taxonID	type	description	language	source
313072224B5BFFA7FEA7FA93965EE433.taxon	type_taxon	TYPE SPECIES. — Handlirschia gelasii Reis, 1909 by monotypy.	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B5BFFA7FEA7FA93965EE433.taxon	diagnosis	DIAGNOSIS. — RP 1 / RP 2 fork and point of origin of the first I- in the area between RP 3 + 4 and MA opposite each other, or nearly so (known in Reisia gelasii, Reisia guillaumei n. comb. and Reisia rubra n. comb.); pons very long (known in Reisia guillaumei n. comb. only); RP / MA fork in a more basal position than the MP / Cu + AA fork (known in Reisia guillaumei n. comb. only); at its origin, Irp 1 - rp 2 closer to RP 2 than to RP 1 (known in Reisia guillaumei n. comb. and Reisia rubra n. comb.); first I- in the RP 3 + 4 area well-developed, dichotomously branched (known in all species); in its distal part, MP with 4 main posterior branches (in addition to the ending of its main stem; known in Reisia gelasii and Reisia rubra n. comb., assumed in Reisia guillaumei n. comb.; probably fewer branches in Reisia rubra n. comb.); in the area between the first and second posterior branches of CuA, occurrence of a well-developed, branched I- (known in Reisia guillaumei n. comb. and Reisia rubra n. comb.); CuP rectilinear (as opposed to curved or sigmoidal), with two main stems, both branched, with a well-formed I + between them (known in Reisia guillaumei n. comb. and Reisia rubra n. comb.); AA (or, anterior branch of) rectilinear (as opposed to curved or sigmoidal; known in Reisia guillaumei n. comb. and Reisia rubra n. comb.); area between AA and the posterior wing margin, basal to the pons, lobe-shaped and filled with poorly organized veinlets, some directed backwards (known in Reisia guillaumei n. comb. only; possibly a feature proper to the hindwing). OTHER SPECIES. — Reisia guillaumei (Grauvogel & Laurentiaux, 1952), n. comb., Reisia nana (Laurentiaux-Vieira, Ricour & Laurentiaux, 1952) and Reisia rubra (Béthoux, De la Horra, Benito, Barrenechea, Galán & López-Gómez, 2009), n. comb.	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B5BFFA7FEA7FA93965EE433.taxon	discussion	REMARKS Whether a number of triadophlebiomorphan species should be included, or not, in this genus, remained a conundrum for decades due to the incompleteness of the available material on one hand, and inadequate data on some of its potential relatives. Preliminarily, in the holotype of Reisia gelasii, circular elevations with a puncture in their middle occur in the area between RP 1 and RP 2 [at least in its basal part; noted by Reis (1909) as ‘ pores’; see Fig. 1 E, F]. This is unique among triadophlebiomorphans but might have been overlooked and / or may be poorly preserved in other members of this taxon. Indeed, similar structures were observed by Carpenter (1947) in Permian, large-sized meganisopterans. According to this author, they might represent setal bases. The occurrence and distribution of these structures would need a better documentation for their systematic relevance to be properly assessed. Our new data and observations led us to concur withBechly (1997; and see Grauvogel & Laurentiaux 1952) that the genus Triadotypus Grauvogel & Laurentiaux, 1952 must be regarded a junior synonym of Reisia Handlirsch, 1912. Indeed, as far as preserved, the respective holotypes (and only known material) of the type-species of each genus (Reisia gelasii [Fig. 1] and ‘ Triadotypus ’ guillaumei [Fig. 2]) are nearly identical, including size (width opposite the RP 1 / RP 2 fork, 22.0 mm and about 22.4 mm, respectively), but for: i) the RP 1 / RP 2 fork and the point of origin of the first I- in the RP 3 + 4, both located in a slightly more basal position in Reisia gelasii; and ii) a higher density of cross-veins in the median part of the wing in Reisia gelasii. Such traits are best regarded as of specific relevance. Among characters advocated by Nel et al. (2001) to maintain Reisia and Triadotypus as distinct genera, only the branching pattern of MP (or lack thereof) is preserved in both holotypes. Even though the holotype of ‘ Triadotypus ’ guillaumei is comparatively poorly preserved in the corresponding area, the course of the CuA distal-most branch suggests that MP indeed possesses genuine posterior branches, as in Reisia gelasii. Another notable point is that in ‘ Triadotypus ’ nana, consistently regarded as closely related to ‘ Triadotypus ’ guillaumei by previous authors (including Nel et al. 2001), MP clearly has several posterior branches. We further noted unexpected similarities between Reisia guillaumei n. comb. and ‘ Rabru ’ rubra. Even if the latter is known from a fragmentary, single wing, its wing venation matches, in nearly every aspect, that of Reisia guillaumei n. comb., including the organisation of CuP, provided with two main stems and an I + between them. Both species are also very similar in size (width opposite the RP 1 / RP 2 fork about 21.4 mm in ‘ Rabru ’ rubra). We therefore propose to assign ‘ Rabru ’ rubra to the genus Reisia. One notable point regards the number of MP and CuA branches, lower in Reisia rubra n. comb. than in any other species herein assigned to Reisia. This trait is considered as of specific relevance. As for Reisia nana (Laurentiaux-Vieira, Ricour & Laurentiaux, 1952), known from a very fragmentary, single wing, there are virtually no differences between it and Reisia guillaumei n. comb., apart from a distinctly smaller size (judging from the morphology of the latter, wing width is about 14.0 mm opposite the RP 1 / RP 2 fork in Reisia nana), which justifies maintaining the species (Bechly 1997). The assignment of ‘ Triadotypus ’ sogdianus Pritykina, 1981 to the genus Reisia, provisionally proposed by Nel et al. (2001), is dubious, as this species has (compared to Reisia species for which the corresponding traits are documented): i) the first fork of the first I- in the area between RP 3 + 4 and MA area located more basally; ii) a more developed MP; and iii) curved CuP and AA. This species should probably be assigned to a genus on its own. The case of ‘ Reisia ’ rieki Deregnaucourt, Wappler, Anderson & Béthoux, 2017 will be addressed below. In summary, available evidence points towards marked similarities between Reisia gelasii, Reisia guillaumei n. comb., Reisia rubra n. comb. and Reisia nana. Yet, it must be acknowledged that most of these species are known from fragmentary material. Future discoveries might reveal unforeseen differences, making it necessary to reinstate some of the genera herein regarded as junior synonyms of Reisia.	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B5AFFA7FE8EFA709032E697.taxon	description	(Fig. 1)	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B5AFFA7FE8EFA709032E697.taxon	materials_examined	TYPE LOCALITY AND STRATIGRAPHY. — Münnerstadt, Germany; Muschelkalk basin, Schaumkalkbänke Member, Jena Formation; Anisian, Triassic (Paleobiology Database; Nel et al. 2001). EMENDED DIAGNOSIS. — Width opposite the RP 1 / RP 2 fork about 22 mm (lower in Reisia nana); in the area delimited anteriorly by RP 2 and posteriorly by the first I- between RP 3 + 4 and MA, approximately in the median area of the wing (between the origin of the first I- in the RP 3 + 4 and its first fork), cross-venation denser than in surrounding areas (autopomorphic trait); MP with 4 main posterior branches (in addition to its anterior stem; lower in Reisia rubra n. comb.).	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B5AFFA7FE8EFA709032E697.taxon	discussion	REMARKS Among triadophlebiomorphans, the comparatively high density of cross-veins in the median area of the wing is unique. We submit that it is relevant to distinguish Reisia gelasii from other Reisia species (area poorly known in Reisia rubra n. comb., but which has fewer MP and CuA branches; area unknown in Reisia nana, which is distinctly smaller). There is little to add to the original description by Reis (1909) of the available material. Yet, the exceptionally wellpreserved vein corrugation provides interesting clues on how intercalaries can mimic main veins. In the distal part of the RP 3 + 4 - MA area four pairs of convex and concave intercalaries (I + and I-) arise from regularly convex cross-veins, close to RP 3 + 4 (Fig. 1 B, C; two of these pairs visible on Fig. 1 C). More basally, a very similar pair of intercalary arises from a cross-vein which elevation can be considered neutral. Even more basally, the concavity typical of the I- of the pair applies to the cross-vein in connection with RP 3 + 4. In other words, the I- of a pair of intercalaries adopts a vein-like origin. This case clearly supports the view that the apparent posterior (concave) branch of RP 3 + 4 is more likely an extensively developed I-. Another relevant aspect of wing venation homology for which the specimen provides some clues regards the number of MP vs CuA branches, a point that could be referred to as the ‘ MP-CuA twilight zone’. In its distal part, MP is undoubtedly provided with three posterior branches (Fig. 1 B, D; the most distal and most basal ones have a clear concave origin; only the most basal one of the three is visible on Fig. 1 D). More basally, concave structures do not display such clear concave origin from MP. As for CuA, even though its basal stem is not preserved, it can be assumed from data on Reisia guillaumei n. comb. (Fig. 2) and Piroutetia liasina (Fig. 3) that it has two branches at least, probably more (in green on Fig. 1 A). Then, the nature of structures encircled in white on Fig. 1 D is not obvious. A first interpretation, favoured herein, predicts that a CuAprlg (see ‘ Material and methods’ section) might begin to form, crossing one posterior branch of MP (which putative base is indicated by ‘ 0 ’ on Fig. 1 D). This vein would then have a total of five main branches (four posterior branches in addition to the ending of its main stem), and CuA three preserved ones, and most likely four in total, as in Reisia guillaumei n. comb. A second interpretation predicts that CuA extends further, with a total of five main branches (four preserved and one inferred), and MP four (including its main stem). The case of Piroutetia liasina (Fig. 3), where no CuAprlg can be assumed, shows that CuA has fewer branches than MP (two vs three). We therefore favour the interpretation that, in Reisia gelasii, a CuAprlg occurs along a short section and that CuA most likely had four branches (three of which are preserved).	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B54FFA9FEAAF8D190FBE56C.taxon	description	(Fig. 3) EXAMINED MATERIAL. — Holotype and only known specimen MNHN. F. B 09711 (left wing in ventral aspect; Muséum national d’Histoire naturelle, Palaeontology collection, Paris, France; Fig. 3).	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B54FFA9FEAAF8D190FBE56C.taxon	discussion	REMARKS The redescription of the holotype (and only known specimen) of this species by Nel (1989) only needs a minor emendation, regarding the occurrence of a cross-vein more oblique than others in the distal part of the RA-RP 1 area. Such a cross-vein commonly occurs in Triadophlebiomorpha (Pritykina 1981). The excellent preservation of veins corrugation in the specimen makes it particularly interesting. It can be assessed that MP is branched distally, posteriorly pectinate, with a total of 4 branches. The first posterior branch of MP as identified by Nel (1989) is herein regarded as an intercalary vein occurring between the two branches of CuA, which is more consistent with: i) the lack of a concave base in connection with MP; ii) the position of this structure relative to the most distal (genuine) MP branches; and iii) the common occurrence of such Icua in Triadophlebiomorpha. The course of Irp 1 - rp 2 (expected to be convex) is not evident if considering veins elevation: our proposal (Fig. 3 A) supposes that it is alternatively convex and concave (Fig. 3 B). A possible explanation is that Irp 1 - rp 2 is fused with a posteriorly pectinate I- that would have occurred between Irp 1 - rp 2 and RP 2. Such a structured I- is known in Cladophlebia parvula Pritykina, 1981 (see original description), in which it seemingly originates from RP 2. Moreover, it is separated from Irp 1 - rp 2 by a very narrow area. A fusion of the two structures, as proposed herein for Piroutetia liasina, is therefore plausible.	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B54FFA9FED1FF3296E3E7D2.taxon	description	(Fig. 2)	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B54FFA9FED1FF3296E3E7D2.taxon	materials_examined	EXAMINED MATERIAL. — Holotype and only known specimen 7885 / 7886 (left wing in dorsal and ventral aspect, respectively; also referred to as ’ 57.32 ’ or ‘ 5732 ’ in the literature; Louis Grauvogel collection at the ‘ Staatliches Museum für Naturkunde Stuttgart’, Stuttgart, Germany; Fig. 2).	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B54FFA9FED1FF3296E3E7D2.taxon	description	TYPE LOCALITY AND STRATIGRAPHY. — Bust (Bas-Rhin, France); ‘ Grès à Voltzia’ Formation; Anisian, Triassic; Gall & GrauvogelStamm 2005).	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B54FFA9FED1FF3296E3E7D2.taxon	diagnosis	EMENDED DIAGNOSIS. — Width opposite the RP 1 / RP 2 fork about 22 mm (lower in Reisia nana); MP with 4 main posterior branches (in addition to its anterior stem; fewer branches in Reisia rubra n. comb.).	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B54FFA9FED1FF3296E3E7D2.taxon	discussion	REMARKS Our investigation revealed a number of discrepancies between previous descriptive accounts and what can be observed in the available material. Most importantly, the part that Nel et al. (2001) regarded as the base of MP (and Grauvogel & Laurentiaux [1952] as the base of CuP) turns out to be the pons, which is very long. Incidentally, it is strongly convex, which is inconsistent with an identification as (part of) MP. The course of the actual base of MP, although poorly preserved, yet is substantiated by a marked concave groove immediately posterior to the pons (black arrow on Fig. 2 E, F). Additionally, there is no evidence of a long RP + MA stem, as suggested by Nel et al. (2001). The corresponding area is not preserved in the side preserving the wing in dorsal aspect (Fig. 2): only a very short section of the stem of RA (or R) is preserved, ending opposite the white arrow on Fig. 2 E, F. The splitting plane then runs at the level of ScP, which is well exposed, with a short section of MA visible (i. e. RA and RP are embedded in the other side of the specimen). Further on, where preserved, RA (blue arrow on Fig. 1 A, B), RP and MA are clearly distinct. Quite unfortunately the corresponding area is totally missing in the side preserving the wing in ventral aspect (7886), probably as a consequence of rock splitting at the time of collection. Given our current knowledge of the wing morphology in triadophlebiomorphans, it is rational to assume that RP and MA run next to each other, but distinct, in Reisia guillaumei n. comb. (Fig. 2 A; and see above).	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B54FFABFC75FBB3918BE753.taxon	materials_examined	TYPE SPECIES. — Triassologus biseriatus Riek, 1976 by original designation. OTHER SPECIES. — Triassologus aveyri (Béthoux & Beattie, 2010), n. comb.	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B54FFABFC75FBB3918BE753.taxon	description	EMENDED DIAGNOSIS. — Hindwing broader than forewing (1.3 broader opposite nodus); area between anterior wing margin (or ScA) and ScP very broad in forewing, moderately broad in hindwing; area between anterior wing margin and RA very narrow opposite the third quarter of wing length, before widening again (as opposed to tapering, or broadening, regularly from the nodus to the apex; uncertain in Triassologus aveyri n. comb.); nodal and sub-nodal cross-veins short, moderately strong, sub-aligned, oblique (condition unknown in Triassologus aveyri n. comb.); RP 1 / RP 2 fork in a position more distal than that of the point of origin of the first I- in the RP 3 + 4 area; first I- in the Irp 1 - rp 2 - RP 2 area seemingly originating from RP 2 (i. e. RP 2 pseudo-forked; large blue arrows on Fig. 4 B-D, F; unknown in Triassologus aveyri n. comb.); point of origin of the first I- in the RP 3 + 4 area basal to the RP 1 / RP 2 fork; RP 3 + 4 very long, parallel to the posterior wing margin for some distance (unknown in Triassologus aveyri n. comb.); pons moderately long; RP / MA fork opposite that of MP / Cu + AA; at least in the basal part of the CuA-CuP area, I- rectilinear and seemingly originating from CuA (possibly as convex elements quickly turning concave; large white arrows on Fig. 6), genuine CuA branches zigzagging and seemingly originating from cross-venation (i. e. as an I +; large green arrows on Fig. 6; and see small green arrows on Figs 4 D, 5 B); CuP reaching GEODIVERSITAS • 2023 • 45 (17) 487 the posterior wing margin opposite the nodus or slightly distal to it; CuP and AA posteriorly pectinate, with a curved and sigmoid course, respectively; area between AA (or Cu + AA, or MP + Cu + AA) and the posterior wing margin, opposite and basal to the pons, narrow (i. e. not forming a lobe).	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B54FFABFC75FBB3918BE753.taxon	discussion	REMARKS It will be shown below that ‘ Reisia ’ rieki Deregnaucourt, Wappler, Anderson and Béthoux, 2017 is a junior synonym of Triassologus biseriatus Riek, 1976. This could imply that Triassologus is a junior synonym of Reisia. However, new data derived from additional material of Triassologus biseriatus (see below) and on Reisia spp. (see above) allows to convincingly differentiate the two genera, including the width of the area between the anterior wing margin (or ScA) and ScP, and the particular organisation of I- and CuA branches in the CuA-CuP area, among others. The more complete data on Triassologus biseriatus (see below) led us to compare it with the Australian Triassic species ‘ Iveryia ’ aveyri Béthoux & Beattie, 2010 (type species of the genus Iverya Béthoux & Beattie, 2010). Indeed, as far as comparison can be made, the two species do not display major differences other than size-related ones, such has the overall number of vein branches (assuming that the only known specimen of ‘ Iverya ’ aveyri is a forewing, it is about 1.4 times larger than in that of Triassologus biseriatus). We therefore propose to newly assign ‘ Iverya ’ aveyri to Triassologus, further exemplifying similarities between South African and Australian Triassic insect faunas (Tierney et al. 2020; Béthoux & Anderson 2021). Bechly (1997) considered that RP 2 is branched in Triassologus biseriatus. However, bearing in mind the propensity of intercalary veins to acquire a main vein habitus points to Odonata in general and Triadophlebiomorpha in particular, an alternative option, favoured herein, is that the apparent anterior branch of RP 2 (large blue arrow on Fig. 4 B-D, F) instead is the first I- in the area between Irp 1 - rp 2 and RP 2.	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B56FFADFC0EF99D912EE2B3.taxon	materials_examined	TYPE LOCALITY AND STRATIGRAPHY. — The holotype and the specimen PRE / F / 10125 were found at the ‘ Birds River’ locality (locality code Bir 111; see Anderson & Anderson 1984); material described by Deregnaucourt et al. (2017), Kapokkraal and Aasvoëlberg localities (Kap 111 and Aas 411, respectively); and specimen PRE / F / 5118, Hlatimbe Valley (Hla 213); all South Africa; Molteno Formation; lower Carnian, Upper Triassic (Anderson et al. 1998). EMENDED DIAGNOSIS. — Maximum width about 17 mm in forewing and about 19.5 mm in hindwing (about 24 mm in Triassologus aveyri n. comb., known from a putative forewing), forewing length about 110 mm.	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B56FFADFC0EF99D912EE2B3.taxon	description	SPECIMEN DESCRIPTIONS Specimen PRE / F / 5118 (Figs 4 D, E, 6 A) Adpression preserving a left wing in dorsal (PRE / F / 5118 b; wing basal and median parts; extreme base, most of posterior area, and apical area missing) and ventral (PRE / F / 5118 a; about the first wing third, with portion of posterior wing margin missing opposite the pons) aspects; preserved length 84.5 mm, estimated total length about 110 mm; preserved maximum width 13.6 mm, estimated total maximum width about 16.0 mm; petiole 3.5 mm wide; area between anterior wing margin broad (maximum width 2.9 mm); MA 1.0 mm long between its origin (from RP + MA) to its point of connection with the pons; distal to the pons and basal to the nodus, RA, RP and MA running very close to each other but distinct; nodus located 33.7 mm from wing base; nodal and sub-nodal cross-veins short, moderately strong, sub-aligned, oblique; RP 1 + 2 / RP 3 + 4 fork located shortly after sub-nodal cross-vein; RP 1 + 2 19.3 mm long; first I- between Irp 1 - rp 2 and RP 2 seemingly emerging from RP 2; first I- in the area between RP 3 + 4 and MA originating 13.6 mm distal to the RP 1 + 2 / RP 3 + 4 fork; pons oblique, 2.5 mm long; MP / Cu + AA fork opposite that of RP / MA; MP without obvious preserved branch; Cu + AA 2.8 mm long, Cu 1.6 mm long; in CuA-CuP area, the first 2 I- rectilinear and seemingly originating from CuA, the third I- originating as convex element shortly turning concave (possibly fused with a CuA branch at its origin); CuA branches zigzagging and seemingly originating from cross-venation; CuP and AA posteriorly pectinate; CuP curved, with 6 branches preserved (probably 7 in total), with intercalary veins between them; AA sigmoidal, with a maximum of 3 cells between it and the posterior wing margin; posterior wing margin forming a sharp angle opposite the RA / RP + MA fork.	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B53FFAEFC78FAF1908CE73A.taxon	materials_examined	TYPE SPECIES. — Nototriadophlebia pritykinae n. sp.; monotypic genus.	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B53FFAEFC78FAF1908CE73A.taxon	etymology	ETYMOLOGY. — The genus name derives from Triadophlebia, a related genus, to which was appended the prefix ‘ Noto’ (‘ south’ in Ancient Greek), referring to the geographic occurrence of its type species.	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B53FFAEFC78FAF1908CE73A.taxon	diagnosis	DIAGNOSIS. — As for the type species.	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B53FFB1FC3AF97394D7E573.taxon	description	(Figs 7 - 9) urn: lsid: zoobank. org: act: BB 1717 F 8 - 0 D 90 - 41 B 5 - 9 D 88 - 68689 FB 9 B 410	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B53FFB1FC3AF97394D7E573.taxon	materials_examined	HOLOTYPE. — Specimen BP / 2 / 20950 ab, adpression preserving four wings (Anderson collection, Evolutionary Studies Institute, University of the Witwatersrand, Johannesburg, South Africa; Figs 7, 8). PARATYPE. — Specimen BP / 2 / 20986 ab, adpression preserving a single wing (Anderson collection, Evolutionary Studies Institute, University of the Witwatersrand, Johannesburg, South Africa; Fig. 9). E ScP RA RP 1 1 - rp 2 CuA ScP F RA RP 1 1 - rp 2	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B53FFB1FC3AF97394D7E573.taxon	diagnosis	DIAGNOSIS. — Area between the anterior wing margin and RA (distal to the nodus) widening gradually to ª / 5 of wing length, where it is as wide as the RA-RP 1 area (most prominent in forewing); first I- in the RP 3 + 4 - MA area (i. e. RP 3 + 4 pseudo-fork; large orange arrows on Fig. 8 A, F) arising before RP 3 + 4 mid-length; CuP + AA stem splitting into CuA and CuP + AA (as opposed to Cu and AA; large red arrow on Figs 8 F, 9 A, C, D); AA reaching the posterior wing margin basal to the end of ScP; between the endings of MA and AA, posterior wing margin comparatively straight.	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B53FFB1FC3AF97394D7E573.taxon	etymology	ETYMOLOGY. — The species epithet honours Lyudmila N. Pritykina, for her contribution to our knowledge of fossil Odonata, and of Triadophlebiomorpha in particular.	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B53FFB1FC3AF97394D7E573.taxon	materials_examined	TYPE LOCALITY AND STRATIGRAPHY. — Both holotype and paratype were collected at Birds River (locality code ‘ Bir 111 ’; see Anderson & Anderson 1984), South Africa; Molteno Formation; lower Carnian, Upper Triassic (Anderson et al. 1998). GENERAL DESCRIPTION Fore- and hindwings subequal, very elongated, broadest at mid-length; wing length about 74 mm (as derived from holotype specimen; possibly down to 70 mm), maximum width about 11.0 mm (possibly down to 10.2 mm), width opposite nodus 7.1 - 7.6 mm); petiole long (about 12.8 mm) and narrow (width about 2.4 mm); ScP reaching the anterior wing margin basal to the first third of wing length; distal to the nodus, area between anterior wing margin and RA broadening gradually, at its broadest opposite the end of Irp 1 + 2 - rp 3 + 4; structure of the nodus and origin of RP + MA, RP and MA unknown; RP 1 / RP 2 fork in a position more distal than that of the point of origin of the first I- in the RP 3 + 4 area; Irp 1 - rp 2 weakly differentiated, zigzagging along RP 1; occurrence of 3 I- (and I + between them) forming RP 2 posterior pseudo-branches in the distal part of the area between RP 2 and Irp 1 + 2 - rp 3 + 4; RP 3 + 4 with a marked angle opposite the origin of the first I- (between RP 3 + 4 and MA; large orange arrows on Fig. 8 A, F); the two first I- in the area between RP 3 + 4 and MA branched; in this area, total of about 8 I- reaching the posterior wing margin; MA simple, rectilinear, very close to RP 3 + 4 in its basal third; marked inflexion of the posterior wing margin opposite the ending of MA; MP originating from MP + Cu + AA with a slight obliquity; branches of MP and CuA not clearly distinguishable (probable occurrence of a CuAprlg), CuA with 2 branches at least, and with a I- distally branched between its two (first posterior) branches; MP and CuA covering more than a quarter (less than a third) of the posterior wing margin length; Cu + AA splitting into CuA and CuP + AA, CuA diverging very obliquely; CuP rectilinear, with about 4 - 5 posterior branches and I + between them; AA gently curved, reaching the posterior wing margin basal to the end of ScP. SPECIMEN DESCRIPTIONS Specimen BP / 2 / 20950 (Figs 7; 8) Adpression preserving four wings, with side BP / 2 / 20950 a preserving the right forewing (isolated) and left wings in ventral aspect, and right hindwing in dorsal aspect (the three latter largely overlapping); and its reverse side BP / 2 / 20950 b. Right forewing (Figs 7; 8 A, B). Well preserved, with area basal to the nodus and small portion of apex missing; preserved length 55.6 mm, maximum width 10.7 mm, width opposite nodus about 7.6 mm.	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
313072224B53FFB1FC3AF97394D7E573.taxon	description	Left forewing (Figs 7; 8 C-E). Only anterior-most area decipherable; preserved length 73.2 mm; estimated total length about 74.2 mm. Right hindwing (Figs 7; 8 C, D, F). Well preserved, with nodal area and part of apex missing, and venation in the petiole difficult to decipher; basal to the nodus, RP and MA not distinguishable from each other (assumed to closely overlap); cubito-anal area preserved, showing a CuP + AA stem. Left hindwing (Figs 7; 8 C, D). Barely decipherable but for the posterior wing margin, with a sharp inflexion opposite the end of MA.	en	Béthoux, Olivier, Anderson, John M. (2023): New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata). Geodiversitas 45 (17): 479-496, DOI: 10.5252/geodiversitas2023v45a17, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf
