taxonID	type	description	language	source
314287B0F635FFA6FF7BF8EC08E5FD04.taxon	vernacular_names	Chinese name: ḧẋ 毛瓢 • ƌ	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F635FFA6FF7BF8EC08E5FD04.taxon	type_taxon	Type species: Scymnus hareja Weise, 1879, by original designation.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F635FFA6FF7BF8EC08E5FD04.taxon	diagnosis	Diagnosis. Sasajiscymnus is similar to Axinoscymnus Kamiya in having three tarsomere tarsi. It also closely resembles Scymnus Kugelann in external appearance. However, it can be easily distinguished from the genera within the tribe Scymnini by its short antennae (nine antennomeres) with 2 or 3 extraordinarily long apical setae.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F635FFA6FF7BF8EC08E5FD04.taxon	description	Description. Body elongate oval, moderately convex, dorsum pubescent (Fig. 1). Body size ranging from 1.3 mm to 3.6 mm in length. The elytral color is usually black and yellow. Head exposed dorsally or withdrawn into prothorax (Fig. 2 a). Eyes dorsally 0.6 – 0.7 × length of head capsule, with ocular canthus extending slightly into eye. Frons wide and flat, not emarginate. Antennae short with 9 antennomeres (Fig. 2 e): antennomere 1 large and bent; antennomere 2 about as broad as 1 st; antennomeres 3 – 4 somewhat triangular, distinctly longer than wide; antennomere 5 transverse, broadening apically, forming a short support for club; antennomeres 6 – 9 forming a compact club, terminal antennomere with 2 or 3 long apical setae. Clypeus with anterior margin slightly arcuate. Labrum entirely exposed, transverse (Fig. 2 d). Mandible robust, with a subapical tooth just behind apical tooth, molar teeth in both mandibles are asymmetric (Figs 2 b – c). Maxilla with cardo subquadrate to weakly transverse; palpomere 2 subquadrate; palpomere 3 short; terminal palpomere short, parallel sided (Fig. 2 f). Labial palps with 3 palpomeres; apical palpomere as long as penultimate one (Fig. 2 g). Mentum cordiform, narrowed basally. Pronotum convex; anterior angle of pronotum indistinct; lateral edge smooth; basal marginal line along the edge distinct and complete (Fig. 2 h). Prosternum T-shaped; prosternal process 0.5 coxal diameter, with complete carinae joined anteriorly forming square (Fig. 2 j). Prosternum in front of coxae distinctly longer than basal width of prosternal process. Procoxal cavity transverse. Mesoventral process broad, meso-metaventral junction forming a straight line, with suture visible. Elytral epipleuron incomplete apically, surface nearly horizontal and flat without distinct foveae. Protibia cylindrical, tarsi trimerous; tarsal claws in both sexes with large basal tooth (Fig. 2 i). Tibiae without apical spurs. Abdomen with six ventrites in both sexes; ventrite 1 slightly longer than ventrite 2, posterior margin straight or weakly arcuate (Fig. 2 k). Abdominal postcoxal lines incomplete, not reaching the hind and lateral margins of ventrite 1. Penis long and siphonate, with distinct penis capsule (Fig. 2 l). Penis guide symmetrical; parameres articulated with phallobase (Figs 2 o – n). Coxites triangular or quadrangular; spermatheca sometimes with very short or indistinct ramus and nodulus (Fig. 3).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F635FFA6FF7BF8EC08E5FD04.taxon	distribution	Distribution. Asia, Africa, Oceania, North America (introduced).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F639FFA9FF7BFDAE0EDCF9D3.taxon	discussion	The Sasajiscymnus sylvaticus species group is characterized by the long and slender penis with short outer arm, apex of penis simple without appendage, penis guide sometimes bifurcated apically, interocular distance greater than eye width in frontal view, abdominal postcoxal lines recurved toward the base of ventrite, and triangular coxites usually elongate.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F639FFA8FF7BF9440B9DFD4C.taxon	description	(Figs 4, 63) Chinese name: 梭叶 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F639FFA8FF7BF9440B9DFD4C.taxon	diagnosis	Diagnosis. This species is similar to S. lishanicus sp. nov., but can be distinguished from the later by the uniformly black elytra, larger and more elongate body size, and parameres about 1 / 3 as long as penis guide (Figs 4 a – b, 4 f). In S. lishanicus, elytra are black with yellowish brown apices, body size is smaller and shorter, and parameres are about 1 / 2 as long as penis guide (Figs 11 a – b, 4 g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F639FFA8FF7BF9440B9DFD4C.taxon	diagnosis	Description. TL: 1.13 – 2.10 mm, TW: 1.34 – 1.61 mm, TH: 0.97 – 1.06 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 4 a – c). Body uniformly black, except antennae, maxillary palpi and legs somewhat yellowish brown. Interocular distance distinctly greater than eye width in frontal view. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 4 d). Male genitalia: Penis long and slender, penis capsule with short outer arm and long inner one, apex of penis pointed (Fig. 4 e). Penis guide triangular in lateral view with pointed apex; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide approximately widest at basal 1 / 3 in ventral view, bifurcated apically (Figs 4 f – g). Female genitalia: Coxites triangular; styli strongly reduced and hardly visible; spermatheca worm-like, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F639FFA8FF7BF9440B9DFD4C.taxon	materials_examined	Type material. Holotype: China: Shanxi: ♂, Xiachuan Village, Lishan National Nature Reserve, 1545 m, 9. VII. 2013, Huo LZ leg (SCAU). Paratypes (SCAU): China: Shanxi: 1 ♂, Same data as holotype. Henan: 2 ♂ 2 ♀, Baotianman National Nature Reserve, 6. VII. 2009, Wang XM leg. Shaanxi: 1 ♀, Cuihuan Mountains, Chang’an District, Xi’an City, 1300 m, 30. VII. 2009, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F639FFA8FF7BF9440B9DFD4C.taxon	distribution	Distribution. China (Shanxi, Henan, Shaanxi).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F639FFA8FF7BF9440B9DFD4C.taxon	etymology	Etymology. The specific epithet fusinus derives from the Latin word “ fusinus ” and refers to the fusiform penis guide.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63BFFABFF7BFCFC0A5FF8A9.taxon	description	(Figs 3 d, 3 h, 5, 63) Chinese name: 尖叶 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63BFFABFF7BFCFC0A5FF8A9.taxon	diagnosis	Diagnosis. This species closely resembles S. anmashanus (Yang, 1971), but can be distinguished from the later by the apically pointed penis guide in ventral view, and yellowish brown antennae and maxillary palpi (Figs 5 c, 5 h). In S. anmashanus, penis guide is apically rounded in ventral view, antennae and maxillary palpi are black (Figs 6 c, 6 h).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63BFFABFF7BFCFC0A5FF8A9.taxon	description	Description. TL: 1.46 – 2.09 mm, TW: 1.06 – 1.46 mm, TH: 0.65 – 0.78 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 5 a – c). Body uniformly black, except antennae, maxillary palpi and tarsi yellowish brown. Interocular distance slightly greater than eye width in frontal view. Dorsal pubescence forming distinct wavy patterns on elytra. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 5 d). Male genitalia: Penis slender, penis capsule with short outer arm and long inner one, apex of penis pointed (Figs 5 e – f). Penis guide triangular in lateral view with pointed apex; parameres about 1 / 2 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, tapering towards pointed apex (Figs 5 g – h). Female genitalia: Coxites triangular; styli strongly reduced and hardly visible; spermatheca worm-like, with basal ramus and nodulus (Figs 3 d, 3 h).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63BFFABFF7BFCFC0A5FF8A9.taxon	materials_examined	Type material. Holotype: China: Yunnan: ♂, Dajian Mountains, Daweishan National Nature Reserve, 2100 m, 20 – 21. IV. 2008, Wang XM leg (SCAU). Paratypes (SCAU): China: Yunnan: 1 ♀, Gaoligongshan National Nature Reserve, 19. IX. 2006, Wang XM leg; 2 ♂ 5 ♀, Xujiaba, Ailaoshan National Nature Reserve, 1 – 3. X. 2006, Wang XM leg; 2 ♂ 1 ♀, Taizhong Town, Jingdong County, 3. X. 2006, Wang XM leg; 4 ♂ 2 ♀, Heiwadi, Gongshan County, 2020 m, 19. VII. 2010, Wang XM leg; 2 ♀, Xujiaba, Ailaoshan National Nature Reserve, 2484 m, 15. VIII. 2013, Chen XS leg; 1 ♂, Xujiaba, Ailaoshan National Nature Reserve, 2484 m, 16. VIII. 2013, Chen XS leg; 5 ♂, Xujiaba, Ailaoshan National Nature Reserve, 2524 m, 17. VIII. 2013, Chen XS leg; 2 ♂, J 22 County Highway, Wuliangshan National Nature Reserve, 2400 m, 18. VIII. 2013, Chen XS leg. Fujian: 2 ♂, Zhangjia, Wuyishan National Nature Reserve, 950 m, 16. IX. 2012, Huo LZ leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63BFFABFF7BFCFC0A5FF8A9.taxon	distribution	Distribution. China (Fujian, Yunnan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63BFFABFF7BFCFC0A5FF8A9.taxon	etymology	Etymology. The specific epithet acutus derives from the Latin word “ acutus ” and refers to the apically pointed penis guide.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63BFFABFF7BFF75092CFD4D.taxon	description	(Fig. 65) Chinese name: 台南 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63BFFABFF7BFF75092CFD4D.taxon	diagnosis	Diagnosis. This species is similar to S. anmashanus (Yang, 1971), but can be distinguished from the later by parameres about 3 / 4 as long as penis guide. In S. anmashanus, parameres are about 1 / 2 as long as penis guide (Fig. 6 g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63BFFABFF7BFF75092CFD4D.taxon	materials_examined	Material examined. None.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63BFFABFF7BFF75092CFD4D.taxon	distribution	Distribution. China (Taiwan); Japan.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63AFFADFF7BFA65095EFEE4.taxon	description	(Figs 6, 63) Chinese name: ḆḤ 山 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63AFFADFF7BFA65095EFEE4.taxon	diagnosis	Diagnosis. This species closely resembles S. acutus Tong & Wang sp. nov., but can be distinguished from the later by the apically rounded penis guide in ventral view, and black antennae and maxillary palpi (Figs 6 c, 6 h). In S. acutus, penis guide is apically pointed in ventral view, antennae and maxillary palpi are yellowish brown (Figs 5 c, 5 h).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63AFFADFF7BFA65095EFEE4.taxon	description	Description. TL: 1.83 – 2.27 mm, TW: 1.30 – 1.67 mm, TH: 0.84 – 1.05 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 6 a – c). Body uniformly black. Interocular distance slightly greater than eye width in frontal view. Dorsal pubescence forming distinct wavy patterns on elytra. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 6 d). Male genitalia: Penis slender, penis capsule with short outer arm and long inner one, apex of penis pointed (Figs 6 e – f). Penis guide triangular in lateral view with pointed apex; parameres about 1 / 2 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, tapering towards blunt apex (Figs 6 g – h). Female genitalia: Coxites triangular; styli strongly reduced and hardly visible; spermatheca worm-like, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63AFFADFF7BFA65095EFEE4.taxon	materials_examined	Material examined. Holotype: China: Taiwan: ♂, Anma Mountains, Taichung City, 2200 m, 21. VII. 1971, Yang CT leg (NCHU). Other material (SCAU). China: Taiwan: 1 ♀, Ali Mountains, Chiayi County, 1800 m, 22. X. 2012, Wang XM leg; 1 ♂, Ali Mountains, Chiayi County, 2200 – 2500 m, 23. X. 2012, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63AFFADFF7BFA65095EFEE4.taxon	distribution	Distribution. China (Taiwan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63DFFACFF7BF9D30A5FFD28.taxon	description	(Figs 7, 64) Chinese name: Ľ 叶 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63DFFACFF7BF9D30A5FFD28.taxon	diagnosis	Diagnosis. This species is similar to S. sylvaticus (Lewis, 1896), but can be distinguished from the later by yellow pronotum, black scutellar shield, and parameres about 1 / 3 as long as penis guide (Figs 7 a – c, 7 g). In S. sylvaticus, pronotum is yellowish brown, scutellar shield is dark brown, and parameres are about 3 / 4 as long as penis guide (Figs 8 a – c, 8 g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63DFFACFF7BF9D30A5FFD28.taxon	description	Description. TL: 1.54 – 2.05 mm, TW: 1.21 – 1.54 mm, TH: 0.77 – 0.91 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 7 a – c). Head yellow with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow. Scutellar shield black. Elytra black with yellow apices. Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 7 d). Male genitalia: Penis long and slender, penis capsule with short outer arm and curved inner one, apex of penis pointed (Figs 7 e – f). Penis guide triangular in lateral view with pointed apex; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, tapering towards pointed apex (Figs 7 g – h). Female genitalia: Coxites distinctly elongate, triangular; styli strongly reduced and hardly visible; spermatheca distinctly curved, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63DFFACFF7BF9D30A5FFD28.taxon	materials_examined	Type material. Holotype: China: Guangxi: ♂, Gaozhai, Mao’ershan National Nature Reserve, 20. X. 2004, Wang XM leg (SCAU). Paratypes (SCAU): China: Guangxi: 5 ♂ 3 ♀, Shili Canyon, Mao’ershan National Nature Reserve, 19. X. 2004, Wang XM leg; 1 ♂, Pinglong Mountains, Shiwandashan National Nature Reserve, 8. XI. 2004, Wang XM leg. Hunan: 1 ♂, Mangshan National Nature Reserve, 3. X. 2004, Ren SX leg. Guangdong: 2 ♂, Nankunshan National Forest Park, 21. X. 2004, Wang XM leg; 1 ♂, Qinshuigu, Nanling National Nature Reserve, 21. IX. 2005, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63DFFACFF7BF9D30A5FFD28.taxon	distribution	Distribution. China (Hunan, Guangdong, Guangxi).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63DFFACFF7BF9D30A5FFD28.taxon	etymology	Etymology. The specific epithet cultratus derives from the Latin word “ cultratus ” and refers to the cultrated penis guide.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63FFFAFFF7BFF7508DBF818.taxon	description	(Figs 1 g, 3 c, 3 g, 8, 65) Chinese name: 枝 Ṅḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63FFFAFFF7BFF7508DBF818.taxon	diagnosis	Diagnosis. This species is similar to S. cultratus sp. nov., but can be distinguished from the later by yellowish brown pronotum, dark brown scutellar shield, and parameres about 3 / 4 as long as penis guide (Figs 8 a – c, 8 g). In S. cultratus, pronotum is yellow, scutellar shield is black, and parameres are about 1 / 3 as long as penis guide (Figs 7 a – c, 7 g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63FFFAFFF7BFF7508DBF818.taxon	description	Description. TL: 2.21 – 2.89 mm, TW: 1.59 – 2.03 mm, TH: 0.86 – 1.17 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 8 a – c). Head yellowish brown with interocular distance distinctly greater than eye width in frontal view. Pronotum yellowish brown. Scutellar shield dark brown. Elytra black with yellowish brown apices. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 8 d). Male genitalia: Penis long and slender, penis capsule with short outer arm and long inner one, apex of penis pointed (Figs 8 e – f). Penis guide long and slender in lateral view, approximately parallel sided for basal 4 / 5, then gradually narrowing to pointed apex; parameres about 3 / 4 as long as penis guide, densely covered with long setae at apex and inner sides; penis guide approximately parallel sided for basal 1 / 2 in ventral view, bifurcated apically (Figs 8 g – h). Female genitalia: Coxites distinctly elongate, triangular; styli reduced with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus (Figs 3 c, 3 g). Materail examined (SCAU). China: Anhui: 2 ♂, Guniujiang National Nature Reserve, 24. IX. 2010, Wang XM leg; 1 ♀, Jinzhai Tianma National Nature Reserve, 600 m, 3 – 5. X. 2010, Wang XM leg. Fujian: 1 ♀, Shipaichang, Longxishan National Nature Reserve, 500 m, 28 – 29. VIII. 2012, Huo LZ leg; 1 ♂, Guadun, Wuyishan National Nature Reserve, 1200 m, 14. IX. 2012, Huo LZ leg; 1 ♂ 1 ♀, Jiangxikou, Tongmu Village, Wuyishan National Nature Reserve, 1130 m, 19. IX. 2012, Huo LZ leg. Jiangxi: 1 ♂ 1 ♀, Huangyangjie, Jinggangshan National Nature Reserve, 16. VIII. 2004, Wang XM leg; 1 ♂, Jingzhu Mountains, Jinggangshan National Nature Reserve, 22. IX. 2004, Wang XM leg; 1 ♂, Xiangzhou, Jinggangshan National Nature Reserve, 23. IX. 2004, Wang XM leg. Hubei: 1 ♂, Lulin Town, Jingshan City, 169 m, 10. VII. 2007, Wang XM leg; 1 ♂ 1 ♀, Laojun Mountains, Shennongjia National Nature Reserve, 1230 m, 5. VIII. 2007, Wang XM leg. Hunan: 4 ♂, Mangshan National Nature Reserve, 3. X. 2004, Ren SX leg; 6 ♀, Shennonggu National Forest Park, 1100 m, 7. X. 2010, Wang XM leg; 1 ♂, Shennonggu National Forest Park, 1000 m, 8. X. 2010, Wang XM leg; 7 ♂ 4 ♀, Shennonggu National Forest Park, 650 – 800 m, 9. X. 2010, Wang XM leg. Guangdong: 1 ♂, Qinshuigu, Nanling National Nature Reserve, 30. IX. 2004, Wang XM leg; 3 ♂ 1 ♀, Xiaohuang Mountains, Nanling National Nature Reserve, 1. X. 2004, Wang XM leg; 2 ♂, Nanling National Nature Reserve, 2. X. 2004, Wang XM leg; 3 ♂ 2 ♀, Qinshuigu, Nanling National Nature Reserve, 21. IX. 2005, Wang XM leg. Guangxi: 3 ♂, Hillside, Mao’ershan National Nature Reserve, 17. X. 2004, Wang XM leg; 17 ♂ 11 ♀, Shili Canyon, Mao’ershan National Nature Reserve, 19. X. 2004, Wang XM leg; 1 ♀, Gaozhai, Mao’ershan National Nature Reserve, 20. X. 2004, Wang XM leg; 1 ♀, Pinglong Mountains, Shiwandashan National Nature Reserve, 6 – 8. XI. 2004, Wang XM leg; 1 ♂ 1 ♀, Hongqi Forest Farm, Shiwandashan National Nature Reserve, 26. VII. 2005, Wang XM leg; 1 ♂, Pinglong Mountains, Shiwandashan National Nature Reserve, 28. VII. 2005, Wang XM leg. Hainan: 1 ♂, Jianfengling National Nature Reserve, 11. XI. 1989, Ren SX leg. Guizhou: 1 ♀, Wudong Village – Maoping, Leigongshan National Nature Reserve, 1350 m, 6. X. 2008, Chen XS leg; 1 ♀, Wudong Village – Fangxiang, Leigongshan National Nature Reserve, 7. X. 2008, Chen XS leg; 6 ♂ 11 ♀, Xiangshuiyan, Leigongshan National Nature Reserve, 980 m, 9. X. 2008, Chen XS leg; 2 ♂ 2 ♀, Jiaomi Village, Taijiang County, 710 m, 14. X. 2008, Chen XS leg. Yunnan: 1 ♂ 1 ♀, Yingpan Town, Jingping County, 1487 m, 5. VIII. 2013, Chen XS leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63EFFB1FF7BFA86095EFEC0.taxon	description	(Figs 9, 63) Chinese name: Ḛ 氏 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63EFFB1FF7BFA86095EFEC0.taxon	diagnosis	Diagnosis. This species is similar to S. hosonaga Kitano, 2012, but can be distinguished from the later by its pronotum yellowish brown with a large black spot at base (Fig. 9 a). In S. hosonaga, pronotum is entirely black (Kitano 2012).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63EFFB1FF7BFA86095EFEC0.taxon	description	Description. TL: 2.7 – 3.0 mm, TW: 2.0 – 2.2 mm, TH: 0.86 – 1.17 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 9 a – d). Head yellowish brown with interocular distance distinctly greater than eye width in frontal view. Pronotum yellowish brown with a large black spot at base. Scutellar shield dark brown. Elytra black with yellow apices. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 9 e). Male genitalia: Penis long and slender, penis capsule with short outer arm and long inner one, apex of penis pointed (Fig. 9 f). Penis guide triangular in lateral view with pointed apex; parameres about 1 / 2 as long as penis guide, with sparsely setae at apex; penis guide widest at base in ventral view, tapering towards pointed apex (Fig. 9 g). Female genitalia: Coxites distinctly elongate, triangular; styli strongly reduced and hardly visible; spermatheca distinctly curved, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63EFFB1FF7BFA86095EFEC0.taxon	materials_examined	Material examined. Holotype: China: Taiwan: ♂, Chutung, Xinzhu County, 3. VI. 1972, Yang CT leg (NCHU). Paratypes: China: Taiwan: 2 ♂ 2 ♀, Neiwan, Xinzhu County, 3. VI. 1972, Yang CT leg (NCHU).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F63EFFB1FF7BFA86095EFEC0.taxon	distribution	Distribution. China (Taiwan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F621FFB1FF7BFB530A8CF83E.taxon	description	(Figs 10, 63) Chinese name: Dz 叶 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F621FFB1FF7BFB530A8CF83E.taxon	diagnosis	Diagnosis. This species is similar to S. nakanei (Araki, 1963), but can be distinguished from the later by parameres distinctly dilated in lateral view (Fig. 10 f). In S. nakanei, parameres are not dilated (Fig. 12 f).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F621FFB1FF7BFB530A8CF83E.taxon	description	Description. TL: 1.89 – 2.21 mm, TW: 1.50 – 1.63 mm, TH: 0.85 – 0.96 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 10 a – c). Head black with antennae and mouthparts yellow, interocular distance distinctly greater than eye width in frontal view. Pronotum and scutellar shield black. Elytra black with yellow apices. Ventral surfaces black. Legs brown. Abdominal postcoxal lines incomplete, slightly recurved toward the base of ventrite (Fig. 10 d). Male genitalia: Penis long and slender, penis capsule with short outer arm and long inner one, apex of penis pointed (Fig. 9 e). Penis guide triangular in lateral view with pointed apex; parameres distinctly dilated in lateral view, about 2 / 3 as long as penis guide, and with a tuft of long setae at apex; penis guide widest at base in ventral view, bifurcated apically (Figs 10 f – g). Female genitalia: Coxites distinctly elongate, triangular; styli strongly reduced and hardly visible; spermatheca distinctly curved, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F621FFB1FF7BFB530A8CF83E.taxon	materials_examined	Type material. Holotype: China: Anhui: ♂, Guniujiang National Nature Reserve, 24. IX. 2010, Wang XM leg (SCAU). Paratype (SCAU): China: Anhui: 1 ♀, Same data as holotype.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F621FFB1FF7BFB530A8CF83E.taxon	distribution	Distribution. China (Anhui).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F621FFB1FF7BFB530A8CF83E.taxon	etymology	Etymology. The specific epithet subulatus derives from the Latin word “ subulatus ” and refers to the subulate penis guide in ventral view.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F620FFB3FF7BFA9D0F85FE08.taxon	description	(Figs 11, 64) Chinese name: 历山 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F620FFB3FF7BFA9D0F85FE08.taxon	diagnosis	Diagnosis. This species is similar to S. fusinus sp. nov., but can be distinguished from the later by the black elytra with yellowish brown apices, smaller and shorter body size, and parameres about 1 / 2 as long as penis guide (Figs 11 a – b, 11 g). In S. fusinus, elytra are uniformly black, body size is larger and more elongate, and parameres are about 1 / 3 as long as penis guide (Figs 4 a – b, 4 f).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F620FFB3FF7BFA9D0F85FE08.taxon	description	Description. TL: 1.47 – 2.04 mm, TW: 1.02 – 1.18 mm, TH: 0.79 – 0.89 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 11 a – c). Head black with antennae and mouthparts yellowish brown, interocular distance distinctly greater than eye width in frontal view. Pronotum black with anterior margin narrowly yellowish brown. Scutellar shield black. Elytra black with brown apices. Ventral surfaces brown to black. Legs brown. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 11 d). Male genitalia: Penis long and slender, penis capsule with short outer arm and long inner one, apex of penis pointed (Figs 11 e – f). Penis guide triangular in lateral view with pointed apex; parameres about 1 / 2 as long as penis guide, with a tuft of long setae at apex; penis guide approximately widest at 1 / 2 length in ventral view, bifurcated apically (Figs 11 g – h). Female genitalia: Coxites distinctly elongate, triangular; styli strongly reduced and hardly visible; spermatheca distinctly curved, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F620FFB3FF7BFA9D0F85FE08.taxon	materials_examined	Type material. Holotype: China: Shanxi: ♂, Lishan National Nature Reserve, 800 – 1400 m, 3. VIII. 2011, Wang XM leg (SCAU). Paratypes (SCAU): China: Shanxi: 5 ♂ 4 ♀, Heilongtan, Dahe Village, Lishan National Nature Reserve, 995 m, 17. VII. 2013, Huo LZ leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F620FFB3FF7BFA9D0F85FE08.taxon	distribution	Distribution. China (Shanxi).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F620FFB3FF7BFA9D0F85FE08.taxon	etymology	Etymology. The specific epithet refers to its type locality, Lishan National Nature Reserve.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F623FFB3FF7BF990095EF86E.taxon	description	Chinese name: 仁爱 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F623FFB3FF7BF990095EF86E.taxon	diagnosis	Diagnosis. This species is similar to S. nakanei (Araki, 1963), but can be distinguished from the later by parameres about 1 / 2 as long as penis guide (Kitano 2012). In S. anmashanus, parameres are about 3 / 4 as long as penis guide (Fig. 6 g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F623FFB3FF7BF990095EF86E.taxon	diagnosis	Material examined. None. Distribution. China (Taiwan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F622FFB5FF7BFF750F99FEE4.taxon	description	(Figs 1 b, 12, 64) Chinese name: AE + ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F622FFB5FF7BFF750F99FEE4.taxon	diagnosis	Diagnosis. This species is similar to S. subulatus sp. nov., but can be distinguished from the later by parameres not dilated in lateral view (Fig. 12 f). In S. subulatus, parameres are distinctly dilated (Fig. 10 f).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F622FFB5FF7BFF750F99FEE4.taxon	description	Description. TL: 1.83 – 2.57 mm, TW: 1.38 – 1.89 mm, TH: 0.71 – 1.03 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 12 a – c). Head black with antennae and mouthparts brown, interocular distance distinctly greater than eye width in frontal view. Pronotum and scutellar shield black. Elytra black with brown apices. Ventral surfaces brown to black. Legs brown. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 12 d). Male genitalia: Penis long and slender, penis capsule with short outer arm and long inner one, apex of penis pointed (Fig. 12 e). Penis guide triangular in lateral view with pointed apex; parameres about 3 / 4 as long as penis guide, densely covered with long setae at apex; penis guide widest at base in ventral view, tapering towards pointed apex (Fig. 12 f – g). Female genitalia: Coxites distinctly elongate, triangular; styli strongly reduced and hardly visible; spermatheca distinctly curved, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F622FFB5FF7BFF750F99FEE4.taxon	materials_examined	Material examined. China: Hubei: 5 ♂ 5 ♀, Lulin Town, Jingshan County, 169 m, 10. VII. 2007, Wang XM leg; 2 ♂, Xingshan County, 16. VII. 2007, Wang XM leg. Chongqing: 1 ♂ 1 ♀, Wuxi County, 2. VII. 1989, Ren SX leg. Guizhou: 4 ♂ 4 ♀, Huaxi District, Guiyang City, 17 – 18. VIII. 1987, Pang XF leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F622FFB5FF7BFF750F99FEE4.taxon	distribution	Distribution. China (Shandong, Hubei, Chongqing, Guizhou, Shaanxi); Japan; South Korea.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F625FFB5FF7BFE55095EFCB2.taxon	description	(Fig. 64)	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F625FFB5FF7BFE55095EFCB2.taxon	diagnosis	Diagnosis. This species is similar to S. ohtai (Yang & Wu, 1972), but can be distinguished from the later by its pronotum entirely black (Kitano 2012). In S. ohtai, pronotum is yellowish brown with a large black spot at base (Fig. 9 a).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F625FFB5FF7BFE55095EFCB2.taxon	materials_examined	Material examined. None.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F625FFB5FF7BFE55095EFCB2.taxon	distribution	Distribution. China (Taiwan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F625FFB4FF7BFC6708D8F9C3.taxon	discussion	The Sasajiscymnus hareja species group is characterized by the stout penis with large penis capsule, apex of penis with membranous appendage, penis guide robust with prominent keel at base, interocular distance distinctly greater than eye width in frontal view, coxites elongate and triangular, styli well developed with terminal setae, and spermathecal with elongate nodulus. 34 species of hareja group are recorded from China, including 12 new species and 1 newly recorded species.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F625FFB4FF7BFC6708D8F9C3.taxon	discussion	Remarks. Sasajiscymnus hareja (Weise, 1879) and Sasajiscymnus changi (Yang, 1971) were not included in the above key since no specimens of these species were available for us.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F624FFB7FF7BF9760908FDBC.taxon	description	(Figs 13, 72) Chinese name: 瘤叶 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F624FFB7FF7BF9760908FDBC.taxon	description	Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis with membranous appendage (Fig. 13 e). Penis guide widest at base in lateral view, apically pointed; parameres inconspicuous, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 3 / 4 in ventral view, then sharply narrowing to pointed apex (Figs 13 f – g). Female genitalia: Unknow.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F624FFB7FF7BF9760908FDBC.taxon	materials_examined	Material examined. Holotype: China: Chongqing: ♂, Wang’erbao, WanZhou District, 28. IX. 1994, Li FS leg (CAU).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F624FFB7FF7BF9760908FDBC.taxon	distribution	Distribution. China (Chongqing).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F627FFB6FF7BF966085CFCDD.taxon	description	Chinese name: Ḃ 兴 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F627FFB6FF7BF966085CFCDD.taxon	diagnosis	Diagnosis. This species is similar to S. bivalvis (Yu, 1997), but can be distinguished from the later by the penis guide widest at base in ventral view, and the penis with several small protuberances at middle (Fig. 14 e). In S. bivalvis, penis guide is approximately parallel sided for basal 1 / 2 in ventral view, penis without protuberances (Fig. 39 e).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F627FFB6FF7BF966085CFCDD.taxon	description	Description. TL: 2.05 – 2.51 mm, TW: 1.50 – 1.89 mm, TH: 0.99 – 1.12 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 14 a – c). Head yellowish brown with interocular distance distinctly greater than eye width in frontal view. Pronotum and scutellar shield yellowish brown. Elytra black with yellowish brown apices and humeral angles. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 14 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, central part of penis with several small protuberances, apex of penis slightly expanded with membranous appendage (Figs 14 e – f). Penis guide with prominent keel at base in lateral view, then gradually narrowing to pointed apex; parameres about 1 / 2 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, apex with a small protuberance (Figs 14 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F627FFB6FF7BF966085CFCDD.taxon	materials_examined	Material examined. Holotype: China: Guangdong: ♂, Shixing County, 8. VII. 1989, Pang XF leg (SCAU); Paratype: China: Guangdong: 1 ♀, Same data as holotype (SCAU). Other material (SCAU). China: Guangdong: 7 ♂ 10 ♀, Huangdong, Shimentai National Nature Reserve, 30. X. 2004, Wang XM leg; 23 ♂ 29 ♀, Huangdong, Shimentai National Nature Reserve, 31. X. 2004, Wang XM leg. Guangxi: 1 ♂, Hillside, Mao’ershan National Nature Reserve, 17. X. 2004, Wang XM leg; 4 ♂, Shili Canyon, Mao’ershan National Nature Reserve, 19. X. 2004, Wang XM leg; 1 ♂, GaoZhai, Mao’ershan National Nature Reserve, 20. X. 2004, Wang XM leg. Taiwan: 2 ♂, Aowanda, Jenai Town, Nantou County, 1280 m, 21. X. 2012, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F627FFB6FF7BF966085CFCDD.taxon	distribution	Distribution. China (Guangdong, Guangxi, Taiwan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F629FFB8FF7BFF75094CFE51.taxon	description	(Figs 15, 72) Chinese name: 条 Ṅḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F629FFB8FF7BFF75094CFE51.taxon	diagnosis	Diagnosis. This species can be distinguished from other species of Sasajiscymnus by its penis strongly constricted at distal 1 / 3 (Fig. 15 e).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F629FFB8FF7BFF75094CFE51.taxon	description	Description. TL: 1.45 – 1.99 mm, TW: 1.01 – 1.44 mm, TH: 0.59 – 0.73 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 15 a – c). Head yellow with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow with a large black spot. Scutellar shield black. Elytra black with yellowish brown apices, and each elytron with a longitudinal reddish brown stripe. Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 15 d). Male genitalia: Penis stout and strongly constricted at distal 1 / 3, penis capsule with stout outer arm and long inner one, apex of penis distinctly constricted with membranous appendage (Figs 15 e – f). Penis guide with prominent keel at base in lateral view, then gradually narrowing to pointed apex; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 3 in ventral view, apically rounded (Figs 15 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F629FFB8FF7BFF75094CFE51.taxon	materials_examined	Type material. Holotype: China: Xizang: ♂, Beibeng Town, Muotuo County, 800 m, 9. X. 2011, Huo LZ leg (SCAU). Paratypes (SCAU): China: Yunnan: 2 ♂ 1 ♀, Qinglandan, Maku Village, Gongshan County, 4 – 7. VIII. 2010, Wang XM leg. Xizang: 1 ♂ 1 ♀, Yarang Village, Muotuo County, 768 – 1030 m, 24. X. 2007, Wang XM leg; 2 ♂ 2 ♀, Beibeng Town, Muotuo County, 600 – 768 m, 25. X. 2007, Wang XM leg; 3 ♂, Zhamo Highroad 140 K, Bomi County, 900 m, 27. X. 2007, Wang XM leg; 5 ♂ 4 ♀, Yihaoqiao, Hanmi, Muotuo County, 14. X. 2009, Wang XM leg; 1 ♂ 2 ♀, Hanmi, Muotuo County, 800 – 2100 m, 5 – 8. X. 2011, Huo LZ leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F629FFB8FF7BFF75094CFE51.taxon	distribution	Distribution. China (Yunnan, Xizang).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F629FFB8FF7BFF75094CFE51.taxon	etymology	Etymology. The specific epithet striatus derives from the Latin word “ striatus ” and refers to the longitudinal stripe on the disc of each elytron.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F628FFBBFF7BFDC00A44FD70.taxon	description	Chinese name: ƮDz 山 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F628FFBBFF7BFDC00A44FD70.taxon	diagnosis	Diagnosis. This species is similar to S. ocelliferus (Canepari, 1997) in structures of the penis, but can be distinguished from the later by the entire yellow pronotum and parameres about 1 / 3 as long as penis guide (Figs 16 a – c, 16 g). In S. ocelliferus, pronotum is black with lateral and anterior margin yellow, parameres are about 3 / 5 as long as penis guide (Figs 17 a – c, 17 g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F628FFBBFF7BFDC00A44FD70.taxon	description	Description. TL: 1.58 –. 92 mm, TW: 1.12 – 1.36 mm, TH: 0.78 – 0.94 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 16 a – c). Head yellow with interocular distance distinctly greater than eye width in frontal view. Pronotum and scutellar yellow. Elytra black with yellow apices, sometimes with two yellow spots on medially. Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 16 d). Male genitalia: Penis stout with a protuberance on inner side at middle, penis capsule with stout outer arm and long inner one, apex of penis distinctly expanded with membranous appendage (Figs 16 e – f). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide stout in ventral view, widest at basal 1 / 3, then gradually narrowing to apex (Figs 16 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F628FFBBFF7BFDC00A44FD70.taxon	materials_examined	Type material. Holotype: China: Yunnan: ♂, J 22 County Highway, Wuliangshan National Nature Reserve, 2400 m, 18. VIII. 2013, Chen XS leg (SCAU). Paratypes (SCAU): China: Yunnan: 2 ♂ 2 ♀, Same data as holotype. Xizang: 1 ♂, Xiachayu Town, Chaiyu County, 1800 m, 14. X. 2007, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F628FFBBFF7BFDC00A44FD70.taxon	distribution	Distribution. China (Yunnan, Xizang).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F628FFBBFF7BFDC00A44FD70.taxon	etymology	Etymology. This new species is named after Wuliangshan National Nature Reserve where the holotype was collected.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62BFFBBFF7BFCA109A6F828.taxon	description	(Figs 17, 68) Chinese name: ẸṄḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62BFFBBFF7BFCA109A6F828.taxon	diagnosis	Diagnosis. This species is similar to S. wuliangshan sp. nov. in structures of the penis, but can be distinguished from the later by the pronotum black with lateral and anterior margin yellow, and parameres about 3 / 5 as long as penis guide (Figs 17 a – c, 17 g). In S. wuliangshan, pronotum is entirely black and parameres are about 1 / 3 as long as penis guide (Figs 16 a – c, 16 g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62BFFBBFF7BFCA109A6F828.taxon	description	Description. TL: 1.62 – 2.32 mm, TW: 1.17 – 1.69 mm, TH: 0.63 – 0.87 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 17 a – c). Head yellowish brown to black, with interocular distance distinctly greater than eye width in frontal view. Pronotum black with lateral and anterior margin yellow. Scutellar shield brown. Elytra black with yellow apices, nd each elytron with an eye-spotted marking on medially extending to the suture. Ventral surfaces yellowish brown to black. Legs yellow. Abdominal postcoxal lines incomplete, slightly recurved toward the base of ventrite (Fig. 17 d). Male genitalia: Penis stout with a large protuberance on inner side near distal 1 / 3, penis capsule with stout outer arm and long inner one, apex of penis distinctly expanded with membranous appendage (Figs 17 e – f). Penis guide approximately parallel sided for basal 1 / 3 in lateral view, then gradually narrowing to pointed apex; parameres about 3 / 5 as long as penis guide, with a tuft of long setae at apex; penis guide stout in ventral view, widest at basal 1 / 3, with a small protuberance at apex (Figs 17 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62BFFBBFF7BFCA109A6F828.taxon	materials_examined	Material examined. China: Xizang: 1 ♀, Zhangmu Town, Nielamu County, 2100 m, 25 – 26. IX. 2009, Ren SX leg; 3 ♀, Zhangmu Town, Nielamu County, 2500 m, 27. IX. 2009, Ren SX leg; 9 ♂ 6 ♀, Linzhi Town, Bayi District, Linzhi City, 2850 m, 20. X. 2011, Huo LZ leg; 1 ♂, Lebu Town, Cuona County, 2400 m, 24. X. 2011, Huo LZ leg; 1 ♀, Zhangmu Town, Nielamu County, 2200 m, 31. X – 2. XI. 2011, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62BFFBBFF7BFCA109A6F828.taxon	distribution	Distribution. China (Xizang); Nepal.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62BFFBBFF7BFCA109A6F828.taxon	discussion	Remarks. This species is recorded from China for the first time. The specimens were in perfect agree well with the descriptions and illustrations of Canepari (1997).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62AFFBDFF7BFA8A095EFD98.taxon	description	(Figs 18, 68) Chinese name: 圆 Ṅḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62AFFBDFF7BFA8A095EFD98.taxon	diagnosis	Diagnosis. This species is similar to S. ocellatus (Yu, 2000), but can be distinguished from the later by the shorter body, the penis guide approximately parallel sided for basal 1 / 3 in ventral view, and the eye-spotted marks on the elytra connecting with the lateral and apical margins (Figs 18 a – b, 18 h). In S. ocellatus, body is more elongate, penis guide is widest at base in ventral view, and the eye-spotted marks on the elytra not touching the lateral and apical margins (Figs 19 a – b, 19 g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62AFFBDFF7BFA8A095EFD98.taxon	description	Description. TL: 2.25 – 2.51 mm, TW: 1.53 – 1.82 mm, TH: 0.95 – 1.07 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 18 a – c). Head yellowish brown with interocular distance distinctly greater than eye width in frontal view. Pronotum yellowish brown with a black spot. Scutellar shield black. Elytra black with two eye-spotted markings connecting with the lateral and apical margins. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 18 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis constricted with membranous appendage (Figs 18 e – f). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 1 / 2 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 3 in ventral view, then gradually narrowing to apex (Figs 18 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62AFFBDFF7BFA8A095EFD98.taxon	materials_examined	Material examined. Holotype: China: Taiwan: ♂, Anma Mountains, Taichung City, 2200 m, 21. VII. 1971, Yang CT leg (NCHU). Paratypes: China: Taiwan: 2 ♂ 2 ♀, Same data as holotype (NCHU). Other material (SCAU). China: Taiwan: 1 ♀, Qingjing, Jenai Town, Nantou County, 720 – 2329 m, 20. X. 2012, Wang XM leg; 1 ♀, Dabang Village, Alishan Mountains, Chiayi County, 960 – 1600 m, 24. X. 2012, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62AFFBDFF7BFA8A095EFD98.taxon	distribution	Distribution. China (Taiwan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62DFFBCFF7BF9F709A0FC94.taxon	description	(Figs 1 e, 19, 66) Chinese name: ḎṄḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62DFFBCFF7BF9F709A0FC94.taxon	diagnosis	Diagnosis. This species is similar to S. orbiculatus (Yang, 1971), but can be distinguished from the later by the more elongate body, the penis guide widest at base in ventral view, and the eye-spotted markings on the elytra not connecting with the lateral and apical margins (Figs 19 a – b, 19 g). In S. orbiculatus, body is shorter, penis guide is approximately parallel sided for basal 1 / 3 in ventral view, and the eye-spotted markings on the elytra connecting with lateral and apical margins (Figs 18 a – b, 18 h). Body oval, moderately convex, surfaces covered with short pubescence (Figs 19 a – c). Head black to brown, with interocular distance distinctly greater than eye width in frontal view. Pronotum yellowish brown with a black mark. Scutellar shield black. Elytron black with two yellow marks: the anterior one O-shaped, situated in the middle part, not touching elytral margin or suture; the other one located at the apex of elytron. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, slightly recurved toward the base of ventrite (Fig. 19 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis constricted with membranous appendage (Fig. 19 e). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 1 / 2 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, then gradually narrowing to apex (Figs 19 f – g). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62DFFBCFF7BF9F709A0FC94.taxon	materials_examined	Material examined. Holotype: China: Sichuan: ♂, Qiaoqi Town, Baoxing County, [102 ° 48 ' E 30 ° 18 ' N], V. 1996, Guo leg (BAAF). Paratypes: China: Sichuan: 1 ♂, Nibagou, Baoxing County, 14. X. 1996, Collector unknown (BAAF). Shaanxi: 1 ♀, Huoditang, Ningshan County, [108 ° 12 ' E 34 ° 0 ' N], 8. IV. 1998, Yao DF leg (BAAF). Other material (SCAU). China: Hubei: 1 ♂ 1 ♀, Yinyuhe, Shennongjia National Nature Reserve, 1600 – 1800 m, 31. VII. 2007, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62DFFBCFF7BF9F709A0FC94.taxon	distribution	Distribution. China (Hubei, Sichuan, Shaanxi).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62FFFBFFF7BFF750953FC14.taxon	description	Chinese name: 矛ū ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62FFFBFFF7BFF750953FC14.taxon	diagnosis	Diagnosis. This species can be easily distinguished from other Sasajiscymnus by the body uniformly yellow without any contrasting markings (Figs 20 a – c).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62FFFBFFF7BFF750953FC14.taxon	description	Description. TL: 2.04 – 2.31 mm, TW: 1.60 – 1.79 mm, TH: 1.10 – 1.23 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 20 a – c). Body uniformly yellow. Interocular distance distinctly greater than eye width in frontal view. Abdominal postcoxal lines incomplete, slightly recurved toward the base of ventrite (Fig. 20 e). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis with membranous appendage (Figs 20 f – g). Penis guide with prominent keel at base in lateral view, then gradually narrowing to pointed apex; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, tapering towards apex (Figs 20 h – i). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62FFFBFFF7BFF750953FC14.taxon	materials_examined	Material examined. Holotype: China: Fujian: ♂, Shaxian District, V. 1979, Huang BK leg (SCAU). Paratypes: China: Fujian: 3 ♀, Same data as holotype (SCAU).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62FFFBFFF7BFF750953FC14.taxon	distribution	Distribution. China (Fujian).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62EFF81FF7BFF750A01FE2D.taxon	description	(Figs 21, 66) Chinese name: ḢÑḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62EFF81FF7BFF750A01FE2D.taxon	diagnosis	Diagnosis. This species is similar to S. seboshii (Ohta, 1929) in the characters of male genitalia, but can be distinguished from the later by the elytra entirely yellow, and the parameres about 1 / 3 as long as penis guide (Figs 21 a – b, 21 g). In S. seboshii, elytra are black with yellow apices and humeral angles, and a large yellowish brown spot presenting in the middle part, parameres are about 2 / 5 as long as penis guide (Figs 34 a – b, 34 g)	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62EFF81FF7BFF750A01FE2D.taxon	description	Description. TL: 1.55 – 2.08 mm, TW: 1.23 – 1.50 mm, TH: 0.85 – 1.01 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 21 a – c). Body uniformly yellow, except ventral surfaces yellow to black. Interocular distance distinctly greater than eye width in frontal view. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 21 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis constricted with membranous appendage (Figs 21 e – f). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, apex with a small protuberance (Figs 21 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62EFF81FF7BFF750A01FE2D.taxon	materials_examined	Type material. Holotype: China: Xizang: ♂, Hanmi, Muotuo County, 800 – 2100 m, 5 – 8. X. 2011, Huo LZ leg (SCAU). Paratypes (SCAU): China: Xizang: 2 ♂ 1 ♀, Same data as holotype. 1 ♂ 1 ♀, Yarang Village, Muotuo County, 768 – 1030 m, 24. X. 2007, Wang XM leg; 1 ♀, Houshan, Muotuo County, 1200 m, 28. X. 2007, Wang XM leg; 1 ♂, Hanmi, Muotuo County, 2100 m, 13. X. 2009, Wang XM leg; 1 ♀, Yihaoqiao, Hanmi, Muotuo County, 14. X. 2009, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62EFF81FF7BFF750A01FE2D.taxon	distribution	Distribution. China (Xizang).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F62EFF81FF7BFF750A01FE2D.taxon	etymology	Etymology. The specific epithet aureus derives from the Latin word “ aureus ” and refers to the aureate pronotum and elytra.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F611FF80FF7BFE1C095EFE74.taxon	description	(Figs 22, 66) Chinese name: 棕OE ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F611FF80FF7BFE1C095EFE74.taxon	diagnosis	Diagnosis. This species can be easily recognized by its uniformly brown elytra (Fig. 22 a).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F611FF80FF7BFE1C095EFE74.taxon	description	Description. TL: 2.05 – 2.51 mm, TW: 1.50 – 1.89 mm, TH: 0.99 – 1.12 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 22 a – d). Head yellow with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow. Scutellar shield and elytra entirely brown. Ventral surfaces yellow to black. Legs yellow to yellowish brown. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 22 e). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis slightly expanded with membranous appendage (Figs 22 f – g). Penis guide with prominent keel at base in lateral view, then gradually narrowing to pointed apex; parameres about 1 / 4 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 2 in ventral view, then gradually narrowing to apex (Fig. 22 h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F611FF80FF7BFE1C095EFE74.taxon	materials_examined	Material examined. Holotype: China: Taiwan: ♂, Anma Mountains, Taichung City, 2200 m, 21. VII. 1971, Yang CT leg (NCHU). Paratypes: China: Taiwan: 5 ♂ 4 ♀, Same data as holotype (NCHU). Other material (SCAU). China: Taiwan: 3 ♀, Qingjing, Jenai Town, Nantou County, 720 – 2329 m, 20. X. 2012, Wang XM leg; 2 ♀, Aowanda, Jenai Town, Nantou County, 1280 m, 21. X. 2012, Wang XM leg; 1 ♂, Ali Mountains, Chiayi County, 1800 m, 22. X. 2012, Wang XM leg; 1 ♂ 2 ♀, Ali Mountains, Chiayi County, 2200 – 2500 m, 23. X. 2012, Wang XM leg; 2 ♂ 1 ♀, Dabang Village, Alishan Mountains, Chiayi County, 960 – 1600 m, 24. X. 2012, Wang XM leg; 1 ♀, Maolin District, Kaohsiung City, 365 – 515 m, 26. X. 2012, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F611FF80FF7BFE1C095EFE74.taxon	distribution	Distribution. China (Taiwan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F610FF83FF7BFDA5095EFC48.taxon	description	(Figs 1 k, 23) Chinese name: 弧 Ṅḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F610FF83FF7BFDA5095EFC48.taxon	diagnosis	Diagnosis. This species is similar to S. quinquepunctatus (Weise, 1923) in the characters of male genitalia, but can be distinguished from the later by parameres slightly shorter than penis guide (Fig. 23 g). In S. quinquepunctatus, parameres are about 3 / 4 as long as penis guide (Fig. 2 n).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F610FF83FF7BFDA5095EFC48.taxon	description	Description. TL: 1.76 – 2.44 mm, TW: 1.34 – 1.86 mm, TH: 0.79 – 1.08 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 23 a – c). Head yellow with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow. Scutellar shield black. Elytra yellowish brown with parenthesis-shaped black markings. Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, slightly recurved toward the base of ventrite (Fig. 23 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis distinctly expanded with membranous appendage (Figs 23 e – f). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres conspicuously inflated, slightly shorter than penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 3 in ventral view, then gradually narrowing to apex (Figs 23 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F610FF83FF7BFDA5095EFC48.taxon	materials_examined	Material examined. China: Taiwan: 1 ♂ 1 ♀, Aowanda, Jenai Town, Nantou County, 1280 m, 21. X. 2012, Wang XM leg; 1 ♀, Ali Mountains, Chiayi County, 1800 m, 22. X. 2012, Wang XM leg; 1 ♂ 2 ♀, Dabang Village, Alishan Mountains, Chiayi County, 960 – 1600 m, 24. X. 2012, Wang XM leg; 1 ♂, Tengjhih, Kaohsiung City, 370 – 1450 m, 25. X. 2012, Wang XM leg; 2 ♀, Maolin District, Kaohsiung City, 365 – 515 m, 26. X. 2012, Wang XM leg; 1 ♂ 4 ♀, Jhihben, Taitung City, 230 m, 28. X. 2012, Wang XM leg; 2 ♂, Nanheng, Chihshang Town, Taitung County, 679 m, 29. X. 2012, Wang XM leg; 1 ♂ 2 ♀, Taroko National Park, Hualien County, 200 – 912 m, 30. X. 2012, Wang XM leg; 1 ♀, Beiheng Highway, Yilan County, 300 – 1500 m, 1. XI. 2012, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F610FF83FF7BFDA5095EFC48.taxon	distribution	Distribution. China (Taiwan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F613FF83FF7BFBF20EDDF870.taxon	description	(Figs 24, 69) Chinese name: Ḵ 叶 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F613FF83FF7BFBF20EDDF870.taxon	diagnosis	Diagnosis. This species is similar to S. quinquepunctatus (Weise, 1923) in the characters of male genitalia, but can be distinguished from the later by the more elongate penis guide and parameres about 1 / 2 as long as penis guide (Figs 24 g – h). In S. quinquepunctatus, penis guide is shorter and parameres are about 3 / 4 as long as penis guide (Figs 2 n – o).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F613FF83FF7BFBF20EDDF870.taxon	description	Description. TL: 2.10 – 2.59 mm, TW: 1.46 – 1.80 mm, TH: 1.03 – 1.28 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 24 a – c). Head yellow with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow. Scutellar shield black. Elytra black with yellow apices. Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 24 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis distinctly expanded with membranous appendage (Figs 24 e – f). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres conspicuously inflated, about 1 / 2 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 2 in ventral view, then gradually narrowing to blunt apex (Figs 24 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F613FF83FF7BFBF20EDDF870.taxon	materials_examined	Type material. Holotype: China: Xizang: ♂, Cibagou National Nature Reserve, 1615 m, 26. IX. 2011, Huo LZ leg (SCAU). Paratypes (SCAU): China: Xizang: 1 ♂, Same data as holotype; 1 ♂ 1 ♀, Cibagou National Nature Reserve, 2000 m, 13. X. 2007, Wang XM leg. Henan: 1 ♂, Baotianman National Nature Reserve, 6. VII. 2009, Wang XM leg. Yunnan: 1 ♂ 3 ♀, Caiyanghe National Forest Park, 4. V. 2009, Wang XM leg; 2 ♂ 3 ♀, Maku Village, Gongshan County, 1600 m, 1. VIII. 2010, Wang XM leg; 2 ♂, Qinglandan, Maku Village, Gongshan County, 4 – 7. VIII. 2010, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F613FF83FF7BFBF20EDDF870.taxon	materials_examined	Distribution. China (Henan, Yunnan, Xizang). Etymology. The specific epithet longus derives from the Latin word “ longus ” and refers to the elongate penis guide.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F612FF87FF7BFA650E3CFBFC.taxon	description	(Figs 2, 25, 73) Chinese name: 五 Ṅḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F612FF87FF7BFA650E3CFBFC.taxon	diagnosis	Diagnosis. This species is similar to S. parenthesis (Weise, 1923) in structures of the male genitalia, but can be distinguished from the later by parameres about 3 / 4 as long as penis guide (Fig. 2 n). In S. parenthesis, parameres are slightly shorter than penis guide (Fig. 23 g). Description. TL: 1.79 – 2.74 mm, TW: 1.28 – 2.07 mm, TH: 0.75 – 1.27 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 25 a – c). Head yellowish brown with interocular distance distinctly greater than eye width in frontal view. Pronotum and scutellar shield yellow. Elytra yellowish brown with five black spots. Sometimes the basal and apical spots are enlarged, or even the entire elytra are black, except for the yellowish brown apical margins. Ventral surfaces yellowish brown to black. Legs yellow. Abdominal postcoxal lines incomplete, slightly recurved toward the base of ventrite (Fig. 2 k). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis distinctly expanded with membranous appendage (Figs 2 m – l). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres conspicuously inflated, about 2 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 3 in ventral view, then gradually narrowing to apex (Figs 2 n – o). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F612FF87FF7BFA650E3CFBFC.taxon	materials_examined	Material examined. China: Zhejiang: 2 ♂ 1 ♀, Qingliangfeng National Nature Reserve, 465 m, 16. IX. 2010, Wang XM leg; 8 ♂ 8 ♀, Zhengjiang Tianmushan National Nature Reserve, 450 – 1000 m, 18. IX. 2010, Wang XM leg. Anhui: 4 ♂ 6 ♀, Tangkou Town, Huangshan District, Huangshan City, 460 m, 12 – 13. IX. 2010, Wang XM leg; 2 ♀, Huangshan Mountain Scenic Spot, 14 – 15. IX. 2010, Wang XM leg; 2 ♂, Hengdu Town, Shitai County, 230 m, 20. IX. 2010, Wang XM leg; 6 ♂ 9 ♀, Guniujiang National Nature Reserve, 24. IX. 2010, Wang XM leg. Fujian: 1 ♂ 1 ♀, Guizhu, Meihuashan National Nature Reserve, 1200 m, 20. VIII. 2012, Huo LZ leg; 2 ♂ 2 ♀, Yuncai, Liangyeshan National Nature Reserve, 735 m, 23. VIII. 2012, Huo LZ leg; 2 ♀, Sanbailiao, Tianboyan National Nature Reserve, 690 m, 26. VIII. 2012, Huo LZ leg; 2 ♂ 1 ♀, Shipaichang, Longxishan National Nature Reserve, 500 m, 28 – 29. VIII. 2012, Huo LZ leg; 2 ♂ 1 ♀, Zonghechang, Longxishan National Nature Reserve, 750 m, 30. VIII. 2012, Huo LZ leg; 2 ♀, Fanjia, Junzifeng National Nature Reserve, 460 m, 1. IX. 2012, Huo LZ leg; 2 ♂ 1 ♀, Masu, Wuyishan National Nature Reserve, 1291 m, 15. IX. 2012, Huo LZ leg; 1 ♂ 1 ♀, Tongmu Village, Wuyishan National Nature Reserve, 750 m, 17. IX. 2012, Huo LZ leg; 1 ♀, Jiangxikou, Tongmu Village, Wuyishan National Nature Reserve, 1130 m, 19. IX. 2012, Huo LZ leg; 2 ♂, Guaxi Village, Fu’an City, 260 m, 21. IX. 2012, Huo LZ leg. Jiangxi: 1 ♂ 1 ♀, Xiangzhou, Jinggangshan National Nature Reserve, 14. VIII. 2004, Wang XM leg; 10 ♂ 1 ♀, Fengshan National Forest Park, 17. VIII. 2004, Wang XM leg; 2 ♂, Tongmuling, Jinggangshan National Nature Reserve, 19. IX. 2004, Wang XM leg; 1 ♂, Xiangzhou, Jinggangshan National Nature Reserve, 23. IX. 2004, Wang XM leg; 3 ♂ 2 ♀, Longtan, Jinggangshan National Nature Reserve, 24. IX. 2004, Wang XM leg; 1 ♀, Hexilong, Jinggangshan National Nature Reserve, 26. IX. 2004, Wang XM leg. Henan: 1 ♂, Laojieling, Xixia County, 1430 m, 9. VII. 2009, Wang XM leg; 2 ♀, Baiyunshan National Nature Reserve, 1375 m, 14. VII. 2009, Wang XM leg. Hubei: 1 ♂ 6 ♀, Banqiao, Shennongjia National Nature Reserve, 1170 m, 21 – 24. VII. 2007, Wang XM leg; 1 ♀, Pingzhan, Shennongjia National Nature Reserve, 1600 m, 27. VII. 2007, Wang XM leg; 1 ♀, Yinyuhe, Shennongjia National Nature Reserve, 1600 – 1800 m, 31. VII. 2007, Wang XM leg; 1 ♀, Laojun Mountains, Shennongjia National Nature Reserve, 1230 m, 5. VIII. 2007, Wang XM leg; 1 ♂, Wudang Mountains, Shiyan City, 700 – 930 m, 8. VIII. 2007, Wang XM leg. Hunan: 3 ♂ 7 ♀, Protection Station, Mangshan National Nature Reserve, 29. IX. 2004, Wang XM leg; 19 ♂ 5 ♀, Mangshan National Nature Reserve, 2 – 3. X. 2004, Ren SX leg; 4 ♂ 5 ♀, Shennonggu National Forest Park, 1100 m, 7. X. 2010, Wang XM leg; 9 ♂ 7 ♀, Shennonggu National Forest Park, 1000 m, 8. X. 2010, Wang XM leg; 18 ♂ 10 ♀, Shennonggu National Forest Park, 650 – 800 m, 9. X. 2010, Wang XM leg. Guangdong: 1 ♂, Chebaling National Nature Reserve, 3. XI. 1989, Yu GY leg; 1 ♂, Chebaling National Nature Reserve, 23. IV. 1991, Yu GY leg; 7 ♂ 1 ♀, Qinshuigu, Nanling National Nature Reserve, 30. IX. 2004, Wang XM leg; 16 ♂ 3 ♀, Xidianzhan, Nanling National Nature Reserve, 1. X. 2004, Wang XM leg; 1 ♂ 1 ♀, Xiaohuang Mountains, Nanling National Nature Reserve, 1. X. 2004, Wang XM; 1 ♂ 1 ♀, Xidianzhan, Nanling National Nature Reserve, 1. X. 2004, Wang XM leg; 3 ♂, Hengshitang, Shimentai National Nature Reserve, 5. X. 2004, Wang XM leg; 29 ♂ 5 ♀, Yangqiu Mountains, Shimentai National Nature Reserve, 6. X. 2004, Wang XM leg; 6 ♂ 2 ♀, Huangniping, Shimentai National Nature Reserve, 7. X. 2004, Wang XM leg; 4 ♂, Wulangzhang, Shimentai National Nature Reserve, 8. X. 2004, Wang XM leg; 4 ♂ 5 ♀, Nankunshan National Forest Park, 21. X. 2004, Wang XM leg; 7 ♂ 10 ♀, Shimen, Shimentai National Nature Reserve, 29. X. 2004, Wang XM leg; 13 ♂ 15 ♀, Huangdong, Shimentai National Nature Reserve, 30. X. 2004, Wang XM leg; 16 ♂ 18 ♀, Huangdong, Shimentai National Nature Reserve, 31. X. 2004, Wang XM leg; 4 ♀, Huangdong, Shimentai National Nature Reserve, 1. XI. 2004, Wang XM leg; 3 ♂ 2 ♀, Qinshuigu, Nanling National Nature Reserve, 21. IX. 2005, Wang XM leg. Guangxi: 4 ♂ 8 ♀, Hillside, Mao’ershan National Nature Reserve, 16 – 17. X. 2004, Wang XM leg; 72 ♂ 72 ♀, Shili Canyon, Mao’ershan National Nature Reserve, 19. X. 2004, Wang XM leg; 2 ♂ 2 ♀, GaoZhai, Mao’ershan National Nature Reserve, 20. X. 2004, Wang XM leg; 4 ♂, Pinglong Mountains, Shiwandashan National Nature Reserve, 6 – 7. XI. 2004, Wang XM leg; 9 ♂ 1 ♀, Hongqi Forest Farm, Shiwandashan National Nature Reserve, 10. XI. 2004, Wang XM leg; 2 ♂ 3 ♀, Hongqi Forest Farm, Shiwandashan National Nature Reserve, 26. VII. 2005, Wang XM leg; 1 ♂ 3 ♀, Pinglong Mountains, Shiwandashan National Nature Reserve, 28. VII. 2005, Wang XM leg. Hainan: 1 ♂, Xin’an, Diaoluoshan National Nature Reserve, 18. IX. 1995, Peng ZQ leg; 1 ♂, Tianchi, Jianfengling National Nature Reserve, IX. 1995, Peng ZQ leg; 1 ♂ 1 ♀, Wufenchang, Limushan National Forest Park, IX. 1995, Peng ZQ leg; 2 ♂ 1 ♀, Limushan National Forest Park, 21. IV. 1996, Peng ZQ leg; 1 ♂, Bawangling National Nature Reserve, 5. V. 2005, Wang XM leg; 1 ♀, Qixianling Hot Springs National Forest Park, 7. V. 2005, Wang XM leg; 2 ♂, Diaoluoshan National Nature Reserve, 8. V. 2005, Wang XM leg; 1 ♂, Tianchi, Jianfengling National Nature Reserve, 18. VII. 2006, Wang XM leg; 1 ♂, Limushan National Forest Park, 22. VII. 2006, Wang XM leg; 1 ♂ 1 ♀, Limushan National Forest Park, 23. VII. 2006, Wang XM 2. VIII. 1997, Peng ZQ leg; 1 ♀, Leigongshan National Nature Reserve, 10. VIII. 2006, Wang XM leg; 1 ♀, Datangwan, Leigongshan National Nature Reserve, 11. VIII. 2006, Wang XM leg; 1 ♀, Xiaodanjiang, Leigongshan National Nature Reserve, 13. VIII. 2006, Wang XM leg; 1 ♂ 2 ♀, Yaogu, Maolan National Nature Reserve, 15. VIII. 2006, Wang XM leg; 1 ♀, Maolan National Nature Reserve, 16. VIII. 2006, Wang XM leg; 1 ♀, Taojiang Town, Rongjiang County, 1000 m, 10 – 11. X. 2008, Chen XS leg; 1 ♂, Xiaodanjiang, Leigongshan National Nature Reserve, 12 – 13. X. 2008, Chen XS leg; 1 ♂, Jiaomi Village, Taijiang County, 710 m, 14. X. 2008, Chen XS leg; 8 ♂ 12 ♀, Dongtang, Maolan National Nature Reserve, 730 m, 15 – 18. X. 2008, Chen XS leg; 1 ♂ 8 ♀, Weng’ang, Maolan National Nature Reserve, 750 m, 20 – 23. X. 2008, Chen XS leg. Yunnan: 1 ♂, Botanic Garden, Jinghong City, 22. VIII. 2005, Wang XM leg; 1 ♂, Ruili City, 27. VIII. 2005, Wang XM leg; 1 ♂, Tongbiguan Town, Yingjiang County, 30. VIII. 2005, Wang XM leg; 1 ♂, Gaoligongshan National Nature Reserve, 31. VIII. 2005, Wang XM leg; 1 ♂, National highway 213 route, Mengla County, 12 – 13. X. 2006, Wang XM leg; 1 ♀, Dajian Mountains, Daweishan National Nature Reserve, 2100 m, 20 – 21. IV. 2008, Wang XM leg; 2 ♂ 2 ♀, Menglun Town, Mengla County, 1160 m, 29. IV. 2008, Wang XM leg; 4 ♂ 3 ♀, Gongxin Village, Menglian County, 1500 m, 8. V. 2008, Wang XM leg; 3 ♂ 3 ♀, Yaoqv Town, Mengla County, 700 m, 7 – 8. V. 2009, Wang XM leg; 1 ♂, Galaxi Village, Lianhuatan Town, Hekou County, 21. V. 2009, Wang XM leg; 1 ♂ 1 ♀, Huayudong National Forest Park, 123 m, 2. VIII. 2013, Chen XS leg. Hongkong: 1 ♂, Shing Mum Country Park, 206 m, 13. V. 2018, Wang XM leg. Taiwan: 1 ♀, Aowanda, Jenai Town, Nantou County, 1280 m, 21. X. 2012, Wang XM leg; 2 ♀, Dabang Village, Alishan Mountains, Chiayi County, 960 – 1600 m, 24. X. 2012, Wang XM leg; 1 ♂ 4 ♀, Shuangli Village, Pingtung County, 200 – 400 m, 27. X. 2012, Wang XM leg; 1 ♂ 3 ♀, Nanheng, Chihshang Town, Taitung County, 679 m, 29. X. 2012, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F612FF87FF7BFA650E3CFBFC.taxon	distribution	Distribution. China (Zhejiang, Anhui, Fujian, Jiangxi, Henan, Hubei, Hunan, Guangdong, Guangxi, Hainan, Chongqing, Sichuan, Guizhou, Yunnan, Hongkong, Taiwan); Japan.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F612FF87FF7BFA650E3CFBFC.taxon	discussion	Remarks. This species varies in elytral patterns. The specimens from Taiwan have yellow pronotum, sometimes with a black spot at base, yellow scutellar shield, and yellow elytra with five black spots. Some specimens from Hubei, Guangxi and Guizhou, elytra are yellow with characteristic black pattern as show in Figure 25 d. This species closely resembles Propipus subopacus Weise, 1901 (Bielawski, 1961; Yu et al., 1993) from Sumatra, the type species of the monotypic genus Propipus Weise, 1901, and can hardly be distinguished from the latter. However, since we were not able to examine the type series of Propipus subopacus deposited in Senckenberg German Entomological Institute in Berlin, the relationship between Propipus and Sasajiscymnus remains unknown.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F617FF89FF7BFB2D08F6FD28.taxon	description	(Figs 26, 68) Chinese name: DZũ ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F617FF89FF7BFB2D08F6FD28.taxon	diagnosis	Diagnosis. This species is similar to S. seboshii (Ohta, 1929), but can be distinguished from the later by more slender penis guide, and the parameres about 1 / 4 as long as penis guide (Figs 26 j – k). In S. seboshii, penis guide is more robust and parameres are about 2 / 5 as long as penis guide (Figs 34 j – k).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F617FF89FF7BFB2D08F6FD28.taxon	description	Description. TL: 1.81 – 2.26 mm, TW: 1.33 – 1.77 mm, TH: 0.75 – 1.04 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 26 a – f). Head yellow to black, with interocular distance distinctly greater than eye width in frontal view. Pronotum yellowish brown. Scutellar shield yellow to black. Elytral patterns varying from mostly black to predominantly yellow with black spots. Ventral surfaces yellow to black. Legs yellow to black. Abdominal postcoxal lines incomplete, slightly recurved toward the base of ventrite (Fig. 26 g). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis constricted with membranous appendage (Figs 2 h – i). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 1 / 4 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, then gradually narrowing to blunt apex (Figs 26 j – k).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F617FF89FF7BFB2D08F6FD28.taxon	materials_examined	Material examined. Holotype: China: Yunnan: ♂, Wenbi Mountain Scenic Spot, Lijiang City, 20. IV. 1997, Yao DF leg (BAAF). Paratypes: China: Yunnan: 1 ♀, Same data as holotype (BAAF); 2 ♀, Wenbi Mountain Scenic Spot, Lijiang City, 13. X. 1996, collector unknown (BAAF); 1 ♀, Wenbi Mountain Scenic Spot, Lijiang City, 15. X. 1996, Yu GY leg (BAAF); 1 ♂, Wenbi Mountain Scenic Spot, Lijiang City, 2600 m, 18. IX. 1997, Yu GY leg (BAAF); 2 ♂, Baishuihe, Yulong Snow Mountain Scenic Spot, Lijiang City, 2800 m, 21. IX. 1997, Montgomery M leg (BAAF); 1 ♀, Yunnan, No. 25, collected details unknown (BAAF); 1 specimen (undissected), Yunnan, No. 22, collected details unknown (BAAF); 1 specimen (undissected), Wenfeng Temples, Lijiang City, 2600 m, 24. IX. 1997, Yu GY leg (BAAF). Sichuan: 1 ♀, Nibagou, Jiajinshan National Forest Park, 14. X. 1996, collector unknown (BAAF). Other material (SCAU). China: Henan: 3 ♂ 4 ♀, Baotianman National Nature Reserve, 6. VII. 2009, Wang XM leg; 3 ♂, Baotianman National Nature Reserve, 800 – 1300 m, 7 – 8. VII. 2009, Wang XM leg; 1 ♀, Laojieling, Xixia County, 1430 m, 9. VII. 2009, Wang XM leg; 6 ♀, Baiyunshan National Nature Reserve, 1375 m, 14. VII. 2009, Wang XM leg. Sichuan: 1 ♂, Puge County, 2000 m, 15. IX. 2007, Wang XM leg; 3 ♀, Meigu Dafengding National Nature Reserve, 2400 m, 21. IX. 2007, Wang XM leg; 3 ♂ 2 ♀, Heizhugou National Nature Reserve, 1900 m, 22 – 23. IX. 2007, Wang XM leg; 1 ♀, Mahuanggou, Jiajinshan National Forest Park, 2330 m, 1. X. 2007, Wang XM leg; 1 ♂ 3 ♀, Labahe National Nature Reserve, 1100 m, 4. X. 2007, Wang XM leg. Guizhou: 2 ♂, Lianhuaping, Leigongshan National Nature Reserve, 8. X. 2008, Wang XM leg. Yunnan: 1 ♂, Dadugang Town, Jinghong City, 1050 m, 5. V. 2009, Wang XM leg; 2 ♂ 3 ♀, Xujiaba, Ailaoshan National Nature Reserve, 2484 m, 15. VIII. 2013, Chen XS leg; 4 ♂, County highway J 22 route, Wuliangshan National Nature Reserve, 2400 m, 18. VIII. 2013, Chen XS leg. Shaanxi: 2 ♂, Taibaishan National Nature Reserve, 1600 m, 18. VII. 2009, Wang XM leg; 1 ♀, Daguping Village, Foping National Nature Reserve, 1170 m, 21. VII. 2009, Wang XM leg; 3 ♀, Fengyu Forest Farm, 1500 m, 28. VII. 2009, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F617FF89FF7BFB2D08F6FD28.taxon	distribution	Distribution. China (Henan, Sichuan, Guizhou, Yunnan, Shaanxi).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F619FF88FF7BFD120A68FC00.taxon	description	Chinese name: ṁ 叶 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F619FF88FF7BFD120A68FC00.taxon	diagnosis	Diagnosis. This species is similar to S. seminigrinus (Yu & Pang, 1993) in structures of the male genitalia, but can be distinguished from the later by elytra entirely black, and the penis guide approximately widest at 1 / 2 length in ventral view (Figs 27 a – b, 27 h). In S. seminigrinus, elytra are yellowish brown with a large, black marking near the base (sometimes absent), and penis guide becomes widest at base in ventral view (Figs 38 a – b, 38 k).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F619FF88FF7BFD120A68FC00.taxon	description	Description. TL: 1.61 – 2.19 mm, TW: 1.13 – 1.56 mm, TH: 0.73 – 0.95 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 27 a – c). Head yellow with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow. Scutellar shield and elytra entirely black. Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 27 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis expanded with membranous appendage (Figs 27 e – f). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 1 / 2 as long as penis guide, with a tuft of long setae at apex; penis guide approximately widest at 1 / 2 length in ventral view, apex with a distinct protuberance (Figs 27 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F619FF88FF7BFD120A68FC00.taxon	materials_examined	Type material. Holotype: China: Xizang: ♂, Beibeng Town, Motuo County, 800 m, 9. X. 2011, Huo LZ leg (SCAU). Paratypes (SCAU): China: Xizang: 2 ♂ 2 ♀, Yarang Village, Muotuo County, 768 – 1030 m, 24. X. 2007, Wang XM leg; 1 ♂, Zhamo Highroad 113 K, Bomi County, 835 m, 30 – 31. X. 2007, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F619FF88FF7BFD120A68FC00.taxon	distribution	Distribution. China (Xizang).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F619FF88FF7BFD120A68FC00.taxon	etymology	Etymology. The specific epithet latus derives from the Latin word “ latus ” and refers to the wide penis guide in ventral view.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F618FF8BFF7BFC310F7BFDBC.taxon	description	(Figs 28, 72) Chinese name: 弯 Ṅḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F618FF8BFF7BFC310F7BFDBC.taxon	diagnosis	Diagnosis. This species is similar to S. bivalvis (Yu, 1997) in structures of the male genitalia, but can be distinguished from the later by elytra with two yellow C-shaped patterns, and parameres about 1 / 3 as long as penis guide (Figs 28 a – b, 28 g). In S. bivalvis, elytra are black with yellowish brown apices, parameres are about 1 / 2 as long as penis guide (Figs 39 a – b, 39 g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F618FF8BFF7BFC310F7BFDBC.taxon	description	Description. TL: 1.55 – 2.45 mm, TW: 1.07 – 1.83 mm, TH: 0.67 – 0.92 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 28 a – c). Head yellowish brown with interocular distance distinctly greater than eye width in frontal view. Pronotum yellowish brown, sometimes with a black spot. Scutellar shield black. Elytra with two yellow C-shaped patterns as show in Figure 28 a. Ventral surfaces yellow to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 28 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis constricted with membranous appendage (Figs 28 e – f). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 3 in ventral view, apically pointed (Figs 28 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F618FF8BFF7BFC310F7BFDBC.taxon	materials_examined	Material examined. Holotype: China: Henan: ♂, Jigongshan National Nature Reserve, 12. VII. 1997, Yu GY leg (BAAF). Other material (SCAU). China: Henan: 2 ♀, Jigongshan National Nature Reserve, 10. VII. 1997, Peng ZQ leg; 1 ♂ 3 ♀, Baotianman National Nature Reserve, 6. VII. 2009, Wang XM leg; 1 ♀, Longyuwan National Forest Park, 1390 m, 11 – 12. VII. 2009, Wang XM leg; 3 ♂ 2 ♀, Baiyunshan National Nature Reserve, 1375 m, 14. VII. 2009, Wang XM leg. Shanxi: 3 ♂, Lishan National Nature Reserve, 800 – 1400 m, 3. VIII. 2011, Wang XM leg. Anhui: 2 ♂ 1 ♀, Yaoluoping National Nature Reserve, 1000 m, 28 – 29. IX. 2010, Wang XM leg; 2 ♂, Chuanling, Yaoluoping National Nature Reserve, 800 m, 2. X. 2010, Wang XM leg; 2 ♂, Jinzhai Tianma National Nature Reserve, 600 m, 3 – 5. X. 2010, Wang XM leg. Hubei: 1 ♂ 1 ♀, Guanmen Mountains, Shennongjia National Nature Reserve, 1260 m, 16 – 19. VII. 2007, Wang XM leg; 1 ♂, Guanmen Mountains, Shennongjia National Nature Reserve, 1260 m, 2. VIII. 2007, Wang XM leg; 1 ♂ 5 ♀, Wudang Mountains, Shiyan City, 700 – 930 m, 8. VIII. 2007, Wang XM leg. Shaanxi: 1 ♀, Songpingsi, Taibaishan National Nature Reserve, 29. VIII. 2006, Wang XM leg; 1 ♂ 1 ♀, Fengyu Forest Farm, 1500 m, 28. VII. 2009, Wang XM leg; 2 ♀, Cuihuan Mountains, Chang’an District, Xi’an City, 1300 m, 30. VII. 2009, Wang XM leg. Gansu: 1 ♂ 1 ♀, Lianfeng Mountains, Weiyuan County, 22. VIII. 2007, Liang JB leg; 2 ♂ 2 ♀, Maijishan National Forest Park, Maiji District, Tianshui City, 1500 m, 10. VIII. 2008, Wang XM leg. Ningxia: 3 ♂ 2 ♀, Hougou, Songdian Village, Liupanshan National Nature Reserve, 1850 m, 8. VIII. 2009, Wang XM leg; 26 ♂ 34 ♀, Baiyunsi, Liupanshan National Nature Reserve, 2300 m, 10. VIII. 2009, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F618FF8BFF7BFC310F7BFDBC.taxon	distribution	Distribution. China (Shanxi, Anhui, Henan, Hubei, Shaanxi, Gansu, Ningxia).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61BFF8AFF7BF8F2092AFCB8.taxon	description	(Figs 1 h, 29, 71) Chinese name: R Ẇḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61BFF8AFF7BF8F2092AFCB8.taxon	diagnosis	Diagnosis. This species can be easily distinguished from other Sasajiscymnus by its black elytra bearing yellowish brown apices with two brown spots close to suture, and the penis apically constricted with fin-shaped appendage (Figs 29 a – b, 19 f).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61BFF8AFF7BF8F2092AFCB8.taxon	description	Description. TL: 1.74 – 2.04 mm, TW: 1.27 – 1.55 mm, TH: 0.65 – 0.84 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 29 a – c). Head yellowish brown with interocular distance distinctly greater than eye width in frontal view. Pronotum yellowish brown. Scutellar shield brown. Elytra black with yellowish brown apices with two brown spots close to suture. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, slightly recurved toward the base of ventrite (Fig. 29 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis distinctly constricted with fin-shaped appendage (Figs 29 e – f). Penis guide with prominent keel at base in lateral view; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide triangular in ventral view, widest at base, then gradually narrowing to apex (Figs 29 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61BFF8AFF7BF8F2092AFCB8.taxon	materials_examined	Material examined. Holotype: China: Fujian: ♂, Xianfengling, Wuyishan National Nature Reserve, 2. X. 1980, Huang BK leg (SCAU). Other material (SCAU). China: Henan: 1 ♂ 2 ♀, Jigongshan National Nature Reserve, 697 m, 4. VII. 2009, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61BFF8AFF7BF8F2092AFCB8.taxon	distribution	Distribution. China (Fujian, Henan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61DFF8CFF7BFF750A36FEE4.taxon	description	(Figs 30, 70) Chinese name: ḦṄḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61DFF8CFF7BFF750A36FEE4.taxon	diagnosis	Diagnosis. This species can be distinguished from other species of Sasajiscymnus by the pointed apex of penis with membranous appendage, and elytra with characteristic pattern (Figs 30 a – b, 30 f).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61DFF8CFF7BFF750A36FEE4.taxon	description	Description. TL: 1.58 – 2.17 mm, TW: 1.17 – 1.61 mm, TH: 0.82 – 1.14 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 30 a – c). Head yellow with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow. Scutellar shield yellowish brown. Elytron with characteristic yellow and black pattern as show in Figure 30 a. Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 30 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis constricted with membranous appendage (Figs 30 e – f). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 2 / 5 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, then gradually narrowing to blunt apex (Figs 30 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus. Type material. Holotype: China: Xizang: ♂, Chayu County, 2320 m, 12. X. 2007, Wang XM leg (SCAU). Paratypes (SCAU): China: Xizang: 7 ♂ 11 ♀, Same data as holotype; 7 ♂ 6 ♀, Cibagou National Nature Reserve, 2000 m, 13. X. 2007, Wang XM leg; 1 ♂ 1 ♀, Xiachayu Town, Chaiyu County, 1800 m, 14. X. 2007, Wang XM leg; 3 ♂ 1 ♀, North Downtown of Chaiyu County, 2300 m, 18. X. 2007, Wang XM leg; 1 ♂ 6 ♀, Cibagou National Nature Reserve, 1615 m, 26. IX. 2011, Huo LZ leg; 1 ♂, Shangchayu Town – Xiachayu Town, Chayu County, 1650 m, 27. IX. 2011, Huo LZ leg; 1 ♀, Chayu County, 2400 m, 29. IX. 2011, Huo LZ leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61DFF8CFF7BFF750A36FEE4.taxon	distribution	Distribution. China (Xizang).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61DFF8CFF7BFF750A36FEE4.taxon	etymology	Etymology. The specific epithet obliquus derives from the Latin word “ obliquus ” and refers to the oblique marks on the elytra.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61CFF8FFF7BFE55087AFD04.taxon	description	Chinese name: DZ Ẹḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61CFF8FFF7BFE55087AFD04.taxon	diagnosis	Diagnosis. This species is similar to S. seboshii (Ohta, 1929), but can be distinguished from the later by the elytra with black humeral angles, abdominal postcoxal lines recurved toward the base of ventrite and more elongate penis guide (Figs 31 a – d, 19 g – h). In S. seboshii, humeral angles of elytra are yellowish brown, abdominal postcoxal lines not recurved toward the base of ventrite and penis guide is shorter (Figs 34 a – g, 19 j – k).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61CFF8FFF7BFE55087AFD04.taxon	description	Description. TL: 1.94 – 2.25 mm, TW: 1.38 – 1.63 mm, TH: 0.90 – 1.09 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 31 a – c). Head yellow with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow. Scutellar shield yellowish brown. Elytra black with yellow apices and a large yellowish brown spot in the middle part. Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 31 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis constricted with membranous appendage (Figs 31 e – f). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base, then gradually narrowing to pointed apex (Figs 31 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61CFF8FFF7BFE55087AFD04.taxon	materials_examined	Type material. Holotype: China: Xizang: ♂, 52 K – 65 K of Zhamo Highroad, Motuo County, 2800 – 3500 m, 13. X. 2011, Huo LZ leg (SCAU). Paratypes (SCAU): China: Xizang: 1 ♀, Zhamo Highroad 113 K, Bomi County, 835 m, 30 – 31. X. 2007, Wang XM leg; 1 ♂, Pailong Town, Bayi District, Linzhi City, 13. X. 2009, Wang XM leg; 1 ♀, Hanmi, Muotuo County, 800 – 2100 m, 5 – 8. X. 2011, Huo LZ leg; 1 ♂, Chilongzangbu, Linzhi City, 2200 m, 19. X. 2011, Huo LZ leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61CFF8FFF7BFE55087AFD04.taxon	distribution	Distribution. China (Xizang).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61CFF8FFF7BFE55087AFD04.taxon	etymology	Etymology. The specific epithet niganulus derives from the Latin words “ niger ” (= black) and “ anulus ” (= annular), and refers to the black ring-spot on the disc of elytra.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61FFF8EFF7BFD35095EFE9C.taxon	description	(Figs 32, 69) Chinese name: Ñġ ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61FFF8EFF7BFD35095EFE9C.taxon	diagnosis	Diagnosis. This species is similar to S. seminigrinus (Yu & Pang, 1993) in body size and external colouration, but can be distinguished from the later by the penis guide apically blunt in ventral view (Fig. 32 d). In S. seminigrinus, penis guide has a small protuberance at apex in ventral view (Fig. 38 k).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61FFF8EFF7BFD35095EFE9C.taxon	description	Description. TL: 2.0 mm, TW: 1.3 mm. Body oval, moderately convex, surfaces covered with short pubescence. Head yellow with interocular distance distinctly greater than eye width in frontal view. Pronotum and scutellar shield yellowish brown. Elytra yellow with a large black marking near the base. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 32 a). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis expanded with distinct membranous appendage (Figs 32 b – c). Penis guide with prominent keel at base in lateral view, then tapering towards apex; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view (Fig. 32 d). Female genitalia: Unknow.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61FFF8EFF7BFD35095EFE9C.taxon	materials_examined	Material examined. Holotype: China: Taiwan: ♂, Puli Town, Nantou County, 17. III. 1972, Yang CT leg (NCHU).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61FFF8EFF7BFD35095EFE9C.taxon	distribution	Distribution. China (Taiwan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61EFF91FF7BFE8D0968FDE0.taxon	description	Chinese name: ỜṄḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61EFF91FF7BFE8D0968FDE0.taxon	diagnosis	Diagnosis. This species is similar to S. aureus sp. nov., but can be distinguished from the later by the elytra with black spots and penis guide approximately parallel sided for basal 1 / 2 in ventral view (Figs 33 a – c, 33 h). In S. aureus, elytra are entirely yellow and the penis guide is widest at base (Figs 21 a – c, 21 h).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61EFF91FF7BFE8D0968FDE0.taxon	description	Description. TL: 1.92 – 2.29 mm, TW: 1.39 – 1.71 mm, TH: 0.72 – 0.84 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 33 a – c). Head yellow with interocular distance distinctly greater than eye width in frontal view. Pronotum and scutellar shield yellow. Elytra yellow and each elytron usually with three black spots: basal spot situated at near humeral calli (sometimes reduced); second spot situated on suture, semicircular and small (sometimes absent); posterior spot situated at apical part of elytron (sometimes absent). Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 33 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis constricted with distinct membranous appendage (Figs 33 e – f). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 2 in ventral view, then gradually narrowing to pointed apex (Figs 33 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61EFF91FF7BFE8D0968FDE0.taxon	materials_examined	Type material. Holotype: China: Xizang: ♂, North Downtown of Chaiyu County, 2300 m, 18. X. 2007, Wang XM leg (SCAU). Paratypes (SCAU): China: Xizang: 2 ♂ 4 ♀ Same data as holotype; 1 ♂, Xiachayu Town, Chaiyu County, 1800 m, 14. X. 2007, Wang XM leg; 1 ♀, 52 K – 65 K of Zhamo Highroad, Motuo County, 2800 – 3500 m, 13. X. 2011, Huo LZ leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61EFF91FF7BFE8D0968FDE0.taxon	distribution	Distribution. China (Xizang).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F61EFF91FF7BFE8D0968FDE0.taxon	etymology	Etymology. The specific epithet maculatus derives from the Latin word “ maculatus ” meaning spotted, and refers to the elytra with black spots.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F601FF90FF7BFD510E2CFF55.taxon	description	(Figs 34, 71)	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F601FF90FF7BFD510E2CFF55.taxon	vernacular_names	Chinese name: AE Ṅḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F601FF90FF7BFD510E2CFF55.taxon	diagnosis	Diagnosis. This species is similar to S. aureus sp. nov. in structures of the male genitalia, but can be distinguished from the later by the black elytra with a large yellowish brown heart-shaped marking in the middle part, and the parameres about 2 / 5 as long as penis guide (Figs 34 a – b, 34 j). In S. aureus, elytra are predominantly yellow with some black spots, and the parameres are about 1 / 3 as long as penis guide (Figs 21 a – b, 21 g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F601FF90FF7BFD510E2CFF55.taxon	description	Description. TL: 1.53 – 2.28 mm, TW: 1.14 – 1.87 mm, TH: 0.76 – 1.12 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 34 a – f). Head yellow to yellowish brown, with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow. Scutellar shield brown. Elytra black with yellow apices and humeral angles, and a large yellowish brown spot presenting in the middle part; sometimes median macula extends to lateral margins of elytra. Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 34 g). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis constricted with membranous appendage (Figs 34 h – i). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 2 / 5 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, then gradually narrowing to pointed apex (Figs 34 j – k). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F601FF90FF7BFD510E2CFF55.taxon	materials_examined	Material examined. China: Anhui: 1 ♂ 1 ♀, Hengdu Town, Shitai County, 230 m, 20. IX. 2010, Wang XM leg; 1 ♂, Yaoluoping National Nature Reserve, 600 – 800 m, 30. IX. 2010, Wang XM leg; 2 ♂, Chuanling, Yaoluoping National Nature Reserve, 800 m, 2. X. 2010, Wang XM leg. Fujian: 1 ♂ 1 ♀, Jiangxikou, Tongmu Village, Wuyishan National Nature Reserve, 1130 m, 19. IX. 2012, Huo LZ leg. Hubei: 1 ♀, Guanmen Mountains, Shennongjia National Nature Reserve, 1260 m, 18. VII. 2007, Wang XM leg; 1 ♂, Banqiao, Shennongjia National Nature Reserve, 1170 m, 21 – 24. VII. 2007, Wang XM leg. Guangxi: 2 ♂, Jiuniutang, Mao’ershan National Nature Reserve, 18. X. 2004, Wang XM leg; 1 ♀, Shili Canyon, Mao’ershan National Nature Reserve, 19. X. 2004, Wang XM leg. Shaanxi: 1 ♂ 1 ♀, Daguping Village, Foping National Nature Reserve, 1170 m, 21. VII. 2009, Wang XM leg; 1 ♂, Xihe, Daguping Village, Foping National Nature Reserve, 1100 m, 22. VII. 2009, Wang XM leg; 1 ♂, Fengyu Forest Farm, 1500 m, 28. VII. 2009, Wang XM leg. Gansu: 5 ♂ 3 ♀, Maijishan National Forest Park, Maiji District, Tianshui City, 1500 m, 10. VIII. 2008, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F601FF90FF7BFD510E2CFF55.taxon	distribution	Distribution. China (Anhui, Fujian, Hubei, Shaanxi, Gansu, Taiwan); Japan.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F601FF90FF7BFD510E2CFF55.taxon	discussion	Remarks. This species varies in elytral patterns. Some specimens from Shaanxi, Gansu and Hubei have predominantly pale yellow elytra, with black markings reducing to small spots or narrow lines near humeral calli.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F603FF92FF7BFF750961FEC0.taxon	description	Chinese name: Ṛūḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F603FF92FF7BFF750961FEC0.taxon	diagnosis	Diagnosis. This species is similar to S. maculatus sp. nov. in structures of the male genitalia, but can be distinguished from the later by the elytra black with yellowish brown apices and humeral angles (Figs 35 a – c). In S. maculatus, elytra are yellow with black spots (Figs 33 a – c).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F603FF92FF7BFF750961FEC0.taxon	description	Description. TL: 2.05 mm, TW: 1.45 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 35 a – c). Head yellowish brown with interocular distance distinctly greater than eye width in frontal view. Pronotum and scutellar shield yellowish brown. Elytra black with yellowish brown apices and humeral angles. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 35 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis with membranous appendage (Figs 35 e – f). Penis guide with prominent keel at base in lateral view, then gradually narrowing to pointed apex; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 3 in ventral view, then gradually narrowing to apex (Figs 34 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F603FF92FF7BFF750961FEC0.taxon	materials_examined	Material examined. Holotype: China: Yunnan: ♂, Heyuan, Lijiang City, 2820 - 2840 m, 1. XI. 1995, Liu SJ leg (BAAF).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F603FF92FF7BFF750961FEC0.taxon	distribution	Distribution. China (Yunnan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F602FF92FF7BFE71095EF828.taxon	description	(Figs 36, 70) Chinese name: 片 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F602FF92FF7BFE71095EF828.taxon	diagnosis	Diagnosis. This species is similar to S. variabilis sp. nov. in structures of the male genitalia, but can be distinguished from the later by the penis guide blunt apically and elytra mostly black with yellowish brown apices (Fig. 36 d). In S. variabilis, penis guide is pointed at apex and elytra have variable markings (Figs 37 a – f, 37 k).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F602FF92FF7BFE71095EF828.taxon	description	Description. TL: 2.1 – 2.3 mm, TW: 1.5 – 1.6 mm. Body oval, moderately convex, surfaces covered with short pubescence. Head yellowish brown with interocular distance distinctly greater than eye width in frontal view. Pronotum yellowish brown. Scutellar shield black. Elytra black with yellowish brown apices. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 36 a). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis with a deep emargination and membranous appendage (Figs 36 b – c). Parameres about 2 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, apically rounded (Fig. 36 d). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F602FF92FF7BFE71095EF828.taxon	materials_examined	Material examined. Holotype: China: Taiwan: ♂, Puli Town, Nantou County, 17. III. 1972, Yang CT leg (NCHU). Paratype: China: Taiwan: 1 ♀, Same data as holotype (NCHU).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F602FF92FF7BFE71095EF828.taxon	distribution	Distribution. China (Taiwan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F605FF94FF7BFF7509ADF83F.taxon	description	(Figs 37, 66) Chinese name: 变 Ṅḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F605FF94FF7BFF7509ADF83F.taxon	diagnosis	Diagnosis. This species is very distinctive by its elytra with variable markings, elongate penis guide, and penis with a deep emargination at apex (Figs 37 a – f, 37 i).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F605FF94FF7BFF7509ADF83F.taxon	description	Description. TL: 1.40 – 2.11 mm, TW: 0.93 – 1.60 mm, TH: 0.59 – 0.98 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 37 a – f). Head yellowish brown with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow sometimes with a large black spot. Scutellar shield black. Elytron with characteristic yellow and black pattern. Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, slightly recurved toward the base of ventrite (Fig. 37 g). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis with a deep emargination and membranous appendage (Figs 37 h – i). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 1 / 2 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 3 in ventral view, apically rounded (Figs 37 j – k). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F605FF94FF7BFF7509ADF83F.taxon	materials_examined	Type material. Holotype: China: Yunnan: ♂, Jinuoshan Town, Jinghong City, 6. V. 2009, Wang XM leg (SCAU). Paratypes (SCAU): China: Yunnan: 1 ♂ 2 ♀, Same data as holotype; 1 ♀, Nansan, Zhenkang County, 28. X. 2000, Peng ZQ leg; 1 ♂, Dongfeng Farm, Jinghong City, 19. IV. 2002, Peng ZQ leg; 1 ♂ 1 ♀, Menglun Town, Mengla County, 21. VIII. 2005, Wang XM leg; 1 ♂, Botanic Garden, Jinghong City, 22. VIII. 2005, Wang XM leg; 1 ♂, Longmen Village, Shangyong Town, Mengla County, 8. X. 2006, Wang XM leg; 1 ♂, Mangbing, Yingjiang City, 29. VIII. 2005, Wang XM leg; 2 ♂ 2 ♀, National highway 213 route, Mengla County, 12 – 13. X. 2006, Wang XM leg; 9 ♂ 13 ♀, Dadugang Town, Jinghong City, 950 m, 26. IV. 2008, Wang XM leg; 2 ♂ 3 ♀, Menglun Town, Mengla County, 1160 m, 29. IV. 2008, Wang XM leg; 7 ♂ 4 ♀, Sangyong Village – Longmen Village, Mengla County, 1. V. 2008, Wang XM leg; 2 ♂ 1 ♀, Lafu Village, Menglian County, 1130 m, 7. V. 2008, Wang XM leg; 1 ♂, Gongxin Village, Menglian County, 1500 m, 8. V. 2008, Wang XM leg; 13 ♂ 7 ♀, Longtan, Ximeng County, 900 m, 9 – 10. V. 2008, Wang XM leg; 2 ♂ 3 ♀, Lancang County, 1300 m, 11. V. 2008, Wang XM leg; 2 ♂ 4 ♀, Banlao Town, Cangyuan County, 1100 m, 16. V. 2008, Wang XM leg; 1 ♂ 1 ♀, Tongbiguan Town, Yingjiang County, 1000 m, 22 – 23. V. 2008, Wang XM leg; 10 ♂ 10 ♀, Caiyanghe National Forest Park, 4. V. 2009, Wang XM leg; 3 ♂ 3 ♀, Dadugang Town, Jinghong City, 1050 m, 5. V. 2009, Wang XM leg; 4 ♂ 8 ♀, Yaoqv Town, Mengla County, 700 m, 7 – 8. V. 2009, Wang XM leg; 1 ♂, Longmen Village, Shangyong Town, Mengla County, 1027 m, 9. V. 2009, Wang XM leg; 2 ♂ 1 ♀, Nanping, Mengman Town, Mengla County, 745 m, 10. V. 2009, Wang XM leg; 1 ♂, Meng’a Town, Menghai County, 1170 m, 12. V. 2009, Wang XM leg; 2 ♂, Daheishan Town, Jiangcheng County, 1030 m, 17. V. 2009, Wang XM leg; 1 ♂, Maku Village, Gongshan County, 1600 m, 1. VIII. 2010, Wang XM leg; 2 ♂ 1 ♀, Yaoqv Town, Mengla County, 818 m, 8. VIII. 2013, Wang XM leg; 1 ♀, Jinuoshan Town, Jinghong City, 992 m, 10. VIII. 2013, Chen XS leg; 1 ♂ 2 ♀, Bulangshan Town, Menghai County, 1188 m, 12. VIII. 2013, Chen XS leg; 1 ♀, Xujiaba, Ailaoshan National Nature Reserve, 2524 m, 17. VIII. 2013, Chen XS leg. Hubei: 2 ♂, Banqiao, Shennongjia National Nature Reserve, 1170 m, 21 – 24. VII. 2007, Wang XM leg; 1 ♀, Yinyuhe, Shennongjia National Nature Reserve, 1600 – 1800 m, 31. VII. 2007, Wang XM leg; 1 ♀, Wudang Mountains, Shiyan City, 700 – 930 m, 8. VIII. 2007, Wang XM leg. Guangdong: 1 ♂, Nankunshan National Forest Park, XI. 1986, Jiang N leg. Guangxi: 1 ♀, Hillside, Mao’ershan National Nature Reserve, 17. X. 2004, Wang XM leg; 3 ♂, Shili Canyon, Mao’ershan National Nature Reserve, 19. X. 2004, Wang XM leg; 3 ♀, GaoZhai, Mao’ershan National Nature Reserve, 20. X. 2004, Wang XM leg; 1 ♂, Pinglong Mountains, Shiwandashan National Nature Reserve, 6. XI. 2004, Wang XM leg; 1 ♂, Hongqi Forest Farm, Shiwandashan National Nature Reserve, 26. VII. 2005, Wang XM leg. Guizhou: 1 ♂, Laodong Village, Danzhai County, 4. X. 2008, Chen XS leg. Taiwan: 1 ♂, Taroko National Park, Hualien County, 200 – 912 m, 30. X. 2012, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F605FF94FF7BFF7509ADF83F.taxon	distribution	Distribution. China (Hubei, Guangdong, Guangxi, Guizhou, Yunnan, Taiwan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F605FF94FF7BFF7509ADF83F.taxon	discussion	Remarks. This species varies in elytral patterns. The specimens from Taiwan have black elytra with yellow apices, and each elytron have a yellow spot on medially; most specimens from Yunnan have yellow pronotum bearing with a large black spot, and have yellow elytra bearing with characteristic black pattern as show in Figure 37 d; other specimens usually have pattern like Figure 37 a.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F607FF96FF7BFF7508ADFAD5.taxon	description	Chinese name: 半DZ ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F607FF96FF7BFF7508ADFAD5.taxon	diagnosis	Diagnosis. This species is similar to S. latus sp. nov. in structures of the male genitalia, but can be distinguished from the later by the elytra yellowish brown with a black marking near the base (sometimes absent), and its penis guide widest near the base in ventral view (Figs 38 a – b, 38 k). In S. latus, elytra are entirely black and penis guide is approximately widest at 1 / 2 length in ventral view (Figs 27 a – b, 27 h).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F607FF96FF7BFF7508ADFAD5.taxon	description	Description. TL: 1.54 – 2.20 mm, TW: 1.09 – 1.64 mm, TH: 0.62 – 0.91 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 38 a – f). Head yellow to yellowish brown, with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow. Scutellar shield brown. Elytra yellowish brown with a black marking situated at the near base (sometimes reduced). Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, slightly recurved toward the base of ventrite (Fig. 38 g). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis expanded with membranous appendage (Figs 38 h – i). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 3 / 5 as long as penis guide, with a tuft of long setae at apex; penis guide nearly widest at base in ventral view, apex with a small protuberance (Figs 38 j – k). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F607FF96FF7BFF7508ADFAD5.taxon	materials_examined	Material examined. Holotype: China: Guangdong: ♂, Chebaling National Nature Reserve, 24. IV. 1991, Yu GY leg (SCAU). Allotype: China: Guangdong: 1 ♀, Same data as holotype (SCAU). Paratypes: China: Guangdong: 5 ♂ 4 ♀, Same data as holotype (SCAU); 1 ♀, Chebaling National Nature Reserve, 23. III. 1989, Yu GY leg (SCAU); 1 ♀, Chebaling National Nature Reserve, 2. XI. 1989, Yu GY leg (SCAU); 1 ♂, Chebaling National Nature Reserve, 3. XI. 1989, Yu GY leg (SCAU). Other material (SCAU). China: Guangdong: 2 ♂ 1 ♀, Chebaling National Nature Reserve, 23. III. 1989, Ren SX leg; 1 ♀, Chebaling National Nature Reserve, 24. IV. 1989, Tong XL leg; 1 ♂, Nankunshan National Forest Park, 21. X. 2004, Wang XM leg. Guangxi: 1 ♂, Hillside, Mao’ershan National Nature Reserve, 16. X. 2004, Wang XM leg; 2 ♂ 3 ♀, Pinglong Mountains, Shiwandashan National Nature Reserve, 6 – 8. XI. 2004, Wang XM leg; 2 ♂ 1 ♀, Hongqi Forest Farm, Shiwandashan National Nature Reserve, 9 – 10. XI. 2004, Wang XM leg; 1 ♂ 1 ♀, Nonggang National Nature Reserve, 3. VIII. 2005, Wang XM leg; 2 ♂, Pinglong Mountains, Shiwandashan National Nature Reserve, 28. VII. 2005, Wang XM leg. Hainan: 3 ♂ 1 ♀, Haikou City, 6. XI. 1989, Ren SX leg; 1 ♀, Maoyang Town, Qiongzhong County, IX. 1995, Peng ZQ leg; 1 ♀, Maoyang Town, Qiongzhong County, IX. 1995, Peng ZQ leg; 1 ♀, East Harbour National Nature Reserve, IX. 1995, Peng ZQ leg; 2 ♂, Honglin Forest Farm, Changjiang County, 10. XII. 1997, Peng ZQ leg; 1 ♂, Bawangling National Nature Reserve, 12. V. 2000, Peng ZQ leg. Guizhou: 2 ♂ 2 ♀, Dongtang, Maolan National Nature Reserve, 730 m, 15 – 18. X. 2008, Wang XM leg; 6 ♀, Weng’ang, Maolan National Nature Reserve, 750 m, 20 – 23. X. 2008, Wang XM leg. Yunnan: 2 ♂, Malipo County, 17. VIII. 2005, Wang XM leg; 4 ♂ 2 ♀, Huayudong National Forest Park, 123 m, 2. VIII. 2013, Chen XS leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F607FF96FF7BFF7508ADFAD5.taxon	distribution	Distribution. China (Guangdong, Guangxi, Hainan, Guizhou, Yunnan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F606FF98FF7BFA440E25FD70.taxon	description	(Figs 3 b, 3 f, 39, 72) Chinese name: R 瓣 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F606FF98FF7BFA440E25FD70.taxon	diagnosis	Diagnosis. This species is similar to S. curvatus (Yu, 1999) in structures of the male genitalia, but can be distinguished from the later by the black elytra with yellowish brown apices, and the parameres about 1 / 2 as long as penis guide (Figs 39 a – b, 39 g). In S. curvatus, elytra are black with two C-shaped yellow marking and parameres are about 1 / 3 as long as penis guide (Figs 28 a – b, 28 g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F606FF98FF7BFA440E25FD70.taxon	description	Description. TL: 1.98 – 2.68 mm, TW: 1.43 – 2.06 mm, TH: 0.95 – 1.21 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 39 a – c). Head yellowish brown with interocular distance distinctly greater than eye width in frontal view. Pronotum yellowish brown. Scutellar shield black. Elytra black with yellowish brown apices. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 39 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis constricted with membranous appendage (Figs 39 e – f). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 1 / 2 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 3 in ventral view, apically pointed (Figs 39 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus (Figs 3 b, 3 f).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F606FF98FF7BFA440E25FD70.taxon	materials_examined	Type material. Holotype: China: Hubei: ♂, Longmenhe National Forest Park, Xingshan County, 1300 m, 12. IX. 1994, Li FS leg (CAU). Other material (SCAU). China: Zhejiang: 1 ♂, Wangdongyang Mountain Wetland Nature Reserve, [119 ° 38 ' 12.09 " E 27 ° 41 ' 19.61 " N], 1261 m, 10. X. 2017, Tong JB leg. Anhui: 1 ♂ 1 ♀, Guniujiang National Nature Reserve, 24. IX. 2010, Wang XM leg; 1 ♂, Yaoluoping National Nature Reserve, 600 – 800 m, 30. IX. 2010, Wang XM leg; 2 ♀, Jinzhai Tianma National Nature Reserve, 600 m, 3 – 5. X. 2010, Wang XM leg. Fujian: 3 ♂ 1 ♀, Sanbailiao, Tianboyan National Nature Reserve, 690 m, 26. VIII. 2012, Huo LZ leg; 1 ♂, Fengtian, Tianboyan National Nature Reserve, 800 m, 27. VIII. 2012, Huo LZ leg; 1 ♂ 5 ♀, Zonghechang, Longxishan National Nature Reserve, 750 m, 30. VIII. 2012, Huo LZ leg; 1 ♂, Guadun, Wuyishan National Nature Reserve, 1200 m, 14. IX. 2012, Huo LZ leg; 3 ♂ 2 ♀, Masu, Wuyishan National Nature Reserve, 1291 m, 15. IX. 2012, Huo LZ leg; 4 ♂ 1 ♀, Tongmu Village, Wuyishan National Nature Reserve, 750 m, 17. IX. 2012, Huo LZ leg. Jiangxi: 2 ♂, Taihe County, 12. VIII. 2004, Wang XM leg; 1 ♂ 2 ♀, Xiangzhou, Jinggangshan National Nature Reserve, 14. VIII. 2004, Wang XM leg; 9 ♂ 4 ♀, Bijia Mountains, Jinggangshan National Nature Reserve, 15. VIII. 2004, Wang XM leg; 4 ♂ 6 ♀, Huangyangjie, Jinggangshan National Nature Reserve, 16. VIII. 2004, Wang XM leg. Hunan: 11 ♂, Mangshan National Nature Reserve, 3. X. 2004, Ren SX leg; 1 ♂, Shennonggu National Forest Park, 1000 m, 8. X. 2010, Wang XM leg; 2 ♂ 1 ♀, Shennonggu National Forest Park, 650 – 800 m, 9. X. 2010, Wang XM leg. Guangdong: 14 ♂ 1 ♀, Qinshuigu, Nanling National Nature Reserve, 30. IX. 2004, Wang XM leg; 2 ♂ 1 ♀, Protection station, Nanling National Nature Reserve, 3. X. 2004, Wang XM leg; 14 ♂ 9 ♀, Xidianzhan, Nanling National Nature Reserve, 1. X. 2004, Wang XM leg; 1 ♂, Huangniping, Shimentai National Nature Reserve, 7. X. 2004, Wang XM leg; 2 ♂, Shimen, Shimentai National Nature Reserve, 29. X. 2004, Wang XM leg; 2 ♂, Huangdong, Shimentai National Nature Reserve, 30. X. 2004, Wang XM leg; 4 ♂ 2 ♀, Dadong Mountains, Nanling National Nature Reserve, 19. VII. 2005, Wang XM leg; 8 ♂ 14 ♀, Qinshuigu, Nanling National Nature Reserve, 21. IX. 2005, Wang XM leg; 2 ♀, Tianjingshan National Forest Park, 22. IX. 2005, Wang XM leg. Guangxi: 2 ♂ 2 ♀, Hillside, Mao’ershan National Nature Reserve, 17. X. 2004, Wang XM leg; 1 ♀, Jiuniutang, Mao’ershan National Nature Reserve, 18. X. 2004, Wang XM leg; 8 ♂ 1 ♀, Shili Canyon, Mao’ershan National Nature Reserve, 19. X. 2004, Wang XM leg; 8 ♂ 3 ♀, GaoZhai, Mao’ershan National Nature Reserve, 20. X. 2004, Wang XM leg. Sichuan: 1 ♂ 1 ♀, Puge County, 2000 m, 15. IX. 2007, Wang XM leg. Guizhou: 1 ♀, Leigongshan National Nature Reserve, 9. VIII. 2006, Wang XM leg; 1 ♂, Datangwan, Leigongshan National Nature Reserve, 1100 m, 5. X. 2008, Wang XM leg; 1 ♂, Xiangshuiyan, Leigongshan National Nature Reserve, 980 m, 9. X. 2008, Wang XM leg; 1 ♂, Taojiang Town, Rongjiang County, 1000 m, 10 – 11. X. 2008, Wang XM leg; 1 ♂, Xiaodanjiang, Leigongshan National Nature Reserve, 12 – 13. X. 2008, Wang XM leg; 3 ♂ 5 ♀, Dongtang, Maolan National Nature Reserve, 730 m, 15 – 18. X. 2008, Wang XM leg; 1 ♂ 1 ♀, Weng’ang, Maolan National Nature Reserve, 750 m, 20 – 23. X. 2008, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F606FF98FF7BFA440E25FD70.taxon	distribution	Distribution. China (Zhejiang, Fujian, Jiangxi, Hubei, Hunan, Guangdong, Guangxi, Sichuan; Guizhou).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F608FF9BFF7BFCA10AC2FC95.taxon	description	Chinese name: 曲叶 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F608FF9BFF7BFCA10AC2FC95.taxon	diagnosis	Diagnosis. This species is similar to S. lancetapicalis (Pang & Gordon, 1986), but can be distinguished from the later by the penis guide apically pointed in ventral view (Fig. 40 h). In S. lancetapicalis, penis guide is apically blunt in ventral view (Fig. 43 h).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F608FF9BFF7BFCA10AC2FC95.taxon	description	Description. TL: 1.90 – 2.18 mm, TW: 1.40 – 1.60 mm, TH: 1.01 – 1.14 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 40 a – c). Head yellow with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow. Scutellar shield yellowish brown. Elytra black with yellow apices and brown humeral angles. Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 40 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis constricted with distinct membranous appendage (Figs 40 e – f). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 2 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 2 in ventral view, then gradually narrowing to pointed apex (Figs 40 g – h). Female genitalia: Unknown.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F608FF9BFF7BFCA10AC2FC95.taxon	materials_examined	Type material. Holotype: China: Hainan: ♂, Bawangling National Nature Reserve, 5. V. 2005, Wang XM leg (SCAU). Paratype (SCAU): China: Yunnan: 1 ♂, Botanic Garden, Jinghong City, 22. VIII. 2005, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F608FF9BFF7BFCA10AC2FC95.taxon	distribution	Distribution. China (Hainan, Yunnan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F608FF9BFF7BFCA10AC2FC95.taxon	etymology	Etymology. The specific epithet flexus derives from the Latin word “ flexus ” and refers to the flexural penis guide in lateral view.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60BFF9AFF7BFC840BB8FD98.taxon	description	(Figs 41, 69) Chinese name: Ûū ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60BFF9AFF7BFC840BB8FD98.taxon	diagnosis	Diagnosis. This species is most similar to S. lancetapicalis (Pang & Gordon, 1986), but can be distinguished from the later by the apically expanded penis (Fig. 41 e). In S. lancetapicalis, penis is not expanded at apex (Fig. 43 e).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60BFF9AFF7BFC840BB8FD98.taxon	description	Description. TL: 1.64 – 2.23 mm, TW: 1.17 – 1.64 mm, TH: 0.64 – 0.92 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 41 a – c). Head yellowish brown with interocular distance distinctly greater than eye width in frontal view. Pronotum yellowish brown. Scutellar shield brown to black. Elytra black with yellowish brown apices. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, slightly recurved toward the base of ventrite (Fig. 41 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis expanded with distinct membranous appendage (Figs 41 e – f). Penis guide with prominent keel at base in lateral view, then tapering towards apex; parameres about 2 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide in ventral view widest near the base, then gradually narrowing to apex (Figs 41 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60BFF9AFF7BFC840BB8FD98.taxon	materials_examined	Type material. Holotype: China: Guangxi: ♂, Hongqi Forest Farm, Shiwandashan National Nature Reserve, 26. VII. 2005, Wang XM leg (SCAU). Paratypes (SCAU): China: Guangxi: 4 ♂ 3 ♀, Same data as holotype; 5 ♂, Hillside, Mao’ershan National Nature Reserve, 17. X. 2004, Wang XM leg; 1 ♂, Jiuniutang, Mao’ershan National Nature Reserve, 18. X. 2004, Wang XM leg; 81 ♂ 74 ♀, Shili Canyon, Mao’ershan National Nature Reserve, 19. X. 2004, Wang XM leg; 9 ♂ 8 ♀, GaoZhai, Mao’ershan National Nature Reserve, 20. X. 2004, Wang XM leg; 13 ♂ 2 ♀, Pinglong Mountains, Shiwandashan National Nature Reserve, 6 – 8. XI. 2004, Wang XM leg; 22 ♂ 2 ♀, Hongqi Forest Farm, Shiwandashan National Nature Reserve, 9 – 11. XI. 2004, Wang XM leg; 11 ♂, Pinglong Mountains, Shiwandashan National Nature Reserve, 28. VII. 2005, Wang XM leg. Jiangxi: 6 ♀, Jingzhu Mountains, Jinggangshan National Nature Reserve, 22. IX. 2004, Wang XM leg; 2 ♂ 7 ♀, Longtan, Jinggangshan National Nature Reserve, 24. IX. 2004, Wang XM leg; 1 ♀, Hexilong, Jinggangshan National Nature Reserve, 26. IX. 2004, Wang XM leg. Hunan: 2 ♂ 3 ♀, Shennonggu National Forest Park, 1100 m, 7. X. 2010, Wang XM leg; 3 ♂ 5 ♀, Shennonggu National Forest Park, 650 – 800 m, 9. X. 2010, Wang XM leg. Guangdong: 2 ♂ 3 ♀, Xidianzhan, Nanling National Nature Reserve, 1. X. 2004, Wang XM leg; 1 ♂, Qinshuigu, Nanling National Nature Reserve, 30. IX. 2004, Wang XM leg; 1 ♂, Nankunshan 1 ♀, Xiaoqikong Town, Libo County, 18. VIII. 2006, Wang XM leg; 2 ♂ 1 ♀, Wangmo County, 21. VIII. 2006, Wang XM leg; 1 ♂ 1 ♀, Wudong Village – Maoping, Leigongshan National Nature Reserve, 1350 m, 6. X. 2008, Wang XM leg; 10 ♂ 10 ♀, Dongtang, Maolan National Nature Reserve, 730 m, 15 – 18. X. 2008, Wang XM leg; 1 ♀, Weng’ang, Maolan National Nature Reserve, 750 m, 20 – 23. X. 2008, Wang XM leg. Yunnan: 1 ♂, Mengla County, 23. VIII. 2005, Wang XM leg; 3 ♂ 10 ♀, Lafu Village, Menglian County, 1130 m, 7. V. 2008, Wang XM leg; 3 ♂, Banlao Town, Cangyuan County, 1100 m, 16. V. 2008, Wang XM leg; 8 ♂ 5 ♀, Dadugang Town, Jinghong City, 1050 m, 5. V. 2009, Wang XM leg; 1 ♂ 6 ♀, Jinuoshan Town, Jinghong City, 6. V. 2009, Wang XM leg; 1 ♂, Longmen Village, Shangyong Town, Mengla County, 1027 m, 9. V. 2009, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60BFF9AFF7BFC840BB8FD98.taxon	distribution	Distribution. China (Jiangxi, Hunan, Guangdong, Guangxi, Hainan, Guizhou, Yunnan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60BFF9AFF7BFC840BB8FD98.taxon	etymology	Etymology. The specific epithet sagittalis derives from the Latin word “ sagittalis ” and refers to its sagittal apex of penis.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60AFF9CFF7BF8E00AC7FEE4.taxon	description	(Figs 42, 71) Chinese name: ffi 瓣 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60AFF9CFF7BF8E00AC7FEE4.taxon	diagnosis	Diagnosis. This species is similar to S. bivalvis (Yu, 1997) in body colouration, but can be distinguished from the later by more robust penis guide, and parameres about 2 / 3 as long as penis guide (Figs 42 g – h) In S. bivalvis, penis guide is more slender, parameres are about 1 / 2 as long as penis guide (Figs 39 g – h).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60AFF9CFF7BF8E00AC7FEE4.taxon	description	Description. TL: 1.64 – 2.56 mm, TW: 1.38 – 1.97 mm, TH: 0.98 – 1.47 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 42 a – c). Head yellowish brown with interocular distance distinctly greater than eye width in frontal view. Pronotum yellowish brown. Scutellar shield black. Elytra black with yellowish brown apices. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 42 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis distinctly expanded with membranous appendage (Figs 42 e – f). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 2 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 3 in ventral view, apically rounded (Figs 42 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60AFF9CFF7BF8E00AC7FEE4.taxon	materials_examined	Type material. Holotype: China: Guangdong: ♂, Nanling National Nature Reserve, [24 ° 54 ' 59 " N 113 ° 2 ' 4 " E], 880 m, 17. XI. 2017, Huo LZ leg (SCAU). Paratypes (SCAU): China: Guangdong: 1 ♂, Qinshuigu, Nanling National Nature Reserve, 30. IX. 2004, Wang XM leg; 3 ♂ 1 ♀, Yangqiu Mountains, Shimentai National Nature Reserve, 6. X. 2004, Wang XM leg; 1 ♂, Wulangzhang, Shimentai National Nature Reserve, 8. X. 2004, Wang XM leg; 1 ♂, Nankunshan National Forest Park, 21. X. 2004, Wang XM leg; 8 ♂ 11 ♀, Huangdong, Shimentai National Nature Reserve, 30. X. 2004, Wang XM leg; 9 ♂ 10 ♀, Huangdong, Shimentai National Nature Reserve, 31. X. 2004, Wang XM leg; 4 ♂, Chebaling National Nature Reserve, 22 – 23. VIII. 2019, Wang XM leg. Fujian: 1 ♂, Guaxi Village, Fu’an City, 260 m, 21. IX. 2012, Huo LZ leg. Jiangxi: 1 ♂, Garden Spot, Xunwu County, 4. X. 1981, collector unknown. Hunan: 1 ♂, Mangshan National Nature Reserve, 3. X. 2004, Ren SX leg. Guangxi: 1 ♂, Hillside, Mao’ershan National Nature Reserve, 17. X. 2004, Wang XM leg; 3 ♂, Shili Canyon, Mao’ershan National Nature Reserve, 19. X. 2004, Wang XM leg. Chongqing: 1 ♂, Wuxi County, 2. VII. 1989, Ren SX leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60AFF9CFF7BF8E00AC7FEE4.taxon	distribution	Distribution. China (Fujian, Jiangxi, Hunan, Guangdong, Guangxi, Chongqing).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60AFF9CFF7BF8E00AC7FEE4.taxon	etymology	Etymology. The specific epithet robustus derives from the Latin word “ robustus ” and refers to the rubust penis guide and parameres.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60CFF9CFF7BFE550FA2F8FE.taxon	description	(Figs 43, 70) Chinese name: Ṙūḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60CFF9CFF7BFE550FA2F8FE.taxon	diagnosis	Diagnosis. This species is most similar to S. disselasmatus (Pang & Huang, 1986), but can be distinguished from the later by the blunt apex of penis guide in lateral view (Fig. 43 h). In S. disselasmatus, apex of penis guide is pointed in lateral view (Fig. 44 h).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60CFF9CFF7BFE550FA2F8FE.taxon	description	Description. TL: 1.58 – 2.31 mm, TW: 1.62 – 1.76 mm, TH: 0.69 – 1.12 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 43 a – c). Head yellow with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow. Scutellar shield yellowish brown. Elytra black with yellow apices and humeral angles. Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, slightly recurved toward the base of ventrite (Fig. 43 d). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis constricted with distinct membranous appendage (Figs 43 e – f). Penis guide with prominent keel at base in lateral view, then tapering towards apex; parameres about 3 / 5 as long as penis guide, with a tuft of long setae at apex; penis guide widest near the base in ventral view, apically rounded (Figs 43 g – h). Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60CFF9CFF7BFE550FA2F8FE.taxon	materials_examined	Material examined. China: Zhejiang: 1 ♂ 1 ♀, Medical Botanical Garden, Dapanshan National Nature Reserve, [120 ° 32 ' 38.51 " E 28 ° 58 ' 40.31 " N], 627 m, 30. IX. 2017, Tong JB leg. Anhui: 2 ♂ 1 ♀, Tangkou Town, Huangshan District, Huangshan City, 460 m, 12 – 13. IX. 2010, Wang XM leg; 1 ♂ 3 ♀, Huangshan Mountain Scenic Spot, 14 – 15. IX. 2010, Wang XM leg; 2 ♂, Hengdu Town, Shitai County, 230 m, 20. IX. 2010, Wang XM leg; 2 ♂ 8 ♀, Guniujiang National Nature Reserve, 24. IX. 2010, Wang XM leg; 4 ♀, Yaoluoping National Nature Reserve, 1000 m, 28 – 29. IX. 2010, Wang XM leg; 1 ♂, Chuanling, Yaoluoping National Nature Reserve, 800 m, 2. X. 2010, Wang XM leg; 10 ♂ 11 ♀, Jinzhai Tianma National Nature Reserve, 600 m, 3 – 5. X. 2010, Wang XM leg. Jiangxi: 2 ♂, Huangyangjie, Jinggangshan National Nature Reserve, 16. VIII. 2004, Wang XM leg; 2 ♂, Fengshan National Forest Park, 17. VIII. 2004, Wang XM leg. Henan: 3 ♂, Jigongshan National Nature Reserve, 10. VII. 1997, Peng ZQ leg; 4 ♂ 6 ♀, Jigongshan National Nature Reserve, 697 m, 4. VII. 2009, Wang XM leg; 1 ♂, Jigongshan National Nature Reserve, 1500 m, 4. VII. 2009, Wang XM leg; 4 ♂ 4 ♀, Purdom Forest Park, Jigongshan National Nature Reserve, 216 m, 5. VII. 2009, Wang XM leg; 2 ♀, Baiyunshan National Nature Reserve, 1375 m, 14. VII. 2009, Wang XM leg. Hubei: 1 ♂, Lulin Town, Jingshan City, 169 m, 10. VII. 2007, Wang XM leg. Hunan: 1 ♂ 1 ♀, Shennonggu National Forest Park, 1000 m, 8. X. 2010, Wang XM leg; 1 ♂, Shennonggu National Forest Park, 650 – 800 m, 9. X. 2010, Wang XM leg. Guangdong: 1 ♂, Chebaling National Nature Reserve, 3. XI. 1989, Yu GY leg; 3 ♂, Wulangzhang, Shimentai National Nature Reserve, 8. X. 2004, Wang XM leg; 1 ♂, Huangdong, Shimentai National Nature Reserve, 30. X. 2004, Wang XM leg; 1 ♂ 1 ♀, Chebaling National Nature Reserve, 22 – 23. VIII. 2019, Wang XM leg. Guizhou: 1 ♂, Dongtang, Maolan National Nature Reserve, 730 m, 15 – 18. X. 2008, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60CFF9CFF7BFE550FA2F8FE.taxon	distribution	Distribution. China (Zhejiang, Anhui, Jiangxi, Henan, Hubei, Hunan, Guangdong, Guizhou).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60FFF9EFF7BFA8508F6FD98.taxon	description	(Figs 44, 67) Chinese name: R Ṙḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60FFF9EFF7BFA8508F6FD98.taxon	diagnosis	Diagnosis. This species is closely similar to S. sagittalis sp. nov., but can be distinguished from the later by the black scutellar shield and the apex of penis with distinct membranous appendage on inner side (Figs 44 a, 44 g). In S. sagittalis, Scutellar shield is yellowish brown and the apex of penis is conspicuously swollen with membranous appendage on both sides (Figs 41 a, 41 f).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60FFF9EFF7BFA8508F6FD98.taxon	description	Description. TL: 1.68 – 2.26 mm, TW: 1.23 – 1.66 mm, TH: 0.68 – 0.86 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 44 a – c). Head yellow with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow. Scutellar shield black. Elytra black with yellow apices. Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, slightly recurved toward the base of ventrite (Fig. 44 e). Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis with distinct membranous appendage (Figs 44 f – g). Penis guide with prominent keel at base in lateral view, then tapering towards apex; parameres about 2 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, then gradually narrowing to blunt apex (Figs 44 g – h). Female genitalia: Unknow.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60FFF9EFF7BFA8508F6FD98.taxon	materials_examined	Material examined. Holotype: China: Fujian: ♂, Nanjing County, 7. III. 1980, Liu SJ leg (SCAU). Other material (SCAU). China: Guangdong: 1 ♂, Shixing County, 8. VII. 1989, Pang XF leg; 1 ♂, Yangqiu Mountains, Shimentai National Nature Reserve, 6. X. 2004, Wang XM leg; 1 ♂, Shimen, Shimentai National Nature Reserve, 29. X. 2004, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60FFF9EFF7BFA8508F6FD98.taxon	distribution	Distribution. China (Fujian, Guangdong).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60FFF9EFF7BFA8508F6FD98.taxon	discussion	Remarks. Yu et al. (1993) reported S. disselasmatus from Chebaling National Nature Reserve. Compared with the holotype, specimen examined by Yu et al. (1993) has penis guide widest near the base and not expanded penis apex, hence we considered it as a misidentification of S. lancetapicalis.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F60EFF9EFF7BF9490843F80C.taxon	description	(Fig. 67) Chinese name: 张氏 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F671FFE1FF7BFC4C0F3BFBD9.taxon	discussion	The Sasajiscymnus atypicus species group is characterized by the long penis with swollen outer arm, apex of penis with membranous area, penis guide elongate and robust, and parameres nearly as long as penis guide.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F671FFE1FF7BFC4C0F3BFBD9.taxon	synonymic_list	The only speices of atypicus group is Sasajiscymnus atypicus Tong & Wang sp. nov.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F671FFE1FF7BFF750A49FCDD.taxon	description	(Fig. 69) Chinese name: R Ṅḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F671FFE1FF7BFF750A49FCDD.taxon	materials_examined	Material examined. None.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F671FFE1FF7BFF750A49FCDD.taxon	distribution	Distribution. China (Taiwan); Russia; Japan; South Korea.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F671FFE1FF7BFF750A49FCDD.taxon	discussion	Remarks. The descriptions of male genitalia of this species vary in different works (Miyatake, 1958; Sasaji, 1971; Park & Yoon, 1994; Kuzuetsov, 1997; Ren et al., 2009) hence it is possible that cryptic species within this species exist. Ren et al. (2009) presented photos of S. hareja with remarkable emargination and distinct membranous appendage at the apex of penis, which differ from all the other descriptions. Based on the examination of that specimen and the revised diagnostic characters, we consider it as a new species, S. variabilis sp. nov., which is described in this study.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F671FFE0FF7BFB4809EEFD4C.taxon	description	(Figs 1 i, 3 a, 3 e, 45, 74) Chinese name: Ǿ 特 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F671FFE0FF7BFB4809EEFD4C.taxon	diagnosis	Diagnosis. This is a very distinctive species of Sasajiscymnus characterized by the yellowish brown head bearing a large black spot, long penis with swollen outer arm and fin-shaped apex, and parameres nearly as long as penis guide (Figs 45 c – h).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F671FFE0FF7BFB4809EEFD4C.taxon	description	Description. TL: 1.76 – 2.44 mm, TW: 1.34 – 1.86 mm, TH: 0.79 – 1.08 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 45 a – c). Head yellowish brown with a large black spot, interocular distance distinctly greater than eye width in frontal view. Pronotum yellowish brown with a black spot at base. Scutellar shield black. Elytra black with yellowish brown apices. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 45 d). Male genitalia: Penis long, penis capsule with swollen outer arm and short inner one, apex of penis fin-shaped (Figs 45 e – f). Penis guide long in lateral view, gradually narrowing to pointed apex; parameres long and slender, nearly as long as penis guide, covered with long setae at apex and inner sides; penis guide stout in ventral view, widest at basal 3 / 4, distinctly narrowed at base, with widest part about 1.42 × of basal width (Figs 45 g – h). Female genitalia: Coxites board; styli strongly reduced and hardly visible; spermatheca worm-like, without clear ramus or nodulus (Figs 3 a, 3 e).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F671FFE0FF7BFB4809EEFD4C.taxon	materials_examined	Type material. Holotype: China: Sichuan: ♂, Fengtongzhai National Nature Reserve, 1560 m, 2 – 3. X. 2007, Wang XM leg (SCAU). Paratypes (SCAU): China: Sichuan: 3 ♂ 2 ♀, Same data as holotype; 2 ♂ 1 ♀, Liziping Reserve, 465 m, 16. IX. 2010, Wang XM leg; 3 ♂ 2 ♀, Zhengjiang Tianmushan National Nature Reserve, 450 – 1000 m, 18. IX. 2010, Wang XM leg. Anhui: 2 ♀, Yaoluoping National Nature Reserve, 600 – 800 m, 30. IX. 2010, Wang XM leg; 3 ♂ 3 ♀, Jinzhai Tianma National Nature Reserve, 600 m, 3 – 5. X. 2010, Wang XM leg. Hubei: 2 ♂, Hongping, Shennongjia National Nature Reserve, 19. VII. 1997, Peng ZQ leg; 1 ♂ 1 ♀, Banqiao, Shennongjia National Nature Reserve, 1170 m, 21 – 24. VII. 2007, Wang XM leg; 1 ♂ 1 ♀, Pingzhan, Shennongjia National Nature Reserve, 1600 m, 27. VII. 2007, Wang XM leg. Hunan: 4 ♂ 2 ♀, Shennonggu National Forest Park, 1100 m, 7. X. 2010, Wang XM leg; 5 ♂ 4 ♀, Shennonggu National Forest Park, 1000 m, 8. X. 2010, Wang XM leg. Guizhou: 1 ♂, Leigongshan National Nature Reserve, 30. VII. 1997, Peng ZQ leg; 1 ♂ 1 ♀, Datangwan, Leigongshan National Nature Reserve, 1100 m, 5. X. 2008, Wang XM leg; 2 ♂, Wudong Village – Maoping, Leigongshan National Nature Reserve, 1350 m, 6. X. 2008, Wang XM leg; 1 ♀, Wudong Village – Fangxiang, Leigongshan National Nature Reserve, 7. X. 2008, Wang XM leg; 4 ♂ 2 ♀, Lianhuaping, Leigongshan National Nature Reserve, 8. X. 2008, Wang XM leg; 1 ♂, Taojiang Town, Rongjiang County, 1000 m, 10 – 11. X. 2008, Wang XM leg. Shaanxi: 10 ♂ 10 ♀, Daguping Village, Foping National Nature Reserve, 1170 m, 21. VII. 2009, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F671FFE0FF7BFB4809EEFD4C.taxon	distribution	Distribution. China (Zhejiang, Anhui, Hubei, Hunan, Sichuan, Guizhou, Shaanxi).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F671FFE0FF7BFB4809EEFD4C.taxon	etymology	Etymology. The specific epithet atypicus derives from the Latin word “ atypicus ” and refers to the atypical head colouration and structures of the male genitalia.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F673FFE3FF7BFF750EDCFD25.taxon	discussion	The Sasajiscymnus amplus species group is characterized by the large body size, slender and strongly curved penis with inconspicuous outer arm, elongate penis guide which is arcuate in lateral view and extremely short and applanate parameres. 3 species of amplus group are recorded from China, including 2 new species.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F673FFE2FF7BFD150A7DFC49.taxon	description	(Figs 1 c, 3 j, 3 n, 46, 75) Chinese name: Üū ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F673FFE2FF7BFD150A7DFC49.taxon	diagnosis	Diagnosis. This species is similar to S. amplus (Yang & Wu, 1972), but can be distinguished from the later by its entirely yellowish brown elytra (Figs 46 a – b). In S. amplus, elytra are black with yellowish brown apices (Figs 47 a – b).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F673FFE2FF7BFD150A7DFC49.taxon	description	Description. TL: 2.56 – 2.87 mm, TW: 1.99 – 2.29 mm, TH: 1.20 – 1.42 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 46 a – c). Body uniformly yellowish brown. Interocular distance distinctly greater than eye width in frontal view. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 46 d). Male genitalia: Penis long and slender, penis capsule with inconspicuous outer arm and long inner one, apex of penis twisted and forming a knot (Figs 46 e – f). Penis guide long and slender in lateral view, arcuate and gradually narrowing to pointed apex; parameres about 1 / 4 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 2 / 3 in ventral view, then tapering towards blunt apex (Figs 46 g – h). Female genitalia: Coxites shaped board plates; styli absent; spermatheca cylinder-shaped, with basal ramus and nodulus (Figs 3 j, 3 n).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F673FFE2FF7BFD150A7DFC49.taxon	materials_examined	Type material. Holotype: China: Xizang: ♂, Shangchayu Town – Xiachayu Town, Chayu County, 1650 m, 27. IX. 2011, Huo LZ leg (SCAU). Paratypes (SCAU): China: Xizang: 6 ♂ 6 ♀, Same data as holotype; 1 ♂, Cibagou National Nature Reserve, 2000 m, 13. X. 2007, Wang XM leg; 1 ♂, Xiachayu Town, Chaiyu County, 1800 m, 14. X. 2007, Wang XM leg. Anhui: 1 ♂, Guniujiang National Nature Reserve, 24. IX. 2010, Wang XM leg. Fujian: 1 ♂, Tongmu Village, Wuyishan National Nature Reserve, 750 m, 17. IX. 2012, Huo LZ leg. Yunnan: 1 ♂, Aini Village, Jinghong City, 11. V. 2009, Wang XM leg; 1 ♀, Dajian Mountains, Daweishan National Nature Reserve, 2100 m, 22 – 23. V. 2009, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F673FFE2FF7BFD150A7DFC49.taxon	distribution	Distribution. China (Anhui, Fujian, Yunnan, Xizang).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F673FFE2FF7BFD150A7DFC49.taxon	etymology	Etymology. The specific epithet intricatus derives from the Latin word “ intricatus ” and refers to the apically intricate penis.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F672FFE2FF7BFBF8095EF874.taxon	description	(Figs 1 a, 3 i, 3 m, 47, 75) Chinese name: k ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F672FFE2FF7BFBF8095EF874.taxon	diagnosis	Diagnosis. This species closely resembles S. pseudoamplus sp. nov., but can be distinguished from the later by its penis guide widest at base in ventral view (Fig. 47 g). In S. pseudoamplus, penis guide is approximately parallel sided for basal 1 / 2 in ventral view (Fig. 48 g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F672FFE2FF7BFBF8095EF874.taxon	description	Description. TL: 2.85 – 3.34 mm, TW: 2.14 – 2.53 mm, TH: 1.23 – 1.52 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 47 a – c). Head yellowish brown, with interocular distance distinctly greater than eye width in frontal view. Pronotum yellowish brown with a large black spot at base. Scutellar shield black. Elytra black with yellowish brown apices. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 47 d). Male genitalia: Penis long and slender, penis capsule with inconspicuous outer arm and long inner one, apex of penis pointed (Figs 47 e – f). Penis guide long and slender in lateral view, arcuate and gradually narrowing to pointed apex; parameres about 1 / 8 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, tapering towards blunt apex (Figs 47 g – h). Female genitalia: Coxites shaped board plates; styli absent; spermatheca worm-like, with basal ramus and nodulus (Figs 3 i, 3 m).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F672FFE2FF7BFBF8095EF874.taxon	materials_examined	Material examined. Holotype: China: Taiwan: ♀, Puli Town, Nantou County, 21. VII. 1971, Yang CT leg (NCHU). Paratype: China: Taiwan: 1 ♀, Same data as holotype (NCHU). Other material (SCAU). China: Taiwan: 1 ♀, Aowanda, Jenai Town, Nantou County, 1280 m, 21. X. 2012, Wang XM leg; 4 ♂, Ali Mountains, Chiayi County, 1800 m, 22. X. 2012, Wang XM leg; 4 ♂ 7 ♀, Tengjhih, Kaohsiung City, 370 – 1450 m, 25. X. 2012, Wang XM leg; 1 ♂ 4 ♀, Maolin District, Kaohsiung City, 365 – 515 m, 26. X. 2012, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F672FFE2FF7BFBF8095EF874.taxon	materials_examined	Distribution. China (Taiwan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F675FFE7FF7BFA670EDCFE9C.taxon	description	(Figs 48, 75) Chinese name: fflk ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F675FFE7FF7BFA670EDCFE9C.taxon	diagnosis	Diagnosis. This species closely resembles S. amplus (Yang & Wu, 1972), but can be distinguished from the later by its penis guide approximately parallel sided for basal 1 / 2 in ventral view (Fig. 48 g). In S. amplus, penis guide is widest at base in ventral view (Fig. 47 g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F675FFE7FF7BFA670EDCFE9C.taxon	description	Description. TL: 2.60 – 3.56 mm, TW: 1.95 – 2.70 mm, TH: 1.27 – 1.48 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 48 a – c). Head yellowish brown, with interocular distance distinctly greater than eye width in frontal view. Pronotum yellowish brown with a large black spot at base. Scutellar shield black. Elytra black with yellowish brown apices. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 48 d). Male genitalia: Penis long and slender, penis capsule with inconspicuous outer arm and long inner one, apex of penis pointed (Figs 48 e – f). Penis guide long and slender in lateral view, arcuate and gradually narrowing to pointed apex; parameres about 1 / 8 as long as penis guide, with a tuft of long setae at apex; Penis guide approximately parallel sided for basal 1 / 2 in ventral view, then gradually narrowing to blunt apex (Figs 48 g – h). Female genitalia: Coxites shaped board plates; styli absent; spermatheca worm-like, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F675FFE7FF7BFA670EDCFE9C.taxon	materials_examined	Type material. Holotype: China: Guizhou: ♂, Luodian County, 20. VIII. 2006, Wang XM leg (SCAU). Paratypes (SCAU): China: Guizhou: 4 ♂ 1 ♀, Same data as holotype; 4 ♂ 1 ♀, Huangguoshu Waterfalls Scenic Area, 15. VIII. 1990, Tian MY leg; 1 ♀, Weng’ang, Maolan National Nature Reserve, 800 m, 15. VIII. 2006, Wang XM leg; 1 ♂ 2 ♀, Wuyan Bridge, Maolan National Nature Reserve, 17. VIII. 2006, Wang XM leg; 7 ♂ 19 ♀, Luodian County, 19. VIII. 2006, Wang XM leg; 19 ♂ 9 ♀, Xianheping Nature Reserve, 14 – 15. IX. 2006, Wang XM leg; 1 ♂ 3 ♀, Dongtang, Maolan National Nature Reserve, 730 m, 15 – 18. X. 2008, Wang XM leg; 1 ♀, Sancha River, Maolan National Nature Reserve, 750 m, 19. X. 2008, Wang XM leg. Hainan: 1 ♀, Baihua Mountains, Qiongzhong County, VIII. 1995, Peng ZQ leg; 1 ♀, Wufenchang, Limushan National Forest Park, IX. 1995, Peng ZQ leg; 1 ♀, Qixianling Hot Springs National Forest Park, 7. V. 2005, Wang XM leg; 1 ♂, Diaoluoshan National Nature Reserve, 8. V. 2005, Wang XM leg; 1 ♂, Limushan National Forest Park, 23. VII. 2006, Wang XM leg; 1 ♀, Yinggezui, Yinggeling National Nature Reserve, 16. IV. 2019, Wang XM leg. Yunnan: 1 ♂, Malipo County, 17. VIII. 2005, Wang XM leg; 5 ♂ 2 ♀, Fengqing County, 27. VIII. 2005, Wang XM leg; 1 ♀, Mang City, 28. VIII. 2005, Wang XM leg; 1 ♀, Mangbing, Yingjiang City, 29. VIII. 2005, Wang XM leg; 1 ♂, Gaoligongshan National Nature Reserve, 19. IX. 2006, Wang XM leg; 5 ♀, Dadugang Town, Jinghong City, 950 m, 26. IV. 2008, Wang XM leg; 4 ♂ 4 ♀, Menglun Town, Mengla County, 1160 m, 29. IV. 2008, Wang XM leg; 2 ♀, Banlao Town, Cangyuan County, 1100 m, 16. V. 2008, Wang XM leg; 10 ♂ 2 ♀, Caiyanghe National Forest Park, 4. V. 2009, Wang XM leg; 1 ♂, Dadugang Town, Jinghong City, 1050 m, 5. V. 2009, Wang XM leg; 1 ♀, Jinuoshan Town, Jinghong City, 6. V. 2009, Wang XM leg; 2 ♂ 2 ♀, Aini Village, Jinghong City, 11. V. 2009, Wang XM leg; 2 ♂ 1 ♀, Meng’a Town, Menghai County, 1170 m, 12. V. 2009, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F675FFE7FF7BFA670EDCFE9C.taxon	distribution	Distribution. China (Hainan, Guizhou, Yunnan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F675FFE7FF7BFA670EDCFE9C.taxon	etymology	Etymology. The specific epithet pseudoamplus refers to the similarity between the new species and S. amplus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F677FFE7FF7BFE8D0EDCFA7B.taxon	discussion	The Sasajiscymnus kurohime species group is characterized by the black elytra, slender penis without outer arm, elongate penis guide, apex of penis simple without appendage, and extremely thin parameres. 14 species of kurohime group are recorded from China, including 7 new species and 1 new recorded species.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F677FFE6FF7BF9AF09FBFDAF.taxon	description	(Figs 49, 77) Chinese name: Ñ Ờḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F677FFE6FF7BF9AF09FBFDAF.taxon	diagnosis	Diagnosis. This is a distinctive species of Chinese Sasajiscymnus by the entire dorsum deeply black with only two small, yellow spots on the elytra (Fig. 49 a).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F677FFE6FF7BF9AF09FBFDAF.taxon	description	Description. TL: 1.35 – 1.62 mm, TW: 0.99 – 1.26 mm, TH: 0.52 – 0.66 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 49 a – c). Head black with antennae and mouthparts yellow, with interocular distance distinctly closer than eye width in frontal view. Pronotum black with anterior margin narrowly yellowish brown. Scutellar shield black. Elytra black with brown apices, and each elytron with a yellow spot on medially but not touching suture. Ventral surfaces black. Legs yellow to yellowish brown. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 49 d). Male genitalia: Penis slender, penis capsule with long inner arm but without outer one, apex of penis pointed (Fig. 49 e). Penis guide long and slender in lateral view, approximately parallel sided for basal 3 / 4, then gradually narrowing to pointed apex; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, tapering towards blunt apex (Figs 49 f – g). Female genitalia: Coxites shortened; styli absent; spermatheca worm-like, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F677FFE6FF7BF9AF09FBFDAF.taxon	materials_examined	Type material. Holotype: China: Yunnan: ♂, Yexianggu, Xishuangbanna National Nature Reserve, 780 m, 23. IV. 2019, Deng LC leg (SCAU). Paratypes (SCAU): China: Yunnan: 1 ♂ 1 ♀, Mengyang Town, Jinghong City, 27. IV. 2008, Wang XM leg; 1 ♂ 4 ♀, Dadugang Town, Jinghong City, 1050 m, 5. V. 2009, Wang XM leg; 3 ♀, Yaoqv Town, Mengla County, 700 m, 7 – 8. V. 2009, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F677FFE6FF7BF9AF09FBFDAF.taxon	distribution	Distribution. China (Yunnan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F677FFE6FF7BF9AF09FBFDAF.taxon	etymology	Etymology. The specific epithet flavostictus derives from the Latin words “ flavus ” (= yellow) and “ stictus ” (= spot), refers to its two yellow spots on the elytra.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F676FFE9FF7BF91E0AD7FD70.taxon	description	(Figs 50, 76) Chinese name: dz 叶 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F676FFE9FF7BF91E0AD7FD70.taxon	diagnosis	Diagnosis. This species is similar to S. applanatus sp. nov. in body size and colouration, but can be distinguished from the later by parameres about 3 / 4 as long as penis guide (Fig. 50 f). In S. applanatus, parameres are about 1 / 3 as long as penis guide (Fig. 60 f).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F676FFE9FF7BF91E0AD7FD70.taxon	description	Description. TL: 1.36 – 1.58 mm, TW: 0.99 – 1.18 mm, TH: 0.52 – 0.67 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 50 a – c). Body uniformly black, except antennae, maxillary palpi and legs yellow to yellowish brown. Interocular distance slightly greater than eye width in frontal view. Abdominal postcoxal lines incomplete, slightly recurved toward the base of ventrite (Fig. 50 d). Male genitalia: Penis slender, penis capsule with long inner arm but without outer one, apex of penis pointed (Fig. 50 e). Penis guide approximately parallel sided for basal 1 / 2 in lateral view, then tapering towards pointed apex; parameres about 3 / 4 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 2 in ventral view, then slightly narrowing to blunt apex (Figs 50 f – g). Female genitalia: Coxites shortened; styli absent; spermatheca worm-like, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F676FFE9FF7BF91E0AD7FD70.taxon	materials_examined	Type material. Holotype: China: Yunnan: ♂, Maku Village, Gongshan County, 1600 m, 1. VIII. 2010, Wang XM leg (SCAU). Paratypes (SCAU): China: Yunnan: 3 ♀, Same data as holotype; 2 ♂, Heiwadi, Gongshan County, 2020 m, 19. VII. 2010, Wang XM leg; 1 ♂ 8 ♀, Longyuan Village, Gongshan County, 1690 m, 24 – 25. VII. 2010, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F676FFE9FF7BF91E0AD7FD70.taxon	distribution	Distribution. China (Yunnan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F676FFE9FF7BF91E0AD7FD70.taxon	etymology	Etymology. The specific epithet falcatus derives from the Latin word “ falcatus ” and refers to the sickle-shaped tegmen in lateral view.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F678FFEBFF7BFF7508B6FD4C.taxon	description	Chinese name: Ẇ 状 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F678FFEBFF7BFF7508B6FD4C.taxon	diagnosis	Diagnosis. This species is similar to S. kuriharai Kitano, 2012 in structures of the male genitalia, but can be distinguished from the later by its pronotum yellowish brown, sometimes with a black spot at base (Figs 51 a – b). In S. kuriharai, pronotum is black with anterior margin narrowly yellowish brown (Figs 52 a – b).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F678FFEBFF7BFF7508B6FD4C.taxon	description	Description. TL: 1.56 – 2.09 mm, TW: 1.17 – 1.54 mm, TH: 0.71 – 0.93 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 51 a – c). Head yellowish brown, with interocular distance distinctly closer than eye width in frontal view. Pronotum yellowish brown, sometimes with a black spot at base. Scutellar shield black. Elytra black with brown apices. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 51 d). Male genitalia: Penis slender, penis capsule with long inner arm but without outer one, apex of penis pointed (Figs 51 e – f). Penis guide apically hook-shaped in lateral view; parameres about 1 / 4 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 5 / 6 in ventral view, then gradually narrowing to blunt apex (Figs 51 g – h). Female genitalia: Coxites shortened; styli absent; spermatheca worm-like, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F678FFEBFF7BFF7508B6FD4C.taxon	materials_examined	Type material. Holotype: China: Guangdong: ♂, Chebaling National Nature Reserve, 23. IV. 1991, Yu GY leg (SCAU). Other material (SCAU). China: Guangdong: 2 ♂, Nankunshan National Forest Park, 21. X. 2004, Wang XM leg; 1 ♂, Qinshuigu, Nanling National Nature Reserve, 21. IX. 2005, Wang XM leg. Hainan: 1 ♂, Diaoluoshan National Nature Reserve, 8. V. 2005, Wang XM leg; 2 ♂, Diaoluoshan National Nature Reserve, 25. VII. 2006, Wang XM leg. Yunnan: 1 ♂, Menglun Town, Mengla County, 21. VIII. 2005, Wang XM leg; 2 ♂, Botanic Garden, Jinghong City, 22. VIII. 2005, Wang XM leg; 1 ♂ 1 ♀, Menghan Town, Jinghong City, 24. VIII. 2005, Wang XM leg; 4 ♂ 1 ♀, National highway 213 route, Mengla County, 12 – 13. X. 2006, Wang XM leg; 1 ♂ 1 ♀, Dadugang Town, Jinghong City, 1050 m, 5. V. 2009, Wang XM leg; 1 ♂, Yaoqv Town, Mengla County, 700 m, 7 – 8. V. 2009, Wang XM leg; 1 ♂, Nanping, Mengman Town, Mengla County, 745 m, 10. V. 2009, Wang XM leg; 4 ♂ 4 ♀, Meng’a Town, Menghai County, 1170 m, 12. V. 2009, Wang XM leg; 1 ♂, Bapo Village, Gongshan County, 1400 m, 28. VII. 2010, Wang XM leg; 1 ♂, Qinglandan, Maku Village, Gongshan County, 4. VIII. 2010, Wang XM leg; 3 ♂, Aini Village, Jinghong City, 700 m, 13. VIII. 2013, Chen XS leg; 1 ♂, Yexianggu, Xishuangbanna National Nature Reserve, 780 m, 23. IV. 2019, Deng LC leg. Xizang: 1 ♂, Hanmi, Muotuo County, 800 – 2100 m, 5 – 8. X. 2011, Huo LZ leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F678FFEBFF7BFF7508B6FD4C.taxon	distribution	Distribution. China (Guangdong, Hainan, Yunnan, Xizang).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F678FFEBFF7BFF7508B6FD4C.taxon	discussion	Remarks. Several individuals have complete abdominal postcoxal lines.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67BFFEDFF7BFCFD083BFE50.taxon	description	(Figs 1 j, 52, 81) Chinese name: B Ẇḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67BFFEDFF7BFCFD083BFE50.taxon	diagnosis	Diagnosis. This species is similar to S. hamatus (Yu & Pang, 1993) in structures of the male genitalia, but can be distinguished from the later by its pronotum black with anterior margin narrowly yellowish brown (Figs 52 a – b). In S. hamatus, pronotum is yellowish brown, sometimes with a black spot at base (Figs 51 a – b).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67BFFEDFF7BFCFD083BFE50.taxon	description	Description. TL: 1.17 – 1.92 mm, TW: 0.94 – 1.47 mm, TH: 0.58 – 0.95 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 52 a – c). Head yellow to black, interocular distance distinctly closer than eye width in frontal view. Pronotum black with anterior margin yellowish brown. Scutellar shield black. Elytra black with yellowish brown apices. Ventral surfaces black. Legs yellow to yellowish brown. Abdominal postcoxal lines almost complete (Fig. 52 d). Male genitalia: Penis slender, penis capsule with long inner arm but without outer one, apex of penis pointed (Fig. 52 e). Penis guide apically hook-shaped in lateral view; parameres about 1 / 5 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 5 / 6 in ventral view, then gradually narrowing to blunt apex (Figs 52 f – g). Female genitalia: Coxites shortened; styli absent; spermatheca worm-like, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67BFFEDFF7BFCFD083BFE50.taxon	materials_examined	Material examined. China: Guangdong: 3 ♂ 1 ♀, South China Agricultural University, Guangzhou City, 26. IV. 1989, Ren SX leg; 1 ♂, South China Agricultural University, Guangzhou City, 28. IV. 1989, Ren SX leg; 10 ♂ 5 ♀, Dinghushan National Nature Reserve, 16. V. 1989, Ren SX leg; 4 ♂ 1 ♀, Dinghushan National Nature Reserve, 17. V. 1989, Ren SX leg; 2 ♂, Qixingyan Scenic Spot, Zhaoqing City, 18. V. 1989, Ren SX leg; 1 ♂, Shipai, Guangzhou City, 30. VII. 1989, Ren SX leg; 7 ♂ 7 ♀, Hengshitang, Shimentai National Nature Reserve, 5. X. 2004, Wang XM leg; 1 ♂ 1 ♀, Yangqiu Mountains, Shimentai National Nature Reserve, 6. X. 2004, Wang XM leg; 8 ♂, Huangniping, Shimentai National Nature Reserve, 7. X. 2004, Wang XM leg; 4 ♂, Wulangzhang, Shimentai National Nature Reserve, 8. X. 2004, Wang XM leg; 1 ♂, Huangdong, Shimentai National Nature Reserve, 30. X. 2004, Wang XM leg. Guangxi: 1 ♂, Xixiangtang District, Nanning City, 24. V. 1982, Pu TS leg; 1 ♂, Gongguang Town, Hepu County, 30. VII. 1985, Pang XF leg; 1 ♂, Longzhou County, 1. VIII. 1985, Pu TS leg; 1 ♂, Hepu County, 30. V. 1986, Pu TS leg; 4 ♀, Shili Canyon, Mao’ershan National Nature Reserve, 19. X. 2004, Wang XM leg; 8 ♂ 8 ♀, Hongqi Forest Farm, Shiwandashan National Nature Reserve, 9 – 11. XI. 2004, Wang XM leg; 1 ♂, Hongqi Forest Farm, Shiwandashan National Nature Reserve, 26. VII. 2005, Wang XM leg; 1 ♀, Pinglong Mountains, Shiwandashan National Nature Reserve, 28. VII. 2005, Wang XM leg; 2 ♀, Baidu Town, Napo County, 4. VIII. 2005, Wang XM leg; 1 ♂, Pinglong Mountains, Shiwandashan National Nature Reserve, 6. XI. 2005, Wang XM leg. Hainan: 1 ♂, Limushan National Forest Park, 14. VII. 2002, Peng ZQ leg; 1 ♂, Qixianling Hot Springs National Forest Park, 7. V. 2005, Wang XM leg; 1 ♂, Diaoluoshan National Nature Reserve, 8. V. 2005, Wang XM leg; 1 ♂, Limushan National Forest Park, 14. VII. 2006, Peng ZQ leg; 1 ♀, Tianchi, Jianfengling National Nature Reserve, 18. VII. 2006, Wang XM leg; 2 ♀, Bawangling National Nature Reserve, 20. VII. 2006, Wang XM leg; 1 ♂, Wuzhishan National Nature Reserve, 24. VII. 2006, Wang XM leg; 1 ♂, Diaoluoshan National Nature Reserve, 26. VII. 2005, Wang XM leg; 1 ♂, Nankai, Yinggeling National Nature Reserve, [109 ° 23 ' 24.70 " E 19 ° 3 ' 2.14 " N], 300 m, 16. IV. 2019, Wang XM leg. Sichuan: 2 ♂, Puge County, 2000 m, 15. IX. 2007, Wang XM leg; 1 ♂, Huanyanding – Qingyinge, E’mei Mountains, E’meishan City, [103 ° 20 ' 43 " E 29 ° 33 ' 40 " N], 1830 m, Zhang XN leg. Guizhou: 4 ♂ 1 ♀, Rongjiang County – Leishan County, 9. VIII. 2006, Wang XM leg; 5 ♂ 5 ♀, Weng’ang, Maolan National Nature Reserve, 800 m, 15. VIII. 2006, Wang XM leg; 7 ♂ 2 ♀, Xiaoqikong Town, Libo County, 18. VIII. 2006, Wang XM leg; 1 ♂ 1 ♀, Luodian County, 20. VIII. 2006, Wang XM leg; 1 ♂ 1 ♀, Xianheping Nature Reserve, 14 – 15. IX. 2006, Wang XM leg; 1 ♀, Lianhuaping, Leigongshan National Nature Reserve, 8. X. 2008, Wang XM leg; 19 ♂ 21 ♀, Dongtang, Maolan National Nature Reserve, 730 m, 15 – 18. X. 2008, Wang XM leg; 6 ♂ 4 ♀, Weng’ang, Maolan National Nature Reserve, 750 m, 20 – 23. X. 2008, Wang XM leg. Yunnan: 2 ♂, Nansan, Zhenkang County, 28. X. 2000, Peng ZQ leg; 3 ♂, Funing County, 15. VIII. 2005, Wang XM leg; 3 ♂, Malipo County, 17. VIII. 2005, Wang XM leg; 2 ♂, Maguan County, 18. VIII. 2005, Wang XM leg; 1 ♀, Dadugang Town, Jinghong City, 950 m, 26. IV. 2008, Wang XM leg; 2 ♀, Dadugang Town, Jinghong City, 1050 m, 5. V. 2009, Wang XM leg; 1 ♂, Bulangshan Town, Menghai County, 1188 m, 12. VIII. 2013, Chen XS leg. Hongkong: 1 ♂, Pat Sin Leng Country Park, 91 m, 13. V. 2018, Wang XM leg. Taiwan: 2 ♂ 13 ♀, Ali Mountains, Chiayi County, 1800 m, 22. X. 2012, Wang XM leg; 3 ♂ 6 ♀, Dabang Village, Alishan Mountains, Chiayi County, 960 – 1600 m, 24. X. 2012, Wang XM leg; 1 ♂ 1 ♀, Tengjhih, Kaohsiung City, 370 – 1450 m, 25. X. 2012, Wang XM leg; 1 ♂, Maolin District, Kaohsiung City, 365 – 515 m, 26. X. 2012, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67BFFEDFF7BFCFD083BFE50.taxon	distribution	Distribution. China (Guangdong, Guangxi, Hainan, Sichuan, Guizhou, Yunnan, Hongkong, Taiwan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67BFFEDFF7BFCFD083BFE50.taxon	discussion	Remarks. This specices shows sexual dimorphism in body colouration. In male, clypeus is yellow to yellowish brown, legs are uniformly yellow, and yellowish brown marks on the pronotum are much larger. In female, clypeus is dark brown, legs are yellow but with a large black marking on femur, yellowish brown marks on the pronotum are smaller. This species closely resembles S. langsonensis (Hoàng, 1982) and S. longisiphonulus (Hoàng, 1985) from Vietnam and they are possibly conspecific, but without the examination of the type specimens of these species, we treat them as valid species in the present study. Several individuals have complete abdominal postcoxal lines.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67DFFECFF7BFDC10930FE50.taxon	description	(Figs 53, 80) Chinese name: 尼泊尔 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67DFFECFF7BFDC10930FE50.taxon	diagnosis	Diagnosis. This species is similar to S. flavostictus sp. nov. in structures of the male genitalia, but can be distinguished from the later by its elytra black with yellowish brown apex, and abdominal postcoxal lines recurved toward the base of ventrite (Figs 53 a – b, 53 d). In S. flavostictus, elytra are black with two yellow spots and abdominal postcoxal lines are not recurved toward the base of ventrite (Figs 49 a – b, 49 d).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67DFFECFF7BFDC10930FE50.taxon	description	Description. TL: 1.49 – 2.11 mm, TW: 1.14 – 1.57 mm, TH: 0.69 – 0.94 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 53 a – c). Head black with antennae and mouthparts yellow, with interocular distance slightly greater than eye width in frontal view. Pronotum black with anterior margin narrowly yellowish brown. Scutellar shield black. Elytra black with yellowish brown apices. Ventral surfaces black. Legs yellow. Abdominal postcoxal lines almost complete (Fig. 53 d). Male genitalia: Penis slender, penis capsule with long inner arm but without outer one, apex of penis pointed (Fig. 53 e). Penis guide apically hook-shaped in lateral view; parameres about 1 / 5 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, tapering towards blunt apex (Figs 53 f – g). Female genitalia: Coxites shortened; styli absent; spermatheca worm-like, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67DFFECFF7BFDC10930FE50.taxon	materials_examined	Material examined. China: Guangxi: 3 ♂ 2 ♀, Pinglong Mountains, Shiwandashan National Nature Reserve, 6 – 8. XI. 2004, Wang XM leg; 2 ♂, Hongqi Forest Farm, Shiwandashan National Nature Reserve, 9 – 11. XI. 2004, Wang XM leg; 4 ♂ 2 ♀, Pinglong Mountains, Shiwandashan National Nature Reserve, 28. VII. 2005, Wang XM leg; 1 ♂, Tongling Canyon, Jingxi City, 5. VIII. 2005, Wang XM leg; 1 ♂, Hillside, Mao’ershan National Nature Reserve, 17. X. 2004, Wang XM leg. Hainan: 1 ♂, Wuzhishan National Nature Reserve, 1. XI. 1997, Peng ZQ leg; 1 ♀, Yinggeling National Nature Reserve, 23. XI. 1997, Peng ZQ leg; 10 ♂ 6 ♀, Limushan National Forest Park, 22. VII. 2006, Wang XM leg; 1 ♂ 1 ♀, Wuzhishan National Nature Reserve, 24. VII. 2006, Wang XM leg; 1 ♂, Hongkan, Yinggeling National Nature Reserve, 700 m, 17. IV. 2019, Wang XM leg. Yunnan: 4 ♂ 4 ♀, Nansan, Zhenkang County, 28. X. 2000, Peng ZQ leg; 1 ♂, Mengpeng Town, Mengla County, 23. IV. 2002, Peng ZQ leg; 1 ♂, Funing County, 16. VIII. 2005, Wang XM leg; 1 ♀, Maguan County, 18. VIII. 2005, Wang XM leg; 3 ♂, Hekou County – Pingbian County, 2005.8.19, Wang XM leg; 1 ♂ 2 ♀, Gongxin Village, Menglian County, 1500 m, 19. VIII. 2005, Wang XM leg; 1 ♀, Menglun Town, Mengla County, 21. VIII. 2005, Wang XM leg; 1 ♀, Ruili City, 27. VIII. 2005, Wang XM leg; 1 ♂, National highway 213 route, Mengla County, 12 – 13. X. 2006, Wang XM leg; 1 ♂, Xiaowei Mountains, Hekou County, 23. IV. 2008, Wang XM leg; 4 ♀, Dadugang Town, Jinghong City, 950 m, 26. IV. 2008, Wang XM leg; 1 ♀, Menglun Town – Mengla County, 1160 m, 29. IV. 2008, Wang XM leg; 3 ♂ 6 ♀, Sangyong Village – Longmen Village, Mengla County, 1. V. 2008, Wang XM leg; 1 ♂ 2 ♀, Lafu Village, Menglian County, 1130 m, 7. V. 2008, Wang XM leg; 2 ♀, Longtan, Ximeng County, 900 m, 9 – 10. V. 2008, Wang XM leg; 1 ♂, Banlao Town, Cangyuan County, 1100 m, 16. V. 2008, Wang XM leg; 1 ♂, Tongbiguan Town, Yingjiang County, 1000 m, 22 – 23. V. 2008, Wang XM leg; 1 ♂ 1 ♀, Caiyanghe National Forest Park, 4. V. 2009, Wang XM leg; 1 ♀, Dadugang Town, Jinghong City, 1050 m, 5. V. 2009, Wang XM leg; 1 ♂, Jinuoshan Town, Jinghong City, 6. V. 2009, Wang XM leg; 2 ♂ 2 ♀, Yaoqv Town, Mengla County, 700 m, 7 – 8. V. 2009, Wang XM leg; 1 ♂ 1 ♀, Longmen Village, Shangyong Town, Mengla County, 1027 m, 9. V. 2009, Wang XM leg; 2 ♂, Aini Village, Jinghong City, 11. V. 2009, Wang XM leg; 1 ♀, Jiangcheng County, 1150 m, 15. V. 2009, Wang XM leg; 1 ♀, Yingpan Town, Jingping County, 1487 m, 5. VIII. 2013, Chen XS leg; 1 ♀, Jinuoshan Town, 992 m, 10. VIII. 2013, Chen XS leg; 2 ♂ 2 ♀, Bulangshan Town, Menghai County, 1188 m, 12. VIII. 2013, Chen XS leg. Xizang: 1 ♂, Yarang Village, Muotuo County, 768 – 1030 m, 24. X. 2007, Wang XM leg; 1 ♂, Beibeng Town, Muotuo County, 600 – 768 m, 25. X. 2007, Wang XM leg; 1 ♂, Zhamo Highroad 113 K, Bomi County, 835 m, 30 – 31. X. 2007, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67DFFECFF7BFDC10930FE50.taxon	distribution	Distribution. China (Guangxi, Hainan, Yunnan, Xizang); Nepal.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67DFFECFF7BFDC10930FE50.taxon	discussion	Remarks. This species is recorded from China for the first time. The specimens perfectly match the descriptions and illustrations of Miyatake (1985).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67CFFEEFF7BF95B0F0BFD70.taxon	description	(Figs 54, 77) Chinese name: Ñae ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67CFFEEFF7BF95B0F0BFD70.taxon	description	Description. TL: 1.60 – 2.21 mm, TW: 1.21 – 1.63 mm, TH: 0.73 – 0.99 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 54 a – c). Head yellowish brown with interocular distance slightly greater than eye width in frontal view. Pronotum yellowish brown. Scutellar shield black. Elytra black with yellowish brown apices. Ventral surfaces yellowish brown to black. Legs yellowish brown. Abdominal postcoxal lines almost complete (Fig. 54 d). Male genitalia: Penis slender, penis capsule with long inner arm but without outer one, apex of penis pointed (Figs 54 e – f). Penis guide widest at base in lateral view, gradually narrowing to blunt apex; parameres about 1 / 4 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, tapering towards blunt apex (Figs 54 g – h). Female genitalia: Coxites shortened; styli absent; spermatheca worm-like, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67CFFEEFF7BF95B0F0BFD70.taxon	materials_examined	Material examined. Holotype: China: Fujian: ♂, Nanjing County 10. III. 1985, Liu SJ leg (SCAU). Paraype: China: Fujian: 1 ♀, Same data as holotype (SCAU). Other material (SCAU). China: Guangdong: 1 ♂, Chebaling National Nature Reserve, 24. IV. 1991, Yu GY leg; 25 ♂ 5 ♀, Hengshitang, Shimentai National Nature Reserve, 5. X. 2004, Wang XM leg; 3 ♂, Yangqiu Mountains, Shimentai National Nature Reserve, 6. X. 2004, Wang XM leg; 1 ♂, Huangniping, Shimentai National Nature Reserve, 7. X. 2004, Wang XM leg; 11 ♂, Wulangzhang, Shimentai National Nature Reserve, 8. X. 2004, Wang XM leg; 7 ♂ 2 ♀, Nankunshan National Forest Park, 21. X. 2004, Wang XM leg. Guangxi: 5 ♂ 3 ♀, Hongqi Forest Farm, Shiwandashan National Nature Reserve, 9 – 11. XI. 2004, Wang XM leg; 1 ♂, Pinglong Mountains, Shiwandashan National Nature Reserve, 28. VII. 2005, Wang XM leg; 1 ♂, Baidu Town, Napo County, 4. VIII. 2005, Wang XM leg. Sichuan: 2 ♂, Puge County, 2000 m, 15. IX. 2007, Wang XM leg. Guizhou: 1 ♂, Huangguoshu Waterfalls Scenic Area, 15. VIII. 1990, Tian MY leg; 5 ♂ 1 ♀, Weng’ang, Maolan National Nature Reserve, 800 m, 15. VIII. 2006, Wang XM leg; 3 ♂ 1 ♀, Maolan National Nature Reserve, 16. VIII. 2006, Wang XM leg; 3 ♀, Wuyan Bridge, Maolan National Nature Reserve, 17. VIII. 2006, Wang XM leg; 35 ♂ 46 ♀, Xiaoqikong Town, Libo County, 18. VIII. 2006, Wang XM leg; 8 ♂ 10 ♀, Luodian County, 19. VIII. 2006, Wang XM leg; 4 ♂ 3 ♀, Luodian County, 20. VIII. 2006, Wang XM leg; 2 ♂ 6 ♀, Xianheping Nature Reserve, 14 – 15. IX. 2006, Wang XM leg; 3 ♀, Xianheping Nature Reserve, 1500 m, 12. IX. 2007, Wang XM leg; 1 ♀, Xiaodanjiang, Leigongshan National Nature Reserve, 12 – 13. X. 2008, Wang XM leg; 2 ♀, Dongtang, Maolan National Nature Reserve, 730 m, 15 – 18. X. 2008, Wang XM leg; 1 ♂, Sancha River, Maolan National Nature Reserve, 750 m, 19. X. 2008, Wang XM leg; 1 ♀, Weng’ang, Maolan National Nature Reserve, 750 m, 20 – 23. X. 2008, Wang XM leg. Yunnan: 6 ♂ 1 ♀, Malipo County, 17. VIII. 2005, Wang XM leg; 1 ♂, Ruili City, 27. VIII. 2005, Wang XM leg; 1 ♂ 3 ♀, Mangbing, Yingjiang City, 29. VIII. 2005, Wang XM leg; 4 ♂, Menglun Town, Mengla County, 1160 m, 29. IV. 2008, Wang XM leg; 1 ♀, Jinuoshan Town, Jinghong City, 6. V. 2009, Wang XM leg; 1 ♀, Aini Village, Jinghong City, 11. V. 2009, Wang XM leg; 2 ♂, Heiwadi, Gongshan County, 2020 m, 19. VII. 2010, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67CFFEEFF7BF95B0F0BFD70.taxon	distribution	Distribution. China (Fujian, Guangdong, Guangxi, Sichuan, Guizhou, Yunnan).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67EFFEEFF7BFCA10F3DF8CC.taxon	description	(Figs 55, 76) Chinese name: 平叶 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67EFFEEFF7BFCA10F3DF8CC.taxon	diagnosis	Diagnosis. This species is similar to S. applanatus sp. nov. in structures of the male genitalia, but can be distinguished from the later by its more robust penis guide in lateral view and yellow pronotum (Figs 55 a – c, 55 f). In S. applanatus, penis guide is more slender and pronotum is black (Figs 60 a – c, 60 f).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67EFFEEFF7BFCA10F3DF8CC.taxon	description	Description. TL: 1.55 – 2.19 mm, TW: 1.20 – 1.76 mm, TH: 0.74 – 0.94 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 55 a – c). Head yellowish brown with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow. Scutellar shield black. Elytra black with yellowish brown apices. Ventral surfaces yellowish brown to black. Legs yellow. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 55 d). Male genitalia: Penis slender, penis capsule with long inner arm but without outer one, apex of penis pointed (Fig. 55 e). Penis guide long and slender in lateral view, widest at base, gradually narrowing to blunt apex; parameres about 1 / 4 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 2 in ventral view, then slightly narrowing to blunt apex (Figs 55 f – g). Female genitalia: Coxites shortened; styli absent; spermatheca worm-like, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67EFFEEFF7BFCA10F3DF8CC.taxon	materials_examined	Type material. Holotype: China: Hainan: ♂, Bawangling National Nature Reserve, 5. V. 2005, Wang XM leg (SCAU). Paratypes (SCAU): China: Hainan: 2 ♂, Jianfengling National Nature Reserve, 12. XI. 1989, Ren SX leg; 4 ♂ 10 ♀, Bawangling National Nature Reserve, 21. III. 1996, Peng ZQ leg. Yunnan: 1 ♂, Menglun Town, Mengla County, 21. VIII. 2005, Wang XM leg; 2 ♂, Tongbiguan Town, Yingjiang County, 30. VIII. 2005, Wang XM leg; 1 ♀, Lafu Village, Menglian County, 1130 m, 7. V. 2008, Wang XM leg; 1 ♀, Lancang County, 1300 m, 11. V. 2008, Wang XM leg; 5 ♂ 1 ♀, Yaoqv Town, Mengla County, 700 m, 7 – 8. V. 2009, Wang XM leg; 1 ♀, Nanping, Mengman Town, Mengla County, 745 m, 10. V. 2009, Wang XM leg; 1 ♂ 4 ♀, Meng’a Town, Menghai County, 1170 m, 12. V. 2009, Wang XM leg; 1 ♀, Aini Village, Jinghong City, 700 m, 13. VIII. 2013, Chen XS leg. Xizang: 1 ♂ 2 ♀, Xiachayu Town, Chaiyu County, 1800 m, 14. X. 2007, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67EFFEEFF7BFCA10F3DF8CC.taxon	distribution	Distribution. China (Hainan, Yunnan, Xizang).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F67EFFEEFF7BFCA10F3DF8CC.taxon	etymology	Etymology. The specific epithet parallelus derives from the Latin word “ parallelus ” and refers to the penis guide which is approximately parallel sided for basal 1 / 2 in both ventral and lateral view.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F661FFF0FF7BFA690A5FF98C.taxon	description	(Figs 1 d, 3 l, 3 p, 56, 77) Chinese name: D 叶 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F661FFF0FF7BFA690A5FF98C.taxon	diagnosis	Diagnosis. This is a very distinctive species of Sasajiscymnus characterized by the dark brown head, stout penis without outer arm, and the penis guide with apical 1 / 3 expanded (Figs 56 a – c, 56 g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F661FFF0FF7BFA690A5FF98C.taxon	description	Description. TL: 2.20 – 2.33 mm, TW: 1.68 – 1.79 mm, TH: 0.97 – 1.05 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 56 a – c). Head dark brown with interocular distance distinctly greater than eye width in frontal view. Pronotum yellowish brown with a black spot at base. Scutellar shield black. Elytra black with brown apices. Ventral surfaces yellowish brown to black. Legs yellowish brown to black. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 56 d). Male genitalia: Penis stout, penis capsule with long inner arm but without outer one, apex of penis pointed (Fig. 56 e). Penis guide expanded apically in lateral view; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide distinctly swollen near apex in ventral view, blunt apically (Figs 56 f – g). Female genitalia: Coxites shortened; styli absent; spermatheca worm-like, without clear ramus or nodulus (Figs 3 l, 3 p).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F661FFF0FF7BFA690A5FF98C.taxon	materials_examined	Type material. Holotype: China: Taiwan: ♂, Beiheng Highway, Yilan County, 300 – 1500 m, 1. XI. 2012, Wang XM leg (SCAU). Paratype: China, Taiwan: 1 ♀, Same data as holotype (SCAU). Distribution. China (Taiwan). Etymology. The specific epithet tumidus derives from the Latin word “ tumidus ” and refers to the expanded penis guide.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F660FFF5FF7BF9BD0991FE9C.taxon	description	(Figs 1 f, 57, 79) Chinese name: ĤW ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F660FFF5FF7BF9BD0991FE9C.taxon	diagnosis	Diagnosis. This species closely resembles S. kurohime (Miyatake, 1959), but can be distinguished from the later by the penis guide blunt apically (Figs 57 h – i). In S. kurohime, penis guide is pointed apically (Figs 58 g – h). Body oval, moderately convex, surfaces covered with short pubescence (Figs 57 a – c). Head black with antennae and mouthparts yellowish brown, interocular distance distinctly greater than eye width in frontal view. Pronotum black with anterior margin narrowly yellowish brown. Scutellar shield black. Elytra black with brown apices. Ventral surfaces black. Legs yellowish brown to black. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 57 e). Abdominal ventrite 6 of females with a deep terminal emargination (Fig. 57 d). Male genitalia: Penis slender, penis capsule with long inner arm but without outer one, apex of penis pointed (Figs 57 f – g). Penis guide long and slender in lateral view, widest at base, gradually narrowing to blunt apex; parameres about 1 / 6 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, tapering towards blunt apex (Figs 57 h – i). Female genitalia: Coxites shortened; styli absent; spermatheca worm-like, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F660FFF5FF7BF9BD0991FE9C.taxon	materials_examined	Material examined. China: Zhejiang: 1 ♀, Xianxialing Nature Reserve, Jiangshan City, [118 ° 40 ' 49.80 " E 28 ° 17 ' 38.04 " N], 617 m, 4. X. 2017, Tong JB leg; 2 ♂ 1 ♀, Wangdongyang Mountain Wetland Nature Reserve, [119 ° 38 ' 12.09 " E 27 ° 41 ' 19.61 " N], 1261 m, 10. X. 2017, Tong JB leg. Anhui: 1 ♂ 3 ♀, Tangkou Town, Huangshan District, Huangshan City, 460 m, 12 – 13. IX. 2010, Wang XM leg; 1 ♀, Guniujiang National Nature Reserve, 24. IX. 2010, Wang XM leg; 1 ♂ 1 ♀, Yaoluoping National Nature Reserve, 1000 m, 28 – 29. IX. 2010, Wang XM leg; 5 ♂ 2 ♀, Jinzhai Tianma National Nature Reserve, 600 m, 3 – 5. X. 2010, Wang XM leg. Fujian: 1 ♂, Shaowu City, IX. 1982, Li P leg; 1 ♂ 1 ♀, Sangang, Wuyishan National Nature Reserve, 20. VIII. 1984, Zhang KC leg; 1 ♂ 2 ♀, Guizhu, Meihuashan National Nature Reserve, 1200 m, 20. VIII. 2012, Huo LZ leg; 2 ♀, Yuncai, Liangyeshan National Nature Reserve, 735 m, 23. VIII. 2012, Huo LZ leg; 16 ♂ 29 ♀, Sanbailiao, Tianboyan National Nature Reserve, 690 m, 26. VIII. 2012, Huo LZ leg; 1 ♂, Fengtian, Tianboyan National Nature Reserve, 800 m, 27. VIII. 2012, Huo LZ leg; 2 ♂, Shipaichang, Longxishan National Nature Reserve, 500 m, 28 – 29. VIII. 2012, Huo LZ leg; 3 ♀, Fanjia, Junzifeng National Nature Reserve, 460 m, 1. IX. 2012, Huo LZ leg; 2 ♂ 1 ♀, Zhangjia, Wuyishan National Nature Reserve, 950 m, 16. IX. 2012, Huo LZ leg; 3 ♀, Tongmu Village, Wuyishan National Nature Reserve, 750 m, 17. IX. 2012, Huo LZ leg; 9 ♂ 1 ♀, Hongshuikeng, Wuyishan National Nature Reserve, 600 m, 18. IX. 2012, Huo LZ leg; 1 ♂ 11 ♀, Guaxi Village, Fu’an City, 260 m, 21. IX. 2012, Huo LZ leg. Jiangxi: 1 ♂, Kenzhichang, Jinggangshan City, 13. VIII. 2004, Wang XM leg. Henan: 1 ♂, Jigongshan National Nature Reserve, 697 m, 4. VII. 2009, Wang XM leg; 2 ♂ 3 ♀, Purdom Forest Park, Jigongshan National Nature Reserve, 216 m, 5. VII. 2009, Wang XM leg; 1 ♀, Baotianman National Nature Reserve, 6. VII. 2009, Wang XM leg. Hubei: 1 ♂, Wudang Mountains, Shiyan City, 17. VII. 1997, Peng ZQ leg; 1 ♂, Banqiao, Shennongjia National Nature Reserve, 1170 m, 21 – 24. VII. 2007, Wang XM leg; 1 ♀, Wudang Mountains, Shiyan City, 700 – 930 m, 8. VIII. 2007, Wang XM leg. Hunan: 10 ♂ 22 ♀, Shennonggu National Forest Park, 1100 m, 7. X. 2010, Wang XM leg. Guangdong: 1 ♂, Nankunshan National Forest Park, XI. 1986, Jiang N leg; 1 ♂, Shixing County, 8. VII. 1989, Pang XF leg; 1 ♀, Shixing County, 9. VII. 1989, Pang XF leg; 1 ♂, Shixing County, 10. VII. 1989, Pang XF leg; 1 ♂ 1 ♀, Chebaling National Nature Reserve, 2. XI. 1989, Yu GY leg; 3 ♂ 2 ♀, Chebaling National Nature Reserve, 3. XI. 1989, Zhang J leg; 1 ♂, Chebaling National Nature Reserve, 19. VI. 1990, Yu GY leg; 1 ♂ 2 ♀, Chebaling National Nature Reserve, 24. IV. 1991, Yu GY leg; 1 ♂ 1 ♀, Chebaling National Nature Reserve, 26. IV. 1991, Yu GY leg; 4 ♂ 1 ♀, Qinshuigu, Nanling National Nature Reserve, 30. IX. 2004, Wang XM leg; 1 ♂ 1 ♀, Xiaohuang Mountains, Nanling National Nature Reserve, 1. X. 2004, Wang XM leg; 1 ♂ 1 ♀, Xidianzhan, Nanling National Nature Reserve, 1. X. 2004, Wang XM leg; 1 ♂ 1 ♀, Yangqiu Mountains, Shimentai National Nature Reserve, 6. X. 2004, Wang XM leg; 1 ♂ 1 ♀, Huangniping, Shimentai National Nature Reserve, 7. X. 2004, Wang XM leg; 2 ♂ 3 ♀, Huangdong, Shimentai National Nature Reserve, 30. X. 2004, Wang XM leg; 1 ♂ 2 ♀, Huangdong, Shimentai National Nature Reserve, 31. X. 2004, Wang XM leg; 1 ♂, Tianjingshan National Forest Park, 17. VII. 2005, Wang XM leg; 5 ♂ 6 ♀, Qinshuigu, Nanling National Nature Reserve, 21. IX. 2005, Wang XM leg. Guangxi: 45 ♂ 29 ♀, Hillside, Mao’ershan National Nature Reserve, 16 – 17. X. 2004, Wang XM leg; 1 ♂ 5 ♀, Jiulongtang, Mao’ershan National Nature Reserve, 18. X. 2004, Wang XM leg; 5 ♂ 4 ♀, Shili Canyon, Mao’ershan National Nature Reserve, 19. X. 2004, Wang XM leg; 18 ♂ 9 ♀, GaoZhai, Mao’ershan National Nature Reserve, 20. X. 2004, Wang XM leg. Hainan: 1 ♂, Limushan National Forest Park, 10. XI. 1989, Ren SX leg; 1 ♀, Bawangling National Nature Reserve, 21. III. 1996, Peng ZQ leg; 16 ♂ 10 ♀, Limushan National Forest Park, 22. VII. 2006, Wang XM leg; 6 ♂ 8 ♀, Diaoluoshan National Nature Reserve, 25. VII. 2006, Peng ZQ leg. Sichuan: 1 ♀, Heizhugou National Nature Reserve, 1900 m, 22 – 23. IX. 2007, Wang XM leg; 1 ♂, Labahe National Nature Reserve, 1100 m, 4. X. 2007, Wang XM leg. Guizhou: 2 ♂ 1 ♀, Fanjingshan National Nature Reserve, 21. VII. 1990, Ren SX leg; 1 ♂, Xiaoqikong Town, Libo County, 18. VIII. 2006, Wang XM leg; 2 ♀, Lianhuaping, Leigongshan National Nature Reserve, 8. X. 2008, Wang XM leg; 2 ♂, Taojiang Town, Rongjiang County, 1000 m, 10 – 11. X. 2008, Wang XM leg; 1 ♂ 5 ♀, Xiaodanjiang, Leigongshan National Nature Reserve, 12 – 13. X. 2008, Wang XM leg; 1 ♀, Jiaomi Village, Taijiang County, 710 m, 14. X. 2008, Wang XM leg; 1 ♂ 2 ♀, Dongtang, Maolan National Nature Reserve, 730 m, 15 – 18. X. 2008, Wang XM leg; 1 ♂, Sancha River, Maolan National Nature Reserve, 750 m, 19. X. 2008, Wang XM leg; 4 ♀, Weng’ang, Maolan National Nature Reserve, 750 m, 20 – 23. X. 2008, Wang XM leg. Yunnan: 3 ♂, Nansan, Zhenkang County, 28. X. 2000, Peng ZQ leg; 1 ♂, Xujiaba, Ailaoshan National Nature Reserve, 1 – 3. X. 2006, Wang XM leg; 1 ♀, Taizhong Town, Jingdong County, 3. X. 2006, Wang XM leg; 1 ♂, National highway 213 route, Mengla County, 12 – 13. X. 2006, Wang XM leg; 3 ♂, Dadugang Town, Jinghong City, 950 m, 26. IV. 2008, Wang XM leg; 10 ♂ 18 ♀, Lafu Village, Menglian County, 1130 m, 7. V. 2008, Wang XM leg; 2 ♂ 1 ♀, Banlao Town, Cangyuan County, 1100 m, 16. V. 2008, Wang XM leg; 1 ♀, Meng’a Town, Menghai County, 1170 m, 12. V. 2009, Wang XM leg; 2 ♂ 2 ♀, Longyuan Village, Gongshan County, 1690 m, 24 – 25. VII. 2010, Wang XM leg; 7 ♂ 3 ♀, Longyuan Village – Kongdang Village, Gongshan County, 1500 m, 26. VII. 2010, Wang XM leg; 1 ♂ 3 ♀, Kongdang Village, Gongshan County, 27. VII. 2010, Wang XM leg; 3 ♂ 3 ♀, Bapo Village – Maku Village, Gongshan County, 1450 m, 29. VII. 2010, Wang XM leg; 2 ♂, Maku Village, Gongshan County, 1600 m, 1. VIII. 2010, Wang XM leg; 1 ♂, Jinuoshan Town, 992 m, 10. VIII. 2013, Chen XS leg; 1 ♀, County highway J 22 route, Wuliangshan National Nature Reserve, 2400 m, 18. VIII. 2013, Chen XS leg. Xizang: 3 ♂ 8 ♀, Cibagou National Nature Reserve, 2000 m, 13. X. 2007, Wang XM leg; 1 ♀, Beibeng Town, Muotuo County, 600 – 768 m, 25. X. 2007, Wang XM leg; 3 ♀, Zhamo Highroad 140 K, Bomi County, 900 m, 27. X. 2007, Wang XM leg; 3 ♀, Zhamo Highroad 113 K, Bomi County, 835 m, 30 – 31. X. 2007, Wang XM leg; 1 ♂ 2 ♀, Pailong Town, Bayi District, Linzhi City, 13. X. 2009, Wang XM leg; 1 ♀, Yihaoqiao, Hanmi, Muotuo County, 14. X. 2009, Wang XM leg; 2 ♀, Hanmi, Muotuo County, 800 – 2100 m, 5 – 8. X. 2011, Huo LZ leg; 2 ♀, Bangxi Town, Muotuo County, 1840 m, 13. X. 2011, Huo LZ leg; 4 ♂ 2 ♀, Chilongzangbu, Linzhi City, 2200 m, 19. X. 2011, Huo LZ leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F660FFF5FF7BF9BD0991FE9C.taxon	distribution	Distribution. China (Zhejiang, Anhui, Fujian, Henan, Hubei, Hunan, Guangdong, Guangxi, Hainan, Chongqing, Sichuan, Guizhou, Yunnan, Xizang); Vietnam.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F665FFF7FF7BFE8D0F84FCDC.taxon	description	(Figs 3 k, 3 o, 58, 78) Chinese name: DZ ḧẋ 毛瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F665FFF7FF7BFE8D0F84FCDC.taxon	diagnosis	Diagnosis. This species closely resembles S. dapae (Hoàng, 1978), but can be distinguished from the later by the penis guide pointed apically (Figs 58 g – h). In S. dapae, penis guide is blunt apically (Figs 57 h – i).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F665FFF7FF7BFE8D0F84FCDC.taxon	description	Description. TL: 1.67 – 2.29 mm, TW: 1.26 – 1.71 mm, TH: 0.69 – 0.99 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 58 a – c). Head black with antennae and mouthparts yellowish brown, interocular distance distinctly greater than eye width in frontal view. Pronotum black with anterior margin narrowly yellowish brown. Scutellar shield black. Elytra black with brown apices. Ventral surfaces black. Legs yellowish brown to black. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 58 d). Male genitalia: Penis slender, penis capsule with long inner arm but without outer one, apex of penis pointed (Figs 58 e – f). Penis guide long and slender in lateral view, widest at base, gradually narrowing to pointed apex; parameres about 1 / 6 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, tapering towards pointed apex (Figs 58 g – h). Female genitalia: Coxites shortened; styli absent; spermatheca worm-like, with basal ramus and nodulus (Figs 3 k, 3 o).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F665FFF7FF7BFE8D0F84FCDC.taxon	materials_examined	Material examined. China: Anhui: 1 ♀, Tangkou Town, Huangshan District, Huangshan City, 460 m, 12 – 13. IX. 2010, Wang XM leg; 2 ♂ 1 ♀, Hengdu Town, Shitai County, 230 m, 20. IX. 2010, Wang XM leg. Fujian: 2 ♂ 2 ♀, Xiangxi, Huboliao National Nature Reserve, 245 m, 18. VIII. 2012, Huo LZ leg; 21 ♂ 47 ♀, Sanbailiao, Tianboyan National Nature Reserve, 690 m, 26. VIII. 2012, Huo LZ leg; 2 ♂ 6 ♀, Fengtian, Tianboyan National Nature Reserve, 800 m, 27. VIII. 2012, Huo LZ leg; 3 ♀, Zonghechang, Longxishan National Nature Reserve, 750 m, 30. VIII. 2012, Huo LZ leg; 2 ♂ 6 ♀, Fanjia, Junzifeng National Nature Reserve, 460 m, 1. IX. 2012, Huo LZ leg; 1 ♂ 1 ♀, Hongshuikeng, Wuyishan National Nature Reserve, 600 m, 18. IX. 2012, Huo LZ leg; 1 ♂, Chishui, Daiyunshan National Nature Reserve, 850 m, 22. IX. 2012, Huo LZ leg. Guangxi: 3 ♂, Hillside, Mao’ershan National Nature Reserve, 17. X. 2004, Wang XM leg; 1 ♂, Jiulongtang, Mao’ershan National Nature Reserve, 18. X. 2004, Wang XM leg; 2 ♀, Shili Canyon, Mao’ershan National Nature Reserve, 19. X. 2004, Wang XM leg; 1 ♂ 1 ♀, GaoZhai, Mao’ershan National Nature Reserve, 20. X. 2004, Wang XM leg; 4 ♂ 3 ♀, Pinglong Mountains, Shiwandashan National Nature Reserve, 6 – 8. XI. 2004, Wang XM leg; 20 ♂ 13 ♀, Hongqi Forest Farm, Shiwandashan National Nature Reserve, 9 – 11. XI. 2004, Wang XM leg. Guangdong: 2 ♂, Nankunshan National Forest Park, XI. 1986, Jiang N leg; 1 ♂ 3 ♀, Chebaling National Nature Reserve, 23. III. 1989, Yu GY leg; 2 ♀, Chebaling National Nature Reserve, 24. IV. 1989, Yu GY leg; 1 ♀, Chebaling National Nature Reserve, 26. IV. 1989, Yu GY leg; 1 ♀, Chebaling National Nature Reserve, 8. VII. 1989, Yu GY leg; 1 ♀, Chebaling National Nature Reserve, 9. VII. 1989, Pang XF leg; 2 ♀, Shixing County, 10. VII. 1989, Pang XF leg; 1 ♂, Chebaling National Nature Reserve, 2. XI. 1989, Yu GY leg; 4 ♂ 7 ♀, Chebaling National Nature Reserve, 3. XI. 1989, Yu GY leg; 1 ♀, Chebaling National Nature Reserve, 4. XI. 1989, Yu GY leg; 1 ♂, Chebaling National Nature Reserve, 19. VI. 1990, Yu GY leg; 2 ♀, Chebaling National Nature Reserve, VI. 1991, Pang XF leg; 2 ♂ 1 ♀, Chebaling National Nature Reserve, 23. IV. 1991, Yu GY leg; 1 ♂ 1 ♀, Yangqiu Mountains, Shimentai National Nature Reserve, 6. X. 2004, Wang XM leg; 1 ♂ 3 ♀, Nankunshan National Forest Park, 21. X. 2004, Wang XM leg; 1 ♀, Huangdong, Shimentai National Nature Reserve, 30. X. 2004, Wang XM leg. Guizhou: 2 ♀, Weng’ang, Maolan National Nature Reserve, 800 m, 15. VIII. 2006, Wang XM leg; 1 ♀, Lianhuaping, Leigongshan National Nature Reserve, 8. X. 2008, Wang XM leg; 4 ♂, Dongtang, Maolan National Nature Reserve, 730 m, 15 – 18. X. 2008, Wang XM leg. Yunnan: 2 ♂ 3 ♀, Nansan, Zhenkang County, 28. X. 2000, Peng ZQ leg; 1 ♀, Funing County, 16. VIII. 2005, Wang XM leg; 1 ♀, Malipo County, 17. VIII. 2005, Wang XM leg; 1 ♂, Menglun Town, Mengla County, 21. VIII. 2005, Wang XM leg; 1 ♂ 1 ♀, Ruili City, 27. VIII. 2005, Wang XM leg; 1 ♀, Mangbing, Yingjiang City, 29. VIII. 2005, Wang XM leg; 1 ♀, Gaoligongshan National Nature Reserve, 19. IX. 2006, Wang XM leg; 1 ♂, Taizhong Town, Jingdong County, 3. X. 2006, Wang XM leg; 1 ♂ 1 ♀, Longmen Village, Shangyong Town, Mengla County, 8. X. 2006, Wang XM leg; 3 ♂ 7 ♀, Dadugang Town, Jinghong City, 950 m, 26. IV. 2008, Wang XM leg; 1 ♀, Menglun Town, Mengla County, 1160 m, 29. IV. 2008, Wang XM leg; 10 ♂ 13 ♀, Sangyong Village – Longmen Village, Mengla County, 1. V. 2008, Wang XM leg; 1 ♂, Lafu Village, Menglian County, 1130 m, 7. V. 2008, Wang XM leg; 4 ♂ 3 ♀, Gongxin Village, Menglian County, 1500 m, 8. V. 2008, Wang XM leg; 2 ♂, Longtan, Ximeng County, 900 m, 9 – 10. V. 2008, Wang XM leg; 1 ♂, Lancang County, 1300 m, 11. V. 2008, Wang XM leg; 1 ♂ 2 ♀, Caiyanghe National Forest Park, 4. V. 2009, Wang XM leg; 4 ♂ 2 ♀, Dadugang Town, Jinghong City, 1050 m, 5. V. 2009, Wang XM leg; 16 ♂ 22 ♀, Yaoqv Town, Mengla County, 700 m, 7 – 8. V. 2009, Wang XM leg; 2 ♀, Longmen Village, Shangyong Town, Mengla County, 1027 m, 9. V. 2009, Wang XM leg; 1 ♂ 3 ♀, Nanping, Mengman Town, Mengla County, 745 m, 10. V. 2009, Wang XM leg; 96 ♂ 57 ♀, Meng’a Town, Menghai County, 1170 m, 12. V. 2009, Wang XM leg; 1 ♂ 4 ♀, Jiangcheng County, 1150 m, 15. V. 2009, Wang XM leg; 2 ♂ 4 ♀, Heiwadi, Gongshan County, 2020 Bapo Village, Gongshan County, 1400 m, 28. VII. 2010, Wang XM leg; 2 ♂ 9 ♀, Bapo Village – Maku Village, Gongshan County, 1450 m, 29. VII. 2010, Wang XM leg; 1 ♂ 3 ♀, Maku Village, Gongshan County, 1600 m, 1. VIII. 2010, Wang XM leg; 19 ♂ 19 ♀, Qinglandan, Maku Village, Gongshan County, 4 – 7. VIII. 2010, Wang XM leg. Xizang: 1 ♀, Cibagou National Nature Reserve, 2000 m, 13. X. 2007, Wang XM leg; 1 ♀, Yihaoqiao, Hanmi, Muotuo County, 14. X. 2009, Wang XM leg; 1 ♂, Muotuo County, 17. X. 2009, Wang XM leg; 1 ♂, Yarang Village, Muotuo County, 768 – 1030 m, 24. X. 2007, Wang XM leg; 1 ♂, Beibeng Town, Muotuo County, 600 – 760 m, 25. X. 2007, Wang XM leg; 1 ♀, Zhamo Highroad 140 K, Bomi County, 900 m, 27. X. 2007, Wang XM leg; 1 ♂, Zhamo Highroad 113 K, Bomi County, 835 m, 30 – 31. X. 2007, Wang XM leg. Taiwan: 5 ♂ 11 ♀, Aowanda, Jenai Town, Nantou County, 1280 m, 21. X. 2012, Wang XM leg; 1 ♂ 1 ♀, Alishan Town, 1800 m, 22. X. 2012, Wang XM leg; 6 ♂ 5 ♀, Dabang Village, Alishan Mountains, Chiayi County, 960 – 1600 m, 24. X. 2012, Wang XM leg; 1 ♂ 1 ♀, Maolin District, Kaohsiung City, 365 – 515 m, 26. X. 2012, Wang XM leg; 2 ♂ 7 ♀, Shuangli Village, Pingtung County, 200 – 400 m, 27. X. 2012, Wang XM leg; 3 ♂ 1 ♀, Jhihben, Taitung City, 230 m, 28. X. 2012, Wang XM leg; 2 ♂ 5 ♀, Nanheng, Chihshang Town, Taitung County, 679 m, 29. X. 2012, Wang XM leg; 2 ♂ 3 ♀, Taroko National Park, Hualien County, 200 – 912 m, 30. X. 2012, Wang XM leg; 1 ♂ 2 ♀, Beiheng Highway, Yilan County, 300 – 1500 m, 1. XI. 2012, Wang XM leg; 1 ♂, Nanhua Village, Xinshe District, Taichung City, 800 m, 17. VIII. 2018, Zhou WY leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F665FFF7FF7BFE8D0F84FCDC.taxon	distribution	Distribution. China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Yunnan, Xizang, Taiwan); Japan; Vietnam; Micronesia.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F665FFF7FF7BFE8D0F84FCDC.taxon	discussion	Remarks. Several individuals have complete abdominal postcoxal lines. Ren et al. (2009) presented photos of S. kurohime, but the penis guide in these photos were blunt apically in both lateral and ventral view. Hence that description is considered as a misidentification of S. dapae.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F667FFF7FF7BFC4D0BA2F960.taxon	description	(Figs 59, 76) Chinese name: 牯牛 Ṟḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F667FFF7FF7BFC4D0BA2F960.taxon	diagnosis	Diagnosis. This species is similar to S. kurohime (Miyatake, 1959), but can be distinguished from the later by parameres about 1 / 3 as long as penis guide (Fig. 59 f). In S. kurohime, parameres are about 1 / 6 as long as penis guide (Fig. 58 g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F667FFF7FF7BFC4D0BA2F960.taxon	description	Description. TL: 1.39 – 1.88 mm, TW: 1.05 – 1.46 mm, TH: 0.74 – 1.00 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 59 a – c). Head black with interocular distance distinctly greater than eye width in frontal view. Pronotum and scutellar shield black. Elytra black with brown apices. Ventral surfaces yellowish black. Legs black to yellowish brown. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 59 d). Male genitalia: Penis slender, penis capsule with long inner arm but without outer one, apex of penis pointed (Fig. 59 e). Penis guide widest at base in lateral view, gradually narrowing to pointed apex; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base in ventral view, then tapering towards pointed apex (Figs 59 f – g). Female genitalia: Coxites shortened; styli absent; spermatheca worm-like, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F667FFF7FF7BFC4D0BA2F960.taxon	materials_examined	Type material. Holotype: China: Anhui: ♂, Guniujiang National Nature Reserve, 24. IX. 2010, Wang XM leg (SCAU). Paratypes (SCAU): China: Anhui: 1 ♂ 3 ♀, Same data as holotype; 3 ♀, Tangkou Town, Huangshan District, Huangshan City, 460 m, 12 – 13. IX. 2010, Wang XM leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F667FFF7FF7BFC4D0BA2F960.taxon	distribution	Distribution. China (Anhui).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F667FFF7FF7BFC4D0BA2F960.taxon	etymology	Etymology. This new species is named after Guniujiang National Nature Reserve where the holotype was collected.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F666FFF9FF7BFA650A15FEE5.taxon	description	(Figs 60, 76) Chinese name: G 叶 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F666FFF9FF7BFA650A15FEE5.taxon	diagnosis	Diagnosis. This species is similar to S. lewisi (Kamiya, 1961), but can be distinguished from the later by its parameres about 1 / 4 as long as penis guide, and penis guide approximately parallel sided for basal 1 / 3 in ventral view (Figs 60 f – g). In S. lewisi, parameres are about 1 / 3 as long as penis guide, and penis guide is approximately parallel sided for basal 2 / 3 in ventral view, then tapering towards blunt apex (Figs 61 f – g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F666FFF9FF7BFA650A15FEE5.taxon	description	Description. TL: 1.36 – 1.58 mm, TW: 0.99 – 1.18 mm, TH: 0.52 – 0.67 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 60 a – c). Body uniformly black, except antennae, maxillary palpi and legs yellow to yellowish brown. Interocular distance slightly closer than eye width in frontal view. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 60 d). Male genitalia: Penis slender, penis capsule with long inner arm but without outer one, apex of penis pointed (Fig. 60 e). Penis guide approximately parallel sided in lateral view; parameres about 1 / 4 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 1 / 3 in ventral view, then slightly narrowing to blunt apex (Figs 60 f – g). Female genitalia: Coxites shortened; styli absent; spermatheca worm-like, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F666FFF9FF7BFA650A15FEE5.taxon	materials_examined	Type material. Holotype: China: Xizang: ♂, Shangchayu Town – Xiachayu Town, Chayu County, 1650 m, 27. IX. 2011, Huo LZ leg (SCAU). Paratypes (SCAU): Xizang: 3 ♂ 8 ♀, Same data as holotype; 1 ♀, Cibagou National Nature Reserve, 1615 m, 26. IX. 2011, Huo LZ leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F666FFF9FF7BFA650A15FEE5.taxon	distribution	Distribution. China (Xizang).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F666FFF9FF7BFA650A15FEE5.taxon	etymology	Etymology. The specific epithet applanatus derives from the Latin word “ applanatus ” and refers to the applanate penis guide.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F669FFF8FF7BF9AD0B9DFCDC.taxon	description	Chinese name: Ħ 氏 ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F669FFF8FF7BF9AD0B9DFCDC.taxon	diagnosis	Diagnosis. This species is similar to S. applanatus sp. nov., but can be distinguished from the later by its parameres about 1 / 3 as long as penis guide, and penis guide approximately parallel sided for basal 2 / 3 in ventral view (Figs 61 f – g). In S. applanatus, parameres are about 1 / 4 as long as penis guide. Penis guide is approximately parallel sided for basal 1 / 3 in ventral view, then slightly narrowing to blunt apex (Figs 60 f – g).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F669FFF8FF7BF9AD0B9DFCDC.taxon	description	Description. TL: 1.51 – 1.78 mm, TW: 1.06 – 1.29 mm, TH: 0.58 – 0.72 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 61 a – c). Body uniformly black, except antennae, maxillary palpi, tibiae and tarsi yellowish brown. Interocular distance slightly greater than eye width in frontal view. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 61 d). Male genitalia: Penis slender, penis capsule with long inner arm but without outer one, apex of penis pointed (Fig. 61 e). Penis guide long and slender in lateral view, widest at base, gradually narrowing to pointed apex; parameres about 1 / 3 as long as penis guide, with a tuft of long setae at apex; penis guide approximately parallel sided for basal 2 / 3 in ventral view, then tapering towards blunt apex (Figs 61 f – g). Female genitalia: Coxites shortened; styli absent; spermatheca worm-like, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F669FFF8FF7BF9AD0B9DFCDC.taxon	materials_examined	Material examined. China: Sichuan: 1 ♀, Mamize Nature Reserve, 2600 m, 19 – 20. IX. 2007, Wang XM leg; 4 ♂ 4 ♀, Heizhugou National Nature Reserve, 1900 m, 22 – 23. IX. 2007, Wang XM leg; 1 ♀, Liziping National Nature Reserve, 2000 m, 26 – 27. IX. 2007, Wang XM leg. Yunnan: 1 ♂ 1 ♀, Xujiaba, Ailaoshan National Nature Reserve, 1 – 3. X. 2006, Wang XM leg; 3 ♂, Xujiaba, Ailaoshan National Nature Reserve, 2524 m, 17. VIII. 2013, Chen XS leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F669FFF8FF7BF9AD0B9DFCDC.taxon	distribution	Distribution. China (Sichuan, Yunnan); Japan; Vietnam.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F669FFF8FF7BF9AD0B9DFCDC.taxon	discussion	Remarks. This species was recorded in Fujian, Henan, Hubei, Shaanxi and Taiwan by Yu (2011). However, it was described as “ yellowish brown head in male and black head in female ”, which is a unique characteristic of S. kuriharai. This characteristic is never reported in S. lewisi and not conforms our examined specimens. According to his description, these records are probably the misidentification of S. kuriharai. Thus, it is still questionable for the existence of this species in the above five provinces, and we tentatively exclude these provinces in the distribution range.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F66BFFFBFF7BFF750FD6FC24.taxon	description	Chinese name: OiAE ḧ 瓢 •	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F66BFFFBFF7BFF750FD6FC24.taxon	diagnosis	Diagnosis. This species is similar to S. lewisi (Kamiya, 1961), but can be distinguished from the later by its penis guide approximately parallel sided for basal 1 / 3 in ventral view (Fig. 62 g). In S. lewisi, penis guide is approximately parallel sided for basal 2 / 3 in ventral view, then tapering towards blunt apex. (Fig. 61 g)	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F66BFFFBFF7BFF750FD6FC24.taxon	description	Description. TL: 1.54 – 1.72 mm, TW: 1.05 – 1.19 mm, TH: 0.70 – 0.84 mm. Body oval, moderately convex, surfaces covered with short pubescence (Figs 62 a – c). Body uniformly black, except antennae, maxillary palpi and legs yellow to yellowish brown. Interocular distance slightly greater than eye width in frontal view. Abdominal postcoxal lines incomplete, not recurved toward the base of ventrite (Fig. 62 d). Male genitalia: Penis slender, penis capsule with long inner arm but without outer one, apex of penis pointed (Fig. 62 e). Penis guide approximately parallel sided in lateral view; parameres about 1 / 3 as long as penis guide, densely covered with long setae at apex; penis guide widest at base in ventral view, then slightly narrowing to apex (Figs 62 f – g). Female genitalia: Coxites shortened; styli absent; spermatheca worm-like, with basal ramus and nodulus.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F66BFFFBFF7BFF750FD6FC24.taxon	materials_examined	Type material. Holotype: China: Xizang: ♂, Lebu Town, Cuona County, 2400 m, 4. X. 2009, Wang XM leg (SCAU). Paratypes (SCAU): China: Xizang: 2 ♂ 1 ♀, Same data as holotype; 1 ♂, Lebu Town, Cuona County, 2450 m, 5. X. 2009, Wang XM leg; 2 ♂, Lebu Town, Cuona County, 2400 m, 24. X. 2011, Huo LZ leg; 1 ♂, Mama Town, Cuona County, 2800 m, 25. X. 2011, Huo LZ leg.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F66BFFFBFF7BFF750FD6FC24.taxon	distribution	Distribution. China (Xizang).	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
314287B0F66BFFFBFF7BFF750FD6FC24.taxon	etymology	Etymology. The name of this new species refers to county of its origin, Cuona County.	en	TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG, WANG, XINGMIN (2022): A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae). Zootaxa 5207 (1): 1-104, DOI: 10.11646/zootaxa.5207.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5207.1.1
