taxonID	type	description	language	source
314487958F66B332FF7DFE09FDFAFBAE.taxon	materials_examined	Type species. Pomatias grandis Godwin-Austen, 1876.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F66B332FF7DFE09FDFAFBAE.taxon	diagnosis	Diagnosis. Shell large, turriform / spindle-shaped, regularly ribbed, ribs very low and weak; shallow " scoring " or groove above the suture; aperture oval with very smooth columellar-parietal and parietal-palatal transitions; peristome very much widened; operculum thin. Differential diagnosis. Differs from Vargapupa n. gen. and Nodopomatias by the absence of a basal keel, and from Pseudopomatias by the presence of a shallow groove above the suture. Csomapupa n. gen. species have conspicuously widened peristome margin, which is wider than in any species of Pseudopomatias. The parietalpalatal transition of the aperture is not angled in Cs. luyorensis, but very slightly angled in Cs. grandis. In Pseudopomatias this transition is usually much more conspicuous.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F66B332FF7DFE09FDFAFBAE.taxon	etymology	Etymology. The new genus Csomapupa (gender feminine) is dedicated to Sándor Kőrösi Csoma (1784? – 1842), also known as Alexander Csoma de Kőrös (approximate pronunciation is “ choma ”), Hungarian- Szekler (Székely) philologist and orientalist, author of the first Tibetan-English dictionary and grammar book. The name Csomapupa is the combination of the family name Csoma and Pupa. Content. Cs. grandis (Godwin-Austen, 1876), Cs. luyorensis (Godwin-Austen, 1917).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F66B332FF7DFE09FDFAFBAE.taxon	distribution	Distribution (Fig. 7): The two species of Csomapupa n. gen. were reported from the Dafla and Abor Hills (north-eastern India) (see also Fig. 2).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F66B332FF7DFEFBFE3BFE9A.taxon	materials_examined	Type genus: Pupina Vignard, 1829	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F79B32DFF7DFF4EFB14FB91.taxon	diagnosis	Diagnosis. A large species with " double " suture, weak sculpture, elongated apical portion, and simple-looking but conspicuously widened peristome margin.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F79B32DFF7DFF4EFB14FB91.taxon	description	Description. Shell yellowish, turriform, with the upper portion of the shell (first five whorls) being slightly concave; the 7.75 – 8.25 (n = 3), rather flat whorls are separated by shallow suture; above the suture there is a groove running parallel with it; this is present on the whole shell except of above the last one to two whorls; protoconch of two whorls; the sculpture of the first whorl is finely granulated, glossy, after that it becomes regularly ribbed; teleoconch regularly, extremely finely ribbed with no spiral lines visible; the ribs are very low and are regular even behind the aperture; aperture rather rounded; the parietal-palatal transition is slightly angled; columellar-parietal transition not angled; peristome white, very much widened and reflexed, consists of two circles; the inner one is slimmer, the outer one is much wider, expanded and reflexed; the boundary between the two circles is hardly visible, the whole periostome seems to be simple. Measurements (in mm). H: 12 – 13.3, D: 4 – 5.6 (n = 3). Differential diagnosis. Cs. luyorensis possesses a smaller shell, deeper suture and more conspicuous parallel groove above the suture. Moreover, the peristome of Cs. luyorensis is clearly separated into two circles, but that of Cs. grandis seems to be rather simple, but widened.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F79B32DFF7DFF4EFB14FB91.taxon	materials_examined	Type material. Dafla Hills, Assam, India, Godwin-Austin Collection, NHMUK 1903.7.1.3438 (9 syntypes).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F79B32DFF7DFF4EFB14FB91.taxon	distribution	Distribution. Cs. grandis is known from the type locality only (see also Fig. 2 and Table 3).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F79B32DFF7DFB1EFB29F865.taxon	diagnosis	Diagnosis. A large, tumid species with " double " suture, weak, but well-visible low ribs and very much widened peristome consisting of two well-visible circles.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F79B32DFF7DFB1EFB29F865.taxon	description	Description. Shell greyish-yellowish, rather spindle-shaped; 7.5 moderately bulging whorls are separated by rather shallow suture; above the suture there is a groove running parallel with it; this is present on the whole shell except for the last quarter of whorl; protoconch consist of about two whorls; the sculpture of the first 0.75 – 1 whorl is finely granulated, rather glossy, after that it becomes regularly ribbed; teleoconch is regularly, extremely finely ribbed with extremely fine spiral lines; the ribs are very low and are regular even behind the aperture; aperture rounded; none of the transitions (columellar-parietal and parietal-palatal) are angled; peristome very much widened and slightly reflexed, consists of two circles; the inner one is slimmer and protruding, the outer one is much wider and a bit expanded. Measurements (in mm). H: 11.3, D: 4.8 (n = 1). Differential diagnosis. See under Cs. luyorensis.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F79B32DFF7DFB1EFB29F865.taxon	materials_examined	Type material. Luyor, Abor Hills, NHMUK 1903.7.1.3377 / 2 syntypes (one adult and one juvenile shell).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F79B32DFF7DFB1EFB29F865.taxon	distribution	Distribution. Cs. luyorensis is known from the type locality only (see also Fig. 2 and Table 3).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7BB32FFF7DFF41FD00FC76.taxon	materials_examined	Type species. Eupomatias sibbumensis Godwin-Austen, 1917 (by original designation).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7BB32FFF7DFF41FD00FC76.taxon	diagnosis	Diagnosis. Shell medium sized to large, slender turriform, regularly ribbed; aperture round with slight upper incision (parietal-palatal transition); peristome double (the boundary between two circles clearly visible); an elevated keel runs around the umbilicus, starts from the most basal point of the peristome lip and terminates near the columellar-parietal transition of the peristome; operculum densely coiled, thin, membranous. Differential diagnosis. Differs from Pseudopomatias and Csomapupa n. gen. by the presence of the basal keel. Tortulosa and Eucataulus have stronger basal keels and a basal canal in the aperture. The keel of Nodopomatias is lower and situated much closer to the umbilicus than that of in Vargapupa n. gen. The keel of Nodopomatias terminates just behind the peristome lip, whereas that of Vargapupa n. gen. flattens gradually on the ventral side of the body whorl. Vargapupa n. gen. has an incision on the basal part of the aperture, where the keel starts from, however this character may not be independent from the position of the basal keel. The peristome of Nodopomatias is " double " (the boundary between the two circles of the peristome clearly visible) and that of Vargapupa n. gen. is " simple " (consists of a single circle, or the boundary is not visible between the two circles of the peristome). Content. oakesi (Godwin-Austen 1917), sibbumensis (Godwin-Austen 1917).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7BB32FFF7DFF41FD00FC76.taxon	distribution	Distribution (Fig. 7): Both species of Nodopomatias are known from the Abor Hills (Arunachal Pradesh, North-eastern India) (see also Fig. 2 and Table 3).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7BB32FFF7DFC3FFA85F902.taxon	diagnosis	Diagnosis. A medium sized, tumid species with very weak peristome margin and a basal keel situated close to the umbilicus.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7BB32FFF7DFC3FFA85F902.taxon	description	Description. Shell light brownish-yellowish; turriform, widest at its base; the 7.25 – 7.5 rather flat whorls (n = 3) are separated by shallow suture; the boundary of the protoconch was not visible on the three specimens examined because of corrosion; approx. the first whorl of the protoconch seemed to be smooth, glossy, the second whorl is extremely finely ribbed; ribs are regular on the whole shell, even behind the aperture; basal keel with blunt ribs; keel starts behind the aperture (a bit left from the lowermost point) and ends behind the columellar-basal meeting point of the aperture; aperture rounded with slight upper incision (parietal-palatal transition); parietal-columellar transition is not angled; apertural rim slightly thickened and not, or very slightly reflexed; it consists of two circles, a wider outer and a very thin inner; the boundary between these visible only from apertural view, from lateral view the aperture seems to be simple. Measurements (in mm). H: 11.0, D: 3.8 (n = 1). Differential diagnosis. Nodopomatias oakesi is much smaller than N. sibbumensis, has a more corpulent shell and weaker basal keel.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7BB32FFF7DFC3FFA85F902.taxon	materials_examined	Type material. Abor Hills, leg. Oakes, Godwin-Austin Collection, NHMUK 1903.7.1.3086. (3 syntypes).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7BB32FFF7DFC3FFA85F902.taxon	distribution	Distribution. Nodopomatias oakesi is known from the type locality only (see also Fig. 2 and Table 3).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7BB32EFF7DF8E3FA47FD29.taxon	diagnosis	Diagnosis. A large, turriform species with widened peristome margin consisting of two well-separated circles and a basal keel situated close to the umbilicus.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7BB32EFF7DF8E3FA47FD29.taxon	description	Description. The three shells available for study were corroded; therefore the shell colour and the fine sculpture of the protoconch could not be observed. It was visible however on the figured syntype, that the first whorl of the protoconch was not ribbed. Shell turriform, widest at its base; the 7.5 – 8.25 (n = 2), rather flat whorls are separated by shallow suture; ribs are regular on the whole shell; the holotype is the only shell with a complete aperture and neck region, but it has a scar (cicatrice) on the neck, therefore the sculpture behind the aperture could not be described properly; basal keel with blunt ribs; keel starts behind the aperture (a bit left from the most basal point), and ends behind the columellar-basal transition of the aperture; aperture round with slight upper incision (parietal-palatal transition); columellar-parietal transition not angled; peristome is very slightly reflexed; it consists of two circles, a wider outer and a very thin inner; the boundary between these is well visible from lateral view. Measurements (in mm). H: 14.5, D: 5.8 (n = 1). Differential diagnosis. See under Nodopomatias oakesi.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7BB32EFF7DF8E3FA47FD29.taxon	materials_examined	Type material. Sibbum, Abor Hills, leg. Oakes, Godwin-Austin Collection, NHMUK 1903.7.1.3146 (3 syntypes).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7BB32EFF7DF8E3FA47FD29.taxon	distribution	Distribution. Nodopomatias sibbumensis is known from the type locality only (see also Fig. 2 and Table 3).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7AB32EFF7DFCF9FAD4F9DA.taxon	materials_examined	Type species. Pseudopomatias amoenus Möllendorff, 1885 by monotypy.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7AB32EFF7DFCF9FAD4F9DA.taxon	diagnosis	Diagnosis. Shell small to large, turriform or spindle-shaped, rather regularly ribbed; aperture rather round with slight columellar-parietal and parietal-palatal transitions; the latter is usually more angled; the parietal side is usually straight; peristome consists of two circles, in some species it appears as " single "; basal keel missing; operculum densely coiled, thin, membranous (Figs 5 C – F). Differential diagnosis. Differs from Eucataulus Kobelt, 1902 (often referred as subgenus of Tortulosa Gray, 1847), Nodopomatias, Tortulosa (syn: Perlisia Tomlin, 1948) and Vargapupa n. gen. by the absence of a basal keel. Moreover, Tortulosa and Eucataulus usually have smooth shells and a basal canal in the aperture. Schistoloma Kobelt, 1902 (synonym: Pinteria Varga, 1972, see Egorov 2013) also has a more corpulent, smooth shell and some species possesses a basal keel. Pseudopomatias lacks the additional line (groove) above the suture, which is present in Csomapupa n. gen. The Australian genus Hedleya Cox, 1892 also has a similar shell in terms of shape and ribbing, but it has two canals in the aperture. Content. P. ab l e t t i n. sp., P. amoenus Möllendorff, 1885, P. e o s Pilsbry & Hirase, 1905, P. harli n. sp., P. himalayae (Benson, 1859), P. linanprietoae n. sp., P. maasseni n. sp., P. nitens n. sp., P. peguensis (Theobald, 1864), P. pleurophorus (Benson, 1857), P. p re s t o ni n. sp., P. re i s c h ue t z i n. sp., P. shanensis n. sp., P. s i y o m e n s i s Godwin-Austen, 1917, P. sophiae n. sp.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7AB32EFF7DFCF9FAD4F9DA.taxon	distribution	Distribution (Fig. 7): The genus is widely distributed in Asia from north-eastern India to Taiwan.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7AB328FF7DF914FED7FBD1.taxon	diagnosis	Diagnosis. A small species with very characteristic triangular aperture shape, " reversely oblique " peristome and flat whorls.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7AB328FF7DF914FED7FBD1.taxon	description	Description. Shell light greyish-yellowish, slender turriform; 7 – 7.75 (n = 2) whorls are flattened, but are separated by rather deep suture; protoconch region was corroded in the available shells; the only visible trait was that the first whorl was not ribbed; teleoconch regularly, finely ribbed; ribs are widely spaced and rather irregular on the last two whorls, their shape is slightly wavy; the last quarter of whorl is with one to two ribs; ribs relatively strong, with fine spiral structure between the ribs on the whole teleoconch; aperture almost triangular, " reversely oblique " from lateral view (upper end is situated posteriorly, basal end anteriorly); parietal part is straight; apertural rim thickened and reflexed, the boundary between the two circles of the rim is hardly visible. Measurements (in mm). H: 6.1 – 6.3, D: 3.0 – 3.2 (n = 2). Differential diagnosis. P. abletti n. sp. differs from P. harli n. sp., P. himalayae, P. sophiae n. sp. by the flatter whorls, triangular aperture and the more widely spaced ribs (especially on the neck region), with usually strong spiral lines inbetween. Moreover, from lateral view the apertural margin of P. abletti n. sp. is strongly oblique. See also under P. siyomensis and Table 5.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7AB328FF7DF914FED7FBD1.taxon	materials_examined	Material. Himalaya, Darjiling, coll. W. Blanford, NHMUK 1860.6.27.32.3 (holotype), NHMUK 1860.6.27.32.4 (1 paratype) (mixed sample with P. himalayae); India, NHMUK 20130409.3 (1 paratype) (mixed sample with P. himalayae); Toruputu Peak, 500, Dafla, NHMUK 1903.7.1.3746.9 / 1 (with weaker spiral lines, not paratype); Indien, Dr. Stoliczka coll, 1880, NHMW 109196 (1 paratype; mixed sample with P. prestoni n. sp.: NHMW 109194 and P. h i m al ay a e: NHMW 109195); Darjiling, Indien, Oberwimmer coll, NHMW 109197 (1 paratype; mixed sample with P. himalayae: NHMW 66304); Darjeeling, leg. Stoliczka, 1870, NHMUK 109198 / 1; Darjiling, coll Ehrmann ex Sowerby + Fulton, SMF 342541 (mixed sample with P. himalayae: SMF 109828). Type locality. Himalaya, Darjiling.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7AB328FF7DF914FED7FBD1.taxon	etymology	Etymology. The new species is dedicated to Jonathan Ablett (NHM), the friend of the first author, for his boundless help during this and other taxonomic revisions.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7AB328FF7DF914FED7FBD1.taxon	distribution	Distribution. Most samples were collected near Darjiling, one sample is labelled as being collected from Toruputu Peak (Dafla Hills) (see also Fig. 2 and Table 3).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7AB328FF7DF914FED7FBD1.taxon	discussion	Remarks. In museum collections all specimens of this species were in mixed lots with Pseudopomatias himalayae.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7CB32BFF7DFB5EFB80FC09.taxon	diagnosis	Diagnosis. A small, turriform, " solid " species with weak sculpture on the neck region.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7CB32BFF7DFB5EFB80FC09.taxon	description	Description. Shell small, greyish-yellowish to light brown, turriform; the 6.25 – 6.75 (n = 5) rather bulging whorls are separated by deep suture; protoconch consists of two whorls, the first 0.75 – 1 whorl is very finely granulated, than it becomes very finely, regularly ribbed; teleoconch regularly, very finely ribbed; ribs gradually weaken on the last whorl; very weak ribs are visible on the neck region of most shells, whereas in the minority of shells the last quarter of whorl is smooth; whole shell with very fine spiral lines; aperture rounded with slightly angled columellar-parietal transition and sharply angled parietal-palatal transition; apertural rim slightly thickened and not, or very slightly reflexed; parietal callus weak; the boundary between the two circles of the rim is usually hardly visible. Measurements (in mm). H: 4.6 – 7.1, D: 2.6 – 3.5 (n = 5). Differential diagnosis. P. harli n. sp. differs from P. h i m al ay a e by the blunter ribs and the smoother area (approximately a quarter of whorl) behind the aperture. In most cases P. himalayae has a thicker apertural rim and stronger callus, more slender shell and deeper suture. P. pleurophorus is larger and has a more corpulent shell, and its shell is almost smooth behind the apertural margin, whereas that area is usually ribbed in P. ha r l i n. sp. See also under P. abletti n. sp., P. reischuetzi n. sp., P. s i y o m e ns i s and P. prestoni n. sp. and Table 5.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7CB32BFF7DFB5EFB80FC09.taxon	materials_examined	Material. Damsang Peak, W Bhutan, Godwin-Austen coll, NHMUK 20130421.1 (holotype), NHMUK 20130421.2 – 229 (228 adult shells = paratypes), NHMUK 20130421.230 – 435 (206 juvenile / broken shells = not paratypes) (mixed sample with P. prestoni n. sp.); Richila Peak, Sikhim, NHMUK 1903.7.1.3468 / 9 paratypes; Sikkim, Himalayas, NHMUK 1888.12.4.338 – 340 / 3 paratypes (mixed sample with P. himalayae); Darjiling, Godwin-Austen coll, NHMUK 1903.7.1.3955.4 (1 paratype; mixed sample with P. himalayae); Rissom Peak, Darjeeling, Bhutan, leg. W. Robert, 1882, Godwin-Austen coll, NHMUK 20130411.3 (one paratype; mixed sample with P. himalayae); Himalayas, E. R. Sykes coll, Acc. no. 1825, NHMUK 20130413.2 (one paratype; mixed sample with P. cf. himalayae and P. prestoni n. sp.); Singtam-Sikhim, NHMUK 1906.1.1.1004.5 – 11 (7 paratypes), NHMUK 1906.1.1.1004.12 – 19 (8 not paratypes = juveniles; mixed sample with P. himalayae); Cherra Poonjee & Kangbun, Godwin-Austen Collection, NHMUK 1903.7.1.3365.1 (1 adult = paratype), NHMUK 1903.7.1.3365.2 – 3 (2 juveniles = not paratypes) (mixed sample with P. pleurophorus); Damsang, Godwin-Austen coll, NHMUK 20130417 / 1 paratype; Rissom, Godwin-Austen coll, NHMUK 20130418.1 – 3 (3 adults = paratypes), NHMUK 20130418.4 – 6 (3 juveniles = not paratypes); Rissom Peak, Sikkim, Godwin-Austen coll, NHMUK 20130419.1 – 4 (4 paratypes), NHMUK 20130419.5 – 11 (7 juveniles = not paratypes); Damsang Peak, Godwin- Austen coll, NHMUK 20130420 / 1 paratype; Barowli Gorge, Aka Hills, NHMUK 1903.7.1.1587 / 10 paratypes; Valley east of Cherra Poonjee, NHMUK 1903.7.1.1588 / 1 (with regular ribs on the neck region, not paratype); Toruputu Peak, 7000, Dafla, NHMUK 1903.7.1.3746 / 1 – 3 (3 paratypes), NHMUK 1903.7.1.3746 / 4 – 8 (5 shells, with few strong ribs on the neck, labelled as cf. harli, not paratypes); Rissom Peak, Sikkim, NHMUK 1903.7.1.3469 / 1 paratype (mixed sample with P. prestoni n. sp.); Damsang Peak, Sikkim, W Bhutan, leg. Godwin- Austen, NHMUK 20130423 / 1 paratype (mixed sample with P. prestoni n. sp.). Type locality. Damsang Peak, W Bhutan.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7CB32BFF7DFB5EFB80FC09.taxon	etymology	Etymology. The new species is dedicated to and named after Josef Harl, Austrian biologist and malacologist, the friend of the first author.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7CB32BFF7DFB5EFB80FC09.taxon	distribution	Distribution. Most samples were collected in Sikkim (Rissom Peak, Richila Peak, Damsang Peak), whereas some others were labelled as being collected in Dafla Hills (southern Arunachal Pradesh), Cherrapunji and Kangbun (Meghalaya state) (see also Fig. 2 and Table 3).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7CB32BFF7DFB5EFB80FC09.taxon	discussion	Remarks. In museum collections Pseudopomatias harli n. sp. was frequently in mixed lots with P. himalayae and P. prestoni n. sp. In one sample, P. h ar l i n. sp. was mixed with P. pleurophorus.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7FB325FF7DFBE6FD95FCDD.taxon	diagnosis	Diagnosis. A small, relatively slender turriform species with strong ribs and thick peristome margin.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7FB325FF7DFBE6FD95FCDD.taxon	description	Description. Shell light brown or yellowish-greenish, slender turriform, widest at its base; the 6.5 – 7.25 (n = 4) bulging whorls are separated by deep suture; protoconch consists of 2 whorls, first whorl is granulated, the second is very finely, regularly ribbed; teleoconch also finely, regularly ribbed without spiral lines; ribs very sharp, but are becoming less regular and sharper on the last half whorl; aperture rounded with slightly angled columellar-parietal transition and sharply angled parietal-palatal transition; apertural rim very much thickened and reflexed; it consists of a slimmer inner and a wider outer circle, but these are usually fused that the transition is not visible. Measurements (in mm). H: 5.6 – 7, D: 2.7 – 3.5 (n = 5). Differential diagnosis. See under P. s i y om en s i s, P. ab l e t t i n. sp. P. ha r l i n. sp. P. linanprietoae n. sp. and P. sophiae n. sp. and Table 5.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7FB325FF7DFBE6FD95FCDD.taxon	materials_examined	Type material. Syntype: UMZC 102695 Darjiling. Museum material. Himalaya, Darjiling, coll. W. Blanford, NHMUK 1860.6.27.32.1 – 2 (2 shells; mixed sample with P. abletti n. sp.); India, NHMUK 20130409.1 – 2 (2 shells; mixed sample with P. abletti n. sp.); Darjeeling, H. Cuming coll, Acc. no.: 1829, NHMUK 20130410 / 3; Sikkim, Himalayas, NHMUK 1888.12.4.336 – 337 (2 shells; mixed sample with P. harli n. sp.); India, NHMUK 1871.9.23.147 / 2; Darjiling, Godwin-Austen coll, NHMUK 1903.7.1.3955.1 – 3 (3 shells; mixed sample with P. harli n. sp.); Rissom Peak, Darjeeling, Bhutan, leg. W. Robert, 1882, Godwin-Austen coll, NHMUK 20130411.1 – 2 (2 shells; mixed sample with P. h ar l i n. sp.); Darjeeling, coll Sowerby, 12 / 10 / 94 / 112, J. J. MacAndrew coll, NHMUK 20130412 / 6; Himalayas, E. R. Sykes coll, Acc. no. 1825, NHMUK 20130413.1 (one very wide shell, labelled as “ cf. himalayae ”; mixed sample with P. harl i n. sp. and P. prestoni n. sp.); Darjeeling, B. R. Lucas coll, Acc. no.: 2351, NHMUK 20130414 / 2; no locality, NHMUK 20130415 / 1; Singtam-Sikhim, NHMUK 1906.1.1.1004.1 – 4 / 4; Indien, Dr. Stoliczka coll, 1880, NHMW 109195 / 3 (mixed sample with P. p res t o ni n. sp.: NHMW 109194 and P. abletti n. sp.: NHMW 109196); Darjiling, Indien, Oberwimmer coll, NHMW 66304 / 2 (mixed sample with P. abletti n. sp.: NHMW 109197); Himalaya, coll. Möllendorff, SMF 109827 / 3; Darjiling, coll Ehrmann ex Sowerby + Fulton, SMF 109828 / 1 (mixed sample with P. abletti n. sp.: SMF 342541).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7FB325FF7DFBE6FD95FCDD.taxon	distribution	Distribution. The species was mentioned from several localities, but the revision revealed that many samples belonged to other species. Confirmed P. himalayae samples are from Darjeeling and Sikkim. See also Fig. 2 and Table 3.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F7FB325FF7DFBE6FD95FCDD.taxon	discussion	Remarks. In museum collections Pseudopomatias himalayae was frequently in mixed lots with P. abletti n. sp., P. harli n. sp. and P. prestoni n. sp.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F71B324FF7DFC55FE15FF1D.taxon	diagnosis	Diagnosis. A small to medium sized, slender turriform species with weak sculpture, bulging whorls (deep suture) and very thin apertural margin.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F71B324FF7DFC55FE15FF1D.taxon	description	Description. Shell yellowish-greenish, very slender turriform, widest at its base; the 7.25 – 8.25 (n = 4), very much bulging whorls are separated by deep suture; protoconch consists of 2 whorls, first half whorl is very finely granulated, glossy; the remaining whorls are very finely, regularly ribbed; teleoconch also finely, regularly ribbed without spiral lines; ribs very low, they become less regular on the last half whorl; aperture rounded with very slightly angled columellar-parietal transition and slightly more sharply angled parietal-palatal transition; apertural rim whitish, slightly thickened and usually not, or very slightly reflexed; it consists of a slimmer inner and a wider outer circle; the boundary between them is hardly visible. Measurements (in mm). H: 7.9 – 10.4, D: 3.6 – 4.1 (NHMUK 1906.5.5.9, n = 3). Differential diagnosis. Pseudopomatias peguensis has deeper suture and more bulging whorls than any other Pseudopomatias species. The most similar species in size, shell shape and morphology of the apertural lip is P. amoenus which has a shallower suture and stronger ribs.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F71B324FF7DFC55FE15FF1D.taxon	materials_examined	Type material. According to Dance (1986) some shells of Theobald’s material is in the NHM, the Indian Museum, and Cambridge Museum, a few types in Cardiff Museum (excoll Fedden). We contacted these museums (except Indian Museum) but we were unable to locate the type specimens. Museum material. Pegu, Burmah, Fedden collection, NMW. 1894.015 / 6; no locality, coll. Godwin-Austen, NHMUK 20130403 / 1; Gwah Hill, Pegu, Burma, E. R. Sykes coll, Acc. no. 1825, NHMUK 20130404 / 2; Gwah Hill, NHMUK 1888.124.331 – 4 / 4; Gwa, Arakan, W. Blanford coll, NHMUK 1906.5.5.9 / 7.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F71B324FF7DFC55FE15FF1D.taxon	distribution	Distribution (Fig. 11): The type locality of the species is Pegu (= Bago, Myanmar). Blanford (1864) mentioned Theobald’s new species from Arakan, and later, Nevill (1878) and Möllendorff (1886) marked " Mai, Sandoway ". Both the Arakan Mts (Rakhin State) and Sandoway are located in western Myanmar, whereas Bago Province lies eastwards. See also Table 3.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F70B324FF7DFE95FA89F875.taxon	diagnosis	Diagnosis. A small to medium sized, widely based turriform species with almost smooth neck region.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F70B324FF7DFE95FA89F875.taxon	description	Description. Shell greyish-yellowish to light brown, widely based turriform; the 6.75 – 7 (n = 3), rather bulging whorls are separated by rather deep suture; last whorl very much bulging; all examined shells had somewhat corroded apical part, therefore the boundary of the smooth and ribbed parts on the protoconch could not be observed; teleoconch regularly, very finely ribbed; ribs gradually weaken on the last whorl, the last half whorl is almost smooth except for the area close to the suture; aperture rounded, with nearly not angled columellar-parietal transition and somewhat sharply angled parietal-palatal transition; apertural rim slightly thickened and not, or very slightly reflexed; it consists of two circles; a thinner inner and a bit wider and stronger outer, but they are not conspicuously different in width. Measurements (in mm). H: 6.1 – 7.9, D: 3.7 – 4.4 (n = 4). Differential diagnosis. The wide-base triangle shape of Pseudopomatias pleurophorus distinguishes it from all other congeneric species. For comparisons with P. p res t o ni n. sp., P. harli n. sp. and P. reischuetzi n. sp., see under those species, and Table 5.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F70B324FF7DFE95FA89F875.taxon	materials_examined	Type material. The original description was based on a single specimen, which we were unable to locate. Museum material. Abor Hills, leg. Capt. Oakes, R. E, Godwin-Austen Coll., NHMUK 1903.7.1.3137 / 2 (" var. elongata "); Khasi Hills, purchased by W. Theobald, NHMUK 1888.12.4.329 / 1; Cherra Poonjee (= Cherrapunji) & Kangbun, Godwin-Austen Collection, NHMUK 1903.7.1.3365.2 – 7 (6 shells; mixed sample with P. harli n. sp.); Rywuk, Garo Hills NHMUK 1903.7.1.3365.1 (1 shell); Harmutti, Dafla Hills, Godwin-Austen Collection, NHMUK 1903.7.1.1536.1 – 4 (4 shells) (" var. daflaensis "); Village of Pachitah (Camp 7), Godwin-Austen Collection, NHMUK 1903.7.1.1536.5 – 7 (3 shells) (" var. daflaensis "); Dafla Hills, Attaram, W. T. Blanford Collection, NHMUK 1906.1.1.975 / 1 (" var. daflaensis "); Khasi Hills (?), W. T. Blanford Collection, NHMUK 1906.5.5.65 / 1.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F70B324FF7DFE95FA89F875.taxon	discussion	Remarks. A sample labelled as P. pleurophorus in the NMW (Psedopomatias [sic] pleurophorus Bens, Khasi Hills, six shells, Melvill-Tomlin collection NMW. 1955.158) contained two species of Cochlostoma; four shells of Cochlostoma cf. scalarinum (Villa, 1841) and two shells of Cochlostoma cf. elegans (Clessin, 1879).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F70B324FF7DFE95FA89F875.taxon	distribution	Distribution. Museum samples are labelled as being collected in different North-eastern Indian localities in Arunachal Pradesh (e. g. Dafla Hills) and Meghalaya Province (e. g. Garo and Khasi Hills). The exact localities of some geographical names (Kangbun, Rywuk, Pachitah, Harmutti) are unknown. See also Fig. 2 and Table 3.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F70B324FF7DFE95FA89F875.taxon	discussion	Remarks. In one museum sample Pseudopomatias pleurophorus was in a mixed lot with P. harli n. sp.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F73B327FF7DFF4EFEE0F9CB.taxon	diagnosis	Diagnosis. A small to medium sized, turriform species with conspicuously thickened ribs (tubercles) around the umbilicus.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F73B327FF7DFF4EFEE0F9CB.taxon	description	Description. Shell greenish-yellowish, rather turriform in shape, widest at its base; the 5.75 – 6.5, rather flat whorls are separated by shallow suture (although the apical whorls can be more bulging with deeper suture); protoconch consist of about 1.5 whorls; the sculpture of the first 0.75 – 1 whorl is granulated, matt, ribs are not detectable; after that it becomes regularly ribbed; teleoconch regularly, finely ribbed with extremely fine spiral lines; the ribs are denser on the apical whorls and become more widely-spaced and lower on the penultimate and last whorls; ribs disappear on the last half whorl, where the very fine, spiral structure is better visible; umbilicus is surrounded by three to five white, radial knots, which represent the lower halfs of the ribs; behind the apertural rim there are one to three (in most cases two) whitish thickenings, usually one of them is much thicker than the others; these thickenings are “ full-length ” ribs near the aperture but became visible only near the umbilicus in posterior direction (= radial knots around the umbilicus); aperture round, with the parietal part being straight; columellarparietal transition slightly, parietal-palatal transition sharply angled; apertural rim slightly reflexed, relatively thin, whitish, with having a slimmer inner and a wider, thicker outer circle; parietal callus well-developed. Measurements (in mm). H: 5.8 – 7.8, D: 3.1 – 3.7. (type series, n = 3); H: 8.6 – 9.4, D: 3.7 – 3.8 (NHMW 109194, n = 2). Differential diagnosis. P. prestoni n. sp. can be distinguished from any other congeneric species by the strong tubercles on the base of the shell. The most similar species in terms of shell shape and aperture morphology is P. harli n. sp., which is, addition to the smooth periumbilical region, usually smaller and has more bulging whorls. P. pleurophorus possesses a more tumid, wider shell without spiral sculpture between the ribs. See also Table 5.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F73B327FF7DFF4EFEE0F9CB.taxon	materials_examined	Material: Damsang Peak, Sikkim, W Bhutan, leg. Godwin-Austen, NHMUK 20130423.1 (holotype), NHMUK 20130423.2 – 46 (46 paratypes), (mixed sample with P. harli n. sp.); Himalayas, E. R. Sykes coll., Acc. no. 1825, NHMUK 20130413.3 (one paratype; mixed sample with P. cf. himalayae and P. harli n. sp.); Damsang Peak, W Bhutan, Godwin-Austen coll., NHMUK 20130421 / 1 paratype (mixed sample with P. ha r l i n. sp.); Rissom Peak, Sikkim, NHMUK 1903.7.1.3469 / 7 paratypes (mixed sample with P. h ar l i n. sp.); Damsang Peak, Sikkim, leg. Godwin-Austen, coll. A. J. Peile, Acc. no. 2242, NHMUK 20130422 / 3 paratypes; Indien, Dr. Stoliczka coll, 1880, NHMW 109194 / 2 (unusually tall shells with strong ribs, not paratypes; mixed sample with P. him al ayae: NHMW 109195 and P. abletti n. sp.: NHMW 109196). Type locality. Damsang Peak, Sikkim, W Bhutan.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F73B327FF7DFF4EFEE0F9CB.taxon	etymology	Etymology. This new species is dedicated to Hugh Berthon Preston (1871 – 1945), British malacologist, who worked on the genus Pseudopomatias at the beginning of the 20 th Century, but did not publish his results.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F73B327FF7DFF4EFEE0F9CB.taxon	distribution	Distribution. According to original labels, the species is known from Sikkim state (India) only. See also Fig. 2 and Table 3.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F73B327FF7DFF4EFEE0F9CB.taxon	discussion	Remarks. Two shells from " India " (NHMW 109194) have taller (6.8 – 9.4 mm), slimmer shell than the type series, and have more conspicuous sculpture, stronger ribs and spiral structure. In museum collections Pseudopomatias prestoni n. sp. was frequently in mixed lots with P. himalayae, P. harli n. sp, and once with P. abletti n. sp.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F73B326FF7DF958FD43FCF0.taxon	diagnosis	Diagnosis. A small to medium sized, glossy, rather spindle-shaped species with almost smooth neck region, rather triangular aperture (strongly angled parietal-palatal transition) and weak peristome margin.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F73B326FF7DF958FD43FCF0.taxon	description	Description. Shell greyish-yellowish, glossy, spindle shaped, widest at the penultimate whorl; the 7.25 – 8 (n = 4) bulging whorls are separated by deep suture; protoconch consists of 1.5 whorls, the first half is very finely granulated, the remaining part is very finely, regularly ribbed; teleoconch also regularly, very finely ribbed; ribs low, not sharp; on the last half whorl the ribbed surface turns into a rather smooth surface having irregular growth lines; shell surface without spiral lines; aperture rather triangular, with smoothly angled columellar-parietal transition and sharply angled parietal-palatal transition; apertural rim slightly thickened and very slightly reflexed, consists of two circles; the inner circle is sharp in front and conspicuously protruding; the outer one being wider; parietal callus normally developed; there are 2 – 3 inner palatal plicae visible above the umbilicus through the translucent shell. Measurements (in mm). H: 7.3 – 8.9, D: 3.3 – 3.7 (n = 3). Differential diagnosis. P. reischuetzi n. sp. differs from P. harli n. sp. and P. pleurophorus by the presence of plicae inside the last whorl, the more slender, spindle-shaped shell, the triangular aperture and the absence of spiral sculpture. P. h i m al ay a e is smaller, has much stronger and sharper ribs, especially on the neck region, it has a rounded aperture and also lacks the plicae. See also under P. si yomensis.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F73B326FF7DF958FD43FCF0.taxon	materials_examined	Material. Lhota Naga, Godwin-Austen coll., NHMUK 1903.7.1.1549 / 1 (holotype), NHMUK 1903.7.1.1549 / 2 – 4 (3 paratypes). Type locality. Lhota Naga.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F73B326FF7DF958FD43FCF0.taxon	etymology	Etymology. The new species is dedicated to and named after Alexander Reischütz, Austrian malacologist, the friend of the first author.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F73B326FF7DF958FD43FCF0.taxon	distribution	Distribution. Only the type series is known, which was collected in " Lhota Naga ", Naga Hills. See also Fig. 2 and Table 3.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F73B326FF7DF958FD43FCF0.taxon	discussion	Remarks. There may be more, deeper situated plicae within the last whorl that are visible only by breaking the shells (see Notes on useful shell characters).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F72B326FF7DFCBEFF35F93A.taxon	diagnosis	Diagnosis. A small to medium sized, slender, almost cylindrical species with strong spiral sculpture between the strong but flattened (blunt) ribs.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F72B326FF7DFCBEFF35F93A.taxon	description	Description. Shell greenish, but all available shells were corroded, therefore the original colour was not visible; shell very slender, rather cylindrical, the last whorl is only very slightly wider than the penultimate one; whorls 7.5 – 8 (n = 2) in number, moderately bulging, separated by rather deep suture; protoconch consists of 2 whorls, first half whorl is very finely granulated, glossy; remaining whorls very finely, regularly ribbed; teleoconch also finely, regularly ribbed even behind the aperture, where the ribs are the most widely-spaced; ribs very well visible, their back is flat, not sharp, between the ribs very fine, regular spiral lines can be seen; aperture rounded drop-shaped with a conspicuous, sharply angled parietal-palatal transition; the columellar-parietal transition is only smoothly angled; apertural rim white, strongly widened and slightly reflexed; it consists of thinner slimmer inner and a wider outer circle, which are completely connected to each other, the boundary is very slightly visible only from apertural (frontal) view. Measurements (in mm). H: 7.8 – 9.1, D: 3.1 – 3.4 (n = 2). Differential diagnosis. P. shanensis n. sp. can be distinguished by other Pseudopomatias species by the very slender, almost cylindrical shell and very fine spiral sculpture between the blunt ribs.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F72B326FF7DFCBEFF35F93A.taxon	materials_examined	Material. Shan States, Woodthorpe, 1895, NHMUK 1903.7.1.1631.1 (holotype), NHMUK 1903.7.1.1631.2 (1 paratype) (note 1903.6.1.1631 is a mislabelling on original box). Type locality. Shan States.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F72B326FF7DFCBEFF35F93A.taxon	etymology	Etymology. The new species was collected in the " Shan States ". The original sample was labelled as " shanense " by Godwin-Austen.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F72B326FF7DFCBEFF35F93A.taxon	distribution	Distribution (Fig. 11): The only sample is labelled as being collected from Shan States (Myanmar). See also Table 3.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F72B321FF7DF8EBFB18FD15.taxon	diagnosis	Diagnosis. A large species with strong, widely-spaced ribs and well-developed peristome consisting of two readily distinguishable circles.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F72B321FF7DF8EBFB18FD15.taxon	description	Description. The only known shell is corroded, the shell colour and the sculpture of the protoconch could not be observed; shell slender turriform; whorls 9 in number and they are moderately bulging, separated by rather deep suture; teleoconch finely, regularly ribbed even behind the aperture, where the spaces between the ribs are wider than on the previous whorls; ribs seem to be stronger behind the aperture; aperture rounded with not angled columellar-parietal transition and slightly angled parietal-palatal transition; peristome very much thickened and slightly reflexed; it consists of a slimmer, protruding inner and a wider outer circle. Measurements (in mm). H: 12.2, D: 4.9 (n = 1). Differential diagnosis. P. siyomensis is larger than P. himalayae, P. abletti n. sp. and P. harl i n. sp. Moreover, P. himalayae has a wider shell and more bulging whorls, and its aperture is oblique in lateral view; P. abletti n. sp. has " reversed oblique ", rather triangular aperture; P. ha r l i n. sp. has turriform shell shape, lower ribs and a thinner apertural lip, which is not divided into two separated circles. P. rei schuetzi n. sp. is slightly more slender, has a thinner apertural rim and almost smooth neck area (last half of whorl).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F72B321FF7DF8EBFB18FD15.taxon	materials_examined	Type material. Siyom, Abor Hills, leg. Captain Oakes, coll. Godwin-Austen, NHMUK 3406.03.7.1. (holotype).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F72B321FF7DF8EBFB18FD15.taxon	distribution	Distribution. P. siyomensis is known from the type locality only. See also Fig. 2 and Table 3.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F74B31CFF7DFF04FC0EFE35.taxon	diagnosis	Diagnosis. A small to large, very variable species in terms of shell shape, peristome thickness and size. It can be recognized by the absence of plicae in the last whorl, the regular, low ribs on the whole shell, even behind the aperture and the rather irregular aperture shape. The shell shape is usually turriform (sometimes rather spindleshaped); peristome very thin in most cases, but can be thickened in some populations.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F74B31CFF7DFF04FC0EFE35.taxon	description	Description of the type series of amoenus: Shell light brown to greenish, slender turriform or rarely spindleshaped; apex blunt; 6.75 – 7.5 (n = 3), rather bulging whorls are separated by deep suture; protoconch consist of about 1.5 whorls; the sculpture of the first half whorl is smooth, glossy, than becomes regularly, very finely ribbed; teleoconch is rather regularly, finely ribbed without spiral lines; the ribs very low, especially on the body whorl; they become somewhat irregular behind the aperture; aperture rather rectangular with very smoothly angled columellar-parietal transition and somewhat sharply angled parietal-palatal transition; all specimens has a bit deformed round shape; apertural rim slim and weak, not reflexed; parietal callus weak. Description of the type series of corneus (synonym of amoenus): Shell yellow, rather turriform with rather blunt apex; 7 – 7.25 (n = 2) bulging whorls are separated by rather deep suture; protoconch consist of one and a half whorls, first whorl rather smooth, the last half whorl is very finely, regularly ribbed (observed in one, not corroded shell); teleoconch regularly, finely ribbed, but ribs becoming lower and somewhat less regular on the last quarter of whorl; no spiral lines visible; aperture rounded with not angled columellar-parietal transition and smoothly angled parietal-palatal transition; apertural rim slightly (MCZ 167038) or strongly (USNM 472343) thickened; the boundary between the two circles is visible, but not conspicuous; the two circles are about of the same strength; parietal callus normally developed but slightly weaker than the apertural rim. Description of the holotype of fulvus (synonym of amoenus): Shell brownish, spindle-shaped with pointed apex; 7.75 (n = 1), rather bulging whorls are separated by deep suture; the sculpture of the first half whorl is smooth, glossy; the remaining two whorls of the protoconch are regularly, very finely ribbed; teleoconch regularly, finely ribbed with extremely fine spiral lines which are consist of a series of minute tubercles; ribs well-pronounced, even on the neck region, but they are slightly lower there; aperture round with not angled columellar-parietal transition and slighlty angled parietal-palatal transition; apertural rim well-developed, rather thick but the boundary between the circles does not seem to be visible; callus well-developed. Measurements (in mm). H: 8.5 – 10, D: 3.5 – 4.0 (" typical " amoenus, NHMUK 20130387, n = 3); H: 7.6 – 9.0, D: 3.7 – 4.1 (corneus, n = 3); H: 9.3, D: 4.1 (holotype of fulvus); H: 8.7, D: 3.4 (shell 13 N of Th ất Khê, identical with typical fulvus); H: 10.9 – 13.2, D: 4.5 – 5.4 (NHMUK 1910.1.21.4 – 6, n = 3); H: 9.5 – 11 mm, D: 3.4 – 4.3 (Kongshan Cun, n = 5). Differential diagnosis. P. maasseni n. sp. has flatter whorls (especially the last), a shallower suture, more elongated shell and every second ribs around the umbilicus are prominent. For comparisons with P. sophiae n. sp. and P. nitens n. sp., see under those species. All these three species mentioned (P. maasseni n. sp. P. sophiae n. sp. and P. nit ens n. sp.) co-occur with P. amoenus. See also under P. e o s and Table 6.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F74B31CFF7DFF04FC0EFE35.taxon	materials_examined	Type material. China: Hupei: Patung; coll. Möllendorff, SMF 40296, (lectotype of amoenus); same data, SMF 40298 (two paralectotypes of amoenus); same data, SMF 40297 (five adult and two juvenile paralectotypes of amoenus); China: Hunan Heng-shan-hsien, coll. Möllendorff, SMF 40299, (lectotype of amoenus tumidula); same data, SMF 40300 (one paralectotype of amoenus tumidula); Tchen-K’eou, USNM 472343 (syntype of cornea); China, Se-chuan, Tchen-K’eou, leg. Farges, MCZ 167038 (2 syntypes of cornea); Tonkin: Manson-Gebirge (mountains), coll. Möllendorff ex Fruhstorfer, SMF 109826 (holotype of fulvus). Museum material. Hupei, Badung, J. J. MacAndrew coll, ex Rolle, NHMUK 20130387 / 6; China, Hunan, Hungerford coll, NHMUK 1891.3.17.838.9; Hupé, Gredler det., NHMW 22924 / 2; Südchina, Gerstenbrandt coll, NHMW 74649; China, Hupé, coll. Klemm, NHMW 109202 / 1; China, Patung, Hupei, coll. Naegele, SMF 342542 / 1 (mixed sample with P. nitens n. sp.: SMF 50008); Tonkin, probably leg. Messager, NHMUK 1907.5.20.86 – 7; Tonkin, Lao Kay, probably leg. Messager, NHMUK 1910.1.21.7 – 9; Tonkin, Trinh Thuong, probably leg. Messager, NHMUK 1909.7.2.35; Tonkin, Nat Son, probably leg. Messager, NHMUK 1909.7.2.23; Tonkin, Muong Hum, probably leg. Messager, NHMUK 1910.1.21.4 – 6; Tonkin, Muong Hum, probably leg. Messager, NHMUK 1909.1.2.33; Tonkin, J. J. MacAndrew coll, NHMUK 20130388 / 2; Tonkin, Lao Kay, probably leg. Messager, E. R. Sykes coll, Acc. no. 1825, NHMUK 20130389 / 2; Tonkin, That Khé, probably leg. Messager, J. J. MacAndrew coll, ex Rolle 27 / 11 / 09, NHMUK 20130390 / 2; Tonkin, Muong Hum, probably leg. Messager, J. J. MacAndrew coll, NHMUK 20130391 / 2; Tonkin, J. J. MacAndrew coll, ex Rolle 27 / 11 / 09, NHMUK 20130392 / 2; Tonkin, Lao Kay, probably leg. Messager, E. R. Sykes coll, Acc. no. 1825, NHMUK 20130393 / 2; Tonkin, Lao Kay, probably leg. Messager, E. R. Sykes coll, Acc. no. 1825, NHMUK 20130394 / 2; Tonkin, J. J. MacAndrew coll, NHMUK 20130395 / 2; Tonkin, Phong Tho, probably leg. Messager, J. J. MacAndrew coll, NHMUK 20130396 / 1; Tonkin, Phong Tho, probably leg. Messager, E. R. Sykes coll, Acc. no. 1825, NHMUK 20130397 / 1; Cam Duong, Tonkin, leg. Messager, excoll Dr. W. Blume, Dr. Leo P. Rušnov coll, NHMW 109200 (1 juvenile shell); Tonkin, That-Khé, coll. C. Dosch, ex H. Rolle, SMF 192250 / 2; Tonkin, That-Khé, coll. Jaeckel, SMF 201186 / 1; Tonkin, Muong-Hum, coll. C. Dosch ex H. Rolle ex Messager, SMF 192248 / 3; Tonkin, coll. Staadt 1969, MNHN 2012 - 26938 / 2; Tonkin, Trinh-Thuong, coll. Denis 1945, MNHN 2012 - 26939 / 6; Tonkin, Muong-Hum, coll. Denis 1945, MNHN 2012 - 26940 / 2 (mixed sample with P. maasseni n. sp.: MNHN 2012 - 26941); Tonkin, coll. Denis 1945, MNHN 2012 - 26945 / 14 (mixed sample with P. nitens n. sp.: MNHN 2012 - 26946); Tonkin, coll. Denis 1945, MNHN 2012 - 26947 / 7; Tonkin, leg. Messager, MNHN 2012 - 26949 / 16; Tonkin, Pac-Kha, leg. Messager, MNHN 2012 - 26950 / 35 (some of them with strange, strong ribs and reflexed peristome); Tonkin, Nat-Son, leg. Messager, MNHN 2012 - 26951 / 16 (1 adult, 15 juveniles); Tonkin, Muong-Bo, Dinh Lu, leg. Messager, MNHN 2012 - 26954 / 39 (including several juveniles); Tonkin, Muong-Hum, leg. Messager, MNHN 2012 - 26956 / 1 (mixed sample with P. maasseni n. sp.: MNHN 2012 - 26956); Tonkin, leg. Messager, MNHN 2012 - 26958 / 21; Tonkin, leg. Messager, MNHN 2012 - 26961 / 1 corroded shell; Indochine, leg. Messager, MNHN 2012 - 26964 / 1; Indochine, leg. Messager, MNHN 2012 - 26967 / 5 damaged shells (mixed sample with P. sophiae n. sp.: MNHN 2012 - 26967); Indochine, leg. Messager, MNHN 2012 - 26968 / 4 (mixed sample with P. sophiae n. sp.: MNHN 2012 - 26970 and P. nitens n. sp.: MNHN 2012 - 26969); Indochine, leg. Messager, MNHN 2012 - 26983 / 2 shells and few opercula; Indochine, leg. Messager, MNHN 2012 - 26984 / 2; Indochine, leg. Messager, MNHN 2012 - 26986 / 23; Tonkin, leg. Messager, MNHN 2012 - 26989 / 4; Tonkin, Muong-Hum, between Muong-Hum and the clouds pass, near the latter one, more than 2000 m a. s. l., leg. Messager, MNHN 2012 - 26990 / 11; same data, MNHN 2012 - 26991 / 3; same data, MNHN 2012 - 26992 / 8 (mixed sample with P. nitens n. sp.: MNHN 2012 - 27022); same data, MNHN 2012 - 26993 / 3; same data, MNHN 2012 - 26994 / 6; same data, MNHN 2012 - 26995 / 7 (with very strong ribs); same data, MNHN 2012 - 26996 / 4; same data, MNHN 2012 - 26997 / 11; same data, MNHN 2012 - 26998 / 5; same data, MNHN 2012 - 26999 / 6; same data, MNHN 2012 - 27000 / 8; same data, MNHN 2012 - 27001 / 2; same data, MNHN 2012 - 27002 / 1; same data, MNHN 2012 - 27003 / 45; Tonkin, Muong-Hum, leg. Messager, MNHN 2012 - 27004 / 132; Indochine, leg. Messager, MNHN 2012 - 27005 / 143 (mixed sample with P. nitens n. sp.: MNHN 2012 - 27005); Tonkin, leg. Messager, MNHN 2012 - 27007 / 78; Tonkin, leg. Messager, MNHN 2012 - 27008 (at least 300 juvenile shells, some of them seems to be P. nitens n. sp.); Tonkin, Pakhé, leg. Messager, MNHN 2012 - 27009 / 263 (mixed sample with P. sophiae n. sp.: MNHN 2012 - 27010 and P. ni t e n s n. sp.: MNHN 2012 - 27011); Tonkin, leg. Messager, MNHN 2012 - 27013 / 84; Tonkin, leg. Messager, MNHN 2012 - 27014 / 47; Tonkin, leg. Messager, MNHN 2012 - 27015 / 263 (mixed sample with P. nitens n. sp.: MNHN 2012 - 27015); Tonkin, Muong-Hum, leg. Messager, MNHN 2012 - 27017 / 38; Tonkin, coll. Denis 1945, MNHN 2012 - 27018 / 1 (mixed sample with P. sophiae n. sp.: MNHN 2012 - 26935 and P. ni t e n s n. sp.: MNHN 2012 - 26936); Tonkin, Muong-Hum, NHMUK 1909.7.2.30 / 1 (labelled as " co-type " of messageri); Tonkin, Nat-Son, 1909.7.2.27 – 28 / 2 (labelled as " co-type " of latebasis); Tonkin, Muong-Hum, NHMUK 1909.7.2.31 / 1 (labelled as " co-type " of foveolata). New material: Vietnam, Ninh Binh, Prov., Cuc Phuong Nat. Park, 2 km road Xom Bong to HQ, 20.21 ° N, 105.54 ° E, primary forest, limestone outcrops in thick clayey soil. Handpicked + soil sample, leg. JJ Vermeulen & L. Deharveng, 10.10.1998, NHMUK 1999.1.4.27 / 1; China, Hubei, Enshi Tujiazu Miaozu Zizhizhou, Enshi Shi, Mufu Cun SW 3 km, Enshi Daxiagu, exit of the stone path, 500 m, leg. Hunyadi, A., 5.11.2010, HA / 1; China, Sichuan, Dujiangyan Shi, Taian Zhen, Qingchenghoushan, Cuiyinghu to upper station of Jinli cable station, 1273 m, 30 ° 56.27110 ' N, 103 ° 28.75198 ' E, leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 0 9.07.2007., OK / 1; China, Sichuan, Dujiangyanshi, Qingchengshan Zhen, below Woyunsi, 1110 m, N 30 ° 54.33919 ', E 103 ° 33.40153 ', leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 10.07.2007., JUO / 1, OK / 1; Vietnam, L ạng Sơn Prov., Na S ầm 12 km towards Th ất Khê, left side of the road 210 m, 22 ° 07.870 ' N 106 ° 35.038 ' E, leg. Hunyadi, A., 12.11.2011, HA / 1; Vietnam, L ạng Sơn Prov., Đèo Lũng Ph ầy, Th ất Khê N 13 km, 475 m, 22 ° 20.363 ' N, 106 ° 27.098 ' E, leg. Hunyadi, A. 15.11.2011., HA / 2; China, Yunnan, Yiliang Xian, Jiuxiang Fengjingqu, 1818 m, 25 ° 04.27306 ' N, 103 ° 22.87032 ' E, leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 10.06.2005., OK / 1, PGB / 1; China, Yunnan, Wenshan Zhuangzu Miaozu Zizhizhou, Guangnan Xian, Shuguang Xiang, Kongshan Cun, 1556 m, 23 ° 42.950 ' N, 105 ° 04.570 ' E, leg. Hunyadi, A., 23.03.2011, HA / 33, PGB / 3; Northern Vietnam, Plateau de Ta Phin, Nam Cay a' Tsin Hō (= Sin Ho), leg. Saurin, MNHN 2012 – 27019 / 1; China, Yunnan, Honghe Hanizu Yizu Zizhizhou, Pingbian Miaozu Zizhixian, Pingbian Daweishan Yuanshi Senlin Gongyuan, 2085 m, 22 ° 54.835 ' N, 103 ° 41.958 ' E, leg. Hunyadi, A., 27.03.2011., HA / 3; Vietnam, Son La Prov., ca. 32 km from Moc Chau to Hanoi (old road), 20 ° 47.351 ' N, 104 ° 50.063 ' E, leg. Hemmen, Ch. & H., 0 2.04.2010., HE / 1; same data, 0 7.10.2010., HE / 1; Vietnam, Lai Chau Prov., ca. 2 km from Patan to Phong To (right off road), 22 ° 28.493 ' N, 103 ° 12.881 ' E, leg. Hemmen, Ch. & H., 0 8.10.2011., HE / 1; Vietnam, Cao Bang Prov., ca. 34.5 km from Cao Bang to Dong Khe (left off road), 22 ° 27.439 ' N, 106 ° 24.994 ' E, leg. Hemmen, Ch. & H., 26.03.2010., HE / 1.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F74B31CFF7DFF04FC0EFE35.taxon	discussion	Remarks. In our understanding, Pseudopomatias amoenus is a very widely distributed species that has several local morphologically distinct forms. Here we handle Fargesia cornea and Pseudopomatias fulvus as synonyms of P. amoenus, but provide the description of the type series. Yen (1939) considered P. amoenus tumidula as the synonym of P. amoenus. The lectotype of P. amoenus tumidula is more corpulent than P. amoenus specimens from Badong, but the paralectotype does not differ from them. The location of the type locality of P. amoenus tumidula is questionable because Hêng-shan-hsien (Hengshan Xian) is situated in eastern Hunan (part of Hengyang Shi), whereas Gui-yang-dshou (Guiyang Xian) is in southeastern Hunan (part of Chenzhou Shi). The two localities are about 130 km from each other (Szekeres M., pers. comm.). P. amoenus was recorded from Nanling Guojia Gongyuan (National Park) in Northern Guangdong by Hu et al. (2004). We have not seen the specimens collected there; therefore this information could not be verified. The specimens collected 13 km north from Th ất Khê are identical with the holotype of P. f u l v us. Typical fulvus has thicker peristome than most Vietnamese and Chinese P. amoenus populations. The species was described from the Manson (Mau Son) Mts., eastern Lang Son Province, Vietnam. We collected the species in two localities in northern Lang Son. We were unable to locate the locality published by Dautzenberg and Fischer (1908) (Deux- Ponts = " two bridges "). Ð ỗ et al. (2010) reported P. fulvus from the Xuan Son National Park (Phu Tho Province, Vietnam). We received photos of three specimens from Dr. Đ ỗ Văn Như ợng. Those specimens are similar to the North Vietnamese shells in terms of shell shape and rib density, but have much thicker peristome and the ribbing on the last quarter of whorl is more widely-spaced than on the preceding areas. Specimens from Jiuxiang Fengjingqu (Fig. 12 C) are very small, spindle-shaped; ribs blunt, very regular on the whole shell, aperture large, deformed rounded, apertural margin slim. Specimens from Kongshan Cun (Fig. 12 D) are tall, elongated turriform; ribs sharp, very regular on the whole shell, aperture large, deformed rounded, apertural margin slim. Most shells collected along the Vietnamese-Chinese border (Figs 12 H – J) are very large, turriform, sometimes with extremely fine spiral structure between the ribs is visible; peristome with angled parietal-columellar transition, apertural margin slim. In museum collections and newly collected material Pseudopomatias amoenus was frequently in mixed lots with P. maasseni n. sp., P. n i t e ns n. sp. and P. s o ph i a e n. sp. In the Cuc Phuong National Park P. amoenus lives sympatrically with Vargapupa oharai n. sp.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F74B31CFF7DFF04FC0EFE35.taxon	distribution	Distribution (Fig. 11): The species inhabits a large geographic area of over 1300 km from Northern Vietnam (Ninh Binh Province) until the Chinese Chongqing and Hubei Provinces.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F48B31FFF7DFDFDFE92FD4D.taxon	diagnosis	Diagnosis. Shell small to medium sized, spindle-shaped; thin, ribs relatively strong, very regular on the whole shell, area between the ribs glossy; aperture " fat drop-shaped " (only the parietal-palatal transition is angled), apertural margin thin but reflexed.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F48B31FFF7DFDFDFE92FD4D.taxon	description	Description. Shell very thin, translucent, yellowish or greenish, spindle-shaped; 6 – 6.75 (n = 3) bulging whorls are separated by deep suture; protoconch consist of about 2 whorls; the sculpture of the first half-first whorl is smooth, finely granulated, than becomes regularly, very finely ribbed; teleoconch is regularly, finely ribbed even on the entire last whorl, no spiral lines visible; ribs normally developed, rather high but flat; on the last quarter of whorl they are sharper; aperture " fat drop-shaped " (columellar-parietal transition smooth, parietal-palatal transition angled), apertural rim slightly thickened and very much reflexed, consists of a weaker, slimmer inner and a wider outer circle; the boundary between the circles is hardly or sometimes not visible; parietal callus normally developed but weaker than the apertural rim; no plicae are visible inside the body whorl. Measurements (in mm). H: 4.9 – 8.6, D: 2.8 – 3.3 (n = 5, from different populations). Reproductive anatomy (Figs 14 A – C): The examined female specimen had very fragile (decayed?) body, therefore only limited information can be given for the reproductive anatomy. The exact boundary of the uterus and the bursa copulatrix could not be seen. The distal, thickened end of the bursa copulatrix was also damaged. Rectum thick, attached to the uterus, all parts of the genitalia white or off-white, the inner wall of the bursa copulatrix is irridescent. Uterus large, tapers towards the distal end; bursa copulatrix long, slowly tapering toward its end; no distinct oviduct was detected (probably because of the decayed condition). A small spermatophore was found in the stalk of the bursa copulatrix (Figs 14 B – C). It consisted of a larger, oval and a smaller, tumid portion. Radula (Figs 14 D – E): Radular teeth arranged in v-shaped rows, each transverse row with seven teeth (2 - 1 - 1 - 1 - 2). Central tooth with well-developed, pointed central cusp and one, only slightly smaller, pointed lateral cusp on each side; lateral teeth with five cusps; the cusps are roughly of the same size, however the second cusp from lateral direction is usually the largest. Differential diagnosis. Pseudopomatias eos has a thinner shell, more regular, stronger ribs and a more strongly reflexed peristome margin than most populations of P. amoenus. The aperture is more rounded than in amoenus, and it has a characteristic " fat drop " shape because of the smooth columellar-parietal transition and the angled parietal-palatal transition. P. amoenus usually possesses irregularly shaped aperture with two recognisable angles.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F48B31FFF7DFDFDFE92FD4D.taxon	materials_examined	Type material. Lectotype, Hotawa, Taiwan, ANSP 89917. Museum material. Asia, Japan, Formosa, Ako, coll. Yoichiro Hirase (date unknown) ex Barbour, MCZ 42909 (5 shells). New material: Taiwan, Xinzhu Xian, Jianshi Xiang, Shangyulao, 1300 m asl, leg. Okubo, K., 7.8.1998., OK / 1; Taiwan, Nantou Xian, Lugu Xiang, Xitou, Fenghuangshan Dengshandao, leg. Okubo, K., 24.8.1989., OK / 2; Taiwan, Nantou Xian, Xinyi Xiang, Dongpu, Caijiangpubu, 1420 m, leg. Okubo, K., 19.8.1991., OK / 2; Taiwan, Nantou Xian, Renai Xiang, Songgang, leg. Okubo, K., 22.8.1990., OK / 2; Taiwan, Nantou Xian, Renai Xiang, Meifeng, leg. Okubo, K., 22.8.1995., OK / 2; Taiwan, Nantou Xian, Renai Xiang, 8 Hao Hehuansuidao, leg. Okubo, K., 6.8.1998., OK / 2; Taiwan, Nantou Xian, Renai Xiang, Dayuling, leg. Okubo, K., 2565 m, 12.8.1998., OK / 2; Taiwan, Gaoxiong Xian, Taoyuan Xiang, Tengzhisenlinyoulequ, leg. Okubo, K., 28.6.1997., OK / 3; Taiwan, Pingdong Xian, Majia Xiang, Majiacun, 780 m, leg. Okubo, K., 21.8.1994., OK / 2; Taiwan, Hualian Xian, Xiulin Xiang, Guanyuan, 2300 m, leg. Okubo, K., 17.8.1991., OK / 2; Hualian Xian, Xiulin Xiang, Bilushenmu, leg. Okubo, K., 10.8.1998., OK / 2; Taiwan, Nantou Xian, Renai Xiang, Songgang, leg. Ohara, K., 0 3.05.1982., OK / 5, PGB / 1; Hualien Xian, Shoufeng Xiang, Chinan Cun, Liyu Tan, Huantan Rd. 800 m, 165 m asl, 23 ° 56.098 ' N 121 ° 30.755 ' E, leg. Hunyadi, 0 7.10.2014. (2014 / 77), HA / 10; Nantou Xian, Renai Xiang, Meifeng SE 600 m, sideroad 600 m, 2065 m asl., 24 ° 05.474 ' N, 121 ° 10.570 ' E, leg. Hunyadi 20.10.2014. (2014 / 107), HA / 7; Nantou Xian, Renai Xiang, Meifeng-Songgang road, around km 14, 2120 m asl., 24 ° 05.103 ' N 121 ° 10.257 ' E, leg. Hunyadi, 20.10.2014. (2014 / 108), HA / 2 (used for molecular study).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F48B31FFF7DFDFDFE92FD4D.taxon	distribution	Distribution (Fig. 11): The type locality (Hotawa) is situated near Jiaxian, Kaohsiung County, southern Taiwan (see Wu et al. 2008). Additional material suggests that the species inhabits almost the whole island.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F48B31FFF7DFDFDFE92FD4D.taxon	discussion	Remarks. The shells from Tengzhisenlinyoulequ are larger (8.5 – 8.7 mm), lighter in colour, their shape is slightly more tumid and have less reflexed apertural lip. More material is necessary to see if they belong to a different species.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4BB31FFF7DFD25FB5BF8A3.taxon	diagnosis	Diagnosis. A medium sized, pale, slender turriform species with strong ribs, conspicuously thickened and reflexed peristome and plicae inside the last whorl.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4BB31FFF7DFD25FB5BF8A3.taxon	description	Description. Shell pale, whitish to corneous, slender turriform; the 8 – 8.5, strongly bulging whorls are separated by very deep suture; first whorl of the protoconch smooth; other whorls very finely, regularly ribbed (similar to the teleoconch); boundary between protoconch and teleoconch hardly visible; ribs are regular on the whole shell, just behind the aperture they became more widely spaced; aperture rounded with very slightly angled columellar-parietal and parietal-palatal transitions; apertural margin white, very much thickened and reflexed, consist of an inner, slimmer circle and an outer, thicker and reflexed circle, with hardly visible boundary; parietal callus well developed. On the dorsal side of the body whorl there are four plicae of about the same size; all of them are visible in an oblique view from the aperture. First and second plicae are standing in the same line but the third and fourth are situated more dorsally. The uppermost plica is almost parallel with the upper suture, slightly descending towards the aperture. The gap between the first and second plica is very large, but the second, third and fourth plicae are situated very close to each other. Second plica horizontal or descending backwards, slim; third and fourth plicae are slightly curved with ends pointing in the direction of the umbilicus. Measurements (in mm). H: 8.1 – 8.7, D: 3.4 – 4 (n = 4). Differential diagnosis. In terms of general shell shape, P. linanprietoae n. sp. is similar to P. himalayae and P. sophiae n. sp., but it can be distinguished from these species on the basis of the pale shell, conspicuously thickened apertural rim and the presence of inner plicae. See also under P. nit ens n. sp.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4BB31FFF7DFD25FB5BF8A3.taxon	materials_examined	Material: Laos, Luang Prabang Prov., Nong Kiau, excoll Jose Ahuir Galindo, HNHM 98835 (holotype), HA / 1 paratype, JAG / 2 paratypes, NHMUK 20140669 / 1 paratype, PGB / 2 paratypes. Type locality. Laos, Luang Prabang Prov., Nong Kiau.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4BB31FFF7DFD25FB5BF8A3.taxon	etymology	Etymology. The new species is dedicated to Carmen Linan Prieto, the wife of Jose Ahuir Galindo, who provided the material.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4BB31FFF7DFD25FB5BF8A3.taxon	distribution	Distribution (Fig. 15): The new species is known from the type locality only.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4BB31FFF7DFD25FB5BF8A3.taxon	discussion	Remarks. Due to the few available shells we have not opened the shells to see possible inner plicae. There may be more, deeper situated plicae within the last whorl (see Remarks on shell characters).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4DB318FF7DF963FAD5F80C.taxon	diagnosis	Diagnosis. A large, slender, almost cylindrical species with thick apertural margin, rather flattened whorls and characteristic ribbing around the umbilicus (every second ribs are prominent).	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4DB318FF7DF963FAD5F80C.taxon	description	Description. Shell brownish or greyish, slender turriform or cylindrical, usually the penultimate whorl is the widest, or the last three whorls are of almost the same width; the 9.75 – 10.25, slowly increasing, moderately bulging or sometimes rather flat whorls are separated by shallow suture; suture is a bit deeper between the apical whorls; last whorl conspicuously flatter than the penultimate whorls; protoconch matt, finely granulated; teleoconch finely, regularly ribbed; last half whorl is irregularly ribbed; on the basal side of the shell (around the umbilicus) every second ribs are wider and stronger; aperture usually rounded with not or very slightly angled columellar-parietal transition and slightly angled parietal-palatal transition; peristome very much thickened but not reflexed, consists of two circles separated from each other; the circles are of different width, the outer is thicker than the inner one; suture shallow between the circles. Measurements (in mm). D: 11.4 – 13.6, H: 4.0 – 4.7 (sample from the type locality, n = 4), H: 14.6 – 14.7, D: 4.4 – 4.5 (NHMUK 20130399, n = 2). Differential diagnosis. P. maasseni n. sp. can be distinguished by other members of the genus by the tall, slender shell, shallow suture, rather quadrangular whorls and the every second elevated ribs on the base. Also see under P. amoenus and Table 6.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4DB318FF7DF963FAD5F80C.taxon	materials_examined	Material. Vietnam, Sơn La Prov., Hà N ội 156 km towards M ộc Châu, right side of the road nr. 6., rocky wall, 1110 m, 20 ° 46.085 ' N 104 ° 53.888 ' E, leg. Hunyadi, A., 0 6.06.2012, HNHM 98833 (holotype) HA / 28 paratypes (2012 / 62; one of them was used for DNA analysis), PGB / 2 paratypes; Vietnam, Sơn La Prov., Hà N ội 156 km towards M ộc Châu, left side of the road nr. 6, rocky wall, 1110 m, 20 ° 45.993 ' N 104 ° 53.868 ' E, leg. Hunyadi, A., 0 6.06.2012, HA / 1 paratype; Vietnam, Sơn La Prov., ca. 32 km from M ộc Châu, to Hà N ội (old road), 20 ° 47.351 ' N, 104 ° 50.063 ' E, leg. Hemmen, Ch. & J., 0 2.04.2010., HE / 6 paratypes, PGB / 2 paratypes; same locality, 15.10.2011., HE / 4 paratypes; same locality, 0 7.10.2010., HE / 3 paratypes; Vietnam, Son La Prov., ca. 34 km from Moc Chau to Mai Chau, 20 ° 45.219 ' N, 104 ° 54.458 ' E, leg. Hemmen, Ch. & H., 15.10.2011., HE / 1 paratype; China, Guangxi, Napo Xian, Pohe Xiang, left side of road to Longhe, 1223 m, 23 ° 20.84445 ' N, 105 ° 54.60324 ' E, leg. Ohara, Okubo & Otani, 10.01.2006. OK / 1 paratype; Tonkin, J. J. MacAndrew coll, NHMUK 20130398 / 1 paratype; Tonkin, Phong Tho, J. J. MacAndrew coll, NHMUK 20130399 / 2 paratype; Tonkin, Phong Tho, E. R. Sykes coll, Acc. no. 1825, NHMUK 20130400 / 2 paratype; Tonkin, Phong Tho, B. R. Lucas coll, Acc. no. 2351, NHMUK 20130401 / 1 paratype; Tonkin, Phong Tho, J. J. MacAndrew coll, Sr. F. 21 / 4 / 09, NHMUK 20130402 / 2 paratypes; Tonkin, Phong-Tho, coll. C. Dosch ex H. Rolle, SMF 192249 / 2 paratypes; Tonkin, Muong-Hum, coll. Denis 1945, MNHN 2012 - 26941 / 4 paratypes (mixed sample with P. amoenus: MNHN 2012 - 26940); Tonkin, Phong-Tho, coll. Denis 1945, MNHN 2012 - 26943 / 10 paratypes; Tonkin, Phong-Tho, coll. Staadt 1969, MNHN 2012 - 26948 / 4 paratypes; Tonkin, leg. Messager, MNHN 2012 - 26955 / 30 paratypes; Tonkin, Muong-Hum, leg. Messager, MNHN 2012 - 26957 / 1 paratype (mixed sample with P. amoenus: MNHN 2012 - 26957); Indochine, leg. Messager, MNHN 2012 - 26985 / 1 paratype; Tonkin, Phong-Tho, NHMUK 1910.9.28.172 – 173, NHMUK 1909.7.2.26, NHMUK 1909.10.19.71 – 72 (altogether 5 paratypes) (labelled as " co-type " of buliminiformis); Vietnam, Bac Can Prov., ca. 1 km from Ba Bé N. P. headquarters to Ba Bé Lake, ca. 286 m, 22 ° 24.829 ' N, 105 ° 37.652 ' E, leg. Hemmen, Ch. & H., 20.10.2010., HE / 2 paratypes; Vietnam, Ha Giang Prov., ca. 7.5 km from Dong Van to Meo Vac (right side off road), 23 ° 14.906 ' N, 105 ° 23.445 ' E, leg. Hemmen, Ch. & H., 18.10.2010., HE / 1 paratype; Vietnam, Son La Prov., 6 km from Son La to Thuan Chau (left side off road), 21 ° 19.502 ' N, 103 ° 53.055 ' E, leg. Hemmen, Ch. & H., 0 8.10.2010., HE / 1; Tonkin, Phong-Tho, NHMUK 1909.7.2.32 / 1 paratype (labelled as " syntype " of buliminiformis var. minor); Tonkin, Phong-Tho, NHMUK 1910.9.28.170 – 171 / 2 paratypes (labelled as " syntype " of buliminiformis var. minor). Type locality. Vietnam, Sơn La Prov., Hà N ội 156 km towards M ộc Châu, right side of the road nr. 6., rocky wall, 1110 m, 20 ° 46.085 ' N 104 ° 53.888 ' E.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4DB318FF7DF963FAD5F80C.taxon	etymology	Etymology. The new species is dedicated to and named after Wim J. M. Maassen, malacologist, the friend of the first author, the outstanding expert of South-East Asian land snail fauna.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4DB318FF7DF963FAD5F80C.taxon	distribution	Distribution (Fig. 15): The species is known from Northern Vietnam (Sơn La, Lai Châu and Lào Cai Provinces) and from one locality in Guangxi, China.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4DB318FF7DF963FAD5F80C.taxon	discussion	Remarks. In museum collections Pseudopomatias maasseni n. sp. was frequently in mixed lots with P. amoenus. Some samples (MNHN 2012 - 26941, MNHN 2012 - 26957, MNHN 2012 - 26985) are characterized by very robust, corpulent shells. The species is still recognisable and can be assigned to this species, and they are distinguishable from P. amoenus on the basis of relatively flat whorls and the every second stronger ribs around the umbilicus. A single, straight, long plica is visible above the aperture in the fresh, translucent shells from the type locality. This plica has not been detected in specimens from other sites. The shells from the Vietnamese Hà Giang and B ắc K ạn Provinces and the single Chinese specimen has the last whorl slightly flattened from " dorsobasal " direction. This results in the formation of a blunt keel, which nevertheless situated closer to the umbilicus than that of the genus Vargapupa gen. n. This keel-like structure in some P. maasseni n. sp. shells and the strong keel of Vargapupa gen. n. are probably not homologous.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4FB31AFF7DFB79FEF0F85C.taxon	diagnosis	Diagnosis. A small to medium sized, glossy, rather turriform species with plicae inside the last whorl.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4FB31AFF7DFB79FEF0F85C.taxon	description	Description. Shell glossy, yellowish to brownish (often " rusty ") in colour, rather turriform (sometimes spindle-shaped); the last or the penultimate whorl is the widest; 6.25 – 6.75 moderately to very much bulging whorls are separated by a rather deep suture which is even deeper between the upper whorls; protoconch consist of 1.5 whorls, the first half is finely tuberculated, glossy, the remaining one whorl is regularly and very finely ribbed; teleoconch regularly, finely ribbed, until the aperture; umbilicus slit-like, very narrow; most specimens with three plicae on the dorsolateral side of the body whorl (for description of the inner plicae see " Remarks on the shell characters " and Fig. 4); aperture rounded, with not of very slightly angled columellar-parietal transition and slightly angled parietal-palatal transition; apertural rim paler than the shell colour, it is thickened and very slightly reflexed; parietal callus slightly thickened; rim double, consists of a weaker inner and a stronger outer circle separated by a deep suture. Measurements (in mm). H: 6.0 – 8.7, D: 2.8 – 3.3 (different Vietnamese samples, n = 4); H: 8.3 – 8.9, D: 3.4 – 3.7 (shells from Changyang, n = 4); H: 7.4 D: 3.4 (HNHM 98834, Sichuan, n = 1). Differential diagnosis. Pseudopomatias nitens n. sp. differs from P. sophiae n. sp. by the more corpulent, spindle-shaped shell, the lower and denser ribs, the thinner apertural margin and the presence of inner plicae above the umbilicus in most nitens specimens. P. linanprietoae n. sp. is lighter in colour, larger and has a thicker apertural margin. P. nitens n. sp. differs from P. amoenus by the smaller, glossy, spindle-shaped shell, more bulging whorls, stronger ribs, the absence of spiral lines between the ribs (sometimes present in P. amoenus), the more slender, relatively smaller shell, the more regularly shaped aperture, thicker peristome and the presence of two to three palatal plicae above the umbilicus that are visible throughout the semi-transparent shell. See also Table 6.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4FB31AFF7DFB79FEF0F85C.taxon	materials_examined	Material. Tonkin, coll. Denis 1945, MNHN 2012 - 27023 / 1 holotype, MNHN 2012 - 26931 / 31 paratypes and 2 juveniles (not paratypes); Muong Kong, Tonkin, NHMUK 1909.7.2.36 / 1 paratype; Long Ping, Tonkin, NHMUK 1909.7.2.40 / 1 paratype; Tonkin, Pac-Kha, probably leg. Messager, NHMUK 1909.7.2.24 / 1 paratype; Tonkin, Long Ping, probably leg. Messager, NHMUK 1909.7.2.25 / 1 paratype; Muong-Hum, Tonkin, E. R. Sykes coll., Acc. no. 1825, NHMUK 20130424 / 1 paratype; Tonkin, Muong-Hum, coll. Letellier 1949, MNHN 2012 - 26932 / 2 paratypes; Tonkin, coll. Denis 1945, MNHN 2012 - 26936 / 3 paratypes (mixed sample with P. sophiae n. sp.: MNHN 2012 - 26935 and P. amoenus: MNHN 2012 - 27018); Tonkin, Coll. Denis 1945, MNHN 2012 - 26942 / 1 not paratype, very slender shell without plicae; Tonkin, Coll. Denis 1945, MNHN 2012 - 26944 / 4 paratypes and 2 juvenile shells (not paratypes, one of them is very large, possibly P. amoenus); Tonkin, coll. Denis 1945, MNHN 2012 - 26946 / 14 paratypes (mixed sample with P. amoenus: MNHN 2012 - 26945); Tonkin, pris du Col de Nuages (2000 m altitude), Tonkin, Muong-Hum, between Muong-Hum and the Clouds Pass (Col de Nuages), near the latter one, more than 2000 m a. s. l., leg. Messager, MNHN 2012 - 27022 / 1 paratype (mixed sample with P. amoenus: MNHN 2012 - 26992); leg. Messager, MNHN 2012 - 26953 / 7 paratypes (mixed sample with P. sophiae n. sp.: MNHN 2012 - 26952); Tonkin, leg. Messager, MNHN 2012 - 26960 / 1 paratype, 2 damaged shells (not paratypes) (mixed sample with P. sophiae n. sp.: MNHN 2012 - 26959); Indochine, leg. Messager, MNHN 2012 - 26962 / 1 corroded shell without plicae (not paratype); Indochine, leg. Messager, MNHN 2012 - 26969 / 3 paratypes and one juvenile shell, which is not paratype (mixed sample with P. amoenus: Messager, MNHN 2012 - 26968 and P. sophiae n. sp. Messager, MNHN 2012 - 26970); Indochine, leg. Messager, MNHN 2012 - 26972 / 2 paratypes (mixed sample with P. sophiae n. sp.: MNHN 2012 - 26971); Indochine, leg. Messager, MNHN 2012 - 26974 / 1 paratype (mixed sample with P. sophiae n. sp.: MNHN 2012 - 26973); Indochine, leg. Messager, MNHN 2012 - 26978 / 3 shells without plicae, not paratypes (mixed sample with P. sophiae n. sp.: MNHN 2012 - 26977); Indochine, leg. Messager, MNHN 2012 - 26979 / 3 paratypes (mixed sample with P. sophiae n. sp.: MNHN 2012 - 26980); Indochine, leg. Messager, MNHN 2012 - 26982 / 4 paratypes (mixed sample with P. sophiae n. sp.: MNHN 2012 - 26981); Tonkin, leg. Messager, MNHN 2012 - 26987 / 1 shell without plicae, not paratype (mixed sample with P. sophiae n. sp.: MNHN 2012 - 26988); Indochine, leg. Messager, MNHN 2012 - 27006 / 20 paratypes (mixed sample with P. amoenus: MNHN 2012 - 27005); Tonkin, Pakhé, leg. Messager, MNHN 2012 - 27011 / 7 paratypes (mixed sample with P. amoenus: MNHN 2012 - 27009 and P. sophiae n. sp.: MNHN 2012 - 27010); Tonkin, leg. Messager, MNHN 2012 - 27016 / 2 paratypes (mixed sample with P. amoenus: MNHN 2012 - 27015); China, Changyang, B. R. Lucas coll, Acc. no. 2351, NHMUK 20130425.1 (not paratype), NHMUK 20130425.2 – 4 (not paratypes); Changya, Hupei, China, coll. Boettger, excoll Schmacker 1893, SMF 45650 / 8 (not paratypes); China, Patung, Hupei, coll. Naegele, SMF 50008 / 2 (not paratypes; mixed sample with P. amoenus: SMF 342542); China, Changya, coll. S. H. Jaeckel, SMF 211185 / 2 (not paratypes); Changya, Carl Bock coll, NHMUK 1894.4.24.94 – 7 / 4 (not paratypes); Changyang, J. J. MacAndrew coll, NHMUK 20130426 / 3 (not paratypes); Changyang, E. R. Sykes coll, Acc. no 1825, NHMUK 20130427 / 2 (not paratypes); Changyang, excoll Dr. W. Blume, Dr. Leo P. Rušnov coll, NHMW 109201 (not paratypes); Changyang, NHMW 83901 / G / 8273 (not paratype); China, Sichuan, Dujiangyan Shi, Taian Zhen, Qingchenghoushan, Cuiyinghu to upper station of Jinli cable station, 1273 m, 30 ° 56.27110 ' N, 103 ° 28.75198 ' E, leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 0 9.07.2007., HNHM 98834 (not paratype), OK / 1; China, Sichuan, Dujiangyan Shi, Taian Zhen, Qingchenghoushan, above lower station of Jinli cable station, 979 m, 30 ° 55.62540 ' N, 103 ° 29.08667 ' E, leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 0 8.07.2007., OK / 1 (not paratype, juvenile specimen, it can be this species or P. amoenus); China, Sichuan, Emeishan Shi, Eshan Zhen, Emeishan, Wanniansi, 865 m, N 29 ° 34.83012 ', E 103 ° 22.97325 ', leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 14.07.2007., OK / 1 (not paratype). Type locality. Tonkin.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4FB31AFF7DFB79FEF0F85C.taxon	etymology	Etymology. We use Preston's manuscript name. The Latin nitens means shiny, glossy, glittering, bright, which is characteristic for P. n i t e ns n. sp.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4FB31AFF7DFB79FEF0F85C.taxon	distribution	Distribution (Fig. 11): The new species is known from Northern Vietnam (only the border region with Yunnan), and the Chinese Sichuan and Hubei Provinces.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F4FB31AFF7DFB79FEF0F85C.taxon	discussion	Remarks. Specimens from Changyang and Badong are taller, have more whorls, rather spindle-shaped shells, more reflexed peristome and more rounded aperture. Shells from Sichuan have tumid (" ovoid ") shape and the two circles of the peristome are equally developed and well separated. In museum collections Pseudopomatias nitens n. sp. was frequently in mixed lots with P. amoenus and P. sophiae n. sp.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F41B315FF7DFF4EFE11F8AB.taxon	diagnosis	Diagnosis. A small, slender turriform species with strong ribs and rounded aperture.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F41B315FF7DFF4EFE11F8AB.taxon	description	Description. Shell light brown to yellowish, slender turriform, apex slightly blunt, the apical part of the shell (3 – 4 th whorls) seems to be concave; shell widest at its base; the 7 – 7.25 very much bulging whorls are separated by deep suture; boundary of the protoconch and teleoconch is not visible, only the first quarter of whorl is not ornamented by ribs, the rest is with sharp, regular ribs; teleoconch also finely, regularly ribbed without spiral lines; ribs very sharp, regular even behind the aperture; aperture almost exactly round with an inconspicuous parietalpalatal transition; columellar-parietal transition not angled; apertural rim thickened and reflexed; the boundary between the two circles of the apertural rim is hardly visible. Measurements (in mm). H: 5.9 – 7.0, D: 2.7 – 3.0 (n = 3). Differential diagnosis. P. s o ph i ae n. sp. is very similar to P. himalayae in terms of rib density and morphology, aperture formation and the building of the aperture lip. It is however smaller, more slender, has deeper suture, a more rounded aperture and regular ribs on the neck region. P. amoenus is much larger, has blunter ribs and a comparatively larger aperture of " deformed " shape. See also under P. abletti n. sp., P. linanprietoae n. sp. and P. nitens n. sp. and Table 5.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F41B315FF7DFF4EFE11F8AB.taxon	materials_examined	Material. Tonkin, Trinh-Thuong, NHMUK 1910.1.21.2 (holotype), NHMUK 1910.1.21.3 (1 paratype); no locality, B. R. Lucas Coll, NHMUK 20130405 / 1 paratype; Muong Hum, Preston 81 (?) / 1.8 / 1 / 09, J. J. MacAndrew coll, NHMUK 20130406 / 1 paratype; Tonkin, Lao Kay, E. R. Sykes coll, Acc. no. 1825, NHMUK 20130407 / 3 paratypes; Tonkin, Preston (?) / 1.8 / 1 / 09, J. J. MacAndrew coll, NHMUK 20130408 / 1 paratype; Tonkin, Col de Nuages, coll. Staadt 1969, MNHN 2012 - 26933 / 3 paratypes; Tonkin, coll. Denis 1945, MNHN 2012 - 26934 / 3 paratypes; Tonkin, coll. Denis 1945, MNHN 2012 - 26935 / 5 paratypes (mixed sample with P. ni t e n s n. sp.: MNHN 2012 - 26936 and P. amoenus: MNHN 2012 - 27018); Tonkin, coll. Denis 1945, MNHN 2012 - 26937 / 3 paratypes; Tonkin, pris du Col de Nuages (2000 m altitude), leg. Messager, MNHN 2012 - 26952 / 37 paratypes (mixed sample with P. n i t e ns n. sp.: MNHN 2012 - 26953); Tonkin, leg. Messager, MNHN 2012 - 26959 / 6 paratypes (mixed sample with P. n i t e ns n. sp.: MNHN 2012 - 26960); Indochine, leg. Messager, MNHN 2012 - 26963 / 1 paratype; Indochine, leg. Messager, MNHN 2012 - 26965 / 4 paratypes; Indochine, leg. Messager, MNHN 2012 - 26966 / 2 paratypes and 1 juvenile shell (not paratype) (mixed sample with P. amoenus: MNHN 2012 - 26967); Indochine, leg. Messager, MNHN 2012 - 26970 / 1 corroded shell (not paratype) (mixed sample with P. amoenus: MNHN 2012 - 26968 and P. nitens n. sp.: MNHN 2012 - 26969); Indochine, leg. Messager, MNHN 2012 - 26971 / 2 paratypes (mixed sample with P. nitens n. sp.: MNHN 2012 - 26972); Indochine, leg. Messager, MNHN 2012 - 26973 / 3 (2 paratypes and one broken specimen which is not paratype; mixed sample with P. nitens n. sp.: MNHN 2012 - 26974); Indochine, leg. Messager, MNHN 2012 - 26975 / 2 paratypes and a juvenile shell, which is not paratype; Indochine, leg. Messager, MNHN 2012 - 26976 / 4 paratypes; Indochine, leg. Messager, MNHN 2012 - 26977 / 7 paratypes (mixed sample with P. n i t e ns n. sp.: MNHN 2012 - 26978); Indochine, leg. Messager, MNHN 2012 - 26980 / 2 shells, the juvenile is not paratypes (mixed sample with P. nitens n. sp.: MNHN 2012 - 26979); Indochine, leg. Messager, MNHN 2012 - 26981 / 14 paratypes (mixed sample with P. n i t e ns n. sp.: MNHN 2012 - 26982); Tonkin, leg. Messager, MNHN 2012 - 26988 / 1 paratype (mixed sample with P. nitens n. sp.: MNHN 2012 - 26987); Tonkin, Pakhé, leg. Messager, MNHN 2012 - 27010 / 2 paratypes (mixed sample with P. amoenus: MNHN 2012 - 27009 and P. n i t e ns n. sp.: MNHN 2012 - 27011); Tonkin, Pac-Kha, NHMUK 1909.7.2.34 / 1 paratype (labelled as " co-type "); Tonkin, Muong-Hum, NHMUK 1909.7.2.29 / 1 paratype (labelled as " co-type "). Type locality. Tonkin, Trinh-Thuong.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F41B315FF7DFF4EFE11F8AB.taxon	etymology	Etymology. This new species is named after Zsófia (Sophia) Kémenes, the mother of the first author.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F41B315FF7DFF4EFE11F8AB.taxon	distribution	Distribution (Fig. 15): The new species was collected in Northern Vietnam (Muong Hum, Lao Kay, Trinh Thuong), near the Chinese boarder by Colonel Messager.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F41B315FF7DFF4EFE11F8AB.taxon	discussion	Remarks. In museum collections Pseudopomatias sophiae n. sp. was frequently in mixed lots with P. amoenus and P. nitens n. sp.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F40B314FF7DFC9BFBEAF93D.taxon	diagnosis	Diagnosis. A medium sized, slender turriform species with a well developed basal keel.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F40B314FF7DFC9BFBEAF93D.taxon	description	Description. Shell turriform, widest at its base; only corroded shell material was available, therefore the original shell colour is unknown; there are 8 – 8.75, regularly increasing, rather flat or moderately bulging whorls; protoconch rather smooth with signs of small regular riblets; suture deeper on the upper whorls, especially at the protoconch; the coarse riblets are regular on the whole shell; sometimes on the last whorl they are less regular; keel blunt; the ribs of the body whorl are visible here too, especially close to the end of the keel; there are deep sutures on both sides of the keel; the keel starts from the back side of the most basal point of the peristome, and ends on the ventrolateral side; keel gradually becomes lower towards its end; keel visible inside the aperture as a shallow canal; aperture round with not angled columellar-parietal transition and slightly angled parietal-palatal transition; the lower incision (at the basal pont of the peristome) is the starting point of the keel; apertural rim thickened and slightly reflexed. Measurements (in mm). H: 8.9 – 10.5, D: 3.5 – 3.9 (n = 3). Differential diagnosis. Vargapupa biheli n. sp. is much smaller than V. oharai n. sp., has narrower umbilicus, more rounded aperture and only one basal keel. The sutures situated on both side of the main keel are deeper.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F40B314FF7DFC9BFBEAF93D.taxon	materials_examined	Material. Laos, Tran Ninh Province, Pa Hia, Pah Xieng Tong, coll. Saurin, MNHN 2012 – 27020 (holotype), MNHN 2012 – 27021 / 9 (paratypes). Type locality. Laos, Tran Ninh Province, Pa Hia, Pah Xieng Tong.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F40B314FF7DFC9BFBEAF93D.taxon	etymology	Etymology. The new species is named after the Biodiversity Heritage Library (www. biodiversitylibrary. org) to thank the multitude of rare literature made available to us. The name “ biheli ” is an acronym derived from the name BI odiversity HE ritage LI brary.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F40B314FF7DFC9BFBEAF93D.taxon	distribution	Distribution (Fig. 15): The new species is known from the type locality only.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F40B314FF7DFF41FC13FD0B.taxon	materials_examined	Type species. Vargapupa oharai n. sp.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F40B314FF7DFF41FC13FD0B.taxon	diagnosis	Diagnosis. Shell slender turriform, regularly ribbed; aperture rounded or slightly rectangular, with not or slightly angled columellar-parietal transition, and slightly angled parietal-palatal transition; there is a lower incision to the right of the bottom point of the aperture, at the starting point of the keel; the parietal side of the aperture usually straight; peristome simple-looking; an elevated keel runs on the basal side of the shell around the umbilicus; keel starts from the lower basal apertural lip and flattens gradually on the ventral side of the body whorl and terminated close to the columellar-palatal transition of the aperture; operculum unknown. Differential diagnosis. Differs from Pseudopomatias and Csomapupa n. gen. by the presence of the basal keel. For the differences with Nodopomatias and see there.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F40B314FF7DFF41FC13FD0B.taxon	etymology	Etymology. The new genus is dedicated to András Varga, prominent Hungarian malacologist, the friend of the authors. His contribution to Vietnamese malacofauna is very important. The name Vargapupa is the combination of the family name Varga and Pupa. Content. V. oharai n. sp., V. biheli n. sp.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F40B314FF7DFF41FC13FD0B.taxon	distribution	Distribution (Fig. 7): The two species of Vargapupa n. gen. species are known in the northern Annamese Mountains (French: Chaîne Annamitique), in Northern Vietnam and Laos.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F40B317FF7DF8F5FEC5FC09.taxon	diagnosis	Diagnosis. A very large, slender turriform, brownish species with a well-developed and a weaker basal keel.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F40B317FF7DF8F5FEC5FC09.taxon	description	Description. Shell brownish corneous or yellowish, slender turriform, widest at its base; there are 10.25 (n = 2) regularly increasing whorls; first whorl of the protoconch rather smooth, the second whorl is finely ribbed; suture deeper on the upper whorls, especially at the protoconch; the coarse riblets are regular on the whole shell; keel blunt, the ribs of the body whorl are visible here too, especially close to the end of the keel (close to the columellarparietal transition of the peristome); there are rather shallow sutures on both sides of the keel where it joins the shell surface of the last whorl; keel starts from the back side of the most basal point apertural rim, and ends on the ventrolateral side; keel gradually becomes lower towards its end; there is a second, lower keel between the main keel and the umbilicus; it also starts from the back side of the apertural rim; main keel visible inside the aperture as a shallow canal; aperture round with angled columellar-parietal and parietal-palatal transitions; the lower incision (basal part of the aperture) is the starting point of the keel; apertural rim thickened and very much reflexed. Measurements (in mm). H: 14.7 – 16.1, D: 5.3 – 5.6 (n = 2). Differential diagnosis. See under Vargapupa biheli n. sp.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F40B317FF7DF8F5FEC5FC09.taxon	materials_examined	Material. Vietnam, Ninh Binh Province, Cuc Phuong National Park, Cave of Prehistoric Man, 237 m, 20 ° 17.56874 ' N, 105 ° 40.01229 ' E, leg. Ohara, K., Okubo, K., Otani, J. U., 16.11.2007., Holotype HNHM 98837, OK / 2, JUO / 2; same data, leg. Okubo, K. & Otani, J. U., 12.04.2007., JUO / 1, PGB / 1; Vietnam, Ninh Binh Province, Cuc Phuong National Park, excoll Jose Ahuir Galindo, PGB / 1; Vietnam, Thanh Hóa Prov., Pu Luong N. P., surroundings of village Am, 20 ° 28.185 ' N, 105 ° 13.309 ' E, collected by native people, 0 5.10.2010., HE / 1 paratype. Type locality. Vietnam, Ninh Binh Province, Cuc Phuong National Park, Cave of Prehistoric Man, 146 m, 20 ° 15.53843 ' N, 105 ° 42.38950 ' E.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F40B317FF7DF8F5FEC5FC09.taxon	etymology	Etymology. Named after our friend, Kenji Ohara, who provided the material.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F40B317FF7DF8F5FEC5FC09.taxon	distribution	Distribution (Fig. 15): The new species is known from two localities in Northern Vietnam. There is approximately 60 km between the two sites.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F40B317FF7DF8F5FEC5FC09.taxon	discussion	Remarks. In the Cuc Phuong National Park Vargapupa oharai n. sp. lives sympatrically with Pseudopomatias amoenus.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F43B317FF7DFA7DFC74F9AE.taxon	discussion	Remarks. This juvenile shell was mentioned by Godwin-Austen (1917) as a probably undescribed new species having coarse ribbing not reported in any other known species.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F43B317FF7DF947FBCBF88F.taxon	materials_examined	Museum material. Shan States, NHMUK 1888.12.4.530 / 1.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
314487958F43B317FF7DF947FBCBF88F.taxon	discussion	Remarks. The single shell is very much corroded. It is much smaller than the shells of the type lot of P. shanensis n. sp. (height: 6.4 mm, width: 2.4 mm), and has flatter whorls. The sculpture is not clearly visible, but spiral lines are not detectable. Probably not conspecific with P. shanensis n. sp.	en	Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András, Asami, Takahiro (2015): Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae). Zootaxa 3937 (1): 1-49, DOI: 10.11646/zootaxa.3937.1.1
