes and behind frontoclypeal suture with a few single short setae, otherwise with minute setae in punctures. Smooth area in front of eyes twice as wide as long. Ocular canthus sparsely finely punctate, narrow and moderately long (1 / 3 of ocular width), with a terminal seta. Eyes moderately large, ratio of diameter / interocular width: 0.6. Antenna with ten antennomeres; club composed of four antennomeres, weakly reflexed, 1.6 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins evenly convex and convergent to rectangular and moderately produced anterior angles, posterior angles blunt and moderately rounded. Anterior margin of pronotum convex, with robust and complete marginal line; basal marginal line absent; surface finely and moderately densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, coarsely and densely punctate, on median base impunctate. Elytra short, widest at middle, striae finely impressed, finely and densely punctate, intervals flat, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd intervals with a few short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface shiny, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.53. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and shiny, with mixed sparse, finer and dense, larger, partly longitudinally rugose punctures, without smooth midline, with numerous short setae and a few longer in the larger punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin finely serrate at apex, only weakly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 3.64; dorsal margin sharply carinate, with two groups of spines, basal group at middle, apical one at three quarter of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally impunctate; metatarsomeres in holotype missing. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 6 E – H. Habitus: Fig. 6 I.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF83FFBAFF3E75432E6ADD8F.taxon	diagnosis	Diagnosis. Gastroserica lucidomarginalis Ahrens, Lukic & Liu, new species is in external appearance and shape of aedeagus similar to Gastroserica ivoi Ahrens, Lukic & Liu, new species Gastroserica lucidomarginalis Ahrens, Lukic & Liu, new species differs by shorter antennal club in male, and by the shape of aedeagus: the ventral lamina of phallobase is more pronounced (lateral view); the basal lobe of the left paramere is distally much longer being almost subequal to the distal lobe of the paramere; the right paramere is preapically more strongly narrowed.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF83FFBAFF3E75432E6ADD8F.taxon	etymology	Etymology. The name of the new species is derived from the combined Latin words “ lucidus ” (shiny) and “ marginalis ” (bordered) (adjective in singular nominative), with reference to the shiny surface and the dark elytral margin.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF83FFBAFF3E75432E6ADD8F.taxon	description	Variation. Length: 4.9 – 5.6 mm, length of elytra: 3.4 – 3.5 mm, width: 3.1 – 3.2 mm. Metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined and twice as long as dorsal tibial spur. The species has an entire dark (black, iridescent shiny) and yellow form with dark elytral margins (holotype). Female: completely yellow, antennal club short, composed of three antennomeres; pygidium flat. Genbank accessions. Table 1.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF80FFBBFF3E734C2E9CDC47.taxon	description	Figs. 6 J – M, 24	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF80FFBBFF3E734C2E9CDC47.taxon	materials_examined	Type material examined. Holotype: Ô “ Phu Rua NP (900 m alt.), Loei P., NE Thai. 26 - 30. IV. 2006 Takakuwa, M. leg. / 1162 Sericini Asia spec. ” (ZFMK). Paratypes: 1 Ô, 3 ♀♀ “ Phu Rua NP (900 m alt.), Loei P., NE Thai. 26 - 30. IV. 2006 Takakuwa, M. leg. ” (ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF80FFBBFF3E734C2E9CDC47.taxon	description	Description of holotype. Length: 4.9 mm, length of elytra: 3.2 mm, width: 2.9 mm. Body oval, yellowish brown, antennal club, frons and margins of elytra blackish, dorsal surface shiny, nearly glabrous. Labroclypeus narrow and almost square, as wide as long, widest at middle, lateral margins convex and weakly convergent to rounded anterior angles; anterior margin moderately emarginated medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface flat, finely and densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons shiny, with dense, fine punctures, beside eyes with a few long setae, otherwise glabrous. Smooth area in front of eyes twice as wide as long. Ocular canthus sparsely finely punctate, narrow and moderately long (1 / 3 of ocular width), with a terminal seta. Eyes very large, ratio of diameter / interocular width: 0.93. Antenna with ten antennomeres; club composed of four antennomeres, reflexed, 2.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at middle, lateral margins evenly convex and convergent to rectangular and moderately produced and almost blunt anterior angles and blunt posterior angles. Anterior margin of pronotum convex, with robust and complete marginal line; basal marginal line absent; surface finely and densely punctate, glabrous; anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, coarsely and densely punctate, on median base impunctate. Elytra short, widest at middle, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine and sparse punctures concentrated along striae, almost entirely glabrous; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface shiny, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.77. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium weakly convex and shiny, with fine and moderately dense punctures, without smooth midline, with numerous short setae and a few longer on apical part. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin finely serrate at apex, only weakly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 3.1; dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical one at two thirds of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, glabrous; ventral margin finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally impunctate; metatarsomeres in holotype missing. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 6 J – L. Habitus: Fig. 6 M.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF80FFBBFF3E734C2E9CDC47.taxon	diagnosis	Diagnosis. Gastroserica loei Ahrens, Lukic & Liu, new species is in external appearance and shape of aedeagus similar to Gastroserica lucidomarginalis Ahrens, Lukic & Liu, new species and G. ivoi Ahrens, Lukic & Liu, new species Gastroserica loei differs by the very large eyes, the long antennal club, and the (in relation to the phallobase) rather short parameres, while the characteristics of the phallobase are more similar to G. piceocoerulea Ahrens, Lukic & Liu, new species.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF80FFBBFF3E734C2E9CDC47.taxon	etymology	Etymology. The name of the new species is named after its occurrence in Loei province (noun in apposition).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF80FFBBFF3E734C2E9CDC47.taxon	description	Variation. Length: 4.5 – 5.4 mm, length of elytra: 3.1 – 3.8 mm, width: 2.6 – 3.4 mm. Metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere little shorter than following two tarsomeres combined and twice as long as dorsal tibial spur. Female: antennal club short, composed of three antennomeres, little shorter than remaining antennomeres combined; eyes little smaller than in male, ratio of diameter / interocular width: 0.6.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF81FFB6FF3E73142E69DA73.taxon	description	Figs. 7 A – E, 26	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF81FFB6FF3E73142E69DA73.taxon	materials_examined	Type material examined. Holotype: Ô “ 839470 / 839470 Microserica spTHAI _ DE 09 _ 1 Thailand L. Dembicky 9. - 13.5.2009 Doi Chiang Dao env. Chiang Mai Prov. 19 ° 24 ’ 45 ’’ N 98 ° 51 ’ 30 ’’ E 1200 m / 22 Sericini Asia spec. ” (ZFMK). Paratypes: Thailand: 1 Ô “ THAI 1. - 8. V. 1993 Soppong Pai 1800 m Pacholatko & Dembicky leg. / TS 31 / 16 Sericini Asia spec. ” (CP), 1 Ô “ Thailand bor. Occ. 30.4. - 14.5.1991 Mae Hong Son env. Ban Huai Po 1800 m ” (ZFMK), 12 ÔÔ “ NE Thailand 1. - 15.5 [?]. 1991 Mae Hong Son Ban Huai Po 800 - 1600 m S. Bily leg. ” (NHMB), 3 ÔÔ “ NE Thailand 28.4.1991 Mae Hong Son S. Bily leg. ” (NHMB), 17 ÔÔ “ NW Thailand, 19.19 N, 97.97 E Mae Hong Son, 1991 Ban Huai Po, 1600 - 2000 m 17. - 23.5., L. Dembicky leg. ” (NHMW), 2 ÔÔ “ NW Thailand 9. - 16.5.1991 Mae Hong Son, Ban Huai Po 1600 - 2000 m J. Horák leg. ” (ZFMK), 1 Ô “ NW Thailand Ban Huai Po 9. - 16.5.1991 I. Horák leg. ” (ZFMK), 1 Ô “ 839469 / 839469 Microserica spTHAI _ DE 09 _ 1 Thailand L. Dembicky 9. - 13.5.2009 Doi Chiang Dao env. Chiang Mai Prov. 19 ° 24 ’ 45 ’’ N 98 ° 51 ’ 30 ’’ E 1200 m ” (ZFMK), 1 Ô “ NW Thailand 1. - 6.5.1991 Soppong Pai 1800 m Leg. Pacholátko / TS 32 ” (CP), 1 Ô “ Chieng Mai N. Thailand V-VI. 1983 / Museum Leiden coll. P. J. J. H. Kuijten ” (RMNH), 2 ÔÔ “ Thai, 17. - 26. V. 1991 Chiang Dao 1000 m 19 ° 25 ’ N 98 ° 52 ’ E Vít Kubáň leg. ” (NMPC, ZFMK), 1 Ô “ Thai, 10. - 16. V. 1991 Chiang Dao 600 m 19 ° 24 ’ N 98 ° 55 ’ E V. Kuban leg. / coll. Milan Nikodym, Praha ” (ZFMK), 2 ÔÔ “ NW Thailand, Chieng Dao, Ban San Pakia, 5. - 10. v. 2004, 1200 m, Sv. Bily ” (NME). China: 1 Ô [China] “ Yunnan 1500 - 2500 m, 25.22 N 98.49 E 17 - 24 / 5 Gaoligong mts. Vit Kubáň leg. 1995 ” (CP), 7 ÔÔ “ China: S-Yunnan (Xishuangbanna) 45 km SW Jinghong vic. Bangzhang vill. / N 21 ° 44 ’ 37 ’’, E 100 ° 27 ’ 02 ’’, 16 - 1700 m, 03. - 05. V. 2009 leg. A. Weigel, blüh Cast ” (NME). 1 Ô [China] “ Meng’e, Xishuangbanna, Yunnan, 10. V. 1958, 1050 - 1080 m, leg. Wang Shuyong / LW- 341 ” (IZAS), 1 Ô [China] “ Meng’e, Xishuangbanna, Yunnan, 22. V. 1958, 1000 m, leg. Pu Fuji ” (IZAS), 3 ÔÔ [China] “ Mengzhe, Xishuangbanna, Yunnan, 15. VI. 1958, 1200 m, leg. Pu Fuji, Chen Zhizi / LW- 377 ” (IZAS). Laos: 1 Ô “ Laos, 1. - 16. v. 1999, Louangphrabang pr., 20 ° 33 - 4 ’ N 102 ° 14 ’ E Ban Song Cha (5 km W) 1200 m, Vit Kubáň leg. ” (CP), 1 Ô “ Laos north, 13. - 24. V. 1997 15 km NW Louang Namtha N 21 ° 07.5 ’, E 101 ° 21.0 alt. 750 + 100 m, M. Štrba & R. Hergovits leg. ” (CP), 1 Ô “ Laos Vientiane Prov. Ban Van Eue 15 - 31. V. 1965 / Native collector Bishop Museum ” (BPBM), 1 Ô “ LAOS, Xieng Khouang prov., 19 ° 37 – 8 ’ N, 103 ° 20 ’ E, Phonsavan (30 km NE) Phou Sane Mt., 1420 m, 10 - 30. v. 2009, D. Hauck leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 3 ÔÔ “ LAOS, Xieng Khouang prov., 19 ° 37 – 8 ’ N, 103 ° 20 ’ E, Phonsavan (30 km NE) Phou Sane Mt., 1420 m, 10 - 30. v. 2009, M. Brancucci leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 16 ÔÔ “ LAOS, Hua Phan prov., 20 ° 13 ’ N, 104 ° 00 ’ E, Phou Pane Mt., 1 - 16. vi. 2009, 1350 - 1500 m, M. Brancucci leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 8 ÔÔ “ LAOS, Houa Phan prov., 20 ° 12 - 13 ’ N, 103 ° 59.5 ’ - 104 ° 01 ’ E, Ban Saluei-> Phou Pane Mt., 1340 - 1870 m, 10. v. - 16. vi. 2009, M. Brancucci & local coll. leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 1 Ô “ LAO-NE, Hua Phan prov., 20 ° 12 ’ N 104 ° 01 ’ E, Phu Phan Mt., 1500 - 1900 m, 17. v. - 3. vi. 2007, M. Brancucci leg. / NHMB Basel expedition to Laos, 2007 ” (NHMB), 5 ÔÔ “ LAOS-NE, Houa Phan prov., Ban Saluei-> Phou Pane Mt., 20 ° 12 - 13.5 ’ N 103 ° 59.5 ’ - 104 ° 01 ’ E, 1340 - 1870 m, 15. iv. - 15. v. 2008, Lao collectors leg. ” (NMPC), 1 Ô “ LAONE, Hua Phan prov., 20 ° 12 ’ N 104 ° 01 ’ E, Phu Phan Mt., 1750 m, 17. v. - 3. vi. 2007, Vít. Kuban leg. / NHMB Basel expedition to Laos, 2007 ” (NHMB), 24 ÔÔ “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ’ N, 104 ° 01 ’ E; 1300 - 1900 m; 01. - 31. v. 2011; leg. C. Holzschuh Ankauf ZFMK Bonn 2011 ” (ZFMK), 178 ÔÔ “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ’ N, 104 ° 01 ’ E; 11. iv. - 15. v. 2012; 1300 - 1900 m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012 ” (ZFMK), 122 ÔÔ “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ’ N, 104 ° 01 ’ E; 14. iv. - 15. v. 2012; 1300 - 1900 m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012 / 13 ” (ZFMK), 3 ÔÔ “ LaosNE Hua Phan prov. 20 ° 13 ’ 09 - 19 ’ N 103 ° 59 ’ 54 ’’ - 104 ° 00 ’ 03 ’’ E 1480 - 1510 m, Phou Pane Mt., 22. iv. - 14. v. 2008,. Vit Kuban leg ” (ZFMK), 4 ÔÔ “ Laos-NE Hua Phan prov. 20 ° 12 ’ N, 104 ° 01 ’ E Phou Phan Mt., 1750 m 17.5. - 3.6.2007, Vít Kubáň leg ” (ZFMK), 1 Ô “ X-DA 4726 / X- DA 4726 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 15. - 30. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica spLAOS Phupane 10 ” (ZFMK), 1 Ô “ X-DA 4934 / X- DA 4934 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 1. - 2. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica spLAOS Phupane 10 ” (ZFMK), 1 Ô “ X-DA 4937 / X- DA 4937 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 1. - 2. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica spLAOS Phupane 10 ” (ZFMK), 1 Ô “ X-DA 4939 / X- DA 4939 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 1. - 2. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica spLAOS Phupane 10 ” (ZFMK), 1 Ô “ X-DA 4720 / X- DA 4720 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 15. - 30. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica spLAOS Phupane 10 ” (ZFMK). Additional material examined. 1 Ô [China] “ Meng’a, Xishuangbanna, Yunnan, 20. V. 1958, 1050 - 1080 m, leg. Pu Fuji / LW- 388 ” (IZAS; not labeled as paratype as specimen lost, only genitalia available).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF81FFB6FF3E73142E69DA73.taxon	description	Description of holotype. Length: 5.0 mm, length of elytra: 3.5 mm, width: 3.0 mm. Body oval, body dark brown to blackish, pronotum with some greenish shine, elytra blue-iridescent, legs and antennal club brown, antennal funiculus yellow, dorsal surface shiny, nearly glabrous. Labroclypeus narrow and almost semicircular, as wide as long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles; anterior margin weakly incised medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface medially weakly convexly elevated, finely and coarsely, densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons shiny, with moderately dense, fine punctures, beside eyes and behind frontoclypeal suture with a few single short setae, otherwise with minute setae in punctures. Smooth area in front of eyes twice as wide as long. Ocular canthus sparsely finely punctate, narrow and moderately long (1 / 3 of ocular width), with a terminal seta. Eyes moderately large, ratio of diameter / interocular width: 0.64. Antenna with ten antennomeres; club composed of four antennomeres, weakly reflexed, 1.8 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest shortly before base, lateral margins evenly convex and convergent to rectangular and moderately produced anterior angles, posterior angles blunt and moderately rounded. Anterior margin of pronotum convex, with robust and complete marginal line; basal marginal line absent; surface finely and moderately densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, coarsely and densely punctate, on median base less densely punctate. Elytra short, widest at middle, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd intervals with a few short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface shiny, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.62. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and shiny, with mixed finer and dense, larger, partly longitudinally rugose punctures, without smooth midline, with numerous short setae and a few longer in the larger punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin finely serrate at apex, only weakly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 2.95; dorsal margin sharply carinate, with two groups of spines, basal group at middle, apical one at three quarter of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally impunctate; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined and little less than twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 7 A – D. Habitus: Fig. 7 E. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF81FFB6FF3E73142E69DA73.taxon	diagnosis	Diagnosis. Gastroserica piceocoerulea Ahrens, Lukic & Liu, new species is in external appearance similar to G. lucens (Ahrens & Fabrizi, 2009), but in shape of aedeagus more similar to Gastroserica lucidomarginalis Ahrens, Lukic & Liu, new species. Gastroserica piceocoerulea differs from G. lucidomarginalis Ahrens, Lukic & Liu, new species by the dark body color, and by the shape of aedeagus: the phallobase is less curved (lateral view), its ventral lamina is reduced in length but present on both sides before apex of ventral phallobase (lateral view); the distal margin of the basal lobe of the left paramere is convex rather than sharply pointed as in G. lucidomarginalis Ahrens, Lukic & Liu, new species. From G. lucens Ahrens & Fabrizi, 2009, G. piceocoerulea differs by the longer antennal club and by the longer basal lobe of the left paramere.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF81FFB6FF3E73142E69DA73.taxon	etymology	Etymology. The name of the new species is derived from the combined Latin words “ piceus ” (black) and “ coeruleus ” (blue) (adjective in singular nominative), with reference to the dark, iridescent shiny body surface. Variation. Length: 4.6 – 6.0 mm, length of elytra: 3.0 – 3.5 mm, width: 2.8 – 3.2 mm. Female unknown. Genbank accessions. Table 1.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8CFFB7FF3E750B288EDAC3.taxon	description	Figs. 7 F – I, 25	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8CFFB7FF3E750B288EDAC3.taxon	materials_examined	Type material examined. Holotype Ô “ Burma (Myanmar) SW Shan state Taunggyi J. Rejsek 1. - 18.6.1997 / Coll. Dirk Ahrens ” (ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8CFFB7FF3E750B288EDAC3.taxon	description	Description of holotype. Length: 5.4 mm, length of elytra: 3.4 mm, width: 2.9 mm. Body oval, body yellowish brown, pronotum and scutellum reddish brown, frons and elytra dark blue to blackish, elytra iridescent, legs and antennal club brown, antennal funiculus yellow, dorsal surface shiny, nearly glabrous. Labroclypeus subrectangular, distinctly wider than long, widest at base, lateral margins basally parallel, moderately convex and convergent to strongly rounded anterior angles; anterior margin almost straight, very weakly incised medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface medially weakly convexly elevated, finely and coarsely, densely punctate, with a few long, erect setae; frontoclypeal suture finely incised and curved medially. Frons shiny, with dense, fine punctures, glabrous. Smooth area in front of eyes 1.5 times as wide as long. Ocular canthus sparsely finely punctate, narrow and moderately long (1 / 3 of ocular width), without terminal seta. Eyes large, ratio of diameter / interocular width: 0.84. Antenna with ten antennomeres; club composed of four antennomeres, weakly reflexed, twice as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at posterior quarter, lateral margins evenly convex and convergent to rectangular and moderately produced anterior angles and to the blunt posterior angles. Anterior margin of pronotum convex, with fine, complete marginal line; basal marginal line absent; surface finely and moderately densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, evenly finely and densely punctate. Elytra short, widest at middle, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, penultimate lateral intervals with a few short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface shiny, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.5. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and shiny, with dense, fine, partly longitudinally rugose punctures, without smooth midline, with a few short setae at apex. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin finely serrate at apex, only weakly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 3.1; dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical one at two thirds of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally impunctate; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere little shorter than following two tarsomeres combined and twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 7 F – H. Habitus: Fig. 7 I. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8CFFB7FF3E750B288EDAC3.taxon	diagnosis	Diagnosis. Gastroserica rubropicea Ahrens, Lukic & Liu, new species is in shape of aedeagus similar to Gastroserica lucidomarginalis Ahrens, Lukic & Liu, new species and Gastroserica piceocoerulea Ahrens, Lukic & Liu, new species. Gastroserica rubropicea new species differs from both by the characteristic coloration. Furthermore, the ventral lamina of phallobase is entirely lacking in Gastroserica rubropicea Ahrens, Lukic & Liu, new species; the distal margin of the basal lobe of the left paramere is sharply pointed as in Gastroserica lucidomarginalis Ahrens, Lukic & Liu, new species, but the phallobase is not as strongly curved as in the latter.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8CFFB7FF3E750B288EDAC3.taxon	etymology	Etymology. The name of the new species is derived from the combined Latin words rubrus (red) and piceus (black) (adjective in singular nominative), with reference to the red and blackish body surface color.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8DFFB0FF3E75902F0ADAC3.taxon	description	Figs. 8 A – E, 24	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8DFFB0FF3E75902F0ADAC3.taxon	materials_examined	Type material examined. Holotype: Ô “ Viet Nam Dalat 6 km S 1400 - 1500 m 9. VI. - 7. VII. 1964 ” (BPBM). Additional material examined. 1 Ô, 1 ♀ “ C-Vietnam, Tira Tien Hue Prov., Phu Loc Bach Ma NP, Top area 05. - 09. V. 2019, 1250 - 1400 m, 16 ° 11 ’ 39 ’’ N 107 ° 51 ’ 12 ’’ E, LF, leg. M. Hartmann / 1132 Sericini Asia spec. ” (NME).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8DFFB0FF3E75902F0ADAC3.taxon	description	Redescription. Length: 5.9 mm, length of elytra: 4.5 mm, width: 3.6 mm. Body oval, body including legs and antenna yellow, multiple symmetric, small dots on pronotum and elytra, abdomen including pygidium dark brown, frons posteriorly slightly darker; dorsal surface dull, nearly glabrous. Labroclypeus subrectangular, distinctly wider than long, widest at base, lateral margins basally slightly narrowed, convex and convergent to strongly rounded anterior angles; anterior margin weakly incised medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface weakly shiny, medially weakly convexly elevated, coarsely, densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons dull, with dense, fine punctures, with a few long setae beside eyes, otherwise with minute setae in punctures. Smooth area in front of eyes narrow, only as wide as long. Ocular canthus wide, short (1 / 3 of ocular width), sparsely finely punctate, without terminal seta. Eyes small, ratio of diameter / interocular width: 0.53. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.1 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at middle, lateral margins evenly convex and convergent to rectangular and moderately produced anterior angles and to well-rounded posterior angles. Anterior margin of pronotum straight, with fine, complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, evenly finely and densely punctate, impunctate along midline. Elytra short, widest at middle, striae finely impressed, finely and densely punctate and darker, intervals weakly convex, with fine and very sparse punctures concentrated along striae, with minute setae in punctures, elytra with a single band of transversal and symmetrical dark dots, of which the ones of the odd intervals are positioned half of their length before the ones of the even intervals, odd intervals with a few short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface shiny, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.7. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and shiny, with dense, fine punctures, without smooth midline, almost glabrous, with a few short setae at apex, otherwise with minute setae in punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin distinctly serrate in apical entire half and widened, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 3.2; dorsal margin sharply carinate, with two groups of spines, basal group at middle, apical one at three quarters of metatibial length, in basal third with a few robust single setae in punctures with serrated margin; lateral face longitudinally convex, finely and sparsely punctate, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally impunctate; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined and twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 8 A – D. Habitus: Fig. 8 E. Variation. Female: Color less dark, dots on elytra less clear, frons yellow; antennal club of female short, as long as remining antennomeres combined, composed of three lamellae.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8AFFB1FF3E7590291ED90F.taxon	description	Figs. 8 F – I, 24	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8AFFB1FF3E7590291ED90F.taxon	materials_examined	Type material examined. Holotype: Ô “ LAOS centr., Bolikhamsai prov., Ban Nok env., 9. - 14.5.1998 Route No. 8, alt. 220 + 50 m, N 18 ° 08.7, E 104 ° 28.1 (GPS), E. Jendek & O. Šauša leg. / 37 Sericini Asia spec. ” (CP). Paratypes: 1 Ô “ LAOS, Sekong prov. ca. 12 km S Sekong, Tad Faek waterfalls (at light) 15 ° 14.7 ’ N, 106 ° 45.1 ’ E, 118 m Jiři Hájek leg. 8. + 12. v. 2010 / 1128 Sericini Asia spec. ” (NMPC).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8AFFB1FF3E7590291ED90F.taxon	description	Description of holotype. Length: 5.0 mm, length of elytra: 3.1 mm, width: 2.8 mm. Body oval, yellow, antennal club brown, margins of elytra and frons blackish, dorsal surface shiny, nearly glabrous. Labroclypeus narrow and almost square, as wide as long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles; anterior margin weakly incised medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface medially weakly convexly elevated, finely and coarsely, densely punctate, with a few long, erect setae; frontoclypeal suture finely incised and curved medially. Frons shiny, with sparse, fine punctures, beside eyes with a few single long setae, otherwise glabrous. Smooth area in front of eyes as wide as long. Ocular canthus sparsely finely punctate, narrow and moderately long (1 / 3 of ocular width), with a terminal seta. Eyes moderately large, ratio of diameter / interocular width: 0.77. Antenna with ten antennomeres; club composed of four antennomeres, weakly reflexed, 1.8 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at posterior third, lateral margins evenly convex and convergent to rectangular and moderately produced anterior angles, posterior angles blunt and moderately rounded. Anterior margin of pronotum convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, coarsely and densely punctate, on midline impunctate. Elytra short, widest at middle, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, penultimate lateral interval with a few short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface shiny, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.73. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and shiny, with mixed sparse, finer and dense, larger, partly longitudinally rugose punctures, without smooth midline, with numerous short setae and a few longer in the larger punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin finely serrate at apex, only weakly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia short and wide, widest at middle, ratio width / length: 1 / 2.9; dorsal margin sharply carinate, with two groups of spines, basal group shortly before middle, apical one at two thirds of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally impunctate; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere little shorter than following two tarsomeres combined and twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 8 F – H. Habitus: Fig. 8 I. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8AFFB1FF3E7590291ED90F.taxon	diagnosis	Diagnosis. Gastroserica bannok Ahrens, Lukic & Liu, new species is in external appearance G. lucidomarginalis Ahrens, Lukic & Liu, new species, while in shape of aedeagus it is more similar to G. cognata (Frey, 1972). From the latter G. bannok Ahrens, Lukic & Liu, new species differs by the lacking dots on the elytra and the shape of parameres: the phallobase is more strongly curved (lateral view) in G. cognata, the left paramere has the basal lobe more developed mesally rather basally, as in G. cognata. From G. lucidomarginalis Ahrens, Lukic & Liu, new species G. bannok Ahrens, Lukic & Liu, new species differs by the basal lobe of the left paramere being not developed distally but only mesally.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8AFFB1FF3E7590291ED90F.taxon	etymology	Etymology. The name of the new species is derived from the type locality, Ban Nok (noun in apposition).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8BFFB1FF3E76D72F52D8AD.taxon	description	Fig. 24	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8BFFB1FF3E76D72F52D8AD.taxon	materials_examined	Material examined. See Ahrens & Fabrizi (2009 b: 267). Aedeagus. See Ahrens & Fabrizi (2009 b: p. 282, fig. 6 D – F).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8BFFB1FF3E76D72F52D8AD.taxon	distribution	Distribution. Eastern Himalaya (Arunachal Pradesh).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8BFFB2FF3E70712EFCD8EF.taxon	description	Figs. 9 A – D, 25	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8BFFB2FF3E70712EFCD8EF.taxon	materials_examined	Type material examined. Paratype: 1 Ô “ Viet Nam 20 km Di Linh (Djiring) 22 - 28. IV. 60 / L. W. Quate collector / Paratype Microserica quateorum G. Frey 1972 ” (CF).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF8BFFB2FF3E70712EFCD8EF.taxon	description	Redescription (of paratype). Length: 5.2 mm, length of elytra: 3.8 mm, width: 3.8 mm. Body oval, body brown, head, legs, antenna, metacoxa, anterior parts of elytra, and margins and a median line of pronotum yellowish, dorsal surface iridescent shiny, partly dull; nearly glabrous. Labroclypeus subrectangular, slightly wider than long, widest at base, lateral margins in basal half subparallel and straight, anteriorly strongly convex and convergent to strongly rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, medially weakly convex, finely and coarsely, densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons dull-iridescent, with sparse, fine punctures, beside eyes and behind lateral frontoclypeal suture with a few single short setae, otherwise with minute setae in punctures. Smooth area in front of eyes as wide as long. Ocular canthus sparsely finely punctate, narrow and moderately long (1 / 3 of ocular width), with a terminal seta. Eyes small, ratio of diameter / interocular width: 0.53. Antenna with ten antennomeres; club composed of four antennomeres, slightly reflexed at apex, 1.5 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins evenly convex and convergent to sharp and distinctly produced anterior angles, posterior angles strongly rounded. Anterior margin of pronotum weakly convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, on median base impunctate. Elytra short, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, penultimate lateral interval with a few short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.86. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium weakly convex and dull, with mixed finer and larger punctures, without smooth midline, with numerous fine adpressed along the apical margin and a few longer erect setae at apex. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin serrate in apical half, weakly widened; dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 2.7; dorsal margin sharply carinate, with two groups of spines, basal group at middle, apical one at three quarter of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere little shorter than following two tarsomeres combined, almost twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 9 A – C. Habitus: Fig. 9 D.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF88FF8CFF3E72452D7AD9EB.taxon	description	Figs. 10 A – D, 27	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF88FF8CFF3E72452D7AD9EB.taxon	materials_examined	Type material examined. Syntypes: 1 Ô “ Burma Myndaik 4500 ft. 5. V. 18 A. G. R. Coll. / Microserica simplex Type Arrow ” (BMNH), 1 ♀ “ Burma Myndaik 4500 ft. 5. V. 18 A. G. R. Coll. ” (BMNH). Additional material examined. Thailand. 18 ex. “ NE Thailand 1. - 15.5 [?]. 1991 Mae Hong Son Ban Huai Po 800 - 1600 m S. Bily leg. ” (NHMB), 10 ex. “ NE Thailand 8. - 17. V. 1992 Mae Hong Son Ban Huai Po 1600 m, S. Bily leg. ” (NHMB), 43 ex. “ NW Thailand, 19.19 N, 97.97 E Mae Hong Son, 1991 Ban Huai Po, 1600 - 2000 m 17. - 23.5., L. Dembicky leg. ” (NHMW), 2 ÔÔ “ NW Thailand 9. - 16.5.1991 Mae Hong Son, Ban Huai Po 1600 - 2000 m J. Horák leg. ” (ZFMK), 7 ex. “ Thai 28 / 4. - 6 / 5.91 Umphang river 1000 m 16 ° 07 ‘ N 99 ° 99 ‘ E David Kral lgt. ” (CK), 4 ex. “ N. Thailand Mae Hong Son, Ban Huai Po 9. - 16.5.1991 1600 - 2000 m I. Horak lgt. ” (ZFMK), 2 ex. “ NW Thailand 19.19 N, 97.59 E Mae Hong Son 1991 Ban Huai Po 1600 - 1900 m 17. - 23.5., L. Dembicky leg. ” (NHMB), 1 ex. “ Thailand Bor. Occ. 30.04. - 14.05.1991 Mae Hong Son env. Ban Huai Po, 1800 m, Jan Farkac leg. ” (NHMB), 1 ex. “ Thai 17. - 24. V. 1991 Chiang Dao 1000 m 19 ° 25 ‘ N 98 ° 52 ‘ E Vit Kuban leg. “ (CN), 1 ex. “ Thailand, 7. - 12. V. 1996 Mae Hong Son Prov. Soppong 1500 m 19 ° 27 ‘ N 98 ° 20 ‘ E lgt. S. Becvar ” (ZFMK), 2 ÔÔ “ Thailand bor. Hang Dong 6.1990 lgt. Wimmer ” (ZFMK), 3 ÔÔ “ Thailand centr. Pa-Sak Fl. IX- 86 Rhtchabunn- 600 m ” (ZFMK). China. 3 ex. (Ô) [China] “ Yunnan 1800 - 2000 m 25.04 N 101.55 E Yipinglang 17 - 20.6. Vit Kubáň leg. 1994 / CS 76 ” (CP), 2 ÔÔ [China] “ Mengzhe, Xishuangbanna, Yunnan, 15. VI. 1958, 1200 m, leg. Wang Shuyong / LW- 380 ” (IZAS), 1 Ô “ Huanian, 15. VI. 1983 ” (IZAS). Laos. 334 ex. “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ’ N, 104 ° 01 ’ E; 14. iv. - 15. v. 2012; 1300 - 1900 m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012 / 13 ” (ZFMK), 760 ex. “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ’ N, 104 ° 01 ’ E; 11. iv. - 15. v. 2012; 1300 - 1900 m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012 ” (ZFMK), 10 ex. “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ’ N, 104 ° 01 ’ E; 1300 - 1900 m; 01. - 31. V. 2011; leg. C. Holzschuh Ankauf ZFMK Bonn 2011 ” (ZFMK), 2 ex. “ Laos-NE Hua Phan prov., 20 ° 12 ’ N, 104 ° 01 ’ E, Phu Phan Mt., 1500 - 1900 m, 17.5. - 3.6.2007, leg. C. Holzschuh ” (ZFMK), 2 ex. “ Laos-NE Hua Phan prov., 20 ° 12 ’ N, 104 ° 01 ’ E, Phu Phan Mt., 1500 - 1900 m, 17.5. - 3.6.2007, leg. Vit. Kuban ” (ZFMK), 1 Ô “ X-DA 4933 / X- DA 4933 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 2. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica simplex ” (ZFMK), 1 Ô “ X-DA 4932 / X- DA 4932 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 2. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica simplex ” (ZFMK), 1 Ô “ X-DA 4718 / X- DA 4718 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 15. - 30. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica simplex ” (ZFMK), 1 Ô “ X-DA 4711 / X- DA 4711 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 15. - 30. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica simplex ” (ZFMK), 1 Ô “ X-DA 4710 / X- DA 4710 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 15. - 30. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica simplex ” (ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFF88FF8CFF3E72452D7AD9EB.taxon	description	Redescription. Length: 6.4 mm, length of elytra: 3.8 mm, width: 3.6 mm. Body oval, ventral face, antennal funiculus, and legs yellowish brown, abdomen and antennal club brown, dorsal surface reddish brown, dorsal surface dull, sometimes partly iridescent shiny, nearly glabrous. Labroclypeus wider than long, widest at middle, lateral margins in basal half subparallel, strongly convex anteriorly and convergent to strongly rounded anterior angles; anterior margin distinctly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, medially convex, finely and coarsely, densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and bent medially. Frons dull, with sparse, fine punctures, beside eyes and behind frontoclypeal suture with a few single short setae, otherwise glabrous. Smooth area in front of eyes 1.5 times as wide as long. Ocular canthus narrow and moderately long (1 / 3 of ocular width), sparsely finely punctate, with a terminal seta. Eyes moderately small, ratio of diameter / interocular width: 0.58. Antenna with ten antennomeres; club composed of six antennomeres, reflexed, 1.7 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins evenly convex and convergent to blunt, moderately produced anterior angles, posterior angles blunt. Anterior margin of pronotum weakly convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, lateral and lateral anterior margins densely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, on midline impunctate. Elytra short oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and moderately dense punctures concentrated along striae, with minute setae in punctures, odd intervals with a few sparse short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.67. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, with mixed finer and larger punctures, without smooth midline, with numerous short setae and a few longer in the larger punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin finely serrate at apex, only weakly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 3.0; dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical one at three quarter of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere little shorter than following two tarsomeres combined, twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 10 A – C. Habitus: Fig. 10 D. Variation. Female: Antennal club composed of three antennomeres, as long as remaining antennomeres combined.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB6FF8DFF3E76B32813DF37.taxon	description	Figs. 11 A – D, 27	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB6FF8DFF3E76B32813DF37.taxon	materials_examined	Type material examined. Holotype: Ô “ Burma (Myanmar) SW Shan state Taunggyi J. Rejsek 1. - 18.6.1997 / Coll. Dirk Ahrens / 10 Asia Sericini spec. ” (ZFMK). Paratypes: Myanmar: 7 ÔÔ “ Maymyo, Burma. H. L. Andrewes. / Andrewes Bequest. B. M. 1922 - 221. ” (BMNH), 5 ÔÔ, 3 ♀♀ “ Myanmar, Shan State, near Nyaung Shwe (Inle lake) 20 ° 39 ’ 18 ’’ N, 96 ° 52 ’ 58 ’’ E, 965 m, 28. VI. 2006, leg. M. Homburg ” (ZFMK). Thailand: 1 Ô “ NW Thailand 9. - 16. V. Mae Hong Son 1991 Ban Huai Po 1600 m leg. P. Pacholátko / TS 72 ” (CP), 23 ÔÔ “ NE Thailand 1. - 15.5 [?]. 1991 Mae Hong Son Ban Huai Po 800 - 1600 m S. Bily leg. ” (NHMW, ZFMK), 1 Ô, 3 ♀♀ “ NW Thailand 9. - 16.5. Mae Hong Son, 1991 Ban Huai Po 1600 - 2000 m J. Horák leg. ” (ZFMK), 3 ÔÔ “ NW Thailand 9. - 16.5.1991 Mae Hong Son, Ban Huai Po 1600 - 2000 m J. Horák leg. ” (ZFMK), 9 ÔÔ, 1 ♀ “ N. Thailand Ban Huai Po 9. - 16.5. 1991 I. Horák leg. / Coll. Milan Nikodym Praha ” (ZFMK), 5 ÔÔ “ NE Thailand 28.4.1991 Mae Hong Son S. Bily leg. ” (NHMW, ZFMK), 6 ÔÔ “ NW Thailand 8. - 17. V. 1992 Mae Hong Son, Ban Huai Po 1500 m S. Bily leg. ” (NHMW, ZFMK), 2 ÔÔ “ Thailand bor. occ. 30.4. - 14.5.1991 Mae Hong Son env. Ban Huai Po 1800 m Jan Farkac leg. ” (NHMW), 35 ÔÔ “ NW Thailand, 19.19 N, 97.59 E Mae Hong Son, 1991 Ban Huai Po 1600 - 2000 m 17. - 23.5., L. Dembicky leg. ” (NHMW, ZFMK), 1 Ô “ Ban Angkhai, alt. 750 m Samoeng Distr. Chiang Mai pref. Thailand 15 - 20 - V- 1998 K. Matsumoto leg. ” (ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB6FF8DFF3E76B32813DF37.taxon	description	Description of holotype. Length: 4.9 mm, length of elytra: 2.9 mm, width: 2.9 mm. Body oval, dark brown, antennal club and tarsi yellowish brown, dorsal surface dull, sometimes partly iridescent shiny, nearly glabrous. Labroclypeus little wider than long, widest at middle, lateral margins in basal half subparallel, strongly convex anteriorly and convergent to strongly rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, almost flat, finely and coarsely, densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and bent medially. Frons dull, with sparse, fine punctures, with minute setae in punctures, otherwise glabrous. Smooth area in front of eyes as wide as long. Ocular canthus narrow and moderately long (1 / 3 of ocular width), sparsely finely punctate, without terminal seta. Eyes small, ratio of diameter / interocular width: 0.52. Antenna with ten antennomeres; club composed of six antennomeres, straight, 1.5 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins evenly convex and convergent to blunt, moderately produced anterior angles, posterior angles blunt. Anterior margin of pronotum weakly convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, lateral and lateral anterior margins glabrous. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, on midline impunctate. Elytra short oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with minute setae in punctures, penultimate lateral interval with a few sparse short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.76. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium moderately convex and dull, with mixed finer and larger punctures, without smooth midline, with numerous short setae and a few longer in the larger punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin entirely smooth, distinctly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 2.78; dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical one at three quarter of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere little shorter than following two tarsomeres combined, almost twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 11 A – C. Habitus: Fig. 11 D. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB6FF8DFF3E76B32813DF37.taxon	diagnosis	Diagnosis. Microserica allovarians Ahrens, Lukic & Liu, new species is in external appearance and shape of aedeagus most similar to Mic. simplex Arrow, 1946. Microserica allovarians Ahrens, Lukic & Liu, new species differs by the darker body color and the parameres being both narrow and long.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB6FF8DFF3E76B32813DF37.taxon	etymology	Etymology. The species name is derived from the combined Greek prefix “ allo ” (other) and the species name “ varians ”, with reference to the certain similarity to M. varians.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB6FF8DFF3E76B32813DF37.taxon	description	Variation. Length: 4.6 – 4.9 mm, length of elytra: 2.9 – 3.0 mm, width: 2.9 – 3.0 mm.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB7FF8EFF3E70CF2E69DC23.taxon	description	Figs. 11 E – H, 27	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB7FF8EFF3E70CF2E69DC23.taxon	materials_examined	Type material examined. Holotype Ô “ THAILAND bor. Chiang Dao env. 21.5. - 4.6.1995 lgt. Snizek M. / 1110 Sericini Asia spec. ” (ZFMK). Paratypes: Thailand: 2 ♀♀ “ THAILAND bor. Chiang Dao env. 21.5. - 4.6.1995 lgt. Snizek M. ” (ZFMK), 1 Ô “ Thai, 17. - 24. V. 1991 Chiang Dao 1000 m 19 ° 25 ’ N 98 ° 52 ’ E Vit Kubáň leg. / Thailand 91 Thanon Thong Chai D. Král & V. Kubáň / Coll. Dirk Ahrens ” (ZFMK), 1 Ô “ THAILAND bor. Chiang Mai 15.5.1997 lgt. M. Snizek ” (CP), 1 Ô “ NW Thailand, 25. iv. - 7. v. Chiang Mai prov., Ban San Pakia 1996 Sv. Bilý leg., 1700 m / TS 125 ” (CP), 1 Ô, 4 ♀♀ “ Thai, 17. - 24. V. 1991 Chiang Dao 1000 m 19 ° 25 ’ N 98 ° 52 ’ E Vit Kubáň leg. / Thailand 91 Thanon Thong Chai D. Král & V. Kubáň ” (ZFMK, NMPC), 1 Ô “ 839472 / 839472 Microserica spThai _ DE 09 _ 3 Thailand L. Dembicky 9. - 13.5.2009 Doi Chiang Dao env. Chiang Mai Prov. 19 ° 24 ’ 45 ’’ N 98 ° 51 ’ 30 ’’ E 1200 m ” (ZFMK), 1 Ô, 1 ♀ “ Thai 17 - 24 / 6.91 Doi Chiang Dao mts. 19.25 N, 98.52 E lgt. D. Kral 1000 m ” (NMPC), 22 ÔÔ, 2 ♀♀ “ Thai, 17. - 26. V. 1991 Chiang Dao 1000 m 19 ° 25 ’ N 98 ° 52 ’ E Vít Kubáň leg. / ex. coll. Milan Nikodym, Praha ” (ZFMK), 5 ÔÔ “ NW Thailand, Chieng Dao, Ban San Pakia, 5. - 10. v. 2004, 1200 m, Sv. Bily ” (NME), 3 ÔÔ “ Thai, 9. - 14. V. 1991 Chiang Dao 350 m 19 ° 22 ’ N 98 ° 57 ’ E V. Kuban lg. / coll. Milan Nikodym, Praha ” (ZFMK), 1 Ô, 1 ♀ “ Thai, 10. - 16. V. 1991 Chiang Dao 600 m 19 ° 24 ’ N 98 ° 55 ’ E V. Kuban leg. / coll. Milan Nikodym, Praha ” (ZFMK). China: 1 Ô [China] “ Meng’a, Xishuangbanna, Yunnan, 23. V. 1958, 1050 - 1080 m, leg. Zhang Yiran / LW- 386 ” (IZAS), 1 Ô [China] “ Northern hill of Jingdong, Yunnan, 30. V. 1956, leg. Zhao Yi ” (IZAS), 1 Ô [China] “ Menghun, Xishuangbanna, Yunnan, 17. VI. 1958, 1200 m, leg. Meng Xuwu / LW- 391 ” (IZAS), 1 Ô [China] “ Meng’e, Xishuangbanna, Yunnan, 23. V. 1958, 1050 - 1080 m, leg. Zhang Yiran ” (IZAS), 1 Ô, 1 ♀ “ Mt. Xishan, Xiaguan, Yunnan, 29. Ⅴ. 1955, 1800 - 1900 m, leg. Krischanovsknja / LW- 439 ” (IZAS), 1 Ô “ Shunbixiang, Yangbi County, Yunnan, 21. Ⅴ. 2004, leg. Yang Xiujuan, Liu Yushuang / LW- 468 ” (HBUM).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB7FF8EFF3E70CF2E69DC23.taxon	description	Description of holotype. Length: 5.1 mm, length of elytra: 3.1 mm, width: 2.9 mm. Body oval, reddish brown, ventral surface including abdomen, frons, anterior central margin of pronotum, and elytral margins blackish brown, antennal club and legs yellowish brown, dorsal surface dull, sometimes partly iridescent shiny, nearly glabrous. Labroclypeus little wider than long, widest at base, lateral margins strongly convex and convergent to strongly rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, flat, finely and coarsely, densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and bent medially. Frons dull, with sparse, fine punctures, with minute setae in punctures, otherwise glabrous. Smooth area in front of eyes as wide as long. Ocular canthus narrow and moderately long (1 / 3 of ocular width), sparsely finely punctate, without terminal seta. Eyes small, ratio of diameter / interocular width: 0.57. Antenna with ten antennomeres; club composed of six antennomeres, straight, 1.5 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins evenly convex and convergent to blunt, moderately produced anterior angles, posterior angles blunt. Anterior margin of pronotum weakly convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, lateral and lateral anterior margins glabrous. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, on midline impunctate. Elytra short oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with minute setae in punctures, penultimate lateral interval with a few sparse short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.67. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium moderately convex and dull, with mixed finer and larger punctures, without smooth midline, with numerous short setae and a few longer in the larger punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin entirely smooth, distinctly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 2.83; dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical one at three quarter of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined, one third of its length longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 11 E – G. Habitus: Fig. 11 H.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB7FF8EFF3E70CF2E69DC23.taxon	diagnosis	Diagnosis. Microserica neosimplex Ahrens, Lukic & Liu, new species is in external appearance and shape of aedeagus most similar to Microserica allovarians Ahrens, Lukic & Liu, new species. Microserica neosimplex Ahrens, Lukic & Liu, new species differs by the right paramere having no mesolateral tooth.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB7FF8EFF3E70CF2E69DC23.taxon	etymology	Etymology. The species name is derived from the combined Greek prefix “ neo ” (new) and the species name “ simplex ”, with reference to the certain similarity to M. simplex.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB7FF8EFF3E70CF2E69DC23.taxon	description	Variation. Length: 4.3 – 5.2 mm, length of elytra: 2.9 – 3.5 mm, width: 2.8 – 3.1 mm. The species varies from almost entire dark (black) to completely yellow. Female: completely yellow, antennal club short, composed of three antennomeres, little shorter than remaining antennomeres combined; pygidium flat, eyes smaller than in male. Genbank accessions. Table 1.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB5FF88FF3E74F328B7DAE7.taxon	description	Figs. 12 A – I, 27	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB5FF88FF3E74F328B7DAE7.taxon	materials_examined	Type material examined. Holotype: Ô “ LAOS, Xieng Khouang prov., 19 ° 37 – 8 ’ N, 103 ° 20 ’ E, Phonsavan (30 km NE) Phou Sane Mt., 1420 m, 10 - 30. v. 2009, Z. Kraus leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň / 1095 Sericini Asia spec. ” (NHMB). Paratypes: Thailand: 1 Ô “ THAI 21. - 26. V. 1993 Namuang Pacholatko & Dembicky leg. / TS 102 / 19 Sericini Asia spec. ” (CP), 3 ÔÔ, 1 ♀ “ Phu Rua NP (900 m alt.), Loei P. NE Thai. 26 - 30. IV. 2006 Takakuwa, M. leg. ” (ZFMK), 1 Ô “ Khak Ngam (820 m) Phu Rua, Loei, NE Thai. 21 - 23. V. 2004 Takakuwa, M. leg. ” (ZFMK). Laos: 3 ÔÔ, 1 ♀ “ LAOS P. D. R., Xieng Khowang 14 - 20. May 1994 K. Miura leg. ” (ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB5FF88FF3E74F328B7DAE7.taxon	description	Description of holotype. Length: 5.2 mm, length of elytra: 3.4 mm, width: 3.1 mm. Body oval, yellowish brown, frons and partly pronotum with some iridescent-greenish shine, antennal club at apex brown, dorsal surface iridescent, partly dull, nearly glabrous. Labroclypeus little wider than long, widest at base, lateral margins convex and convergent to strongly rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, weakly convex, finely and coarsely, densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and bent medially. Frons with sparse, fine punctures, with a few longer setae beside eyes, otherwise with minute setae in punctures or glabrous. Smooth area in front of eyes 1.5 times as wide as long. Ocular canthus narrow and moderately long (1 / 3 of ocular width), sparsely finely punctate, with a terminal seta. Eyes small, ratio of diameter / interocular width: 0.49. Antenna with ten antennomeres; club composed of six antennomeres, straight, 1.5 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest shortly before base, lateral margins evenly convex and convergent to blunt, moderately produced anterior angles, posterior angles blunt. Anterior margin of pronotum straight, with robust and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, on midline impunctate. Elytra short oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with minute setae in punctures, odd intervals with a few sparse short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.81. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, with mixed finer and larger punctures, without smooth midline, with numerous short setae and a few longer in the larger punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin finely serrate at apex, distinctly widened in apical half, dorsal posterior margin serrate, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 2.79; dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical one at two thirds of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined, twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 12 A-C, E – H. Habitus: Fig. 12 D, I. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB5FF88FF3E74F328B7DAE7.taxon	diagnosis	Diagnosis. Microserica parasimplex Ahrens, Lukic & Liu, new species is in shape of aedeagus most similar to is in external appearance and shape of aedeagus most similar to Microserica allovarians Ahrens, Lukic & Liu, new species. Microserica parasimplex Ahrens, Lukic & Liu, new species differs by the lacking basidorsal process of the right paramere, the left paramere is dorsally enlarged and covers almost completely the right one.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB5FF88FF3E74F328B7DAE7.taxon	etymology	Etymology. The species name is derived from the combined Greek prefix “ para - ” (false) and the species name “ simplex ”, with reference to the certain similarity to Mic. simplex.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB5FF88FF3E74F328B7DAE7.taxon	description	Variation. Length: 4.7 – 5.2 mm, length of elytra: 3.0 – 3.3 mm, width: 3.0 – 3.1 mm. The species varies from almost entire dark (black) to completely yellow. Female: yellow, antennal club short, composed of three antennomeres, little shorter than remaining antennomeres combined; pygidium flat, eyes smaller than in male.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB2FF89FF3E717C2E49DE63.taxon	description	Figs. 13 A – D, 28	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB2FF89FF3E717C2E49DE63.taxon	materials_examined	Type material examined. Holotype Ô [Vietnam] “ Hoa-Binh Tonkin A. de Cooman B. M. 1934 - 95 ” (BMNH). Additional material examined. 1 Ô, 1 ♀ [Vietnam] “ Museum Paris Tonkin Reg. de Hoa-Binh A. de Cooman 1929 / Museum Paris ex. Coll. R. Oberthür 1952 / 1115 Asia Sericini spec. ” (MNHN).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB2FF89FF3E717C2E49DE63.taxon	description	Redescription. Length: 5.0 mm, length of elytra: 3.9 mm, width: 3.5 mm. Body oval, body yellowish brown, lateral intervals, frons, two large, pairs of spots on pronotum, and multiple dots on elytra brown, dorsal surface dull and nearly glabrous. Labroclypeus trapezoidal, wider than long, widest at base, lateral margins almost straight and convergent to moderately rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce an indistinct, blunt angle; surface shiny, medially weakly convex, finely and coarsely, densely punctate, with a few long, erect setae anteriorly in transversal row; frontoclypeal suture finely incised and bent medially. Frons dull, with dense, fine punctures, beside eyes with a few single short setae, otherwise glabrous. Smooth area in front of eyes twice as wide as long. Ocular canthus impunctate, narrow and short (1 / 4 of ocular width), with a terminal seta. Eyes small, ratio of diameter / interocular width: 0.56. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins weakly evenly convex and convergent to distinctly produced and sharp anterior angles, posterior angles strongly rounded. Anterior margin of pronotum convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron indistinctly carinate, carina not produced ventrally. Scutellum triangular, finely and densely punctate, on midline impunctate. Elytra short oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd lateral intervals with a few short setae; with two incomplete bands of spots on elytra, one anterior with spots on interval 2, 4, and 6, and one posterior behind the middle on intervals 2, 4 to 6; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose; apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, laterally with numerous long, yellow, and robust setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.76. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, with mixed finer and larger punctures, without smooth midline, with minute setae in punctures and numerous short setae along apical margin. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin weakly widened in apical half, smooth, dorsal posterior margin also smooth, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 2.75; dorsal margin longitudinally convex, with two groups of spines, basal group at first third, apical one at three quarter of metatibial length, basally almost glabrous; lateral face strongly longitudinally convex, finely and sparsely punctate, partly with minute setae in punctures; ventral margin carinate and finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined, almost twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 13 A – C. Habitus: Fig. 13 D. Variation. Female: antennal club short, distinctly shorter than the remaining antennomeres combined, composed of three antennomeres; pygidium flat.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB3FF8AFF3E71302D96DC6B.taxon	description	Figs. 13 E – H, 28	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB3FF8AFF3E71302D96DC6B.taxon	materials_examined	Type material examined. Holotype Ô [China] “ Huawangshanzhuang, Jinxiu, Guangxi, 19. V. 1999, 550 m, leg. Gao Mingyuan / LW- 633 ” (IZAS). Paratypes: 3 ÔÔ “ Fukien, S. China Kienyang: Liutun 11. VI. 42 T. C. Maa / 536 Asia Sericini spec. ” (BPBM, ZFMK), 1 ♀ “ China: prov. Fujian bor. Occ. Shaowu env. 25 km road ShaowuTaining 13. - 16.6. M. Nikodým lgt. 1991 ” (ZFMK), 1 ♀ [China] “ Kuatun (2300 m) 27, 40 n. Br. 117, 40 ö. L. J. Klapperich 23.6.1938 (Fukien) / ex coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC), 1 ♀ [China] “ Kuatun, Fukien China 15.6.46 (Tschung Sen.) / ex coll. V. Balthasar National Museum Prague, Czech Republic ” (NMPC).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB3FF8AFF3E71302D96DC6B.taxon	description	Description of holotype. Length: 5.6 mm, length of elytra: 4.1 mm, width: 3.8 mm. Body oval, body yellowish brown, lateral intervals, frons, two large, pairs of spots on pronotum, and multiple dots on elytra brown, dorsal surface dull and nearly glabrous. Labroclypeus trapezoidal, wider than long, widest at base, lateral margins almost straight and convergent to moderately rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce an indistinct, blunt angle; surface shiny, medially weakly convex, finely and coarsely, densely punctate, with a few long, erect setae anteriorly in transversal row; frontoclypeal suture finely incised and bent medially. Frons dull, with dense, fine punctures, beside eyes with a few single short setae, otherwise glabrous. Smooth area in front of eyes twice as wide as long. Ocular canthus impunctate, narrow and short (1 / 4 of ocular width), with a terminal seta. Eyes small, ratio of diameter / interocular width: 0.59. Antenna with ten antennomeres; club composed of four antennomeres, straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins weakly evenly convex and convergent to distinctly produced and sharp anterior angles, posterior angles strongly rounded. Anterior margin of pronotum convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron indistinctly carinate, carina not produced ventrally. Scutellum triangular, finely and densely punctate, on midline impunctate. Elytra short oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd lateral intervals with a few short setae; with two incomplete bands of spots on elytra, one anterior with spots on interval 2, 4, and 6, and one posterior behind the middle on intervals 2, 4 to 6; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose; apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with numerous long, yellow, and robust setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.63. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, with mixed finer and larger punctures, without smooth midline, with minute setae in punctures and numerous short setae along apical margin. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin weakly widened in apical half, smooth, dorsal posterior margin also smooth, with a few short setae basally. Metatibia lacking in holotype. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 13 E – G. Habitus: Fig. 13 H.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB3FF8AFF3E71302D96DC6B.taxon	diagnosis	Diagnosis. Microserica paravicula Ahrens, Lukic & Liu, new species is in external appearance and morphology of aedeagus very similar to Mic. avicula (Arrow, 1946). The new species differs by the shape of the parameres and aedeagus: the dorsal phallobasal apophysis is separated at base into two separate filiform branches, while in Mic. avicula it is split shortly before apex; the dorsal lobe of the right paramere is narrow and split into a distal and a median branch, while in Mic. avicula the dorsal lob is wide and long, directed distally and not further subdivided; the left paramere is in the new species longer and strongly bent ventrally while it is straight in Mic. avicula.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB3FF8AFF3E71302D96DC6B.taxon	etymology	Etymology. The species name is derived from the combined Greek prefix “ para - ” (false) and the species name “ avicula ”, with reference to the certain similarity to Mic. avicula.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB3FF8AFF3E71302D96DC6B.taxon	description	Variation. Length: 5.6 – 6.2 mm, length of elytra: 4.1 – 4.4 mm, width: 3.7 – 3.8 mm. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 2.86; dorsal margin longitudinally convex, with two groups of spines, basal group at first third, apical one at three quarter of metatibial length, basally almost glabrous; lateral face strongly longitudinally convex, finely and sparsely punctate, partly with minute setae in punctures; ventral margin carinate and finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined, almost twice as long as dorsal tibial spur. Female: antennal club short, distinctly shorter than the remaining antennomeres combined, composed of three antennomeres; pygidium flat.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB0FF84FF3E73332963DA73.taxon	description	Figs. 13 I – L, 28	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB0FF84FF3E73332963DA73.taxon	materials_examined	Type material examined. Holotype Ô [China] “ Luoluoxinzhaishan Mountains, Caiyanghe River, Simao, Yunnan, 21. V. 2000, 1500 m, Bu Wenjun leg. / LW- 374 ” (NKU). Paratypes: 1 Ô [China] “ Menghun, Xishuangbanna, Yunnan, 23. V. 1958, 1200 - 1400 m, leg. Meng Xuwu ” (IZAS), 1 ♀ [China] “ Meng’e, Xishuangbanna, Yunnan, 23. V. 1958, 1050 - 1080 m, leg. Zhang Yiran ” (IZAS).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB0FF84FF3E73332963DA73.taxon	description	Description of holotype. Length: 5.4 mm, length of elytra: 4.9 mm, width: 3.2 mm. Body oval, body yellowish brown, lateral intervals, frons, two large, pairs of spots on pronotum, and multiple dots on elytra brown, dorsal surface dull and nearly glabrous. Labroclypeus trapezoidal, wider than long, widest at base, lateral margins almost straight and convergent to moderately rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce an indistinct, blunt angle; surface shiny, medially weakly convex, finely and coarsely, densely punctate, with a few long, erect setae anteriorly in transversal row; frontoclypeal suture finely incised and bent medially. Frons dull, with dense, fine punctures, beside eyes with a few single short setae, otherwise glabrous. Smooth area in front of eyes twice as wide as long. Ocular canthus impunctate, narrow and short (1 / 4 of ocular width), without a terminal seta. Eyes moderately large, ratio of diameter / interocular width: 0.69. Antenna with ten antennomeres; club composed of four antennomeres, strongly reflexed, twice as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins weakly evenly convex and convergent to distinctly produced and sharp anterior angles, posterior angles strongly rounded. Anterior margin of pronotum convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron indistinctly carinate, carina not produced ventrally. Scutellum triangular, finely and densely punctate, on midline impunctate. Elytra short oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd lateral intervals with a few short setae; with two incomplete bands of spots on elytra, one anterior with spots on interval 2, 4, and 6, and one posterior behind the middle on intervals 2 to 6; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose; apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, laterally with numerous long, yellow, and robust setae. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.62. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium moderately convex and dull, with mixed finer and larger punctures, without smooth midline, with minute setae in punctures and numerous short setae along apical margin. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin weakly widened in apical half, smooth, dorsal posterior margin also smooth, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio width / length: 1 / 3.22; dorsal margin longitudinally convex, with two groups of spines, basal group at first third, apical one at three quarter of metatibial length, basally almost glabrous; lateral face strongly longitudinally convex, finely and sparsely punctate, partly with minute setae in punctures; ventral margin carinate and finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined, almost twice as long as dorsal tibial spur. Protibia short, bidentate; anterior tarsi missing in holotype. Aedeagus: Fig. 13 I – K. Habitus: Fig. 13 L.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB0FF84FF3E73332963DA73.taxon	diagnosis	Diagnosis. Microserica caiyangheana Ahrens, Lukic & Liu, new species is in external appearance and morphology of aedeagus very similar to Mic. avicula (Arrow, 1946) and Mic. paravicula Ahrens, Lukic & Liu, new species. Microserica caiyangheana Ahrens, Lukic & Liu, new species differs by much longer antennal club, the continuous posterior band of spot on the elytra and the shape of the aedeagus: the dorsal phallobasal apophysis is absent; the dorsal lobe of the right paramere is weakly sclerotized and short; the left paramere is strongly curved ventrally.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB0FF84FF3E73332963DA73.taxon	etymology	Etymology. The species name, caiyangheana, is derived from the type locality, Caiyanghe River (adjective in nominative singular case).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB0FF84FF3E73332963DA73.taxon	description	Variation. Length: 5.4 – 5.6 mm, length of elytra: 4.9 – 5.0 mm, width: 3.2 – 3.3 mm. Female: antennal club short, little shorter than remaining antennomeres combined, composed of three antennomeres; pygidium flat.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBEFF85FF3E750B297ED9EB.taxon	description	Figs. 14 A – D, 28	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBEFF85FF3E750B297ED9EB.taxon	materials_examined	Type material examined. Holotype: Ô “ Laos centr., Bolikhamsai prov. Ban Nape-Kaew Nua Pass, 18.4. - 1.5.1998, alt. 600 + 100 m, N 18 ° 22,3, E 105 ° 09,1 ‘ (GPS), E. Jendek & O. Šauša leg. / M. Štrba & R. Hergovits / 45 Sericini Asia spec. ” (CP). Paratype: Ô “ Laos CE, Boli Kham Xai prov. Ban Nape (8 km NE), 600 m, 18 ° 21 ‘ N, 105 ° 08 ’ E P. Pachlátko leg.: 1. - 18. v. 2001 ” (ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBEFF85FF3E750B297ED9EB.taxon	description	Description of holotype. Length: 5.9 mm, length of elytra: 4.1 mm, width: 3.6 mm. Body oval, body yellowish brown, lateral elytral intervals, frons, two large, pairs of spots on pronotum, and multiple dots on elytra brown, dorsal surface dull and nearly glabrous. Labroclypeus trapezoidal, wider than long, widest at base, lateral margins weakly convex and strongly convergent to moderately rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce an indistinct, blunt angle; surface shiny, medially convex, finely and coarsely, densely punctate, with a few long, erect setae anteriorly in transversal row; frontoclypeal suture finely incised and bent medially. Frons dull, with dense, fine punctures, beside eyes with a few single short setae, otherwise glabrous. Smooth area in front of eyes 1.3 times as wide as long. Ocular canthus impunctate, narrow and short (1 / 4 of ocular width), with a terminal seta. Eyes moderately large, ratio of diameter / interocular width: 0.61. Antenna with ten antennomeres; club composed of four antennomeres, weakly reflexed, 1.5 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins weakly evenly convex and convergent to distinctly produced and sharp anterior angles, posterior angles strongly rounded. Anterior margin of pronotum convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron indistinctly carinate, carina not produced ventrally. Scutellum triangular, finely and densely punctate, on midline impunctate. Elytra short oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd lateral intervals with a few short setae; with two incomplete bands of spots on elytra, one anterior with spots on interval 2, 4, and 6, and one posterior behind the middle on intervals 2 to 6; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose; apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with numerous long, yellow, and robust setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.91. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium moderately convex and dull, with mixed finer and larger punctures, without smooth midline, with minute setae in punctures and numerous short setae along apical margin. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin weakly widened in apical half, smooth, dorsal posterior margin also smooth, with a few short setae basally. Metatibia moderately long and narrow, widest at middle, ratio width / length: 1 / 3.2; dorsal margin longitudinally convex, with two groups of spines, basal group at first third, apical one at three quarter of metatibial length, basally almost glabrous; lateral face strongly longitudinally convex, finely and sparsely punctate, partly with minute setae in punctures; ventral margin carinate and finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined, almost twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 14 A – C. Habitus: Fig. 14 D. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBEFF85FF3E750B297ED9EB.taxon	diagnosis	Diagnosis. Microserica multipunctata Ahrens, Lukic & Liu, new species is in external appearance and morphology of aedeagus similar to Mic. avicula (Arrow, 1946). Microserica multipunctata Ahrens, Lukic & Liu, new species differs by slightly longer antennal club, the continuous posterior band of spot on the elytra and the shape of the aedeagus: the dorsal phallobasal apophysis is dorsoventrally flatted beside being also curved; the right paramere is more narrowed towards aped (lateral view); the left paramere is strongly sickle-like curved (dorsal view). From Microserica caiyangheana Ahrens, Lukic & Liu, new species, Microserica multipunctata Ahrens, Lukic & Liu, new species differs by the presence of a dorsal apical phallobasal apophysis and the shorter antennal club.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBEFF85FF3E750B297ED9EB.taxon	etymology	Etymology. The species name is derived from the combined Latin words, “ multi ” (multiple) and “ punctatus ” (with punctures), with reference to the multiple dots on elytra (adjective in nominative singular case).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBFFF86FF3E76B32911D857.taxon	description	Figs. 14 E – H, 28	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBFFF86FF3E76B32911D857.taxon	materials_examined	Type material examined. Holotype: Ô “ LAOS, 21 ° 09 ’ N 101 ° 19 ’ E Louangnamtha pr., Namtha -> Muang Sing, 5 - 31. v. 1997, 900 - 1200 m Vit Kubáň leg. / LS 38 ” (CP).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBFFF86FF3E76B32911D857.taxon	description	Description of holotype. Length: 6.4 mm, length of elytra: 4.1 mm, width: 3.4 mm. Body oval, body yellowish brown, lateral elytral intervals, frons, two large, pairs of spots on pronotum, and multiple dots on elytra brown, dorsal surface dull and nearly glabrous. Labroclypeus semicircular, little wider than long, widest at base, lateral margins convex and strongly convergent to convex anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, medially convex, finely and coarsely, densely punctate, with a few long, erect setae anteriorly in transversal row; frontoclypeal suture finely incised and curved. Frons dull, with dense, fine punctures, beside eyes with a few single short setae, otherwise glabrous. Smooth area in front of eyes 1.5 times as wide as long. Ocular canthus impunctate, narrow and short (1 / 4 of ocular width), with a terminal seta. Eyes small, ratio of diameter / interocular width: 0.58. Antenna with ten antennomeres; club composed of four antennomeres, strongly reflexed, 1.7 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins weakly evenly convex and convergent to distinctly produced and sharp anterior angles, posterior angles strongly rounded. Anterior margin of pronotum convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron not visibly carinate. Scutellum triangular, finely and densely punctate, on midline impunctate. Elytra short oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd lateral intervals with a few short setae; with two incomplete bands of spots on elytra, one anterior with spots on interval 2, 4, and 6, and one posterior behind the middle on intervals 2 to 6; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose; apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with numerous long, yellow, and robust setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.63. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium moderately convex and dull, with mixed finer and larger punctures, without smooth midline, with minute setae in punctures and numerous short setae along apical margin. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin weakly widened in apical half, smooth, dorsal posterior margin also smooth, with a few short setae basally. Metatibia moderately long and narrow, widest at middle, ratio width / length: 1 / 3.16; dorsal margin longitudinally convex, with two groups of spines, basal group at first third, apical one at three quarter of metatibial length, basally almost glabrous; lateral face strongly longitudinally convex, finely and sparsely punctate, partly with minute setae in punctures; ventral margin carinate and finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined, almost twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 14 E – G. Habitus: Fig. 14 H. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBFFF86FF3E76B32911D857.taxon	diagnosis	Diagnosis. Microserica hieroglyphica Ahrens, Lukic & Liu, new species is in external appearance and morphology of aedeagus similar to Microserica caiyangheana Ahrens, Lukic & Liu, new species Both species lack the dorsal apical phallobasal apophysis. Microserica hieroglyphica Ahrens, Lukic & Liu, new species differs by slightly shorter antennal club and the shape of the aedeagus: the left paramere is shorter and stouter, in both species it is strongly curved; the right paramere is mesoventrally more extended (ventrodistal view).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBFFF86FF3E76B32911D857.taxon	etymology	Etymology. The species name is derived from the Latin word, “ hieroglyphicus ” (hieroglyphic), with reference to the pattern of multiple dots on elytra resembling hieroglyphic signs (adjective in nominative singular case).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBCFF87FF3E77E428A7D8A7.taxon	description	Figs. 14 I – L, 28	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBCFF87FF3E77E428A7D8A7.taxon	materials_examined	Type material examined. Holotype: Ô “ N-Vietnam Cao Bang Pr. vic. Vin Den Nui pia Oac Nature res. 10. - 15. V. 2014 N 22 ° 33.53 ’; E 105 ° 52.53 ’ 900 - 1300 m, leg. A. Weigel / collection Naturkundemuseum Erfurt / 1114 Sericini Asia spec. ” (NME). Paratypes: 4 ÔÔ “ N-Vietnam Cao Bang Pr. vic. Vin Den Nui pia Oac Nature res. 10. - 15. V. 2014 N 22 ° 33.53 ’; E 105 ° 52.53 ’ 900 - 1300 m, leg. A. Weigel / collection Naturkundemuseum Erfurt / 1114 Sericini Asia spec. ” (NME, ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBCFF87FF3E77E428A7D8A7.taxon	description	Description of holotype. Length: 6.9 mm, length of elytra: 5.5 mm, width: 4.1 mm. Body oblong-oval, body brown, antenna and elytral intervals yellowish brown, elytral striae, scutellum, frons, and pronotum darker, with strong greenish shine, dorsal surface dull but partly iridescent, except large single white, scale-like setae on odd intervals glabrous. Labroclypeus wider than long, widest at base, lateral margins convex and strongly convergent to convex anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, medially convex, finely and coarsely, densely punctate, with a few long, erect setae anteriorly in transversal row; frontoclypeal suture finely incised and curved. Frons dull, with dense, fine punctures, beside eyes with a few single short setae, otherwise glabrous. Smooth area in front of eyes 1.5 times as wide as long. Ocular canthus impunctate, narrow and moderately long (1 / 3 of ocular width), with a terminal seta. Eyes moderately large, ratio of diameter / interocular width: 0.65. Antenna with ten antennomeres; club composed of four antennomeres, strongly reflexed, 1.8 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide, widest shortly before base, lateral margins evenly convex and convergent to distinctly produced and sharp anterior angles, posterior angles strongly rounded. Anterior margin of pronotum convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron not visibly carinate. Scutellum triangular, finely and densely punctate, on basal midline impunctate. Elytra rather elongate, widest at posterior third, striae distinctly and broadly impressed, finely and densely punctate, intervals strongly convex, dark, with fine and sparse punctures concentrated only along striae, with very minute setae in punctures, odd lateral intervals with large single white, scale-like setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose; apical border broadly membraneous, with a broad fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum covered with fine, short, or very minute setae; metacoxa glabrous, with numerous long, white, and robust setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.57. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium moderately convex and dull, with mixed finer and larger punctures, without smooth midline, with short and long setae. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta present; setae of anterior and posterior row long, dense, and fine; ventral posterior margin weakly widened in apical half, smooth, dorsal posterior margin also smooth, with a few short setae basally. Metatibia moderately long and narrow, widest at middle, ratio width / length: 1 / 3.6; dorsal margin longitudinally convex, with two groups of spines, basal group at first third, apical one at three quarter of metatibial length, basally with no other robust setae; lateral face strongly longitudinally convex, densely and coarsely punctate, with short or long setae in punctures; ventral margin carinate and finely serrate, with three robust setae of which the apical one is more distant; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, dense setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous ventrally, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere little shorter than following two tarsomeres combined, little more than twice as long as dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 14 I – K. Habitus: Fig. 14 L. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBCFF87FF3E77E428A7D8A7.taxon	diagnosis	Diagnosis. Microserica vinden Ahrens, Lukic & Liu, new species is in external appearance and morphology of aedeagus similar to Microserica costisquamosa (Ahrens, Fabrizi & Liu, 2019) new combination The new species differs by the much longer left paramere.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBCFF87FF3E77E428A7D8A7.taxon	etymology	Etymology. The species name is named after its type locality, Vin Den (noun in apposition).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBDFF87FF3E707F2D9EDE1D.taxon	description	Fig. 28	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBDFF87FF3E707F2D9EDE1D.taxon	materials_examined	Material examined. 3 ÔÔ, 3 ♀♀ “ China, S Guizhou Maolan, 700 - 800 m, 25 ° 19 ’ N 107 ° 52 ’ E Jatua leg., 5. VI. 2012 ” (ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBDFF87FF3E707F2D9EDE1D.taxon	discussion	Remarks. Based on the genital morphology, Microserica costisquamosa (Ahrens, Fabrizi & Liu, 2019) new combination appears to be closely related to the other species of the M. avicula complex, whose systematic placement is yet, nevertheless, uncertain.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBDFF80FF3E71A12E6ADE8B.taxon	description	Figs. 15 A – D, 28	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBDFF80FF3E71A12E6ADE8B.taxon	materials_examined	Type material examined. Holotype Ô “ Yunnan, Lvchun shuiku, 2012 - V- 13 / LW- 1264 ” (ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBDFF80FF3E71A12E6ADE8B.taxon	description	Description of holotype. Length: 6.0 mm, length of elytra: 4.4 mm, width: 3.8 mm. Body oval, body including legs and antenna yellow, two spots on pronotum and multiple symmetric smaller spots on elytra, and abdomen including pygidium brown, frons posteriorly slightly darker; dorsal surface dull, brown areas with some greenish shine, surface nearly glabrous. Labroclypeus trapezoidal, distinctly wider than long, widest at base, lateral margins straight and convergent to bluntly rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface weakly shiny, medially convexly elevated, coarsely, densely punctate, with a few long, erect setae anteriorly in a transversal line; frontoclypeal suture finely incised and curved medially. Frons dull, with sparse, fine punctures and a few setae beside eyes, otherwise glabrous. Smooth area in front of eyes narrow, 1.5 times as wide as long. Ocular canthus wide, short (1 / 4 of ocular width), sparsely finely punctate, with a terminal seta. Eyes small, ratio of diameter / interocular width: 0.5. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.1 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins evenly moderately convex and convergent to sharp and distinctly produced anterior angles; posterior angles blunt. Anterior margin of pronotum weakly convex, with fine, complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, evenly finely and densely punctate, impunctate along midline, glabrous. Elytra elongate, widest at posterior third, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and dense punctures only along striae, with minute setae in punctures, elytra with a single band of transversal and symmetrical dark dots at posterior third on interval 2 - 6, odd intervals with a few short setae; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.58. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium weakly convex and dull, with sparse, fine punctures, without smooth midline, with a few short adpressed setae on apical half, otherwise with minute setae in punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin distinctly not serrate but moderately widened, dorsal posterior margin smooth, with short setae. Metatibia moderately long and wide, widest at apical third, ratio width / length: 1 / 3.4; dorsal margin longitudinally convex, with two groups of spines, basal group at one third, apical one at three quarters of metatibial length, in basal third without robust setae; lateral face longitudinally convex, finely and sparsely punctate, glabrous; ventral margin finely serrate, with four robust setae of which the distal one is more distant; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally impunctate; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined and little less than twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 15 A – C. Habitus: Fig. 15 D. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBDFF80FF3E71A12E6ADE8B.taxon	diagnosis	Diagnosis. The species differs from all other Microserica species by the lacking carina of the hypomeron, as well as the non carinate dorsal margin of the metatibia; of course, also the male genitalia are different from all other species. The systematic position of this species is based on Eberle et al. (2017) close to the species of the Microserica quadrimaculata group.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBDFF80FF3E71A12E6ADE8B.taxon	etymology	Etymology. The name of the new species is derived from the combined Larin prefix “ fere- ” (almost) and the adjective, “ sticticus ” (line, file), with reference to the imperfect stripe-dotted color pattern of the elytra. Genbank accessions. Table 1.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBAFF81FF3E725329F5DD1B.taxon	description	Figs. 16 A – D, 28	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBAFF81FF3E725329F5DD1B.taxon	materials_examined	Type material examined. Holotype: Ô “ N Vietnam Cuc Phuong Nat. Park 21. - 22. v. 1996 Pacholátko & Dembický leg. / VS 98 / 49 Sericini Asia spec. ” (CP).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBAFF81FF3E725329F5DD1B.taxon	description	Description of holotype. Length: 5.9 mm, length of elytra: 3.9 mm, width: 3.4 mm. Body oval, body yellowish brown, lateral elytral intervals, center of frons, two large, pairs of spots on pronotum, and multiple dots on elytra brown, dorsal surface dull and nearly glabrous. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins convex and strongly convergent to convex anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, medially convex, finely and coarsely, densely punctate, with a few long, erect setae anteriorly in transversal row; frontoclypeal suture finely incised and curved. Frons dull, with dense, fine punctures, beside eyes with a few single short setae, otherwise glabrous. Smooth area in front of eyes as wide as long. Ocular canthus impunctate, narrow and short (1 / 4 of ocular width), with a terminal seta. Eyes small, ratio of diameter / interocular width: 0.58. Antenna with ten antennomeres; club composed of four antennomeres, strongly reflexed, 1.5 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins weakly evenly convex and convergent to distinctly produced and sharp anterior angles, posterior angles strongly rounded. Anterior margin of pronotum convex, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron not visibly carinate. Scutellum triangular, finely and densely punctate, on midline impunctate. Elytra short oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd lateral intervals with a few short setae; with two incomplete bands of spots on elytra, one anterior with spots on interval 2, 4, and 6, and one posterior behind the middle on intervals 2 to 6; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose; apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with numerous long, yellow, and robust setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.62. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium moderately convex and dull, with mixed finer and larger punctures, without smooth midline, with minute setae in punctures and numerous short setae along apical margin. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin weakly widened in apical half, smooth, dorsal posterior margin also smooth, with a few short setae basally. Metatibia moderately long and narrow, widest at middle, ratio width / length: 1 / 3.2; dorsal margin longitudinally convex, with two groups of spines, basal group at first third, apical one at three quarter of metatibial length, basally almost glabrous; lateral face strongly longitudinally convex, finely and sparsely punctate, partly with minute setae in punctures; ventral margin carinate and finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined, twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 16 A – C. Habitus: Fig. 16 D. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBAFF81FF3E725329F5DD1B.taxon	diagnosis	Diagnosis. Microserica cucphuong Ahrens, Lukic & Liu, new species is in external appearance and morphology of aedeagus similar to Microserica hieroglyphica Ahrens, Lukic & Liu, new species and Mic. caiyangheana Ahrens, Lukic & Liu, new species All species lack the dorsal apical phallobasal apophysis. Microserica cucphuong Ahrens, Lukic & Liu, new species differs from both latter species by the left paramere being curved upwards (lateral view); the basal process of the right paramere is extended more basally than mesally (ventrodistal view).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFBAFF81FF3E725329F5DD1B.taxon	etymology	Etymology. The species name is derived from the type locality, Cuc Phuong Nat. Park (noun in apposition).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB8FF83FF3E76192E8BD87B.taxon	description	Figs. 17 A – D, 29	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB8FF83FF3E76192E8BD87B.taxon	materials_examined	Type material examined. Holotype: Ô “ Yunnan 1800 - 2000 m 25.04 N 101.55 E Vipinglang 17 - 20.6. Vit Kuban leg. 1994 / 830 Sericini: Asia spec. ” (CP). Paratypes: 2 ÔÔ “ Yunnan 1800 - 2000 m 25.04 N 101.55 E Vipinglang 17 - 20.6. Vit Kuban leg. 1994 / CS 39 / 23 Asia Sericini spec. ” (CP, ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB8FF83FF3E76192E8BD87B.taxon	description	Description of holotype. Length: 5.2 mm, length of elytra: 3.1 mm, width: 3.2 mm. Body oval, black, antennal funiculus, anterior and middle legs brown, dorsal surface simply shiny, ventral surface dull, dorsal surface glabrous. Labroclypeus little wider than long, widest at base, lateral margins convex and convergent to moderately rounded anterior angles; anterior margin distinctly and widely emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, strongly convex, coarsely and densely punctate, with a few long, erect setae anteriorly; frontoclypeal indistinctly incised, curved. Frons with moderately dense, coarse punctures, with a few long setae beside eyes and at middle behind frontoclypeal suture, otherwise glabrous. Smooth area in front of eyes strongly convex, as wide as long. Ocular canthus wide and short (1 / 3 of ocular width), densely coarsely punctate, without terminal seta. Eyes very small, ratio of diameter / interocular width: 0.4. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.1 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins in basal half straight and subparallel, anteriorly evenly convex and convergent to sharp, distinctly produced anterior angles, posterior angles blunt. Anterior margin of pronotum straight, with robust and complete marginal line; basal marginal line absent; surface coarsely and densely punctate, with very minute setae in punctures, anterior and lateral anterior margins sparsely setose. Hypomeron carinate, not produced ventrally. Scutellum triangular and dull, finely and densely punctate, on mesal base impunctate. Elytra short oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with minute setae in punctures, odd intervals with a few sparse short setae; epipleural edge fine, ending convex external apical angle of elytra, epipleura densely setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.55. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium shiny and strongly convex, with mixed finer and larger punctures, without smooth midline, with numerous longer setae along apical margin, otherwise with minute setae in punctures. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; ventral posterior margin distinctly widened in apical half and entirely smooth, dorsal posterior margin finely serrate, with a few short setae basally. Metatibia short and wide, widest at apex, ratio width / length: 1 / 2.78; dorsal margin weakly carinate, with two groups of spines, basal group at anterior quarter, apical one at two thirds of metatibial length, in basal third with a few fine, single setae; lateral face longitudinally convex, finely and densely punctate in basal half, in apical half punctation sparse, almost entirely smooth, with minute setae in punctures; ventral margin finely serrate, with two robust setae in apical half; medial face densely finely punctate, apex interiorly near tarsal articulation concavely emarginate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere little shorter than following two tarsomeres combined, almost twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 17 A – C. Habitus: Fig. 17 D. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB8FF83FF3E76192E8BD87B.taxon	diagnosis	Diagnosis. Microserica vipinglangensis Ahrens, Lukic & Liu, new species is in shape of aedeagus most similar to Microserica soppongensis Ahrens, 2005. Microserica vipinglangensis Ahrens, Lukic & Liu, new species differs by the shiny black dorsal surface, the short antennal club composed only of four antennomeres, and shape of the parameres: the right paramere is strongly reflexed (dorsal view), while being almost straight in Mic. soppongensis; the left paramere is shorter, being distinctly shorter than the right one, and strongly curved (dorsal view); in Mic. soppongensis the latter is subequal to the right one and straight over its entire length except at apex.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB8FF83FF3E76192E8BD87B.taxon	etymology	Etymology. The species name is derived from the name of its type locality, Vipinglang (adjective nominative singular case).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB8FF83FF3E76192E8BD87B.taxon	description	Variation. Paratypes of the same size as holotype.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB9FF9CFF3E77032E4ADA73.taxon	description	Fig. 29	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB9FF9CFF3E77032E4ADA73.taxon	materials_examined	Material examined. See Ahrens (2005 a: 32). 1 Ô “ Mt. Phu Phan, Xan Neua, Houapan, N. Laos, 19 - 21 V. 2005 Takakuwa, M. leg. ” (ZFMK), 155 ex. “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ’ N, 104 ° 01 ’ E; 14. iv. - 15. v. 2012; 1300 - 1900 m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012 / 13 ” (ZFMK), 387 ex. “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) - 20 ° 12 ’ N, 104 ° 01 ’ E; 11. iv. - 15. v. 2012; 1300 - 1900 m; leg. C. Holzschuh Ankauf ZFMK Bonn 2012 ” (ZFMK), 23 ex. “ NE-Laos: Hua Phan prov., Ban Saleui, Phou Pan (Mt.) ~ 20 ° 12 ’ N, 104 ° 01 ’ E; 1300 - 1900 m; 01. - 31. V. 2011; leg. C. Holzschuh Ankauf ZFMK Bonn 2011 ” (ZFMK), 78 ex. “ Laos-NE, Houa Phan prov., 20 ° 13 ’ 09 - 19 ’’ N 103 ° 59 ’ 54 ’’ - 104 ° 00 ’ 03 ’’ E, 1480 - 1510 m Phou Pane Mt., 22. IV. - 14. V. 2008 Vit Kuban leg. ” (NMPC), 13 ex. “ Laos-NE Hua Phan prov., 20 ° 12 ’ N, 104 ° 01 ’ E, Phu Phan Mt., 1500 - 1900 m, 17.5. - 3.6.2007, leg. Vit Kuban ” (ZFMK), 14 ex. “ Laos-NE Hua Phan prov., 20 ° 12 ’ N, 104 ° 01 ’ E, Phu Phan Mt., 1500 - 1900 m, 17.5. - 3.6.2007, leg. C. Holzschuh ” (ZFMK), 10 ex. “ LAOS, Xieng Khouang prov., 19 ° 37 – 8 ’ N, 103 ° 20 ’ E, Phonsavan (30 km NE) Phou Sane Mt., 1420 m, 10 - 30. v. 2009, Z. Kraus leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 2 ex. “ LAOS, Xieng Khouang prov., 19 ° 37 – 8 ’ N, 103 ° 20 ’ E, Phonsavan (30 km NE) Phou Sane Mt., 1420 m, 10 - 30. v. 2009, M. Brancucci leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 4 ex. “ LAOS, Hua Phan prov., 20 ° 13 ’ N, 104 ° 00 ’ E, Phou Pane Mt., 1 - 16. vi. 2009, 1350 - 1500 m, M. Brancucci leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 1 ex. “ LAOS, Houa Phan prov., 20 ° 12 - 13 ’ N, 103 ° 59.5 ’ - 104 ° 01 ’ E, Ban Saluei-> Phou Pane Mt., 1340 - 1870 m, 10. v. - 16. vi. 2009, M. Brancucci & local coll. leg. / NHMB Basel, NMPC Prague, Laos 2009, Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň ” (NHMB), 14 ex. “ LAOS-NE, Houa Phan prov., Ban Saluei-> Phou Pane Mt., 20 ° 12 - 13.5 ’ N 103 ° 59.5 ’ - 104 ° 01 ’ E, 1340 - 1870 m, 15. iv. - 15. v. 2008, Lao collector leg. ” (NHMB), 2 ex. “ LAO-NE, Hua Phan prov., 20 ° 12 ’ N 104 ° 01 ’ E, Phu Phan Mt., 1500 - 1900 m, 17. v. - 3. vi. 2007, M. Brancucci leg. / NHMB Basel expedition to Laos, 2007 ” (NHMB), 2 ex. “ LAOS-NE; Hua Phan prov., Ban Saleui, Phou Pan (Mt.), 20 ° 15 ’ N, 104 ° 02 ’ E; 1500 - 2000 m; J. Bezděk leg.; 26. iv. - 11. v. 2001 ” (CP), 1 ex. “ NE Laos Houaphan Prov. Phu Pan (Mt.) 15 - 26. V. 2004 T. Mizusawa leg. ” (ZFMK), 1 ex. (♀) “ Laos, Attapeau prov.; Annam Highlands Mts Dong Amphan; NBCA, ca. 1160 m Nong Fa (crater lake) env., 15 ° 05.9 ’ N, 107 ° 25.6 ’ E St. Jakl lgt, 30.4. - 6.5.2010 ” (ZFMK), 1 Ô “ X-DA 4854 / X-DA 4854 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 3. - 5. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica panzona ” (ZFMK), 1 Ô “ X-DA 4940 / X- DA 4940 LAOS Hua Phua prov., Ban Saluei, Phou Pan (Mt.) 20 ° 12 ’ N, 104 ° 01 ’ E 2. iv. 2014 leg. C. Holzschuh – ZFMK Ankauf 2014 Microserica panzona ” (ZFMK). Aedeagus. See Ahrens (2005 a: p. 33; Fig. 1 A – C).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFB9FF9CFF3E77032E4ADA73.taxon	distribution	Distribution. Northeastern Laos.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA6FF9CFF3E750B2C90DAC3.taxon	description	Fig. 29	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA6FF9CFF3E75BF2E69D9C7.taxon	materials_examined	Material examined. See Ahrens (2005 a: p. 34). Aedeagus. See Ahrens (2005 a: p. 33; Fig. 1 D – F).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA6FF9CFF3E75BF2E69D9C7.taxon	distribution	Distribution. Thailand. Genbank accessions. Table 1.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA6FF9DFF3E769F2D4ED87B.taxon	description	Figs. 17 E – H, 29	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA6FF9DFF3E769F2D4ED87B.taxon	materials_examined	Type material examined. Holotype Ô “ Banlao, Cangyuan, Yunnan, 17. V. 1980, 1100 m, leg. Gao Ping / LW- 588 ” (IZAS).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA6FF9DFF3E769F2D4ED87B.taxon	description	Description of holotype. Length: 9.0 mm, length of elytra: 6.4 mm, width: 5.2 mm. Body oval, reddish brown, antenna yellow, dorsal and ventral surface dull, labroclypeus and anterior frons shiny; dorsal surface glabrous. Labroclypeus trapezoidal, distinctly wider than long, widest at base, lateral margins convex and convergent to moderately rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, flat, coarsely and densely punctate, entirely glabrous; frontoclypeal indistinctly incised, bent medially. Frons in anterior third shiny, with irregularly, moderately dense, fine punctures, being in dull part more superficial; with a few single setae beside eyes, otherwise glabrous. Smooth area in front of eyes flat, twice as wide as long. Ocular canthus wide and short (1 / 3 of ocular width), its external margin strongly convex rather than straight, densely coarsely punctate, with a terminal seta. Eyes small, ratio of diameter / interocular width: 0.52. Antenna with ten antennomeres; club composed of four antennomeres, straight, 0.9 times as long as remaining antennomeres combined, the first lamella of club only half as long as club. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins weakly convex but strongly convergent to sharp, distinctly produced anterior angles, posterior angles blunt, well-rounded at tip. Anterior margin of pronotum convex, with robust and complete marginal line; basal marginal line absent; surface finely and densely punctate, with very minute setae in punctures, anterior margin glabrous, lateral margin sparsely setose. Hypomeron strongly carinate, strongly produced ventrally, deeply transversely grooved. Scutellum large and triangular, dull, finely and densely punctate. Elytra oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and moderately dense punctures concentrated along striae, glabrous; epipleural edge fine, ending convex external apical angle of elytra, epipleura densely setose, apical border membraneous, with a fine fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum on disc with long erect setae, otherwise sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.27. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium weakly convex and dull, with fine dense punctures, without smooth midline, with numerous longer setae along apical margin, otherwise glabrous or with minute setae in punctures. Legs moderately wide and long; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures reduced; ventral posterior margin distinctly widened in apical half and entirely smooth, dorsal posterior margin also smooth, with a few short setae basally. Metatibia short and wide, widest at middle but only weakly narrowed towards apex, ratio width / length: 1 / 2.58; dorsal margin carinate, with two groups of spines, basal group at anterior quarter, apical one at two thirds of metatibial length, in basal third with a few fine, single setae; lateral face longitudinally convex, finely and very sparsely punctate, glabrous; ventral margin finely serrate, with four equidistant setae; medial face smooth and glabrous, apex interiorly near tarsal articulation bluntly truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally, but without parallel subventral smooth carina; first metatarsomere little shorter than following two tarsomeres combined, almost twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 17 E – G. Habitus: Fig. 17 H. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA6FF9DFF3E769F2D4ED87B.taxon	diagnosis	Diagnosis. Microserica banlaoana Ahrens, Lukic & Liu, new species is external appearance and in shape of aedeagus most similar to Microserica lampungensis Ahrens, 2004 from Southern Sumatra. Microserica banlaoana Ahrens, Lukic & Liu, new species differs by the less stout and bent phallobase (lateral view), and the left paramere being at base mesally not widened towards a basal lobe.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA6FF9DFF3E769F2D4ED87B.taxon	etymology	Etymology. The species name is derived from the name of its type locality, Banlao (adjective nominative singular case).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA7FF9EFF3E77032F53DF7F.taxon	description	Fig. 18 A – D	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA7FF9EFF3E77032F53DF7F.taxon	materials_examined	Type material examined. Holotype: Ô “ Haut Mekong Vieng Vai 23. V. 1918 R. V. de Salvaza / Brit Mus. 1921 - 89 / 1111 Sericini Asia spec. ” (BMNH). Paratypes: 4 ÔÔ, 2 ♀♀ “ Haut Mekong Vieng Vai 23. V. 1918 R. V. de Salvaza ” (BMNH, ZFMK), 7 ÔÔ, 4 ♀♀ “ Haut Mekong Vieng Vien 3. VI. 1918 R. V. de Salvaza ” (BMNH, ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA7FF9EFF3E77032F53DF7F.taxon	description	Description of holotype. Length: 4.8 mm, length of elytra: 4.4 mm, width: 3.2 mm. Body oval, dark brown, head and pronotum with some greenish shine, antenna yellow, dorsal and ventral surface dull but with iridescent shine; dorsal surface glabrous. Labroclypeus trapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and convergent to strongly rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce an indistinct blunt angle; surface shiny, weakly convex medially, coarsely and densely punctate, with a few short erect setae; frontoclypeal fine, weakly curved. Frons dull, with dense, fine punctures; with a few single setae beside eyes, otherwise glabrous. Smooth area in front of eyes flat, 1.5 times as wide as long. Ocular canthus wide and short (1 / 3 of ocular width), its external margin straight, impunctate, with a terminal seta. Eyes small, ratio of diameter / interocular width: 0.57. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.1 times as long as remaining antennomeres combined, all lamella of club equal in length. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins moderately convex and convergent to sharp, distinctly produced anterior angles, posterior angles blunt. Anterior margin of pronotum weakly convex, with fine, complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, lateral anterior margin and lateral margin sparsely setose. Hypomeron weakly carinate, not produced ventrally, not grooved. Scutellum small and triangular, finely and densely punctate, smooth along midline. Elytra oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and dense punctures only concentrated along striae, along middle almost impunctate, with minute setae in punctures, odd intervals with a few single, short white, adpressed setae; epipleural edge fine, ending convex external apical angle of elytra, epipleura densely setose; apical border membraneous, with a fine fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum on disc with long erect setae, otherwise sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.83. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a large, robust seta. Pygidium weakly convex and dull, with fine, dense punctures, without smooth midline, with a few long setae along apical margin, otherwise glabrous. Legs moderately wide and long; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures reduced; ventral posterior margin distinctly widened in apical half and entirely smooth, dorsal posterior margin also smooth, with a few short setae basally. Metatibia short and wide, widest at middle but only weakly narrowed towards apex, ratio width / length: 1 / 3.1; dorsal margin carinate, with two groups of spines, basal group at anterior third, apical one at three quarters of metatibial length, in basal third with a few fine, single setae; lateral face longitudinally convex, finely and sparsely punctate on dorsal and basal portion, glabrous; ventral margin finely serrate, with four equidistant setae; medial face smooth and glabrous, apex interiorly near tarsal articulation bluntly truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally, but without parallel subventral smooth carina; first metatarsomere much shorter than following two tarsomeres combined, little longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 18 A – C Habitus: Fig. 18 D.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA7FF9EFF3E77032F53DF7F.taxon	diagnosis	Diagnosis. Microserica viengvai Ahrens, Lukic & Liu, new species has not similar species on continual Asian mainland; its external appearance is similar to Microserica martini Ahrens, 2001 from Sumatra, differs, however, by the uniform dark and smaller body as well as by the shape of aedeagus: the phallobase is more compact and curved (lateral view), the parameres are inserted more asymmetrically; the left paramere is shorter and simple, and the right one is strongly widened and dorsoventrally flattened.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA7FF9EFF3E77032F53DF7F.taxon	etymology	Etymology. The species name is derived from the name of its type locality, Vieng Vai (noun in apposition).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA7FF9EFF3E77032F53DF7F.taxon	description	Variation. Length: 4.8 – 5.5 mm, length of elytra: 3.6 – 4.4 mm, width: 3.2 – 3.3 mm. Female: eyes slightly smaller, ratio of diameter / interocular width: 0.55; pygidium flat, antennal club short, composed of three antennomeres, distinctly shorter than remaining antennomeres combined.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA4FF9FFF3E70072900DF37.taxon	description	Figs 18 E-H, 29	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA4FF9FFF3E70072900DF37.taxon	materials_examined	Type material examined. Holotype: Ô “ X-DA 5510 / X-DA 5510 Thailand Phetchabun Nam Nao NP Hill evergreen forest 16 ° 44.371 ’ N, 101 ° 34.549 ’ E 834 m leg. T 2429 Leng Jantteab Malaise trap 26. v. - 2. vi. 2007 Microserica sp. TigerThai 09 / 1171 Sericini Asia spec. ” (QSBG).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA4FF9FFF3E70072900DF37.taxon	description	Description of holotype. Length: 5.3 mm, length of elytra: 3.5 mm, width: 3.2 mm. Body oval, dark brown, head and pronotum with some greenish shine, antenna yellow, dorsal and ventral surface dull but with iridescent shine; dorsal surface glabrous. Labroclypeus trapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and convergent to strongly rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce an indistinct blunt angle; surface shiny, weakly convex medially, coarsely and densely punctate, with a few short erect setae; frontoclypeal fine, weakly curved. Frons dull, with dense, fine punctures; with a few single setae beside eyes, otherwise glabrous. Smooth area in front of eyes flat, 1.5 times as wide as long. Ocular canthus wide and short (1 / 3 of ocular width), its external margin straight, impunctate, with a terminal seta. Eyes small, ratio of diameter / interocular width: 0.53. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.2 times as long as remaining antennomeres combined, all lamella of club equal in length. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins moderately convex and convergent to sharp, distinctly produced anterior angles, posterior angles blunt. Anterior margin of pronotum weakly convex, with fine, complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, lateral anterior margin and lateral margin sparsely setose. Hypomeron weakly carinate, not produced ventrally, not grooved. Scutellum small and triangular, finely and densely punctate, smooth along midline. Elytra oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and dense punctures only concentrated along striae, along middle almost impunctate, with minute setae in punctures, odd intervals with a few single, short white, adpressed setae; epipleural edge fine, ending convex external apical angle of elytra, epipleura densely setose; apical border membraneous, with a fine fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum on disc with long erect setae, otherwise sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.88. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a large, robust seta. Pygidium weakly convex and dull, with fine, dense punctures, without smooth midline, with a few long setae along apical margin, otherwise glabrous. Legs moderately wide and long; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures reduced; ventral posterior margin distinctly widened in apical half and entirely smooth, dorsal posterior margin also smooth, with a few short setae basally. Metatibia short and wide, widest at middle but only weakly narrowed towards apex, ratio width / length: 1 / 3.3; dorsal margin carinate, with two groups of spines, basal group at anterior third, apical one at three quarters of metatibial length, in basal third with a few fine, single setae; lateral face longitudinally convex, finely and sparsely punctate on dorsal and basal portion, glabrous; ventral margin finely serrate, with four equidistant setae; medial face smooth and glabrous, apex interiorly near tarsal articulation bluntly truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally, but without parallel subventral smooth carina; first metatarsomere little shorter than following two tarsomeres combined, little longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 18 E – G Habitus: Fig. 18 H. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA4FF9FFF3E70072900DF37.taxon	diagnosis	Diagnosis. Microserica namnao Ahrens, Lukic & Liu, new species is very similar to Mic. viengvai Ahrens, Lukic & Liu, new species. Microserica namnao Ahrens, Lukic & Liu, new species differs by the more triangular left paramere (lateral view), and the apically less narrowed right one (lateral view) which is broudly rounded at apex.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA4FF9FFF3E70072900DF37.taxon	etymology	Etymology. The species name is derived from the name of its type locality, Nam Nao (noun in apposition).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA5FF98FF3E70CF2813DD67.taxon	description	Figs 18 I-L, 29	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA5FF98FF3E70CF2813DD67.taxon	materials_examined	Type material examined. Holotype: Ô “ X-DA 5202 / X- DA 5202 Thailand Chaiyaphum Pha Hin Ngam NP Dry evergreen forest near stream 15 ° 40.569 ’ N, 101 ° 26.705 ’ E 461 m leg. T 2476 Katae Sa-nog & Buakaw Adnafai Malaise traps 7. - 13. vi. 2007 Microserica sp. TigerThai 09 / 1172 Sericini Asia spec. ” (QSBG). Paratypes: 1 Ô “ XDA 5201 / X- DA 5201 Thailand Chaiyaphum Pha Hin Ngam NP Dry evergreen forest near stream 15 ° 40.569 ’ N, 101 ° 26.705 ’ E 461 m leg. T 2476 Katae Sa-nog & Buakaw Adnafai Malaise traps 7. - 13. vi. 2007 Microserica sp. TigerThai 09 ” (ZFMK), 1 Ô “ X-DA 5200 / X- DA 5200 Thailand Chaiyaphum Pha Hin Ngam NP Dry evergreen forest near stream 15 ° 40.569 ’ N, 101 ° 26.705 ’ E 461 m leg. T 2476 Katae Sa-nog & Buakaw Adnafai Malaise traps 7. - 13. vi. 2007 Microserica sp. TigerThai 09 ” (QSBG), 1 Ô “ X-DA 5193 / X-DA 5193 Thailand Chaiyaphum Pha Hin Ngam NP Deciduous forest 15 ° 39.966 ’ N, 101 ° 27.198 ’ E 357 m leg. T 2467 Katae Sa-nog & Buakaw Adnafai Pan traps 8. - 9. vi. 2007 Microserica sp. TigerThai 09 ” (QSBG).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA5FF98FF3E70CF2813DD67.taxon	description	Description of holotype. Length: 5.2 mm, length of elytra: 3.5 mm, width: 3.1 mm. Body oval, dark brown, head and pronotum with some greenish shine, antenna yellow, dorsal and ventral surface dull but with iridescent shine; dorsal surface glabrous. Labroclypeus trapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and convergent to strongly rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce an indistinct blunt angle; surface shiny, weakly convex medially, coarsely and densely punctate, with a few short erect setae; frontoclypeal fine, weakly curved. Frons dull, with dense, fine punctures; with a few single setae beside eyes, otherwise glabrous. Smooth area in front of eyes flat, twice as wide as long. Ocular canthus wide and short (1 / 3 of ocular width), its external margin straight, impunctate, with a terminal seta. Eyes small, ratio of diameter / interocular width: 0.49. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.1 times as long as remaining antennomeres combined, all lamella of club equal in length. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins moderately convex and convergent to sharp, distinctly produced anterior angles, posterior angles blunt. Anterior margin of pronotum weakly convex, with fine, complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, lateral anterior margin and lateral margin sparsely setose. Hypomeron weakly carinate, not produced ventrally, not grooved. Scutellum small and triangular, finely and densely punctate, smooth along midline. Elytra oval, widest at middle, striae distinctly impressed, finely and densely punctate, intervals convex, with fine and dense punctures only concentrated along striae, along middle almost impunctate, with minute setae in punctures, odd intervals with a few single, short white, adpressed setae; epipleural edge fine, ending convex external apical angle of elytra, epipleura densely setose; apical border membraneous, with a fine fringe of short microtrichomes (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum on disc with long erect setae, otherwise sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.93. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a large, robust seta. Pygidium weakly convex and dull, with fine, dense punctures, without smooth midline, with a few long setae along apical margin, otherwise glabrous. Legs moderately wide and long; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures reduced; ventral posterior margin distinctly widened in apical half and entirely smooth, dorsal posterior margin also smooth, with a few short setae basally. Metatibia short and wide, widest at middle but only weakly narrowed towards apex, ratio width / length: 1 / 3.4; dorsal margin carinate, with two groups of spines, basal group at anterior third, apical one at three quarters of metatibial length, in basal third with a few fine, single setae; lateral face longitudinally convex, finely and sparsely punctate on dorsal and basal portion, glabrous; ventral margin finely serrate, with four equidistant setae; medial face smooth and glabrous, apex interiorly near tarsal articulation bluntly truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally, but without parallel subventral smooth carina; first metatarsomere distinctly shorter than following two tarsomeres combined, little longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 18 I – K Habitus: Fig. 18 L. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA5FF98FF3E70CF2813DD67.taxon	diagnosis	Diagnosis. Microserica chaiyaphum Ahrens, Lukic & Liu, new species is very similar to Mic. viengvai Ahrens, Lukic & Liu, new species and Mic. namnao Ahrens, Lukic & Liu, new species. Microserica chaiyaphum Ahrens, Lukic & Liu, new species differs from both by the shape of parameres: the right one is even wider (lateral view) than that of Mic. namnao, while the left paramere is not triangular, but almost evenly wide as that of Mic. viengvai, however, its apex is not bifid as in the latter but evenly broadly rounded.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA5FF98FF3E70CF2813DD67.taxon	etymology	Etymology. The species name is named after its occurrence in the Chaiyaphum province (noun in apposition).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA5FF98FF3E70CF2813DD67.taxon	description	Variation. Length: 5.1 – 5.3 mm, length of elytra: 3.5 – 3.6 mm, width: 3.1 – 3.2 mm.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA2FF98FF3E723F2EF8DC05.taxon	description	Fig. 29	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA2FF98FF3E723F2EF8DC05.taxon	materials_examined	Material examined. See Ahrens (2003: p. 39). Aedeagus. See Ahrens (2003: p. 41, figs 6 – 8).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA2FF98FF3E723F2EF8DC05.taxon	distribution	Distribution. Malay Pensinsula (Perak).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA3FF99FF3E77DA2F25DEEF.taxon	description	Fig. 29	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA3FF99FF3E77DA2F25DEEF.taxon	materials_examined	Material examined. See Ahrens (2003: p. 53). Aedeagus. See Ahrens (2003: p. 56, figs 26 – 33).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA3FF99FF3E77DA2F25DEEF.taxon	distribution	Distribution. Malay Pensinsula, Sumatra, Java, Thailand.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA3FF99FF3E74F32862DA84.taxon	description	Fig. 29	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA3FF99FF3E74F32862DA84.taxon	materials_examined	Material examined. See Ahrens (2003: p. 36). 1 Ô “ Thailand-S, 200 m, Nakhon Si Thammarat pr. Khao Luang, Kiriwong, 8 ° 27 ’ N 99 ° 44 ’ E, 1. - 6. iv. 1997, Jiří Kolibáč leg. / 1113 Sericini Asia spec. ” (NHMW). Aedeagus. See Ahrens (2003: p. 41, figs 3 – 5).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA3FF99FF3E74F32862DA84.taxon	distribution	Distribution. Malay Pensinsula, southern and eastern Thailand, Cambodia.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA3FF99FF3E76582E42D802.taxon	description	Fig. 29	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA3FF99FF3E76582E42D802.taxon	materials_examined	Material examined. See Ahrens (2003: p. 36). Aedeagus. See Ahrens (2003: p. 37, figs 1 – 5).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA3FF99FF3E76582E42D802.taxon	distribution	Distribution. Malay Pensinsula.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA0FF9AFF3E77612C90D82D.taxon	description	Fig. 30	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA0FF9AFF3E77092F80DF78.taxon	materials_examined	Material examined. See Ahrens (2002 b: p. 384). Aedeagus. See Ahrens (2002 b: p. 406, figs 1 – 3).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA0FF9AFF3E77092F80DF78.taxon	distribution	Distribution. India (Assam, Arunachal Pradesh) and northern Myanmar.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA0FF9AFF3E70392E9BDE00.taxon	description	Fig. 30	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA0FF9AFF3E70392E9BDE00.taxon	materials_examined	Material examined. See Ahrens (2002 b: p. 402). Aedeagus. See Ahrens (2002 b: p. 411, figs 35 – 37).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA0FF9AFF3E70392E9BDE00.taxon	distribution	Distribution. Malaysia (Kelantan, Benom Mts.).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA0FF9AFF3E71D12DADDDA8.taxon	description	Fig. 30	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA0FF9AFF3E71D12DADDDA8.taxon	materials_examined	Material examined. See Ahrens (2002 b: p. 395). Aedeagus. See Ahrens (2002 b: p. 409, figs 23 – 25).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA0FF9AFF3E71D12DADDDA8.taxon	distribution	Distribution. Myanmar.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA0FF9BFF3E737C2DADDBBB.taxon	description	Fig. 30	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA0FF9BFF3E737C2DADDBBB.taxon	materials_examined	Material examined. See Ahrens (2002 b: p. 389). Aedeagus. See Ahrens (2002 b: p. 407, figs 11 – 12).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA0FF9BFF3E737C2DADDBBB.taxon	distribution	Distribution. Myanmar.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA1FF9BFF3E75432EC5D959.taxon	description	Fig. 30	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA1FF9BFF3E75432EC5D959.taxon	materials_examined	Material examined. See Ahrens (2002 b: p. 394). Aedeagus. See Ahrens (2002 b: p. 409, figs 20 – 22).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA1FF9BFF3E75432EC5D959.taxon	distribution	Distribution. Myanmar, northern Thailand.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA1FF9BFF3E76ED2FC9D8F4.taxon	description	Fig. 30	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA1FF9BFF3E76ED2FC9D8F4.taxon	materials_examined	Material examined. See Ahrens (2002 b: p. 394). Aedeagus. See Ahrens (2002 b: p. 410, figs 26 – 28).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA1FF9BFF3E76ED2FC9D8F4.taxon	distribution	Distribution. Southern Vietnam, southern Laos, southern Thailand.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA1FF9BFF3E77882E92DE41.taxon	description	Fig. 30	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA1FF9BFF3E77882E92DE41.taxon	materials_examined	Material examined. See Ahrens (2002 b: p. 388). Aedeagus. See Ahrens (2002 b: p. 407, figs 7 – 10).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA1FF9BFF3E77882E92DE41.taxon	distribution	Distribution. Myanmar, northwestern Thailand.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA1FF9BFF3E71132E4DDD30.taxon	materials_examined	Material examined. See Ahrens (2002 b: p. 392). Aedeagus. See Ahrens (2002 b: p. 408, figs 17 – 19).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA1FF9BFF3E71132E4DDD30.taxon	distribution	Distribution. Southern Vietnam.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA1FF9BFF3E72C12EDDDCBB.taxon	description	Fig. 30	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA1FF9BFF3E72C12EDDDCBB.taxon	materials_examined	Material examined. See Ahrens (2002 b: p. 401). Aedeagus. See Ahrens (2002 b: p. 411, figs 32 – 34).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA1FF9BFF3E72C12EDDDCBB.taxon	distribution	Distribution. Malay Peninsula, Singapore.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAEFF94FF3E74F32E02DAF6.taxon	materials_examined	Material examined. See Ahrens (2002 b: p. 391). Aedeagus. See Ahrens (2002 b: p. 408, figs 14 – 16).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAEFF94FF3E74F32E02DAF6.taxon	distribution	Distribution. Myanmar (Tenasserim).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAEFF94FF3E75BB2E61D99E.taxon	description	Fig. 30	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAEFF94FF3E75BB2E61D99E.taxon	materials_examined	Material examined. See Ahrens (2002 b: p. 399). Aedeagus. See Ahrens (2002 b: p. 410, figs 29 – 31).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAEFF94FF3E75BB2E61D99E.taxon	distribution	Distribution. Thailand, Laos.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAEFF94FF3E77532FD6DF53.taxon	description	Fig. 30	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAEFF94FF3E77532FD6DF53.taxon	materials_examined	Material examined. See Ahrens (2002 b: p. 386); Fabrizi & Ahrens (2016: p. 134). Aedeagus. See Ahrens (2002 b: p. 406, figs 4 – 6).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAEFF94FF3E77532FD6DF53.taxon	distribution	Distribution. India (Assam, Meghalaya, and Darjeeling district).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAEFF95FF3E70B328D4DDD3.taxon	description	Figs. 19 A – D, 32	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAEFF95FF3E70B328D4DDD3.taxon	materials_examined	Type material examined. Holotype: Ô “ N-Thailand 14. - 28. VIII. 1990 Doi Inthanon lg. Malicky / Zoologische Staatssammlung München / 40 Sericini Asia spec. ” (ZSM). Paratypes: Thailand: 20 ÔÔ, 7 ♀♀ “ N-Thailand 14. - 28. VIII. 1990 Doi Inthanon lg. Malicky / Zoologische Staatssammlung München ” (ZSM, ZFMK), 9 ÔÔ, 7 ♀♀ “ N-Thailand 27. III. - 3. IV. 1990 Doi Inthanon lg. Malicky / Zoologische Staatssammlung München ” (ZSM, ZFMK), 5 ÔÔ, 1 ♀ “ N-Thailand 10. - 12. IV. 1990 Doi Inthanon lg. Malicky / Zoologische Staatssammlung München ” (ZSM, ZFMK), 2 ÔÔ “ N-Thailand 12 - 14. V. 1990 Doi Inthanon lg. Malicky / Zoologische Staatssammlung München ” (ZSM), 1 Ô “ LAOS, 21 ° 09 ’ N 101 ° 19 ’ E Louangnamtha pr., Namtha -> Muang Sing, 5 - 31. v. 1997, 900 - 1200 m Vit Kubáň leg. / LS 38 ” (CP), 3 ÔÔ “ NW Thailand, 19.19 N 97.59 E Mae Hong Son 1991 Ban Huai Po, 1600 - 2000 m, 17. - 23.5. L. Dembicky ” (NHMW), 11 ÔÔ “ W. Thailand: 300 m, Thung Yai Wildlife Sanctuary, 15 ° 28 ’ N – 98 ° 48 ’ E / Kanchaburi Prov., Sankhla Buri Dist. Mae Kasa Stream. iv / v. 1988 / M. J. D. Brendell B. M. 1988 - 183 ” (BMNH), 1 Ô “ W. Thailand: 300 m, Thung Yai Wildlife Sanctuary, 15 ° 28 ’ N – 98 ° 48 ’ E / Tak Province Umphang District Song Bae Stream. 18 - 27. iv. 1988 / M. J. D. Brendell B. M. 1988 - 183 ” (BMNH). Laos: 1 Ô “ LAOS north, 4. V. 1997 Louang Namtha env. N 21 ° 00.3, E 101 ° 24.6., E. Jendek & O. Sausa leg. ” (CP).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAEFF95FF3E70B328D4DDD3.taxon	description	Description of holotype. Length: 3.6 mm, length of elytra: 2.6 mm, width: 2.3 mm. Body oval, body yellowish brown, lateral and sutural elytral intervals and frons brown, dorsal surface dull and nearly glabrous. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins convex and convergent to moderately rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface shiny, flat, finely, densely punctate, with two long, erect setae on each side; frontoclypeal suture finely incised and curved, short, as long as smooth area in front of eye. Frons dull, with sparse, fine punctures, beside eyes with two single setae, otherwise glabrous. Smooth area in front of eyes large, mesally strongly enlarged, convex, 1.2 times as wide as long. Ocular canthus impunctate, narrow and short (1 / 4 of ocular width), with a terminal seta. Eyes moderately large, ratio of diameter / interocular width: 0.64. Antenna with ten antennomeres; club composed of five antennomeres, strongly reflexed, three times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins in basal half straight and weakly convergent, in anterior third convex and moderately convergent to moderately produced and blunt anterior angles, posterior angles blunt. Anterior margin of pronotum straight, with fine and complete marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures, lateral margin sparsely setose, anterior margin glabrous. Hypomeron distinctly carinate, carina not ventrally produced. Scutellum large, triangular, finely and densely punctate. Elytra short oval, widest at posterior third, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine and moderately dense punctures concentrated along striae, with minute setae in punctures, otherwise glabrous; epipleural edge robust, ending at slightly rounded external apical angle of elytra, epipleura densely setose; apical border broadly membraneous (visible at ca 100 x magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with numerous long, yellow, and robust setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 2.3. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium weakly convex and dull, with fine, dense punctures, without smooth midline, with minute setae in punctures and numerous long robust setae along apical margin. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur dull, anterior margin acute, with fine and continuous submarginal serrated line; anterior row of setabearing punctures reduced; ventral posterior margin strongly widened in apical half, smooth, dorsal posterior margin also smooth, with a few short setae basally. Metatibia moderately long and narrow, widest at middle, ratio width / length: 1 / 2.47; dorsal margin sharply carinate, with two groups of spines, basal group at two thirds, apical one at four fifths of metatibial length, basal to the first group over the entire length with a fine and straight, serrated carina, beside which are a few single and short setae; lateral face flat, finely and densely punctate, with minute setae in punctures; ventral margin carinate and finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally with a few single fine punctures; metatarsomeres glabrous ventrally, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined, almost twice as long as dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 19 A – C. Habitus: Fig. 19 D.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAEFF95FF3E70B328D4DDD3.taxon	diagnosis	Diagnosis. Microserica planiforceps Ahrens, Lukic & Liu, new species is in external appearance rather similar to Microserica neosimplex Ahrens, Lukic & Liu, new species However, Microserica planiforceps Ahrens, Lukic & Liu, new species differs from this and all other species by the presence of a serrated longitudinal carina in the metatibia and beside the anterior margin of metafemur and as well as by the dorsoventrally flattened parameres.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAEFF95FF3E70B328D4DDD3.taxon	etymology	Etymology. The species name is derived from the combined Latin words, “ planus ” (flat) and “ forceps ” (forceps) (adjective in nominative case), with references to the dorsoventrally flattened aedeagus including parameres.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAEFF95FF3E70B328D4DDD3.taxon	description	Variation. Length: 3.6 – 4.5 mm, length of elytra: 2.6 – 3.5 mm, width: 2.3 – 3.0 mm. Female: slightly larger in general than males, eyes smaller, ratio of diameter / interocular width: 0.57; pygidium flat, antennal club short, composed of three antennomeres, distinctly shorter than remaining antennomeres combined.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAFFF96FF3E73602EA0DCD7.taxon	description	Figs. 19 E – I, 32	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAFFF96FF3E73602EA0DCD7.taxon	materials_examined	Type material examined. Holotype: Ô “ Dalat, 1966 S. Vietnam / Type Ophthalmoserica septemfoliata G. Frey 1971 ” (CF / NHMB).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAFFF96FF3E73602EA0DCD7.taxon	description	Redescription of the holotype. Length: 4.7 mm, length of elytra: 3.0 mm, width: 2.5 mm. Body oblong oval, yellowish brown; antenna yellow; frons, small spots elytral intervals, and two large spots on each side of pronotum darker brown; dorsal surface dull, labroclypeus and anterior frons simply shiny; dorsal surface almost glabrous. Labroclypeus subrectangular and narrow, widest in anterior third, lateral margins subparallel straight and weakly convergent basally, anterior angles convex, lateral border and ocular canthus produce a distinct blunt angle, anterior and lateral margins moderately reflexed, anterior margin weakly emarginate medially; surface flat, finely and densely punctate, with a few erect setae; frontoclypeal suture short, only one quarter of clypeal width, finely impressed and weakly curved; smooth area in front of eye very large, 1.5 times as wide as long; ocular canthus long and narrow, impunctate, without a terminal seta. Frons posteriorly dull, anteriorly shiny, with fine and moderately dense punctures and with a few long setae on disc and sides. Eyes very large, ratio of diameter / interocular width: 0.93. Antenna yellow, with ten antennomeres; club with seven antennomeres equal in length, strongly reflexed, club 3.5 times as long as remaining antennomeres combined. Mentum weakly convexly elevated anteriorly. Pronotum narrow, widest at middle, lateral margins anteriorly curved and convergent anteriorly, in posterior half straight but convergent towards blunt posterior angles, anterior angles moderately produced and blunt; anterior margin weakly convex medially, with a distinct and fine marginal line; basal margin without marginal line; surface with dense and fine punctures, with minute setae in punctures, otherwise glabrous on disc; anterior and lateral borders sparsely but robustly setose; hypomeron carinate, its basal margin weakly produced ventrally. Scutellum narrow and short, with fine, irregular, and moderately dense punctures, with minute setae in punctures. Elytra oblong oval, widest at middle, striae well impressed and finely densely punctate; intervals finely punctate, punctures concentrated long striae, odd intervals convex, even intervals weakly convex, throughout with minute setae in punctures, penultimate lateral interval with a single robust long setae before apical quarter; interior apical angle of elytron with a robust seta; epipleural edge robust but convex, ending at strongly convex external apical angle of elytra; epipleura sparsely setose, apical border membraneous, with a fine rim of microtrichomes. Ventral surface dull, with fine and dense punctures, with dense short setae, setae adpressed; metacoxa minutely setose, laterally with robust adpressed setae; each abdominal sternite with indistinct transversal row of coarse and dense punctures, sternite bearing short strong setae between fine and dense punctation, otherwise almost glabrous except partly minute setae in punctures. Mesosternum between mesocoxae almost as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.5. Pygidium moderately convex, dull, with fine and dense punctures, without smooth midline, with longer setae along on apical half. Legs moderately slender; femora on ventral surface dull, finely and moderately densely punctate, glabrous, with two longitudinal rows of setae; anterior edge of metafemur acute, without adjacent continuously serrated line; posterior margin weakly convex and glabrous, ventrally weakly widened in apical half and distinctly serrate, posterior margin dorsally also finely serrate. Metatibia moderately slender and long, widest at apical third, ratio width / length: 1 / 3.5, dorsally sharply carinate, with one group of spines at five sixths of metatibial length, basally with a few single spines; with a straight and continuous serrated line from base to group of spines; lateral face weakly convex, with moderately dense and coarse punctures ,, punctures with minute setae or with a few short, robust setae; ventral margin carinate and serrated, with three robust spines of which the distal is more distant; medial face impunctate and smooth, apex interiorly (near tarsal articulation) distinctly truncate. Tarsomeres dorsally glabrous and sparsely finely punctate, ventrally with a sparse, short setae; metatarsomeres dorsally carinate, with fine punctures bearing short setae, ventrally with a single strongly serrated carina, adjacent subventral carina present, but interiorly; first metatarsomere distinctly shorter than following two tarsomeres combined and twice as long as dorsal metatibial spur. Protibia short, bidentate, protarsal claws symmetrical. Aedeagus: Figs. 19 E – H. Habitus: Fig. 20 I. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAFFF96FF3E73602EA0DCD7.taxon	discussion	Remarks. Despite the labelling on the holotype, the species was originally treated by Frey (1972) as a Triochoserica species. The systematic placement of this species is uncertain. Its morphology, particularly the serrated metatibia, the divergent genital morphology, and the constitution of the antenna are not retrieved in any other species of Microserica. Some affinities even with the species of the Neoserica calva group exist. However, given the reported strong homoplasy of the serrated metatibia in several other Sericini lineages and its considerable variation within some of these (e. g., Ahrens 2006; Bohacz & Ahrens 2020) but also of the number of antennomeres composing the antennal club, we do not see very strong arguments for a robust classification of this species, neither to erect an independent genus for it at this moment.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFADFF97FF3E74F32E62DAEA.taxon	materials_examined	Material examined. See Ahrens & Fabrizi (2016: p. 131). Aedeagus. See Ahrens & Fabrizi (2016: p. 298, Fig. 10 T – W).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFADFF97FF3E74F32E62DAEA.taxon	distribution	Distribution. Southern India.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFADFF90FF3E75B229EBD926.taxon	type_taxon	Type species: Microserica lineata Moser, 1915 (by present designation).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFADFF90FF3E75B229EBD926.taxon	diagnosis	Diagnosis. Moronoserica Ahrens, Lukic & Liu, new genus differs from its sister clade (Fig. 1), Trioserica Moser, 1922, by the bidentate protibia, the dorsal surface being more or less strongly covered with white scale-like setae, the strongly convex elytral intervals, and most of all by the strongly derived shape of aedeagus. The latter have the left paramere almost entirely reduced, the dorsoapical emargination of phallobase between the parameres extremely deep, it exceeds at least the half of the phallobasal length (the genital characters appear to be likely the only unambiguous synapomorphies). In contrast to all other Microserica species, the dorsal margin of metatibia is not carinate but longitudinally convex. The phylogeny based on molecular characters has shown that the group is not monophyletic with the remainder Microserica species. So far, the species of this group have been treated as Microserica lineata group (Ahrens 2004; Bohacz & Ahrens 2020). They are so far known only from Indochina.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFADFF90FF3E75B229EBD926.taxon	etymology	Etymology. The genus name is composed by the root “ Serica ” (type genus of Sericini) and the name Morón. The new genus is dedicated to Miguel-Angel Morón (1952 – 2017), Mexican specialist of Melolonthinae, who passed away by far much too early.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAAFF90FF3E76E92EF4D8F0.taxon	description	Fig. 31	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAAFF90FF3E76E92EF4D8F0.taxon	materials_examined	Material examined. See Bohacz & Ahrens (2020: p. 268). Aedeagus. See Bohacz & Ahrens (2020: p. 269, Figs. 3 A – D).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAAFF90FF3E76E92EF4D8F0.taxon	distribution	Distribution. Northern Laos, Thailand.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAAFF90FF3E77942E48DF95.taxon	description	Fig. 31	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAAFF90FF3E77942E48DF95.taxon	materials_examined	Material examined. See Ahrens (2004 c: p. 10); Fabrizi & Ahrens (2016: p. 130). Aedeagus. See Ahrens (2004 c: p. 11, Figs 11 – 13).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAAFF90FF3E77942E48DF95.taxon	distribution	Distribution. India (Meghalaya).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAAFF90FF3E71262E49DD1C.taxon	description	Fig. 31	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAAFF90FF3E71262E49DD1C.taxon	materials_examined	Material examined. See Ahrens (2004 c: p. 7). 1 Ô “ NE Thailand 1. - 15.5.1991 Ban Huai Po 1600 m, S. Bily leg. ” (NHMB). Aedeagus. See Ahrens (2004 c: p. 8, Figs 7 – 10).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAAFF90FF3E71262E49DD1C.taxon	distribution	Distribution. Northern Thailand.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAAFF90FF3E72DD2D88DCAB.taxon	description	Fig. 31	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAAFF90FF3E72DD2D88DCAB.taxon	materials_examined	Material examined. See Bohacz & Ahrens (2020: p. 267). Aedeagus. See Bohacz & Ahrens (2020: p. 266, Fig. 2 E – H).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFAAFF90FF3E72DD2D88DCAB.taxon	distribution	Distribution. Central Laos.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFABFF91FF3E74F32E02DAC6.taxon	materials_examined	Material examined. See Bohacz & Ahrens (2020: p. 270). 1 Ô “ Mus. Pragense Tenasserim Coll. Helfer ” (NMPC). Aedeagus. See Bohacz & Ahrens (2020: p. 269; Fig. 3 E – G).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFABFF91FF3E74F32E02DAC6.taxon	distribution	Distribution. Myanmar (Tenasserim).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFABFF91FF3E759E2E5AD865.taxon	description	Fig. 31	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFABFF91FF3E759E2E5AD865.taxon	materials_examined	Material examined. See Ahrens (2004 c: p. 2). Aedeagus. See Ahrens (2004 c: p. 3, Figs 1 – 3).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFABFF91FF3E759E2E5AD865.taxon	distribution	Distribution. Myanmar (Pegu).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFABFF91FF3E77362E48DFEF.taxon	description	Fig. 31	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFABFF91FF3E77362E48DFEF.taxon	materials_examined	Material examined. See Ahrens (2004 c: p. 5); Bohacz & Ahrens (2020: p. 265). Aedeagus. See Ahrens (2004 c: p. 5; Figs 4 – 6).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFABFF91FF3E77362E48DFEF.taxon	distribution	Distribution. Northern Thailand.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFABFF91FF3E70B72E02DEAE.taxon	materials_examined	Material examined. See Bohacz & Ahrens (2020: p. 271). Aedeagus. See Bohacz & Ahrens (2020: p. 269, Fig. 3 I – L).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFABFF91FF3E70B72E02DEAE.taxon	distribution	Distribution. Myanmar (Tenasserim).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFABFF91FF3E72762EC8DC29.taxon	description	Fig. 31	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFABFF91FF3E72762EC8DC29.taxon	materials_examined	Material examined. See Bohacz & Ahrens (2020: p. 265). 1 Ô “ Mandian (Forest), Nabanhe Nature Reserve, Jinghong, Xishuangbannna, Yunnan, 6. VI. 2009, 746 m, leg. Meng L. Z. / LW- 371 ” (IZAS). Aedeagus. See Bohacz & Ahrens (2020: p. 266, Fig. 2 A – D;).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFABFF91FF3E72762EC8DC29.taxon	distribution	Distribution. Central Laos, China (Yunnan).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA8FF93FF3E74F328E0DAE7.taxon	description	Figs. 20. A – E, 31	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA8FF93FF3E74F328E0DAE7.taxon	materials_examined	Type material examined. Holotype: Ô “ Thai 28 / 4 - 6. / 5.91 Umphang river 1000 m 16 ° 07 ’ N 88 ° 00 ’ E David Kral lgt. / 1161 Sericini Asia spec. ” (ZFMK). Paratypes: 4 ÔÔ, 3 ♀♀ “ Thai 28 / 4 - 6. / 5.91 Umphang river 1000 m 16 ° 07 ’ N 88 ° 00 ’ E David Kral lgt. ” (ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA8FF93FF3E74F328E0DAE7.taxon	description	Description of the holotype. Length: 8.4 mm, length of elytra: 5.4 mm, width: 5.0 mm. Body oval, brown, scutellum lighter brown; frons and pronotum with greenish shine, dorsal surface dull, labroclypeus simply shiny; dorsal surface except center of elytral intervals almost evenly and densely covered with fine, white setae. Labroclypeus subtrapezoidal and moderately short, widest at base, lateral margins convex and strongly convergent anteriorly, anterior angles broadly rounded, lateral border and ocular canthus produce an indistinct obtuse angle, anterior and lateral margins moderately reflexed, anterior margin distinctly sinuate medially; surface flat and shiny, finely and very densely punctate, a few larger punctures mixed with generally small ones, with numerous long, erect setae; frontoclypeal suture distinct and weakly elevated, weakly curved; smooth area in front of eye 2.5 times as wide as long; ocular canthus short and triangular (1 / 3 of ocular diameter), sparsely finely punctate, with a short single terminal seta. Frons with fine and dense punctures, with minute white setae in punctures, beside eyes with a few long erect setae. Eyes small, ratio of diameter / interocular width: 0.53. Antenna yellow, with ten antennomeres; club with four antennomeres subequal in length, club as long as remaining antennomeres combined. Mentum weakly convexly elevated anteriorly. Pronotum moderately wide, widest at base, lateral margins distinctly evenly curved and convergent anteriorly; anterior angles distinctly produced and acute, posterior angles blunt; anterior margin convexly produced medially, with a distinct and fine marginal line; basal margin without marginal line; surface with dense and fine punctures, with adpressed white minute setae in punctures; anterior and lateral borders sparsely setose; hypomeron carinate, its basal margin not produced ventrally. Scutellum narrow and short, with fine and dense punctures along lateral margins, otherwise at middle widely impunctate, minute setae present in punctures. Elytra short oval, widest at middle, striae distinctly impressed and finely densely punctate; intervals strongly convex and densely finely punctate, punctures concentrated along striae, center of intervals impunctate, with fine white setae in punctures, with a few long, yellow setae on odd intervals; interior apical angle of elytron with a robust seta; before apex with an indistinct, dark, glabrous spot; epipleural edge robust but convex, ending at strongly convex external apical angle of elytra; epipleura densely setose, apical border broadly membraneous, with a rim of short microtrichomes. Ventral surface dull, with fine and dense punctures, with dense short setae, setae adpressed; metacoxa minutely setose, laterally with robust adpressed setae; each abdominal sternite with indistinct transversal row of coarse and dense punctures, sternite bearing short strong setae between fine and dense punctation, all sternites bearing minute white setae. Mesosternum between mesocoxae almost as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.5. Pygidium strongly convex, with fine and dense punctures bearing fine, white setae, with a few longer yellowish setae, without smooth midline. Legs moderately slender; femora on ventral surface dull, finely and moderately densely punctate, glabrous, with two longitudinal rows of setae; anterior edge of metafemur acute, with a robust adjacent continuously serrated line, posterior margin weakly convex and glabrous, ventrally weakly widened in apical half but not serrate, posterior margin dorsally not serrate. Metatibia moderately wide and long, dorsal and ventral margin almost subparallel, ratio width / length: 1 / 3.3, dorsally longitudinally convex, with two groups of spines, the basal group shortly before middle, apical one at three quarters of metatibial length, basally with a few single spines; lateral face longitudinally convex, with dense and moderately coarse punctures, along the middle of lateral face narrowly impunctate, punctures with minute to short setae, before the basal group of spines with a continuously serrated line; ventral margin sharp and serrated, with five strong and long, equidistant spines; medial face impunctate and smooth, apex interiorly (near tarsal articulation) distinctly truncate. Tarsomeres dorsally glabrous and impunctate, ventrally with a very few sparse, short setae; metatarsomeres ventrally with a single strongly serrated carina, a subventral carina lacking, first metatarsomere as long as following two tarsomeres combined and one third of its length longer than dorsal metatibial spur. Protibia short, bidentate, protarsal claws symmetrical. Aedeagus: Fig. 20 A – D. Habitus: Fig. 20 E.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA8FF93FF3E74F328E0DAE7.taxon	diagnosis	Diagnosis. Moronoserica squamulatoides Ahrens, Lukic & Liu, new species differs from Mor. squamulata (Moser, 1915) new combination by fine white setae covering the elytral intervals instead of scales, and by the narrower and longer right paramere (lateral view).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA8FF93FF3E74F328E0DAE7.taxon	etymology	Etymology. The name of the new species (noun in nominative singular case) is derived from the combined words, “ squamulatus ” (with scales) and the Greek suffix “ - oides ”, with reference to the similarity to Mor. squamulata.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA8FF93FF3E74F328E0DAE7.taxon	description	Variation. Length: 8.3 – 8.4 mm, length of elytra: 5.3 – 5.4 mm, width: 4.9 – 5.0 mm. Female: Antennal club composed of three antennomeres, little shorter than the remaining antennomeres combined.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA9FFECFF3E75B42D4CD953.taxon	description	Figs. 20 F – J, 31	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA9FFECFF3E75B42D4CD953.taxon	materials_examined	Type material examined. Holotype: Ô “ Viet Nam: 10 km E of BanMe Thout 570 m, 9. V. 1060 / S. Quate Collector / 509 Asia Sericini spec. ” (BPBM).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA9FFECFF3E75B42D4CD953.taxon	description	Description of the holotype. Length: 6.4 mm, length of elytra: 4.4 mm, width: 3.9 mm. Body oval, yellowish brown, elytra, pronotum and frons with smaller, irregular but symmetric, dark spots, head and pronotum partly with greenish shine, otherwise surface dull; labroclypeus simply shiny, dorsal surface glabrous. Labroclypeus subtrapezoidal and short, widest at base, lateral margins convex and strongly convergent anteriorly, anterior angles convex, lateral border and ocular canthus produced into an indistinct, obtuse angle, anterior and lateral margins moderately reflexed, anterior margin weakly emarginate medially; surface weakly convex medially and shiny, finely and densely punctate, with a few long, erect setae; frontoclypeal suture indistinctly impressed and curved; smooth area in front of eye 2.5 times as wide as long; ocular canthus short and triangular, impunctate, with a short terminal seta. Frons dull, with fine and moderately dense punctures, beside eyes with a few long setae, otherwise with minute setae in punctures. Eyes small, ratio of diameter / interocular width: 0.5. Antenna yellow, with ten antennomeres; club with four antennomeres subequal in length, club little shorter than remaining antennomeres combined. Mentum weakly convexly elevated anteriorly. Pronotum wide, widest shortly before base; with one c-shaped central and two lateral, dark spots on each side; lateral margins convex and convergent anteriorly and posteriorly, anterior angles moderately produced and blunt, posterior angles blunt; anterior margin convexly produced medially, with a distinct and fine marginal line, basal margin without marginal line; surface with dense and fine punctures, only with minute setae in punctures, otherwise glabrous; anterior and lateral borders sparsely setose; hypomeron weakly carinate, carina not produced ventrally. Scutellum narrow and short, with fine and sparse punctures, impunctate on midline, with minute setae in punctures. Elytra short oval, widest at middle, striae finely impressed and finely densely punctate, intervals convex and finely densely punctate, punctures concentrated long striae, with a few long white setae on penultimate lateral interval; interior apical angle of elytron with a robust seta; epipleura and smaller spots on odd intervals at base and middle, sutural interval completely yellow; epipleural edge robust but convex, ending at strongly convex external apical angle of elytra; epipleura densely setose, apical border broadly membraneous, with a rim of short microtrichomes. Ventral surface dull, with fine and dense punctures, with dense short setae, setae adpressed; metacoxa minutely setose, laterally with robust adpressed setae; each abdominal sternite with indistinct transversal row of coarse and dense punctures, sternite bearing short strong setae between fine and dense punctation, all sternites bearing minute white setae. Mesosternum between mesocoxae almost as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.84. Pygidium moderately convex, with fine and dense punctures bearing only minute setae, with a few long, fine setae beside apical margin, without smooth midline. Legs moderately slender; femora on ventral surface dull, finely and moderately densely punctate, glabrous, with two longitudinal rows of setae; anterior edge of metafemur acute, with a robust adjacent continuously serrated line, posterior margin weakly convex and glabrous, ventrally weakly widened in apical half but not serrate, posterior margin dorsally not serrate. Metatibia moderately slender and long, widest at middle, dorsal and ventral margin almost subparallel, ratio width / length: 1 / 3.3, dorsally longitudinally convex, with two groups of spines, basal one shortly before middle, apical one at three quarters of metatibial length, basally with a few single spines in the punctures; lateral face longitudinally convex, with dense and fine punctures, with minute setae, before the basal group of spines without serrated line; ventral margin sharp and serrated, with four robust and long, equidistant spines; medial face impunctate and smooth, apex interiorly (near tarsal articulation) distinctly truncate. Tarsomeres dorsally glabrous and impunctate, ventrally with a very few sparse, short setae. Metatarsomeres ventrally with a single strongly serrated carina, a subventral carina lacking; first metatarsomere as long as following two tarsomeres combined and almost twice as long as dorsal metatibial spur. Protibia short, bidentate, protarsal claws symmetrical. Aedeagus: Fig. 20 F-I. Habitus: Fig. 20 J. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA9FFECFF3E75B42D4CD953.taxon	diagnosis	Diagnosis. Moronoserica banmethout Ahrens, Lukic & Liu, new species differs from Mor. helferi (Bohacz & Ahrens, 2020) by the distally narrowed left lamina of phallobase, which is in Mor. helferi widened towards apex; the left paramere in the new species is straight and convex at apex, while in Mor. helferi it is sharply pointed at apex and curved (lateral view).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFA9FFECFF3E75B42D4CD953.taxon	etymology	Etymology. The name of the new species (noun in apposition) is derived from the type locality of the species, BanMe Thout.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD6FFEDFF3E76EB2813D833.taxon	description	Figs. 20 K – O, 31	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD6FFEDFF3E76EB2813D833.taxon	materials_examined	Type material examined. Holotype: Ô “ Siam Renong / Doherty / Fry Coll. 1905.100. / 529 Asia Sericini spec. ” (BMNH). Paratype: 1 Ô “ Mt. Phu Rua Summit Loei Prov. NE Thailand 24. V. 2004 Y. Komiya leg. ” (ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD6FFEDFF3E76EB2813D833.taxon	description	Description of the holotype. Length: 5.2 mm, length of elytra: 3.6 mm, width: 3.2 mm. Body oval, brown; antenna yellow; frons, small spots elytral intervals, and multiple smaller, symmetric spots on each side of pronotum darker, with greenish shine; dorsal surface iridescent shiny, partly dull, labroclypeus simply shiny; dorsal surface almost evenly and moderately densely covered with small white setae, elytra with single long erect setae, dark spots glabrous. Labroclypeus subtrapezoidal and narrow, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles convex, lateral border and ocular canthus produce an indistinct obtuse angle, anterior and lateral margins moderately reflexed, anterior margin weakly emarginate medially; surface weakly convex medially and shiny, finely and densely punctate, with a few erect setae; frontoclypeal suture indistinctly impressed and weakly curved; smooth area in front of eye 1.5 times as wide as long; ocular canthus short and triangular, almost sparsely finely punctate, with a short terminal seta. Frons dull, anteriorly narrowly shiny, glabrous, with fine and moderately dense punctures, beside eyes with a few long, single setae. Eyes moderately large, ratio of diameter / interocular width: 0.61. Antenna yellow, with ten antennomeres; club with four antennomeres subequal in length, club as long as remaining antennomeres combined. Mentum weakly convexly elevated anteriorly. Pronotum narrow, widest at base, lateral margins weakly curved and convergent anteriorly, anterior angles moderately produced and acute, posterior angles moderately blunt; anterior margin convex, with a distinct and fine marginal line; basal margin without marginal line; surface with dense and fine punctures, with adpressed, moderately dense, white, short setae, darker sport with less dense punctures; anterior and lateral borders sparsely setose; hypomeron carinate, its basal margin weakly produced ventrally. Scutellum narrow and short, apex weakly rounded, with fine and moderately dense punctures, with minute setae in punctures. Elytra oval, widest at middle, striae finely impressed and finely densely punctate; intervals moderately convex and finely punctate, punctures concentrated long striae, throughout with fine, white setae in punctures, odd intervals with single, long, erect setae; before apex a large dark spot, also smaller dark spots impunctate; interior apical angle of elytron with a robust seta; epipleural edge robust but convex, ending at strongly convex external apical angle of elytra; epipleura densely setose, apical border broadly membraneous. Ventral surface dull, with fine and dense punctures, with dense short setae, setae adpressed; metacoxa minutely setose, laterally with robust adpressed setae; each abdominal sternite with indistinct transversal row of coarse and dense punctures, sternite bearing short strong setae between fine and dense punctation, all sternites bearing minute white setae. Mesosternum between mesocoxae almost as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.62. Pygidium weakly convex, with fine and moderately dense punctures bearing fine white setae, with larger setae along on apical half, without smooth midline. Legs moderately slender; femora on ventral surface dull, finely and moderately densely punctate, glabrous, with two longitudinal rows of setae; anterior edge of metafemur acute, with a robust adjacent continuously serrated line, posterior margin weakly convex and glabrous, ventrally weakly widened in apical half but not serrate, posterior margin dorsally not serrate. Metatibia moderately slender and long, widest at middle, ratio width / length: 1 / 2.7, dorsally longitudinally convex, with two groups of spines, the basal group at middle, apical one at three quarters of metatibial length, basally with a few single spines in the punctures; lateral face longitudinally convex, with dense and moderately coarse punctures, along the middle of lateral face narrowly impunctate, punctures with minute setae, with a straight serrated line from base to posterior group of spines which is interrupted by anterior group of spines; ventral margin sharp and serrated, with four robust, equidistant spines; medial face impunctate and smooth, apex interiorly (near tarsal articulation) distinctly truncate. Tarsomeres dorsally glabrous and impunctate, ventrally with a very few sparse, short setae; metatarsomeres ventrally with a single strongly serrated carina, a subventral carina lacking, first metatarsomere as long as following two tarsomeres combined and distinctly longer than dorsal metatibial spur. Protibia short, bidentate, protarsal claws symmetrical. Aedeagus: Fig. 20 K – N. Habitus: Fig. 20 O. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD6FFEDFF3E76EB2813D833.taxon	diagnosis	Diagnosis. Moronoserica renong Ahrens, Lukic & Liu, new species differs from Mor. helferi (Bohacz & Ahrens, 2020) by the long, erect setae on odd intervals of elytra, the narrower and more strongly curved right paramere as well as by the left lamina of phallobase, which has in Mor. renong Ahrens, Lukic & Liu, new species a large hooked before apex (lateral view); in Mor. helferi the preapical ventral hook of phallobase is only small.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD6FFEDFF3E76EB2813D833.taxon	etymology	Etymology. The name of the new species (noun in apposition) is derived from the type locality of the species, Renong.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD6FFEDFF3E76EB2813D833.taxon	description	Variation. Length: 5.1 – 5.2 mm, length of elytra: 3.5 – 3.6 mm, width: 3.1 – 3.2 mm.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD7FFEEFF3E77C02D2ADF37.taxon	description	Figs. 21 A – D, 31	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD7FFEEFF3E77C02D2ADF37.taxon	materials_examined	Type material examined. Holotype: Ô “ W. Thailand: 300 m Thung Yai Wildlife Sanctuary, 15 ° 28 ’ N - 98 ° 48 ’ E / Tak Province Umphang District Song Bae Stream. 18 - 27. iv. 1988 / at light / M. J. D. Brendell B. M. 1988 - 183. / 486 Asia Sericini spec. ” (BMNH).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD7FFEEFF3E77C02D2ADF37.taxon	description	Description of the holotype. Length: 6.1 mm, length of elytra: 3.8 mm, width: 3.2 mm. Body oval, brown, elytra yellow with numerous dark spots; frons, pronotum and spots on elytra with greenish shine, dorsal surface dull, partly iridescent; labroclypeus simply shiny; pronotum and elytra partly moderately densely covered with white, robust, scale-like setae. Labroclypeus subtrapezoidal and moderately short, widest at base, lateral margins convex and strongly convergent anteriorly, anterior angles broadly rounded, lateral border and ocular canthus produced into an indistinct obtuse angle, anterior and lateral margins moderately reflexed, anterior margin weakly emarginate medially; surface weakly convex medially and shiny, finely and densely punctate, with numerous long, erect setae; frontoclypeal suture indistinctly impressed and weakly curved; smooth area in front of eye twice as wide as long; ocular canthus short and triangular, almost sparsely finely punctate, with a short single terminal seta. Frons dull, anteriorly narrowly shiny, with fine and moderately dense punctures, beside eyes and behind frontoclypeal suture with a few long erect setae. Eyes small, ratio of diameter / interocular width: 0.57. Antenna yellow, with ten antennomeres; club with four antennomeres subequal in length, club as long as remaining antennomeres combined. Mentum weakly convexly elevated anteriorly. Pronotum wide, widest at middle, lateral margins convex and convergent anteriorly and posteriorly, anterior angles moderately produced and acute, posterior angles convex; anterior margin convexly produced medially, with a distinct and fine marginal line, basal margin without marginal line; surface with dense and fine punctures, on disc and lateral portions with adpressed white, partly scale-like setae; anterior and lateral borders sparsely setose; hypomeron weakly carinate, carina not produced ventrally. Scutellum narrow and short, with fine and dense punctures, impunctate along midline, with minute setae in punctures. Elytra short oval, widest at middle, striae finely impressed and finely densely punctate, intervals moderately convex and finely punctate, punctures concentrated long striae, with a few long white setae on odd intervals, and on lateral intervals with sparse small, white setae; interior apical angle of elytron with a robust seta; margins, smaller spots on odd intervals, and a glabrous large spot before apex dark; epipleural edge robust but convex, ending at strongly convex external apical angle of elytra; epipleura densely setose, apical border broadly membraneous, with a rim of short microtrichomes. Ventral surface dull, with fine and dense punctures, with dense short setae, setae adpressed; metacoxa minutely setose, laterally with robust adpressed setae; each abdominal sternite with indistinct transversal row of coarse and dense punctures, sternite bearing short strong setae between fine and dense punctation, all sternites bearing minute white setae. Mesosternum between mesocoxae almost as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.49. Pygidium weakly convex, with fine and dense punctures bearing fine white setae, on apex with numerous long, fine setae, without smooth midline. Legs moderately slender; femora on ventral surface dull, finely and moderately densely punctate, glabrous, with two longitudinal rows of setae; anterior edge of metafemur acute, with a robust adjacent continuously serrated line, posterior margin weakly convex and glabrous, ventrally weakly widened in apical half but not serrate, posterior margin dorsally not serrate. Metatibia moderately slender and long, dorsal and ventral margin almost subparallel, ratio width / length: 1 / 3.4, dorsally longitudinally convex, with two groups of spines, the basal group at one third, apical one at four fifths of metatibial length, basally with a few single spines in the punctures; lateral face longitudinally convex, with dense and moderately coarse punctures, with minute setae, before the basal group of spines without serrated line; ventral margin sharp and serrated, with four robust and long, equidistant spines; medial face impunctate and smooth, apex interiorly (near tarsal articulation) distinctly truncate. Tarsomeres dorsally glabrous and impunctate, ventrally with a very few sparse, short setae. Metatarsomeres ventrally with a single strongly serrated carina, a subventral carina lacking; first metatarsomere as long as following two tarsomeres combined and twice as long as dorsal metatibial spur. Protibia short, bidentate, protarsal claws symmetrical. Aedeagus: Figs. 21 A – C. Habitus: 21 D. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD7FFEEFF3E77C02D2ADF37.taxon	diagnosis	Diagnosis. Moronoserica songbae Ahrens, Lukic & Liu, new species differs from all other species of Moronoserica by the relatively small dorsomedian emargination of phallobase which is in this species almost only half of the phallobasal length; in all other species the dorsomedian emargination is reaching almost the phallobasal apodeme (Ahrens 2006).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD7FFEEFF3E77C02D2ADF37.taxon	etymology	Etymology. The name of the new species (noun in apposition) is derived from the type locality of the species, Song Bae stream.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD4FFEFFF3E70CF2D9CDEF3.taxon	description	Figs. 21 E – I, 31	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD4FFEFFF3E70CF2D9CDEF3.taxon	materials_examined	Type material examined. Holotype: Ô “ W. Thailand: 300 m Thung Yai Wildlife Sanctuary, 15 ° 28 ’ N - 98 ° 48 ’ E / Tak Province Umphang District Song Bae Stream. 18 - 27. iv. 1988 / at light / M. J. D. Brendell B. M. 1988 - 183. ” (BMNH).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD4FFEFFF3E70CF2D9CDEF3.taxon	description	Description of the holotype. Length: 4.4 mm, length of elytra: 3.1 mm, width: 2.9 mm. Body oval, brown, elytra yellow with numerous dark spots; frons, pronotum and spots on elytra with greenish shine, dorsal surface dull, partly iridescent; labroclypeus simply shiny; pronotum and elytra partly moderately densely covered with white, robust, scale-like setae. Labroclypeus subtrapezoidal and short, widest at base, lateral margins convex and strongly convergent anteriorly, anterior angles convex, lateral border and ocular canthus produced into an indistinct, obtuse angle, anterior and lateral margins moderately reflexed, anterior margin weakly emarginate medially; surface weakly convex medially and shiny, finely and densely punctate, with numerous long, erect setae; frontoclypeal suture indistinctly impressed and weakly curved; smooth area in front of eye twice as wide as long; ocular canthus short and triangular, densely and finely punctate, with a short single terminal seta. Frons dull, anteriorly narrowly shiny, with fine and moderately dense punctures, beside eyes and behind frontoclypeal suture with a few long erect setae, otherwise with minute setae in punctures. Eyes small, ratio of diameter / interocular width: 0.59. Antenna yellow, with ten antennomeres; club with four antennomeres subequal in length, club as long as remaining antennomeres combined. Mentum weakly convexly elevated anteriorly. Pronotum wide, widest shortly before base, lateral margins convex and convergent anteriorly and posteriorly, anterior angles moderately produced and acute, posterior angles convex; anterior margin convexly produced medially, with a distinct and fine marginal line, basal margin without marginal line; surface with dense and fine punctures, lateral portions with adpressed white, partly scale-like setae; anterior and lateral borders sparsely setose; hypomeron weakly carinate, carina not produced ventrally. Scutellum narrow and short, with fine and sparse punctures, impunctate on base, with minute setae in punctures. Elytra short oval, widest at middle, striae finely impressed and finely densely punctate, intervals moderately convex and finely punctate, punctures concentrated long striae, with a few long white setae on odd intervals, and on lateral intervals with sparse small, white setae; interior apical angle of elytron with a robust seta; margins, smaller spots on odd intervals, and a glabrous large spot before apex dark; epipleural edge robust but convex, ending at strongly convex external apical angle of elytra; epipleura densely setose, apical border broadly membraneous, with a rim of short microtrichomes. Ventral surface dull, with fine and dense punctures, with dense short setae, setae adpressed; metacoxa minutely setose, laterally with robust adpressed setae; each abdominal sternite with indistinct transversal row of coarse and dense punctures, sternite bearing short strong setae between fine and dense punctation, all sternites bearing minute white setae. Mesosternum between mesocoxae almost as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.66. Pygidium weakly convex, with fine and dense punctures bearing white adpressed, partly scalelike setae, on apex with numerous long, fine setae, without smooth midline. Legs moderately slender; femora on ventral surface dull, finely and moderately densely punctate, glabrous, with two longitudinal rows of setae; anterior edge of metafemur acute, with a robust adjacent continuously serrated line, posterior margin weakly convex and glabrous, ventrally weakly widened in apical half but not serrate, posterior margin dorsally not serrate. Metatibia moderately slender and long, widest at middle, dorsal and ventral margin almost subparallel, ratio width / length: 1 / 2.9, dorsally longitudinally convex, with two groups of spines, the basal group at one third, apical one at four fifths of metatibial length, basally with a few single spines in the punctures; lateral face longitudinally convex, with dense and moderately coarse punctures, with minute setae, before the basal group of spines without serrated line; ventral margin sharp and serrated, with four robust and long, equidistant spines; medial face impunctate and smooth, apex interiorly (near tarsal articulation) distinctly truncate. Tarsomeres dorsally glabrous and impunctate, ventrally with a very few sparse, short setae. Metatarsomeres ventrally with a single strongly serrated carina, a subventral carina lacking; first metatarsomere as long as following two tarsomeres combined and twice as long as dorsal metatibial spur. Protibia short, bidentate, protarsal claws symmetrical. Aedeagus: Figs. 21 E – H. Habitus: Fig. 21 I. Female unknown.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD4FFEFFF3E70CF2D9CDEF3.taxon	diagnosis	Diagnosis. Moronoserica thungyai Ahrens, Lukic & Liu, new species differs from Mor. tenasserimensis (Bohacz & Ahrens, 2020) by the sinuated dorsal margin of the right paramere, which as at apex a well-pronounced convexity (lateral view). From the syntopically occurring previous species Moronoserica thungyai new species differs as mentioned above by the deeper dorsomedian emargination of phallobase which almost exceeds up to the phallobasal apodeme.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD4FFEFFF3E70CF2D9CDEF3.taxon	etymology	Etymology. The name of the new species (noun in apposition) is derived from the type locality of the species, Thung Yai Wildlife Sanctuary.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD2FFE8FF3E77B92E7CDE06.taxon	materials_examined	Material examined. See Ahrens (1995 b: p. 49; 2001 a: p. 220; 2004 b: p. 191; 2006 a: p. 413); Ahrens & Fabrizi (2011: p. 163). Aedeagus. See Ahrens (1995 b: p. 50, figs 43 – 35).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD2FFE8FF3E77B92E7CDE06.taxon	distribution	Distribution. Central Nepal.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD2FFE8FF3E71DE2F06DDA8.taxon	materials_examined	Material examined. See Ahrens (2001 a: p. 223; 2004 b: p. 191). Aedeagus. See Ahrens (2001 a: p. 222, figs 5 – 7).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD2FFE8FF3E71DE2F06DDA8.taxon	distribution	Distribution. Central Nepal, India (Darjeeling / Sikkim area).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD2FFE9FF3E737C2EB0DBBB.taxon	materials_examined	Material examined. See Ahrens (1995 b: p. 52; 2001 a: p. 220; 2004 b: p. 192; 2006 a: p. 413). Aedeagus. See Ahrens (1995 b: p. 53, figs 52 – 54).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD2FFE9FF3E737C2EB0DBBB.taxon	distribution	Distribution. India (Darjeeling / Sikkim area).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD3FFE9FF3E75432E7CD97D.taxon	materials_examined	Material examined. See Sreedevi et al. (2019: p. 17). Aedeagus. See Sreedevi et al. (2019: p. 19; Fig. 3 E – H).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD3FFE9FF3E75432E7CD97D.taxon	distribution	Distribution. Central Nepal.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD3FFE9FF3E76012EFDD81F.taxon	materials_examined	Material examined. See Ahrens (1995 b: p. 49; 2001 a: p. 219; 2004 b: p. 191). Aedeagus. See Ahrens (1995 b: p. 50, figs 40 – 42).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD3FFE9FF3E76012EFDD81F.taxon	distribution	Distribution. Eastern central Himalaya.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD3FFE9FF3E77A72E7ADFA0.taxon	materials_examined	Material examined. See Ahrens (1995 b: p. 51; 2004 b: p. 192; 2006 a: p. 413). Aedeagus. See Ahrens (1995 b: p. 50, figs 46 – 48).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD3FFE9FF3E77A72E7ADFA0.taxon	distribution	Distribution. Eastern Nepal.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD3FFE9FF3E71442E19DD45.taxon	materials_examined	Material examined. See Ahrens (1995 b. p. 51; 2004 b: p. 192). Aedeagus. See Ahrens (2001 a: p. 222; figs 4, 8).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD3FFE9FF3E71442E19DD45.taxon	distribution	Distribution. India (Darjeeling area).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD3FFE9FF3E72192E7CDCC9.taxon	materials_examined	Material examined. See Ahrens (1995 b: p. 48; 2001 a: p. 222). Aedeagus. See Ahrens (1995 b: p. 47, figs 37 – 39).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD3FFE9FF3E72192E7CDCC9.taxon	distribution	Distribution. Central Nepal.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD0FFEAFF3E76342E06D8F0.taxon	materials_examined	Material examined. See Ahrens (1995 b: p. 46; 2001 a: p. 218; 2004 b: p. 191; 2006 a: p. 413); Ahrens & Fabrizi (2011: p. 162). Aedeagus. See Ahrens (1995 b: p. 47, figs 34 – 36).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD0FFEAFF3E76342E06D8F0.taxon	distribution	Distribution. Inner central Himalaya.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD0FFEAFF3E74F32EB6DA90.taxon	materials_examined	Material examined. See Ahrens (1995 b: p. 45; 2001 a: p. 217; 2004 b: p. 190; 2006 a: p. 412); Ahrens & Fabrizi (2009 b: p. 273); Shrestha et al. (2012: p. 283); Fabrizi & Ahrens (2016: p. 136). Aedeagus. See Ahrens (1995 b: p. 47, figs 31 – 33).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD0FFEAFF3E74F32EB6DA90.taxon	distribution	Distribution. Western and central Himalaya.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD0FFEAFF3E77942DB7DF92.taxon	materials_examined	Material examined. See Ahrens (1995 b: p. 52; 2001 a: p. 220; 2004 b: p. 192). Aedeagus. See Ahrens (1995 b: p. 53, figs 49 – 51).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD0FFEAFF3E77942DB7DF92.taxon	distribution	Distribution. Bhutan.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD1FFEBFF3E77B42E7ADFD2.taxon	materials_examined	Material examined. See Ahrens (1998 a: p. 45). Aedeagus. See Ahrens (1998 a: p. 46, figs 35 – 37).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD1FFEBFF3E77B42E7ADFD2.taxon	distribution	Distribution. Eastern Nepal.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD1FFEBFF3E71602EB1DE9E.taxon	materials_examined	Material examined. See Ahrens (1998 a: p. 37); Fabrizi & Ahrens (2016: p. 130). Aedeagus. See Ahrens (1998 a: p. 36, figs 14 – 16).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD1FFEBFF3E71602EB1DE9E.taxon	distribution	Distribution. Bhutan and India (Meghalaya).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD1FFEBFF3E725C2E7ADC4A.taxon	materials_examined	Material examined. See Ahrens (1999 a: p. 243). Aedeagus. See Ahrens (1999 a: p. 244, figs 1 – 3).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD1FFEBFF3E725C2E7ADC4A.taxon	distribution	Distribution. Eastern Nepal.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD1FFE4FF3E73082DB7DB97.taxon	materials_examined	Material examined. See Ahrens (1998 a: p. 38). Aedeagus. See Ahrens (1998 a: p. 40, figs 20 – 22).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD1FFE4FF3E73082DB7DB97.taxon	distribution	Distribution. Bhutan.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDEFFE4FF3E752F2E10D935.taxon	materials_examined	Material examined. See Ahrens (1998 a: p. 39; 2004 b: p. 183; 2006 a: p. 412); Ahrens & Fabrizi (2009 b: p. 272; 2011: p. 163). Aedeagus. See Ahrens (1998 a: p. 40, figs 23 – 25).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDEFFE4FF3E752F2E10D935.taxon	distribution	Distribution. Western central Nepal.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDEFFE4FF3E76C92DB7D8F4.taxon	materials_examined	Material examined. See Ahrens (1998 a: p. 37; 2004 b: p. 183). Aedeagus. See Ahrens (1998 a: p. 38, figs 17 – 19).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDEFFE4FF3E76C92DB7D8F4.taxon	distribution	Distribution. Bhutan.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDEFFE4FF3E77882F21DE78.taxon	materials_examined	Material examined. See Ahrens (1998 a: p. 34; 2004 b: p. 180; 2006 a: p. 412); Ahrens & Fabrizi (2009 b: p. 273). Aedeagus. See Ahrens (1998 a: p. 33, figs 8 – 10).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDEFFE4FF3E77882F21DE78.taxon	distribution	Distribution. Kumaon-Himalaya to eastern central Nepal.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDEFFE4FF3E710C2F05DDDF.taxon	materials_examined	Material examined. See Ahrens (1998 a: p. 32; 2004 b: p. 179; 2006 a: p. 412). Aedeagus. See Ahrens (1998 a: p. 31, figs 1 – 3).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDEFFE4FF3E710C2F05DDDF.taxon	distribution	Distribution. Eastern Nepal, India (Darjeeling / Sikkim area).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDEFFE4FF3E73672E7ADC81.taxon	materials_examined	Material examined. See Ahrens (1998 a: p. 43; 2004 b: p. 183). Aedeagus. See Ahrens (1998 a: p. 44, figs 31 – 33).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDEFFE4FF3E73672E7ADC81.taxon	distribution	Distribution. Eastern Nepal.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDFFFE5FF3E74F32E10DAEA.taxon	materials_examined	Material examined. See Ahrens (1998 a: p. 46). Aedeagus. See Ahrens (1998 a: p. 47, figs 40 – 42).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDFFFE5FF3E74F32E10DAEA.taxon	distribution	Distribution. Western central Nepal.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDFFFE5FF3E75B22EB0D9AD.taxon	materials_examined	Material examined. See Ahrens (1998 a: p. 50; 2006 a: p. 412). Aedeagus. See Ahrens (1998 a: p. 47, figs 43 – 45).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDFFFE5FF3E75B22EB0D9AD.taxon	distribution	Distribution. India (Darjeeling / Sikkim area).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDFFFE5FF3E77702F53DF0C.taxon	materials_examined	Material examined. See Ahrens (1998 a: p. 39; 2004 b: p. 183; 2006 a: p. 412); Shrestha et al. (2012: p. 381); Fabrizi & Ahrens (2016: p. 132). Aedeagus. See Ahrens (1998 a: p. 42, figs 26 – 30).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDFFFE5FF3E77702F53DF0C.taxon	distribution	Distribution. Himalaya, Sutlej river to eastern Nepal.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDFFFE5FF3E70D02E10DECF.taxon	materials_examined	Material examined. See Ahrens (1998 a: p. 43; 2004 b: p. 182). Aedeagus. See Ahrens (1998 a: p. 44, figs 34 – 36).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDFFFE5FF3E70D02E10DECF.taxon	distribution	Distribution. Western central Nepal.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDFFFE5FF3E71972E10DC4C.taxon	materials_examined	Material examined. See Ahrens (1998 a: p. 32; 2004 b: p. 181; 2006 a: p. 412); Ahrens & Fabrizi (2011: p. 163); Shrestha et al. (2012: p. 381). Aedeagus. See Ahrens (1998 a: p. 33, figs 5 – 7).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDFFFE5FF3E71972E10DC4C.taxon	distribution	Distribution. Western central Nepal.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDFFFE6FF3E73102FE5DA73.taxon	materials_examined	Material examined. See Ahrens (2004 b: p. 186). 1 Ô “ Burma Mishmi Hills, Dingliang. 2450 ft. 13. iii. 1935 M. Steele. / Brit. Mus. 1935 - 312 ” (BMNH). Aedeagus. See Ahrens (2004 b: p. 424, figs 284 – 286).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDFFFE6FF3E73102FE5DA73.taxon	distribution	Distribution. Southeastern Tibet and Myanmar (Mishmi Hills).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDCFFE6FF3E75002EB0D9CC.taxon	materials_examined	Material examined. See Ahrens (1998 a: p. 35; 2004: p. 181); Ahrens & Fabrizi (2009 b: p. 273). Aedeagus. See Ahrens (1998 a: p. 36, figs 11 – 13).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDCFFE6FF3E75002EB0D9CC.taxon	distribution	Distribution. India (Sikkim / Darjeeling area).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDCFFE7FF3E768D2E03D883.taxon	description	Figs. 22 A – D, 32	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDCFFE7FF3E768D2E03D883.taxon	materials_examined	Type material examined. Lectotype (here designated): Ô “ Chang Yang near Ichang 4 - 6000 ft. V, VII, VIII / nigropicta Fairm. Cotype [handwriting. Fairm.] ” (MNHN). Paralectotype: 1 ♀ “ Chang Yang / Museum Paris 1906 Coll. Leon Faimaire / Serica nigropicta F. ” (MNHN). Additional material examined. 1 ex. “ Chang Yang, near Ichang 4 - 6000 ft. V. VII. VIII. / Museum Paris Yang- Tse-Kiang Chang-Yang Brenske 1898 / S. nigropicta Typ. Comp. Fairm. Brenske det. 1898 ” (MNHN), 2 ex. “ Museum Paris Kouy-Tchéou Gan Chouen Fou, Kiang-Long et Tun-Lin-Tchéou, P. Cavalerie 1912 ” (MNHN), 2 ÔÔ, 2 ♀♀ “ Chang Yang near Ichang 6 - 8000 ft. June 88. / Coll. Kraatz / Moser det. ” (DEIC), 5 ex. “ Museum Paris KouyTchéou Reg. de Pin-Fa, Père Cavalerie 1908 ” (MNHN), 2 ex. “ Museum Paris Kouy-Tchéou, P. Cavalerie 1910 ” (MNHN), 2 ex. “ Museum Paris Kouy-Tchéou Gan Chouen Min y Fou et Loyang, P. Cavalerie 1912 ” (MNHN), 1 ex. “ Museum Paris Kouy-Tchéou, Kouy-Yang, P. P. Cavalerie et Fortunat 1906 ” (MNHN), 1 ex. “ Yunnan-Sen Mt. Excoffier 1898 ” (MNHN), 2 ex. “ Chine Yunnan Recu de Lou-Nan ” (MNHN).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDCFFE7FF3E768D2E03D883.taxon	description	Redescription. Length: 5.0 mm, length of elytra: 3.4 mm, width: 3.1 mm. Body oval, black, legs brown, elytra reddish brown with margins and one transversal mesolateral and one anteromedial spot black, dorsal surface dull and nearly glabrous. Labroclypeus transversal, wider than long, shiny, widest at base, lateral margins in basal half almost straight, anteriorly strongly convex and convergent to rounded anterior angles; anterior margin weakly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface almost flat, coarsely and densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons dull, anterior quarter not covered with tomentum and shiny, with dense, fine punctures, beside eyes and behind frontoclypeal suture with numerous long setae, otherwise with minute setae in punctures. Smooth area in front of eyes convex, as wide as long. Ocular canthus finely densely punctate, wide and moderately long (1 / 3 of ocular diameter), with two terminal setae. Eyes very small, ratio of diameter / interocular width: 0.38. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.5 times as long as than remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide, widest at base, lateral margins in basal half subparallel, anteriorly convex and convergent towards sharp and produced anterior angles; posterior angles blunt. Anterior margin of pronotum moderately convex, with fine but medially widely interrupted marginal line; basal marginal line absent; surface finely and densely punctate, glabrous, with minute setae in punctures; setae of lateral and anterior margins dense, extending on the disc around anterior angles. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, on base along middle narrowly impunctate. Elytra elongate, widest at posterior third, striae weakly impressed, finely and densely punctate; sutural interval and lateral 2 – 3 intervals black, with a continuous black dot at anterior quarter from interval 2 to 4, and on posterior elytra between interval 4 to 7; intervals weakly convex, with fine and sparse punctures, with minute setae in punctures, penultimate lateral interval with a few short setae; epipleural edge fine, ending at rounded external apical angle of elytra, epipleura sparsely setose; apical border narrowly membranous, with a fine rim of short microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.6. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium weakly convex and dull, with dense, coarse punctures but without smooth midline, in punctures with numerous minute setae, with a few longer setae along apical margin. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; posterior margin smooth ventrally, widened in apical half, posterior margin smooth dorsally, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio of width / length: 1 / 2.84; dorsal margin sharply carinate, with two groups of spines, basal group at anterior third, apical one at two thirds of metatibial length, in basal third without further robust setae; lateral face longitudinally convex, finely and sparsely punctate, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation shallowly concavely truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere little shorter than following two tarsomeres combined, little longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 22 A – C. Habitus: 22 D.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDDFFE0FF3E70502E04DCDD.taxon	description	Figs. 22 E – H, 32	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDDFFE0FF3E70502E04DCDD.taxon	materials_examined	Type material examined. Syntypes: 1 Ô “ Tchao Pin-io Chine / Microserica diversicornis Type Ô Mos ” (ZMHB), 1 ♀ “ Tchao Pin-io Chine / Microserica diversicornis Type Ô Mos ” (ZMHB). Additional material examined. 3 ex. “ Yünnan China ” (ZMHB), 6 ex. “ Yun-nan sen ” (ZMHB), 1 ex. “ W. Yunnan Chao-chow-fu 23.8. - 21.9.1914 2300 m. Hohe / Berghang mit viel Busch, gekaeschert Meil. S. ” (ZMHB), 2 ex. “ Chine Kong-Tchéou R. P. Esquirol 1912 “ (MNHN), 3 ex. “ Museum Paris Haut Plateau Yunnanais Distr. de Yunnan-Fou 1850 - 2000 m, 25 Lat. Dr. Legendre 1915 / Houa Ting-Se, bord du lac 23 Mars 1914 “ (MNHN), 1 ex. “ China, W. Yunnan Baoshan env. 5 - 8. VI. 1993 E. Jendek & O. Sausa leg. ” (CP), 2 ex. “ China Yunnan Lijiang 8 - 24. VI. 1994 leg. B. Šiška & T. Spevár ” (CP), 1 ex. “ China VI. 1990 Lijiang leg. Bocak ” (CP), 2 ex. “ China-Yunnan 22.5. - 2.6. 100 km W Kunming Diaolin Nat. Res. 1993 E. Jendek & O. Sausa leg. [sic!] ” (NHMW), 1 ex. “ China, Yunnan, 19. VII. 1993 Dali, Hua Dian Ba 50 km N of Dali C. Holzschuh leg. ” (CP), 1 Ô, 3 ♀♀ “ China: Yunnan above Dali, 2000 - 2200 m 4. - 17. IV. 1999 leg. W. Schawaller ” (SMNS), 1 ♀ “ China-Yünnan W Dali, 1900 - 2400 m 25 ° 45 ‘ N 100 ° 06 ‘ E 8. - 11. VII. 1996 Chr. Häuser leg. ” (SMNS), 1 Ô “ China: Yunnan Tse-Kou, 2000 m 28 ° 02 ’ N, 98 ° 54 ’ E 7. + 8.6.2006 leg. Jiri Klir ” (NMPC), 1 ex. “ Yunnan-North 30 km E Lijiang, N 26 ° 50 ’ - E 100 ° 24 ’ 2500 m, 4.6.2006 V. Major leg. ” (CP), 1 ex. “ S China, Yunnan prov., 18 km SW Baoshan, 2100 m, 16 - 19. v. 2003 leg. S. Murzin ” (CP), 3 ex. “ China (N. Yunnan) Dali Bai Nat. Aut. Pref., 1 km W Dali old town, creek valley at foothill of Diancang Shan, 2170 m, 25 ° 41,9 ’ N / 100 ° 08.4 ’ E (ruderal place) 28. VIII / 1. IX. / 3. IX. 2003 Wrase [18] ” (ZFMK), 1 ex. “ Yunnan 2000 - 2800 m 25.11 N 100.24 E Weibaoshan mts. W slope 25 - 28 / 6.92 Vit Kubáň leg. ” (CN), 1 ex. “ China pr. Yunnan c. occ. Dali-Teiheqiao 12.6.1993 R. Cervenka lgt. ” (ZFMK), 3 ÔÔ “ Micangshan, Hanzhong S. Shanxi, III- 2004, leg. Ying ” (ZFMK), 1 Ô “ Baimianshan Yanling F-Hunugu, 1. X. 2003, leg. Li et al. ” (ZFMK), 1 Ô “ China, E Hubei, 7. - 10. V. Dabie Shan, 31.1 N 115.8 E Wujiashan forest park Jaroslav Turna leg., 2004 ” (ZFMK), 12 ex. “ China, NE Jiangxi, 18. IV. Sanqingshan, 450 - 600 m, 28 ° 52 ’ N, 118 ° 04 ’ E Jaroslav Turna leg., 2006 ” (ZFMK). Uncertain identification (♀): 1 ex. “ China, W. Yunnan Diaolin 18. VI. 1993 E. Jendek & O. Sausa leg. ” (CP), 2 ex. “ Yunnan 1500 - 3200 m 26.07 N 103.14 E Dongchuan 28 / 6 - 3 / 7 Vit Kuban leg. 1994 ” (CP), 1 ex. „ China-Yunnan Dali / 30.5.94 / 2100 - 2500 m Paulus leg. ” (CP), 1 ex. “ Yunnan, 20 - 23. vii. 1996, 27 ° 05 ’ N, 100 ° 13 ’ E, 3400 m, Yulong mts. - Ganhaizi, Vit Kubáň leg. ” (CP), 1 ex. (♀) “ Yunnan, 2300 m Jizu Mts. 18. - 20 Jul. 1995 Bolm lgt. ” (CP), 1 ex. (♀) “ China, Yunnan Lijiang 8. - 24. VI. 1994 leg. B. Šiška & T. Spevár ” (CP), 1 ex. (♀) „ C-China, Shaanxi, Quinling Shan, 6 km N of Foping, 1150 - 1300 m, 20 - 21. VI. leg. C. Holzschuh 2000 ” (CP), 1 ex. (♀) “ China, 1000 - 1300 m, Shaanxi, Quinling mts., Xunyangba (6 km E) 23.5. - 13. vi. 1998 J. H. Marshal leg. ” (CP), 1 ex. (♀) “ China, N. Yunnan, env. Xiaguan, 2400 m, 29. vii. 2002, leg. S. Murzin, J. Shokhin ” (CP), 1 ex. “ Kiautschou China ” (ZMHB).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDDFFE0FF3E70502E04DCDD.taxon	description	Redescription. Length: 5.2 mm, length of elytra: 3.4 mm, width: 3.1 mm. Body oval, black, legs brown, elytra reddish brown with margins and one transversal mesolateral and one anteromedial spot black, dorsal surface dull and nearly glabrous. Labroclypeus transversal, wider than long, shiny, widest at base, lateral margins in basal half almost straight, anteriorly strongly convex and convergent to rounded anterior angles; anterior margin distinctly emarginate medially, margins weakly reflexed; lateral margins and ocular canthus produce a blunt angle; surface flat, coarsely and densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons dull, anteriorly narrowly shiny, with dense, fine punctures, beside eyes and behind frontoclypeal suture with numerous long setae, otherwise with minute setae in punctures. Smooth area in front of eyes convex, as wide as long. Ocular canthus finely densely punctate, wide and moderately long (1 / 3 of ocular diameter), with two terminal setae. Eyes very small, ratio of diameter / interocular width: 0.39. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.2 times as long as than remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide, widest at base, lateral margins in basal half subparallel, anteriorly convex and convergent towards sharp and produced anterior angles; posterior angles blunt. Anterior margin of pronotum moderately convex, with fine but medially widely interrupted marginal line; basal marginal line absent; surface finely and densely punctate, glabrous, with minute setae in punctures; setae of lateral and anterior margins dense, extending on the disc around anterior angles. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, on base along middle narrowly impunctate. Elytra elongate, widest at posterior third, striae weakly impressed, finely and densely punctate; sutural interval and lateral 2 – 3 intervals black, with a continuous black dot at anterior quarter from interval 2 to 4, and on posterior elytra between interval 4 to 7; intervals weakly convex, with fine and sparse punctures, with minute setae in punctures, penultimate lateral interval with a few short setae; epipleural edge fine, ending at rounded external apical angle of elytra, epipleura sparsely setose; apical border narrowly membranous, with a fine rim of short microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.77. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, with dense, coarse punctures but without smooth midline, in punctures with numerous minute setae, with a few longer setae along apical margin. Legs moderately wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; posterior margin smooth ventrally, widened in apical half, posterior margin smooth dorsally, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio of width / length: 1 / 2.94; dorsal margin sharply carinate, with two groups of spines, basal group at shortly before middle, apical one at three quarters of metatibial length, in basal third without further robust setae; lateral face longitudinally convex, finely and sparsely punctate, with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation shallowly concavely truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere little shorter than following two tarsomeres combined, little longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 22 E – G. Habitus: 22 H.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDBFFE2FF3E74F328E0DAC3.taxon	description	Figs. 23 A – D, 32	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDBFFE2FF3E74F328E0DAC3.taxon	materials_examined	Type material examined. Holotype: Ô “ Birmah Ruby Mes. / Doherty / Fry Coll. 1905.100. / 896 Asia Sericini spec. ” (BMNH). Paratypes: 1 Ô, 1 ♀ “ Hte Birmanie Mines des Rubies 1200 - 2300 m Doherty 1890 / Museum Paris ex. Coll. R. Oberthur ” (MNHN, ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDBFFE2FF3E74F328E0DAC3.taxon	description	Description of holotype. Length: 7.2 mm, length of elytra: 5.4 mm, width: 4.4 mm. Body oblong-oval, dark reddish brown, elytra and antennal funiculus yellowish brown, elytral margins and one transversal mesolateral spot dark brown, dorsal surface dull or partly with iridescent shine, nearly glabrous. Labroclypeus little wider than long, shiny, widest at base, lateral margins strongly convex and convergent to convex anterior angles; anterior margin deeply emarginate medially, margins moderately reflexed; lateral margins and ocular canthus produce a blunt angle; surface convex, coarsely and densely punctate, with numerous long, erect setae; frontoclypeal suture finely incised and curved medially. Frons entirely shiny, with irregularly dense, fine punctures, beside eyes with some long setae, otherwise glabrous. Smooth area in front of eyes flat, twice as wide as long. Ocular canthus narrow and moderately long (1 / 3 of ocular diameter), almost impunctate, with a terminal seta. Eyes moderately large, ratio of diameter / interocular width: 0.73. Antenna with ten antennomeres; club composed of four antennomeres, straight, as long as than remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide, widest at base, lateral margins in basal half straight and distinctly convergent, anteriorly moderately convex and convergent towards sharp and produced anterior angles; posterior angles blunt. Anterior margin of pronotum weakly convex, with fine but medially widely interrupted marginal line; basal marginal line absent; surface finely and densely punctate, glabrous, with minute setae in punctures; setae of lateral and anterior margins sparse, not extending on pronotal disc around anterior angles. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, glabrous. Elytra elongate, widest at middle, striae weakly impressed, finely and densely punctate; sutural interval and lateral 2 – 3 intervals dark, dark color on elytra on mesal intervals near scutellum extended, as behind middle on sublateral intervals; intervals weakly convex, with fine and sparse punctures, with minute setae in punctures, penultimate lateral interval with a few short setae; epipleural edge fine, ending at weakly rounded, and almost blunt external apical angle of elytra, epipleura sparsely setose; apical border narrowly membranous, with a fine rim of short microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.59. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium weakly convex and dull, with dense, coarse punctures but without smooth midline, with a few longer setae along apical margin. Legs almost slender and long; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; posterior margin smooth ventrally, not serrate, widened in apical half, posterior margin smooth dorsally, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio of width / length: 1 / 3.55; dorsal margin weakly carinate in apical half, in basal half longitudinally convex, with two groups of spines, basal group at shortly before middle, apical one at three quarters of metatibial length, in basal third without further robust setae; lateral face longitudinally convex, finely and sparsely punctate, sometimes with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation shallowly concavely truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere little shorter than following two tarsomeres combined, distinctly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 23 A – C. Habitus: 23 D.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDBFFE2FF3E74F328E0DAC3.taxon	diagnosis	Diagnosis. Oxyserica bifascipennis Ahrens, Lukic & Liu, new species resembles in external appearance the O. marginata (Brenske, 1898), while in shape of aedeagus it is similar to O. nigropicta (Fairmaire, 1891) and O. diversicornis (Moser, 1915). Oxyserica bifascipennis Ahrens, Lukic & Liu, new species differs from all of them by the larger body size, and, of course, in shape of parameres: the distal margin of phallobase (lateral view) is reflexed, the ventral lobes of parameres which are in the new species as well as in O. nigropicta and O. diversicornis subsymmetric, are longer than half of the phallobase in Oxyserica bifascipennis.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDBFFE2FF3E74F328E0DAC3.taxon	etymology	Etymology. The name of the new species is derived from the combined Latin prefix “ bi ” (two-), and the words “ fascis ” (stripes) and “ pennis ” (wing).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFDBFFE2FF3E74F328E0DAC3.taxon	description	Variation. Length: 7.0 – 7.2 mm, length of elytra: 5.1 – 5.4 mm, width: 4.3 – 4.4 mm. Female: Antennal club composed of three antennomeres, little shorter than the remaining antennomeres combined.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD8FFE3FF3E759B2916DA9B.taxon	description	Figs. 23 E – H, 32	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD8FFE3FF3E759B2916DA9B.taxon	materials_examined	Type material examined. Holotype: Ô “ Myanmar N (Burma) 25 km E Putao, H = 800 m Nan Sa Bon vill. 06 - 09.05.1998 leg. S. Murzin & V. Sinaev ” (ZFMK).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD8FFE3FF3E759B2916DA9B.taxon	description	Description of holotype. Length: 4.7 mm, length of elytra: 3.2 mm, width: 3.2 mm. Body short-oval, light reddish brown, elytra and antennal funiculus yellowish brown, head, antennal club, elytral margins and one transversal mesolateral spot dark brown, dorsal surface dull, nearly glabrous. Labroclypeus trapezoidal, distinctly wider than long, shiny, widest at base, lateral margins weakly convex and strongly convergent to weakly convex, blunt anterior angles; anterior margin weakly emarginate medially, margins moderately reflexed; lateral margins and ocular canthus produce an indistinct angle; surface convex, finely and densely punctate, with a few long, erect setae; frontoclypeal suture finely incised and curved medially. Frons anteriorly narrowly shiny, with dense, fine punctures, beside eyes with some long setae, otherwise glabrous. Smooth area in front of eyes convex, as wide as long. Ocular canthus narrow and short (1 / 4 of ocular diameter), almost impunctate, with a terminal seta. Eyes very small, ratio of diameter / interocular width: 0.38. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.1 times as long as than remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins evenly and moderately convex and convergent towards sharp and produced anterior angles; posterior angles blunt. Anterior margin of pronotum straight, with fine but medially widely interrupted marginal line; basal marginal line absent; surface finely and densely punctate, with minute setae in punctures; setae of lateral and anterior margins sparse, not extending on pronotal disc around anterior angles. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, glabrous. Elytra short, widest at middle, striae weakly impressed, finely and sparsely punctate; sutural interval and epipleura dark, behind middle with a large dark spot from external margin to 5 th interval; intervals flat, with fine and sparse punctures, with minute setae in punctures, penultimate lateral interval with a few short setae; epipleural edge fine, ending at weakly rounded, and almost blunt external apical angle of elytra, epipleura sparsely setose; apical border chitinous, without a rim of microtrichomes (magnification 100 x). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum / metacoxa: 1 / 1.67. Abdominal sternites finely punctate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium moderately convex and dull, with dense, coarse punctures but without smooth midline, with minute setae in punctures, with a few longer setae along apical margin. Legs short and wide; femora finely and sparsely punctate, with two longitudinal rows of longer setae. Metafemur moderately shiny, anterior margin acute, without submarginal serrated line; anterior row of seta-bearing punctures present; posterior margin smooth ventrally, not serrate, strongly widened in apical half, posterior margin smooth dorsally, with a few short setae basally. Metatibia moderately long and wide, widest at middle, ratio of width / length: 1 / 2.7; dorsal margin weakly carinate, with two groups of spines, basal group at one third, apical one at two thirds of metatibial length, in basal third without further robust setae; lateral face longitudinally convex, finely and sparsely punctate, sometimes with minute setae in punctures; ventral edge finely serrate, with three robust equidistant setae; medial face smooth, apex interiorly near tarsal articulation shallowly concavely truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it; first metatarsomere as long as following two tarsomeres combined, distinctly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 23 E – G. Habitus: 23 H.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD8FFE3FF3E759B2916DA9B.taxon	diagnosis	Diagnosis. Oxyserica serena Ahrens, Lukic & Liu, new species resembles in external appearance Oxyserica pygidialis Brenske, 1900, however, the aedeagus is not completely symmetric as in the Oxyserica species group around O. pygidialis. In fact, morphology of aedeagus it more similar to the three previous species, O. nigropicta (Fairmaire, 1891), O. diversicornis (Moser, 1915), and O. bifascipennis Ahrens, Lukic & Liu, new species which differs from all the species of the Oxyserica marginata group by the subsymmetric ventral lobes of parameres.	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
32553F1BFFD8FFE3FF3E759B2916DA9B.taxon	etymology	Etymology. The name of the new species is derived from the combined Latin adjective “ serenus ” (light).	en	Ahrens, Dirk, Liu, Wangang, Lukic, Daniel, Bai, Ming (2023): A taxonomic review of Microserica Brenske, 1894 from continental Asia (Coleoptera: Scarabaeidae: Melolonthinae: Sericini). Zootaxa 5241 (1): 1-115, DOI: 10.11646/zootaxa.5241.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5241.1.1
