taxonID	type	description	language	source
305C878DEB2FFFFCFC7EFB65FCDAA566.taxon	type_taxon	TYPE SPECIES. — By original designation, Stenoterommata platensis Holmberg, 1881.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB2FFFFCFC7EFB65FCDAA566.taxon	diagnosis	DIAGNOSIS. — See Goloboff (1995: 57) and Indicatti et al. (2017: 436).	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB2FFFF7FC2EF9C3FDD5A29F.taxon	description	(Figs 1 A-C; 2 - 9) urn: lsid: zoobank. org: act: D 5921 D 02 - 2 B 53 - 4879 - B 850 - 822 C 1675 C 920	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB2FFFF7FC2EF9C3FDD5A29F.taxon	materials_examined	TYPE MATERIAL. — Holotype. Brazil • 1 ♂; São Paulo, Itirapina, Estação Ecológica de Itirapina; 22 ° 14 ’ 47.1 ” S, 47 ° 49 ’ 18.2 ” W; 07 - 08. X. 2018; V. M. Ghirotto, E. F. Trova & R. P. Indicatti leg.; excavated from burrow; CAD 763. Paratypes. Brazil • 1 ♀, 1 ♂; same data as the holotype; CAD (1 ♀ CAD 764); IBSP (1 ♂) • 1 ♂; same data as the holotype; 07 - 08. X. 2018; kept alive, became adult in 01. X. 2019, died in XI. 2019; MCTP • 1 ♀; São Paulo, Itirapina, Estação Ecológica de Itirapina; 22 ° 14 ’ 47.1 ” S, 47 ° 49 ’ 18.2 ” W; 09. X. 2018; E. F. Trova & R. P. Indicatti leg.; excavated from burrow; MZUSP • 2 ♂, 3 ♀; São Paulo, Iperó, Floresta Nacional de Ipanema, near Mirante do Cruzeiro; 23 ° 26 ’ 01.4 ” S, 47 ° 36 ’ 54.8 ” W; 10 - 14. X. 2019; J. P. L. Guadanucci, A. Galleti-Lima, R. P. Indicatti leg.; CAD (1 ♂ CAD 765, 1 ♀ CAD 766), IBSP (1 ♀ IBSP), MNRJ (1 ♂ MNRJ 7687, 1 ♀ MNRJ 7688) • 1 ♂; São Paulo; São Paulo; Morumbi; 10. XII. 2016; R. P. Indicatti leg.; MZUSP ADDITIONAL MATERIAL EXAMINED. — Brazil. 1 ♀, 1 juv.; same data as the holotype; CAD 767 • 1 ♀; same locality as holotype; 22 ° 13 ’ 53.7 ” S, 47 ° 53 ’ 40.8 ” W; 06. X. 2018; E. F. Trova & R. P. Indicatti leg.; ‘ ’ campo sujo’ ’; CAD 768 • 3 ♂, 3 juv.; São Paulo, Iperó, Floresta Nacional de Ipanema, near Mirante do Cruzeiro; 23 ° 26 ’ 01.4 ” S, 47 ° 36 ’ 54.8 ” W; 10 - 14. X. 2019; J. P. L. Guadanucci, A. Galleti-Lima, R. P. Indicatti leg.; CAD (3 ♂ CAD 769 - 771, 3 juv. CAD 772) • 1 juv.; São Paulo; Ipeúna; 2017; V. M. Ghirotto leg.; immature male; CAD 773.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB2FFFF7FC2EF9C3FDD5A29F.taxon	diagnosis	DIAGNOSIS. — Males, females and juveniles of S. neodiplornata Ghirotto & Indicatti, n. sp. can be easily distinguished from all the species of the genus by the combination of striped coloration on tibiae and metatarsi (Fig. 1 A-C) and the dark brown stains reaching all abdomen ventrally (Figs 2 D; 4 D), more evident in live specimens. Males resemble those of S. arnolisei Indicatti, Lucas, Ott & Brescovit, 2008, S. curyi Indicatti, Lucas, Ott & Brescovit, 2008, S. grimpa Indicatti, Lucas, Ott & Brescovit, 2008, S. gugai Indicatti, Chavari, Zucatelli-Júnior, Lucas & Brescovit, 2017, S. leticiae Indicatti, Chavari, Zucatelli-Júnior, Lucas & Brescovit, 2017 and S. uruguai Goloboff, 1995 by the palpal bulb with thick, conical embolus, but can be distinguished by the duct slightly curved in the basal region (Fig. 3 C), embolus bearing strong and slightly sinuous keels, and the blunt, keeled tip, not acutely ending (Fig. 3 A-D, J). Females differ from those of S. crassimana (Mello-Leitão, 1923) by the wider base and shorter or thinner basal dome projection of the spermathecae (Fig. 5 A-F); from S. grimpa by the unfused spermathecal basal dome (Fig. 5 A-F); from S. gugai, S. pavesii Indicatti, Chavari, Zucatelli-Júnior, Lucas & Brescovit, 2017 and S. pescador Indicatti, Chavari, Zucatelli-Júnior, Lucas & Brescovit, 2017 by the curved receptacula ducts (Fig. 5 A-F); from S. iguazu Goloboff, 1995 by having a single basal dome on each side of the spermathecae (Fig. 5 A- F); from S. arnolisei by the shorter basal dome and more rounded receptacula (Fig. 5 A-F); from S. tenuistyla Goloboff, 1995 by the longer and developed ducts (Fig. 5 A-F); from S. palmar Goloboff, 1995 and S. sevegnaniae Indicatti, Chavari, Zucatelli-Júnior, Lucas & Brescovit, 2017 by the longer ducts and shorter basal dome, apically connecting to the ducts (Fig. 5 A-F); from all remaining Stenoterommata by having one or two receptacula on each side of the spermathecae (Fig. 5 A-F).	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB2FFFF7FC2EF9C3FDD5A29F.taxon	etymology	ETYMOLOGY. — The specific epithet is a composite name combining half the name of a genus of Neotropical Barychelidae spiders, Neodiplothele Mello-Leitão, 1917, and the latin word ornatus, meaning ornate as a Neodiplothele, in reference to their very similar general color pattern. At first sight in nature, before closer examination the first two authors though they were collecting Neodiplothele spiders.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB2FFFF7FC2EF9C3FDD5A29F.taxon	description	DESCRIPTION Male (holotype) Color pattern: in life, chelicerae and carapace black covered with golden brown setae, femora black covered on 5 / 6 with black setae, 1 / 6 apical with golden brown setae (Fig. 1 A), trochanter, patellae, tibiae, metatarsi and tarsi dark brown mostly covered with golden brown setae or black setae combined with darker patches mainly on basal region of segments, lighter in the patellae (Fig. 1 A). Sternum, maxillae and coxae yellowish light brown. Abdomen dorsally brownish with black stains covered by golden brown setae, laterally and ventrally brownish with many black stains (Fig. 1 A); in ethanol, carapace (Fig. 2 A) and legs reddish brown with symmetric dark brown mottles covered by golden setae. Abdomen dorsally yellowish light brown with black mottling forming discrete chevron (Fig. 2 C), laterally and ventrally yellowish light brown with many symmetric black stains (Fig. 2). Femora, patellae, tibiae and metatarsi brown yellowish with some darker patches, yellower in the patellae. Total length 12.20. Chelicerae 1.28 long, 0.88 wide (only left side). Carapace 5.50 long, 4.11 wide, with very narrow, procurved fovea, 0.35 wide. Abdomen 5.42 long, 3.45 wide. Thoracic region slightly raised. Clypeus narrow, 0.04 long. Eye tubercle 0.67 long, 1.04 wide, slightly elevated. Anterior eye row slightly procurved, posterior recurved (Fig. 2 A). Eye sizes: AME 0.33, ALE 0.35, PME 0.22, PLE 0.22. Chelicerae with 7 teeth in prolateral row, with c. 10 basal smaller teeth, rastellum weak formed by long thin setae (Figs 2 B; 3 H). Intercheliceral tumescence large, pale yellow, covered with few setae on basal region (Fig. 3 H). Labium 0.40 long, 0.75 wide, without cuspules (Fig. 2 B). Maxillae with c. 36 blunt cuspules on internal basal angle (Fig. 2 B), becoming thin and very elongated at inner margin edge. Serrula developed (Fig. 3 I). Sternum oval, 2.87 long, 2.23 wide. Labial sigilla distant from margin by c. 0.1 × its length, larger than sternal sigilla. Sternal sigilla (Fig. 2 B): anterior slightly smaller than medium, posterior the largest; anterior distant from margin by c. 1 × length, medium and posterior by c. 0.5 × length. Measurements: palp: femur 2.30 / patella 1.35 / tibia 1.43 / cymbium 0.74 / total 5.82; legs: I: femur 4.43 / patella 2.83 / tibia 3.44 / metatarsus 3.21 / tarsus 2.84 / total 16.75; II: 4.25 / 2.45 / 3.06 / 3.25 / 2.50 / 15.51; III: 3.74 / 1.96 / 2.62 / 3.77 / 2.39 / 14.48; IV: 5.20 / 2.52 / 4.14 / 5.04 / 2.60 / 19.50. Spination: palp: femur: d 0 - 0 - 2; legs: femora: I: d 0 - 0 - 0 - 1 p- 1 p; II: d 0 - 0 - 0 - 1 p- 1 p; III: d 0 - 2 - 2 - 2; IV: d 1 - 0 - 2 - 2; patellae: III: p 1 - 1 - 1; IV: r 0 - 1 - 0; tibiae: I: v 2 - 1 r- 1 p- 1 ap + 1 r megaspine (Fig. 3 G), p 0 - 1 - 1 - 0; II: v 1 r- 1 r- 0 - 2 ap, p 0 - 1 - 1 - 0; III: d 0 - 1, v 2 - 1 r- 2 ap, p 0 - 1 - 1 - 0, r 0 - 1 - 1 - 0; IV: v 1 r- 1 r- 2 ap, p 1 - 0 - 1 - 0, r 1 - 0 - 1 - 0; metatarsi: I: v 0 - 1 r- 0 - 1 rap, p 0 - 1 - 1, r 0 - 1 - 0; II: v 0 - 1 r- 1 r- 1 p- 0 - 2 ap, p 0 - 1 - 0 - 0 - 1 ap, r 0 - 0 - 1 - 0 - 1; III: d 1 r- 1 p- 0 - 2, v 2 - 2 - 3 ap, p 1 - 1 - 0 - 1 - 0, r 0 - 1 - 0; IV: d 1 r- 0 - 1 p- 0 - 2, v 2 - 1 r- 1 p- 3 ap, p 1 - 1 - 1, r 0 - 1 - 0. Metatarsal preening combs: III: 3 VR, 3 VP; IV: 3 VR, 3 VP. Combs of leg IV are formed by thicker setae. Tarsi I-IV flexible (Fig. 3 G, tarsus I). Scopulae on tarsi I-IV light and symmetric; I, II divided by sparse row of thin setae; III, IV divided by three rows of thicker setae. Scopulae on full length of metatarsi I and on 1 / 5 of metatarsi II; I, II divided by 3 - 5 sparse rows of thicker setae; III, IV absent. STC large with double row of teeth: I: 6, 7, 8, 7; II: 7 in each row; III: 7, 7, 8, 7; IV: 7, 8, 8, 7. ITC on tarsus IV. Tricobothria with rounded, elevated bases as in S. pavesii (Indicatti et al. 2017: fig. 34). Around 40 epiandric spigots. Four spinnerets (Fig. 2 D): PMS 0.37 long, with pumpkiniform spigots on apical half. PLS: basal segment 1.05, median 0.80, apical triangular, 0.46 long, with band of pumpkiniform spigots on inner edge of all segments. Palp (Fig. 3 E-F): cymbium with elongate dense setae, denser at tip (Fig. 3 E, F); tibia with shallow ventral excavation on apical third (Fig. 3 E, F); tibial excavation and basal region of tegulum with grooves (Fig. 3 A-D); bulb piriform, ventrally curved, thick embolus, with c. 13 retrodorsal parallel keels (Fig. 3 A-D, J). Female (paratype CAD 764) Color pattern as in male, but slightly darker in general (Fig. 4 A-D), and carapace bordered with golden red setae in live specimen (Fig. 1 B). Total length 18.20. Chelicerae 2.75 long, 1.60 wide (only left side). Carapace 6.90 long, 5.32 wide, with narrow, procurved fovea, 0.58 wide. Abdomen 8.55 long, 5.52 wide. Thoracic region slightly raised. Clypeus narrow, 0.17 long. Eye tubercle 0.88 long, 1.32 wide, slightly elevated. Anterior eye row slightly procurved, posterior recurved (Fig. 4 A). Eye sizes: AME 0.32, ALE 0.39, PME 0.24, PLE 0.32. Chelicerae with 7 teeth in prolateral row, with c. 10 basal smaller teeth, rastellum weak, formed by long thin setae (Fig. 4 B). Labium 0.60 long, 1.20 wide, with 1 cuspule (Fig. 4 B). Maxillae with c. 80 blunt cuspules on internal basal angle (Fig. 4 B). Serrula developed (similar to male, Fig. 3 I). Sternum oval, 3.63 long, 2.92 wide. Labial sigilla distant from margin by c. 0.1 × its length, larger than anterior sternal sigilla. Sternal sigilla (Fig. 4 B): anterior slightly smaller, medium and posterior about the same size; anterior distant from margin by c. 1 × length, medium by c. 0.5 × length, and posterior c. 1.2 × length. Measurements: palp: femur 2.92 / patella 1.99 / tibia 1.88 / tarsus 1.78 / total 8.57; legs: I: femur 4.24 / patella 2.98 / tibia 3.06 / metatarsus 2.54 / tarsus 1.78 / total 14.60; II: 4.10 / 2.84 / 2.64 / 2.40 / 1.90 / 13.88; III: 3.67 / 2.32 / 2.05 / 3.10 / 1.89 / 13.03; IV: 4.92 / 2.88 / 3.63 / 4.17 / 2.10 / 17.70. Spination: palp: femur: d 0 - 0 - 0 - 0 - 1 p- 0; tibia: p 0 - 1 - 0, v 1 r- 1 r- 1 p- 0 - 3 ap; legs: femora: I, II: d 0 - 0 - 0 - 0 - 1 p- 0; III, IV: 0; patellae: III: p 1 - 1 - 1; tibiae: III: v 0 - 0 - 0 - 2 ap, p 0 - 1 - 1 - 0, r 0 - 1 - 0; IV: v 0 - 1 r- 0 - 2 ap, r 1 - 0 - 1 - 0; metatarsi: I: v 0 - 1 - 0 - 0 - 1 ap; II: v 0 - 2 - 0 - 0 - 2 ap; III: d 0 - 2 - 0 - 1 p 2, v 0 - 1 - 2 - 0 - 2 - 0 - 3 ap, p 1 - 1 - 1 - 1 - 0, r 0 - 1 - 0; IV: d 0 - 1 r- 1 p- 0 - 1 p- 0 - 2, v 0 - 2 - 0 - 1 r- 1 p- 0 - 0 - 3 ap, p 1 - 1 - 1 - 0, r 0 - 0 - 1 - 0. Metatarsal preening combs: II: 3 VP thin setae; III: 4 VP, 4 VR; IV: 3 VP, 3 VR. Retrolateral combs are formed by thicker setae. Tarsi I, II rigid; III, IV flexible. Scopulae on tarsi I undivided, II divided by 1 - 2 rows of thin setae; III, IV by 3 - 4 rows of thicker setae; palp and legs I, II dense and symmetric; III, IV less dense than anterior tarsi. Scopulae on full length of metatarsi I, II; III only on 1 / 6 of lateral sides; I, II divided on basal third by 3 - 4 rows of thicker setae; IV absent. STC with double row of teeth: I: 5, 7, 6, 5; II: 6, 6, 7, 6; III: 5, 7, 6, 5; IV: 5, 8, 6, 5. ITC on tarsus IV. Palpal claw with 5 teeth on promargin. Tricobothria with rounded, elevated bases. Four spinnerets (Fig. 4 D): PMS 0.76 long, with pumpkiniform spigots on distal half. PLS: basal segment 1.57, median 1.04, apical triangular, 0.56 long, with band of pumpkiniform spigots on inner edge of all segments. Spermathecae with a single receptacula on each side; base inclined outwards; basal dome elevated, ducts curved, receptacula rounded, basally constricted at the ducts (Fig. 5 A). VARIATION. — Males (n = 7): total length 8.02 - 12.40; carapace 3.73 - 5.56 long; maxillae with 32 - 51 cuspules, sometimes elongate and thin or clavate; palpal tibia with spine r 0 - 0 - 1. Females (n = 6): total length 11.60 - 20.36; carapace 4.67 - 7.41; labium with 1 - 3 cuspules; maxillae with 80 - 95 cuspules. Spermathecae either with one or two receptacula on each side; basal dome inclined outwards, either elevated to ducts base, rounded or pointed; curved ducts; receptacula rounded, basally constricted at the ducts (Fig. 5 A-F). NOTE. — Examination of two exuviae from consecutive molts of a juvenile female from ESEC Itirapina showed little variation in the spermathecae, even in size (Fig. 6 A, B). The cephalothorax size of the exuvia of the first molt is 3.21 long; second molt, 4.46 long; the current size in life is 5.54 long.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB2FFFF7FC2EF9C3FDD5A29F.taxon	distribution	DISTRIBUTION. — Southeastern Brazil, state of São Paulo: Itirapina, Ipeúna, Iperó and São Paulo. HABITAT. — This species inhabits distinct environments, showing habitat plasticity (Fig. 7 A-E). Specimens occurs in anthropized zones of seasonal forest (Fig. 7 A), bordered in part by exotic Pinus and Eucalyptus forestry, as well in native savanna formations (Fig. 7 B) at ESEC Itirapina (see Brasileiro et al. 2005; IF 2006 for more details). At FLONA Ipanema, specimens occur in savanna formations within rock outcrops (Fig. 7 C-D), bordered by seasonal forest (mostly Atlantic Forest (Floresta Estacional semidecidual), see Albuquerque & Rodrigues 2000; IBAMA 2006 for more details) and anthropized areas (Fig. 7 E). The species also occur in the city of São Paulo, in a hygrophilous Atlantic Forest urban fragment; historically, this area had great influence and presence of savanna formations (Hueck 1956; Catharino & Aragaki 2008), which could explain the spider’s current distribution in this more humid area.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB2FFFF7FC2EF9C3FDD5A29F.taxon	biology_ecology	NATURAL HISTORY These spiders construct a single open burrow lined with fine silk in the soil (Fig. 8 A, B), within fallen logs (Fig. 8 C, D), rocks crevices (Fig. 8 E, F) and loose rugose tree bark (Fig. 9 A- C). At ESEC Itirapina, most specimens were found in small ravines slopes in an anthropized area presenting earthy soil covered with leaf litter (Fig. 7 A), contrasting with the typical sandy open areas of savanna formations of the remaining of the reserve (Fig. 7 B), where only one individual was found. The anthropized area is similar to close by seasonal forest areas. In some areas, spiders occurred in high density populations of at least five individuals per square meter, where large and some small open burrows (Fig. 8 A-D) are visible even during the day. Many more small to medium sized juvenile spiders were revealed by carefully disassembling the soil from the ravines. At FLONA Ipanema, specimens were found in savanna formations within rock outcrops, bordered by Atlantic Forest and seasonal forest areas (Fig. 7 C, D). Few specimens were found in ravines, in an anthropized area. Most burrows were found in tree trunks (Fig. 9 A-F) and between the rock crevices (Fig. 8 E, F). In the trees, the burrows occurred at different heights, from the ground near roots up to six meters. Most specimens were found in Cambará trees, Gochnatia polymorpha (Less.) Cabrera (Fig. 9 D-F), a typical Cerrado species with very rugose thick bark (ridges with 20 - 50 mm). This allow the spiders to survive fires, which naturally occurs in savannas or forests, even when burning the tree bark (R. P. Indicatti, personal observ.) (Fig. 9 D). Although not common for the genus, the behavior of inhabiting live trees also can be observed in S. arnolisei and S. palmar (Indicatti et al. 2008; 2017), as well as an undescribed species from Parque Nacional do Itatiaia, Rio de Janeiro state, indicating that activity flow between soil and upper branches of the highest part of the trees may take place. This behavior has not yet been observed in any other Pycnothelidae genus, in which other known representatives always inhabits burrows associated with the soil (Goloboff 1995; Lucas et al. 2008; Indicatti 2013; Pérez-Miles et al. 2014; Ferretti 2015; R. P. Indicatti, pers. obs.). Within live tree barks the spiders behaves the same as those that live in the ground or ravines, as observed when the tree barks are removed (Fig. 9 C), exposing remains of prey exoskeleton and old exuviae in end of chamber (Fig. 9 C). The same occurs with some males, which could be observed resting near juvenile and female burrows (Fig. 9 D, E). Individuals kept alive in the laboratory constructed burrows in the soil, lined with fine silk, with 1 - 2 openings. The openings were sometimes observed to be closed with silk and soil for some days to few weeks, when the spider reopened the burrow. They were able to take down large prey (roaches, moths, crickets), c. 1.5 × their total body size.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB24FFEAFEC5FDA7FB1CA484.taxon	description	(Figs 1 D, E; 10 - 12) urn: lsid: zoobank. org: act: 041 B 0 EC 6 - EED 9 - 41 B 7 - 9 F 63 - E 7328 D 0068 F 7	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB24FFEAFEC5FDA7FB1CA484.taxon	materials_examined	TYPE MATERIAL. — Holotype. Brazil • 1 ♂; Mato Grosso do Sul, Bonito, Parque Nacional da Serra da Bodoquena, Três Morros, Fazenda Marambaia trail; 20 ° 58 ’ 13.4 ” S, 56 ° 43 ’ 03.3 ” W; 05. VIII. 2019; V. M. Ghirotto, R. F. Ferreira, J. F. J. Mendes & M. J. A. Morales leg.; under fallen log; CAD 774. Paratype. Brazil • 1 ♂; Mato Grosso do Sul, Bonito, near Jaraguá cave; 21 ° 05 ’ 26.9 ” S, 56 ° 34 ’ 30.6 ” W; 03. VIII. 2019; V. M. Ghirotto, R. F. Ferreira, J. F. J. Mendes & M. J. A. Morales leg.; under fallen log; CAD 775.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB24FFEAFEC5FDA7FB1CA484.taxon	diagnosis	DIAGNOSIS. — Males of S. bodoquena Ghirotto & Indicatti, n. sp. resemble those of S. pavesii, S. gugai, S. peri Indicatti, Chavari, Zucatelli-Júnior, Lucas & Brescovit, 2017 and S. egric Ghirotto & Indicatti, n. sp. by the elongated and thicker embolus (Fig. 11 A-D, J), but can be distinguished by palpal duct with basal and middle region strongly curved (Fig. 11 B-D) and embolus with dorsoventrally compressed keel at the tip (Fig. 11 A-D, J). Additionally, it can be distinguished by having 6 spines (p 1 - 2 - 2 - 1) on prolateral region of patellae III.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB24FFEAFEC5FDA7FB1CA484.taxon	etymology	ETYMOLOGY. — The specific name is a noun in apposition from the type locality, Parque Nacional da Serra da Bodoquena, in the Bodoquena mountain range, in Mato Grosso do Sul, Brazil.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB24FFEAFEC5FDA7FB1CA484.taxon	description	DESCRIPTION Male (holotype) Color pattern: in life, chelicerae, carapace black, legs dark brown, all covered with golden and brown setae (similar to paratype, Fig. 1 D). Abdomen dorsally brown with many black stains not forming a chevron, covered with black setae; in ethanol, chelicerae, carapace and legs reddish brown covered with golden setae (Fig. 10 A). Sternum, maxillae and coxae reddish light brown (Fig. 10 B). Abdomen dorsally yellowish light brown with many dark brown stains not forming a chevron, covered with golden and black setae (Fig. 10 C), and ventrally yellowish light brown with few posterior black stains (Fig. 10 D). Total length 15.99. Chelicerae 2.20 long, 1.27 wide (only left side). Carapace 6.94 long, 5.33 wide, with narrow, procurved fovea, 0.75 wide. Abdomen 6.45 long, 4.41 wide. Thoracic region slightly raised. Clypeus narrow, 0.11 long. Eye tubercle 0.82 long, 1.01 wide, slightly elevated. Anterior eye row procurved, posterior recurved (Fig. 10 A). Eye sizes: AME 0.33, ALE 0.36, PME 0.24, PLE 0.26. Chelicerae with 8 teeth in prolateral row (Fig. 11 H), with c. 25 basal smaller teeth, rastellum weak formed by long thin setae (Figs 10 B; 11 H). Intercheliceral tumescence medium, with few setae on basal region (Fig. 11 H). Labium 0.60 long, 1.16 wide, without cuspules (Fig. 10 B). Maxillae with 105 / 110 blunt and clavate cuspules on internal basal angle (Fig. 10 B), becoming thin and elongated at inner margin edge. Serrula developed (Fig. 11 I). Sternum oval, 3.46 long, 2.86 wide. Labial sigilla distant from margin by c. 0.1 × its length, same size as anterior sternal sigilla. Sternal sigilla (Fig. 10 B): anterior slightly smaller than medium, posterior the largest; anterior and medium distant from margin by c. 0.5 × length, posterior by c. 1 × length. Measurements: palp: femur 2.95 / patella 1.43 / tibia 1.67 / cymbium 0.84 / total 6.89; legs: I: femur 5.01 / patella 3.41 / tibia 3.84 / metatarsus 3.63 / tarsus 2.51 / total 18.40; II: 4.73 / 2.95 / 2.96 / 3.57 / 2.36 / 16.57; III: 4.59 / 2.46 / 2.73 / 4.23 / 2.32 / 16.33; IV: 6.16 / 2.95 / 4.37 / 6.12 / 2.73 / 22.33. Spination: palp: femur: d 0 - 0 - 0 - 0 - 2, tibia: p 1 ap, r 1 ap; legs: femora: I: d 0 - 0 - 0 - 2 - 2; II: d 0 - 1 p- 1 p- 1 p; III: d 0 - 0 - 2 - 2 - 2; IV: d 1 - 1 p- 2 - 2; patellae: II: p 0 - 0 - 1 s- 1 s; III: p 1 - 2 - 2 - 1, r 0 - 1 - 0; IV: r 0 - 1 - 0; tibiae: I: v 2 r- 2 - 1 ap + 1 r megaspine (Fig. 11 G), p 0 - 1 - 1 - 0; II: v 1 p- 2 r- 0 - 1 - 0 - 2 ap, p 0 - 1 - 1 - 0; III: d 0 - 0 - 1 - 0, v 3 - 3 - 3 ap, p 0 - 1 - 1 - 0, r 0 - 1 - 1 - 0; IV: v 3 - 3 - 3 ap, p 0 - 1 - 1 - 0, r 1 - 1 - 1; metatarsi: I: v 0 - 1 - 0 - 0 - 1 ap, p 0 - 1 - 0 - 1; II: d 0 - 0 - 1 p- 0 - 0, v 0 - 1 r- 1 r- 0 - 0 - 3 ap, p 1 - 1 - 0 - 0 - 1, r 0 - 0 - 1 - 0; III: d 1 r- 1 - 1 - 0 - 2 ap, v 0 - 2 - 2 - 2, p 1 - 1 - 1, r 1 - 1 - 0 - 1; IV: d 1 r- 1 - 1 - 0 - 1 rap, v 0 - 2 - 1 p- 1 r- 3 ap, p 1 - 1 - 1 - 1, r 1 - 0 - 1 - 0 - 1. Metatarsal preening combs: III: 5 VR, 6 VP; IV: 5 VR, 4 VP. Combs of leg IV are formed by thicker setae. Tarsi I-IV flexible (Fig. 11 G, tarsus I). Scopulae on tarsi I-IV light and symmetric; I, II divided by 2 rows of thicker setae; III divided by 3 - 4 rows; IV divided by 4 - 5 rows. Scopulae on 1 / 8 of metatarsi I, II; divided by 4 - 5 sparse rows of thicker setae; III, IV absent. STC large with double row of teeth: I: 8, 5, 5, 8; II: 8, 6, 6, 7; III: 7, 5, 5, 7; IV: 8, 5, 5, 8. ITC on tarsus IV. Tricobothria with rounded, elevated bases. Around 32 epiandric spigots. Four spinnerets (Fig. 10 D): PMS 0.77 long, with pumpkiniform spigots on apical half. PLS: basal segment 1.41, median 1.02, apical triangular, 0.66 long, with band of pumpkiniform spigots on inner edge of all segments. Palp (Fig. 11 E-F): cymbium with elongate dense setae, denser at tip (Fig. 11 E, F); tibia with shallow ventral excavation on apical third (Fig. 11 E, F); tibial excavation and basal region of tegulum (Fig. 11 A-D) with grooves; bulb piriform, ventrally curved, embolus with c. 15 parallel keels mainly located retrodorsally (Fig. 11 A-D, J). VARIATION. — Males (n = 2): total length 13.00 - 15.99; carapace 6.44 - 6.94 long; maxillae with 82 - 110 cuspules.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB24FFEAFEC5FDA7FB1CA484.taxon	distribution	DISTRIBUTION. — Known only from Bonito, state of Mato Grosso do Sul, Central Brazil.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB24FFEAFEC5FDA7FB1CA484.taxon	biology_ecology	HABITAT. — Individuals were found in two nearby although distinct forest areas, 20 km apart. The Parque Nacional da Serra da Bodoquena consists of a relictual Atlantic Forest patch (semidecidual forest), surrounded and heavily influenced by the Cerrado. The area near Jaraguá cave is a fragment of deciduous Cerrado forest (seasonal forest), with low canopy and dryer conditions (Fig. 12 A-C). NATURAL HISTORY Only two males and two juveniles were found by active search. The juveniles were found inside fallen rotting logs (Fig. 12 D, E), which were already soft and loose, where they supposedly constructed burrows (as they were visible after splitting the log, no such burrows could be seen). Both adult males were found underneath fallen logs. In the areas they were found, no burrows were visible in the soil.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB38FFEFFF4FFF46FAD1A55B.taxon	description	(Figs 1 F; 13 - 17) urn: lsid: zoobank. org: act: 42 DEC 204 - EE 4 C- 4001 - 81 B 7 - E 6 A 946 F 23 BE 9	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB38FFEFFF4FFF46FAD1A55B.taxon	materials_examined	TYPE MATERIAL. — Holotype. Brazil • 1 ♂; São Paulo, Botucatu, Fazenda Experimental Edgárdia; 22 ° 48 ’ 28.5 ” S, 48 ° 23 ’ 36.6 ” W; 02. VI. 2014; R. C. B. Paradero leg.; restauration area, collected with pitfall traps; CAD 585. Paratypes. Brazil • 1 ♀; same data as the holotype; CAD 776 • 6 ♂; same data as the holotype; CAD (1 ♂ CAD 777), IBSP (1 ♂), MZUSP (1 ♂), MNRJ (1 ♂ MNRJ 7689), MCTP (1 ♂), MCN (1 ♂) • 1 ♂; same locality and leg. as holotype; 21. XI. 2013; pasture area; CAD 591 • 1 ♂, same locality and collector as the holotype; 18. VIII. 2015; forest area; CAD 577. ADDITIONAL MATERIAL EXAMINED. — Brazil • 8 ♂; same data as the holotype; CAD (2 ♂ CAD 778, 2 ♂ CAD 779), IBSP (2 ♂), MZUSP (1 ♂), MNRJ (1 ♂ MNRJ 7690) • 2 ♂; same locality as the holotype; 11. XI. 2013; CAD 587.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB38FFEFFF4FFF46FAD1A55B.taxon	diagnosis	DIAGNOSIS. — Males of S. chavarii Ghirotto & Indicatti, n. sp. resemble those of S. crassimana, S. palmar, S. sevegnaniae and S. tenuistyla by the palpal bulb with thin, filiform embolus (Fig. 14 A-D, J). It can be distinguished from S. sevegnaniae and S. tenuistyla by the curved embolus at the tip (Fig. 14 A-D, J); from S. crassimana by the palpal duct much less curved in medial region (Fig. 14 B, D) and lacking stains ventrally on abdomen (Fig. 14 D); from S. palmar by the palpal tibia with 2 spines on prolateral-medial region (Fig. 14 E) and presence of ITC on legs III, IV. Females can be distinguished from the remaining species of the genus, except from S. quena Goloboff, 1995, by the presence of metatarsal preening combs on leg I and on VP and VR sides of leg II; differs from S. quena and resembles S. platensis Holmberg, 1881 and S. iguazu by the spermathecae with two independent bases on each side (Fig. 16) and prolateral spines 1 - 1 - 1 on patella III; differs from them by the elongated basal dome, metatarsal preening combs on all legs and anterior eye row slightly recurved (Fig. 15 A).	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB38FFEFFF4FFF46FAD1A55B.taxon	etymology	ETYMOLOGY. — The specific name is a patronym in honor of João Lucas Chavari, friend of the last authors and collector of several mygalomorph spiders from Brazil.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB38FFEFFF4FFF46FAD1A55B.taxon	description	DESCRIPTION Male (holotype) Color pattern: in ethanol, chelicerae, carapace reddish brown covered with golden setae (Fig. 13 A), legs brown, covered with black and golden setae. Sternum, maxillae and coxae yellowish brown (Fig. 13 B). Abdomen yellowish light brown with dark brown mottling forming discrete chevron (Fig. 13 C). Total length 10.23. Chelicerae 0.90 long, 0.75 wide (only left side). Carapace 5.08 long, 3.64 wide, with narrow, procurved fovea, 0.48 wide. Abdomen 3.98 long, 2.25 wide. Thoracic region raised. Clypeus very narrow, 0.06 long. Eye tubercle 0.59 long, 0.86 wide, slightly elevated. Anterior eye row slightly procurved, posterior slightly recurved (Fig. 13 A). Eye sizes: AME 0.19, ALE 0.28, PME 0.19, PLE 0.21. Chelicerae with 8 teeth in prolateral row, with c. 20 basal smaller teeth, rastellum weak formed by long thin setae (Figs 13 B; 14 H). Intercheliceral tumescence small, with few setae on basal region (Fig. 14 H). Labium 0.35 long, 0.76 wide, without cuspules (Fig. 13 B). Maxillae with 48 / 54 blunt and clavate cuspules on internal basal angle (Fig. 13 B), becoming thin and very elongated at inner margin edge. Serrula developed (Fig. 14 I). Sternum oval, 2.38 long, 1.99 wide. Labial sigilla distant from margin by c. 0.1 × its length, same size as anterior sternal sigilla. Sternal sigilla (Fig. 13 B): anterior slightly smaller than medium, posterior the largest; anterior and posterior distant from margin by c. 1 × length, medium by c. 0.5 × length. Measurements: palp: femur 1.97 / patella 1.00 / tibia 1.17 / cymbium 0.56 / total 4.70; legs: measurements: I: femur 3.46 / patella 2.18 / tibia 2.45 / metatarsus 2.36 / tarsus 1.72 / total 12.17; II: 3.13 / 1.92 / 1.95 / 2.24 / 1.76 / 11.00; III: 3.09 / 1.67 / 1.78 / 2.78 / 1.82 / 11.14; IV: 4.04 / 2.08 / 2.77 / 4.04 / 2.02 / 14.95. Spination: palp: femur: d 0 - 0 - 0 - 0 - 0 - 0 - 2 - 2; patella: p 0 - 0 - 1; tibia: p 0 - 0 - 2 - 0 - 2, r 1 sap; legs: femora: I: d 0 - 0 - 0 - 0 - 1 r- 2; II: d 0 - 1 r- 1 r- 2 - 2; III: d 0 - 0 - 2 - 2 - 2; IV: d 0 - 0 - 2 - 0 - 2 - 2; patellae: I: p 0 - 0 - 1; II: p 0 - 0 - 1; III: p 1 - 1 - 1, r 0 - 0 - 1 - 0; IV: r 0 - 0 - 1 - 0; tibiae: I: v 1 - 0 - 1 - 2 - 0 - 1 p + 1 r megaspine (Fig. 14 G), p 0 - 0 - 1 - 1 - 0; II: v 1 - 1 - 0 - 3 ap, p 0 - 0 - 1 - 1 - 0; III: d 0 - 1, v 2 - 3 - 0 - 3 ap, p 0 - 1 - 1 - 0, r 0 - 1 - 0; IV: v 2 - 2 - 0 - 3 ap, p 0 - 1 - 1 - 0, r 1 - 1 - 0 - 1 - 0; metatarsi: I: v 0 - 2 - 0 - 1 ap, p 0 - 1 - 0 - 1 ap, r 0 - 0 - 0 - 1 - 0; II: d 0 - 1 - 0, v 0 - 1 r- 1 r- 1 p- 0 - 3 ap, p 1 - 1 - 0 - 1, r 0 - 0 - 1 - 0; III: d 1 r- 1 r- 1 p- 0 - 2 - 0 - 0 - 2, v 2 - 2 - 3 ap, p 1 - 0 - 1 - 1, r 1 - 1 - 0; IV: d 2 - 1 p- 0 - 1 r- 1 p- 0 - 2, v 0 - 2 - 0 - 1 p- 1 r- 1 p- 3 ap, p 1 - 1 - 1, r 1 - 0 - 1 - 0. Metatarsal preening combs: III: 3 VR, 4 VP; IV: 4 VR, 3 VP. Combs of leg IV are formed by thicker setae. Tarsi I-IV flexible (Fig. 14 G, tarsus I). Scopulae on tarsi I-IV light and symmetric; I, II divided by 2 sparse rows of thin setae; III divided by 4 - 5 rows; IV divided by 5 rows, III, IV divided by thicker setae. Scopulae on 1 / 3 of metatarsi I, II, divided by 2 rows of thicker setae; III, IV absent. STC large with double row of teeth: I: 6, 6, 5, 6; II: 7, 6, 6, 6; III: 8, 8, 6, 7; IV: 9, 8, 7, 8. ITC on tarsi III, IV. Tricobothria with rounded, elevated bases. Around 20 epiandric spigots. Four spinnerets (Fig. 13 D): PMS 0.44 long, with pumpkiniform spigots on apical half. PLS: basal segment 0.92, median 0.56, apical triangular short, 0.36 long, with wide band of pumpkiniform spigots on inner edge of all segments. Palp (Fig. 14 E, F): cymbium with elongate dense setae, denser at tip (Fig. 14 E, F); tibia ventrally with a shallow excavation on apical third (Fig. 14 E, F); tibial excavation and basal region of tegulum with grooves (Fig. 14 A-D); bulb piriform, thin embolus curved at the tip, with c. 20 parallel keels (Fig. 14 A-D, J). Female (paratype CAD 776) Color pattern: in ethanol, chelicerae reddish dark brown, carapace (Fig. 15 A) and legs brown, covered with few black and golden setae, Sternum, coxae, and maxillae yellowish brown (Fig. 15 B). Abdomen dorsally light yellowish brown with dark brown mottling forming a chevron (Fig. 15 C), ventrally light yellowish brown (Fig. 15 D). Total length 12.68. Chelicerae 1.70 long, 1.12 wide (only left side). Carapace 5.12 long, 3.50 wide, with narrow, procurved fovea 0.45 wide. Abdomen 5.86 long, 3.58 wide. Thoracic region slightly raised. Clypeus narrow 0.10 long. Eye tubercle 0.45 long, 0.85 wide, slightly elevated. Anterior eye row slightly recurved, posterior recurved (Fig. 15 A). Eye sizes: AME 0.20, ALE 0.27, PME 0.17, PLE 0.20. Chelicerae with 8 teeth in prolateral row, with c. 20 basal smaller teeth, rastellum weak, formed by long thin setae (Fig. 15 B). Labium 0.47 long, 0.84 wide, with 2 cuspules (Fig. 15 B). Maxillae with 72 / 73 blunt cuspules on internal basal angle (Fig. 15 B). Serrula weak. Sternum oval, 2.68 long, 2.15 wide. Labial sigilla distant from margin by c. 0.1 × its length, larger than anterior sternal sigilla. Sternal sigilla (Fig. 15 B): anterior almost round, slightly smaller than medium; posterior c. 2 × larger; all distant from margin by c. 1 × its length. Measurements: palp: femur 2.34 / patella 1.32 / tibia 1.51 / tarsus 1.46 / total 6.63; legs: I: femur 3.56 / patella 2.25 / tibia 2.42 / metatarsus 2.13 / tarsus 1.44 / total 11.80; II: 3.16 / 2.02 / 2.10 / 2.00 / 1.54 / 10.82; III: 2.70 / 1.81 / 1.50 / 1.85 / 1.55 / 9.41; IV: 3.53 / 2.05 / 2.34 / 3.12 / 1.27 / 12.31. Spination: palp: tibia: v 0 - 2 - 0 - 3 ap; tarsus: r 1 i- 0 - 0 - 0; legs: femora: I: d 1 p sap; II-IV: 0; patellae: I, II: 0; III: p 1 - 1 - 1, r 0 - 1 - 0; IV: r 0 - 1 - 0; tibiae I, II: 0; III: d 0 - 1, v 2 ap, p 1 - 1, r- 0 - 1 s- 0; IV: v 2 ap, p 1 i- 1 i, r 1 s- 1 s; metatarsi: I: v 1 ap; II: v 1 r- 1 r- 1 p- 0 - 2 ap; III: d 1 r- 1 r- 1 p- 2 - 0 - 2, v 2 - 2 - 0 - 3 ap, p 1 - 1 - 1 - 1; IV: d 0 - 1 r- 1 p- 2 - 0 - 2, v 2 - 0 - 1 p- 1 r- 1 p- 3 ap, r 1 s- 1 s- 0, p 1 - 1 - 1 - 0. Metatarsal preening combs: I: 2 VP thin setae; II: 4 VP in one line, 4 VR in two lines; III: 4 VP, 5 VR; IV: 4 VP, 4 VR, III, IV in one line. Tarsi I-IV rigid. Scopulae on tarsi I divided by 2 - 3 rows of setae, II by 4 rows, III, IV divided by 8 rows of thicker setae; palp and I, II dense and symmetric; III, IV light, only lateral sides. Scopulae on 4 / 5 of metatarsi I and 2 / 5 on II, divided by 1 - 2 rows of thicker setae on leg I and 2 - 3 on II; III, IV absent. STC with double row of teeth: I: 4, 5, 5, 4; II: 4, 5, 5, 5; III: 4, 6, 6, 4; IV: 6, 7, 7, 6. ITC on tarsi III, IV. Palpal claw with 3 teeth on promargin. Tricobothria with rounded, elevated bases. Four spinnerets (Fig. 15 D): PMS 0.38 long, with pumpkiniform spigots on apical 2 / 3. PLS: basal segment 0.89, median 0.58, apical triangular, short, 0.35 long, with wide band of pumpkiniform spigots on inner edge of all segments. Spermathecae with two independent bases on each side (Fig. 16); the external side has an elongated basal dome with one globose receptaculum arising from the median area; the internal side has an elongated basal dome not presenting a receptaculum; both very sclerotized (Fig. 16). VARIATION. — Males (n = 15): total length 8.26 - 10.94; carapace 3.71 - 5.10 long; labium with 0 - 2 cuspules; maxillae with 46 - 58 cuspules, sometimes elongated.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB38FFEFFF4FFF46FAD1A55B.taxon	distribution	DISTRIBUTION. — Known only from the type locality, Fazenda Experimental Edgárdia, Botucatu, São Paulo state, Brazil.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB38FFEFFF4FFF46FAD1A55B.taxon	biology_ecology	HABITAT. — These spiders were collected by pitfall traps in ecotonal zones between seasonal forest (Cerrado) and Atlantic Forest (Fig. 17 A, B), in Fazenda Experimental Edgárdia, where both vegetation types are present. In addition, they were also captured in restoration and pasture areas, which have similar local conditions to Cerrado formations, indicating that the species probably also occur in strictly Cerrado areas, as S. bodoquena Ghirotto & Indicatti, n. sp.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB3CFFE5FC8BF9E3FDC9A7A0.taxon	description	(Figs 1 G, H; 18 - 22) urn: lsid: zoobank. org: act: B 04 C 1562 - E 4 F 5 - 478 A-B 836 - FE 850178 AB 6 E	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB3CFFE5FC8BF9E3FDC9A7A0.taxon	materials_examined	TYPE MATERIAL. — Holotype. Brazil • 1 ♂; Minas Gerais, Lima Duarte, Conceição do Ibitipoca, Ibitipoca mountain range; 21 ° 41 ’ 14.7 ” S, 43 ° 53 ’ 46.9 ” W; 11. I. 2017; V. M. Ghirotto leg.; under stone, kept alive, became adult in 01. X. 2017; CAD 780. Paratypes. Brazil • 2 ♀; same data as the holotype; CAD (1 ♀ CAD 781, 1 ♀ CAD 782) • 8 ♂, 1 ♀; Parque Estadual do Ibitipoca; 21 ° 42 ’ 55.4 ” S, 43 ° 53 ’ 56.9 ” W; IX. 1997; 1380 m a. s. l.; A. de Oliveira & B. M. Sousa leg.; sucupira area, collected with pitfall traps; IBSP (6 ♂, 1 ♀ IBSP 108383, 2 ♂ IBSP 108458). ADDITIONAL MATERIAL EXAMINED. — Brazil • 39 ♂, 1 j; Minas Gerais, Lima Duarte, Conceição do Ibitipoca, Parque Estadual do Ibitipoca; 21 ° 42 ’ 55.4 ” S, 43 ° 53 ’ 56.9 ” W; 1380 m a. s. l.; IX. 1997; A. de Oliveira & B. M. Sousa leg.; sucupira area, collected with pitfall traps; IBSP (2 ♂ IBSP 108377, 1 ♂ IBSP 108379, 1 ♂ IBSP 108380, 1 ♂ IBSP 108405, 1 ♂ IBSP 108407, 1 ♂ IBSP 108415, 4 ♂ IBSP 108431, 1 ♂ IBSP 108433, 1 ♂ IBSP 108434, 1 ♂ IBSP 108437, 9 ♂ IBSP 108448, 5 ♂ IBSP 108463, 1 ♂ IBSP 108465, 3 ♂ IBSP 108475, 2 ♂, 1 j IBSP 108476, 3 ♂ IBSP 108477 - 108479, 1 ♂ IBSP 108486, 1 ♂ IBSP 110531).	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB3CFFE5FC8BF9E3FDC9A7A0.taxon	diagnosis	DIAGNOSIS. — Males of S. egric Ghirotto & Indicatti, n. sp. resembles those of S. pavesii, S. gugai, S. peri and S. bodoquena Ghirotto & Indicatti, n. sp. by the elongated and thicker embolus (Fig. 19 A-D, J); differ from S. pavesii by the thicker embolus, not curved at the tip (Fig. 19 A-D, J); from S. gugai, S. peri and S. bodoquena Ghirotto & Indicatti, n. sp. by the thicker embolus tip (Fig. 19 A-D, J). Females differ from S. gugai and S. pescador by the receptacula with shorter and thicker ducts (Fig. 21 A, B); from those of S. arnolisei, S. iguazu and S. tenuistyla by having 2 - 3 receptacula on each side (Fig. 21 A, B); from S. pavesii by the basal dome slight inclined outwards (Fig. 21 A, B); from all remaining Stenoterommata species by the shorter and less sclerotized basal dome (Fig. 21 A, B). Additionally, individuals can be distinguished from most species by the dark coloration in life (Fig. 1 G, H) and by the greater size, both sexes averaging significantly larger than most species of the genus (Fig. 1, note scales).	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB3CFFE5FC8BF9E3FDC9A7A0.taxon	etymology	ETYMOLOGY. — The specific epithet is an acronym taken from “ Espeleo Grupo Rio Claro - EGRIC ” which organized the expedition to Ibitipoca mountain range that led the first author to collect the type specimens, and in recognition of their efforts in the study and preservation of Brazilian caves.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB3CFFE5FC8BF9E3FDC9A7A0.taxon	description	DESCRIPTION Male (holotype) Color pattern: in life, chelicerae, carapace and legs black, abdomen dorsally yellowish brown with dark brown mottling forming a chevron with wide bands. Body covered by black setae (Fig. 1 G); in ethanol, chelicerae, carapace (Fig. 18 A) and legs reddish brown covered with black and golden setae. Sternum, maxillae and coxae reddish light brown (Fig. 18 B). Abdomen dorsally yellowish brown with dark brown mottling forming a chevron with wide bands, densely covered with dark brown setae (Fig. 18 C), and ventrally yellowish light brown with few posterior brown stains (Fig. 18 D). Total length 15.35. Chelicerae 1.55 long, 1.36 wide (only left side). Carapace 7.56 long, 5.92 wide, with narrow, procurved fovea, 0.77 wide. Abdomen 6.24 long, 3.40 wide. Thoracic region slightly raised. Clypeus narrow, 0.05 long. Eye tubercle 0.78 long, 1.30 wide, slightly elevated. Anterior eye row recurved, posterior slightly recurved (Fig. 18 A). Eye sizes: AME 0.34, ALE 0.42, PME 0.26, PLE 0.27. Chelicerae with 10 teeth in prolateral row (Fig. 19 H), with c. 35 basal smaller teeth, rastellum weak formed by long thin setae (Figs 18 B, 17 H). Intercheliceral tumescence small with few setae on basal region (Fig. 19 H). Labium 0.57 long, 1.13 wide, with 2 cuspules (Fig. 18 B). Maxillae with 84 / 90 blunt cuspules on internal basal angle. Serrula weak (Fig. 19 I). Sternum oval, 3.58 long, 2.92 wide. Labial sigilla distant from margin by c. 0.1 × it is length, larger than sternal sigilla. Sternal sigilla (Fig. 18 B): anterior and medium of the same size, posterior c. 2 × larger; anterior and medium distant from margin by c. 0.5 × length, posterior c. 1 × length. Measurements: palp: femur 3.68 / patella 2.29 / tibia 2.51 / cymbium 1.17 / total 9.65; legs: I: femur 6.32 / patella 3.69 / tibia 5.17 / metatarsus 4.68 / tarsus 3.66 / total 23.52; II: 6.03 / 3.38 / 4.30 / 5.26 / 3.10 / 22.07; III: 5.65 / 2.78 / 3.60 / 5.38 / 3.15 / 20.56; IV: 7.01 / 3.19 / 5.20 / 7.25 / 3.29 / 25.94. Spination: palp: femur: d 0 - 0 - 2; tibia: r 1 sap; legs: femora: I: d 0 - 0 - 0 - 2 - 2; II: d 0 - 1 r- 1 p- 1 r- 2 - 2; III: d 1 - 2 - 2 - 2; IV: d 1 - 1 p- 0 - 2 - 2; patellae: I: p 0 - 0 - 1; II: p 0 - 1 - 1; III: p 1 - 1 - 1, r 0 - 1; tibiae: I: v 3 - 0 - 1 r- 1 p- 2 p ap + 1 r megaspine (Fig. 19 G), p 0 - 1 - 1 - 0; II: v 3 - 0 - 1 r- 1 p- 3 ap, p 0 - 1 - 1 - 0; III: d 0 - 1, v 3 - 2 - 3 ap, p 0 - 1 - 1 - 0, r 0 - 1 - 1 - 0; IV: v 4 - 2 - 3 ap, p 1 - 1, r 1 - 1; metatarsi: I: d 0 - 1 p- 0 - 1 p, v 0 - 1 r- 0 - 0 - 1 ap; II: d 0 - 1 p- 0 - 1 p, v 2 - 2 - 0 - 2 ap, r 0 - 0 - 1 - 0; III: d 1 - 1 r- 1 p- 2 - 0 - 2, v 2 - 2 - 3 ap, p 1 - 1 - 0 - 1; IV: d 2 - 2 - 1, v 2 - 0 - 2 - 3 ap, p 1 - 1 - 1. Metatarsal preening combs: III: 5 VR, 5 VP; IV: 7 VR, 7 VP. Tarsi I-IV flexible (Fig. 19 G, tarsus I). Scopulae on tarsi I-IV light and symmetric; I, II divided by 3 rows of thin setae; III, IV divided by 4 rows of thicker setae. Scopulae on full length of metatarsi I and on apical 1 / 5 of metatarsi II, both divided by 3 sparse rows; III, IV absent. STC large with double row of teeth: I: 9, 6, 4, 8; II: 8, 6, 6, 8; III: 8, 6, 7, 8; IV: 9, 7, 6, 7. ITC on tarsus IV. Tricobothria with rounded, elevated bases. Around 60 epiandric spigots. Four spinnerets (Fig. 18 D): PMS 0.81 long, with pumpkiniform spigots on 2 / 3 apical. PLS: basal segment 1.19, median 0.71, apical triangular, 0.60 long, with band of pumpkiniform spigots on inner edge of all segments. Palp: cymbium with elongate dense setae, denser at tip (Fig. 19 E, F); tibia ventrally with a shallow excavation on apical third (Fig. 19 E, F); tibial excavation and basal region of tegulum with grooves (Fig. 19 A-D); bulb piriform, ventrally curved, with embolus thick, dorso-ventrally flattened, with c. 12 parallel keels (Fig. 19 A-D). Female (paratype CAD 781) Color pattern: in life as in male; in ethanol, chelicerae reddish dark brown, carapace (Fig. 20 A) and legs reddish brown, covered with black and golden setae, Sternum, coxae, and maxillae slightly lighter than the carapace (Fig. 20 B). Abdomen dorsally yellowish brown with dark brown mottling forming a chevron with wide bands (Fig. 20 C), ventrally yellowish brown (Fig. 20 D). Total length 25.01. Chelicerae 3.90 long, 2.65 wide (only left side). Carapace 9.35 long, 7.78 wide, with narrow, procurved fovea 1.10 wide. Abdomen 11.76 long, 7.69 wide. Thoracic region slightly raised. Clypeus narrow 0.24 long. Eye tubercle 1.14 long, 1.59 wide, slightly elevated. Anterior eye row slightly recurved, posterior slightly recurved (Fig. 20 A). Eye sizes: AME 0.38, ALE 0.55, PME 0.39, PLE 0.35. Chelicerae with 11 teeth in prolateral row, with c. 35 basal smaller teeth, rastellum weak, formed by long thin setae (Fig. 20 B). Labium 1.06 long, 1.92 wide, with 2 cuspules (Fig. 20 B). Maxillae with c. 110 blunt cuspules on internal basal angle (Fig. 20 B). Serrula weak. Sternum oval, 4.67 long, 4.32 wide. Labial sigilla distant from margin by c. 0.1 × its length, larger than anterior sternal sigilla. Sternal sigilla (Fig. 20 B): anterior almost round, slightly smaller than medium; posterior c. 2 × larger; all distant from margin by c. 0.5 × length. Measurements: palp: femur 5.11 / patella 3.01 / tibia 3.43 / tarsus 3.16 / total 14.71; legs: I: femur 7.26 / patella 4.79 / tibia 5.09 / metatarsus 4.36 / tarsus 3.20 / total 24.70; II: 6.34 / 4.27 / 4.26 / 4.02 / 3.30 / 22.19; III: 5.48 / 3.22 / 3.30 / 4.51 / 2.48 / 18.99; IV: 7.46 / 4.01 / 5.45 / 6.40 / 2.62 / 25.94. Spination: palp: tibia: v 2 - 1 r- 1 p- 4 ap, p 0 - 1 - 0; legs: femora: I, II: d 1 p sap; III: d 0 - 0 - 2; patellae: I, II: p 0 - 0 - 1; III: p 1 - 1 - 1, r 0 - 1 - 0; tibiae: I: v 1 r- 1 r- 2 ap, p 0 - 1 - 1 - 0; II: v 1 r- 1 r- 2 ap, p 0 - 1 - 1 - 0; III: d 0 - 1, v 2 - 1 r- 2 ap, p 1 - 1, r- 0 - 1 - 0; IV: v 1 - 1 - 2 ap, r 1 - 1; metatarsi: I: v 1 r- 0 - 1 r ap; II: v 1 r- 2 - 0 - 0 - 2 ap, p 0 - 1 - 0 - 0; III: d 1 r- 1 r- 1 p- 2 - 0 - 2, v 2 - 2 - 3 ap, p 1 - 1 - 1; IV: d 0 - 1 r- 1 p- 2 - 0 - 2, v 2 - 1 p- 1 r- 1 p- 0 - 3 ap, p 0 - 1 - 1. Metatarsal preening combs: III: 12 VP, 12 VR, joined; IV: 4 VP, 8 VR. Tarsi I, II rigid; III, IV flexible. Scopulae on tarsi I undivided; on tarsi II divided by 2 rows of sparse setae; III divided by 6 rows, IV by 8 rows; III, IV divided by thicker setae; palp and I, II dense and symmetric; III, IV moderate. Scopula on full lenght of metatarsi I, II, undivided; III, IV absent. STC with double row of teeth: I: 7, 5, 3, 5; II: 6, 4, 3, 7; III: 7, 5, 4, 6; IV: 6, 4, 4, 7. STC on legs III, IV, the latter larger. ITC on tarsus IV. Palpal claw with 5 teeth on promargin. Tricobothria with rounded, elevated bases. Four spinnerets (Fig. 20 C, D): PMS 1.03 long, with pumpkiniform spigots on apical 2 / 3. PLS: basal segment 1.86, median 1.23, apical triangular, 0.81 long, with band of pumpkiniform spigots on inner edge of all segments. Spermathecae with 2 receptacula on the right side, 3 on the left side; very small basal dome, inclined outwards, sclerotized at the top, ducts very short, receptacula rounded (Fig. 21 A). VARIATION. — Females (n = 2): total length 25.24 - 28.74; carapace 9.35 - 9.92; maxillae with 100 - 110 cuspules. Spermathecae with 2 - 3 receptacula (Fig. 21 B).	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB3CFFE5FC8BF9E3FDC9A7A0.taxon	distribution	DISTRIBUTION. — Known only from Ibitipoca mountain range, Lima Duarte, Minas Gerais state, Brazil.	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
305C878DEB3CFFE5FC8BF9E3FDC9A7A0.taxon	biology_ecology	HABITAT. — This species inhabits different formations at the type locality. They were found in small forests patches of the Atlantic Forest biome in deeper soils and also savanna formations at the border of the main formation of the Ibitipoca mountain range. These forests patches penetrate and are surrounded by savannas of the Cerrado domain (Fig. 22 A). In the Parque Estadual do Ibitipoca, all the specimens of S. egric Ghirotto & Indicatti, n. sp. were collected in similar habitats as for the holotype (Sousa & Cruz 2008: 261). Individuals of this species were seen (but not collected) at cave entrances in the Parque Estadual do Ibitipoca. NATURAL HISTORY Individuals were found in burrows lined with dense silk underneath stones. The burrow constitutes of one or two entrances (Fig. 22 B). Due to few opportunities of active search at night, it is not unlikely that individuals also construct burrows directly in the soil, as does several Stenoterommata species (e. g. S. neodiplornata Ghirotto & Indicatti, n. sp.). In the Parque Estadual do Ibitipoca, the specimens of S. egric Ghirotto & Indicatti, n. sp. were collected with pitfall traps by Sousa & Cruz (2008); they represented one of the most common mygalomorph species in the region (47 ♂, 1 ♀, 1 juv.), after Homoeomma montanum (Mello-Leitão, 1923) (153 ♂, 2 ♀, 2 juv.), Theraphosidae Thorell, 1869 and Idiops camelus (Mello-Leitão, 1937) (109 ♂, 1 ♀, 1 juv.), Idiopidae Simon, 1889 (Sousa & Cruz 2008; R. P. Indicatti pers. obs.). Stenoterommata spiders, despite being a potential prey for small vertebrates, were not found within the stomach contents of the lizards Enyalius perditus Jackson, 1978 (Sousa & Cruz 2008). On the other hand, curiously, 13.33 % (n = 4) of the sampled lizards had juveniles of the trapdoor spider genus Idiops Perty, 1833 in its stomach contents (Sousa & Cruz 2008). In addition, these lizards are diurnal, sleeping on the tip of the branches of shrubs and trees at night (Sturaro & Silva 2010), while juveniles and females of Idiops usually are nocturnal, living within trapdoor burrows (Indicatti 2013: figs 12, 13; Ferretti et al. 2017: fig. 3 b, c), making the encounter even more difficult. Probably encounters occurred at dusk, during which the two could be active. Although the Idiops have a nocturnal habit, some specimens can be found at dusk on the edge, or even entirely outside their burrows, as observed in several areas of Cerrado and Atlantic Forest in southeastern Brazil (R. P. Indicatti pers. obs.).	en	Ghirotto, Victor Morais, Guadanucci, José Paulo Leite, Indicatti, Rafael Prezzi (2021): The genus Stenoterommata Holmberg, 1881 (Araneae, Pycnothelidae) in the Cerrado and Atlantic Forest from Southeastern and Central Brazil: description of four new species. Zoosystema 43 (17): 311-339, DOI: 10.5252/zoosystema2021v43a17
