taxonID	type	description	language	source
374618679B44FF9451EAF949C4B9F93E.taxon	diagnosis	Diagnosis. This species is distinguished by the following combination of characters: scapular tubercles with setae located almost at posterior margin of prodorsal shield, presence of incomplete median line, non granular prodorsal shield margin, prodorsal shield shows symmetrical cells and lines, presence of 7 columnar cells at anterior margin of prodorsal shield, 5 rayed tarsal empodium, smooth dorsal annuli, h 1 seta present, surface of coxa I ornamented with curved lines where as that of coxa II is almost smooth except two curved parallel lines at distal ends, 1 b tubercles with setae located ahead of anterior coxal approximation, epigynium is almost smooth except a median sinuate line.	en	Chakrabarti, Samiran, Sarkar, Sanjay (2011): Three new species of eriophyoid mites (Acari: Eriophyoidea) infesting fruit yielding plants from India. Zootaxa 2988: 28-36, DOI: 10.5281/zenodo.203722
374618679B44FF9451EAF949C4B9F93E.taxon	description	FEMALE (n = 10): Body 140.9 (139.8 – 140.9) long, 60.6 (58.3 – 61.2) wide. Gnathosoma 17.2 (15.2 – 17.2) long, curved down, dorsal pedipalp genual seta d 5.6 (5.3 – 5.9). Prodorsal shield 40.1 (40.1 – 41.2) long, 56.0 (55.3 – 56.0) wide, nongranular and with an acuminate shield lobe; anterolateral region of prodorsal shield shows 7 columnar cells running parallel to each others, between the lateral margin of prodorsal shield and submedian line there are 2 cells of unequal size, median line incomplete and present only on rear end, two carinae from the admedian line meet the anterior tip of the median line, admedian lines complete, sinuate and connected with each others at two sites by transverse lines, the transverse connection between admedian line at middle region has two lateral extensions, submedian lines sinuate, diverge and bifurcate above the base of dorsal tubercles, dorsal tubercles almost on rear shield margin, scapular seta sc 4.5 (4.5 – 5.6). Leg I from base of trochanter 43.8 (43.8 – 44.8); femur 10.2 (10.2 – 11.2), basiventral femoral seta bv 11.2 (9.3 – 11.2); genu 5.6 (5.1 – 5.6), antaxial genual seta 1 ʺ 14.9 (14.9 – 15.8); tibia 5.6 (5.6 – 6.5), paraxial tibial seta 1 ʹ 3.7 (3.7 – 4.1); tarsus 6.5 (5.6 – 6.5); paraxial fastigial tarsal setae ft ʹ 17.7 (16.9 – 17.7), antaxial fastigial tarsal seta ft ʺ 14.0 (13.9 – 14.0); paraxial unguinal tarsal seta u ʹ 3.7 (3.7 – 4.4), tarsal solenidion ω highly curved, knobbed and 7.4 (6.8 – 7.4), 5 rayed, tarsal empodium em 6.5 (5.6 – 6.5). Leg II from base of trochanter 40.1 (39.2 – 40.1); femur 9.3 (8.4 – 9.3), basiventral femoral seta bv 5.6 (5.6 – 6.5), genu 4.5 (4.1 – 4.5), antaxial genual seta 1 ʺ 9.3 (9.3 – 9.8); tibia 10.2 (9.8 – 10.2) without paraxial tibial seta 1 ʹ; tarsus 6.5 (5.6 – 6.5), paraxial fastigial tarsal setae ft ʹ 19.6 (18.4 – 19.6), antaxial fastigial tarsal seta ft ʺ absent, paraxial unguinal tarsal seta u ʹ 4.5 (4.1 – 4.5), tarsal solenidion ω strongly curved, knobbed and 7.4 (6.8 – 7.4); 5 rayed, tarsal empodium em 6.5 (6.5 – 5.6). Coxae I 18.6 (17.9 – 18.6) and contiguous; coxal surface ornamented with curve lines; 1 b tubercles and seta present well above the sternal line; seta 1 b 7.4 (7.4 – 8.4) and 10.1 (9.4 – 10.1) apart, 1 a tubercles with seta present a little ahead of line across the 2 a tubercles; seta 1 a 13.0 (13.0 – 14.0) and 6.9 (6.9 – 7.3) apart; the distance between setae 1 a and 1 b is 8.1 (7.9 – 8.2); coxa II comparatively smooth except two curve lines along the anterior margin of coxae and 14.9 (14.0 – 14.9), seta 2 a 39.2 (37.3 – 39.2) and 18.2 (17.2 – 18.2) apart. Opisthosoma with 39 (37 – 39) smooth dorsal annuli and 72 (70 – 72) ventral annuli; microtubercles rounded and located on first 63 annular lines; last 9 ventral annuli have microstriations, seta c 2 30.8 (28.0 – 30.8) present on annulus 14 (13 – 14), seta d 42.0 (41.0 – 42.0) on annulus 28 (28 – 29); seta e 27.0 (26.1 – 27.0) on ventral annulus 40 (38 – 40); seta f 26.1 (25.9 – 26.1) on ventral annulus 61 (59 – 62); seta h 1 1.8 (1.2 – 1.8), seta h 2 39.2 (39.2 – 37.3). Epigynium 9.3 (8.4 – 9.3) long, 14.9 (13.8 – 14.9) wide, smooth except a median sinuate line and semi circular in shape; seta 3 a 22.4 (19.6 – 22.4). MALE: Not observed	en	Chakrabarti, Samiran, Sarkar, Sanjay (2011): Three new species of eriophyoid mites (Acari: Eriophyoidea) infesting fruit yielding plants from India. Zootaxa 2988: 28-36, DOI: 10.5281/zenodo.203722
374618679B44FF9451EAF949C4B9F93E.taxon	materials_examined	Type material. Holotype: Female (marked) on slide (no. 1300 / 19 / 2006), India: West Bengal: Malda, Englishbazar, Latitude: 24 ° 50 ʹ 40 ʺ N and Longitude: 87 ° 55 ʹ 50 ʺ E, 23 June, 2006 from Spondias pinnata Kurz (Anacardiaceae), Coll. S. Sarkar. Paratypes: 11 females on slide bearing holotype and 42 females on 5 slides (nos. 1301 - 1305 / 85 / 2006); collection data same as in holotype. Relation to host. Light brown, fusiform mites are found as under surface leaf vagrants.	en	Chakrabarti, Samiran, Sarkar, Sanjay (2011): Three new species of eriophyoid mites (Acari: Eriophyoidea) infesting fruit yielding plants from India. Zootaxa 2988: 28-36, DOI: 10.5281/zenodo.203722
374618679B44FF9451EAF949C4B9F93E.taxon	etymology	Etymology. The specific epithet derived from ‘ Spondias ’ genus of the host plant.	en	Chakrabarti, Samiran, Sarkar, Sanjay (2011): Three new species of eriophyoid mites (Acari: Eriophyoidea) infesting fruit yielding plants from India. Zootaxa 2988: 28-36, DOI: 10.5281/zenodo.203722
374618679B44FF9451EAF949C4B9F93E.taxon	discussion	Remarks. The new species shows similarities with A. morindae Ghosh & Chakrabarti, 1989 by having ornamented coxae, 5 rayed tarsal empodium, incomplete median line, and 1 b tubercles ahead of anterior coxal approximation, but differs from the latter by having smooth epigynium and in location of scapular tubercles. The new species also resembles A. privae Mohanasundaram, 1980 by incomplete median line on prodorsal shield, ornamented coxae and presence of h 1 setae but differs from the latter by its smooth prodorsal shield margin and dorsal annuli.	en	Chakrabarti, Samiran, Sarkar, Sanjay (2011): Three new species of eriophyoid mites (Acari: Eriophyoidea) infesting fruit yielding plants from India. Zootaxa 2988: 28-36, DOI: 10.5281/zenodo.203722
374618679B46FF9251EAF88AC5DEF8F6.taxon	diagnosis	Diagnosis. This species is distinguished by the following combination of characters: prodorsal shield shows a median line at the cell formed between the dorsal tubercles by the admedians and V like cross-line, prodorsal shield lobe shows only admedian lines, direction of scapular setae upward and centrad, presence of seta h 1, 4 rayed, tarsal empodium, curved tarsal solenidion, 14 longitudinal scorings on epigynium and both coxal surface ornamented with wavy parallel lines.	en	Chakrabarti, Samiran, Sarkar, Sanjay (2011): Three new species of eriophyoid mites (Acari: Eriophyoidea) infesting fruit yielding plants from India. Zootaxa 2988: 28-36, DOI: 10.5281/zenodo.203722
374618679B46FF9251EAF88AC5DEF8F6.taxon	description	FEMALE (n = 10): Body 113.8 (113.8 – 135.3) long, 63.4 (57.8 – 68.1) wide. Gnathosoma 21.4 (21.4 – 22.4) long, evenly curved downward. Dorsal pedipalp genual seta d 3.7 (3.2 – 3.7) and pedipalp coxal seta ep present. Prodorsal shield 42.0 (42.0 – 42.9) long, 50.4 (50.4 – 54.14) wide with an anterior lobe overhanging gnathosoma; prodorsal shield design represents clear pattern of curved lines. Median line extends between the middle and posterior V like cross line of admedians; admedian lines wavy with three cross lines first one on 9.2, second one on 22.2 from anterior margin of prodorsal shield, and the third one on 4.7 from posterior margin of the prodorsal shield forming a V-shaped line, widely apart at middle forming an almost rhomboid shaped gap; submedian lines sinuate, extend from the anterior cross line of admedian up to the posterior shield margin touching the base of the scapular setae sc, submedian lines touch the admedian at middle. Dorsal tubercles ahead of rear shield margin, 25.2 apart; scapular seta sc 5.6 (4.6 – 5.6) long, directed upward and centered. Leg I from base of trochanter 28.0 (28.0 – 28.9), femur 13.0 (12.1 – 13.0), basiventral femoral seta bv 13.1 (12.8 – 13.1), genu 3.7 (3.7 – 4.6), antaxial genual seta 1 ʺ 23.3 (22.4 – 24.2), tibia 4.6, paraxial tibial seta 1 ʹ 4.6 (4.6 – 5.6), tarsus 5.6 (5.1 – 5.6), paraxial fastigial tarsal seta ft ʹ 14.9 (14.9 – 18.6), antaxial fastigial tarsal seta ft ʺ 25.2 (21.4 – 25.2), paraxial unguinal tarsal seta u ʹ 3.2 (3.2 – 3.7), tarsal empodium em 4.6 (4.1 – 4.6) and 4 rayed, tarsal solenidion ω knobbed, slightly curved and 6.5 (6.5 – 7.0). Leg II from base of trochanter 28.9 (28 – 28.9), femur 14.0 (13.0 – 14.0), basiventral femoral seta bv 13.0 (12.1 – 13.0), genu 3.7 (3.7 – 4.6), antaxial genual seta 1 ʺ 7.4 (7.4 – 8.4), tibia 3.7 (3.7 – 4.6); paraxial tibial seta 1 ' 5.6 (4.6 – 5.6); tarsus 5.6 (5.1 – 5.6), paraxial fastigial tarsal setae ft ʹ 10.2 (10.2 – 11.2), antaxial fastigial tarsal seta ft ʺ 23.3 (19.6 – 23.3), paraxial unguinal tarsal seta u ʹ 3.7 (3.2 – 3.7), 4 - rayed, tarsal empodium em 4.7 (4.1 – 4.7), tarsal solenidion ω 6.5 (6.1 – 6.5). Coxae I contiguous, 11.2 (11.2 – 12.1) long, ornamented with parallel lines between seta 1 b and seta 1 a, seta 1 b 4.6 (4.6 – 5.6) and 9.2 (9.2 – 9.8) apart, 1 a tubercles with seta a little ahead of the line between the 2 a tubercles; seta 1 a 12.1 (12.1 – 13.2) long and 9.2 (9.2 – 9.8) apart; the distance between setae 1 a and 1 b is 4.8 (4,8 – 5.2); coxa II 12.1 (12.1 – 13.0) and ornamented with wavy parallel lines just below the setae 2 a, seta 2 a 25.2 (25.2 – 28.9) long and 22.8 (21.2 – 22.8) apart. Opisthosoma with 36 (32 – 36) dorsal annuli with a broad dorsal trough, flanked on either side with ridges, ventral annuli 61 (61 – 64). Ventral annuli microtuberculated on anterior margin of annuli whereas dorsal annuli devoid of microtubercles, dorsal annuli broader and larger than ventral annuli, seta c 2 14.0 (14.0 – 16.8) on annulus 11 (10 – 11), seta d 48.5 (46.6 – 48.5) on ventral annulus 52 (23 – 25), seta e 4.6 (4.6 – 5.6) on ventral annulus 39 (39 – 40), seta f 14.9 (14.0 – 14.9) on ventral annulus 57 (57 – 59), seta h 1 4.6 (3.7 – 4.6), seta h 2 43.8 (42.0 – 43.8); last 9 (9 – 13) annuli micro striated. Epigynium almost triangular in shape, 13.0 (13.0 – 15.8) long and 22.4 (20.5 – 22.4) wide, with 14 scorings, seta 3 a 36.2 (29.8 – 39.2). MALE: Not observed.	en	Chakrabarti, Samiran, Sarkar, Sanjay (2011): Three new species of eriophyoid mites (Acari: Eriophyoidea) infesting fruit yielding plants from India. Zootaxa 2988: 28-36, DOI: 10.5281/zenodo.203722
374618679B46FF9251EAF88AC5DEF8F6.taxon	materials_examined	Type material. Holotype: Female (marked) on slide (no. 1434 / 85 / 2006), India: West Bengal: Dakshindinajpur, Doulatpur, Latitude: 25 ° 10 ʹ 20 ʺ N and Longitude: 88 ° 14 ʹ 50 ʺ E, 24 December, 2006 from Citrus maxima (Rutaceae), Coll. S. Sarkar. Paratypes: 6 females on slide bearing holotype and 72 females on 7 slides (nos. 1435 - 1441 / 85 / 2006); collection data same as in holotype. Relation to host. White, spindle form mites are found as vagrants on undersurface of leaf showing yellow spots on leaf under surface.	en	Chakrabarti, Samiran, Sarkar, Sanjay (2011): Three new species of eriophyoid mites (Acari: Eriophyoidea) infesting fruit yielding plants from India. Zootaxa 2988: 28-36, DOI: 10.5281/zenodo.203722
374618679B46FF9251EAF88AC5DEF8F6.taxon	etymology	Etymology. The specific epithet derived from ‘ Citrus ’ genus of the host plant, ‘ cola ’ from Latin ‘ Colus ’ meaning dwelling in or inhabitant.	en	Chakrabarti, Samiran, Sarkar, Sanjay (2011): Three new species of eriophyoid mites (Acari: Eriophyoidea) infesting fruit yielding plants from India. Zootaxa 2988: 28-36, DOI: 10.5281/zenodo.203722
374618679B46FF9251EAF88AC5DEF8F6.taxon	discussion	Remarks: Among the species of this genus having 4 rayed tarsal empodium the new species comes close to P. daturae Mohanasundaram & Ranganath, 1985 by longitudinal scorings on epigynium and to P. comorensis Keifer, 1974 by curved tarsal solenidion and similar epigynium. However, the new species differs from P. d a t u r a e by having ornamented coxal surface, presence of h 1 seta and dissimilar direction of scapular setae and from P. comorensis by dissimilar shield pattern and direction of scapular seta with tubercles. The new species also shows its closeness to Phyllocoptruta oleivora (Ashmead, 1879) in sharing common host genus and by having similar body shape, opisthosomal trough, location and direction of scapular seta (upward and centrad), scoring on epigynium but differs from it by having 4 rayed tarsal empodium, presence of seta h 1, smooth basal area of epigynium and the prodorsal shield pattern.	en	Chakrabarti, Samiran, Sarkar, Sanjay (2011): Three new species of eriophyoid mites (Acari: Eriophyoidea) infesting fruit yielding plants from India. Zootaxa 2988: 28-36, DOI: 10.5281/zenodo.203722
374618679B43FF9151EAFF2BC686F839.taxon	diagnosis	Diagnosis. This species is distinguished by the following combination of characters: numerous irregular broken lines on prodorsal shield, frontal lobe of prodorsal shield has clear median and admedian lines, bifurcated dorsal pedipalp genual setae d, scapular tubercles much ahead of the posterior shield margin, seta on tibia I absent, 4 rayed, tarsal empodium, presence of two types of ventral annuli i. e. first 14 – 16 ventral annuli with microtubercles and rest of the ventral annuli with microstriations, both the coxae smooth, proximal region of female genitalia has small striations, two horn like lateral projections emerge from mid lateral margin of epigynium.	en	Chakrabarti, Samiran, Sarkar, Sanjay (2011): Three new species of eriophyoid mites (Acari: Eriophyoidea) infesting fruit yielding plants from India. Zootaxa 2988: 28-36, DOI: 10.5281/zenodo.203722
374618679B43FF9151EAFF2BC686F839.taxon	description	FEMALE (n = 10): Body 169.9 (165.4 – 169.9) long, 56.0 (55.2 – 56.0) wide. Gnathosoma 23.3 (22.3 – 23.3) long, curved down, dorsal pedipalp genual seta d bifurcated and 9.3 (9.1 – 9.3). Prodorsal shield 65.3 (64.8 – 65.3) long, 56.0 (55.2 – 56.0) wide with a prominent shield lobe of 19.6 (18.6 – 19.6) long over the base of gnathosoma, frontal shield lobe triangular, with distinct median and admedian lines and numerous broken lines; prodorsal shield shows granulations and a prominent ‘ V’ shaped mark extended from anterior shield margin up to middle encompassing prominent median line, faint impressions of median and admedian lines are found below the ‘ V’ mark, numerous broken lines are also found at two lateral sides of prodorsal shield, near the base of each dorsal tubercle small semicircular arcs are found. Scapular tubercles 13.0 (12.6 – 13.0) conspicuous, ahead of rear shield margin and 26.1 (26.1 – 26.9) apart, scapular seta sc 2.8 (2.1 – 2.8) and directed upward and centrad. Leg I from base of trochanter 25.2 (24.2 – 25.2); femur 10.2 (10.2 – 11.2), basiventral femoral seta bv 9.3 (8.9 – 9.3); genu 3.7 (3.7 – 4.2) with antaxial genual seta 1 ʺ 28.0 (27.0 – 28.0); tibia 6.5 (5.6 – 6.5) without paraxial tibial seta 1 ʹ; tarsus 4.6 (3.7 – 4.6); two identical tarsal setae-paraxial fastigial tarsal setae ft ʹ and antaxial fastigial tarsal seta ft ʺ 14.0 (11.3 – 14.0); paraxial unguinal tarsal seta u ʹ 3.7 (2.6 – 3.7), tarsal solenidion ω curved, knobbed and 4.6 (3.7 – 4.6), 4 rayed, tarsal empodium em 3.7 (3.7 – 4.4). Leg II from base of trochanter 24.2 (23.2 – 24.2); femur 10.2 (10.2 – 11.2), basiventral femoral seta bv 9.3 (8.9 – 9.3), genu 3.7 (3.7 – 4.2) with antaxial genual seta 1 ʺ 28.0 (27.0 – 28.0); tibia 5.6 (4.5 – 5.6) without paraxial tibial seta 1 ʹ; tarsus 3.7 (3.7 – 4.2) with two identical tarsal setae-paraxial fastigial tarsal setae ft ʹ and antaxial fastigial tarsal seta ft ʺ 14.0 (11.3 – 14.0), paraxial unguinal tarsal seta u ʹ 2.8 (2.1 – 2.8), tarsal solenidion ω curved, knobbed and 4.6 (4.1 – 4.6); 4 rayed, tarsal empodium em 3.7 (3.7 – 4.4). Coxae I 18.6 (17.5 – 18.6) long, smooth and contiguous with a mid sternal line, seta 1 b 6.5 (5.6 – 6.5) and 5.9 (5.1 – 5.9) apart; 1 a tubercles with seta present a little ahead of line across the 2 a tubercles; seta 1 a 14.0 (13.0 – 14.0) and 6.9 (6.1 – 6.9) apart, the distance between setae 1 a and 1 b is 6.2 (6.2 – 6.4); coxa II smooth, 13.0 (13.0 – 13.8), seta 2 a 29.8 (28.8 – 29.8) and 23.6 (22.8 – 23.6) apart. Opisthosoma with 29 (28 – 29) smooth dorsal annuli and 56 (56 – 57) narrow ventral annuli, micro tubercles rounded and located on first 15 (14 – 16) ventral annuli; rest of the ventral annuli starting from posterior margin of genitalia up to last ventral annulus have microstriations, seta c 2 15.8 (14.9 – 15.8) on annulus 15 (14 – 15), seta d 52.2 (52.2 – 53.2) on annulus 23 (23 – 24); seta e 4.6 (4.6 – 5.6) on ventral annulus 29 (28 – 29); seta f 14.0 (14.0 – 14.9) on ventral annulus 49 (48 – 49); seta h 1 absent, seta h 2 42.0 (42.0 – 43.2). Epigynium 16.8 (16.8 – 17.5) long, 19.6 (19.6 – 21.1) wide; smooth except small striations at anterior margin, two horn like lateral projections emerge from mid lateral margin of epigynium; seta 3 a 12.1 (10.2 – 12.1). MALE. Not observed.	en	Chakrabarti, Samiran, Sarkar, Sanjay (2011): Three new species of eriophyoid mites (Acari: Eriophyoidea) infesting fruit yielding plants from India. Zootaxa 2988: 28-36, DOI: 10.5281/zenodo.203722
374618679B43FF9151EAFF2BC686F839.taxon	materials_examined	Type material. Holotype: Female (marked) on slide (no. 1378 / 91 / 2006), India: West Bengal: Malda, Amriti, Latitude: 24 ° 20 ʹ 38 ʺ N and Longitude: 87 ° 05 ʹ 29 ʺ E, 24 December, 2006 from Mangifera indica (L.) (Anacardiaceae), Coll. S. Sarkar. Paratypes: 8 females on slide bearing holotype and 49 females on 6 slides (nos. 1376 - 1377 / 91 / 2006 and 1379 - 1382 / 91 / 2006); collection data same as in holotype. Relation to host. Pinkish brown, fusiform mites are vagrants on the undersurface of the leaves.	en	Chakrabarti, Samiran, Sarkar, Sanjay (2011): Three new species of eriophyoid mites (Acari: Eriophyoidea) infesting fruit yielding plants from India. Zootaxa 2988: 28-36, DOI: 10.5281/zenodo.203722
374618679B43FF9151EAFF2BC686F839.taxon	etymology	Etymology. The specific epithet ‘ fisus ’, a Latin word, meaning split, and refers to the bifurcate dorsal pedipalp genual seta.	en	Chakrabarti, Samiran, Sarkar, Sanjay (2011): Three new species of eriophyoid mites (Acari: Eriophyoidea) infesting fruit yielding plants from India. Zootaxa 2988: 28-36, DOI: 10.5281/zenodo.203722
374618679B43FF9151EAFF2BC686F839.taxon	discussion	Remarks. The new species resembles T. schleicherae Ghosh & Chakrabarti, 1985, T. ferrugeniae Mohanasundaram, 1985, T. tricarinatus Fletchmann, 1996 in having 4 rayed tarsal empodium and scapular tubercles much above the shield margin. However, it differs from T. schleicherae by not having granulated prodorsal shield, from T. ferrugeniae by not having lateral spine of dorsal annuli and from T. tricarinatus in overall prodorsal shield structure, shorter setae 3 a and smooth coxae. The new species also closely resembles T. convolvuli (Channabasavanna, 1966) in location of scapular seta well ahead of rear margin of prodorsal shield and having 4 rayed tarsal empodium but differs from it by having numerous irregular broken lines on prodorsal shield besides bifurcated dorsal pedipalp genual setae and not having seta on tibia I.	en	Chakrabarti, Samiran, Sarkar, Sanjay (2011): Three new species of eriophyoid mites (Acari: Eriophyoidea) infesting fruit yielding plants from India. Zootaxa 2988: 28-36, DOI: 10.5281/zenodo.203722
