taxonID	type	description	language	source
312287F0B7070E1B8924E3D5FBFF4DED.taxon	description	Fig. 1	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7070E1B8924E3D5FBFF4DED.taxon	materials_examined	Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 3 ♂♂, 1 ♀; El Silencio, Laguna del Silencio; 10 ° 10 ′ 34.8 ″ N, 75 ° 44 ′ 26.6 ″ W; 24 Aug. 2017; C. M. López-Orozco, R. Borja-Arrieta and Y. M. Carpio-Díaz leg.; CUDC-CRU 141 • 3 ♂♂, 2 ♀♀; same collection data as for preceding; 10 ° 10 ′ 35 ″ N, 75 ° 44 ′ 26.4 ″ W; CUDC-CRU 142 • 1 juv.; same collection data as for preceding; 10 ° 10 ′ 35.4 ″ N, 75 ° 44 ′ 26.2 ″ W; CUDC-CRU 143 • 1 ♂, 1 ♀; same collection data as for preceding; 10 ° 10 ′ 34.3 ″ N, 75 ° 44 ′ 26.7 ″ W; CUDC-CRU 144 • 1 ♂; same locality and collectors as for preceding; 10 ° 10 ′ 32.9 ″ N, 75 ° 44 ′ 27.3 ″ W; 25 Aug. 2017; CUDC-CRU 145 • 1 ♀; same collection data as for preceding; 10 ° 10 ′ 32.3 ″ N, 75 ° 44 ′ 27.2 ″ W; CUDC-CRU 146 • 1 ♂; same collection data as for preceding; 10 ° 10 ′ 32.5 ″ N, 75 ° 44 ′ 27.1 ″ W; CUDC-CRU 147 • 1 ♀; Caño Ratón, Laguna Palmar; 10 ° 10 ′ 24.5 ″ N, 75 ° 44 ′ 39.7 ″ W; 24 Nov. 2017; same collectors as for preceding; CUDC-CRU 148 • 2 ♂♂, 1 ♀; El Silencio, Laguna del Silencio; 10 ° 10 ′ 35.4 ″ N, 75 ° 44 ′ 26 ″ W; 24 Nov. 2017; same collectors as for preceding; CUDC-CRU 149 • 1 ♀; same locality and collectors as for preceding; 10 ° 10 ′ 48 ″ N, 75 ° 43 ′ 39.3 ″ W; 25 Aug. 2017; CUDC-CRU 150 • 1 ♂; same collection data as for preceding; 10 ° 10 ′ 34 ″ N, 75 ° 44 ′ 26.8 ″ W; CUDC-CRU 151 • 1 juv.; same collection data as for preceding; 10 ° 10 ′ 47.74 ″ N, 75 ° 43 ′ 39.93 ″ W; CUDC-CRU 152 • 1 ♂, 1 ♀; same collection data as for preceding; 10 ° 10 ′ 35 ″ N, 75 ° 44 ′ 26.9 ″ W; ICN- CR-is 260 • 1 ♂, 2 ♀♀; Caño Ratón; 10 ° 10 ′ 44.91 ″ N, 75 ° 44 ′ 53.01 ″ W; 24 Nov. 2017, same collectors as for preceding; ICN-CR-is 261 • 1 juv.; Paraíso Secreto; 10 ° 10 ′ 15.9 ″ N, 75 ° 44 ′ 38.8 ″ W; 4 Apr. 2018, C. M. López-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 222. Previous records Valle del Cauca: Bahía Málaga (Pacific); Bolívar: Cartagena de Indias (Caribbean); Magdalena: Santa Marta (Ríos & Ramos 1990; Lazarus-Agudelo & Cantera-Kintz 2007; López-Orozco et al. 2014; Santamaria et al. 2014).	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7070E1B8924E3D5FBFF4DED.taxon	distribution	Distribution This species was originally described from Veracruz, Mexico (but see Discussion). Molecular phylogenetic analyses, however, revealed that what has been denominated as this species corresponds to a clade composed of highly divergent lineages (probably a complex of cryptic species), distributed in southern Florida, across the Caribbean Sea, Bahamas, Bermuda, and probably in northern Brazil, and the Pacific coast from Costa Rica, Panamá, Colombia, probably to Ecuador (Santamaria et al. 2014). Although this species has been reported in the Gulf of California and California, extensive sampling for phylogeographic studies conducted by some of us suggest that these previous records are likely misidentifications (Hurtado et al. 2010; Eberl et al. 2013; Hurtado et al. 2018).	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7060E148954E00CFB0A4971.taxon	description	Fig. 1	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7060E148954E00CFB0A4971.taxon	materials_examined	Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 3 ♂♂, 1 ♀; La Punta, Laguna Encantada; 10 ° 10 ′ 48.1 ″ N, 75 ° 43 ′ 37.5 ″ W; 24 Aug. 2017; Y. M. Carpio-Díaz, C. M. López- Orozco and R. Borja-Arrieta leg.; CUDC-CRU 153 • 4 ♂♂, 5 ♀♀; same locality and collectors as for preceding; 10 ° 10 ′ 48.4 ″ N, 75 ° 43 ′ 38.4 ″ W; 25 Aug. 2017; CUDC-CRU 154 • 2 ♂♂, 1 ♀; same collection data as for preceding; 10 ° 10 ′ 48.3 ″ N, 75 ° 43 ′ 38.9 ″ W; CUDC-CRU 155 • 5 ♂♂, 7 ♀♀; same collection data as for preceding; 10 ° 10 ′ 48.1 ″ N, 75 ° 43 ′ 39.1 ″ W; CUDC-CRU 156 • 4 ♂♂, 1 ♀; same collection data as for preceding; 10 ° 10 ′ 48 ″ N, 75 ° 43 ′ 39.3 ″ W; CUDC-CRU 157 • 5 ♂♂, 2 ♀♀; same collection data as for preceding; 10 ° 10 ′ 47.72 ″ N, 75 ° 43 ′ 40.09 ″ W; CUDC-CRU 158 • 2 ♂♂, 1 ♀; same collection data as for preceding; 10 ° 10 ′ 47.76 ″ N, 75 ° 43 ′ 40.27 ″ W; CUDC-CRU 159 • 5 ♂♂, 2 ♀♀; same collection data as for preceding; 10 ° 10 ′ 47.7 ″ N, 75 ° 43 ′ 39.93 ″ W; CUDC-CRU 160 • 9 ♂♂, 17 ♀♀; El Silencio, Laguna del Silencio; 10 ° 10 ′ 32.9 ″ N, 75 ° 44 ′ 27.3 ″ W; 24 Aug. 2017, same collectors as for preceding; CUDC-CRU 161 • 3 ♂♂, 5 ♀♀; same collection data as for preceding; 10 ° 10 ′ 35 ″ N, 75 ° 44 ′ 26.4 ″ W; CUDC-CRU 162 • 10 ♂♂, 6 ♀♀; same locality and collectors as for preceding; 10 ° 10 ′ 48.3 ″ N, 75 ° 43 ′ 38.9 ″ W; 25 Aug. 2017; CUDC-CRU 163 • 15 ♂♂, 12 ♀♀; La Punta, El Terminal; 10 ° 10 ′ 55.75 ″ N, 75 ° 43 ′ 36.57 ″ W; 6 Oct. 2017; C. M. López-Orozco leg.; ICN-CR-is 263 • 2 ♀♀; La Punta, Laguna Encantada; 10 ° 10 ′ 47.8 ″ N, 75 ° 43 ′ 39.5 ″ W; 25 Aug. 2017; Y. M. Carpio- Díaz, C. M. López-Orozco and R. Borja-Arrieta leg.; CUDC-CRU 164 • 2 ♂♂, 2 ♀♀; La Punta, Hotel Cocoliso; 10 ° 10 ′ 42.19 ″ N, 75 ° 43 ′ 32.88 ″ W; 6 Oct. 2017; same collectors as for preceding; CUDC- CRU 165 • 4 ♀♀; Hotel La Pola; 10 ° 10 ′ 19.4 ″ N, 75 ° 44 ′ 48.9 ″ W; 24 Nov. 2017; same collectors as for preceding; CUDC-CRU 166 • 1 ♂, 2 ♀♀; Caño Ratón, Laguna Palmar; 10 ° 10 ′ 24.5 ″ N, 75 ° 44 ′ 39.7 ″ W; same date and collectors as for preceding; CUDC-CRU 167 • 4 ♂♂, 5 ♀♀; Playa Libre; 10 ° 10 ′ 52.3 ″ N, 75 ° 43 ′ 55.7 ″ W; same date as for preceding; C. M. López-Orozco leg.; CUDC-CRU 168 • 3 ♂♂, 2 ♀♀; El Silencio, Laguna del Silencio; 10 ° 10 ′ 34.3 ″ N, 75 ° 44 ′ 26.7 ″ W; 24 Aug. 2017; C. M. López-Orozco, Y. M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC-CRU 169 • 1 ♀; El Silencio, Laguna del Silencio; 10 ° 10 ′ 35 ″ N, 75 ° 44 ′ 26.4 ″ W; 25 Aug. 2017; same collectors as for preceding; CUDC-CRU 170 • 3 ♂♂, 6 ♀♀, 1 juv.; Paraíso Secreto; 10 ° 10 ′ 15.9 ″ N, 75 ° 44 ′ 38.8 ″ W; 24 Nov. 2017; C. M. López-Orozco leg.; CUDC-CRU 171 • 1 ♂, 6 ♀♀; El Silencio, Laguna del Silencio; 10 ° 10 ′ 31.9 ″ N, 75 ° 44 ′ 26.9 ″ W; 7 Sep. 2017; same collector as for preceding; CUDC-CRU 172 • 4 ♂♂, 6 ♀♀; La Punta, Laguna Encantada; 10 ° 10 ′ 48.1 ″ N, 75 ° 43 ′ 37.5 ″ W; 7 Sep. 2017; C. M. López-Orozco, Y. M. Carpio-Díaz and R. Borja- Arrieta leg.; CUDC-CRU 173 • 9 ♂♂, 17 ♀♀; El Silencio, Mirador del Silencio; 10 ° 10 ′ 33.36 ″ N, 75 ° 44 ′ 24.14 ″ W; 5 Oct. 2017; same collectors as for preceding; CUDC-CRU 174 • 4 ♂♂, 6 ♀♀, 1 juv.; Caño Ratón; 10 ° 10 ′ 44.91 ″ N, 75 ° 44 ′ 53.01 ″ W; 24 Nov. 2017; C. M. López-Orozco leg.; CUDC- CRU 175 • 29 ♂♂, 38 ♀♀; Paraíso Secreto; 10 ° 10 ′ 15.9 ″ N, 75 ° 44 ′ 38.8 ″ W; 6 Sep. 2017; same collector as for preceding; CUDC-CRU 176 • 18 ♂♂, 17 ♀♀; El Silencio, Laguna Caracolí; 10 ° 10 ′ 28.4 ″ N, 75 ° 44 ′ 29.96 ″ W; 6 Oct. 2017; Y. M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC-CRU 177 • many ♂♂ and ♀♀; Caño Ratón; 10 ° 10 ′ 42.6 ″ N, 75 ° 44 ′ 52.6 ″ W; 24 Nov. 2017; C. M. López-Orozco leg.; CUDC- CRU 178 • 1 ♂; La Punta, Laguna Encantada; 10 ° 10 ′ 48.2 ″ N, 75 ° 43 ′ 38 ″ W; 24 Aug. 2017; Y. M. Carpio- Díaz, C. M. López-Orozco and R. Borja-Arrieta leg.; CUDC-CRU 179 • 5 ♂♂, 8 ♀♀; El Silencio, Laguna del Silencio; 10 ° 10 ′ 35.4 ″ N, 75 ° 44 ′ 26 ″ W; same date and collectors as for preceding; CUDC- CRU 180 • 4 ♂♂, 6 ♀♀; same collection data as for preceding; 10 ° 10 ′ 34.8 ″ N, 75 ° 44 ′ 26.6 ″ W; CUDC- CRU 181 • 10 ♂♂, 5 ♀♀; 10 ° 10 ′ 32.3 ″ N, 75 ° 44 ′ 27.2 ″ W; 25 Aug. 2017; C. M. López-Orozco leg.; CUDC-CRU 182 • 19 ♂♂, 21 ♀♀; La Punta, Laguna Encantada; 10 ° 10 ′ 47.72 ″ N, 75 ° 43 ′ 40.09 ″ W; 25 Aug. 2017; Y. M. Carpio-Díaz, C. M. López-Orozco and R. Borja-Arrieta leg.; CUDC-CRU 183 • 14 ♂♂, 7 ♀♀; Paraíso Secreto; 10 ° 10 ′ 15.9 ″ N, 75 ° 44 ′ 38.8 ″ W; 4 Apr. 2018; C. M. López-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 184 • 10 ♂♂, 17 ♀♀; El Silencio, Laguna del Silencio; 10 ° 10 ′ 35.4 ″ N, 75 ° 44 ′ 26.2 ″ W; 24 Aug. 2017; Y. M. Carpio-Díaz, C. M. López-Orozco and R. Borja- Arrieta leg.; ICN-CR-is 264. Previous records Bolívar: Barú, Cartagena de Indias (Caribbean) (Carpio-Díaz et al. 2016).	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7060E148954E00CFB0A4971.taxon	distribution	Distribution Widely distributed in Atlantic coastal areas of Mexico (Kensley & Schotte 1989); Florida, Cuba (Van Name 1936); Bahamas, Tobago, Bonaire (Schultz 1974); Dominica, Curacao, Virgin Islands, Belize, Venezuela (Kensley & Schotte 1989); Puerto Rico (Hurtado et al. 2014); Colombia (Carpio-Díaz et al. 2016); and Brazil (Silva & Alves 2000). It is also present on the Pacific coast of Costa Rica (Taiti et al. 2018). First record of the species for continental islands of the Colombian Caribbean.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7090E148931E76DFEC14B8A.taxon	description	Figs 1, 2 A	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7090E148931E76DFEC14B8A.taxon	materials_examined	Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 1 ♀; Playa Libre; 10 ° 10 ′ 52.4 ″ N, 75 ° 43 ′ 56.1 ″ W; 7 Sep. 2017; C. M. López-Orozco, R. Borja-Arrieta and Y. M. Carpio- Díaz leg.; CUDC-CRU 136 • 11 ♂♂, 39 ♀♀, 1 juv.; La Punta, Hotel Majagua; 10 ° 10 ′ 56.41 ″ N, 75 ° 43 ′ 33.51 ″ W; 6 Oct. 2017; same collectors as for preceding; CUDC-CRU 137.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7090E148931E76DFEC14B8A.taxon	distribution	Distribution Florida, Bahamas, Leeward Islands, Belize, Venezuela and Cuba (Schultz 1974; Kensley & Schotte 1989; Schmalfuss & Vergara 2000; Schmidt 2001; Schmalfuss 2003; Hurtado et al. 2014). First record for Colombia.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B70B0E108A86E337FBAC4F58.taxon	description	urn: lsid: zoobank. org: act: DFCDBD 2 C-AEBD- 4754 - 951 A- 1 D 927 E 81 E 6 DC Figs 1, 2 B, 3 – 5	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B70B0E108A86E337FBAC4F58.taxon	etymology	Etymology The new species is named after Mr Hernando Gómez Molina, also known as Mr Negro, for his efforts to conserve the biodiversity of the Rosario Islands.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B70B0E108A86E337FBAC4F58.taxon	materials_examined	Material examined Holotype COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • ♂; Playa Libre; 10 ° 10 ′ 52.3 ″ N, 75 ° 43 ′ 54.8 ″ W; 5 Oct. 2017; C. M. López-Orozco, R. Borja-Arrieta and Y. M. Carpio- Díaz leg.; CUDC-CRU 127. Paratypes COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 1 ♂ (parts in micropreparations); same collection data as for holotype; CUDC-CRU 128 • 2 ♀♀ (parts in micropreparations); same collection data as for holotype; CUDC-CRU 129 • many ♂♂ and ♀♀; same collection data as for holotype; CUDC-CRU 130 • 7 ♂♂, 13 ♀♀; El Silencio, Playa La Cocotera; 10 ° 10 ′ 49.3 ″ N, 75 ° 44 ′ 16.7 ″ W; 6 Oct. 2017; same collectors as for holotype; CUDC-CRU 131 • 8 ♂♂, 10 ♀♀; La Punta, El Terminal; 10 ° 10 ′ 55.75 ″ N, 75 ° 43 ′ 36.57 ″ W; same collectors as for holotype; CUDC-CRU 132 • 5 ♀♀; Playa Libre; 10 ° 10 ′ 52.3 ″ N, 75 ° 43 ′ 55.7 ″ W; 7 Sep. 2017; same collectors as for holotype; CUDC-CRU 133 • 1 ♀; same collection data as for holotype; CUDC-CRU 134 • 4 ♂♂, 3 ♀♀; same collection data as for holotype; 5 Oct. 2017; ICN-CR-is 262 • 1 ♂; La Punta, Hotel Cocoliso; 10 ° 10 ′ 42.19 ″ N, 75 ° 43 ′ 32.88 ″ W; 6 Oct. 2017; same collectors as for holotype; CUDC-CRU 135.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B70B0E108A86E337FBAC4F58.taxon	description	Description MEASUREMENTS. Maximum body length: male 10 mm; female 8.1 mm. BODY. Colour yellowish with many irregular dark spots (Fig. 2 B). Body outline as in Fig. 3 A. Endoantennal conglobation. Dorsum covered with lanceolate scale-setae (Fig. 3 B). CEPHALON. Frontal lamina triangular (Fig. 3 C – D); eyes consisting of 26 – 27 ommatidia. PEREON. Pereonite 1 epimera grooved on lateral margins, schisma with inner lobe triangular, slightly surpassing outer lobe (Fig. 3 E); pereonite 2 with ventral lobe triangular, shorter than outer lobe (Fig. 3 E); epimera of pereonites 2 – 4 subtriangular, 5 – 7 subquadrangular (Fig. 3 A). PLEON. Epimera of pleonites 3 – 5 with rounded distal margin (Fig. 3 F). Ventral phylacomera present on pleonites 3 – 5; phylacomera 5 oar-shaped, with wide distal part, medial margin oblique and slightly sinuous (Fig. 3 G). Telson rectangular, posterior margin slightly folded upwards (Fig. 3 F). ANTENNULA. Composed of one article, flattened and triangular, several aesthetascs inserted on slightly rounded lobe (Fig. 3 H). ANTENNA. Short and thickset, flagellum of three articles, apical article as long as first and second articles (Fig. 4 A). MOUTH. Mandibles with strong molar process, left mandible (Fig 4 B) with two penicils on molar process, two on hairy lobe and five free penicils; right mandible (Fig. 4 C) with two penicils on molar process, one on hairy lobe and five free penicils. Maxillula (Fig. 4 D) inner endite with three long penicils densely covered with thin setae, apical penicil longest; outer endite composed of 5 + 5 simple teeth plus one sensory seta, inner set bearing two teeth slightly serrate, median portion bearing three short setae. Maxilla (Fig. 4 E) of two lobes, inner lobe rounded, covered with thick setae, outer lobe covered with thin setae. Maxilliped (Fig. 4 F) basis with rounded outer margin, palp proximal article triangular without setae; endite rectangular, bearing five apical and two subapical penicils. UROPOD. As in Fig 3 G. PEREOPODS. Pereopod 1 (Fig. 5 A) basis with triangular latero-apical process. Pereopod 7 (Fig. 5 B) propodus slightly inflated; dactylus of one claw, dactylar seta stout, not surpassing claw and bearing several small sensory setae on median to apical portion. Male PEREOPODS 1 AND 7. Without any sexual dimorphism (Fig. 5 A – B). PLEOPODS. Pleopod 2 exopod rectangular (Fig. 5 C) with respiratory folds; endopod with distal portion slightly directed inwards, apex covered with cleft scales of 2 – 4 branches (Fig. 5 C).	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B70B0E108A86E337FBAC4F58.taxon	discussion	Remarks Presently, the genus Tylos has a worldwide distribution and comprises 21 coastal species (Schmalfuss & Vergara 2000; Schmalfuss 2003). In Colombia, only T. niveus Budde-Lund, 1885 was recorded from the Caribbean region (Carpio-Díaz et al. 2016). Tylos negroi sp. nov. can be distinguished from all species of the genus by the shape of the ventral phylacomera 5. Moreover, it differs from T. niveus in the pereonite 1 epimeron having the inner lobe of the schisma surpassing the outer lobe (vs inner lobe not surpassing outer lobe) and the pereopod 7 propodus being inflated (vs not inflated).	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B70B0E108A86E337FBAC4F58.taxon	distribution	Distribution Presently known only from Isla Grande, Cartagena de Indias, Colombia.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B70C0E0E8A99E424FBAC4F5B.taxon	description	urn: lsid: zoobank. org: act: 045 F 7904 - E 6 DE- 4912 - 9073 - 1473 A 522 FAC 7 Figs 1, 2 C, 6 – 8	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B70C0E0E8A99E424FBAC4F5B.taxon	etymology	Etymology The new species is named after Dr Nestor Hernando Campos, for his contribution to the knowledge of Crustacea from Colombia.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B70C0E0E8A99E424FBAC4F5B.taxon	materials_examined	Material examined Holotype COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • ♂ (parts in micropreparations); La Punta, Laguna Encantada; 10 ° 10 ′ 48.1 ″ N, 75 ° 43 ′ 37.5 ″ W; 7 Sep. 2017; C. M. López-Orozco, R. Borja-Arrieta and Y. M. Carpio-Díaz leg.; CUDC-CRU 185. Paratypes COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 1 ♂, 1 ♀ (parts in micropreparations); same collection data as for holotype; CUDC-CRU 186 • 2 ♀♀; same collection data as for holotype; CUDC-CRU 187 • 1 ♀; same locality and collectors as for holotype; 24 Aug. 2017; CUDC-CRU 188.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B70C0E0E8A99E424FBAC4F5B.taxon	description	Description MEASUREMENTS. Maximum body length: male 1.8 mm; female 2.4 mm. BODY. Body pigments absent (Fig. 2 C). Body (Fig. 6 A – C) slender, about four times as long as wide, parallel sides. Dorsal surface bearing ribs and tubercles (Fig. 6 A – C, E – G), as follows: cephalon with two tubercles on anterior portion and three on posterior portion; pereonite 1 with two transversal ribs; pereonites 1 – 4 with three longitudinal ribs (middle rib split in two on pereonites 2 and 3), one lateral tubercle and one tubercle between two lateral ribs per side; pereonites 5 – 7 with two longitudinal ribs and one lateral tubercle per side; pleonites 2 – 5 and telson with two paramedian tubercles. Dorsum bearing rectangular scale-setae with fringed apex (Fig. 6 D). CEPHALON. Sub-rectangular (Fig. 6 E – F), median lobe triangular, lateral lobes well-developed, directed frontwards, not surpassing median lobe; eyes absent. PEREON. Epimera of pereonites 2 – 7 laterally grooved (Fig. 6 B). Pereonites 4 – 5 ventrally with lateral sternal thickenings small, triangular; pereonites 6 and 7 ventrally with sub-rectangular sternal plates (Fig. 6 C). PLEON. Pleonite 2 dorsally visible, 3 – 5 with rectangular epimera (Fig. 6 A – C). Telson (Fig. 6 G) trapezoidal, distal margin broadly rounded. ANTENNULA. Composed of two articles (Fig. 6 H), proximal article stout with shield-like projection on distal margin, distal article comma-shaped bearing two distal aesthetascs. ANTENNA. Short (Fig. 6 I), peduncle articles more or less subequal in length; flagellum of two articles, distal article twice as long as proximal article, bearing two lateral aesthetascs. MOUTH. Mandibles with molar penicil consisting of two plumose setae, left mandible (Fig. 7 A) with two penicils on incisor process, right mandible (Fig. 7 B) with one penicil. Maxillula (Fig. 7 C) inner endite with two short apical penicils; outer endite bearing 4 + 4 teeth, inner set with two teeth apically cleft. Maxilla (Fig. 7 D) of two rounded lobes, inner lobe bearing three thick setae. Maxilliped (Fig. 7 E) basis with sparse setae; endite stout, distal margin rounded, bearing thin setae, one large penicil inserted near medial margin. UROPOD. Protopod stout and conical, exopod short, bearing two long apical setae, endopod inserted proximally (Fig. 7 F). PEREOPODS. Pereopods 1 – 7 short, stout, merus to propodus bearing sparse setae on sternal margins; pereopod 1 carpus with distal seta fringed at apex; dactylus of two claws, inner claw very short, ungual seta simple, not surpassing outer claw, dactylar seta stout, reaching distal margin of outer claw, apex bearing many small setae. Male PEREOPODS 1 AND 7. Without sexual dimorphism (Fig. 8 A – B). PLEOPODS. Pleopod 1 exopod absent; endopod (Fig. 8 C) enlarged on proximal portion, distal portion tapering. Pleopod 2 (Fig. 8 D) exopod triangular, inner margin grooved, bearing one strong seta; endopod about twice as long as exopod. Pleopod 3 and 4 exopods (Fig. 8 E – F) sub-quadrangular, inner margin grooved, distal margin slightly serrate. Pleopod 5 exopod (Fig. 8 G) sub-rectangular, inner margin slightly serrate, distal margin straight, bearing one seta.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B70C0E0E8A99E424FBAC4F5B.taxon	discussion	Remarks Presently, Stenoniscus comprises four species (Schmalfuss 2003): S. pleonalis Aubert & Dollfus, 1890 from the northern coasts of the Mediterranean east to the Aegean, Bermuda, Bulgaria, Madeira and Mexico; S. carinatus Silvestri, 1897 from coastal areas of Croatia, Italy, Portugal and Spain; S. aenariensis (Verhoeff, 1942) and S. plutonis (Verhoeff, 1942) from Ischia Island, Italy, both most probably junior synonyms of S. pleonalis according to the short descriptions by Verhoeff (1942). The genus can be defined by animals of reduced size (up to 3.5 mm), body slender with lateral sides parallel, dorsum bearing longitudinal ribs and tubercles, sometimes reduced, epimera of pereonites 2 – 7 with lateral margins grooved, pereonites 4 – 5 ventrally with lateral sternal thickenings and 6 – 7 with sternal plates, antennula of two articles, pleonite 1 absent, uropods of conical shape, pleopod exopods without respiratory organs, and pleopod 1 absent on females (see Vandel 1962; Schmidt 2003). Stenoniscus nestori López-Orozco, Taiti & Campos-Filho sp. nov. is morphologically similar to Stenoniscus carinatus, from which it differs mainly in the number and disposition of the dorsal ribs and tubercles on the cephalon and pereonites (compare Fig. 6 A – B with Schmidt 2003: fig. 67 and Taiti & Ferrara 1982: fig. 6 a for S. carinatus). The new species is readily distinct from Stenoniscus pleonalis in having a strong dorsal ornamentation (vs very weak) and lacking the tomentose appearance due to long dorsal scale-setae (see Vandel 1944: fig. 1 a – c).	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B70C0E0E8A99E424FBAC4F5B.taxon	distribution	Distribution Presently known only from Isla Grande, Cartagena de Indias, Colombia.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7130E0E8932E15FFE1849BA.taxon	discussion	Remarks The family Detonidae Budde-Lund, 1904 comprises 39 littoral species in the genera Armadilloniscus Uljanin, 1875 (32 spp.), Deto Guérin-Méneville, 1836 (5 spp.) and Detonella Lohmander, 1927 (2 spp.), including two dubious species, Armadilloniscus minutus Uljanin, 1875 and Deto spinicornis Brandt, 1851 (Schultz 1984; Paoletti & Stinner 1989; Taiti & Ferrara 1989; Schmidt 2002; Schmalfuss 2003; Schmidt & Leistikow 2004; WoRMS 2020 b). The family can be characterized by animals having a clinger habitus (sensu Schmalfuss 1984), cephalon with well-developed lateral lobes, dorsal surface more or less tuberculated or with longitudinal ridges, exopods of pleopods 1 – 5 without respiratory structures and lateral margins covered with thin setae conferring a fringe-like appearance (see Taiti & Ferrara 1989; Schmidt 2002). Recently, Reinoso-Flórez et al. (2016) recorded the family from Páramo de Estambul (Tolima), Colombia. However, after the re-examination of the material deposited at Colección Zoológica de la Universidad del Tolima (CZUT-IT 1423 – 1426, see Reinoso-Flórez et al. 2016), the specimens belong to the families Styloniscidae and Philosciidae, and therefore, our work constitutes the first record of the family in the country.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7130E0F8AE8E47FFC604EB4.taxon	description	Figs 1, 2 D, 9	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7130E0F8AE8E47FFC604EB4.taxon	materials_examined	Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 2 ♀♀; Paraíso Secreto; 10 ° 10 ′ 15.9 ″ N, 75 ° 44 ′ 38.8 ″ W; 4 Apr. 2018; C. M. López-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 189 • 4 ♂♂, 10 ♀♀; same locality as for preceding; 6 Sep. 2017; C. M. López-Orozco, Y. M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC-CRU 190 • 1 ♀; Caño Ratón; 10 ° 10 ′ 33.4 ″ N, 75 ° 44 ′ 52.4 ″ W; 24 Nov. 2017; same collectors as for preceding; CUDC-CRU 191 • 2 ♀♀; 10 ° 10 ′ 42.6 ″ N, 75 ° 44 ′ 52.6 ″ W; same collection data as for preceding; CUDC-CRU 192.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7130E0F8AE8E47FFC604EB4.taxon	discussion	Remarks Paoletti & Stinner (1989) described A. caraibicus from Parque Marrocoy, Falcón State, Venezuela. In the description, no sexual dimorphism was noted in the dorsal ornamentation. However, the male specimens show four dorsal tubercles on pleonite 2, while females have only two (see Fig. 9 and Paoletti & Stinner 1989: fig. 11).	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7130E0F8AE8E47FFC604EB4.taxon	distribution	Distribution Venezuela: Coralline cays of the Parque Natural Morrocoy (Paoletti & Stinner 1989), and north coast of the Península de Paria (Schmidt 2001). First record for Colombia.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7150E058A9AE337FBAC4D32.taxon	description	urn: lsid: zoobank. org: act: 88600 E 5 C-BFF 9 - 49 BA- 976 C- 8853 E 33 A 6 E 58 Figs 1, 2 E, 10 – 12	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7150E058A9AE337FBAC4D32.taxon	etymology	Etymology The new species is named after Mr Luis Felipe Arrieta Zúñiga, for his interest in nature and support to Ricardo Borja-Arrieta as a biologist.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7150E058A9AE337FBAC4D32.taxon	materials_examined	Material examined Holotype COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • ♂; Laguna del Silencio, El Silencio; 10 ° 10 ′ 35.4 ″ N, 75 ° 44 ′ 26 ″ W; 24 Aug. 2017; C. M. López-Orozco, Y. M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC-CRU 193. Paratypes COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 1 ♂, 1 ♀ (parts in micropreparations); same collection data as for holotype; CUDC-CRU 194 • 10 ♂♂, 5 ♀♀; same collection data as for holotype; CUDC-CRU 195 • 1 ♀; 10 ° 10 ′ 32.9 ″ N, 75 ° 44 ′ 27.3 ″ W; same date and collectors as for holotype; CUDC-CRU 206 • 1 ♀; 10 ° 10 ′ 33.5 ″ N, 75 ° 44 ′ 26.9 ″ W; same date and collectors as for holotype; CUDC-CRU 207 • 1 ♂; 10 ° 10 ′ 35 ″ N, 75 ° 44 ′ 26.4 ″ W; same date and collectors as for holotype; CUDC-CRU 208 • 1 ♂; 10 ° 10 ′ 34.8 ″ N, 75 ° 44 ′ 26.6 ″ W; same date and collectors as for holotype; CUDC-CRU 209 • 6 ♂♂, 12 ♀♀; same collection data as for holotype; 10 ° 10 ′ 35 ″ N, 75 ° 44 ′ 16.4 ″ W; CUDC-CRU 210 • 14 ♂♂, 10 ♀♀; 10 ° 10 ′ 34 ″ N, 75 ° 44 ′ 26.8 ″ W; 25 Aug. 2017; same locality and collectors as for holotype; CUDC-CRU 211 • 3 ♂♂; La Punta, Laguna Encantada; 10 ° 10 ′ 48.4 ″ N, 75 ° 43 ′ 38.4 ″ W; 25 Aug. 2017; C. M. López-Orozco, Y. M. Carpio-Díaz and R. Borja- Arrieta leg.; CUDC-CRU 196 • 1 ♂; same locality as for preceding; 10 ° 10 ′ 47.72 ″ N, 75 ° 43 ′ 40.09 ″ W; same date and collectors as for holotype; CUDC-CRU 197 • 1 ♂, 1 ♀; same locality as for preceding; 10 ° 10 ′ 48 ″ N, 75 ° 43 ′ 39.3 ″ W; same date and collectors as for holotype; CUDC-CRU 198 • 1 ♀; same locality as for preceding; 10 ° 10 ′ 47.7 ″ N, 75 ° 43 ′ 39.93 ″ W; same date and collectors as for holotype; CUDC-CRU 199 • 2 ♂♂, 3 ♀♀; same locality as for preceding; 10 ° 10 ′ 48.3 ″ N, 75 ° 43 ′ 38.9 ″ W; same date and collectors as for holotype; CUDC-CRU 200 • 3 ♀♀; same locality as for preceding; 10 ° 10 ′ 48.1 ″ N, 75 ° 43 ′ 39.1 ″ W; same date and collectors as for holotype; CUDC-CRU 201 • 2 ♀♀; same locality as for preceding; 10 ° 10 ′ 48.1 ″ N, 75 ° 43 ′ 37.5 ″ W; same date and collectors as for holotype; CUDC-CRU 202 • 4 ♂♂, 4 ♀♀; same locality as for preceding; 10 ° 10 ′ 47.72 ″ N, 75 ° 43 ′ 40.09 ″ W; same date and collectors as for holotype; CUDC-CRU 203 • 2 ♂♂, 2 ♀♀; same locality as for preceding; 7 Sep. 2017; same date and collectors as for holotype; CUDC-CRU 204 • 3 ♂♂, 4 ♀♀; Laguna La Charca; 10 ° 10 ′ 47.8 ″ N, 75 ° 44 ′ 0.4 ″ W; 7 Sep. 2017; C. M. López-Orozco, Y. M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC- CRU 205.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7150E058A9AE337FBAC4D32.taxon	description	Description MEASUREMENTS. Maximum length: male 2.5 mm; female 3.1 mm. BODY. Color brown with usual pale muscle spots; antennae brown with pale flagellum (Fig. 2 E). Body elongated and elliptical-shaped (Fig. 10 A, H). Dorsal surface bearing elongated scale-setae (Fig. 10 B). Dorsum covered with broad tubercles (Fig. 10 A, C – E, H): cephalon bearing seven tubercles in two rows, anterior row with four and posterior row with three; ♂ pereonites 1 – 3 with one row of 10 tubercles, 4 – 7 with one row of eight tubercles; ♀ pereonites 1, 5 and 6 with one row of 10 tubercles, 2 – 4 and 7 with one row of eight tubercles; ♂ pleonites 1 – 5 and telson with two paramedian tubercles, pleonite 2 with two broad tubercles at sides, pleonite 3 with two small additional tubercles; ♀ pleonites 1 – 5 and telson with two paramedian tubercles, pleonites 2 and 3 with two additional small tubercles at sides. CEPHALON. Lateral lobes well developed and directed outwards, median lobe triangular, slightly surpassing distal margin of lateral lobes, frontal and suprantennal lines absent; eyes consisting of four ommatidia (Fig. 10 C – D). PLEON. Outline continuous with that of pereonite 7, epimera of pleonites 3 – 5 rectangular (Fig. 10 A, E, H). Telson (Fig. 10 E) triangular, lateral margins almost straight, distal part truncate and not surpassing distal margin of pleonite 5 epimera. ANTENNULA. Composed of two articles, second article bearing a few lateral setae, distal margin bearing one flagellar seta and two aesthetascs (Fig. 10 F). ANTENNA. When extended posteriorly slightly surpassing pereonite 1, flagellum of four articles (Fig. 10 G). MOUTH. Mandibles with molar penicil of several plumose setae, left mandible with 2 + 1 penicils (Fig. 11 A), right with 1 + 1 penicils (Fig. 11 B). Maxillula (Fig. 11 C) inner endite with two penicils; outer endite of 4 + 8 teeth, five of them pectinated, and one slender stalk between outer and inner set, outer distal margin bearing fringe of long thin setae. Maxilla (Fig. 11 D) with lobes fused, inner middle portion with five long setae, distal margin rounded, bearing several thin and thick setae. Maxilliped (Fig. 11 E) palp bearing two tufts of setae on first article, distal articles of palp fused, bearing several thin and thick setae; endite rectangular, inner margin bearing fringe of long thin setae plus four long thick setae, distal margin bilobed, bearing broad rounded penicil with long thick setae. UROPOD. Protopod enlarged with distal margin rounded, exopod not surpassing protopod, endopod longer than exopod (Fig. 11 F). PEREOPODS. Pereopods 1 – 7 stout, merus to propodus bearing sparse setae on sternal margin, carpus 1 with strong distal seta bearing long sensilla, dactylus with very long ungual seta, dactylar seta elongated, apically cleft and plumose. Male PEREOPODS 1 AND 7. Without any sexual modifications (Fig. 12 A – B). GENITAL PAPILLA. Ventral shield triangular, papilla slightly cleft at apex, bearing small setae (Fig. 12 C). PLEOPODS. Pleopod 1 (Fig. 12 D) exopod ovoid; endopod three times as long as exopod, apical portion bent outwards. Pleopod 2 (Fig. 12 E) exopod subtriangular; endopod with distal article flagelliform. Exopods of pleopods 3 – 5 as in Fig. 12 F – H.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7150E058A9AE337FBAC4D32.taxon	discussion	Remarks In the Caribbean region, four species of Armadilloniscus have been recorded: A. caraibicus from Venezuela, A. ellipticus (Harger, 1878) from the Atlantic coasts of USA and Bermuda, A. ninae Schultz, 1984 from Belize, and A. steptus Schotte & Heard, 1991 from Turks and Caicos Islands (British West Indies) (Schultz 1984; Paoletti & Stinner 1989; Taiti & Ferrara 1989; Schotte & Heard 1991). For a definition of the genus see Arcangeli (1957), Vandel (1962), Taiti & Ferrara (1989), and Schmidt (2002). Armadilloniscus luisi Carpio-Díaz, Taiti & Campos-Filho sp. nov. is readily distinguished from all the other species of the genus in the number and arrangement of the dorsal tubercles on the cephalon, pereon and pleon. Moreover, it can be distinguished from the other four Caribbean species in having two broad tubercles on pleonite 2 (vs small in A. caraibicus and A. steptus, absent in A. ninae), the cephalon with the median lobe triangular (vs rounded in A. caraibicus) and lateral lobes subquadrangular (vs subtriangular in A. ninae and A. steptus), eyes composed of four ommatidia (vs 4 – 5 in A. caraibicus, 6 in A. ellipticus and 5 in A. ninae), telson with distal margin truncate (vs rounded in A. caraibicus and A. ellipticus, triangular in A. ninae), and ♂ pleopod 1 exopod ovoid (vs subtriangular in A. caraibicus, A. ellipticus and A. ninae, almost rounded in A. steptus) (Paoletti & Stinner 1989; Schultz 1984; Schotte & Heard 1991; Garthwaite et al. 1992; Schmidt 2002).	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7150E058A9AE337FBAC4D32.taxon	distribution	Distribution Presently known only from Isla Grande, Cartagena de Indias, Colombia.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7180E008934E3B1FDC54F92.taxon	description	Figs 1, 2 F, 13 – 14	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7180E008934E3B1FDC54F92.taxon	materials_examined	Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 1 ♂ (parts in micropreparations); Paraíso Secreto; 10 ° 10 ′ 15.9 ″ N, 75 ° 44 ′ 38.8 ″ W; 24 Nov. 2017; C. M. López- Orozco leg.; CUDC-CRU 212 • 1 ♂, 4 ♀♀; same collection data as for preceding; CUDC-CRU 214 • 21 ♂♂, 61 ♀♀; same locality as for preceding; 4 Apr. 2018; C. M. López-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 213 • 3 ♀♀; same locality as for preceding; 6 Sep. 2017; C. M. López- Orozco leg.; CUDC-CRU 215 • 1 ♂, 6 ♀♀ (one with parts in micropreparations); Caño Ratón; 10 ° 10 ′ 33.4 ″ N, 75 ° 44 ′ 52.4 ″ W; 24 Nov. 2017; C. M. López-Orozco leg.; CUDC-CRU 216 • 5 ♀♀; 10 ° 10 ′ 44.91 ″ N, 75 ° 44 ′ 53.01 ″ W; same collection data as for preceding; CUDC-CRU 217. CUBA – Cayo Piedras del Norte, Varadero • 2 ♂♂, 2 ♀♀; 8 Aug. 1985; A. Poggesi leg.; MZUF 1824.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7180E008934E3B1FDC54F92.taxon	description	Redescription BODY. Color pale brown; cephalon, antennae, epimera of pereonites 1 – 7, epimera of pleonites 1 – 5 and uropod protopods more pigmented; pereon and pleon with median and paramedian region more depigmented (Fig. 2 F). Body elongated and elliptical-shaped (Fig. 13 A). Dorsal surface bearing elongated scale-setae (Fig. 13 B). Cephalon and pereon covered with small tubercles (Fig. 13 A, C – D): cephalon bearing six tubercles in two rows, anterior row with two and posterior row with four; pereonites 1 – 7 with one row of 8 tubercles. CEPHALON. Lateral lobes well developed and directed outwards, median lobe triangular, slightly surpassing distal margin of lateral lobes, frontal and suprantennal lines absent; eyes consisting of 5 ommatidia (Fig. 13 C – D). PLEON. Outline continuous with that of pereonite 7, epimera of pleonites 3 – 5 rectangular (Fig. 13 A, E). Telson (Fig. 13 E) triangular, wider than long, lateral margins almost straight, broadly rounded apex. ANTENNULA. Composed of two articles, second article bearing many lateral setae, distal margin bearing one flagellar seta and two aesthetascs (Fig. 13 F). ANTENNA. When extended posteriorly slightly surpassing pereonite 1, flagellum of four articles (Fig. 13 G). BUCCAL PIECES. As in Armadilloniscus luisi sp. nov. UROPOD. Protopod enlarged with distal margin slightly rounded, exopod not surpassing protopod, endopod longer than exopod (Fig. 13 E). PEREOPODS. Pereopods 1 – 7 stout, merus to propodus bearing sparse setae on sternal margin, carpus 1 with strong distal seta bearing long sensilla, dactylus with elongated and digitform ungual seta, dactylar seta elongated, apically cleft and plumose. Male PEREOPODS 1 AND 7. Without any sexual modifications (Fig. 14 A – B). GENITAL PAPILLA. Ventral shield stout triangular, papilla rounded at apex, bearing small setae (Fig. 14 C). PLEOPODS. Pleopod 1 (Fig. 14 D) exopod ovoid, wider than long; endopod stout, three times as long as exopod, apical portion slightly bent inwards. Pleopod 2 (Fig. 14 E) exopod ovoid, wider than long; endopod with flagelliform distal article. Exopods of pleopods 3 – 5 as in Fig. 14 F – H.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7180E008934E3B1FDC54F92.taxon	discussion	Remarks Schultz (1984) described A. ninae from San Pedro beach, Ambergris Cay, Belize. Comparing Shultz’s description with the specimens examined here, it was possible to observe that almost all characters mentioned, including the pale dorsal pigmentation, are quite similar. However, Schultz (1984) mentioned that A. ninae has eyes composed of 14 ommatidia and antennula with three articles, and herein the specimens showed the eyes composed of 5 ommatidia and antennula of two articles. Most probably, the author misinterpreted the composition of the eyes of this species, since the illustrations of the cephalon clearly show a smaller number. The same statement can be applied to the antennula. In general, within Oniscidea, the composition of the antennula does not vary within the genera, with just few exceptions (see Schmidt 2002, 2003). Moreover, analyzing other representatives of Armadilloniscus, it is possible to observe that the antennula is always composed of two articles (see Taiti & Ferrara 1989; Kwon & Wang 1996). Therefore, we identify our specimens as A. ninae.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7180E008934E3B1FDC54F92.taxon	distribution	Distribution This species was previously recorded only from San Pedro beach, Ambergris Cay, Belize (Schultz 1984). First record for Colombia and Cuba.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B71D0E008921E62AFD55498E.taxon	discussion	Remarks The family Halophilosciidae comprises 34 species distributed in three genera, Halophiloscia Verhoeff, 1908 (9 spp.), Littorophiloscia Hatch, 1947 (22 spp.), and Stenophiloscia Verhoeff, 1908 (3 spp.) (WoRMS 2020 c). In Colombia, Pearse (1915) recorded Philoscia culebrae Moore, 1901 [= Littorophiloscia c.] in coffee plantations from Hacienda Cincinati and Hacienda La Rosa, Sierra Nevada de Santa Marta (ca 1400 m a. s. l.), which is considered to be a misidentification by Van Name (1936). Littorophiloscia culebrae (Moore, 1901) is recorded from coastal areas of Angola, Brazil, Canary Islands, Caribbean islands, Florida (USA), Hawaii, Iraq, Madagascar, Socotra Archipelago and Yemen (Schmalfuss 2003; Taiti & Ferrara 2004; Araujo & Taiti 2007; Taiti & Lopez 2008; Naser et al. 2015; Lisboa et al. 2017; Campos-Filho et al. 2018). Recent expeditions to the north coast of Bolívar revealed the existence of this species in Colombia, inhabiting the coastal areas of the Barú Peninsula, Cartagena de Indias (C. López-Orozco & Y. Carpio-Díaz, pers. obs.).	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B71D0E038ACFE4AFFD4E4B4A.taxon	description	Figs 1, 2 G, 15 – 17	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B71D0E038ACFE4AFFD4E4B4A.taxon	materials_examined	Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 3 ♂♂; Paraíso Secreto; 10 ° 10 ′ 15.9 ″ N, 75 ° 44 ′ 38.8 ″ W; 4 Apr. 2018; C. M. López-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 218, 220, 221 • 1 ♂, 1 ♀ (parts in micropreparations); same locality as for preceding; 6 Sep. 2017; C. M. López-Orozco, Y. M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC-CRU 219. CUBA – Matanzas, Bahia de Conchinos • 4 ♂♂, 10 ♀♀; near Cueva de los Peces, between Playa Larga and Playa Giron; 10 Apr. 2002; S. Taiti leg.; MZUF 9607. – Pinar del Rio • 2 ♂♂, 3 ♀♀; Penisula Guanahacabibes, La Bajada; 7 Apr. 2002; S. Taiti leg.; MZUF 9608.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B71D0E038ACFE4AFFD4E4B4A.taxon	description	Redescription MEASUREMENTS. Maximum body length: male 2.8 mm and female 4 mm. BODY. Color light brown (Fig. 2 G), mottled with the usual pale muscle spots; posterior margins of pereonites 4 – 7, pleonites 3 – 5 and uropod branches more pigmented. Body outline as in Fig. 15 A. Back smooth with numerous triangular scale-setae (Fig. 15 B). One line of noduli laterales per side on the pereonites inserted at same distance from lateral margins (Fig. 15 A, C). CEPHALON. Without frontal line, suprantennal line straight on median portion; eyes consisting of 10 ommatidia (Fig. 15 D – E). PLEON. Narrower than pereon; epimera of pleonites 3 – 5 not developed. Telson with lateral sides slightly concave, distal margin broadly rounded (Fig. 15 A). ANTENNULA. Composed of three articles subequal in length, distal article with two lateral aesthetascs plus apical pair (Fig. 15 F). ANTENNA. When extended posteriorly reaches posterior margin of pereonite 5; flagellum as long as fifth article of peduncle, first article slightly longer than second and third, second and third articles with three and two lateral aesthetascs, respectively (Fig. 15 G). MOUTH. Mandibles with molar penicil consisting of at least five branches, left mandible with 2 + 1 penicils (Fig. 16 A), right mandible with 1 + 1 penicils (Fig. 16 B). Maxillula (Fig. 16 C) inner endite with two long penicils inserted transversely; outer endite of 5 + 5 teeth, inner set apically cleft. Maxilla (Fig. 16 D) with bilobate apex, inner lobe rounded, bearing thick setae, outer lobe rounded, covered with thin setae plus three thick setae. Maxilliped (Fig. 16 E) base bearing sparse setae, proximal article of palp with two stout setae slightly distinct in length; endite subrectangular, inner and distal margins bearing several thin setae, medial seta not surpassing distal margin, distal margin with one small hairy penicil. UROPOD. Protopod and exopod grooved on outer margin, bearing many glandular pores, exopod almost three times as long as endopod (Fig. 16 F). PEREOPODS. Rather slender, pereopod 1 carpus bearing transverse antennal grooming brush and distal seta apically cleft; dactylus with ungual seta simple and surpassing outer claw, dactylar seta simple, not surpassing outer claw. Male PEREOPODS 1 AND 7. Pereopod 1 (Fig. 17 A) propodus slightly flattened, carpus with transverse antennal grooming brush and distal seta hand-like. Pereopod 7 (Fig. 17 B) without any sexual dimorphism. GENITAL PAPILLA. Ventral shield ovoid, papilla bifurcate with genital orifices opening at apex (Fig. 17 C). PLEOPODS. Pleopod 1 (Fig. 17 D) exopod triangular, inner margin almost straight, outer margin sinuous, distal portion triangular with distal margin rounded; endopod with thickest distal part with sides almost parallel, inner distal part with long pointed process concave on outer margin, prominent subquadrangular apical lobe on outer distal portion bearing thin setae. Pleopod 2 (Fig. 17 E) exopod triangular, outer margin concave, bearing seven stout setae; endopod robust, distinctly longer than exopod. Exopods of pleopods 3 – 5 as in Fig. 17 F – H.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B71D0E038ACFE4AFFD4E4B4A.taxon	discussion	Remarks Vandel (1973) erected the genus Microphiloscia to allocate the new species M. trichoniscoides from Cueva de la Colorada, Provincia de Oriente, Sierra Maestra, Colorada del Maso, Cuba. After the examination of material belonging to this species, M. trichoniscoides shows the shape of the genital papilla and male pleopod 1 endopod typical of members of Halophiloscia (see also Taiti & Lopez 2008). Halophiloscia includes 9 halophilic species mainly distributed along the Mediterranean and Atlantic coasts of Europe and Africa (Schmalfuss 2003; Taiti & Lopez 2008; Taiti & Argano 2009, 2011). In America, only H. couchii (Kinahan, 1858) was recorded, from Argentina, Bermuda, and USA (see Schmidt 2003). The genus is mainly characterized by the runner-type habitus (sensu Schmalfuss 1984), epimera of pereonites 1 – 7 with one or more lines of noduli laterales, male pereopod 1 and sometimes pereopod 2 carpus and propodus enlarged and covered with several scales on frontal side, male genital papilla distally bifurcated, and male pleopod 1 endopod stout bearing a long, pointed process on the apex (see Schmidt 2003; Taiti & Lopez 2008). The species is included within the genus since it shows all the previously mentioned characters, except that the male pereopods 1 and 2 do not have the carpus and propodus enlarged. Thus, Microphiloscia is considered to be a junior synonym of Halophiloscia.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B71D0E038ACFE4AFFD4E4B4A.taxon	distribution	Distribution Cuba (Vandel 1973). First record for Colombia.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7200E3D8AE4E374FDE449D2.taxon	description	Figs 1, 2 H	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7200E3D8AE4E374FDE449D2.taxon	materials_examined	Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 5 ♂♂, 4 ♀♀; El Silencio, Laguna Caracolí; 10 ° 10 ′ 28.4 ″ N, 75 ° 44 ′ 29.96 ″ W; 6 Oct. 2017; Y. M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC-CRU 223 • 1 ♂, 1 ♀; El Silencio, Laguna del Silencio; 10 ° 10 ′ 34 ″ N, 75 ° 44 ′ 26.8 ″ W; 25 Aug. 2017; leg. Y. M. Carpio-Díaz, C. M. López-Orozco and R. Borja-Arrieta; CUDC- CRU 224 • 1 ♂, 4 ♀♀; Laguna La Charca; 10 ° 10 ′ 47.8 ″ N, 75 ° 44 ′ 0.4 ″ W; 7 Sep. 2017; same collectors as for preceding; CUDC-CRU 225 • 2 ♂♂, 3 ♀♀; Hotel La Pola; 10 ° 10 ′ 19.4 ″ N, 75 ° 44 ′ 48.9 ″ W; 24 Nov. 2017; same collectors as for preceding; CUDC-CRU 226 • 1 ♂, 1 ♀; Caño Ratón; 10 ° 10 ′ 42.6 ″ N, 75 ° 44 ′ 52.6 ″ W; 24 Nov. 2017; C. M. López-Orozco leg.; CUDC-CRU 227 • 1 ♂, 1 ♀; 10 ° 10 ′ 33.4 ″ N, 75 ° 44 ′ 52.4 ″ W; 24 Nov. 2017; C. M. López-Orozco, Y. M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC- CRU 228 • 1 ♂, 3 ♀♀; same collection data as for preceding; 10 ° 10 ′ 44.91 ″ N, 75 ° 44 ′ 53.01 ″ W; CUDC- CRU 229 • 1 ♂, 1 ♀; La Punta, Laguna Encantada; 10 ° 10 ′ 48.3 ″ N, 75 ° 43 ′ 38.9 ″ W; 25 Aug. 2017; C. M. López-Orozco, Y. M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC-CRU 230 • 5 ♂♂, 6 ♀♀; same collection data as for preceding; 10 ° 10 ′ 48.1 ″ N, 75 ° 43 ′ 39.1 ″ W; CUDC-CRU 231 • 9 ♀♀; same collection data as for preceding; 10 ° 10 ′ 48.1 ″ N, 75 ° 43 ′ 37.5 ″ W; CUDC-CRU 232 • 1 ♀; same collection data as for preceding; 10 ° 10 ′ 48.4 ″ N, 75 ° 43 ′ 38.4 ″ W; CUDC-CRU 233 • 1 ♀; same collection data as for preceding; 10 ° 10 ′ 48 ″ N, 75 ° 43 ′ 39.3 ″ W; CUDC-CRU 234 • 1 ♂, 5 ♀♀; same collection data as for preceding; 10 ° 10 ′ 48.1 ″ N, 75 ° 43 ′ 37.5 ″ W; 7 Sep. 2017; CUDC-CRU 235 • 3 ♂♂, 1 ♀; Paraíso Secreto; 10 ° 10 ′ 15.9 ″ N, 75 ° 44 ′ 38.8 ″ W; 24 Nov. 2017; C. M. López-Orozco leg.; CUDC-CRU 236 • 13 ♂♂, 13 ♀♀, 8 juvs; same locality and collector as for preceding, 6 Sep. 2017; CUDC-CRU 237 • 14 ♂♂, 32 ♀♀, 3 juvs; same locality as for preceding; 4 Apr. 2018; C. M. López-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 238.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7200E3D8AE4E374FDE449D2.taxon	distribution	Distribution Tanzania (Zanzíbar), Maldives, Comoro Islands, Indonesia (Bali) and Taiwan (Schmalfuss 2003; Taiti 2014). First record for Colombia.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7200E3E8AEFE7D0FE9B4D14.taxon	description	Figs 1, 2 I	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7200E3E8AEFE7D0FE9B4D14.taxon	materials_examined	Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 1 ♂, 2 ♀♀; El Silencio, Laguna Caracolí; 10 ° 10 ′ 28.4 ″ N, 75 ° 44 ′ 29.96 ″ ′ W; 6 Oct. 2017; Y. M. Carpio-Díaz and R. Borja- Arrieta leg.; CUDC-CRU 239 • 3 ♂♂, 3 ♀♀; Paraíso Secreto 10 ° 10 ′ 15.9 ″ N, 75 ° 44 ′ 38.8 ″ W; 6 Sep. 2017; C. M. López-Orozco leg.; CUDC-CRU 240 • 13 ♂♂, 18 ♀♀; same locality as for preceding, 4 Apr. 2018; C. M. López-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 241.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7200E3E8AEFE7D0FE9B4D14.taxon	distribution	Distribution Andaman Islands, Brazil and Guatemala (Schmalfuss 2003; Lisboa et al. 2017). First record for Colombia.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7230E3E8AF4E38BFD2548A5.taxon	description	Figs 1, 2 J	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7230E3E8AF4E38BFD2548A5.taxon	materials_examined	Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • many ♂♂ and ♀♀; Playa Libre; 10 ° 10 ′ 52.3 ″ N, 75 ° 43 ′ 55.7 ″ W; 7 Sep. 2017; C. M. López-Orozco, R. Borja-Arrieta and Y. M. Carpio-Díaz leg.; CUDC-CRU 242 • 1 ♂, 6 ♀♀; same locality as for preceding; 10 ° 10 ′ 52.3 ″ N, 75 ° 43 ′ 54.8 ″ W; 5 Oct. 2017; C. M. López-Orozco leg.; CUDC-CRU 243 • 2 ♂♂; La Punta, Hotel Majagua; 10 ° 10 ′ 56.41 ″ N, 75 ° 43 ′ 33.51 ″ W; 6 Oct. 2017; C. M. López-Orozco and L. Vides leg.; CUDC- CRU 244 • 1 ♂, 2 ♀♀; El Silencio, Laguna Caracolí; 10 ° 10 ′ 28.4 ″ N, 75 ° 44 ′ 29.96 ″ W; 6 Oct. 2017; Y. M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC-CRU 245 • 1 ♂, 2 ♀♀, 1 juv.; El Silencio, Playa La Cocotera; 10 ° 10 ′ 49.3 ″ N, 75 ° 44 ′ 16.7 ″ W; 6 Oct. 2017; C. M. López-Orozco leg.; CUDC-CRU 246 • 1 ♂, 2 ♀♀; Caño Ratón; 10 ° 10 ′ 44.91 ″ N, 75 ° 44 ′ 53.01 ″ W; 24 Nov. 2017; C. M. López-Orozco, R. Borja- Arrieta and Y. M. Carpio-Díaz leg.; CUDC-CRU 247 • 2 ♂♂, 2 ♀♀; same locality, date and collectors; as for preceding; 10 ° 10 ′ 33.4 ″ N, 75 ° 44 ′ 52.4 ″ W; CUDC-CRU 248 • 1 ♂, 3 ♀♀; Paraíso Secreto; 10 ° 10 ′ 15.9 ″ N, 75 ° 44 ′ 38.8 ″ W; 24 Nov. 2017; C. M. López-Orozco leg.; ICN-CR-is 265.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7230E3E8AF4E38BFD2548A5.taxon	distribution	Distribution Circumtropical (Schmalfuss 2003; Taiti & Ferrara 2004; Taiti & Checcucci 2009; Taiti 2014; Lisboa et al. 2017). First record for Colombia.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7230E3F8A47E754FB924F60.taxon	description	Fig. 1	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7230E3F8A47E754FB924F60.taxon	materials_examined	Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 1 ♂; La Punta (TDF); 10 ° 10 ′ 48.42 ″ N, 75 ° 43 ′ 41.36 ″ W; 8 Sep. 2017; Y. M. Carpio-Díaz, C. M. López-Orozco and R. Borja- Arrieta leg.; CUDC-CRU 267 • 1 ♂, 1 ♀; same collection data as for preceding; 10 ° 10 ′ 47.36 ″ N, 75 ° 43 ′ 42.5 ″ W; CUDC-CRU 268 • 4 ♂♂, 6 ♀♀; same collection data as for preceding; 10 ° 10 ′ 47.55 ″ N, 75 ° 43 ′ 42.09 ″ W; CUDC-CRU 269 • 2 ♂♂, 7 ♀♀; same collection data as for preceding; 10 ° 10 ′ 46.79 ″ N, 75 ° 43 ′ 43.51 ″ W; CUDC-CRU 270 • 2 ♂♂, 6 ♀♀; same collection data as for preceding; 10 ° 10 ′ 48.02 ″ N, 75 ° 43 ′ 41.58 ″ W; CUDC-CRU 271 • 1 ♂, 6 ♀♀; same collection data as for preceding; 10 ° 10 ′ 47.74 ″ N, 75 ° 43 ′ 41.8 ″ W; CUDC-CRU 272 • 1 ♂, 6 ♀♀; same collection data as for preceding; 10 ° 10 ′ 48.1 ″ N, 75 ° 43 ′ 37.5 ″ W; CUDC-CRU 273 • 1 ♂; La Punta (TDF); 10 ° 10 ′ 47.76 ″ ′ N, 75 ° 43 ′ 40.27 ″ W; 25 Aug. 2017; Y. M. Carpio-Díaz, C. M. López-Orozco and R. Borja-Arrieta leg.; CUDC-CRU 274 • 1 ♂, 1 ♀; La Punta, Casa Sra. Miriam; 10 ° 10 ′ 46.91 ″ N, 75 ° 43 ′ 46.73 ″ W; 8 Sep. 2017; C. M. López-Orozco, R. Borja-Arrieta and Y. M. Carpio-Díaz leg.; CUDC-CRU 275 • 2 ♀♀; El Silencio, Laguna Caracolí; 10 ° 10 ′ 28.4 ″ N, 75 ° 44 ′ 29.96 ″ W; 6 Oct. 2017; Y. M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC- CRU 276 • 4 ♂♂, 4 ♀♀, 3 juvs; El Silencio (TDF); 10 ° 10 ′ 31.91 ″ N, 75 ° 44 ′ 23.68 ″ W; 24 Aug. 2017; A. Figueroa and B. Valencia leg.; CUDC-CRU 277 • 1 ♂, 1 ♀; El Silencio, Laguna del Silencio; 10 ° 10 ′ 32.3 ″ N, 75 ° 44 ′ 27.2 ″ W; 24 Nov. 2017; C. M. López-Orozco leg.; CUDC-CRU 278 • 7 ♂♂, 12 ♀♀; El Paraiso (TDF); 10 ° 10 ′ 22.9 ″ N, 75 ° 44 ′ 32.8 ″ W; 6 Sep. 2017; C. M. López-Orozco, R. Borja-Arrieta and Y. M. Carpio-Díaz leg.; CUDC-CRU 279 • 3 ♂♂; Caño Ratón; 10 ° 10 ′ 42.9 ″ N, 75 ° 44 ′ 52.8 ″ W; 24 Nov. 2017; C. M. López-Orozco leg.; CUDC-CRU 280. Previous records Botanical Garden “ Guillermo Piñeres ”, Turbaco, Bolívar (Carpio-Díaz et al. 2018).	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7230E3F8A47E754FB924F60.taxon	distribution	Distribution Colombia. First record for continental islands of the Colombian Caribbean.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7220E3F8AE9E17EFA844AA0.taxon	description	Fig. 1	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7220E3F8AE9E17EFA844AA0.taxon	materials_examined	Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 3 ♂♂, 9 ♀♀; La Punta, Casa Sra. Miriam; 10 ° 10 ′ 46.91 ″ N, 75 ° 43 ′ 46.73 ″ W; 8 Sep. 2017; C. M. López-Orozco, R. Borja-Arrieta and Y. M. Carpio-Díaz leg.; CUDC-CRU 260 • 3 ♀♀; Caño Ratón; 10 ° 10 ′ 42.9 ″ N, 75 ° 44 ′ 52.8 ″ W; 24 Nov. 2017; C. M. López-Orozco leg.; CUDC-CRU 261 • 3 ♀♀; El Silencio, Laguna del Silencio; 10 ° 10 ′ 31.91 ″ N, 75 ° 44 ′ 23.68 ″ W; 25 Aug. 2017; Alison Figueroa leg.; CUDC-CRU 262 • 1 ♂, 1 ♀; La Punta (TDF), Laguna Encantada; 10 ° 10 ′ 47.36 ″ N, 75 ° 43 ′ 42.5 ″ W; 8 Sep. 2017; C. M. López-Orozco, Y. M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC-CRU 263 • 2 ♂♂, 3 juvs; same locality, date and collectors as for preceding; 10 ° 10 ′ 47.55 ″ N, 75 ° 43 ′ 42.89 ″ W; CUDC-CRU 264 • 2 ♀♀; same locality and collectors as for preceding; 10 ° 10 ′ 47.8 ″ N, 75 ° 43 ′ 37.5 ″ W; 25 Aug. 2017; CUDC-CRU 265 • 2 ♂♂, 2 ♀♀; La Punta; 10 ° 10 ′ 47.74 ″ N, 75 ° 43 ′ 41.85 ″ W; 8 Sep. 2017, same collectors as for preceding; CUDC-CRU 266. Previous records Botanical Garden “ Guillermo Piñeres ”, Turbaco, Bolívar (Carpio-Díaz et al. 2018).	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7220E3F8AE9E17EFA844AA0.taxon	distribution	Distribution Pantropical (Schmalfuss 2003). First record for continental islands of the Colombian Caribbean.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7220E388921E558FE9F482F.taxon	description	Fig. 1	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7220E388921E558FE9F482F.taxon	materials_examined	Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 3 ♂♂, 7 ♀♀; Orika; 10 ° 10 ′ 37.2 ″ N, 75 ° 44 ′ 27.59 ″ W; 8 Sep. 2017; A. Figueroa leg.; CUDC-CRU 249 • 2 ♀♀; El Silencio, Playa La Cocotera; 10 ° 10 ′ 49.3 ″ N, 75 ° 44 ′ 16.7 ″ W; 6 Oct. 2017; C. M. López-Orozco, R. Borja-Arrieta and Y. M. Carpio-Díaz leg.; CUDC-CRU 250 • 1 ♂, 11 ♀♀; El Silencio, Laguna del Silencio; 10 ° 10 ′ 33.5 ″ N, 75 ° 44 ′ 26.9 ″ W; 24 Aug. 2017, same collectors as for preceding; CUDC-CRU 251 • 1 ♀; same collection data as for preceding; 10 ° 10 ′ 32.9 ″ N, 75 ° 44 ′ 27.3 ″ W; CUDC-CRU 252 • 18 ♂♂, 15 ♀♀, 1 juvenile; same collection data as for preceding; 10 ° 10 ′ 35 ″ N, 75 ° 44 ′ 26.4 ″ W; CUDC-CRU 253 • 3 ♀♀; La Punta, Laguna Encantada; 10 ° 10 ′ 47.76 ″ N, 75 ° 43 ′ 40.27 ″ W; 25 Aug. 2017; C. M. López-Orozco, Y. M. Carpio- Díaz and R. Borja-Arrieta leg.; CUDC-CRU 254. Previous records San Andrés (Van Name 1936); Angelópolis, Cafetal La Camelia Antioquia Dept, and near Nevado del Ruíz, Caldas Dept (Richardson 1912); Cartagena de Indias, Barú, Bolívar Dept (Caribbean region) (Carpio-Díaz et al. 2016).	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7220E388921E558FE9F482F.taxon	distribution	Distribution Cosmopolitan species (Schmalfuss 2003). First record for continental islands of the Colombian Caribbean.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7250E388909E6C0FAFA4B8A.taxon	description	Figs 1, 2 K	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7250E388909E6C0FAFA4B8A.taxon	materials_examined	Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 5 ♂♂, 6 ♀♀; Caño Ratón, Casa Comunal; 10 ° 10 ′ 39.06 ″ N, 75 ° 44 ′ 50.4 ″ W; 8 Sep. 2017; C. M. López-Orozco, R. Borja- Arrieta, Y. M. Carpio-Díaz and A. Figueroa leg.; CUDC-CRU 138 • 2 ♀♀; Orika; 10 ° 10 ′ 37.2 ″ N, 75 ° 44 ′ 27.59 ″ W; 8 Sep. 2017; A. Figueroa leg.; CUDC-CRU 139 • 13 ♂♂, 49 ♀♀; La Punta, Casa Sra. Miriam; 10 ° 10 ′ 46.91 ″ N, 75 ° 43 ′ 46.73 ″ W; 8 Sep. 2017; C. M. López-Orozco, R. Borja-Arrieta and Y. M. Carpio-Díaz leg.; CUDC-CRU 140.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7250E388909E6C0FAFA4B8A.taxon	distribution	Distribution Ascension Island, Senegal, Guinea Bissau, Arabian Peninsula, Madagascar, southern China, Taiwan and Venezuela (Ashmole & Ashmole 2000; Schmidt 2001; Schmalfuss 2003). First record for Colombia.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7240E39892AE34EFB7E48E3.taxon	description	Figs 1, 2 L	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7240E39892AE34EFB7E48E3.taxon	materials_examined	Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 1 ♂; Caño Ratón, Casa Comunal; 10 ° 10 ′ 39.06 ″ N, 75 ° 44 ′ 50.4 ″ W; 8 Sep. 2017; C. M. López-Orozco, R. Borja-Arrieta, Y. M. Carpio-Díaz and A. Figueroa leg.; CUDC-CRU 255 • 1 ♂; Orika; 10 ° 10 ′ 39 ″ N, 75 ° 44 ′ 26.8 ″ W; 25 Aug. 2017; C. M. López-Orozco, R. Borja-Arrieta and Y. M. Carpio-Díaz leg.; CUDC-CRU 256 • 2 ♂♂, 5 ♀♀; La Punta, Casa Sra. Miriam; 10 ° 10 ′ 46.91 ″ N, 75 ° 43 ′ 46.73 ″ W; 8 Sep. 2017; same collectors as for preceding; CUDC-CRU 257 • 4 ♂♂; El Silencio, Laguna del Silencio; 10 ° 10 ′ 33.5 ″ N, 75 ° 44 ′ 26.9 ″ W; 24 Aug. 2017, same collectors as for preceding; CUDC-CRU 258 • 2 ♂♂; same collection data as for preceding; 10 ° 10 ′ 32.9 ″ N, 75 ° 44 ′ 27.3 ″ W; CUDC-CRU 259 • 3 ♂, 11 ♀♀; same collection data as for preceding; 10 ° 10 ′ 34.3 ″ N, 75 ° 44 ′ 26.7 ″ W; ICN-CR-is 266.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7240E39892AE34EFB7E48E3.taxon	distribution	Distribution Brazil and Haiti (Schmalfuss 2003; Campos-Filho et al. 2017). First record for Colombia.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
312287F0B7240E398BB5E693FB074B8A.taxon	description	Sequences obtained for the three species of Tylos from Isla Grande and the specimen from Florida have been deposited under GenBank Accession Numbers MW 532964 – MW 532970 and MW 533069 – MW 533075. Only the 12 S rDNA sequence was obtained for the specimen from Isla Grande assigned to T. marcuzzii. Phylogenetic analyses using only 12 S rDNA group this sequence with the two specimens of T. marcuzzii from Cuba reported in Hurtado et al. (2014); see Fig. 18. The 12 S rDNA p-distance between T. marcuzzii from Isla Grande and T. marcuzzii from Cuba is 2.0 – 2.7 %, whereas the distance between the specimens from the two Cuban localities is 2.0 % (Table 2). Only the 16 S rDNA sequence was obtained for the specimen of T. niveus from the Florida Keys, the type locality of this species. Phylogenetic analyses of 16 S rDNA show that specimens identified as T. niveus from Isla Grande are most closely related to the T. niveus specimen from the Florida Keys. The 16 S rDNA p-distance between T. niveus from Isla Grande and the Florida Keys is 2.7 % (Table 2). The concatenated (12 S rDNA and 16 S rDNA) dataset was comprised of 43 taxa and 740 characters (excluding characters with ambiguous homology), 236 of which were parsimony-informative. The specimens assigned to T. marcuzzii were used as outgroup. Five main lineages were identified for the ingroup taxa in the concatenated phylogenetic analyses, which formed a basal polytomy (Fig. 18): (1) T. niveus from Puerto Rico; (2) Tylos sp. from Yaguanabo, Cuba; (3) T. niveus from Florida and Isla Grande; (4) T. negroi sp. nov.; and (5) Tylos punctatus sensu lato from the Northeastern Pacific. The range of 16 S rDNA and 12 S rDNA p-distances between the four basal lineages excluding T. punctatus is 11.1 – 13.6 % and 9.7 – 15.5 %, respectively. The range of 16 S rDNA and 12 S rDNA p-distances between these ingroup lineages and T. marcuzzii is 19.4 – 22.2 % and 19.5 – 23.8 %, respectively.	en	López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, Gómez-Estrada, Harold, Hurtado, Luis A. (2022): A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island. European Journal of Taxonomy 793: 1-50, DOI: 10.5852/ejt.2022.793.1643, URL: http://dx.doi.org/10.5852/ejt.2022.793.1643
