taxonID	type	description	language	source
2DFF5087A2A771552CB7E3A0FFDD7EE6.taxon	diagnosis	Diagnosis. Small beetles, body length nearly 2.8 mm. Body shape elongate oval in dorsal view, moderately convex in lateral view, with dorsal outline nearly straight along anterior 2 / 3 of elytra (Fig. 17). Surface of head and pronotum granulate (Fig. 17 C). Body pale / yellowish brown, with head slightly darker. Eyes with anterior margin straight in lateral view (not emarginate), in dorsal view slightly projecting from outline of head (Fig. 17 C). Labrum not concealed by clypeus (Fig. 17 C). Antennae with nine antennomeres, with strongly asymmetric cupule, with longer side acute. Maxillary palps elongate, with palpomere 4 nearly as long as palpomere 3 (d'Orchymont 1943 f: 7, in key). Elytra without sutural striae, narrowly explanate laterally, serial punctures strongly marked, arranged in rows (Fig. 17 A). Prosternum flat, rather sharply carinate medially, with angulate anteromedian projection. Posterior elevation of mesoventrite only weakly bulging. Metaventrite with hydrofuge pubescence. Metafemora without distinct tibial grooves, mostly pubescent, only glabrous at apex. Metatarsomeres 1 - 4 similar in length; metatarsomere 5 similar in length to metatarsomeres 1 - 4 combined. Fifth abdominal ventrite apically emarginate, with stout setae.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
2DFF5087A2A771552CB7E3A0FFDD7EE6.taxon	distribution	Distribution. Afrotropical: Mozambique; Fig. 4.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
6D67DF6FCEFE45AE7C558CAABACCCD82.taxon	diagnosis	Diagnosis. Small beetles, body length 1.4 - 4.8 mm. Body shape elongate to broadly oval in dorsal view, weakly to moderately convex in lateral view, rarely strongly convex (Figs 18, 19). Surface of head and pronotum smooth, usually with shallow ground punctation. Body ranging from pale / yellowish to dark brown (Figs 18, 19), either uniform across body regions or with different regions colored differently (e. g., darker head, paler elytra and margins of pronotum; Fig. 18 A, B). Eyes with anterior margin straight in lateral view (not emarginate), in dorsal view slightly projecting from outline of head. Clypeus moderately convex, with distinct systematic punctures, with anterior margin slightly to clearly emarginate. Labrum not concealed by clypeus. Mentum nearly 1.5 x wider than long, with variable surface, with wide and moderate median anterior depression limited by low transverse carina. Antennae with eight or nine antennomeres, with slightly asymmetric cupule, round in outline. Maxillary palps elongate, 0.7 - 1.5 x width of head, with inner margin of palpomere 2 usually straight and palpomere 4 nearly as long to slightly longer than palpomere 3 (Fig. 12 G). Pronotum with ground punctation usually moderate. Elytra without sutural striae, not laterally explanate, with serial punctures usually absent; systematic punctures usually rather sparse and aligned in four rows along elytra. Prosternum slightly convex, not carinate medially. Posterior elevation of mesoventrite variable, from simply bulged, to bearing variously shaped elevations; anapleural sutures variable in shape and orientation. Metaventrite with posteromedian glabrous patch. Metafemora without distinct tibial grooves, either mostly pubescent (only glabrous at apex), or with pubescence reduced to small basal area (" Gymnhelochares "). Metatarsomere 1 shorter than 2; metatarsomere 2 slightly shorter than 5; metatarsomere 5 similar in length to metatarsomeres 3 and 4 combined. Fifth abdominal ventrite apically emarginate, sometimes very slightly, or rounded, with or without fringe of stout setae. Aedeagus trilobed in form (Fig. 20); basal piece shorter to longer than parameres; outline of apical region of parameres variable; median lobe triangular, with well-developed lateral basal apodemes, usually rounded at apex; gonopore well developed.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
6D67DF6FCEFE45AE7C558CAABACCCD82.taxon	distribution	Distribution. Afrotropical: Angola, Botswana, Cameroon, Democratic Republic of the Congo, Djibouti, Eritrea, Eswatini, Ethiopia (in doubt), Gabon, Ghana, Guinea, Ivory Coast, Kenya, Madagascar, Malawi, Mozambique, Namibia, Nigeria, Oman, Republic of South Africa, Saudi Arabia, Sudan, Tanzania, United Arab Emirates, Yemen, Zimbabwe. Australasian: Australia (New South Wales, Northern Territory, Queensland, Western Australia), Indonesia (Java, Papua), Papua New Guinea. Indo-Malayan: Bhutan, Brunei, China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Himachal, Hong Kong, Hunan, Jiangxi, Taiwan, Yunnan, Zhejiang), India (Arunachal Pradesh, Assam, Goa, Himachal Pradesh, Kerala, Karnataka, Madhya Pradesh, Maharashtra, Meghalaya, North Andaman Island, Sikkim, Tamil Nadu, Uttar Pradesh, Uttarakhand), Laos, Malaysia, Myanmar, Nepal, Philippines, Sri Lanka, Thailand, Vietnam. Palearctic: China (Anhui, Gansu, Hubei, Shaanxi, Shandong, Sichuan, Tibet), Iran, Japan, Korea, Pakistan, South Korea; Fig. 4.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
CBBF4016F7853CE690B58F1281F3EB70.taxon	diagnosis	Diagnosis. Medium sized beetles, total body length 5.8 - 7.5 mm. Body shape elongated oval in dorsal view; weakly convex in lateral view (Fig. 21). Color orange brown to dark brown; ventral surface covered with rather long golden setae, especially on abdominal ventrites, and more densely so (with shorter setae) on surface of femora. Head subquadrate in dorsal view, seemingly constricted at anterior margin of eyes (Fig. 11 J). Eyes relatively small, separated by distance nearly 6.5 x the maximum width of an eye (Fig. 11 J). Clypeus with lateral margins nearly parallel, slightly convex, with anterior margin only slightly narrower than posterior margin (Fig. 11 J). Labrum fully exposed. Mentum and submentum roughly punctate (Fig. 21 C). Antennae with nine antennomeres, with cupule slightly asymmetrical and round in outline. Maxillary palps long, nearly 1.5 x longer than maximum width of head, with inner and outer margins of maxillary palpomere 2 evenly curved (Fig. 21 A). Pronotum with ground punctation shallow and uniformly sparse. Elytra without sutural striae, with outer margins slightly flared; serial punctures, ground punctures and systematic punctures similar in size, shallowly impressed. Surface of prosternum flat (slightly carinate only along midline of antero-mesal projection of anterior margin). Posterior elevation of mesoventrite simple, without carinae or ridges; anapleural sutures concave, anteriorly converging, anteriorly separated by distance nearly 0.3 x as wide as anterior margin of mesepisternum. Metaventrite densely and uniformly pubescent. Protibiae with spines of anterior row very small and appressed (Fig. 21 C); apical spurs of protibiae very short (not exceeding the length of the first tarsomere) and stout. Hydrofuge pubescence covering most surface of metafemora (Fig. 21 C). Ventral face of tarsomeres 1 - 4 densely covered by stiff setae. Apex of fifth abdominal ventrite strongly emarginate; emargination fringed by stout setae. Aedeagus tubular (Fig. 22 A-C), somewhat cylindrical, with parameres forming a 5 - 7 x longer than wide tube; basal piece very short and strongly concave; gonopore reduced, located at apex of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
CBBF4016F7853CE690B58F1281F3EB70.taxon	distribution	Distribution. Neotropical: Brazil (Amazonas, Roraima), French Guiana, Guyana, Suriname, Venezuela; Fig. 4.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
1B49368E86973E9AC7370F8F6DC7C1B5.taxon	diagnosis	Diagnosis. Body length between 3 - 4 mm. Body shape oval in dorsal view, moderately convex in lateral view, with dorsal outline nearly straight along basal 2 / 3 (Fig. 23). Dorsal surfaces smooth, uniformly covered by short setae, brown to pale brown in coloration, either uniform or with yellowish patches along margins of pronotum and elytra, or scattered throughout surface giving spotted appearance (Fig. 23 A, B); ground punctation fine and shallow; ventral surfaces rather densely covered by rather long and fine golden setae. Head rather oval in dorsal view, clearly constricted at anterior margin of eyes (Fig. 11 E). Eyes not emarginate, moderate in size, separated by nearly 3.8 x width of eye, strongly projected from outline of head (Fig. 11 E). Clypeus with anterior margin broadly emarginate, with medial region of emargination nearly straight; anterior corners round. Labrum fully exposed, with apical region anteriorly flattened, thus forming fine transverse carina across anterior region (Fig. 11 D). Mentum rather flat, surface laterally punctate, mesally and anteriorly striate, with anteromedial region depressed. Submentum finely and shallowly punctate. Antennae with nine antennomeres, with strongly asymmetric and round cupule. Maxillary palps nearly 1.5 x longer than maximum width of head, with palpomere 4 0.8 x as long as palpomere 3 (Fig. 23 C); inner margin of maxillary palpomere 2 nearly straight, outer margin apically slightly curved. Pronotum medially evenly convex, explanate and somewhat bending upwards along antero-lateral areas; posterior margin of pronotum clearly narrower than anterior margin of elytra combined. Elytra without sutural striae, with outer margins explanate, especially along anterior third; serial punctures well developed, forming longitudinal rows, at least well defined along outer areas, or visible along entire length of elytra; seta bearing systematic punctures irregularly distributed. Surface of prosternum slightly elevated along midline, with anterior margin acutely triangular and slightly projected anteriorly. Posterior elevation of mesoventrite rather flat; intercoxal process of mesoventrite broad (nearly as wide as antennal club), apically truncate; anapleural sutures sinuate, separated at anterior margin by distance slightly shorter than anterior margin of mesepisternum. Metaventrite with medial surface elevated as platform, densely covered with hydrofuge pubescence, except for posterolateral patches (Fig. 23 C). Protibiae with spines of anterior row very fine and erect; apical spurs of protibiae small (larger spur similar in size and shape to tarsal claws). Metafemora without tibial grooves; metafemora with hydrofuge pubescence covering at least basal 2 / 3 of anterior surface (Fig. 23 C). Metatarsomere 5 1.5 x longer than metatarsomere 2, metatarsomere 2 nearly as long as metatarsomeres 3 and 4 combined; tarsomeres 1 to 4 with sparse long setae on dorsal surface, and spiniform dense setae on ventral surface; tarsomere 5 with few setae along apical margin. Abdomen with five pubescent ventrites. Fifth abdominal ventrite with apex broadly truncate, without stout setae. Aedeagus trilobed, with basal piece nearly as long as parameres (Fig. 22 D); parameres somewhat triangular, slender and apically narrowing; median lobe tapering to round apex; gonopore well-developed.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
1B49368E86973E9AC7370F8F6DC7C1B5.taxon	distribution	Distribution. Afrotropical: Burundi / Rwanda, Central African Republic, Democratic Republic of the Congo, Gabon, Guinea, Kenya, Republic of the Congo, Uganda; Fig. 4.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
98ECB81B738A46C9C860D9572E16A8B5.taxon	diagnosis	Diagnosis. Body length ranging from 2.5 - 5.0 mm. Body shape oval in dorsal view, parallel-sided to broader around midlength, dorsoventrally flattened, weakly to moderately convex in lateral view (Fig. 24), either evenly convex or flattened along anterior half. Surface of head, pronotum and elytra smooth, with usually shallow ground punctation. Coloration ranging from yellowish orange to dark brown, usually uniform along body, sometimes darker on head or only frons. Shape of head trapezoid (Fig. 11 H). Eyes varying in size, usually subquadrate in dorsal view, only very weakly emarginated anteriorly, and usually projected from outline of head. Clypeus trapezoid, with anterior margin mesally weakly to strongly emarginated; membranous preclypeal area visible when clypeus strongly emarginated (Fig. 11 H). Labrum fully exposed, semioval, anteriorly mesally emarginated. Mentum usually rather smooth, with anterior depression often reaching midlength of mentum, sometimes limited by low transverse carina. Antennae with eight antennomeres, with cupule slightly asymmetric and rounded. Maxillary palps usually slender and slightly longer than width of head, with inner margin slightly and evenly curved, and outer margin curved along apical half. Pronotum evenly convex. Elytra with sutural striae, with outer margins slightly flared; ground punctures usually only shallowly marked, serial punctures absent and at least one median row of systematic punctures clearly visible on each elytron (Fig. 24). Surface of prosternum usually flat, only rarely with low medial carina along intercoxal process. Posterior elevation of mesoventrite with an either blunt or sharp longitudinal elevation; anapleural sutures sinuate, separated at anterior margin by distance similar or slightly shorter than anterior margin of mesepisternum. Metaventrite with posteromesal and posterolateral glabrous patches (Fig. 24 C). Protibiae with spines of anterior row semi erect, relatively long, thick and sparse; apical spurs of protibiae moderately long and thick, reaching apex of protarsomere 2. Metafemora with tibial grooves moderately developed, with sharp posterior margin; hydrofuge pubescence covering at least basal 3 / 4 of anterior surface of metafemora (Fig. 24 C, F). Metatarsomeres 2 - 4 with two rows of spiniform setae on ventral surface; metatarsomere 5 nearly as long as 3 and 4 combined; metatarsomere 2 shorter to nearly as long as 5. Apex of fifth abdominal ventrite emarginate, with fringe of flat and stout setae. Aedeagus trilobed (Fig. 25); basal piece shorter to nearly as long as parameres; outline of apical region of parameres variable; sometimes parameres asymmetrical; median lobe triangular, either simple or bearing additional sclerite, with well-developed lateral basal apodemes and gonopore.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
98ECB81B738A46C9C860D9572E16A8B5.taxon	distribution	Distribution. Neotropical: Argentina, Brazil (Amapa, Amazonas, Minas Gerais, Para, Piaui, Rio de Janeiro, Roraima, Sao Paulo), Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Panama, Paraguay, Suriname, Venezuela; Fig. 4.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
E8AD7A7218B6224DFB8CD4C42252E1FD.taxon	etymology	Etymology. From the Latin word colossus, meaning extremely large, in reference to the comparatively large and robust bodies of the members of the genus, combined with the ending chares, expressing affinity with Helochares. Masculine.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
E8AD7A7218B6224DFB8CD4C42252E1FD.taxon	diagnosis	Diagnosis. Body length 8.5 - 14.0 mm. Body shape broadly oval in dorsal view, strongly and uniformly convex in lateral view (Fig. 26). Dorsal surfaces even and smooth, uniformly dark brown (nearly black) in coloration with reddish antennae, palps and tarsi; ground punctation extremely fine and shallow (Fig. 26 A); ventral surfaces rather densely covered by rather long and fine golden setae (Fig. 26 C). Eyes not emarginate, moderate in size, subquadrate in dorsal view, separated by nearly 4 x width of eye, projected from outline of head (Fig. 11 I). Frons with large (and somewhat fused together) systematic punctures along inner margin of eye. Clypeus with anterior margin broadly roundly emarginate. Labrum fully exposed, medially convex (Fig. 11 I). Antennae with nine antennomeres, with strongly asymmetric and round cupule. Maxillary palps slender, slightly longer than maximum width of head, with palpomere 4 0.7 x as long as palpomere 3 (Fig. 11 I). Mentum medially broadly depressed, laterally punctate, mesally and anteriorly striate; sculpture of mentum ranging from shallow to strong. Pronotum evenly convex, and very smooth, with ground punctation very fine and shallow; systematic punctures of pronotum reduced to paired depressions near anterior margin and at midlength of lateral margins. Elytra without sutural striae, with margins slightly flared; serial punctures either absent or only visible along outer lateral area and posterior third of elytra; systematic punctures enlarged, broadly separated longitudinally, forming five rows mostly visible along outer lateral area and posterior third of elytra (Fig. 26 A, B). Surface of prosternum flat to broadly convex, with anterior margin slightly projected anteriorly (Fig. 26 C). Posterior elevation of mesoventrite with broad longitudinal elevation; anapleural sutures concave, anteriorly converging and separated by distance nearly 1 / 3 of anterior margin of mesepisternum. Metaventrite uniformly densely covered by with hydrofuge pubescence, medial surface elevated as platform. Protibiae with anterior row of spines extremely reduced to tiny and scanty, appressed denticles; apical spurs of protibiae large, outer nearly as thick and reaching apex of protarsomere 2. Metafemora with tibial grooves well-developed; metafemora with hydrofuge pubescence covering basal 4 / 5 of anterior surface (Fig. 26 C). Metatarsomeres laterally compressed, metatarsomere 2 longer than 5, metatarsomere 5 nearly as long as 3 and 4 combined; all tarsomeres with rows spiniform setae covering ventral surface. Fifth abdominal ventrite with apex emarginate, with fringe of flat and stout setae. Aedeagus symmetrical, either trilobed (C. satoi; Hebauer 2003 a: fig. 1) or highly modified (Fig. 27 A), with basal piece shorter than parameres; median lobe variable.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
E8AD7A7218B6224DFB8CD4C42252E1FD.taxon	distribution	Distribution. Afrotropical: Benin, Burkina Faso, Cameroon, Democratic Republic of the Congo, Ethiopia, Gabon, Ghana, Guinea, Ivory Coast, Liberia, Malawi, Nigeria, Republic of the Congo, Uganda; Fig. 4.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
691A14D47234D37E43742D58EE233E5F.taxon	diagnosis	Diagnosis. Body length ranging from 2.5 - 4.8 mm. Body shape oval in dorsal view, dorsoventrally slightly flattened, moderately convex in lateral view, with dorsal outline nearly evenly convex (Fig. 28); surface even and smooth, with usually shallow ground punctation (Fig. 28). Coloration usually dark brown seldom yellowish, uniform across body regions. Head trapezoid (Fig. 11 G). Eyes relatively large, at most only slightly emarginated anteriorly, and not or only slightly projected from outline of head. Clypeus trapezoid, with anterior margin mesally emarginate; membranous preclypeal area visible when clypeus strongly emarginated. Labrum fully exposed. Mentum punctate or punctate laterally and medially obliquely striate; medial surface flat to depressed (Fig. 28 C); anteromedial depression sometimes limited by low transverse carina. Antennae with nine antennomeres, with cupule slightly asymmetric and rounded. Maxillary palps slender, 1.2 - 1.5 x longer than width of head; maxillary palpomere 4 nearly 0.7 x length of maxillary palpomere 3; inner margin of maxillary palpomere 2 nearly straight, and outer margin curved along apical half. Pronotum evenly convex. Elytra with sutural striae, with outer margins slightly flared; ground punctures usually only shallowly marked, serial punctures absent and at least one median row of systematic punctures visible on each elytron (Fig. 28). Surface of prosternum usually flat, sometimes tectiform. Posterior elevation of mesoventrite with longitudinal carina; anapleural sutures sinuate, separated at anterior margin by distance similar to slightly shorter than anterior margin of mesepisternum. Metaventrite with posteromesal and posterolateral glabrous patches (Fig. 28 C). Protibiae with spines of anterior row semi erect, relatively long, thick and sparse; apical spurs of protibiae relatively short and stout, not reaching apex of protarsomere 2. Metafemora with tibial grooves moderately developed; hydrofuge pubescence covering basal 4 / 5 of anterior surface of metafemora (Fig. 28 C). Metatarsomeres 2 - 4 gradually decreasing in size, with two rows of spines on ventral surface; metatarsomere 2 slightly longer than 5, 5 shorter than 3 and 4 combined. Fifth abdominal ventrite emarginate at apex, with fringe of flat and stout setae. Aedeagus (Fig. 27 B-D) with parameres at most only fused at base on dorsal surface; median lobe divided in dorsal and ventral plates; dorsal plate sclerotized along margins, medially membranous, membranes with papillae or denticles along apico-medial region; ventral plate as inverted Y, sometimes accompanied by basal median laminar sclerite; basal piece nearly as long as or longer than ventral length of parameres, always noticeable; gonopore not clearly visible.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
691A14D47234D37E43742D58EE233E5F.taxon	distribution	Distribution. Afrotropical: Angola, Benin, Botswana, Burundi, Cameroon, Democratic Republic of the Congo, Gabon, Gambia, Ghana, Guinea, Kenya, Liberia, Madagascar, Mauritius, Mozambique, Namibia, Niger, Nigeria, Rwanda, Senegal, Seychelles (Aldabra), Sierra Leone, Somalia, Republic of South Africa, Sudan, Tanzania, Uganda, Zambia, Zimbabwe. Australasian: Australia (New South Wales, Northern Territory, Queensland), Fiji (Vanua Levu, Viti Levu), New Caledonia, Papua New Guinea. Indo-Malayan: Cambodia, China (Guangdong, Hong Kong, Taiwan, Yunnan), Indonesia (Borneo, Java, Papua, Sulawesi, Sumatra), Laos, Malaysia, Sri Lanka, Thailand, Vietnam. Palearctic: Bosnia, Croatia, Greece, Israel, Italy, Japan, Serbia and Montenegro, Spain, Tunisia, Turkey; Fig. 4.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
27E3B78C69A2531C19096C0C8CD73E9D.taxon	diagnosis	Diagnosis. Small beetles, body length 2.0 - 2.5 mm. Body shape elongated oval in dorsal view; moderately convex in lateral view (Fig. 29). Color orange brown to dark brown. Head trapezoid. Eyes moderate to small, projected from outline of head. Clypeus trapezoid, with anterior margin broadly and roundly emarginate. Labrum fully exposed. Mentum with lateral oblique ridges; anterior median depression marked by transverse carina (Fig. 29 C). Antennae with nine antennomeres, with cupule only slightly asymmetrical and rounded. Maxillary palps moderately long, slightly longer than width of head (Fig. 29 A). Elytra without sutural striae, with outer margins of elytra slightly flared; serial punctures, ground punctures and systematic punctures similar in size and degree of impression, either shallow or rather sharply marked; all punctures seemingly arranged in rows (Fig. 29 A). Prosternum with well-developed median, longitudinal, laminar carina (Fig. 29 C). Posterior elevation of mesoventrite with a strongly produced, anteriorly pointed transverse ridge, longitudinally carinate (Fig. 14 C); anapleural sutures sinuate, separated by distance nearly 0.6 x width of anterior margin of mesepisternum. Metaventrite densely pubescent, except for median and postero-lateral glabrous patches (Fig. 29 C). Protibiae with spines of anterior row long and thick; apical spurs of protibiae short and stout, almost reaching apex of protarsomere 2. Metafemora covered by hydrofuge pubescence along basal 4 / 5 (Fig. 29 C). Metatarsomeres 2 - 4 gradually slightly decreasing in size; metatarsomere 5 slightly longer than 2; ventral coverage of tarsomeres composed of fine and spiniform setae. Fifth abdominal ventrite apically rounded, truncate, or slightly emarginate, without stout setae. Aedeagus trilobate (Fig. 30 A-E); basal piece 0.2 - 0.25 x the length of parameres; median lobe with well-developed lateral basal apodemes, and acute to narrowly rounded apex; parameres nearly as long as median lobe, with outer margins usually sinuate; gonopore situated distad of midlength of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
27E3B78C69A2531C19096C0C8CD73E9D.taxon	distribution	Distribution. Neotropical: Brazil (Minas Gerais, Para), French Guiana, Guyana, Suriname, Venezuela; Fig. 5.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
2B870C4118EE4448ACC5E3B17FD4EB32.taxon	diagnosis	Diagnosis. Small beetles, body length 1.8 - 3.3 mm. Body shape oval in dorsal view, moderate to strongly convex in lateral view (Fig. 31); with ground punctation usually moderately marked. Color yellowish brown to dark brown, usually uniform across body regions (Fig. 31). Shape of head trapezoid. Eyes relatively small, at most only slightly emarginated anteriorly, usually moderately projected from outline of head. Clypeus trapezoid, with anterior margin from broadly to only slightly emarginate. Labrum fully exposed. Mentum with strong median anterior depression sometimes limited by low transverse carina; surface of mentum mostly smooth and undulated. Antennae with nine antennomeres; cupule slightly asymmetric, with rounded outline. Maxillary palps short, nearly 2 / 3 width of head, and stout (Fig. 31 C); inner margin of maxillary palpomere 2 nearly straight, outer margin strongly curved along apical half. Elytra without sutural striae, and only rarely with impressed striae; ground punctures moderate to sharply marked, uniformly and rather densely distributed; systematic punctures slightly larger and deeper than remainder punctures; serial punctures usually not clearly differentiated; outer margins of elytra only slightly flared (Fig. 31 A, D). Prosternum flat, sometimes only slightly elevated along longitudinal midline (Fig. 31 C). Posterior elevation of mesoventrite either with transverse ridge, or with well-developed tooth that extends anteriorly as longitudinal carina; anapleural sutures concave, separated at anterior margin by distance nearly 0.3 x anterior margin of mesepisternum. Metaventrite densely pubescent, except for large median teardrop-shaped glabrous patch (Fig. 31 C, F); anteromedian area of metaventrite with a deep and narrow transverse depression before anterior intercoxal process. Protibiae with spines of anterior row hair-like, semi erect, relatively long and thick (Fig. 31 C). All tarsomeres bearing long apical hair-like setae on dorsal face, and two lateral rows of hair-like spines on ventral face of tarsomeres 2 - 4. Posterior femora mostly glabrous, with few scattered setae along basal half to basal 2 / 3, with hydrofuge pubescence along anterodorsal margin (Fig. 31 C, F); tibial grooves well-developed, sometimes covered by hydrofuge pubescence. Fifth abdominal ventrite apically truncate, with stout setae. Aedeagus trilobed (Fig. 30 F-I), with outer margins convex, straight or sinuate, with basal piece 0.45 - 0.9 x length of parameres; median lobe somewhat triangular in shape, with well-developed lateral basal apodemes; apex of median lobe widely to narrowly acute, sometimes " pinched "; parameres nearly as long as median lobe; well-developed gonopore, preapically situated.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
2B870C4118EE4448ACC5E3B17FD4EB32.taxon	distribution	Distribution. Neotropical: Brazil (Bahia, Minas Gerais); Fig. 5.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
02E38F608953FAA3FF90B1ED37DD0E62.taxon	diagnosis	Diagnosis. Small beetles, body length 1.9 - 2.3 mm. Body shape rounded in dorsal view, strongly convex in lateral view (Fig. 32). Surface of head, pronotum and elytra smooth, with moderate to shallow ground punctation. Coloration yellow to dark brown, uniform along body, with paler mouthparts and tarsi (Fig. 32). Shape of head relatively oval. Eyes relatively small, anteriorly emarginated (Fig. 32 B), not projected from outline of head. Clypeus trapezoid, with anterior margin mesally broadly emarginate. Labrum fully exposed. Mentum with anterior depression limited by low transverse carina; surface of mentum only slightly striate. Antennae with eight antennomeres, with cupule only slightly asymmetric and rounded in outline. Maxillary palps slender, slightly shorter than width of head (Fig. 32 C). Pronotum evenly convex. Elytra without sutural or other distinct striae, with outer margins slightly flared; elytral ground punctation shallow to moderate, uniformly distributed (Fig. 32). Surface of prosternum flat. Mesoventrite with transverse ridge, usually elevated medially into acute tooth (Fig. 32 C); anapleural sutures concave, separated at anterior margin by distance nearly as width of anterior margin of mesepisternum. Metaventrite uniformly covered by hydrofuge pubescence, with small, longitudinal posteromesal glabrous patch, and reduced posterolateral glabrous patches (Fig. 32 C). Protibiae with spines of anterior row long, thick, semi erect and sparse; apical spurs of protibiae short and of moderate thickness. Metafemora with moderate tibial grooves; hydrofuge pubescence covering basal 4 / 5 of anterior surface (Fig. 32 C). Tarsomeres 1 - 4 ventrally with rows of long and thick setae. Metatarsomeres 2 - 4 gradually decreasing in size, 5 nearly as long as 2 - 4 combined. Fifth abdominal ventrite with small truncation at apex, with fringe of flat and stout setae. Aedeagus trilobed (Fig. 30 J); with short basal piece, less than 1 / 3 length of parameres; median lobe wider than width of parameres; gonopore well differentiated.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
02E38F608953FAA3FF90B1ED37DD0E62.taxon	distribution	Distribution. Neotropical: Brazil (Amazonas, Para), Colombia, Guyana, Suriname, Venezuela; Fig. 5.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
A31E3BC7EE538638DD8BBA79A4B08822.taxon	diagnosis	Diagnosis. Medium sized beetles, body length 4 - 7 mm. Body shape oval in dorsal view, dorsoventrally flattened, with dorsal outline nearly straight along medial third in lateral view (Fig. 33); surface even and granulate. From yellowish, orange brown to dark brown in coloration, usually with patterns along elytra, with different areas of head and pronotum darkened. Shape of head somewhat trapezoid (Fig. 11 L). Anterior corners of frons extended laterally and posteriorly, emarginating anterior margin of eyes. Eyes of moderate size, somewhat oval, anteriorly deeply emarginated, not projected from outline of head. Clypeus somewhat pentagonal, laterally explanate, with anterior margin usually straight (Fig. 11 L). Labrum concealed by clypeus (Fig. 11 L). Mentum with surface variably sculptured, usually with oblique and transverse striae (Fig. 33 C). Antennae with eight antennomeres, with cupule strongly asymmetric and oval in outline. Maxillary palps slender, slightly longer than greatest width of head; inner margin of maxillary palpomere 2 weakly and evenly curved, and outer margin weakly curved along apical third (Fig. 33 C). Pronotum with surface of lateral areas flat. Elytra without sutural striae, with outer margins laterally explanate; serial punctures clearly aligned in longitudinal rows (Fig. 33 A). Scutellar shield U-shaped. Surface of prosternum flat, to medially bulging, smooth to irregularly sculptured. Posterior elevation of mesoventrite only weakly bulging, with pair of lateral, longitudinal, low ridges; anapleural sutures nearly parallel along anterior section, separated anteriorly by distance slightly shorter than anterior margin of mesepisternum. Metaventrite uniformly covered by hydrofuge pubescence, with medial, narrow, and slightly carinate glabrous patch; posterolateral glabrous patches reduced. Protibiae with spines of anterior row short and semi erect; apical spurs of protibiae reduced, much shorter than protarsomere 1. Metafemora with tibial grooves moderately developed; hydrofuge pubescence covering 5 / 6 of anterior surface (Fig. 33 C). Tarsomeres 1 - 4 ventrally densely covered by setae; metatarsomere 2 longer than 3 and 4 combined, 1 nearly as long as 3, and 5 nearly as long as 2 - 4 combined. Fifth abdominal ventrite apically emarginate, with fringe of flat and stout setae. Aedeagus divided (Fig. 34), parameres separated from each other for most of their lengths; median lobe divided in dorsal and ventral plates; dorsal plate usually strongly sclerotized; ventral plate bilaterally bifurcated, forming thick lateral lobes along apical region; basal piece nearly 0.2 x the length of parameres, always noticeable; gonopore not clearly visible.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
A31E3BC7EE538638DD8BBA79A4B08822.taxon	distribution	Distribution. Nearctic: United States (California, Florida, Louisiana, Mississippi, North Carolina, South Carolina, Texas, Virginia). Neotropical: Argentina, Bolivia, Brazil (Amazonas, Ceara, Mato Grosso, Mato Grosso do Sul, Para, Rio de Janeiro, Roraima), Cuba, Guatemala, Mexico, Paraguay, Suriname, Venezuela; Fig. 5.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
BD3F78871BF9B0EBFB01ED802A043F68.taxon	diagnosis	Diagnosis. Small to medium sized beetles, body length 2 - 7 mm. Body shape oval in dorsal view; slightly to moderately convex in lateral view, with dorsal outline nearly flat along anterior half of elytra, or somewhat evenly curved (Figs 35, 36). Coloration usually yellowish brown, sometimes orange brown, pale brown to medium brown; ground punctation shallow (e. g., Fig. 35 D) to strongly marked (e. g., Fig. 36 D). Shape of head trapezoid to oval (e. g., Fig. 11 F). Eyes medium sized to large, not or moderately emarginated anteriorly, usually projected from outline of head. Clypeus trapezoid, with anterior margin broadly and roundly emarginate; sometimes lateral margins of clypeus slightly bent upwards. Labrum fully exposed. Mentum rather flat, sparsely punctate, coarsely to shallowly, rarely striate (e. g., Figs 35 C, 36 C); median anterior depression of mentum relatively shallow; submentum shallowly punctate to smooth. Antennae with nine antennomeres; cupule strongly asymmetric, with rounded outline; antennomere 9 slightly, to 3 x longer than antennomere 7. Maxillary palps slender, moderately long, 0.6 - 1.2 x the width of head (e. g., Figs 35 C, 36 C); inner margin of maxillary palpomere 2 weakly and evenly curved to nearly straight, outer margin evenly curved to curved along apical 2 / 3; maxillary palpomere 3 slightly longer than 4. Prosternum flat to medially bulging to tectiform. Elytra without sutural striae, with ground punctures usually moderately marked; often with serial punctures forming ten longitudinal rows along elytra (e. g., Fig. 35 A). Posterior elevation of mesoventrite, flat to simply bulging (e. g., Fig. 35 C); bulge usually with long fine setae; anapleural sutures strongly concave, nearly parallel along anterior section, separated anteriorly by distance 0.6 - 1.0 x anterior margin of mesepisternum. Metaventrite densely covered by hydrofuge pubescence, without glabrous patches (e. g., Figs 35 C, 36 C). Protibiae with spines of anterior row either nearly absent (e. g., Fig. 35 C) or as long thick semi-erect setae. Metafemora with tibial grooves weakly developed to absent; hydrofuge pubescence covering basal 6 / 7 of anterior surface. Tarsomeres 1 - 4 with pair of lateral rows of long fine spines on ventral face, sometimes ventral face densely covered by hair-like spines; tarsomere 5 with medial row of long fine spines; metatarsomeres variable in proportions (2 - 4 gradually decreasing in size with 5 nearly as long as 3 and 4 combined; 2 and 5 similar in length, each slightly longer than 3 and 4 combined). Fifth abdominal ventrite apically emarginate, with fringe of stout setae. Aedeagus tubular (Fig. 37 A-H); parameres fused to each other for most of their lengths, with apex either simple or bifurcate / bilobate; median lobe with very long basal apodemes (as long or longer than main piece of median lobe), often extending beyond base of parameres in repose; median lobe either simple (without subdivisions), or with multiple and different kinds of sclerotizations of inner membranes; basal piece usually much shorter than parameres; gonopore of variable development, usually visible when median lobe is simple.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
BD3F78871BF9B0EBFB01ED802A043F68.taxon	distribution	Distribution. Afrotropical: Angola, Benin, Botswana, Burkina Faso, Burundi, Cameroon, Chad, Democratic Republic of the Congo, Eritrea, Ethiopia, Gabon, Gambia, Ghana, Guinea, Guinea Bissau, Ivory Coast, Kenya, Liberia, Madagascar, Malawi, Mali, Mauritania, Mauritius (incl. Mascarene Is., Rodrigues), Morocco [in doubt], Mozambique, Namibia, Niger, Nigeria, Oman, Republic of the Congo, Reunion, Rwanda, Sao Tome and Principe, Saudi Arabia, Senegal, Seychelles (incl. Aldabra), Sierra Leone, Republic of South Africa, South Sudan, Sudan, Tanzania, Togo, Uganda, United Arab Emirates, Yemen (incl. Socotra), Zambia, Zimbabwe. Australasian: Australia (Australian Capital Territory, New South Wales, Northern Territory, Queensland, South Australia, Tasmania, Victoria, Western Australia), Fiji, Papua New Guinea (incl. Duke of York), Vanuatu. Indo-Malayan: Bangladesh, Burma, Cambodia, China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan, Jiangxi, Macao, Taiwan, Yunnan, Zhejiang), India (Andaman Is., Assam, Bihar, Karnataka, Madhya Pradesh, Nicobar Is., Uttarakhand, Uttar Pradesh, Tamil Nadu, West Bengal), Indonesia (Bali, Borneo, Java, Lombok, Papua, Sumatra), Laos, Malaysia (Peninsula, Sabah), Nepal, Philippines (Manila), Singapore, Sri Lanka, Thailand, Vietnam. Nearctic: U. S. A. (Alabama, Arkansas, Arizona, California, Delaware, District of Columbia, Florida, Georgia, Illinois, Indiana, Iowa, Kansas, Kentucky, Louisiana, Maryland, Mississippi, Missouri, Nevada, North Carolina, North Carolina, Ohio, Oklahoma, Oregon, Pennsylvania, South Carolina, Tennessee, Texas, Virginia). Neotropical: Costa Rica, Ecuador, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Peru, Venezuela. Oceanian: Samoa, Tonga. Palearctic: Algeria, Austria, Azerbaijan, Belarus, Bosnia Herzegovina, Bulgaria, Canary Islands, China (Chongqing, Jilin, Hubei, Jiangsu, Shanghai, Shaanxi, Sichuan, Xinjiang, Xizang [Tibet]), Croatia, Czech Republic, Denmark, Egypt, Estonia, Finland, France, Germany, Georgia, Great Britain, Greece, Hungary, Iran, Iraq, Ireland, Israel, Italy, Japan, Latvia, Lebanon, Lithuania, Luxembourg, Macedonia, Morocco, Netherlands, Norway, Pakistan, Poland, Portugal, Russia, Serbia and Montenegro, Slovakia, Slovenia, South Korea, Spain, Sweden, Switzerland, Syria, Tunisia, Turkey, Ukraine; Fig. 5.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
C46AFB08B3766F94741F6E8A1724ED56.taxon	diagnosis	Diagnosis. Small beetles, body length nearly 3.5 mm. Body broadly oval and explanate in dorsal view, rather flat in lateral view, with dorsal outline nearly straight along median region (Fig. 38). Surface smooth (without granulations or reticulations), with ground punctation strongly marked. Body orange brown, slightly paler along margins (Fig. 38). Shape of head somewhat trapezoid. Anterior corners of frons extended laterally and posteriorly, emarginating anterior margin of eyes. Eyes relatively small, with anterior margin markedly emarginate in lateral view, in dorsal view not projecting from outline of head. Clypeus laterally expanded in front of eyes; anterior margin of clypeus slightly emarginate. Labrum concealed under clypeus. Mentum with surface obliquely striate (Fig. 38 C). Antennae with nine antennomeres, cupule strongly asymmetric, with rounded outline. Maxillary palps short and moderately stout, hardly 3 / 4 as long as width of head; maxillary palpomere 4 nearly as long as palpomere 3; inner margin of maxillary palpomere 2 nearly straight, outer margin curved along apical half (Fig. 38 C). Elytra without sutural striae, broadly explanate laterally, serial punctures absent, ground punctures sharply marked, densely and uniformly distributed (Fig. 38 A). Prosternum slightly convex, not carinate medially (Fig. 38 C). Posterior elevation of mesoventrite only bulging (Fig. 38 C); anapleural sutures only slightly concave, separated at anterior margin by distance similar to anterior margin of mesepisternum. Metaventrite uniformly covered by hydrofuge pubescence (Fig. 38 C). Protibiae with spines of anterior row long, thick, and semi-erect; apical spurs of protibiae stout, extending to apex of protarsomere 2. Metafemora without distinct tibial grooves; hydrofuge pubescence covering basal 3 / 4 of anterior surface of metafemora (Fig. 38 C). Tarsomeres 2 - 4 ventrally densely covered by setae; metatarsomere 1 much shorter than 2; metatarsomere 5 nearly as long as metatarsomere 2 or 3 and 4 combined. Fifth abdominal ventrite apically emarginate, with fringe of flat and stout setae. Aedeagus tubular (Fig. 37 I); distal region of each paramere diverging; apex of parameres rounded; basal piece nearly half as long as parameres; median lobe broad, apically tapering to rounded tip; gonopore not clearly visible.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
C46AFB08B3766F94741F6E8A1724ED56.taxon	distribution	Distribution. Indo-Malayan: Myanmar (formerly Burma); Fig. 5.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
A58E2BA21C04E7D1257E011A2B9DBB0C.taxon	diagnosis	Diagnosis. Medium to small beetles, body length 2.7 - 4.5 mm. Body shape oval to elongated in dorsal view; moderately and evenly convex in lateral view (Fig. 39). Color orange brown to dark brown, rather uniform along body regions (Fig. 39). Shape of head trapezoid. Eyes relatively small, subquadrate, at most only slightly emarginated anteriorly, moderately projected from outline of head. Clypeus trapezoid, with anterior margin broadly emarginate. Labrum fully exposed. Mentum with strong median anterior depression sometimes limited by low transverse carina; surface of mentum with lateral oblique ridges (Fig. 39 C, F). Antennae with nine antennomeres; cupule slightly asymmetric, with rounded outline. Maxillary palps moderately long, 0.7 x to nearly as long as width of head; inner margin of maxillary palpomere 2 slightly curved near apex, outer margin curved, sometimes strongly, along apical half (Fig. 39 C, F). Each elytron with five rows of deep / large systematic punctures; elytra without sutural striae, with outer margins slightly flared; serial punctures absent (Fig. 39 A, D). Prosternum slightly convex to tectiform. Posterior elevation of mesoventrite, with a well-defined, curved transverse ridge; anapleural sutures forming an obtuse angle, separated at anterior margin by distance 0.2 - 0.3 x the width of anterior margin of mesepisternum. Metaventrite densely pubescent, except for large median rhomboid glabrous patch (Fig. 39 C, F). Protibiae with spines of anterior row hair-like, semi erect, relatively long and thick. All tarsomeres bearing long apical hair-like setae on dorsal face, and hair-like spines on ventral face of tarsomeres 2 - 4. Posterior femora glabrous at most along apical third (Fig. 39 C, F). Fifth abdominal ventrite apically truncate to slightly emarginate, with fringe of stout setae. Aedeagus trilobed (Fig. 40 A-D), nearly parallel sided, with basal piece between 0.5 and 1.1 x length of parameres; median lobe wider than each paramere, gradually narrowing apically, with Peltochares conspicuous median longitudinal sclerotization, and well-developed lateral basal apodemes; apex of median lobe acute; parameres nearly as long as median lobe, with apical setae; gonopore preapically situated.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
A58E2BA21C04E7D1257E011A2B9DBB0C.taxon	distribution	Distribution. Neotropical: Ecuador, Peru, Venezuela; Fig. 5.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
4E32588B5754A0ABA237203807BC067B.taxon	diagnosis	Diagnosis. Very small beetles, body length 1.15 - 1.45 mm. Body shape oval in dorsal view; slightly to moderately, and evenly convex in lateral view (Fig. 41). Coloration uniformly brown, to variable along the body; ground punctation shallow to moderately marked (Fig. 41). Shape of head trapezoid and relatively wide. Eyes moderate in size, slightly emarginated anteriorly, not projected from outline of head. Clypeus trapezoid, with anterior margin broadly emarginate. Labrum fully exposed. Mentum with lateral oblique ridges. Antennae with eight antennomeres; cupule slightly asymmetric, with rounded outline. Maxillary palps slender, moderately long nearly 0.7 x the width of head; inner margin of maxillary palpomere 2 nearly straight, outer margin curved along apical half (e. g., Fig. 41 C, F). Each elytron with ground punctures usually only shallowly marked, seemingly forming longitudinal rows, with irregularly distributed systematic punctures bearing rather long setae, denser along lateral and posterior regions; elytra without sutural striae. Prosternum flat, at most only weakly convex. Posterior elevation of mesoventrite, usually projected as low and short longitudinal carina between mesocoxae; anapleural sutures only weakly curved, separated at anterior margin by distance nearly 0.9 x width of anterior margin of mesepisternum. Metaventrite with posterolateral and mesal glabrous patches (e. g., Fig. 41 C, F). Protibiae with spines of anterior row hair-like, semi erect, relatively long, thick and sparse. Metafemora mostly densely covered by hydrofuge pubescence (e. g., Fig. 41 C, F). All tarsomeres with long and thick spines on ventral faces of tarsomeres 2 - 4; metatarsomeres 2 - 4 gradually decreasing in size, metatarsomere 5 as long as 3 and 4 combined, 2 slightly shorter. Fifth abdominal ventrite apically emarginate, with fringe of stout setae. Aedeagus trilobed (Fig. 40 E-H), nearly parallel sided, with basal piece 0.3 - 0.6 x length of parameres; median lobe with well-developed lateral basal apodemes, wider at base than base of each paramere, usually narrower at apex than preapical width of parameres; apex of median lobe rounded; parameres from slightly shorter to longer than median lobe, and only narrowing at apex; gonopore situated beyond midpoint of median lobe.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
4E32588B5754A0ABA237203807BC067B.taxon	distribution	Distribution. Neotropical: Brazil (Para), Guyana, Suriname; Fig. 5.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
4C3858378807689BD015E34B5CDD4933.taxon	diagnosis	Diagnosis. Body length 1.2 - 14.0 mm. Body shape oval in dorsal view, dorsoventrally flattened, or weakly to strongly convex in lateral view (Fig. 1); surface even (without elevations or depressions), granulate (e. g., Figs 17, 33) or smooth on head and pronotum. From yellowish to dark brown in coloration (Fig. 1), usually uniform, sometimes different regions of body colored differently. Shape of head variable (trapezoid, subquadrate, round; Fig. 11 E-L). Anterior corners of frons sometimes extended posteriorly forming Tobochares canthus and emarginating anterior margin of eyes (e. g., Tobochares, Helobata; e. g., Fig. 11 B, C). Eyes varying in size, shape, degree of emargination, and degree of projection from outline of head (Fig. 11 E-L); absent only in cavernicolous genus Troglochares Spangler, 1981 a. Clypeus variable in shape (rectangular to trapezoid; Fig. 11 E-L), with anterior margin from straight to mesally emarginate. Labrum usually exposed; concealed by clypeus in Helobata (Fig. 11 L) and Helopeltarium (Fig. 1 H). Mentum usually wider than long, with strong median anterior depression, may be limited by low transverse carina (Fig. 12 A-C); surface of mentum with variable sculpture, ranging from smooth (Fig. 12 A) to roughly punctate or obliquely striate (Fig. 12 B). Antennae with eight or nine antennomeres (Fig. 12 D, E), with cupule varying in symmetry and shape. Maxillary palps curved inward, ranging from very short (nearly half width of the head; e. g., Quadriops reticulatus, Fig. 12 C) and stout, to very long and slender (nearly twice the width of the head; e. g., Peltochares, Fig. 11 K). Pronotum evenly convex, usually with systematic punctures forming paired anterolateral semicircles and paired short posterolateral transverse bands. Elytra with or without sutural striae, with outer margins simple, slightly flared, or laterally explanate; elytral punctation variable (Fig. 13). Hind wings usually well developed (Fig. 15 A-F), seldom reduced along apical region (Fig. 15 G). Surface of prosternum flat (e. g., Fig. 14 A, B), convex or rarely medially carinate (e. g., Crucisternum; Fig. 29 C), with anterior margin straight or anteriorly projected. Posterior elevation of mesoventrite either only weakly bulging or with transverse (e. g., Fig. 14 E, G) or longitudinal ridge (e. g., Fig. 14 D, F); with strongly produced, anteriorly pointed and longitudinally carinate transverse ridge in Crucisternum (Fig. 14 C). Anapleural sutures variable in shape and orientation. Metaventrite rather uniformly covered by hydrofuge pubescence (e. g., Fig. 14 E), sometimes with posteromesal glabrous patch (e. g., Fig. 14 D, F, G), sometimes also with posterolateral glabrous patches (e. g., Fig. 14 D). Protibiae with anterior row of spines varying in shape and development; apical spurs of protibiae varying in development. Metafemora with tibial grooves of varying development; hydrofuge pubescence on anterior surface of metafemora absent, reduced to only basal or dorsal patch, or increasingly covering most of surface. Tarsomeres 5 - 5 - 5; tarsomeres variable in size, proportions, and dorsal and ventral coverage. Abdomen with five pubescent ventrites, density of setae ranging from sparse to very dense. Fifth abdominal ventrite with apex either rounded (Fig. 15 I), truncate (Fig. 15 J), or emarginate (Fig. 15 H); apex with or without fringe of flat and stout setae. Aedeagus usually symmetrical (Fig. 16), with basal piece varying in size from longer than parameres (e. g., Primocerus, Fig. 47; Batochares, Fig. 22 D), to reduced and virtually absent (e. g., Peltochares, Fig. 45); parameres highly variable in shape, either slender and only connected to each other at base of ventral surface (e. g., Fig. 16 A-D, G, H), or fused together forming tube-like structure (e. g., Fig. 16 E, F); apex of parameres either simple, or bifurcated and modified with hooks and spines (e. g., Fig. 16 C, D); median lobe either simple or with dorsal and ventral lobes, with well-developed lateral basal apodemes; further modifications (longitudinal divisions, presence of internal hooks and spines, development of gonopore) widespread.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
860DF4E4B70AE877796325D295B04CDA.taxon	etymology	Etymology. From the Latin word novus, meaning new, in reference to the genus being restricted to the New World, combined with the ending chares, expressing affinity with Helochares. Masculine.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
860DF4E4B70AE877796325D295B04CDA.taxon	diagnosis	Diagnosis. Medium sized beetles, body length 4.5 - 9.0 mm. Body shape oval in dorsal view; slightly to moderately convex in lateral view, with dorsal outline nearly flat along anterior half of elytra, or somewhat evenly curved (Fig. 42). Coloration usually uniformly dark brown, sometimes orange or pale brown; ground punctation shallow to moderately marked (Fig. 42). Shape of head trapezoid. Eyes relatively large, not emarginated anteriorly, usually projected from outline of head. Clypeus trapezoid, with anterior margin broadly and roundly emarginate. Labrum fully exposed. Mentum with lateral longitudinal crenulations, lateral oblique ridges, and transverse crenulations along antero-medial area (Fig. 42 C, F). Antennae with nine antennomeres; cupule strongly asymmetric, with rounded outline; antennomere 9 slightly to 2 x longer than antennomere 7. Maxillary palps slender, moderately long, 1.1 - 1.5 x the width of head; inner margin of maxillary palpomere 2 weakly and evenly curved to nearly straight, outer margin evenly curved or curved along apical half; maxillary palpomere 3 slightly longer than 4 (Fig. 42 C, F). Prosternum flat to weakly convex. Elytra without sutural striae, with ground punctures usually shallowly marked; usually at least one row of systematic punctures visible along midline of each elytron; serial punctures sometimes visible along posterior half of elytra (e. g., Fig. 42 D). Posterior elevation of mesoventrite, usually simply bulging, sometimes bulge impressed posteriorly, sometimes bulge extends anteriorly as low, shiny, and glabrous longitudinal ridge; anapleural sutures concave, separated at anterior margin by distance 0.6 - 0.9 x the width of anterior margin of mesepisternum. Metaventrite with medial glabrous patch, sometimes very narrow and extending along entire length of metaventrite (e. g., Fig. 42 C, F). Protibiae with spines of anterior row extremely reduced to tiny appressed denticles. Metafemora with tibial grooves well developed; hydrofuge pubescence covering basal 6 / 7 of anterior surface. Tarsomeres 1 - 4 with long, thick, and rather dense setae on ventral face, sometimes with only rows of short spines on metatarsomeres 2 - 4; metatarsomere 2 as long or slightly longer than 5 and as 3 and 4 combined. Fifth abdominal ventrite apically emarginate, with fringe of stout setae. Aedeagus divided (Fig. 43); parameres separated from each other for most of their lengths; median lobe divided in dorsal and ventral plates; dorsal plate usually strongly sclerotized and elongated, often bifurcated or otherwise shaped along apical region; ventral plate sometimes reduced, usually simple and of variable length; basal piece 0.3 x or less than length of parameres, usually clearly noticeable; gonopore usually clearly visible.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
860DF4E4B70AE877796325D295B04CDA.taxon	distribution	Distribution. Nearctic: U. S. A. (Florida; thought to be introduced). Neotropical: Argentina, Belize, Bolivia, Brazil (Amazonas, Espirito Santo, Mato Grosso, Mato Grosso do Sul, Minas Gerais, Pernambuco, Piaui, Rio de Janeiro, Sao Paulo), Colombia, Costa Rica, Cuba, Ecuador, French Guiana, Guatemala, Lesser Antilles (Grenada, Guadeloupe, St. Vincent), Mexico, Panama, Paraguay, Suriname, Uruguay, Venezuela; Fig. 6.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
FF7F93BC9EC80936914DF9E2CB86ECE3.taxon	diagnosis	Diagnosis. Body length 6 - 14 mm. Body shape oval in dorsal view, weakly to moderately convex in lateral view (Fig. 44). Dorsal surfaces even and smooth, either uniformly covered by short setae (Fig. 44 A), or with scarce long setae along particular areas of surface (associated with systematic punctures; Fig. 44 D), dark brown in coloration, usually uniform; ground punctation fine and shallow to moderate; ventral surfaces densely covered by fine golden setae (Fig. 44 C, F). Head subquadrate (Fig. 11 K). Eyes not emarginate, moderate in size, subquadrate, separated by 4.5 - 5.5 x width of eye, strongly projected from outline of head. Clypeus with anterior margin broadly emarginate, either roundly or acutely, sometimes further medially notched; membranous preclypeal area visible when clypeus strongly emarginated. Labrum fully exposed, often medially convex. Antennae with nine antennomeres, with moderately asymmetric and round cupule; antennomere 9 slightly to 2 x longer than antennomere 7. Maxillary palps slender, 1.3 - 1.8 x longer than maximum width of head, with palpomere 4 nearly 0.8 x as long as palpomere 3; maxillary palpomere 2 with inner margin slightly and evenly curved, and outer margin curved along apical half (Fig. 44 C, F). Mentum slightly depressed mesally, surface laterally punctate, mesally and anteriorly striate, with anteromedial region depressed (Fig. 44 C, F). Submentum punctate to crenulate. Pronotum evenly convex, usually with systematic punctures forming distinct anterolateral semicircles. Elytra without sutural striae, with margins usually only slightly flared (explanate in P. conspicuus; Fig. 44 A); serial punctures usually absent (visible along entire length of elytra in P. conspicuus; Fig. 44 A); ground punctation usually shallow (moderate to strongly marked in P. foveicollis). Surface of prosternum flat to broadly convex, with anterior margin roundly projected anteriorly (Fig. 44 C, F). Posterior elevation of mesoventrite usually with longitudinal or somewhat longitudinal elevation, sometimes forming acute posterior point; apical region of elevation usually with long fine setae; anapleural sutures forming obtuse angle, nearly parallel along anterior section, separated anteriorly by distance 0.3 - 0.7 x anterior margin of mesepisternum. Metaventrite densely covered by hydrofuge pubescence, except for posterolateral patches (Fig. 44 C, F). Protibiae with anterior row of spines reduced to extremely reduced (Fig. 44 C); apical spurs of protibiae stout, ranging from very large (larger spur considerably larger and thicker than tarsal claws, e. g., P. foveicollis), or very short (barely reaching apex of protarsomere 1, e. g., P. conspicuus); pro- and mesotarsal claws are sexually dimorphic in some species (e. g., P. foveicollis). Metafemora with tibial grooves sharply marked; metafemora with hydrofuge pubescence covering at least basal 3 / 4 of anterior surface (Fig. 44 C, F). Metatarsomeres 5 and 2 similar in length or 2 slightly longer, metatarsomere 2 slightly longer than metatarsomeres 3 and 4 combined; all tarsomeres with ventral surface rather densely covered by long spiniform setae on ventral surface (sparser on tarsomere 5). Abdomen with five pubescent ventrites. Fifth abdominal ventrite with apex emarginate, fringed by stout setae. Aedeagus spiked (Figs 16 C, D, 45); main component of median lobe strongly sclerotized, slender, and apically acute, usually accompanied by additional shorter slender sclerotizations; apical region of parameres usually partly heavily sclerotized and partly membranous, often bifurcated; basal piece strongly reduced; gonopore usually not clearly visible.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
FF7F93BC9EC80936914DF9E2CB86ECE3.taxon	distribution	Distribution. Afrotropical: Angola, Benin, Botswana, Burkina Faso, Burundi, Cameroon, Central African Republic, Chad, Democratic Republic of the Congo, Ethiopia, Gabon, Gambia, Ghana, Guinea, Ivory Coast, Kenya, Liberia, Madagascar, Malawi, Mozambique, Namibia, Niger, Nigeria, Republic of the Congo, Rwanda, Senegal, Sierra Leone, Somalia, Republic of South Africa, South Sudan, Tanzania, Togo, Uganda, Western Sahara, Zambia, Zimbabwe. Australasian: Australia (Australian Capital Territory, New South Wales, Northern Territory, Queensland, Western Australia), Indonesia (Papua), New Caledonia, Papua New Guinea. Indo-Malayan: Bangladesh, Cambodia, China (Guangdong, Guangxi, Guizhou, Hong Kong, Jiangxi, Macao), Indonesia (Borneo, Sumatra), Laos, Malaysia, Nepal, Sri Lanka, Thailand, Vietnam. Palearctic: Canary Islands, Egypt, Israel, Japan (Nansei Islands); Fig. 6.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
D98EEF00EC35CE87682604F14B9CFFF9.taxon	diagnosis	Diagnosis. Small to medium sized beetles, body length 2.4 - 4.9 mm. Body shape elongated oval in dorsal view; moderate to strongly convex in lateral view; dorsal outline uniformly convex or nearly straight and anteriorly inclined along anterior half (Fig. 46). Color brown, dark brown, reddish brown, or rather orange, usually uniform along body regions, but sometimes with slightly paler margins, pronotum or ventral surfaces and appendages; ground punctation shallow to moderately marked (Fig. 46). Shape of head trapezoid. Eyes small to moderate, seldom very small, not emarginated anteriorly, usually projected from outline of head. Clypeus trapezoid, with anterior margin broadly and roundly emarginate. Labrum fully exposed. Mentum rather flat and smooth, sometimes with lateral oblique ridges, and few crenulations; median anterior depression sometimes marked by a transverse carina (Fig. 46 C, F, I). Antennae with eight antennomeres; cupule slightly asymmetric, with rounded outline. Maxillary palps moderately stout, shorter to nearly as long as width of head; inner margin of maxillary palpomere 2 nearly straight, outer margin curved along apical 2 / 3; maxillary palpomeres 3 and 4 similar in length (Fig. 46 C, F, I). Prosternum flat to mesally only slightly produced (Fig. 46 C, F, I). Elytra with sutural striae; elytral punctures from shallow to sharply marked; ground punctures rather uniformly distributed; some species with serial punctures; outer margins of elytra slightly flared (Fig. 46 A, D, G). Posterior elevation of mesoventrite usually with curved transverse ridge, rather sharp and low, or bearing sharp, pyramidal (triangular) projection; anapleural sutures concave to forming obtuse angle, separated at anterior margin by distance 0.3 - 0.4 x width of anterior margin of mesepisternum (Fig. 46 C, F, I). Metaventrite with posteromesal glabrous patch nearly as wide as long (Fig. 46 C, F, I). Protibiae with spines of anterior row as thick, long semi-erect setae; apical spurs of protibiae moderately stout, reaching midlength of protarsomere 3. Metafemora with tibial grooves moderately developed; hydrofuge pubescence coverage ranging from sparse (nearly glabrous metafemora) to dense along basal 3 / 4 (Fig. 46 C, F, I). Tarsomeres 1 - 4 with long spiniform setae on ventral face; metatarsomere 2 nearly as long as 5 and as 3 and 4 combined. Fifth abdominal ventrite apically rounded, truncate, or slightly emarginate, usually with fringe of stout setae. Aedeagus trilobed (Fig. 47); basal piece as long or longer than parameres; median lobe triangular, nearly as wide at base as basal width of each paramere, with apical projection; gonopore absent.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
D98EEF00EC35CE87682604F14B9CFFF9.taxon	distribution	Distribution. Neotropical: Brazil (Para), Guyana, Suriname, and Venezuela; Fig. 6. We have seen additional specimens that slightly expand the range of the genus, but all still fall within the Guiana Shield region of South America.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
5EB1AAFB20AAC3BD840794A3A11F4BB5.taxon	diagnosis	Diagnosis. Small to very small beetles, body length 1.6 - 2.6 mm. Body shape oval in dorsal view; moderate to strongly convex in lateral view, dorsal outline evenly convex or nearly straight along median region (Fig. 48). Color orange brown to dark brown, uniform along body regions; ground punctation shallow to moderately marked (Fig. 48). Shape of head somewhat rectangular. Frons lateral and posteriorly expanded, forming Tobochares canthus completely dividing eyes in dorsal and ventral portions (Fig. 11 C). Eyes very small in dorsal view. Clypeus laterally expanded in front and around outer margin of eyes; anterior margin of clypeus straight (Fig. 11 C). Labrum partly exposed. Mentum rather smooth and medially depressed; median anterior depression marked by a transverse carina (Fig. 48 C, F). Antennae with nine antennomeres, cupule slightly asymmetric with rounded outline. Maxillary palps rather short and stout, nearly half as long as width of head; maxillary palpomere 4 slightly longer than palpomere 3; inner margin of maxillary palpomere 2 straight to convex, outer margin strongly curved along apical 2 / 3. Elytra without sutural striae, with punctures either irregularly distributed or forming well defined longitudinal rows; elytra narrowly explanate anteriorly, explanation gradually broader towards apex (Fig. 48). Surface of prosternum flat. Posterior elevation of mesoventrite, usually with well-defined transverse ridge, seldom with acute tooth; anapleural sutures concave, separated at anterior margin by distance nearly 0.7 x width of anterior margin of mesepisternum. Metaventrite usually uniformly densely pubescent, sometimes with reduced posteromedian glabrous patch. Protibiae with spines of anterior row hair-like, semi erect, relatively long, and thick; apical spurs of protibia moderately stout, reaching apex of protarsomere 3. All tarsomeres with thick hair-like spines on ventral face of tarsomeres 2 - 4; metatarsomeres 1 - 4 similar in length, 5 nearly as long as 3 and 4 combined. Metafemora with tibial grooves moderately developed; anterior surface of metafemora mostly glabrous, with few very scattered small setae (Fig. 48 C, F). Fifth abdominal ventrite apically rounded and without fringe of stout setae. Aedeagus trilobed (Fig. 49 A-D), with basal piece about half length of parameres; median lobe wider than base of each paramere, with narrow, triangular, longitudinal sclerite, usually extending along apical third; parameres as long as, to longer than median lobe, and nearly half as wide; gonopore situated preapically; basal piece with lateral margins straight to sinuate, apically slightly diverging.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
5EB1AAFB20AAC3BD840794A3A11F4BB5.taxon	distribution	Distribution. Neotropical: Brazil (Amazonas), Costa Rica, Ecuador, French Guiana, Guyana, Panama, Peru, Suriname, Venezuela; Fig. 6.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
6E1EF8D87ED362D57EC42B6336468F3A.taxon	diagnosis	Diagnosis. Medium sized beetles, body length 4.5 - 6.2 mm. Body shape oval in dorsal view; moderate to strongly convex in lateral view; dorsal outline nearly straight and anteriorly inclined along anterior half (Fig. 50). Color dark brown, usually uniform along body regions, sometimes margins of pronotum and elytra slightly paler; ground punctation fine, moderately marked (Fig. 50 A, D). Shape of head trapezoid and rather wide. Eyes moderate in size, not emarginated anteriorly, slightly projected from outline of head. Clypeus trapezoid, with anterior margin broadly, roundly, and weakly emarginate. Labrum fully exposed. Mentum medially rather broadly depressed, laterally longitudinally elevated; median anterior depression marked by transverse nearly straight carina (Fig. 50 C, F). Antennae with nine antennomeres; cupule slightly asymmetric, with rounded outline. Maxillary palps short and stout, nearly as long as half width of head (e. g., Fig. 50 C); inner margin of maxillary palpomere 2 nearly straight, outer margin strongly curved along apical 2 / 3; maxillary palpomere 4 slightly shorter than 3. Prosternum flat, only slightly carinate along midline of anterior projection. Elytra without sutural striae; elytral punctures shallow to moderately marked; ground punctures rather uniformly distributed; some species with serial punctures clearly visible along posterior third of elytra; outer margins of elytra slightly flared (Fig. 50 A, D). Posterior elevation of mesoventrite with median longitudinal carina elevated and forming posteriorly pointing process; anapleural sutures strongly concave, separated at anterior margin by distance nearly half width of anterior margin of mesepisternum. Metaventrite sometimes with posteromesal glabrous patch. Protibiae with anterior row of spines completely reduced; apical spurs of protibiae stout, reaching apex of protarsomere 3. Metafemora with tibial grooves very sharply marked and covered by hydrofuge pubescence; hydrofuge pubescence restricted to dorsal half on basal three-quarters of anterior surface of metafemora (Fig. 50 C, F). Tarsomeres 1 - 4 with long spiniform setae on ventral face; metatarsomere 2 nearly as long as 5 and as 3 and 4 combined. Fifth abdominal ventrite evenly rounded, without apical emargination or fringe of stout setae. Aedeagus either trilobed (Fig. 49 I-L) or divided (Fig. 49 G, H), with basal piece short and rather simple parameres separated from each other for most of their lengths; gonopore well developed.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
6E1EF8D87ED362D57EC42B6336468F3A.taxon	distribution	Distribution. Neotropical: Guyana, Suriname, Venezuela; Fig. 6.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
1297A93F7C9DFAABF03D8BA3354DB234.taxon	diagnosis	Diagnosis. Small to medium sized beetles, body length 2.5 - 5.0 mm. Body shape oval in dorsal view, moderately to strongly convex in lateral view (Fig. 51); dorsal outline usually evenly curved. Dorsal surfaces even and smooth, yellowish, orange brown to brown and rather uniform in coloration; ground punctation fine and extremely shallow (Fig. 51 A). Shape of head trapezoid. Eyes not emarginate, moderate to relatively large in size, subquadrate, separated by nearly 5 x width of eye, only slightly projected from outline of head. Clypeus trapezoid, with anterior margin broadly and slightly emarginate. Labrum fully exposed, convex, and anteriorly emarginate. Mentum rather flat, with few shallow transverse crenulations on anterior region; median anterior depression relatively shallow, sometimes marked by transverse carina (Fig. 51 C). Submentum smooth to very shallowly sculptured. Antennae with nine antennomeres, with strongly asymmetric and round cupule; antennomere 9 nearly 3 x longer than antennomere 8. Maxillary palps slender, 1.2 - 1.5 x longer than maximum width of head; inner margin of maxillary palpomere 2 usually evenly weakly curved, outer margin curved along apical third; palpomere 4 nearly 0.8 x as long as palpomere 3 (Fig. 51 C). Pronotum evenly convex, usually with systematic punctures forming distinct anterolateral semicircles. Elytra without sutural striae, with margins only slightly flared; serial punctures absent; scarce systematic punctures, bearing moderately long setae (Fig. 51 A). Surface of prosternum somewhat longitudinally elevated, sometimes with low and blunt longitudinal carina; anterior margin acutely to roundly projected anteriorly. Posterior elevation of mesoventrite with sharp and strongly elevated (laminar) longitudinal carina, with the ventral edge of the carina usually straight and parallel to the body (Fig. 51 C); anapleural sutures concave, separated at anterior margin by distance nearly half width of anterior margin of mesepisternum. Metaventrite densely and uniformly covered by hydrofuge pubescence (Fig. 51 C). Protibiae with anterior row of spines reduced (short appressed spines) to extremely reduced (tiny denticles); apical spurs of protibiae moderate, broad and reaching apex of protarsomere 2. Metafemora with tibial grooves sharply marked, and hydrofuge pubescence covering at least basal four fifths of anterior surface (Fig. 51 C). Metatarsomere 2 slightly shorter or similar in length to metatarsomere 5, metatarsomere 2 similar in length to metatarsomeres 3 and 4 combined; ventral surface of all tarsomeres with long setiform setae on ventral surface (tarsomeres 1 and 2 with small stout spines). Abdomen with five pubescent ventrites. Fifth abdominal ventrite emarginate at apex; emargination fringed by stout setae. Aedeagus divided (Fig. 49 E, F), somewhat pear-shaped, with basal piece nearly 0.3 x length of parameres; parameres slender, narrowing apically, with outer margins at least slightly sinuated, usually apically rounded; median lobe divided into dorsal and ventral plates; dorsal plate of median lobe medially bifurcate, with narrow, slender and apically rounded lobes; ventral lobe of median lobe varying in width and length, usually very lightly sclerotized; gonopore well-developed, usually positioned at midlength of aedeagus.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
1297A93F7C9DFAABF03D8BA3354DB234.taxon	distribution	Distribution. Neotropical: Argentina, Bolivia, Brazil (Amazonas, Mato Grosso do Sul, Rio de Janeiro, Rio Grande do Sul), Colombia [in doubt; d'Orchymont, 1943 d: 56], Costa Rica, French Guiana [in doubt; d'Orchymont, 1943 d: 56], Guatemala, Lesser Antilles (Antigua), Mexico, Nicaragua, Paraguay, Uruguay; Fig. 6.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
045BFFE789799959DDD435C8F3AC09F0.taxon	diagnosis	Diagnosis. Small beetles, total body length 1.5 - 2.6 mm. Body shape oval in dorsal view; moderately to strongly convex in lateral view (Fig. 52 - 54); dorsal outline usually evenly curved. Color yellowish brown, orange brown to dark brown, sometimes with paler spots on head, or paler margins of pronotum and elytra; ground punctation moderate to shallow. Shape of head somewhat oval. Eyes not emarginate (e. g., Fig. 11 A) to strongly emarginate (e. g., Fig. 11 B), moderate to small in size, somewhat oval, slightly to strongly projected from outline of head. Clypeus trapezoid, with anterior margin broadly emarginate; membranous preclypeal area often visible. Labrum fully exposed, convex, and anteriorly emarginate. Mentum rather smooth, often medially depressed, or anteriorly shallowly crenulated; median anterior depression marked by transverse carina (e. g., Fig. 53 C). Submentum anteriorly smooth and shiny. Antennae with eight antennomeres, cupule slightly asymmetric with rounded outline. Maxillary palps from short and slender (slightly shorter than the width of the head; e. g., Fig. 53 C) to very short and stout (nearly half the width of the head; Fig. 54 E); maxillary palpomere 4 similar in length to slightly longer than palpomere 3; inner margin of maxillary palpomere 2 straight, outer margin strongly curved along apical 2 / 3. Elytra without sutural striae (in some species, stria 1 more strongly impressed along posterior half of elytra; Fig. 54 C); elytral punctures seemingly arranged in rows, in some species more pronounced; interserial punctures occasionally longitudinally aligned; serial punctures sometimes impressed into distinct grooves (e. g., Fig. 52 A). Prosternum flat. Posterior elevation of mesoventrite either flat, bulging or with transverse or longitudinal ridge (Fig. 14 F, G); anapleural sutures concave, separated at anterior margin by distance nearly 0.3 - 0.5 x width of anterior margin of mesepisternum. Metaventrite densely pubescent, except for median glabrous patch, either ovoid and broad (Fig. 14 G) or longitudinal and narrow (Fig. 14 F). Protibiae with spines of anterior row hair-like, semi erect, relatively long and thick; apical spurs of protibia from very short and stout, to enlarged to reach apex of protarsomere 3. Tarsomeres 2 - 4 densely covered by hair-like spines on ventral face; metatarsomeres 1 - 4 similar in length, 5 nearly as long as 3 and 4 combined, or metatarsomere 2 similar in length to 5. Metafemora mostly glabrous, with only few scattered setae, sometimes with hydrofuge pubescence along basal half of anterodorsal margin (e. g., Figs 52 C, F, 53 C, F). Fifth abdominal ventrite apically evenly rounded, without fringe of stout setae. Aedeagus trilobed (Fig. 55), with basal piece usually very short (nearly 1 / 3 length of parameres); median lobe usually broader than each paramere; median lobe and parameres apically rounded to truncate; apex of median lobe seldom medially emarginated; gonopore well developed.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
045BFFE789799959DDD435C8F3AC09F0.taxon	distribution	Distribution. Neotropical: Brazil (Amapa, Amazonas, Goias, Roraima), French Guiana, Guyana, Suriname, Venezuela; Fig. 6.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
B8136D6F704C9A23DBC94DEE71C425A1.taxon	diagnosis	Diagnosis. Small beetles, body length 1.9 mm. Body shape oval in dorsal view; moderately convex in lateral view (Hansen 1991: fig. 39). Color yellowish light brown; ground punctation extremely shallowly marked. Shape of head somewhat oval. Eyes absent (Fig. 56 B). Clypeus trapezoidal, with anterior margin broadly emarginate, with medial region of emargination nearly straight (Fig. 56 B). Labrum fully exposed, convex. Mentum rather smooth and antero-medially depressed; median anterior depression broad. Antennae with nine antennomeres (Spangler 1981 a: fig. 3); cupule slightly asymmetric, with rounded outline. Maxillary palps slender, nearly as long as width of head; inner margin of maxillary palpomere 2 nearly straight, outer margin curved along apical third; maxillary palpomere 3 slightly shorter than 4. Prosternum non carinate, slightly convex. Elytra without sutural striae; ground punctation fine, shallow; outer margins slightly flared (Fig. 56 A). Posterior elevation of mesoventrite with curved, transverse ridge (Spangler 1981 a: fig. 8); anapleural sutures concave, separated at anterior margin by distance 0.7 x width of anterior margin of mesepisternum. Metaventrite densely pubescent except for median short and narrow posterior glabrous patch; metaventrite short (nearly as long as first abdominal ventrite; Spangler 1981 a: fig. 8). Protibiae with spines of anterior row long; apical spurs of protibiae moderately slender, reaching apex of protarsomere 2; metatarsomeres 2 - 4 slightly decreasing in size; metatarsomere 5 nearly as long as 2 - 4 combined. Posterior femora densely covered by hydrofuge pubescence along basal 2 / 3 (Spangler 1981 a: fig. 8). Fifth abdominal ventrite apically truncate, without stout setae (Spangler 1981 a: fig. 9).	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
B8136D6F704C9A23DBC94DEE71C425A1.taxon	distribution	Distribution. Neotropical: Ecuador; Fig. 6.	en	Giron, Jennifer C., Short, Andrew Edward Z. (2021): The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species. ZooKeys 1045: 1-236, DOI: http://dx.doi.org/10.3897/zookeys.1045.63810, URL: http://dx.doi.org/10.3897/zookeys.1045.63810
