taxonID	type	description	language	source
373F87D7FFBCFFADFF5FAD3FFE7D471C.taxon	materials_examined	Type species (by original designation): Conus selenae van Mol, Tursch & Kempf, 1967; Recent, Western Atlantic. Note. According to Tucker & Tenorio (2009), this genus is characterised by short and rounded shells with convex spire whorls with cords. Nodules on spire whorls are absent or become obsolete very early. The subsutural flexure is shallow and asymmetrical. In addition, we consider the presence of more or less prominent beads on the last whorl as a characteristic feature, being also present on the type species. The genus Artemidiconus was not treated by Puillandre et al. (2014 a, b).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFBCFFA6FF5FAEC8FCAB43EE.taxon	description	Conus Granularis — Borson 1820: 196, pl. 1, fig. 3. Conus (Stephanoconus) Stachei nov. form. — Hoernes & Auinger 1879: 16, pl. 6, figs 14 – 16. [Conus granuliferus Grat. var.] Drnowitzensis de Greg. — De Gregorio 1885: 376.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFBCFFA6FF5FAEC8FCAB43EE.taxon	description	Conus (Hemiconus) granularis Bors. var. stachei Hörn. u. Auin. — Meznerics 1932 / 1933: 346, pl. 14, fig. 2. Conus (Stephanoconus) stachei Hoernes et Auinger — Csepreghy-Meznerics 1956: 421, pl. 3, fig. 7.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFBCFFA6FF5FAEC8FCAB43EE.taxon	description	Conus (Hemiconus) granularis stachei Hoernes et Auinger — Csepreghy-Mezneric 1972: 34, pl. 17, figs 23 – 24.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFBCFFA6FF5FAEC8FCAB43EE.taxon	materials_examined	Type material. Lectotype (following ICZN Article 74.6), illustrated in Borson (1820, pl. 5, fig. 3) and Pavia (1976, pl. 2, fig. 12), stored in the Museo Regionale di Scienze Naturali, Torino. The type locality is given as Valle Andona (Italy), which is of Pliocene age. Pavia (1876) assumed that this was an error by Borson (1820) and pointed out that the preservation corresponds to material from the Messinian of Borelli (Turin Hills). The type localities of the taxa considered to be junior synonyms of A. granularis are: Conus stachei Hoernes & Auinger, 1879, Baden (Austria); Conus drnowitzensis De Gregorio, 1885, Drnovice u Vyškova (Czech Republic); Conus opellus De Gregorio, 1885, Baden (Austria); Conus lissitzensis Sacco, 1893, Lysice (Czech Republic); all are of middle Miocene (Badenian / Langhian) age. Studied material. 6 spec. NHMW 1864 / 0001 / 0441, Drnovice u Vyškova (Czech Republic), including holotype of Conus granuliferus drnowitzensis De Gregorio, 1885; 3 spec. NHMW 1865 / 0015 / 0004, Lysice (Czech Republic), including holotype of Conus granularis lissitzensis Sacco, 1893; 2 spec. NHMW 1865 / 0036 / 0103, Lysice (Czech Republic); 4 spec. NHMW 1871 / 0011 / 0001, Sedlec (Czech Republic); 1 spec. NHMW 1861 / 0040 / 0 0 28, Hrušovany (Czech Republic); 1 spec. NHMW 1999 z 0077 / 0025, Baden (Austria), designated herein as lectotype of Conus stachei Hoernes & Auinger, 1879, holotype of Conus granuliferus opellus De Gregorio, 1885; 20 spec. NHMW 1863 / 0015 / 0604, Niederleis (Austria); 1 spec. NHMW 1887 / 0018 / 0014, Steinebrunn (Austria), illustrated in Meznerics (1932 / 1933, pl. 14, fig. 2); 5 spec. NHMW 1846 / 0037 / 0038, 17 spec. NHMW 1868 / 0001 / 0 0 93, Steinebrunn (Austria). Illustrated material. Figs 3 A, 4 A 1 – A 3: Lysice (Czech Republic): SL: 15.1 mm, MD: 8.0 mm, NHMW 1865 / 0015 / 0004, specimen illustrated in Hoernes & Auinger (1879, pl. 6, fig. 16) (= holotype of Conus granularis lissitzensis Sacco, 1893); Figs 4 B 1 – B 3: Baden (Austria): SL: 15.2 mm, MD: 7.9 mm, NHMW 1999 z 0077 / 0025, specimen illustrated in Hoernes & Auinger (1879, pl. 6, fig. 14) (= holotype of Conus granuliferus opellus De Gregorio, 1885). Revised description. Small shells of about 13 – 15 mm in length, biconical, with moderately high conical spire and stout, ventricose last whorl; slightly constricted at base. Protoconch high conical, comprising at least three whorls. Early spire whorls forming a narrow, beaded upper sutural band and a tuberculate lower part. Subsequent spire whorls consisting of two adsutural spiral bands separated by a narrow groove; a secondary spiral cord appears between the spiral bands on late spire whorls; upper band usually bearing small beads. Subsutural flexure very shallow, moderately curved, nearly symmetrical. Last whorl weakly angulated in subadult shells, rounded in adults; broad and weak nodes appear along the shoulder in some specimens resulting in a wavy appearance. Widely spaced pustulose spiral ridges on lower half of last whorl grading into sharp spiral cords on ventral side. Upper half of last whorl with sharp spiral cords, which rarely bear beads. Base faintly constricted. Aperture straight, slightly broadening in abapical half. Fasciole short, moderately swollen, covered with spiral threads. No colour pattern observed.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFBCFFA6FF5FAEC8FCAB43EE.taxon	description	Shell measurements and ratios. n = 13 adult specimens; largest specimen: SL: 15.2 mm, MD: 7.9 mm, mean SL: 13.6 mm (σ = 0.8), mean MD: 7.2 (σ = 0.4), spire angle: µ = 68 ° (σ = 2.9 °), last whorl angle: µ = 41 ° (σ = 4.6 °), LW: µ = 1.9 (σ = 0.07), RD: µ = 0.74 (σ = 0.04), PMD: µ = 0.9 (σ = 0.05), RSH: µ = 0.29 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFBCFFA6FF5FAEC8FCAB43EE.taxon	discussion	Discussion. This species is synonymized with the Italian early Miocene to Pliocene Conus granularis (Borson, 1820) (e. g. Bałuk 1997). Conus granuliferus Grateloup, 1835, from the early Miocene of the Aquitaine Basin, was also considered a synonym of C. granularis by most authors (e. g. Hall 1966, Lozouet et al. 2001). The specimens from the Badenian of the Paratethys seemingly differ from the Italian and French specimens in their mode of spire formation. Hall (1966) and Davoli (1972) described and figured a spire sculpture consisting of two cords separated by a deep groove. The same sculpture is seen in the somewhat abraded holotype of Conus granularis as illustrated in Pavia (1976) and in the French specimen illustrated in Lozouet et al. (2001). The Paratethyan shells, however, develop a secondary cord on the groove separating the two main cords. Some specimens of A. granularis from the Tortonian of Italy in the NHMW-collection, however, show a clear secondary cord as well. Therefore, this feature seems to reflect intraspecific variability and consequently we do not separate the Paratethyan specimens as a distinct species. Sacco (1893 b) placed the Italian Conus granularis Borson, 1820 in Hemiconus Cossmann, 1889, which in our opinion is an exclusively Eocene genus. Consequently, Friedberg (1911) and Bałuk (1997) treated the Paratethyan species also as Hemiconus. Its type species, Hemiconus stromboides (Lamarck, 1803), from the Eocene of the Paris Basin, is high spired fusiform species with blunt nodes on the spire whorls. Other species placed in Hemiconus by Cossmann & Pissarro (1911) show a similar spire sculpture of nodes and a granulose cord along the upper suture. These morphologies have nothing in common with the Miocene species and therefore the placement in Hemiconus is herein rejected. Tucker & Tenorio (2009) listed Conus granularis Borson, 1820 within Conilithes Swainson, 1840, but the mode of spire formation in C. granularis is different from the scalariform spire of Conilithes. Moreover, the spirally elongate beads on the last whorl of C. granularis do not occur in any Conilithes species and we follow the recommendation by John K. Tucker (pers. comm.) to treat Conus stachei (= granularis) tentatively as Artemidiconus, despite the large geographic gap between the European Miocene record and the extant Western Atlantic occurrence.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFBCFFA6FF5FAEC8FCAB43EE.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Baden, Steinebrunn, Niederleis (Austria), Hrušovany nad Jevišovkou, Sedlec (Czech Republic) (Hoernes & Auinger 1879; Sieber 1956); Bükk Mountains: Borsodbóta (Hungary); Pannonian Basin: Szob (Hungary) (Csepreghy-Mecnerics 1956; Strausz 1966; Kovács & Vicián 2013); Carpathian Foredeep: Drnovice u Vyškova, Lysice (Czech Republic), Korytnica, Małoszów, Dryszczów (Poland), Sboriw (Ukraine) (Hoernes & Auinger 1879; Zelinskaya et al. 1968; Bałuk 1997); Transylvanian Basin: Coşteiu de Sus, Lăpugiu de Sus (Romania) (Hoernes & Auinger 1879; Boettger 1902); Caransebeş-Mehadia Basin: Valea Bela Reca (Romania) (Hinculov 1968); Dacian Basin: Portitovci (Bulgaria) (Kojumdgieva & Strachimirov 1960); southern Pannonian Basin: Hrvaćani (Bosnia and Herzegovina) (Atanacković 1985). Proto-Mediterranean Sea and north eastern Atlantic. Aquitanian (France: Aquitaine Basin) (Lozouet et al. 2001), Burdigalian (Italy: Colli Torinesi; France Aquitaine Basin) (Peyrot 1931; Hall 1966), Tortonian (Italy: Sant'Agata Fossili, Stazzano Montegibbio) (Davoli 1972).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB7FFA6FF5FAB68FB9E44B7.taxon	materials_examined	Type species (by subsequent designation, Tucker & Tenorio 2009): Conus pagoda Kiener, 1847. Recent, Pacific, Philippines. Note. According to Tucker & Tenorio (2009) the genus Conasprella Thiele 1929 is characterised by relatively small-shelled species, the spire is relatively elevated, with beaded early whorls. Cords may be present or absent on the spire whorls, the last whorl has sulci that reach mid-whorl and often the shoulder; the subsutural sinus is deep; an anterior notch is absent; and the protoconch is multispiral. This genus has a Miocene to Recent Indo-Pacific distribution. It is similar in shell characters to the Western Atlantic genus Jaspidiconus Petuch 1993, which also has a Miocene to Recent record in tropical America (Landau & da Silva 2010), but differs in having tubercles developed along the shoulder carina of the last whorl. Jaspidiconus, together with Perplexiconus Tucker & Tenorio, 2009, were considered synonyms of Ximeniconus Emerson & Old, 1962 by Puillandre et al. (2014 a), although one can see that each of these groups is distinct, with Perplexiconus and Ximeniconus more closely related than Jaspidiconus. Conus jaspideus Gmelin, 1791, the type species of Jaspidiconus, has a symmetrical curved subsutural flexure (Hendricks 2009), requiring a revised diagnosis of Conasprella, if Jaspidiconus is considered as synonym of it, although it is more likely that they are distinct genera; Conasprella is restricted to the Indo-Pacific, while Jaspidiconus is tropical American. All Paratethyan species placed in Conasprella herein have tuberculate early spire whorls and beaded cords on the last whorl. Their subsutural flexures are shallow, moderately to strongly curved and moderately asymmetrical. This is an interesting record as it suggests, amongst others, a Paratethyan occurrence of a taxon nowadays restricted to the Indo-West Pacific.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB7FFA7FF5FAE59FD80467A.taxon	description	Conus (Leptoconus) Berwerthi nov. form. — Hoernes & Auinger 1879: 35, pl. 5, figs 11 – 12.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB7FFA7FF5FAE59FD80467A.taxon	materials_examined	Type material. Syntype NHMW 1860 / 0001 / 0074 a, Steinebrunn (Austria), illustrated in Hoernes & Auinger (1879, pl. 5, fig. 11); syntype NHMW 1860 / 0001 / 0074 b, Steinebrunn (Austria), illustrated in Hoernes & Auinger (1879, pl. 5, fig. 12); middle Miocene, Badenian (late Langhian). Studied material. Syntypes and 1 spec. NHMW 1846 / 0037 / 0038, Steinebrunn (Austria), illustrated in Hörnes (1851, pl. 5, figs 4 a – c), holotype of Conus excatenatus Sacco, 1893; 1 spec. NHMW 1860 / 0001 / 0074, Steinebrunn (Austria), illustrated in Hörnes (1851, pl. 5, fig. 4); 1 spec. NHMW 2015 / 0416 / 0001, 1 spec. NHMW 2015 / 0416 / 0002; 1 spec. NHMW 2015 / 0416 / 0003, Steinebrunn or Gainfarn (Austria). Illustrated material. Figs 4 C 1 – C 2: Steinebrunn (Austria): SL: 10 mm, MD: 4.7 mm, NHMW 1860 / 0001 / 0074 b, illustrated in Hoernes & Auinger (1879, pl. 5, fig. 12); Figs 4 D 1 – D 3: Steinebrunn or Gainfarn (Austria): SL: 20.5 mm, MD: 9.4 mm, NHMW 2015 / 0416 / 0002; Figs 4 E 1 – E 3: Steinebrunn or Gainfarn (Austria): SL: 20.9 mm, MD: 9.7 mm, NHMW 2015 / 0416 / 0001; Figs 3 B, 4 F 1 – F 4: Steinebrunn (Austria): SL: 22.7 mm, MD: 11.8 mm, NHMW 1846 / 0037 / 0038, illustrated in Hörnes (1851, pl. 5, figs 4 a – c). Revised description. Small biconical shells of about 20 – 23 mm height; protoconch high conical, comprising at least three whorls. Spire height moderate to high, coeloconoid. Whorls with broad, straight to weakly concave sutural ramp, periphery placed a short distance above suture, below whorl tapering in towards lower suture. Shoulder bearing more or less well defined beads, starting at carina and extending to suture during ontogeny. Delicate and finely beaded spiral threads appear below shoulder on penultimate whorls. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Last whorl conical with 13 to 16 wide spaced spiral rows of small but prominent, spirally elongate beads. Beads merge into spiral ribs close to siphonal canal and fasciole. Distinct growth lines between the glossy spiral rows. Base weakly constricted. Siphonal fasciole narrow and indistinct; aperture with parallel margins, moderately narrow. Siphonal canal short, weakly twisted, poorly delimited from base. Colour pattern in UV light consisting of light dots coinciding with the beads on last whorl; short opisthocline stripes, following the subsutural flexure, appear close below the shoulder and merge with the dots on the uppermost spiral row of beads.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB7FFA7FF5FAE59FD80467A.taxon	description	Shell measurements and ratios. n = 4 adult specimens: largest specimen: SL: 22.7 mm, MD: 11.8 mm, mean SL: 21.1 mm (σ = 1.1), mean MD: 10.2 mm (σ = 1.1), spire angle: µ = 78 ° (σ = 14.9 °), last whorl angle: µ = 32.3 ° (σ = 2.2 °), LW: µ = 2.1 (σ = 0.1), RD: µ = 0.6 (σ = 0.05), PMD: µ = 0.9 (σ = 0.04), RSH: µ = 0.2 (σ = 0.05).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB7FFA7FF5FAE59FD80467A.taxon	discussion	Discussion. Hoernes & Auinger (1879) doubted that the specimen illustrated by Hörnes (1851, pl. 5) as Fig. 4 was conspecific with the specimen illustrated by Hörnes (1851, pl. 5, figs 4 a – c) as Conus catenatus and separated it as Conus berwerthi. They based the separation on the smaller size, the more slender shape and the lack of pustulose spiral threads in the upper half of the last whorl of C. berwerthi. Sacco (1893 a) recognized that the Viennese Miocene Conus catenatus of Hörnes (1851) and Hoernes & Auinger (1879) was not conspecific with the extant American C. catenatus Sowerby III, 1 879 [= Tenorioconus granarius (Kiener, 1847)] and proposed excatenata as new variation name. In our opinion, the differences between both morphologies are only ontogenetic stages of a single species. Conus berwerthi is based on three juvenile to subadult shells, which lack the broad and strongly sculptured last whorl of C. excatenatus. Moreover, both types were collected at the same locality. Consequently, Conus berwerthi gains priority over C. excatenatus. The Paratethyan species is also not conspecific with the American Conus catenatus Sowerby I, 1850, which has concave spire whorls. A very rare species, which is easily recognized by its characteristic beaded sculpture. Paleoenvironment. The Austrian occurrences point to shallow sublittoral settings as the preferred paleoenvironment of this species.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB7FFA7FF5FAE59FD80467A.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn, Steinebrunn, Drasenhofen (Austria); Eisenstadt-Sopron Basin: Marz, Mattersburg (Austria) (Hoernes & Auinger 1879; Sieber 1958 b); Alpine-Carpathian Foredeep: Grund (Austria), Drnovice u Vyškova (Czech Republic), Sboriw (Ukraine) (Hoernes & Auinger 1879; Friedberg 1938; Sieber 1947); Transylvanian Basin: Lăpugiu de Sus (Romania) (Hoernes & Auinger 1879; Moisescu 1955 b).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB6FFA4FF5FAF15FCEF4478.taxon	materials_examined	Holotype: Figs 4 I 1 – I 5: SL: 13.1, MD: 6.3 mm, NHMW 1869 / 0001 / 0331 a, Steinebrunn (Austria). Paratype: Figs 4 H 1 – H 3: SL: 10.6, MD: 5.1 mm, NHMW 1869 / 0001 / 0331 b, Steinebrunn (Austria).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB6FFA4FF5FAF15FCEF4478.taxon	description	Paratype: Figs 3 E, 4 G 1 – G 3: SL: 10.4, MD: 4.7 mm, NHMW 1869 / 0001 / 0331 c, Steinebrunn (Austria).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB6FFA4FF5FAF15FCEF4478.taxon	materials_examined	Additional material: 2 spec. NHMW 1869 / 0001 / 0331 d, 1 spec. NHMW 1858 / 0029 / 0017, Steinebrunn (Austria); 1 spec. NHMW 1870 / 0054 / 0044, Niederleis (Austria); 1 spec. NHMW 1869 / 0001 / 0627, Marz (Austria). Type stratum: Bioclastic sand and marl of the Baden Group. Type locality: Steinebrunn (Austria). Age: Middle Miocene, middle Badenian (= late Langhian).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB6FFA4FF5FAF15FCEF4478.taxon	etymology	Etymology. Referring to the small size.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB6FFA4FF5FAF15FCEF4478.taxon	description	Description. Very small, slender biconical shells. High, scalariform spire of at least 7 teleoconch whorls. Spire whorls carinate just above suture, weakly tuberculate; sutural ramp distinctly striate. Impressed, somewhat irregular suture. Last whorl with flat to faintly concave sutural ramp, weakly striate. Subsutural flexure shallow, strongly curved, moderately asymmetrical. Last whorl elongate conical to weakly ventricose below angulated shoulder; slightly constricted at base. Aperture with subparallel margins; short wide siphonal canal. Siphonal fasciole obsolete. Sculpture consisting of about 15 – 20 spiral cords with spirally elongate beads. Spiral sculpture overlays much weaker growth lines forming a faintly cancellate pattern. Spirals become crowded on base where beads are only weakly developed. Colour pattern in UV light consisting of a few large, widely-spaced subquadratic blotches on sutural ramp of last whorl and weaker blotches on spire whorls; beads on last whorl appear as dots on thin spirals. Shell measurements and ratios. n = 5: largest specimen: SL: 13.1 mm, MD: 6.3 mm, mean SL: 10.4 (σ = 0.8), mean MD: 5.1 (σ = 0.7), spire angle: µ = 63.2 ° (σ = 8.5 °), last whorl angle: µ = 32.4 ° (σ = 2.4 °), LW: µ = 2.04 (σ = 0.13), RD: µ = 0.68 (σ = 0.03), PMD: µ = 0.81 (σ = 0.09), RSH: µ = 0.28 (σ = 0.04).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB6FFA4FF5FAF15FCEF4478.taxon	discussion	Discussion. This tiny species was mixed with Artemidiconus stachei in the NHMW collections, probably due to its small size and beaded sculpture. The scalariform spire and the lack of spiral cords on the spire whorls, however, allow a clear separation. Conasprella berwerthi (Hoernes & Auinger, 1879) develops a similar but coarser sculpture on the last whorl. Moreover, it differs in its lower spire, the much broader last whorl and it lacks subquadratic blotches on the sutural ramp. Phasmoconus ottiliae (Hoernes & Auinger, 1879) lacks the scalariform spire outline, has an elongate last whorl, a denser spiral sculpture, lacks the prominent beads and differs in its colour pattern of dense spirals. Paleoenvironment. The occurrence at Steinebrunn suggests seagrass meadows in shallow marine nearshore settings as habitat.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB6FFA4FF5FAF15FCEF4478.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Niederleis (Austria), Steinebrunn (Austria); Eisenstadt Sopron Basin: Marz, Forchtenau (Austria).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB5FFA4FF5FAD13FD2A4605.taxon	materials_examined	Type species (by monotypy): Conus antidiluvianus Bruguière, 1792. Miocene-Pliocene, Europe. Note. According to Tucker & Tenorio (2009) this genus is characterised by species with a scalariform spire, the shoulder is angular and carinate and often bearing tubercles, but devoid of spiral sculpture. The subsutural flexure is deep and symmetrical. This is an extinct genus, with an Eocene to Pliocene stratigraphical distribution and is found in Europe and North America. Tucker & Tenorio (2009) erected the subfamily Conilithinae for this and other related genera. The molecular phylogeny presented by Puillandre et al. (2014) confirmed this deeply rooted phylogenetic group and used the genus Conasprella Thiele, 1929, with related groups relegated to subgenus level. Based on the fossil Paratethyan material, we can confirm this generic shell description. The shells have a tall to very tall scalariform, carinate spire, with beading at least on the earliest teleoconch whorls but devoid of spiral sculpture. Last whorl without beads; along with typical spiral cords, a reticulate pattern may rarely be formed. The subsutural flexures are deep (or rarely moderately deep), strongly curved and moderately asymmetrical and the last whorl relative diameter is medium to elongate.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB4FFA5FF5FA8E1FB0C46EC.taxon	description	Conus Allioni mihi — Michelotti 1847: 338, pl. 17, fig. 17. Conus oblitus mihi — Michelotti 1847: 340, pl. 14, figs 2 – 2 a. [Conus] Leptoconus Allionii [sic] (Micht.) — Sacco 1893 a: 32, pl. 4, fig. 6.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB4FFA5FF5FA8E1FB0C46EC.taxon	description	Conus brochii [sic] Bronn, 1828 — Zunino & Pavia 2009: 365, pl. 2, fig. 1 [non Conilithes brocchii (Bronn, 1828)].	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB4FFA5FF5FA8E1FB0C46EC.taxon	materials_examined	Type material. Syntype or holotype illustrated by Michelotti (1847, pl. 17, fig. 17), Turin Hills, Italy; the specimen might have been stored in the collections of the Dipartimento di Scienze della Terra, Sapienza Università di Roma, but parts of the Michelotti collection were destroyed during World War II (Manni 2005); early Miocene, Burdigalian. Studied material. 6 spec. NHMW 1976 / 1785 / 0044, Kleinebersdorf (Austria); 8 spec. NHMW 1976 / 1785 / 0 0 47, Kleinebersdorf (Austria); holotype of Conus raulini, Saint-Jean-de-Marsacq (France), SL: 32 mm, MD: 18.5 mm Neuville collection, Université Bordeaux n ° 30.2.7. Illustrated material. Figs 5 A 1 – A 3: Kleinebersdorf (Austria): MD: 16.1 mm, NHMW 1976 / 1785 / 0044. Revised description. Small shells with moderately low conical spire; early spire with beaded carina. Late spire whorls smooth, deeply concave near upper suture and with prominent rim at lower suture; suture narrowly incised. Subsutural flexure of medium depth, strongly curved, moderately asymmetrical. Last whorl elongate conical, slightly constricted at base. A few deep and broad spiral grooves are restricted to the base. Siphonal fasciole very weak. Aperture largely destroyed; no information on colour patterns.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB4FFA5FF5FA8E1FB0C46EC.taxon	description	Shell measurements and ratios. n = 7: largest specimen (fragmented): MD: 16.1 mm, mean SL: 23.9 mm (σ = 1.9), mean MD: 12.7 (σ = 1.0), spire angle: µ = 100.1 ° (σ = 5.7 °), last whorl angle: µ = 33.7 ° (σ = 1.1 °), LW: µ = 1.89 (σ = 0.06), RD: µ = 0.66 (σ = 0.02), PMD: µ = 0.9 (σ = 0.04), RSH: µ = 0.21 (σ = 0.02).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB4FFA5FF5FA8E1FB0C46EC.taxon	discussion	Discussion. We do not follow Hall (1966), who synonymised Conus allioni with Conus brocchii Bronn, 1828. Conilithes brocchii is a Pliocene species, which differs in its larger size, higher and sometimes slightly coeloconoid spire, prominent sutural ridge and the broader last whorl, lacking the slight constriction of C. allioni, which has a low and sometimes cyrtoconoid spire. Conus raulini Peyrot, 1931, based on a specimen from the Burdigalian of the Aquitaine Basin, is most probably a subjective junior synonym of Conus allioni. This species was introduced by Michelotti (1847) as Conus Allioni. Therefore, the prevailing use as Conus allionii is an incorrect subsequent spelling. Paleoenvironment. The Kleinebersdorf section represents coastal mudflat environments with Avicennia - mangroves (Zuschin et al. 2004).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB4FFA5FF5FA8E1FB0C46EC.taxon	distribution	Distribution in Paratethys.? Ottnangian (early Miocene): North Alpine Foreland Basin: Bad Tölz (Germany) (Steininger 1973); Karpatian (early Miocene): Korneuburg Basin: Kleinebersdorf (Austria). Proto-Mediterranean Sea and North eastern Atlantic. Burdigalian (early Miocene): Aquitaine Basin: Saint- Jean-de-Marsacq, Saubrigues (France) (Peyrot 1931); Turin Hills: Baldissero, Valle Ceppi (Italy) (Sacco 1893 a; Zunino & Pavia 2009). Occurrences from the early and middle Miocene of the North Sea Basin, mentioned by Beyrich (1853) and Kautsky (1925) seem to represent another species (see Janssen 1984).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB2FFA1FF5FAEE3FBD842A2.taxon	description	Conus antediluvianus Brug. — Hörnes 1851: 38, pl. 5, figs 2 a – e.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB2FFA1FF5FAEE3FBD842A2.taxon	materials_examined	Type material. Neotype: collection of Museo Civico di Storia Naturale (Milano), registration number MSNM i 28027 Badagnano, Rio dei Carbonari (Italy), designated by Janssen et al. (2014); Pliocene. Studied material. 3 spec. NHMW A 1615, Windpassing (Austria); 5 spec. NHMW 1973 / 1615 / 0300, 4 spec. NHMW 1989 / 0089 / 0058, 14 spec. NHMW 2013 / 0078 / 0524, 4 spec. NHMW 1869 / 0001 / 0161, Baden (Austria), including illustrated specimens of Hörnes (1851, pl. 5, figs 2 a – e); 4 spec. NHMW 1997 z 0178 / 1615, 5 spec. NHMW 1872 / 0030 / 0019, 13 spec. NHMW 2013 / 0300 / 0524, Baden-Sooß (Austria); 11 spec. NHMW 1855 / 0045 / 0 860, 30 spec. NHMW 2010 / 0004 / 1338, 3 spec. NHMW 1970 / 1396 / 1225, 4 spec. NHMW 1997 z 0178 / 1199, 15 spec. NHMW 2010 / 0004 / 1338, Bad Vöslau (Austria); 10 spec. NHMW 2012 / 0229 / 0223, Traiskirchen (Austria); 4 spec. NHMW 2013 / 0479 / 1604, Gainfarn (Austria); 9 spec. NHMW 1997 z 0178 / 1590, 20 NHMW spec. NHMW 1869 / 0001 / 0247, Möllersdorf (Austria), including specimen illustrated in Schultz (1998, pl. 29, fig. 9). Illustrated material. Figs 5 J 1 – J 3: Baden (Austria): SL: 56.3 mm, MD: 20.5 mm, NHMW 1869 / 0001 / 0161, illustrated in Hörnes (1851, pl. 5, fig. 2); Fig. 5 K 1: Möllersdorf (Austria): SL: 66.9 mm, MD: 23.7 mm, NHMW 1869 / 0001 / 0247; Figs 6 A 1 – A 3: Bad Vöslau (Austria): SL: 61.6 mm, MD: 23.0 mm, NHMW 2010 / 0004 / 1338; Fig. 3 C: Möllersdorf (Austria): SL: 25.6 mm, MD: 9.1 mm, NHMW 1869 / 0001 / 0247.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB2FFA1FF5FAEE3FBD842A2.taxon	description	Description. Medium sized to moderately large, elongate shells. Protoconch high and conical comprising 3.5 smooth, moderately convex whorls with delicate spiral thread at upper suture; depressed, slightly sunken nucleus. First teleoconch whorl with indistinct angulation passing into a carina with blurred nodes on the first 2 spire whorls. Teleoconch consisting of 9 – 11 whorls; spire moderately high; gradate to scalariform with distinctly beaded carina placed below mid-whorl. Maximum diameter at carina or slightly below. Sutural ramp flat to slightly concave, devoid of spiral sculpture, bearing densely spaced, raised growth lines. Two to four spiral threads below carina may intersect the slightly axially elongated beads. The beads are very prominent and regular on early spire whorls and may become subobsolete on the last two teleoconch whorls. Last whorl elongate conical, weakly constricted at base; siphonal canal long, slightly twisted. Outer, lip simple, straight, prosocyrt; columella straight, columella callus not thickened, restricted to narrow glossy margin. Deep spiral grooves on lower third of last whorl; apical part of last whorl glossy and faintly striate in some specimens. Shell measurements and ratios. n = 17 adult specimens: largest specimen: SL: 81.9 mm, MD: 30.1 mm, mean SL: 59.2 mm (σ = 2.4), mean MD: 21.8 mm (σ = 4.6), spire angle: µ = 61.4 ° (σ = 7.0 °), last whorl angle: µ = 29.4 ° (σ = 1.4 °), LW: µ = 2.69 (σ = 0.2), RD: µ = 0.52 (σ = 0.03), PMD: µ = 0.88 (σ = 0.05), RSH: µ = 0.27 (σ = 0.04).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB2FFA1FF5FAEE3FBD842A2.taxon	discussion	Discussion. The complex taxonomic history of this species was discussed in great detail by Janssen et al. (2014), who proposed a neotype from the Pliocene of Badagnano (Italy). Still, however, it remains to be proven if all the Miocene and Pliocene specimens listed as C. antidiluvianus are conspecific. The adult shells of Pliocene Mediterranean specimens range around 69.7 mm (n = 14, σ = 10.8, NHMW collection) and are distinctly larger than those from the middle Miocene Paratethys. A further difference is the regularly beaded sculpture of the carina on the earliest teleoconch whorls in C. antidiluvianus from the Pliocene of Italy and Sicily, whereas the majority of Paratethyan specimens develop a carina with indistinct and more irregular nodes. The protoconch of Miocene North Sea specimens develop 5.5 whorls (Janssen et al. 2014), whereas the Pliocene Italian ones have 3.5 whorls (own data and Muñiz-Solís 1999). Similarly, the few Paratethyan Miocene specimens with preserved protoconch develop only 3.5 protoconch whorls. This may indicate that that the North Sea specimens represent a closely related but distinct species. A statistical analysis of the shell ratios (LW, RD, PMD, RSH) of Pliocene Mediterranean and Miocene Paratethyan specimens, however, does not support a clear separation of both groups. Moreover, establishing a new (chrono-sub) species name for the Paratethyan specimens is not advisable given the large number of available names introduced by Sacco (1893 a) for Miocene specimens from Italy. Therefore, we refrain from separating the Miocene specimens as a distinct taxon. Two subspecies names are found in Paratethyan literature: Moisescu (1955 a) introduced Conus antidiluvianus buiturica for an incomplete specimen from Bujtur in Romania. This specimen is clearly unrelated to Conilithes antidiluvianus and might rather represent a stout C. exaltatus (Eichwald, 1830), which lacks the anterior canal. Conus antediluvianus [sic] anomalus was proposed by Csepreghy-Meznerics (1972) for a single fragment from the Bükk Mountains in Hungary, without description. Therefore, Conus antidiluvianus anomalus is a nomen nudum. The specimen is most probably an aberrant C. exaltatus with a high spire. Paleoenvironment. The species is mainly found in offshore clays suggesting middle shelf to upper bathyal settings (e. g. Harzhauser et al. 2011).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB2FFA1FF5FAEE3FBD842A2.taxon	distribution	Distribution in Paratethys. Ottnangian (early Miocene): North Alpine Foreland Basin: Ottnang-Schanze (Austria) (Hoernes 1875; Steininger 1973); Karpatian (early Miocene): Vienna Basin: Cerova (Slovakia) (Harzhauser et al. 2011); Badenian (middle Miocene): Vienna Basin: Bad Vöslau, Baden, Baden-Sooß, Möllersdorf, Niederleis (Austria), Hrušovany, Rudice (Czech Republic); Alpine-Carpathian Foredeep: Grund, Windpassing (Austria), Lysice, Jaroměřice nad Rokytnou (Czech Republic); Eisenstadt-Sopron Basin: Forchtenau (Austria) (Hoernes & Auinger 1879; own data); Pannonian Basin: Balaton, Borsodbóta, Csermely, Csokvaomány, Hidas, Letkés (Hungary) (Strausz 1966; Csepreghy-Meznerics 1972; Kovács & Vicián 2013); southern Pannonian Basin: Miljevići (Bosnia and Herzegovina) (Atanacković 1969); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus (Romania) (Chira & Voia 2001; own data); Dacian Basin: Staropatica, Opanec (Bulgaria) (Kojumdgieva & Strachimirov 1960). Proto-Mediterranean Sea and north eastern Atlantic. Conilithes antidiluvianus (Bruguière, 1792) sensu stricto occurs in the Tortonian of Italy (Sant'Agata Fossili, Stazzano, Montegibbio) (Sacco 1893 a; Davoli 1972) and becomes very common at Pliocene localities in Italy (Hall 1966; Davoli 1972; Janssen et al. 2014). Further Pliocene occurrences are mentioned from southern France (Biot), the Hatay Basin in Turkey, Sicily and Syria (Erünal-Erentöz 1958; Janssen et al. 2014) (see Robba 1968 for further occurrences).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB0FFBFFF5FAC6AFC0942CF.taxon	description	Conus (Leptoconus) Brezinae nov. form. — Hoernes & Auinger 1879: 36. Conus Brezinae R. Hoern. i Auing. — Friedberg 1911: 51, pl. 2, figs 13 – 14. Conus (Conospira) dujardini Desh. — Strausz 1954: 113, pl. 7, fig. 144. Conus (Conospira) dujardini brezinae H. et Au. — Strausz 1954: 113, pl. 4, fig. 80. Conus (Conuspira) brezinae R. Hörnes und Auinger, 1879 — Moisescu 1955: 161, pl. 14, figs 9, 12 – 14.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB0FFBFFF5FAC6AFC0942CF.taxon	materials_examined	Type material. Syntype NHMW 1999 z 0077 / 0023 a, illustrated in Hörnes (1851, pl. 5, fig. 8 a); syntype NHMW 1999 z 0077 / 0023 b, illustrated in Hörnes (1851, pl. 5, fig. 8 b); syntype NHMW 1999 z 0077 / 0023 c, illustrated Hörnes (1851, pl. 5, fig. 8 c); syntype NHMW 1999 z 0077 / 0023 d, illustrated in Hörnes (1851, pl. 5, fig. 8 d); syntype NHMW 1999 z 0077 / 0023 e, illustrated in Hörnes (1851, pl. 5, fig. 8 e), all Steinebrunn (Austria); middle Miocene, Badenian (late Langhian). Studied material. Syntypes and 37 spec. NHMW 1846 / 0037 / 0040, 25 spec. NHMW 1846 / 0037 / 0039, Gainfarn or Enzesfeld (Austria); 12 spec. NHMW 1853 / 0010 / 0008, 9 spec. NHMW A 1614, Enzesfeld (Austria), including specimen illustrated in Schultz (1998 pl. 29, fig. 10); 2 spec. NHMW 1997 z 0178 / 1848, 3 spec. NHMW 1985 / 0083 / 0159 a,> 100 spec. NHM 2013 / 0479 / 1583 - 1602, Gainfarn (Austria); 10 spec. NHMW A 451, 29 spec. NHMW 1860 / 0001 / 0076, Steinebrunn (Austria); 2 spec. NHMW 2012 / 0213 / 0104, Pfaffstätten (Austria); 11 spec. NHMW A 994, Guntersdorf (Austria); 16 spec. NHMW A 995, Grund (Austria); 24 spec. NHMW 1854 / 0035 / 0 0 54, Lăpugiu de Sus (Romania). Illustrated material. Figs 5 D 1 – D 3: Steinebrunn (Austria), SL: 37.9 mm, MD: 15.9 mm, syntype NHMW 1999 z 0077 / 0023 a, illustrated in Hörnes (1851, pl. 5, fig. 8 a); Figs 5 E 1 – E 3: Steinebrunn (Austria), SL: 36.9 mm, MD: 14.5 mm, syntype NHMW 1999 z 0077 / 0023 b, illustrated in Hörnes (1851, pl. 5, fig. 8 b); Figs 5 F 1 – F 3: Steinebrunn (Austria), SL: 35.1 mm, MD: 14.1 mm, syntype NHMW 1999 z 0077 / 0023 c, illustrated in Hörnes (1851, pl. 5, fig. 8 c); Figs 5 G 1 – G 3: Gainfarn or Enzesfeld (Austria): SL: 37.3 mm, MD: 17.8 mm, NHMW 1846 / 0037 / 0039; Fig. 5 H 1: Gainfarn or Enzesfeld (Austria); SL: 36.2 mm, MD: 16.1 mm, NHMW 1846 / 0037 / 0039; Fig. 3 F: Steinebrunn (Austria), SL: 26.6 mm, MD: 12.2 mm, syntype NHMW 1999 z 0077 / 0023 e, illustrated in Hörnes (1851, pl. 5, fig. 8 e). Revised description. Moderately small to medium sized, biconical shells, consisting of 10 – 12 teleoconch whorls. Protoconch high, conical and multispiral. Very high conical spire; early spire whorls with weakly tuberculate angulation just above suture; nodes become obsolete on 3 rd to 4 th spire whorl grading into a prominent carina, which successively migrates towards the lower suture abapically. Sutural ramp flat to slightly concave with prominent, raised growth lines. Subsutural flexure deep, strongly curved, moderately asymmetrical. Faint spiral threads appear on the sutural ramp of the early spire whorls in a few specimens. Last whorl slightly allometric with wider and less steep sutural ramp; weakly ventricose with slight constriction at base; siphonal canal very short and moderately wide. Aperture broadening slightly abapically, with thin, prosocyrt outer lip. Surface smooth, except for deep, broad spiral grooves on base, confined to the lower quarter of last whorl. Colour pattern in UV light consisting of irregularly spaced thin spiral lines on last whorl intercalated by 2 – 3 spirals of delicate dots. These stripes appear on the sutural ramp parallel to the subsutural flexure. Beads on spire coincide with short prosocyrt dots.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB0FFBFFF5FAC6AFC0942CF.taxon	description	Shell measurements and ratios. n = 25: largest specimen: SL: 37.9 mm, MD: 17.8 mm, mean SL: 32.8 mm (σ = 2.9), mean MD: 14.3 (σ = 1.5), spire angle: µ = 58.4 ° (σ = 5.2 °), last whorl angle: µ = 32.7 ° (σ = 2.4 °), LW: µ = 2.3 (σ = 0.13), RD: µ = 0.63 (σ = 0.03), PMD: µ = 0.9 (σ = 0.03), RSH: µ = 0.31 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB0FFBFFF5FAC6AFC0942CF.taxon	discussion	Discussion. There is considerable confusion about the taxonomic status of Conus exaltatus Eichwald, 1830, Conus dujardini Deshayes, 1845 and Conus brezinae Hoernes & Auinger, 1879 and in the literature all combinations of synonymizations can be found. The validity of C. brezinae as distinct species was especially doubted by many authors (e. g. Bałuk 1997; Landau et al. 2013; Kovács & Vicián 2013). After a re-examination of the type material we reject this decision. A principal components analysis (PCA) of the shell ratios (Fig. 7 A) and measurements (Fig. 7 B) reveals a very clear separation of Conilithes brezinae from C. exaltatus. Despite some variability in slenderness and spire height, C. brezinae is characterized by its marked shoulder and somewhat broader sutural ramp of the last whorl, the absence of striae or nodes on late spire whorls and the spiral grooves confined to the base. Conilithes brezinae further differs from C. exaltatus in the lower position of the carina, which is also less sharp, the lower height of the spire whorls, the shorter last whorl, which lacks a pronounced basal constriction, and the lack of the punctate spiral grooves. With respect to this re-definition, the Serravallian specimens from the Turkish Karaman Basin, described by us (Landau et al. 2013) as Conilithes dujardini [which we consider to be a subjective junior synonym of Conilithes exaltatus (Eichwald, 1830), see below], should be treated as C. brezinae. Kovács & Vicián (2013) proposed to synonymize Conilithes brezinae with the Pliocene C. canaliculatus (Brocchi, 1814). We do not accept this conclusion because C. canaliculatus has a more ventricose last whorl, its maximum diameter is below the shoulder, the sutural ramp of the last whorl is narrower and the spiral grooves cover a larger part of the base (see Malatesta 1974; Pinna & Spezia 1978; Chirli 1997). Paleoenvironment. The occurrences in the Vienna Basin represent shallow water settings with sea grass meadows (e. g. Gainfarn; Zuschin et al. 2007). This ecological preference is an additional argument for a separation from C. exaltatus, which is typically found in offshore clays.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFB0FFBFFF5FAC6AFC0942CF.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn, Enzesfeld, Steinebrunn, Grinzing, Pötzleinsdorf, Bad Vöslau, Niederleis (Austria), Mikulov, Kienberk, Hrušovany, Sedlec (Czech Republic), Devínska Nová Ves (Slovakia) (Hoernes & Auinger 1879; Sieber 1958 b; Švagrovský 1981); Alpine- Carpathian Foredeep: Grund, Windpassing (Austria), Lysice, (Czech Republic) (Hoernes & Auinger 1879; Sieber 1947, 1949); Korytnica, Babice, Błonie, Hołubica, Podhorce, Zukowce, Biłka, Tarnopol (Friedberg 1911; Bałuk 1997); Oberpullendorf Basin: Ritzing (Austria) (Hoernes & Auinger 1879: Sieber 1956); Bükk Mountains: Borsodbóta (Hungary); Pannonian Basin: Várpalota, Balaton, Diósd, Hont, Letkés, Mátraverebély, Pécsszabolcs, Szob, Budapest: Illés street Szob, Hidas, (Hungary) (Csepreghy-Meznerics 1954; Strausz 1966; Kovács & Vicián 2013); Styrian Basin: Pöls (Austria) (Hoernes & Auinger 1879); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus (Romania); Caransebeş-Mehadia Basin: Valea Bela Reca (Romania); Buzău Basin: Crivineni, Valea Muscel (Romania) (Hoernes & Auinger 1879; Hinkulo 1968; Ionesi & Nicorici 1994); Banja Luka Basin: Jazovac (Bosnia and Herzegovina) (Atanacković 1985); Krka Basin: Dolenja Brezovica (Slovenia) (Mikuž 2009). Proto-Mediterranean Sea and north eastern Atlantic. This species is clearly also present in the Mediterranean area (e. g. Serravallian, Karman Basin, Turkey, Landau et al. 2013) but the confusion with C. exaltatus makes a critical evaluation of the literature data difficult.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFAEFFBCFF5FAB81FCF5409F.taxon	description	C. [onus] exiguus m. — Eichwald 1830: 222 [non Phasmoconus exiguus (Lamarck, 1810)]. Conus exiguus — Eichwald 1852: plate captions, pl. 9, figs 4 a – b. Con. exiguus m. — Eichwald 1853: 208. Syntypes: Two potential syntypes are stored in the Zoological Museum of St. Petersburg (Russia); inventory number ZISP 62085.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFAEFFBCFF5FAB81FCF5409F.taxon	materials_examined	Type locality. Eichwald (1830) mentioned Bilozirka (= Bialazurka, Bialozurka) and Shushkivtsi (= Shukowze) as type localities but on the label of the available specimens the locality is given as Zalisce. Therefore, it is unclear if these specimens are syntypes. In any case, all localities are located within a small area about 50 km NE of Ternopil (Ukraine). Age: Middle Miocene, Badenian (Langhian or Serravallian).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFAEFFBCFF5FAB81FCF5409F.taxon	etymology	Etymology. Referring to the Russian palaeontologist Karl Eduard von Eichwald (1795 – 1876), who was a pioneer in geosciences in the Carpathian Foredeep.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFAEFFBCFF5FAB81FCF5409F.taxon	materials_examined	Illustrated material. Figs 5 B 1 – B 2, 5 C 1 – C 2: 2 spec. ZISP 62085, Zalisce (Ukraine).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFAEFFBCFF5FAB81FCF5409F.taxon	description	Description. Small strombiform shells with high spire; teleoconch consisting of at least 9 whorls; spire gradate to scalariform with beaded carina placed mid-whorl or close to the lower suture; sutural ramp faintly striate. Shoulder broadly rounded, weakly subangulate; position of maximum diameter below shoulder; shoulder striate; subsutural flexure unknown. Last whorl stout, concave below maximum diameter, constricted; lower third of last whorl with sharp spiral cords; siphonal fasciole indistinct. Aperture largely missing in both specimens. Shell measurements and ratios. 2 specimens are available: SL: 15 mm, MD: 7 mm, spire angle: 60 °, last whorl angle: 33 °; SL: 11 mm, MD: 6 mm, spire angle: 70 °, last whorl angle: 40 °	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFAEFFBCFF5FAB81FCF5409F.taxon	discussion	Discussion. This species is unique among Paratethyan cones for its high spire and rounded shoulder, resulting in a strombiform profile. It is reminiscent of Plagioconus burdigalensis (Mayer, 1858), from the Burdigalian of the Aquitaine Basin, which also has a gradate to scalariform spire with carina and a last whorl with a broadly rounded, weakly subangulate shoulder and prominent spiral cords on the base. It differs from C. eichwaldi in its larger size and elongate last whorl. Therefore, it is unlikely that the Polish species is just a juvenile Plagioconus burdigalensis. The high spire, ventricose last whorl and striate sutural ramp would also fit in Lautoconus but the tuberculate and scalariform spire outline, are rather untypical for this genus (although Lautoconus desidiosus (Adams, 1853) may form a scalariform spire). Therefore, we tentatively place this species in Conilithes. Eichwald (1830) described this small and rare species as Conus exiguus. This name was already preoccupied by the Recent Conus exiguus Lamarck, 1810 from New Caledonia. Therefore, we propose Conilithes eichwaldi as replacement name. Paleoenvironment. Unknown; other specimens from the localities mentioned by Eichwald (1839, 1853) are typical nearshore taxa.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFAEFFBCFF5FAB81FCF5409F.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Carpathian Foredeep: Zalisce, Bilozirka (= Bialazurka, Bialozurka) and Shushkivtsi (= Shukowze) (Ukraine).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFADFFBAFF5FAA34FA1447B7.taxon	description	[Conus] exaltatus m. — Eichwald 1830: 222.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFADFFBAFF5FAA34FA1447B7.taxon	description	Con. exaltatus m. — Eichwald 1853: 208.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFADFFBAFF5FAA34FA1447B7.taxon	description	Conus (Leptoconus) dujardini Desh. — Boettger 1906: 2. Conus cf. avellana Lam. — Friedberg 1911: 56 (partim), pl. 2, fig. 20. Conus Dujardini Desh. — Friedberg 1911: 47, pl. 2, fig. 11. Conus Dujardini Desh. var. exaltatus Eichw. — Friedberg 1911: 51, pl. 2, fig. 12. Conus (Conuspira) [sic] dujardini Deshayes, 1831 — Moisescu 1955 a: 159, pl. 14, figs 15 – 18. Conus (Conuspira) antideluvianus Bruguière var. buiturica var. nov. — Moisescu 1955 a: 162, pl. 14, figs 7 – 8.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFADFFBAFF5FAA34FA1447B7.taxon	description	Conus (Conospira) dujardini brezinae Hoernes & Auinger — Csepreghy-Meznerics 1972: 34, pl. 17, figs 15 – 17.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFADFFBAFF5FAA34FA1447B7.taxon	materials_examined	Type material. Syntypes described by Eichwald (1830) from Bilozirka (= Bialazurka, Bialozurka) and Shushkivtsi (= Shukowze) about 50 km NE of Ternopil (Ukraine) (see map in Dubois de Montpéreux 1831). The specimens were most probably stored in the Zoological Museum of St. Petersburg (Russia) but could not be identified on request; middle Miocene, Badenian (Langhian). Studied material. 33 spec. NHMW 1846 / 0037 / 0037, Baden (Austria) including specimen illustrated in Hörnes (1851, pl. 5, fig. 3 d), 32 spec. NHMW 1846 / 0037 / 0036, Baden (Austria) including specimens illustrated in Hörnes (1851, pl. 5, figs 3 a, 3 b, 3 c), 12 spec. NHMW 1866 / 0011 / 0182, 12 spec. NHMW 2007 z 0078 / 007032, 24 spec. NHMW 2013 / 0078 / 052 - 2013 / 0078 / 0523, Baden (Austria); 22 spec. NHMW 1862 / 0001 / 0310,> 100 spec. NHMW 2010 / 0004 / 1343 - 2010 / 0004 / 1345, Bad Vöslau (Austria); 5 spec. NHMW 1973 / 1615 / 0214; 19 spec. NHMW A 1616, Baden-Sooß (Austria) including specimen illustrated in Schultz (1998, pl. 29, fig. 11); 1 spec. NHMW 2013 / 0479 / 1603, 1 spec. NHMW 1970 / 1396 / 1231, Gainfarn (Austria); 26 spec. NHMW 1867 / 0019 / 0013, Coşteiu de Sus (Romania); 59 spec. NHMW 1870 / 0033 / 0012, 16 NHMW A 1617, Lăpugiu de Sus (Romania). Illustrated material. Figs 6 B 1 – B 3: Möllersdorf (Austria): SL: 18.6 mm, MD: 7.2 mm, NHMW 2016 / 0002 / 0001; Figs 6 C 1 – C 3: Baden (Austria): SL: 48.5 mm, MD: 17.9 mm, NHMW 1866 / 0011 / 0182; Figs 6 D 1 – D 3: Baden (Austria): SL: 44.9 mm, MD: 17.6 mm, NHMW 1846 / 0037 / 0036, illustrated in Hörnes (1851, pl. 5, fig. 3), Figs 6 E 1 – E 3: Baden (Austria): SL: 48.6 mm, MD: 19.6 mm, NHMW 1846 / 0037 / 0036; Figs 6 F 1 – F 2: SL: 26.2 mm, MD: 11.3 mm, MNHN A 31836, Coşteiu de Sus (Romania) (pic: P. Loubry, MNHN); 3 D: Bad Vöslau (Austria), SL: 27.7 mm, MD: 11.6 mm, NHMW 1862 / 0001 / 0310. Revised description. Moderately small to medium-sized elongate shells with very high, scalariform spire and long last whorl. Protoconch high and conical, consisting of 3 whorls. Spire whorls faintly striate with sharp carina slightly below mid-whorl; sutural ramp flat to weakly concave. Tubercles on carina present only on first 3 – 4 spire whorls. Several deeply incised, punctate spiral grooves below carina on spire whorls and last whorl. Subsutural flexure deep, strongly curved, moderately asymmetrical. Last whorl strongly angular at shoulder with maximum diameter at carina; constricted at base with long, slightly widening and weakly recurved canal. Punctate spiral grooves on lower half of last whorl, extending over the adapical half in occasional specimens. Last whorl of subadult shells often completely covered by grooves. Colour pattern showing irregular blotches on the spire and shoulder and spirally arranged subquadratic dots on the cords on the last whorl, amalgamating to irregular blotches.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFADFFBAFF5FAA34FA1447B7.taxon	description	Shell measurements and ratios. n = 26 adult specimens: largest specimen: SL: 48.6 mm, MD: 19.6 mm, mean SL: 40.8 mm (σ = 4.9), mean MD: 16.4 mm (σ = 0.4), spire angle: µ = 59.7 ° (σ = 4.9 °), last whorl angle: µ = 30.9 ° (σ = 1.5 °), LW: µ = 2.5 (σ = 0.15), RD: µ = 0.56 (σ = 0.02), PMD: µ = 0.9 (σ = 0.03), RSH: µ = 0.29 (σ = 0.04).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFADFFBAFF5FAA34FA1447B7.taxon	discussion	Discussion. The status of Conus dujardini Deshayes, 1845 and Conus exaltatus Eichwald, 1830 is controversially discussed in the literature (e. g. Bałuk 1997; Kovács & Vicián 2013). Conus dujardini was introduced by Deshayes (1845) without clearly stating type material or type area because he referred to material from Ukraine, Austria and France. From the context, it is obvious, that he had French specimens at hand. In any case he referred to the illustration of Dubois de Montpéreux (1831, pl. 1, fig. 1), which was later designated the lectotype of Conus dujardini by Glibert (1952 b). The specimen of Dubois de Montpéreux originates from one of three villages in the area of Bilozirka (= Bialazurka, Bialozurka), 50 km ENE of Ternopil (= Tarnopol) in Ukraine (see map in Dubois de Montpéreux 1831). The specimens described by Eichwald (1830, 1853) as Conus exaltatus were collected at the same localities. As already discussed by Kovács & Vicián (2013), both specimens seem to be conspecific, which is also supported by the PCA; based on shell measurements, both specimens clearly plot with the specimens described herein as Conilithes exaltatus (Fig. 7 B). In a PCA plot based on shell-ratios (Fig. 7 A), both specimens plot close to C. brezinae, together with other stout morphologies of C. exaltatus (see remarks paragraph for C. brezinae for the separating features). Hence, it is highly likely that C. exaltatus and C. dujardini are conspecific, making C. dujardini a subjective junior synonym. The name C. dujardini has been extensively used in the literature, however it cannot be favoured over C. exaltatus as Friedberg (1911) used the name as a variety of C. dujardini, and Bałuk (1997) figured a specimen under this name. D’Orbigny (1852) introduced Conus subacutangulus as new name for the specimen, which was illustrated by Dubois de Montpéreux (1831, pl. 1, fig. 1) erroneously as Conus antidiluvianus. Conus subacutangulus is thus a subjective junior synonym of C. exaltatus Eichwald, 1830. Paleoenvironment. The majority of specimens in the NHMW collection derive from offshore clays suggesting inner to outer shelf environments as preferred habitat.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFADFFBAFF5FAA34FA1447B7.taxon	distribution	Distribution in Paratethys. Karpatian (early Miocene): Korneuburg Basin: Kleinebersdorf (Harzhauser 2002); Badenian (middle Miocene): Vienna Basin: Gainfarn, Enzesfeld, Steinebrunn, Bad Vöslau, Baden-Sooß, Möllersdorf, Pfaffstätten (Austria), Sedlec (Czech Republic); Alpine-Carpathian Foredeep: Grund, Windpassing (Austria) (Hoernes & Auinger 1879; Schultz 1998); Carpathian Foredeep: Węglinek, Łychów, Korytnica (Poland); Hołubica, Biłka, Jasinów, Bilozirka, Shushkivtsi (Ukraine) (Dubois de Montpéreux 1831; Bałuk 1997); Southern Carpathians: Dubova (Romania); Bükk Mountains: Borsodbóta (Hungary); Pannonian Basin: Bánd, Diósd, Herend, Hetvehely, Hidas, Kovácsvágás, Letkés, Márkháza, Mátraverebély, Nógrádszakál, Sámsonháza, Szob, Szokolya, Várpalota, Zebegény, Budapest: Rákos, Illés street (Hungary) (Strausz 1966; Csepreghy- Meznerics 1972; Kovács & Vicián 2013); Banja Luka Basin: Jazovac (Bosnia and Herzegovina) (Atanacković 1963); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus, Bujtur (Romania); Zârand Basin: Minişul de Sus (Romania); Caransebeş-Mehadia Basin: Valea Bela Reca (Romania) (Moisescu 1955 a; Stancu et al. 1971; Chira & Voia 2001; Caze et al. 2010); Dacian Basin: Opanec (Bulgaria) (Kojumdgieva & Strachimirov 1960). Proto-Mediterranean Sea and northeastern Atlantic. This species is clearly also present in the Miocene of the Mediterranean area but the confusion with C. brezinae makes a critical evaluation of the literature data difficult.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFABFFBBFF5FAF5CFE1F455D.taxon	materials_examined	Type material. Syntype, inventory number SMF XII. 2245 a coll. O. Boettger ex M. v. Kimakowicz 1883, Senckenberg Museum, Frankfurt (Germany); Lăpugiu de Sus (Romania); middle Miocene, Badenian (Langhian). Studied material. Syntype Illustrated material. Figs 5 I 1 – I 3: Syntype, SL: 13.5 mm, MD: 6.6 mm, Lăpugiu de Sus (Romania); picture taken by Sigrid Hof, provided by Ronald Janssen, section Malacology, Senckenberg Forschungsinstitut Frankfurt / Main. Revised description. Small biconical shell with moderately high scalariform spire. Early spire whorls probably tuberculate; later spire whorls smooth, carinate, weakly concave. Subsutural flexure deep, nearly symmetrical. Last whorl with angulated shoulder; position of maximum diameter slightly below shoulder; faintly ventricose, weakly constricted at base. Siphonal canal short, rather narrow. Few deeply incised spiral grooves on base. Colour pattern consisting of axially arranged zig-zag stripes (according to Boettger 1887; the illustrated syntype shows only vague traces of this pattern).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFABFFBBFF5FAF5CFE1F455D.taxon	description	Shell measurements and ratios. Syntype: SL: 13.5 mm, MD: 6.6 mm, spire angle: µ = 68 °, last whorl angle: 44 °, LW: 2.05, RD: 0.70, PMD: 0.89, RSH: 0.30.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFABFFBBFF5FAF5CFE1F455D.taxon	discussion	Discussion. Boettger (1906) emphasised that Conilithes brezinae (Hoernes & Auinger, 1879) did not occur at Coşteiu de Sus and united all “ brezinae - like ” specimens from that locality in his Conus sceptophorus. In the collections of the Natural History Museum Vienna, however, numerous specimens from Coşteiu de Sus undoubtedly represent C. brezinae. In contrast, not a single of these specimens agrees with the syntypes of Conilithes sceptophorus in the stout biconical outline and the zig-zag colour pattern. Whilst one might argue that Boettger’s syntypes are just stout specimens of C. brezinae, the zig-zag colour pattern supports a separation because C. brezinae develops thin spiral lines. For the same reason we reject a synonymization with the Pliocene Italian Conilithes canaliculatus (Brocchi, 1814), as proposed by Kovács & Balász (2016), because this species has densely spaced spirals (see Panganelli 2014). Paleoenvironment. Unknown.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFABFFBBFF5FAF5CFE1F455D.taxon	distribution	Distribution in Paratethys. Transylvanian Basin: Lăpugiu de Sus (Romania), Coşteiu de Sus (Romania) (Boettger 1887, 1906).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFAAFFBBFF5FACFDFA0046B3.taxon	materials_examined	Type species: Conus pulcher [Lightfoot, 1786], by original designation. Recent, West Africa. Note. According to Tucker & Tenorio (2009), the genus Kalloconus da Motta, 1991 is characterised by large to very large and obconic shells, with a broad, rounded shoulder. The spire whorls can be smooth, striate with the sculpture disappearing on later whorls, or persisting as crowded, weak spirals. The protoconch is multispiral. The subsutural flexure is moderately deep to deep in larger specimens, and shallower in smaller specimens. The shell is ornamented with spots and dashes in spiral rows. In their molecular phylogeny, Puillandre et al. (2014 a) recognised this group as being monophyletic, albeit at subgeneric level, and a sister group to Lautoconus. Both of these genera today have a West African and European distribution. However, whereas Tucker & Tenorio (2009) included only two extant species within the genus, C. pulcher [Lightfoot, 1786] and C. byssinus (Röding, 1798), the molecular phylogeny by Puillandre et al. (2014 a) included six further species: C. ateralbus Kiener, 1850, C. genuanus Linnaeus, 1758, C. trochulus Reeve, 1844, C. venulatus Hwass in Bruguière, 1792, C. atlanticoselvagem Afonso & Tenorio, 2004 and Conus pseudonivifer Monteiro, Tenorio & Poppe, 2004. Although the molecular phylogenetics have led to a wider generic concept, the generic description of the shell remains unchanged. Based on Paratethyan material here reviewed, we can add that Kalloconus species are small to very large, squat to moderately elongate. The spire is always low to very low; spire whorls are convex and usually smooth except for occasional striae on early spire whorls. The proto-Mediterranean Kalloconus betulinoides (Lamarck, 1810) is a fossil species with striate spire whorls like the extant type species Kalloconus pulcher ([Lightfoot], 1786). Spiral sculpture on last whorl very reduced. The depth of the subsutural flexures in all our fossil species is highly variable, ranging from very shallow to deep; they are usually moderately curved and moderately asymmetrical. The last whorl is of medium to wide width. As with the living species, the colour pattern in most of the fossil representatives is also composed of spiral rows of spots and dashes; only few species develop continuous spirals.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA9FFB6FF5FA8E1FAC8474D.taxon	description	Conus Berghausi mihi, Michelotti 1847: 342, pl. 13, figs 9 – 9 ’. Conus Berghausi Micht. — Hörnes 1851: 19, pl. 1, figs 3 a – c [non fig 3 d = Kalloconus hendricksi nov. sp.]. [Dendroconus] [Conus] Vaceki n. f. — Hoernes 1878 a: 195 (nomen nudum). C. [onus] Vaceki — Hoernes 1878 b: 206 (nomen nudum). Conus (Dendroconus) Vaceki nov. form. — Hoernes & Auinger 1879: 22 [nov. nom. pro C. berghausi in Hörnes 1851, pl. 1, fig. 3].	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA9FFB6FF5FA8E1FAC8474D.taxon	description	Conus Berghausi Micht. — Friedberg 1928: 566, text-fig. 80.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA9FFB6FF5FA8E1FAC8474D.taxon	materials_examined	Type material. The specimen illustrated by Michelotti (1847) is lost. Therefore, Hall (1966) proposed the specimen illustrated by Sacco (1893 a, pl. 1, fig. 16) from the late Miocene (Tortonian) of Stazzano in Italy as neotype (stored in the Museo Regionale di Scienze Naturali, Torino, catalogue number BS. 039.40.006). The type locality of Conus vaceki Hoernes & Auinger, 1879 (considered to be junior synonym of C. berghausi) is Niederkreuzstetten (Austria); the type stratum is lower Miocene coastal sand of the Korneuburg Formation. Studied material. 9 spec. NHMW 1849 / 0004 / 0017, Niederkreuzstetten (Austria), including specimen illustrated in Hörnes (1851, pl. 1, figs 3 a – c) and Harzhauser (2002, pl. 10, fig. 5); 5 spec. NHMW 1864 / 0001 / 0498, Niederkreuzstetten (Austria); 1 spec. private collection, Anton Breitenberger, Weitendorf (Austria); 3 spec. NHMW 1856 / 0050 / 0413, Lăpugiu de Sus (Romania). Illustrated material. Figs 8 A 1 – A 3: Niederkreuzstetten (Austria), SL: 39.1, MD: 28.9 mm, NHMW 1849 / 0004 / 0017, specimen illustrated in Hörnes (1851, pl. 1, figs 3 a – c), syntype of Conus vaceki Hoernes & Auinger, 1879; Figs 8 B 1 – B 3: Niederkreuzstetten (Austria), SL: 32.2, MD: 21.4 mm, NHMW 1864 / 0001 / 0498; Fig. 8 C: Niederkreuzstetten (Austria), SL: 31.4, MD: 21.7 mm, NHMW 1849 / 0004 / 0017; Figs 8 D 1 – D 3: Weitendorf (Austria), SL: 34.5 mm, MD: 23.4 mm, private collection, Anton Breitenberger (this specimen shows its natural coloration due to heating by basalt flows); Figs. 8 E, 3 G 1 – G 2: Lăpugiu de Sus (Romania): SL: 42.0 mm, MD: 29.8 mm, NHMW 1856 / 0050 / 0413. Revised description. Medium-sized, stout club-shaped shells; spire low and conical, weakly coeloconoid. Early spire whorls slightly concave with strong spiral cords. Suture impressed, emphasized by two raised cords, resulting in a narrowly channelled suture in some specimens. Spiral cords become obsolete within 3 rd – 4 th spire whorls, which develop broad, irregular swellings along shoulder causing undulations of the shoulder. Subsequent spire whorls smooth, becoming successively more convex. Last whorl distinctly broadening with prominent, rounded shoulder coinciding with maximum diameter. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Last whorl short, conical, not constricted. Aperture moderately wide; slightly widening towards short and straight canal; siphonal fasciole indistinct, short; inner lip short, slightly twisted. Deep spiral grooves demarcate broad spiral cords on base; grooves stop abruptly within lower third of whorl and do not become obsolete adapically. Colour pattern highly variable, consisting of about 13 – 20 rows of spirally arranged, often subquadratic dots. Size of dots highly variable; small dots tend to be densely spaced, whilst large dots are widely spaced; often two rows of dots form close-spaced pairs separated by wider interspaces from the next pair.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA9FFB6FF5FA8E1FAC8474D.taxon	description	Shell measurements and ratios. n = 14: largest specimen: SL: 44 mm, MD: 31.1 mm, mean SL: 35.0 mm (σ = 5.4), mean MD: 24.3 mm (σ = 4.2), spire angle: µ = 128.0 ° (σ = 10.2 °), last whorl angle: µ = 40.1 ° (σ = 1.1 °), LW: µ = 1.4 (σ = 0.06), RD: µ = 0.75 (σ = 0.04), PMD: µ = 0.87 (σ = 0.03), RSH: µ = 0.08 (σ = 0.04).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA9FFB6FF5FA8E1FAC8474D.taxon	discussion	Discussion. Despite the numerous papers dealing with this species, no author so far has described the conspicuous sculpture of the early spire whorls with broad shoulder nodules and prominent striae. Similarly, the concave tops of early spire whorls have not been mentioned before. A comparison with specimens from the Tortonian of Modena and Tortona, showed the same features and fully support the identification of the Paratethyan specimens. This species was placed in Kalloconus da Motta, 1991 by Tucker & Tenorio (2009) and Landau et al. (2013). The concave tops do not contradict this assignment as the type species Kalloconus pulcher ([Lightfoot, 1786]) and also K. byssinus (Röding, 1798) — the second extant species included by Tucker & Tenorio (2009) in the genus — have also slightly concave tops. Clear deviation from the diagnosis is the presence of very prominent spiral cords on early spire whorls and the broad nodules. Consequently, either the diagnosis of Kalloconus has to be emended or the Miocene species has to be placed in another genus. Provisionally, we follow Tucker & Tenorio (2009) and tentatively treat the species as Kalloconus. The colouration of the specimens from Weitendorf (Styria, Austria) seems to reflect the original pattern and several other mollusc species from this site reveal similarly well preserved patterns (NHMW collection). Although we do not know the chemical mechanism, it is likely that heating by a synsedimentary basalt flow (Krainer 1987) was responsible for this preservation.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA9FFB6FF5FA8E1FAC8474D.taxon	distribution	Distribution in Paratethys. Karpatian (early Miocene): Alpine-Carpathian Foredeep: Niederkreuzstetten (Austria) (Harzhauser 2002); Badenian (middle Miocene): Vienna Basin: Steinebrunn (Austria), Mikulov-Kienberk (Czech Republic) (Hörnes 1851; Sieber 1958 b); Styrian Basin: Pöls, Weitendorf (Austria) (own data); Carpathian Foredeep: Korytnica, Białogon (Poland) (Friedberg 1911; Bałuk 1997); Pannonian Basin System: Pécsszabolcs, Letkés, Mátraverebély, Sámsonháza, Várpalota, Bánd, Diósd, Hidas, Márkháza, Zebegény, Budapest: Rákos, Illés street (Hungary) (Kovács & Vicián 2013); southern Pannonian Basin: Milijevići (Bosnia and Herzegovina) (Atanacković 1969); Transylvanian Basin: Lăpugiu de Sus (Romania) (Hoernes & Auinger 1879); Şimleu Basin: Tusa (Romania) (Nicorici 1972); Caransebeş-Mehadia Basin: Valea Strinii, Valea Bela Reca, Valea Calvei, Valea Satului (Romania); Krka Basin: Orehovica (Slovenia) (Miku 2009). Proto-Mediterranean Sea and north eastern Atlantic. Burdigalian: Colli Torinesi, Italy (Sacco 1893 a; Hall 1966); Langhian and Serravallian: Aquitaine Basin, France (Peyrot 1931); Langhian: Loire Basin (Glibert 1952 a); Serravallian: Karaman Basin, Turkey (Landau et al. 2013); Tortonian: Cacela Basin, Portugal (Pereira da Costa 1866); Po Basin, Italy (Sacco 1893 a; Davoli 1972, 1990; Ruggieri & Davoli 1984; Caprotti 2011). An occurrence from the Pliocene of Italy mentioned by Sacco (1893 a) needs confirmation and is most probably incorrect.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA7FFB4FF5FAE0CFDB04356.taxon	materials_examined	Holotype: Figs 8 F 1 – F 5: Lăpugiu de Sus (Romania), SL: 25.2, MD: 16.9 mm, NHMW 1870 / 0033 / 0005 a. Paratype: Figs 8 G 1 – G 3: Pöls (Austria), SL: 21.2 mm, MD: 13.0 mm NHMW 1861 / 0001 / 0226.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA7FFB4FF5FAE0CFDB04356.taxon	description	Additional paratypes: Fig. 8 H: Weitendorf (Austria), SL: 24.6 mm, MD: 16.4 mm, private collection, Anton Breitenberger (this specimen shows its natural coloration due to heating by basalt flows); Fig. 3 H: Pöls (Austria), SL: 21.1 mm, MD: 13.4 mm NHMW 1861 / 0001 / 0226; Fig. 3 I: Lăpugiu de Sus (Romania), SL: 21.4 mm, MD: 13.6 mm, NHMW 1870 / 0033 / 0005 c; 1 spec. NHMW 1849 / 0023 / 0003, Baden (Austria), illustrated in Hörnes (1851, pl. 1, fig. 3 d); 31 spec. NHMW 1870 / 0033 / 0005, Lăpugiu de Sus (Romania), including specimen illustrated in Hoernes & Auinger (1879, pl. 1, fig. 21); 24 spec. NHMW 1861 / 0001 / 0226, 12 spec. NHMW 2002 / 0181, Weitendorf (Austria).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA7FFB4FF5FAE0CFDB04356.taxon	materials_examined	Type stratum: Badenian marly-clayey deposits with thin interlayers of sand and corallinacean limestones. Type locality: Lăpugiu de Sus (Romania). Age: Middle Miocene, early Badenian (= Langhian).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA7FFB4FF5FAE0CFDB04356.taxon	etymology	Etymology. In honour of Jonathan R. Hendricks, a specialist for Neogene cone snails, in recognition of his contributions to the knowledge of the group.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA7FFB4FF5FAE0CFDB04356.taxon	description	Description. Small shells; spire very low conical with pointed, coeloconoid initial part. Early spire whorls weakly concave, distinctly striate with impressed, sometimes narrowly canaliculated suture. Within 3 rd - 5 th teleoconch whorl, shoulder of spire whorls marked by irregular swellings and blunt beads, resulting in undulating suture; last spire whorl weakly convex, smooth, distinctly broadening. Position of maximum diameter coinciding with shoulder or slightly below. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Last whorl conical to faintly ventricose, weakly constricted. Aperture narrow; slightly widening towards short and weakly reflected siphonal canal; siphonal fasciole narrow, rather indistinct; inner lip short, slightly twisted. Deep spiral grooves demarcate broad spiral cords on base. Shell surface glossy. Colour pattern consisting of about 13 – 16 regularly spaced rows of spirally arranged, subquadratic dots. Rows usually consisting of dots of more or less equal size; rarely single rows are formed by smaller dots. Shell measurements and ratios. n = 18: largest specimen: SL: 26.8 mm, MD: 17.2 mm, mean SL: 22.6 mm (σ = 1.7), mean MD: 14.4 mm (σ = 1.1), spire angle: µ = 128.3 ° (σ = 10.6 °), last whorl angle: µ = 39.9 ° (σ = 1.4 °), LW: µ = 1.6 (σ = 0.07), RD: µ = 0.70 (σ = 0.02), PMD: µ = 0.90 (σ = 0.02), RSH: µ = 0.09 (σ = 0.04).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA7FFB4FF5FAE0CFDB04356.taxon	discussion	Discussion. Kalloconus berghausi is an eye-catching species due to its frequently preserved colour pattern of spirally arranged dots. Typical specimens are medium-sized, squat and club-shaped with prominent shoulder (e. g. Landau et al. 2013). Along with this morphology, a distinctly smaller, less club-shaped morphotype is usually identified as Conus berghausi in collections and the literature, mainly due to its very similar colour pattern. A Principal Component Analysis of the shell measurements and ratios of both morphotypes shows a very good separation of both groups (Fig. 9). Both types co-occur also in the late Miocene of Montegibbio (Davoli 1972) and are represented in NHMW collections from Modena and Tortona (Italy). The conspicuous spire sculpture strongly suggests a close relationship between both types. Aside from size and outline, typical berghausi differs from the smaller species in the less pointed early spire, the much stronger spiral cords on the spire whorls, the broader, less defined shoulder nodules, the shallower suture and the wider suture. In addition, both groups differ also in the colour pattern, which is less variable in the K. hendricksi, consisting of regularly spaced rows of regularly sized dots. A further difference is the row of dots on the last spire whorl of K. hendricksi whereas K. berghausi develops irregular, thin flammulae. The lack of intermediate specimens and the very constant size in the collection lots does not support an interpretation of the K. hendricksi as juveniles of the larger K. berghausi. Therefore, the smaller species seems to represent a distinct but overlooked Miocene species. Nevertheless, the fact that both morphotypes frequently cooccur at some Paratethyan and proto-Mediterranean sections is striking and we cannot exclude that we are dealing with ontogenetic stages or sexual dimorphism as discussed for Conus centurio Born, 1778 for which shells of females are reported to be larger and more obtuse (Percharde 1984). Despite the numerous names introduced by Sacco (1893 a), we were not able to find any available name for this small berghausi - like species. Conus broteri Pereira da Costa, 1866, from the Tortonian of Portugal, may partly correspond to this species (e. g. Pereira da Costa 1866, pl. 49, fig. 26), but other syntypes of C. broteri differ clearly in their very squat shape with flat spire (e. g. Pereira da Costa 1866, pl. 49, fig. 30; Gonçalves & Monteiro 2012, p. 33, unnumbered fig.). Paleoenvironment. Shallow marine near shore settings.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA7FFB4FF5FAE0CFDB04356.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Baden (Austria) (Hörnes 1851); Styrian Basin: Pöls, Weitendorf (Austria) (own data); Carpathian Foredeep: Korytnica (Poland) (Bałuk 1997); Pannonian Basin System: Várpalota, Mecsek Mts. (Hungary) (Strausz 1966; Bohn-Havas 1973); Transylvanian Basin: Lăpugiu de Sus (Romania) (Hoernes & Auinger 1879); Dacian Basin: Târnene, Staropatica, Radomirci (Bulgaria) (Kojumdgieva & Strachimirov 1960). Proto-Mediterranean Sea and north eastern Atlantic.? Tortonian: Cacela Basin, Portugal (Pereira da Costa 1866); Po Basin, Italy (Davoli 1972).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA5FFB5FF5FADEFFF564319.taxon	description	Conus Cacellensis Costa — Pereira da Costa 1866: 15, pl. 3, figs 4 – 6.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA5FFB5FF5FADEFFF564319.taxon	materials_examined	Type material: Syntypes illustrated in Pereira da Costa (1866: pl. 3, figs 4 – 6); late Miocene, Tortonian; Cacella, Portugal. Studied material: 1 spec. NHMW 1855 / 0043 / 0002 Lăpugiu de Sus (Romania). Illustrated material. Figs 8 I 1 – I 4, 3 J: Lăpugiu de Sus (Romania): SL: 70.3 mm, MD: 43.6 mm, NHMW 1855 / 0043 / 0002. Revised description. Moderately large shell with low spire; spire whorls weakly convex, smooth except for faint striae in the adapical thirds; suture impressed. Last whorl with additional faint spiral cords just above shoulder of last whorl. Subsutural flexure very shallow, weakly curved, moderately asymmetrical. Broadly rounded shoulder; last whorl elongate conical with weak constriction on base. No spiral cords on base. Siphonal fasciole weakly swollen, short; siphonal canal moderately wide, short. Colour pattern consisting of numerous delicate, continuous spiral stripes under UV light and about 10 narrow, whitish spiral bands in normal light (not clearly visible under UV light).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA5FFB5FF5FADEFFF564319.taxon	description	Shell measurements and ratios. SL: 70.3 mm, MD: 43.6 mm, spire angle: 130 °, last whorl angle: 30 °, LW: 1.61, RD: 0.69, PMD: 0.93, RSH: 0.10.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA5FFB5FF5FADEFFF564319.taxon	discussion	Discussion. Only a single specimen is available, which agrees well with the specimens illustrated by Pereira da Costa (1866) and Gonçalves & Monteiro (2012) in shape, the shallow subsutural flexure, the colour pattern and the faint concavity of the last whorl. This species is reminiscent of Kalloconus betulinoides (Lamarck, 1810), from the late Miocene and Pliocene of the Proto-Mediterranean Sea, which differs in its even more elongate shell, the longer siphonal canal, the channelled suture, the striate early spire whorls and the prominent spiral cords on the base (based on NHMW collection). Paleoenvironment. Unknown.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA5FFB5FF5FADEFFF564319.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Transylvanian Basin: Lăpugiu de Sus. Proto-Mediterranean Sea and northeastern Atlantic. Tortonian: Cacela Basin, Portugal (Pereira da Costa 1866).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA4FFB5FF5FAACCFD9E469C.taxon	description	Conus gallicus May. - Eym. — Mayer-Eymar 1890: 295.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA4FFB5FF5FAACCFD9E469C.taxon	materials_examined	Type material. Holotype, ETH Zurich (Swiss Federal Institute of Technology in Zurich, German: Eidgenössische Technische Hochschule Zürich), inventory number S 2776 (fide Hall 1966); Larriey-Saucats, France; early Miocene, Aquitanian. Studied material. 1 spec. NHMW 2016 / 0036 / 0001, Nemeşeşti (Romania). Illustrated material. Figs 10 A 1 – A 6, 3 K: Nemeşeşti (Romania), SL: 50.1 mm, MD: 31.9 mm, NHMW 2016 / 0036 / 0001. Revised description. Medium-sized shell; low conical spire with coeloconoid initial part; early spire whorls faintly striate, slightly concave; later whorls weakly convex; suture impressed. Subsutural flexure of medium depth, moderately curved, moderately asymmetrical. Narrow, rounded shoulder, coinciding with position of maximum diameter. Last whorl conical, straight sided with faint constriction at base. Aperture narrow with parallel margins; Siphonal canal short, nearly straight; siphonal fasciole very weak. Few wide-spaced spiral cords on base with weaker cords intercalating. Colour pattern on spire and last whorl consisting of irregular, strongly amalgamating dark blotches and few a light blotches.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA4FFB5FF5FAACCFD9E469C.taxon	description	Shell measurements and ratios. SL: 50.1 mm, MD: 31.9 mm, spire angle: 123 °, last whorl angle: 37 °, LW: 1.57, RD: 0.73, PMD: 0.90, RSH: 0.12.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA4FFB5FF5FAACCFD9E469C.taxon	discussion	Discussion. This species is characterised by the sloping last spire whorl, the narrow rounded shoulder and the narrow aperture. Nevertheless, the lack of any “ extravagant ” conchological features might raise doubts about the identification of the middle Miocene Paratethyan shell with an early to middle Miocene species from the northeastern Atlantic. Fortunately, the identification is supported by the unusual colour pattern, which was documented by Caze et al. (2011) also for an early Miocene specimen from the Aquitaine Basin.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA4FFB5FF5FAACCFD9E469C.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Transylvanian Basin: Nemeşeşti (Romania) (own data). Proto-Mediterranean Sea and northeastern Atlantic. early Miocene (Aquitanian, Burdigalian): Aquitaine Basin: Larriey, Saucats, Merignac (Mayer-Eymar 1890); Turin Hills: Termofourà (Italy) (Hall 1966);? Pliocene: Po Basin: Asti (Italy) (Hall 1966).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA3FFB2FF5FA8E1FA0846A6.taxon	description	[Lithoconus] [Conus] hungaricus n. f. — Hoernes 1878 a: 195 (nomen nudum). Conus (Lithoconus) Hungaricus nov. form. — Hoernes & Auinger 1879: 29 (partim), pl. 2, figs 6 a – b, [non pl. 4, figs 1 a – c].	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA3FFB2FF5FA8E1FA0846A6.taxon	materials_examined	Type material. 2 syntypes NHMW 1851 / 0013 / 0018, Bad Vöslau (Austria), including specimen illustrated in Hoernes & Auinger (1879, pl. 2, fig. 6), 1 syntype NHMW 1867 / 0029 / 0004, Coşteiu de Sus (Romania), illustrated herein as Figs 10 B 1 – B 3, designated herein as lectotype. Studied material. Syntypes and 6 spec. NHMW A 990; Grund (Austria), 4 spec. NHMW 1870 / 0037 / 0002, Ritzing (Austria). Illustrated material. Figs 10 B 1 – B 3, 3 L: Coşteiu de Sus (Romania), 55.9 mm, MD: 40.4 mm, NHMW 1867 / 0029 / 0004; Figs 10 C 1 – C 3: Bad Vöslau (Austria), SL: 66.2 mm, MD: 46.1 mm, NHMW 1851 / 0013 / 0018; Figs 10 D 1 – D 2: Bad Vöslau, SL: 65.8 mm, MD: 43.9 mm, NHMW 2010 / 0004 / 1578. Revised description. Moderately large, stout, broadly club-shaped, robust shells. Spire depressed and conical, initial part low coeloconoid; low dome-shaped in large, fully grown specimens. Spire whorls flat to weakly convex, smooth; subsutural flexure of medium depth, moderately curved, moderately asymmetrical. Rounded shoulder, coinciding with position of maximum diameter; last whorl broad, ventricose, not constricted. Aperture wide, distinctly broadening abapically, terminating in a wide, short and straight canal. Inner lip short, broad, strongly twisted, demarcated from weakly swollen fasciole by a shallow groove, causing an angulation at the transition from inner lip to columella. Spiral cords on base. Colour pattern poorly preserved, consisting of numerous densely spaced spiral rows of narrow dashes.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA3FFB2FF5FA8E1FA0846A6.taxon	description	Shell measurements and ratios. n = 7: largest specimen: SL: 69.3 mm, MD: 55.5 mm, mean SL: 62.0 mm (σ = 5.4), mean MD: 44.2 mm (σ = 6.0), spire angle: µ = 141.6 ° (σ = 6.5 °), last whorl angle: µ = 43.3 ° (σ = 2.0 °), LW: µ = 1.41 (σ = 0.08), RD: µ = 0.76 (σ = 0.04), PMD: µ = 0.89 (σ = 0.02), RSH: µ = 0.07 (σ = 0.01). Paleoenvironment. The sediment infill in the apertures of the specimens from Bad Vöslau and Grund suggests a shallow marine silty-sandy environment.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA3FFB2FF5FA8E1FA0846A6.taxon	discussion	Discussion. Conus hungaricus is a problematic taxon. Hoernes & Auinger (1879) based their description on specimens, which — in our opinion — represent three different species. Unfortunately, they considered the specimen from Coşteiu de Sus, illustrated by them as pl. 4, fig. 1, as characteristic specimen. This specimen, however, has concave and striate spire whorls and a distinctly angulate shoulder (like a second specimen in the type lot). We consider these specimens to represent Monteiroconus daciae (Hoernes & Auinger, 1879). The second specimen illustrated by Hoernes & Auinger (1879, pl. 2, fig. 6) and other syntypes from the Vienna Basin, differ considerably from those specimens in their rounded shoulder and weakly convex, smooth spire whorls. All subsequent authors identified this morphotype as Conus hungaricus. Hoernes & Auinger (1879) did not designate a type specimen in accordance to our modern understanding and therefore we select the syntype from Coşteiu de Sus (Romania), illustrated herein as Figs 10 B 1 – B 3 as lectotype of Conus hungaricus. Due the ambiguous situation, Bałuk (1997) synonymized Conus daciae and Conus hungaricus. Although subadult M. daciae are morphologically reminiscent of Kalloconus hungaricus, it differs from M. daciae in the ventricose last whorl and the shallower subsutural flexure. Further specimens from Lăpugiu de Sus (Romania) treated as Conus hungaricus by Hoernes & Auinger (1879), are described herein as Kalloconus pseudohungaricus. The juvenile specimen from the Badenian of the Bükk Mountains in Hungary illustrated by Csepreghy- Meznerics (1972) has deep spiral grooves on the base, which would be atypical for K. hungaricus.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA3FFB2FF5FA8E1FA0846A6.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Alpine-Carpathian Foredeep: Grund (Austria) (Sieber 1949); Eisenstadt-Sopron Basin: Forchtenau (Austria) (Sieber 1956);? Bükk Mountains: Borsodbóta (Hungary) (Csepreghy-Meznerics 1972); Pannonian Basin System: Várpalota (Hungary) (Strausz 1966); Transylvanian Basin: Coşteiu de Sus (Romania) (Hoernes & Auinger 1879; NHMW collection). Proto-Mediterranean Sea. Serravallian (middle Miocene): Karman Basin (Turkey) (Landau et al. 2013). The occurrence of this species in the Tortonian of Barcelona, mentioned by Faura I Sans (1908), needs confirmation.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA1FFB0FF5FA8E1FB7F4788.taxon	materials_examined	Holotype: Figs 10 E 1 – E 3: SL: 29.5 mm, MD: 20.6 mm, NHMW 2016 / 0006 / 0001. Paratype: Figs 10 F 1 – F 4: SL: 30.1 mm, MD: 21.0 mm, NHMW 2016 / 0006 / 0002. Additional material: 1 spec. NHMW 2016 / 0006 / 0004, 3 spec. private collection Anton Breitenberger: Figs 10 G 1 – G 4: SL: 40.9 mm, MD: 23.3 mm; Figs 10 H 1 – H 3: SL: 35.5 mm, MD: 23.2 mm, Fig. 3 M: SL: 37.4 mm, MD: 26.6 mm; all specimens from Letkés, Hungary. Type stratum: fossil-rich marly sand with coral blocks and andesite boulders of the Sámsonháza Formation (Császár 1997). Type locality: Letkés at the western part of the Börzsöny Mts. (Hungary); see Kovács & Vicián (2013) for a map and description. Age: Middle Miocene, early Badenian (= Langhian).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA1FFB0FF5FA8E1FB7F4788.taxon	etymology	Etymology: Referring to the type locality Letkés in Hungary.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA1FFB0FF5FA8E1FB7F4788.taxon	description	Description. Moderately small to medium sized, stout shells with very low to low spire and broad conical last whorl. Early spire weakly coeloconoid with faintly concave and striate whorls; broad, weak nodules along the shoulder of 3 rd – 5 th spire whorl resulting in undulating suture. Later spire whorls narrowly coiled, weakly convex, weakly striate; suture narrow; last spire whorl broad and flat with shallow central broad groove, striate, passing via rounded shoulder into broad conical to faintly ventricose last whorl. Position of maximum diameter in posterior quarter of last whorl. Subsutural flexure shallow, moderately curved, strongly asymmetrical. Siphonal fasciole very weak but well demarcated from short, narrow inner lip. Siphonal canal short, straight. Aperture straight, moderately narrow, slightly broadening towards siphonal canal. Delicate spiral grooves on lower third of last whorl. Very prominent colour pattern in UV light consisting of very densely spaced spirals of thin, delicate dashes covering the entire last whorl. Below the maximum diameter, slightly below mid-whorl and on the base, the dashes area arranged in three darker bands. Broad flammulae appear on spire whorls. Shell measurements and ratios. n = 6: largest specimen: SL: 40.9 mm, MD: 26.6 mm, mean SL: 33.8 mm (σ = 4.8), mean MD: 22.3 mm (σ = 2.6), spire angle: µ = 138.5 ° (σ = 10.1 °), last whorl angle: µ = 41.5 ° (σ = 1.9 °), LW: µ = 1.52 (σ = 0.13), RD: µ = 0.73 (σ = 0.03), PMD: µ = 0.87 (σ = 0.11), RSH: µ = 0.10 (σ = 0.06).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA1FFB0FF5FA8E1FB7F4788.taxon	discussion	Discussion. This species could be easily mistaken for Kalloconus berghausi (Michelotti, 1847) or Kalloconus hendricksi nov. sp. from which it differs in the constantly striate spire whorls, the faintly ventricose last whorl and especially in the conspicuous spiral colour pattern, which differs completely from the large dots and blotches of K. berghausi and K. hendricksi. Therefore, some specimens in the Paratethyan literature might rather represent Kalloconus letkesensis than K. berghausi (e. g. Kojumdgieva in Kojumdgieva & Strachimirov 1960, pl. 51, fig. 3). Another morphologically similar species is Kalloconus neumayri (Hoernes & Auinger, 1879), which is distinguished easily by its colour pattern consisting of large rectangular blotches. This species was mixed by Kovács & Vicián (2013) with Kalloconus tietzei (Hoernes & Auinger, 1879). It differs from K. tietzei in its smaller size, more slender outline, striate spire whorls, shallower subsutural flexure, broader last spire whorl, much weaker siphonal fasciole and the dense spirals of dashes, whereas K. tietzei shows much coarser and wider-spaced dashes. Paleoenvironment. The locality Letkés comprises a mixed assemblage of partly reworked taxa (Kovács & Vicián 2013); the frequent occurrence of corals suggests a shallow marine depositional environment.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA1FFB0FF5FA8E1FB7F4788.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Pannonian Basin: Letkés (Hungary).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA1FFB1FF5FAF43FF5647A4.taxon	description	Conus fusco-cingulatus Bronn — Hörnes 1851: 21 (partim), pl. 1, figs 4 a – c [non Phasmoconus fuscocingulatus (Hörnes, 1851)]. [Lithoconus] [Conus] moravicus n. f. — Hoernes 1878 a: 195 (nomen nudum). Conus (Lithoconus) Moravicus nov. form. — Hoernes & Auinger 1879: 29 (partim) [nov. nom. pro Conus fusco-cingulatus in Hörnes 1851, pl. 1, figs 4 a – c].	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA1FFB1FF5FAF43FF5647A4.taxon	materials_examined	Type material. 7 syntypes NHMW 1851 / 0010 / 0001, including specimen illustrated in Hörnes (1851, pl. 1, fig. 4); 3 syntypes NHMW 1851 / 0020 / 0001, all Mikulov-Kienberk (Czech Republic); middle Miocene, Badenian (late Langhian). Studied material. Syntypes and 12 specimens, NHMW 2016 / 0033 / 0001, Mikulov-Kienberk (Czech Republic); 15 spec. NHMW 1846 / 0037 / 0057, Gainfarn (Austria); 4 spec. NHMW 1848 / 0003 / 0004, Ritzing (Austria); 1 spec. NHMW 1865 / 0035 / 0025, Szob (Hungary). Illustrated material. Figs 11 A 1 – A 5: Mikulov-Kienberk (Czech Republic): SL: 40.2 mm, MD: 26.2 mm, NHMW 2016 / 0033 / 0001; Figs 11 B 1 – B 3: syntype, Mikulov-Kienberk (Czech Republic): SL: 47.5 mm, MD: 32.0 mm, NHMW 1851 / 0010 / 0001, illustrated in Hörnes (1851, pl. 1, fig. 4); Fig. 3 N: Mikulov-Kienberk (Czech Republic): SL: 35.2 mm, MD: 23.1 mm, NHMW 2016 / 0033 / 0001. Revised description. Medium sized solid shells with very low conical spire and short, broad conical last whorl. Spire whorls nearly flat to weakly concave in adapical half and slightly convex abapically; early spire whorls distinctly striate, with two striae demarcating impressed suture. Very broad, poorly defined swellings along shoulder of 3 rd - 6 th teleoconch whorl, causing undulating suture. Last two whorls smooth with very irregular suture. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Last whorl broad with rounded, prominent shoulder. A weak concavity appears in many specimens slightly below shoulder, resulting in stout, ficoid outline. Last whorl straight-sided or weakly ventricose below concavity; indistinctly constricted at base. Siphonal canal short and wide; fasciole short, moderately swollen, well demarcated from short inner lip; aperture moderately wide with subparallel margins. Densely spaced wavy spiral grooves and cords on base. Colour pattern frequently preserved, consisting of 12 – 16 thin, continuous spirals separated by broader interspaces.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA1FFB1FF5FAF43FF5647A4.taxon	description	Shell measurements and ratios. n = 13 specimens: largest specimen: SL: 47.5 mm, MD: 32.0 mm, mean SL: 39.4 mm (σ = 4.4), mean MD: 26.2 (σ = 3.3), spire angle: µ = 132.1 ° (σ = 8.0 °); last whorl angle: µ = 37.9 ° (σ = 1.6 °), LW: µ = 1.5 (σ = 0.04), RD: µ = 0.72 (σ = 0.02), PMD: µ = 0.91 (σ = 0.03), RSH: µ = 0.08 (σ = 0.03); the large specimen of 70 mm height, mentioned by Hoernes & Auinger (1879), is based on a misidentification.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA1FFB1FF5FAF43FF5647A4.taxon	discussion	Discussion. The colour pattern is highly reminiscent of Phasmoconus fuscocingulatus (Hörnes, 1851), which was the reason why Hörnes (1851) mixed both species. The broad conical and often fig-shaped last whorl of K. moravicus, however, allows a separation of both species (Fig. 12). Nevertheless, the original species concept of Hoernes & Auinger (1879) was blurred by lumping different species. Especially juvenile shells of Monteiroconus supracompressus (Sacco, 1893) were mixed with K. moravicus in the type lots. Similarly, Švagrovský (1982) misidentified a Monteiroconus supracompressus as K. moravicus (both species are found at the locality described by Švagrovský 1982). The more elongate last whorl of M. supracompressus and its nearly flat spire, separate both species distinctly. Paleoenvironment. Typically found in shallow marine, sandy nearshore environments with seagrass.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA1FFB1FF5FAF43FF5647A4.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Niederleis, Gainfarn, Enzesfeld, Steinebrunn, Pötzleinsdorf, Baden, Bad Vöslau (Austria), Mikulov-Kienberk (Czech Republic), Borský Mikuáš (Slovakia) (Hoernes & Auinger 1879; Švagrovský 1982); Eisenstadt-Sopron Basin: Forchtenau (Austria) (Hoernes & Auinger 1879); Oberpullendorf Basin: Ritzing (Austria) (Hoernes & Auinger 1879); Styrian Basin: Gamlitz (Austria) (Hoernes & Auinger 1879); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus (Romania) (Boettger 1906).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA0FFCFFF5FAF57FBF745A5.taxon	description	[Lithoconus] [Conus] Neumayri n. f. — Hoernes 1878 a: 195 (nomen nudum). Conus (Lithoconus) Neumayri nov. form. — Hoernes & Auinger 1879: 27, pl. 1, figs 17 – 18.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA0FFCFFF5FAF57FBF745A5.taxon	materials_examined	Type material. Syntype NHMW 1999 z 0077 / 0027, Lăpugiu de Sus (Romania), illustrated in Hoernes & Auinger (1879, pl. 1, fig. 17), 3 syntypes NHMW 1854 / 0035 / 0056, Lăpugiu de Sus (Romania), including specimen illustrated in Hoernes & Auinger (1879, pl. 1, fig. 18); middle Miocene, Badenian (Langhian). Studied material. Syntypes and 1 spec. NHMW 1843 / 0099 / 0058, Gainfarn (Austria), 29 spec. NHMW A 1628 Lăpugiu de Sus (Romania). Illustrated material. Figs 11 C 1 – C 3: Lăpugiu de Sus (Romania): SL: 27.9 mm, MD: 17.5 mm, NHMW A 1628; Figs 11 D 1 – D 3, 3 D: syntype, Lăpugiu de Sus (Romania): NHMW 1999 z 0077 / 0027, SL: 30.6 mm, MD: 20.8 mm, illustrated in Hoernes & Auinger (1879, pl. 1, fig. 17); Figs 11 E 1 – E 2: Gainfarn (Austria), SL: 30.3, MD: 19.2 mm, NHMW 1843 / 0099 / 0058; Figs 11 F 1 – F 2: Lăpugiu de Sus (Romania): SL: 29.2 mm, MD: 20.3 mm, NHMW A 1628. Revised description. Moderately small shells with low spire and broad conical last whorl. Early spire pointed; suture of early spire whorls impressed, undulating, resulting in coeloconoid, gradate outline. Later spire whorls weakly convex with deep but narrower suture. Early spire whorls striate; later whorl tops glossy, faintly striate (only visible under high magnification). Last spire whorl distinctly broadening, forming a prominent, rounded shoulder, coinciding with position of maximum diameter. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Last whorl stoutly conical to weakly ventricose; aperture moderately narrow, with subparallel margins. Siphonal canal very short, straight; fasciole short, twisted; inner lip broad, short, straight. Surface glossy with few pitted spiral grooves on base demarcating broad spiral cords. Colour pattern consisting of large rectangular blotches roughly axially and spirally arranged.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA0FFCFFF5FAF57FBF745A5.taxon	description	Shell measurements and ratios. n = 10: largest specimen: SL: 36.1 mm, MD: 23.6 mm, mean SL: 28.8 mm (σ = 3.5), mean MD: 18.5 (σ = 2.9), spire angle: µ = 128.1 ° (σ = 6.2 °), last whorl angle: µ = 37.9 ° (σ = 2.0 °), LW: µ = 1.6 (σ = 0.1), RD: µ = 0.71 (σ = 0.03), PMD: µ = 0.91 (σ = 0.03), RSH: µ = 0.09 (σ = 0.04).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA0FFCFFF5FAF57FBF745A5.taxon	discussion	Discussion. The highly characteristic colour pattern is among the most intense in Paratethyan cones under UV light and allows a quick identification in the collection, even in mixed lots. None of the specimens tends to form dots or dashes. Therefore, we reject the synonymization with Kalloconus berghausi (Michelotti, 1847) as proposed by Hall (1966) and Bałuk (1997). Moreover, the striae on the early spire whorl become obsolete much earlier and it lacks the nodules along the shoulder of the early spire whorls seen in K. berghausi. Paleoenvironment. At least the lot NHMW A 1628 from Lăpugiu de Sus (Romania) derives from sandy nearshore environments with miliolid and elphidiid foraminifers (based on sedimentfill of the shells).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFA0FFCFFF5FAF57FBF745A5.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn (Austria) (NHMW collection), Transylvanian Basin: Lăpugiu de Sus (Romania) (Hoernes & Auinger 1879).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDEFFCDFF5FAD55FC8B4034.taxon	description	? Conus (Rhizoconus) ponderosus steinabrunnensis (Sacco) — Csepreghy-Meznerics 1956: plate captions, pl. 11, figs 11 – 12. Conus (Chelyconus) ponderosus var. ponderoaustriaca (Sacco) — Kojumdgieva in Kojumdgieva & Strachimirov 1960: 214, pl. 51, fig. 1.? Conus (Chelyconus) ponderosus var. steinabrunnensis (Sacco) — Kojumdgieva in Kojumdgieva & Strachimirov 1960: 213, pl. 50, fig. 8 [non Lautoconus steinabrunnensis (Sacco, 1893)].	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDEFFCDFF5FAD55FC8B4034.taxon	description	? Conus (Chelyconus) aff. ponderoaustriacus Sacco, 1893 — Strausz 1966: 462, pl. 69, figs 7 – 8. Conus (Chelyconus) ponderosus ponderoaustriacus (Sacco, 1893) — Nicorici & Sagatovici 1973: 176, pl. 27, figs 6 – 7.? Conus (Chelyconus) ponderosus ponederoaustriacus [sic] (Sacco, 1893) — Atanacković 1985: 177, pl. 39, figs 10 – 11. Conus panderosus [sic] ponderoaustriacus (Sacco) — Ionesi & Nicorici 1994: 62, pl. 5, figs 5, 7.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDEFFCDFF5FAD55FC8B4034.taxon	materials_examined	Type material. Holotype NHMW 1854 / 0035 / 0041, Lăpugiu de Sus (Romania); middle Miocene, Badenian (Langhian). Studied material. Holotype and 1 spec. NHMW 1854 / 0035 / 0041, 1 spec. NHMW 1854 / 0035 / 0037, Lăpugiu de Sus (Romania); 2 spec. NHMW 1856 / 0050 / 0109, Gainfarn (Austria); 5 spec. NHMW 1855 / 0045 / 0358, Gainfarn (Austria); 9 spec. NHMW 1861 / 0033 / 0005, Bujtur (Romania). Illustrated material. Figs 11 G 1 – G 3, 3 P: holotype, Lăpugiu de Sus (Romania), SL: 50.4 mm, MD: 27.1 mm, NHMW 1854 / 0035 / 0041; Figs 11 H 1 – H 3: Gainfarn (Austria), SL: 49.6 mm, MD: 27.5 mm, NHMW 1855 / 0045 / 0 358. Revised description. Medium-sized shells with low conical to broad coeloconoid spire; early spire whorls nearly flat, later whorls weakly convex with last spire whorl often slightly more convex; spire whorls smooth; suture channelled. Subsutural flexure shallow, weakly curved, moderately asymmetrical. Spire passing via a rounded shoulder into conical to weakly ventricose last whorl; position of maximum diameter close below shoulder. Last whorl faintly constricted; prominent to weak spiral grooves may appear on lower third; siphonal canal short; siphonal fasciole weak and indistinct. Aperture moderately narrow, posteriorly narrowing and only weakly broadening anteriorly. Colour pattern poorly preserved, but seems to consist of densely spaced continuous spirals.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDEFFCDFF5FAD55FC8B4034.taxon	description	Shell measurements and ratios. n = 10 adult specimens: largest specimen: SL: 59.3 mm, MD: 34.1 mm, mean SL: 49.8 mm (σ = 4.6), mean MD: 27.6 mm (σ = 2.9), spire angle: µ = 115.2 ° (σ = 8.1 °), last whorl angle: µ = 34.6 ° (σ = 2.5 °), LW: µ = 1.8 (σ = 0.1), RD: µ = 0.64 (σ = 0.03), PMD: µ = 0.9 (σ = 0.02), RSH: µ = 0.1 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDEFFCDFF5FAD55FC8B4034.taxon	discussion	Discussion. Differs from Lautoconus steinabrunnensis in the rounded shoulder, convex spire whorls, lower spire and the higher position of the maximum diameter. Its aperture is narrower especially in the posterior part. Paleoenvironment. Shallow marine environments; at least at Gainfarn, the species was associated with seagrass.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDEFFCDFF5FAD55FC8B4034.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn (Austria) (Hoernes & Auinger 1879); Transylvanian Basin: Lăpugiu de Sus (Romania) (Hoernes & Auinger 1879); Pannonian Basin: Hidas, Szob (Hungary) (Csepreghy-Meznerics 1950, 1956; Strausz 1966); Zârand Basin: Minişul de Sus (Romania) (Nicorici & Sagatovici 1973); Caransebeş-Mehadia Basin: Valea Satului (Romania) (Hinculov 1968); Buzău Basin: Valea Muscelului (Romania) (Ionesi & Nicorici 1994); Dacian Basin: Dobruša (Bulgaria) (Kojumdgieva & Strachimirov 1960); Banja Luka Basin: Jazovac (Bosnia and Herzegovina) (Atanacković 1985); Karlovac-Glina Basin: Glina (Croatia) (Eremija 1959).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDCFFCDFF5FA9D9FC3046B3.taxon	materials_examined	Type material. Holotype NHMW 1849 / 0004 / 0016, Niederkreuzstetten (Austria), illustrated in Hörnes (1851, pl. 3, fig. 8). Studied material. Holotype and 5 spec. NHMW 1853 / 0003 / 0003, 2 spec. NHMW A 991, all Niederkreuzstetten (Austria). Illustrated material. Figs 13 A 1 – A 5, 3 Q: Holotype, SL: 34.5 mm, MD: 21.3 mm, NHMW 1849 / 0004 / 0016, Niederkreuzstetten (Austria); Figs 13 B 1 – B 3: SL: 53.9 mm, MD: 34.0 mm, NHMW A 991, Niederkreuzstetten (Austria). Revised description. Medium-sized, robust shells; moderately high, broad conical to weakly cyrtoconoid, spire. Convex, bulgy spire whorls with deeply incised, channelled suture; early whorls striate with undulating suture. Last spire whorl less convex; subsutural flexure deep, moderately curved, strongly asymmetrical. Distinct, subangulate shoulder coinciding with position of maximum diameter. Conical last whorl rapidly contracting, rarely with weak concavity below shoulder. Weakly constricted at base; siphonal fasciole weak; siphonal canal moderately long, slightly reflected. Aperture narrow with subparallel margins. Very weak spiral cords on base; irregularly spaced growth lines on last whorl may form prominent grooves. Colour pattern consisting of broad flammulae on spiral whorls, parallel to subsutural flexure; two broad dark bands below shoulder and on the posterior third with spirally arranged, somewhat irregularly spaced light dots on entire last whorl. Major growth lines coincide with thin, light axial threads in UV light.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDCFFCDFF5FA9D9FC3046B3.taxon	description	Shell ratios and measurement. n = 7 adult specimens: largest specimen: SL: 53.9 mm, MD: 35.5 mm, mean SL: 48.6 mm (σ = 3.6), mean MD: 31.4 mm (σ = 3.2), spire angle: µ = 106.1 ° (σ = 8.6 °), last whorl angle: µ = 38.2 ° (σ = 1.4 °), LW: µ = 1.55 (σ = 0.09), RD: µ = 0.76 (σ = 0.05), PMD: µ = 0.89 (σ = 0.02), RSH: µ = 0.14 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDCFFCDFF5FA9D9FC3046B3.taxon	discussion	Discussion. This species was mixed with Conus steindachneri Hoernes, 1879 by Hoernes & Auinger (1879) and subsequent authors (e. g. Harzhauser 2002). It differs in the narrower last spire whorl, the higher, broad conical to cyrtoconoid spire, the broader last whorl, the weaker siphonal fasciole and the shorter siphonal canal. A further difference is the colour pattern of broad flammulae on the spire whorls, which are absent in Lautoconus steindachneri. Kalloconus ponderovagus is also reminiscent of Kalloconus tschermaki (Hoernes & Auinger, 1879) but differs in its smaller size and the colour pattern, which consist of wide-spaced spirals in K. tschermaki. Sacco (1893 b) introduced this name for the specimen illustrated by Hörnes (1851, pl. 3, fig. 8), which he recognised as “ totally different from Conus steindachneri ”, thus rejecting the synonymization by Hoernes & Auinger (1879). Although we agree with Sacco (193 b), his act was correct just by chance, as the specimen of Hörnes (1851) is a subadult specimen, lacking the characteristic broad shoulder. Therefore, we provide illustrations of adult specimens from the type locality. Conus parvicaudatus Sacco, 1893, from the early Miocene of the Turin Hills, differs in the pronounced constriction of the last whorl and the higher position of the adapical part of the aperture, which reaches above the shoulder.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDCFFCDFF5FA9D9FC3046B3.taxon	description	Paleoenvironment. The species occurs in sandy shallow water deposits.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDCFFCDFF5FA9D9FC3046B3.taxon	distribution	Distribution in Paratethys. Karpatian (early Miocene): Alpine-Carpathian Foredeep: Niederkreuzstetten (Austria); Korneuburg Basin: Karnabrunn (Austria) (Harzhauser 2002).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDBFFCAFF5FA8E1FA0B4762.taxon	materials_examined	Holotype: Figs 13 E 1 – E 3: SL: 55.2 mm, MD: 38.4 mm, NHMW 1868 / 0001 / 0380 b. Paratype: Figs 13 C 1 – C 4: SL: 49.4 mm, MD: 34.5 mm, NHMW 1868 / 0001 / 0380 a.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDBFFCAFF5FA8E1FA0B4762.taxon	description	Paratype: Figs 13 D: SL: 53.2 mm, MD: 35.4 mm, NHMW 1868 / 0001 / 0380 d. Paratype: Figs 13 F 1 – F 2, 3 R: SL: 49.9 mm, MD: 35.9 mm, NHMW 1868 / 0001 / 0380 c. Additional paratypes: 2 specimens, NHMW 1868 / 0001 / 0380, all specimens from Lăpugiu de Sus (Romania).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDBFFCAFF5FA8E1FA0B4762.taxon	materials_examined	Type stratum: Badenian marly-clayey deposits with thin interlayers of sand and corallinacean limestones. Type locality: Lăpugiu de Sus (Romania). Age: Middle Miocene, early Badenian (= Langhian).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDBFFCAFF5FA8E1FA0B4762.taxon	etymology	Etymology: Referring to Kalloconus hungaricus with which this species was mixed by Hoernes & Auinger (1879).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDBFFCAFF5FA8E1FA0B4762.taxon	description	Description. Robust, medium-sized shells; spire depressed and conical with low coeloconoid early whorls. Spire whorls nearly flat, smooth; last spire whorl forming a very shallow concavity close to the prominent, subangulate shoulder. Suture narrow but deeply incised. Subsutural flexure deep, strongly curved, strongly asymmetrical. Last whorl rapidly contracting, straight sided; position of maximum diameter close below shoulder. Lower third of last whorl with weak spiral grooves. Aperture narrow, with subparallel margins, only weakly broadening towards siphonal canal. Siphonal fasciole weak; siphonal canal moderately long, weakly reflected. Highly variable colour pattern consisting of broad flammulae on spire whorls and spirally arranged rows of small to moderately large subquadrate blotches on last whorl; delicate spirals of small dots may be intercalated; large blotches are indistinct. Shell measurements and ratios. n = 6: largest specimen: SL: 55.2 mm, MD: 38.4 mm, mean SL: 52.4 mm (σ = 2.3), mean MD: 35.6 mm (σ = 1.5), spire angle: µ = 134.7 ° (σ = 6.4 °), last whorl angle: µ = 40.5 ° (σ = 0.6 °), LW: µ = 1.47 (σ = 0.06), RD: µ = 0.74 (σ = 0.02), PMD: µ = 0.90 (σ = 0.01), RSH: µ = 0.08 (σ = 0.03). Paleoenvironment. Unknown; maybe offshore environments as represented by the clays of the type locality.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDBFFCAFF5FA8E1FA0B4762.taxon	discussion	Discussion. The specimens from Lăpugiu de Sus (Romania) were mixed by Hoernes & Auinger (1879) with Kalloconus hungaricus from which they differ in their straight-sided, conical last whorl and the narrower shoulder. In addition, the conspicuous colour pattern is unknown from K. hungaricus.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDBFFCAFF5FA8E1FA0B4762.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Transylvanian Basin: Lăpugiu de Sus (Romania).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDBFFC8FF5FAE2DFDC5443D.taxon	description	[Lithoconus] [Conus] Tietzei n. f. — Hoernes 1878 a: 195 (nomen nudum). Conus (Lithoconus) Tietzei nov. form. — Hoernes & Auinger 1879: 28 (partim), pl. 1, fig. 3. Conus (Lithoconus) tietzei (Hoernes & Auinger) — Atanacković 1969: 215, pl. 12, figs 14 – 15.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDBFFC8FF5FAE2DFDC5443D.taxon	materials_examined	Type material. Syntype NHMW 1847 / 0046 / 0004, Szob (Hungary), illustrated in Hoernes & Auinger (1879 pl. 1, fig. 3); middle Miocene, Badenian (Langhian). The specimen from Lăpugiu de Sus (NHMW 1870 / 0033 / 0002), mentioned by Hoernes & Auinger (1879) in their description of Conus tietzei, is most probably not conspecific with K. tietzei and may represent a juvenile M. hoernesi. Studied material. Syntype. Illustrated material. Figs 13 G 1 – G 5, 3 S: sytype, Szob (Hungary): SL: 62.1 mm, MD: 41.2 mm, NHMW 1847 / 0046 / 0004, illustrated in Hoernes & Auinger (1879 pl. 1, fig. 3). Revised description. Moderately large shell with depressed spire; early spire coeloconoid, later spire nearly flat; spire whorls faintly concave; smooth except for delicate growth lines; irregular, impressed suture. Subsutural flexure of medium depth, moderately curved, strongly asymmetrical. Rounded but prominent shoulder with position of maximum diameter closely below shoulder. Last whorl straight-sided and conical, not constricted; broad, low spiral cords on base. Aperture moderately narrow, widening abapically into weakly recurved, short siphonal canal; siphonal fasciole weak, twisted, poorly demarcated from base and inner lip. Colour pattern consisting of narrowly spaced spiral dashes.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDBFFC8FF5FAE2DFDC5443D.taxon	description	Shell measurements and ratios. Only one syntype is available: SL: 62.1 mm, MD: 41.2 mm, spire angle: 141 ° (without coeloconoid early spire), last whorl angle: 41 °, LW: 1.51, RD: 0.72, PMD: 0.85, RSH: 0.08.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDBFFC8FF5FAE2DFDC5443D.taxon	discussion	Discussion. Kalloconus hungaricus (Hoernes & Auinger, 1879) is very similar in colour-pattern and general shape to K. tietzei, which could be considered simply a more slender morphotype of K. hungaricus. However, the coeloconoid profile of the early spire, the better-defined shoulder, the narrower last spire whorl and the narrower angle of the last whorl seem to justify the separation. A rare species, which is at first sight also similar to several Paratethyan Monteiroconus species, but can but can be separated from these by the absence of spiral sculpture on the spire whorls. The only Monteiroconus species with smooth spire whorls is M. boeckhi (Halaváts, 1884), which can be immediately separated by its rather odd profile, constricted both just below the shoulder and the base and by its more twisted and slightly longer siphonal canal. Among the specimens from Letkés described by Kovács & Vicián (2013) the specimen illustrated as fig. 93 might represent K. tietzei, although it is slightly ventricose and its shoulder is slightly more rounded. The other specimens (figs. 92, 94) are distinctly smaller and more slender. Specimens of this species from Letkés in the collections of the NHMW have striate spire whorls, a broad last spire whorl and a densely spaced spiral colour pattern of delicate dashes, thus differing from K. tietzei and are described herein as Kalloconus letkesensis nov. sp. Paleoenvironment. The mollusc fauna and lithofacies of Szob suggest a sandy coastal environment with sea grass (Dulai 1996).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFDBFFC8FF5FAE2DFDC5443D.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Pannonian Basin: Letkés, Szob, Diósd, Kemence (Hungary) (Csepreghy-Meznerics 1956; Kovács & Vicián 2013); southern Pannonian Basin: Miljevići (Bosnia and Herzegovina) (Atanacković 1985).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD9FFC9FF5FADDCFA9742A4.taxon	description	[Rhizoconus] Conus Tschermaki n. f. — Hoernes 1878 a: 195 (nomen nudum). [Rhizoconus] [Conus] Bittneri n. f. — Hoernes 1878 a: 195 (nomen nudum).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD9FFC9FF5FADDCFA9742A4.taxon	materials_examined	Type material. Syntype NHMW 1864 / 0001 / 0499, Niederkreuzstetten (Austria), illustrated in Hoernes & Auinger (1879, pl. 1, fig. 2); early Miocene, Karpatian (late Burdigalian); syntype NHMW 1853 / 0003 / 0003 a Gainfarn (Austria) illustrated in Hoernes & Auinger (1879, pl. 5, fig. 2); middle Miocene, Badenian (late Langhian). Gainfarn (Austria) is also the type locality of Conus bittneri Hoernes & Auinger 1879 (considered to be a junior synonym of C. tschermaki). Studied material. Syntypes and 2 spec. Niederkreuzstetten (Austria), NHMW 1846 / 0037 / 0063; 5 spec. Niederkreuzstetten (Austria), NHMW 1846 / 0037 / 0044; 1 spec. Gainfarn (Austria) NHMW 1853 / 0003 / 0003 b, 1 spec. Gainfarn (Austria) NHMW 1853 / 0010 / 0003, 3 spec. Gainfarn (Austria) NHMW 1853 / 0010 / 0002; 4 spec. Enzesfeld (Austria) 4 NHMW A 460. Illustrated material. Figs 14 A 1 – A 3: syntype, Gainfarn (Austria): SL: 63.4 mm, MD: 37.0 mm, NHMW 1853 / 0003 / 0003 a, illustrated in Hoernes & Auinger (1879, pl. 5, fig. 2); Figs 14 B 1 – B 4, 3 T: syntype, Niederkreuzstetten (Austria), SL: 75.3 mm, MD: 43.9 mm, NHMW 1864 / 0001 / 0499, illustrated in Hoernes & Auinger (1879, pl. 1, fig. 2); NHMW 1846 / 0037 / 0044. Revised description. Moderately large, robust shells. Spire regularly conical, elevated, with moderately convex, smooth, bulgy whorls; suture deeply impressed to channelled, wavy and irregular. Last spire whorl less convex, forming a low-angled sutural ramp structured by prominent growth lines. Subsutural flexure of medium depth, moderately curved, moderately asymmetrical. Shoulder with rounded angulation coinciding with position of maximum diameter; last whorl regularly conical, not constricted; aperture straight, rather narrow, only weakly broadening towards short, weakly recurved canal; fasciole weak, slightly twisted and not well demarcated from base and lip. Faint spiral cords may appear on base. Colour pattern of last whorl consisting of slightly wavy and frequently interrupted spiral lines, separated by wider interspaces.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD9FFC9FF5FADDCFA9742A4.taxon	description	Shell measurements and ratios. n = 8: largest specimen: SL: 75.3 mm, MD: 43.9 mm, mean SL: 62.7 mm (σ = 7.5), mean MD: 36.8 mm (σ = 3.9), spire angle: µ = 108.3 ° (σ = 7.5 °), last whorl angle: µ = 38.5 ° (σ = 2.0 °), LW: µ = 1.7 (σ = 0.06), RD: µ = 0.7 (σ = 0.02), PMD: µ = 0.93 (σ = 0.02), RSH: µ = 0.16 (σ = 0.02).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD9FFC9FF5FADDCFA9742A4.taxon	discussion	Discussion. Monteiroconus hoernesi (Doderlein, 1863) is superficially similar but differs in its broader shape, the deeper subsutural flexure and the wider and straight siphonal canal. Conus bittneri Hoernes & Auinger, 1879, from Gainfarn, is most probably just an aberrant Kalloconus tschermaki with a pathologic spire. Paleoenvironment. All occurrences point to shallow marine coastal habitats (e. g. Harzhauser et al. 2002; Zuschin et al. 2007).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD9FFC9FF5FADDCFA9742A4.taxon	distribution	Distribution in Paratethys. Karpatian (early Miocene): Alpine-Carpathian Foredeep: Niederkreuzstetten (Austria) (Hoernes & Auinger 1879); Badenian (middle Miocene): Vienna Basin: Gainfarn, Enzesfeld, Steinebrunn (Austria) (Hoernes & Auinger 1879); Eisenstadt-Sopron Basin: Forchtenau (Hoernes & Auinger 1879).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD8FFC7FF5FAC57FEF6439A.taxon	description	Conus clavatus Lam. — Hörnes 1851: 25 (partim), pl. 2, figs 5 a – b. [Dendroconus] [Conus] Vöslauensis n. f. — Hoernes 1878 a: 195 (nomen nudum). Conus (Dendroconus) Voeslauensis nov. form. — Hoernes & Auinger 1879 (partim): 22, pl. 1, fig. 8, pl. 3, fig. 4. [Conus (Chelyconus) mediterraneus] var. ponderoclavata Sacc. — Sacco 1893 b: 104 (nov. nom. pro Conus clavatus in Hörnes 1851, pl. 2, fig. 5). Conus (Dendroconus) voeslauensis Hoernes et Auinger — Csepreghy-Meznerics 1956: 122, pl. 11, figs 3 – 4.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD8FFC7FF5FAC57FEF6439A.taxon	materials_examined	Type material. 6 syntypes NHMW 1849 / 0023 / 0005, Bad Vöslau (Austria) including specimen illustrated in Hoernes & Auinger (1879, pl. 3, fig. 4), 2 syntypes NHMW 1851 / 0010 / 0002, Mikulov-Kienberk (Czech Republic), including specimen illustrated in Hoernes & Auinger (1879, pl. 1, fig. 8); all middle Miocene, Badenian (Langhian). Studied material. Syntypes and 3 spec. NHMW 1853 / 0010 / 0005, Enzesfeld (Austria), 6 spec. NHMW A 1633, Enzesfeld (Austria), 3 spec. NHMW 1849 / 0033 / 0002, Bad Vöslau; 1 spec. NHMW 1846 / 0037 / 0049, Gainfarn (Austria), illustrated in Hörnes (1851, pl. 2, fig. 5). Illustrated material. Figs 14 C 1 – C 3: syntype, Bad Vöslau (Austria): SL: 55.5 mm, MD: 34.4 mm, NHMW 1849 / 0023 / 0005 a, specimen illustrated in Hoernes & Auinger (1879, pl. 3, fig. 4); Figs 14 D 1 – D 3: syntype, Enzesfeld (Austria): SL: 52.0 mm, MD: 31.0 mm, NHMW 1853 / 0010 / 0005; Fig. 3 U: syntype, Bad Vöslau (Austria): SL: 42.8 mm, MD: 27.4 mm, NHMW 1849 / 0023 / 0005 b. Revised description. Medium-sized solid shells; low conical to weakly coeloconoid spire; spire whorls bulgy and convex, delicate spiral threads on early whorls, later smooth except for prominent growth lines, suture deeply channelled. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Shoulder rounded, rarely subangulate, position of maximum diameter slightly below; last whorl only weakly ventricose or straight-sided, slightly constricted at base, typically with several strongly incised growth lines. Faint spiral cords on base; entire whorl covered with delicate spiral sculpture, visible only in tangential light. Siphonal canal short, slightly recurved; siphonal fasciole weakly swollen, twisted with prominent growth lines. Aperture moderately narrow, widening towards siphonal canal. Colour pattern consisting of numerous spirals (no subquadratic dots can be observed as illustrated by Hoernes & Auinger 1879).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD8FFC7FF5FAC57FEF6439A.taxon	description	Shell measurements and ratios. n = 10: largest specimen: SL: 61.8 mm, MD: 35.0 mm, mean SL: 50.2 mm (σ = 5.7), mean MD: 31.2 (σ = 2.8), spire angle: µ = 118.8 ° (σ = 4.7 °), last whorl angle: µ = 38.9 ° (σ = 1.0 °), LW: µ = 1.6 (σ = 0.08), RD: µ = 0.71 (σ = 0.03), PMD: µ = 0.87 (σ = 0.02), RSH: µ = 0.12 (σ = 0.01).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD8FFC7FF5FAC57FEF6439A.taxon	discussion	Discussion. Most shells which can be reliably identified as Kalloconus voeslauensis are characterised by several deeply incised growth lines on the last whorl. In their original description of this species, Hoernes & Auinger (1879) referred also to specimens from Lăpugiu de Sus, which, however, represent Kalloconus hungaricus. The specimen illustrated by Hörnes (1851, pl. 2, figs 5 a – b) as Conus clavatus is an aberrant specimen of Kalloconus voeslauensis. Sacco (1893 b) introduced ponderoclavata as varietal name for this aberrant specimen, which thus is a subjective junior synonym of Conus voeslauensis. Paleoenvironment. The specimens from Bad Vöslau are filled with clay and might have dwelled in basinal settings. Mikulov-Kienberk and Enzesfeld, in contrast, represent shallow marine environments (own data).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD8FFC7FF5FAC57FEF6439A.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Alpine-Carpathian Foredeep: Grund (Austria); Vienna Basin: Bad Vöslau, Gainfarn (Austria), Mikulov-Kienberk (Czech Republic) (Hoernes & Auinger 1879); Eisenstadt-Sopron Basin: Forchtenau (Austria) (Sieber 1956); Transylvanian Basin: Lăpugiu de Sus (Romania) (Chira & Voia 2001); Pannonian Basin: Szob, Budapest, Mecsek Mountains (Hungary) (Csepreghy-Meznerics 1956; Strausz 1966).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD6FFC7FF5FAB32FC7C4797.taxon	materials_examined	Type species (by original designation): Conus mediterraneus Hwass in Bruguière, 1792. Recent, Mediterranean Sea. Note. Tucker & Tenorio (2009) gave few shell features by which to distinguish members of the genus Lautoconus Monterosato, 1923. The shells are turgid in shape with convex sides. The protoconch is paucispiral. The whorl tops are ornamented with cords that reach the middle spire whorls and often persist. The subsutural flexure is shallow to moderate in depth. Puillandre et al. (2014 a, b) synonymized Varioconus da Motta, 1991 with Lautoconus Monterosato, 1923 based on molecular data. According to Tucker & Tenorio (2009) the genus Varioconus da Motta, 1991 is characterised by shells with smooth spire whorls, a rounded, indistinct shoulder, a shallow subsutural sinus and a paucispiral protoconch. For their molecular phylogeny Puillandre et al. (2014 a, b) analysed numerous species, traditionally placed in Lautoconus but only two species, which are listed as Varioconus by Tucker & Tenorio (2009) (Conus xicoi Röckel, 1987 and C. franciscoi Rolán & Röckel, 2000). Unfortunately, Conus variegatus Kiener, 1845, the type species of Varioconus, is missing from their analysis. Therefore, the synonymization remains debateable. The main difference in shell characters listed by Tucker & Tenorio (2009) is the presence of cords on Lautoconus and their absence in Varioconus. The characteristic pyriform outline of most species placed in Varioconus is not a distinguishing feature but may also be developed by Lautoconus species such as Lautoconus mercator (Linnaeus, 1758). The radula morphologies as described by Tucker & Tenorio (2009) do not require a separation and their West African geographic distribution is identical. Consequently, we follow Puillandre et al. (2014 a, b) and treat Varioconus as subjective junior synonym of Lautoconus. The genus Lautoconus is represented in the middle Miocene Paratethys Sea by several species, which comprise moderately small to moderately large and medium-wide to wide shells with low to medium high spires; spiral cords on the sutural ramp may be present or replaced by faint spiral threads. The subsutural flexures are variable in depth, ranging from very shallow to deep, are usually moderately curved and moderately asymmetrical. To our knowledge, none of the extant Lautoconus (and Varioconus) species has a tuberculate early spire or develops nodes. Therefore, we exclude tuberculate species from Lautoconus.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD6FFC5FF5FAF27FE844191.taxon	description	Conus Andréei n. f. — Kittl 1887: 241, pl. 8, figs 2 a – b.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD6FFC5FF5FAF27FE844191.taxon	materials_examined	Type material. Holotype: NHMW 1888 / 0001 / 0008, Jaklowetz at Ostrava (Czech Republic); middle Miocene, early Badenian (Langhian). Studied material. Holotype. Illustrated material. 15 A 1 – A 3: Jaklowetz at Ostrava (Czech Republic): SL: 70 mm, MD: 39.5 mm, NHMW 1888 / 0001 / 0008. Revised description. Large solid shell with cyrtoconoid, bulbous spire with high, convex, weakly angulated whorls; deep suture. Flat to slightly concave, steeply sloping sutural ramp on last whorl; shoulder angulated. Subsutural flexure not preserved. Last whorl cylindrical below shoulder and distinctly constricted at base. Siphonal canal not preserved. Aperture moderately wide, anteriorly flaring. Shell surface smooth.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD6FFC5FF5FAF27FE844191.taxon	discussion	Discussion. Only a single fragmentary specimen with recrystallized shell is available. As already pointed out by Kittl (1887), the barrel-shaped last whorl characterises this rare species and separates it from all other Paratethyan cones. The poor preservation and lack of important conchological features make our generic placement provisional. Paleoenvironment. The mollusc assemblages from the Ostrava region suggest a deep water environment. Distribution in Paratethys. Badenian (middle Miocene): Carpathian Foredeep: Jaklowetz at Ostrava (Czech Republic).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD4FFC2FF5FA924FF56445C.taxon	description	Conus Eschewegi Costa — Pereira da Costa 1866: 29, pl. 9, figs 18, 19 a – b, 21, 23 a – b [non figs 20, 22, 34].	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD4FFC2FF5FA924FF56445C.taxon	description	[Conus] D. [endroconus] Eschewegi var. depressoastensis Sacc. — Sacco 1893 a: 13, pl. 1, fig. 25. Dendroconus pyruloides (Dod. Sacc.) — Sacco 1893 a: 13, pl. 1, fig. 26. Dendroconus cf. Eschewegi Da Cost. var. caelata Dod., Sacc. — Szalay 1926: 334.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD4FFC2FF5FA924FF56445C.taxon	description	Lautoconus eschewegi (da Costa) — Kovács & Vicián 2013: 70, figs 7, 56 – 61.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD4FFC2FF5FA924FF56445C.taxon	materials_examined	Type material. Specimens illustrated in Pereira da Costa (1866, pl. 9, figs 23 a – b, selected as lectotype proposed by Sacco (1893 a); Tortonian, Cacela Basin, Portugal. The whereabouts of the collection are unknown to us. Studied material. 7 spec. NHMW 1858 / 0043 / 0007, Lăpugiu de Sus (Romania), including specimen illustrated in Hoernes & Auinger (1879, pl. 1, fig. 20), 7 spec. NHMW 1854 / 0035 / 0035, Lăpugiu de Sus (Romania), 1 spec NHMW A 1609, Lăpugiu de Sus (Romania); 11 spec. NHMW 2016 / 0007 / 0001, Letkés (Hungary). Illustrated material. Figs 16 A 1 – A 2,: Lăpugiu de Sus (Romania), SL: 39.9 mm, MD: 22.9 mm, NHMW 1858 / 0043 / 0007; Figs 16 B 1 – B 2, 3 V: Lăpugiu de Sus (Romania), SL: 42.8 mm, MD: 23.8 mm, NHMW 1858 / 0043 / 0007, specimen illustrated in Hoernes & Auinger (1879, pl. 1, fig. 20); Figs 16 C 1 – C 2: Letkés (Hungary), SL: 32.6 mm, MD: 18.7 mm, NHMW 2016 / 0007 / 0001 b; Figs 16 D 1 – D 2: Letkés (Hungary), SL: 39.6 mm, MD: 23.2 mm, NHMW 2016 / 0007 / 0001 a. Revised description. Medium-sized, pyriform shells; extraordinarily pointed, turreted early spire (but poorly preserved in all specimens); later spire low conical to cyrtoconoid with channelled suture; spiral whorls slightly convex; subsutural flexure very shallow, weakly curved, moderately asymmetrical; shoulder strongly rounded, slightly subangulate in fully grown shells. Last whorl elongate and conical, constricted at base. Moderately long, broad and reflected siphonal canal; aperture moderately wide, slightly widening anteriorly. Siphonal fasciole swollen, strongly twisted, not well demarcated from base and straight, broad inner lip. Few wavy spiral grooves on base demarcating broad spiral cords. Colour pattern consisting of slightly irregularly spaced spirals formed by short dashes and subquadratic dots; size of dots rather homogenous within a spiral but may vary in size and density between spirals; sometimes two spirals form a close-spaced pair with nearly amalgamating dots. Dots become axially elongate on shoulder, rarely forming an inverted “ Y ”; narrow axially elongated stripes on spire whorls.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD4FFC2FF5FA924FF56445C.taxon	description	Shell measurements and ratios. n = 7: largest specimen: SL: 42.8 mm, MD: 23.8 mm, mean SL: 37.4 mm (σ = 3.5), mean MD: 20.8 mm (σ = 2.1), spire angle: µ = 113.3 ° (σ = 8.6 °), last whorl angle: µ = 37.0 ° (σ = 1.4 °), LW: µ = 1.8 (σ = 0.04), RD: µ = 0.63 (σ = 0.01), PMD: µ = 0.86 (σ = 0.01), RSH: µ = 0.12 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD4FFC2FF5FA924FF56445C.taxon	discussion	Discussion. Tucker & Tenorio (2009) proposed a placement of this species in Lautoconus Monterosato, 1923. The completely smooth spire whorls, which lack any sculpture even on earliest teleoconch whorls and the mode of siphonal canal formation would perfectly fit in Varioconus da Motta, 1991, which however is currently considered to be a junior synonym of Lautoconus (Puillandre et al. 2014 a, b). Hoernes & Auinger (1979) illustrated three shells as “ Dendroconus subraristriatus ”, which represent three different species, of which none is conspecific with Lautoconus subraristriatus (Pereira da Costa, 1866). The reason for uniting these different morphologies in a single species was the superficial similarity in colour patterns and the also much too broad species concept of Pereira da Costa (1866) (see Landau et al. 2013 and discussion of Lautoconus subraristriatus). The original description of Lautoconus eschewegi (Pereira da Costa, 1866) united several morphotypes, which are most probably not conspecific. Therefore, Sacco (1893 a) designated the slender, pyriform specimen illustrated by Pereira da Costa (1866, fig. 23) as type of this species. This species was also described as Conus pyruloides Sacco, 1893, from the late Miocene of Italy, which was considered to be a junior synonym of L. eschewegi by Hall (1966) and Davoli (1972). In the material from Letkés in the collection of the NHMW, this species is easily recognised under UV light by its conspicuous and intense colour pattern. This also allows the identification of subadult shells, which have a weaker siphonal fasciole, shorter siphonal canal and an evenly rounded shoulder. Therefore, we consider the specimens identified by Kovács & Vicián (2013) as Lautoconus belus (d’Orbigny, 1852) and L. pyrula (Brocchi, 1814) to represent different morphotypes and growth stages of L. eschewegi. Moreover, Lautoconus belus sensu Hall (1966, pl. 22, figs 15 – 16) and Muñiz-Solís (1999, figs 7 A – B) differs from the Paratethyan species in its broader outline, better defined shoulder and the conspicuous spiral sculpture on the last whorl and L. pyrula has a strong spiral sculpture on the base, a much weaker fasciole and a well-defined shoulder. Muñiz-Solís (1999) described Conus eschewegi from the lower upper Pliocene of Estepona. These specimens develop a comparatively broad conical last whorl and are most probably not conspecific with L. eschewegi. The superficially similar Kalloconus fuscocingulatus (Hörnes, 1851) differs in the colour pattern of continuous spirals and the tuberculate spire whorls. Paleoenvironment. Probably shallow marine environments based on the coral-associated assemblages at species Letkés in Hungary (Kovács & Vicián 2013).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD4FFC2FF5FA924FF56445C.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Transylvanian Basin: Lăpugiu de Sus (Romania); Pannonian Basin: Pannonian Basin: Várpalota, Letkés (Hungary) (Szalay 1926; Kovács & Vicián 2013). Proto-Mediterranean Sea and northeastern Atlantic. Tortonian: Cacela Basin: Cacela Velha (Portugal) (Pereira da Costa 1866); Sant'Agata Fossili, Stazzano, Montegibbio (Po Basin, Italy) (Sacco, 1893 a; Davoli 1972); Casa Nova Calises, Forlì (Apennines, Italy) (Ruggieri & Davoli 1984); Messinian: Borelli (Turin Hills) (Davoli 2003). The early Pliocene occurrences from Vezza d’Alba and Asti (Po Basin, Italy) need confirmation (Davoli 1972).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD3FFC0FF5FADFBFE3E47EC.taxon	materials_examined	Holotype: Figs 16 E 1 – E 3: SL: 19.8 mm, MD: 11.4 mm, NHMW 2016 / 0003 / 0001. Paratype: Figs 16 F 1 – F 3: SL: 18.8 mm, MD: 11.8 mm, NHMW 2016 / 0003 / 0003. Paratype: Figs 16 G 1 – G 2: SL: 20.1 mm, MD: 12.1 mm, NHMW 2016 / 0003 / 0002. Paratype: Figs 16 H 1 – H 3: SL: 18.7 mm, MD: 11.7 mm, NHMW 2016 / 0003 / 0004 a. Paratype: Figs 16 I 1 – I 3: SL: 17.2 mm, MD: 10.0 mm, NHMW 2016 / 0003 / 0004 a. Additional material. 7 specimens NHMW 2016 / 0003 / 0004, Fig. 3 W: SL: 18.3 mm, MD: 10.6 mm, all specimens from Letkés, Hungary. Type stratum. fossil-rich marly sand with coral blocks and andesite boulders of the Sámsonháza Formation (Császár 1997). Type locality. Letkés at the western part of the Börzsöny Mts. (Hungary); see Kovács & Vicián (2013) for a map and description. Age: Middle Miocene, early Badenian (= Langhian).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD3FFC0FF5FADFBFE3E47EC.taxon	etymology	Etymology. In honour of Zoltán Kovács (Franz Liszt Academy of Music, Budapest), who published the milestone paper “ Kovács & Vicián 2013 ” on the Badenian Conidae from Letkés.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD3FFC0FF5FADFBFE3E47EC.taxon	description	Description: Small, globose shells with mammillate, moderately high spires and convex last whorls. Early spire whorls pointed but usually eroded; later spire whorls faintly striate to smooth, weakly to strongly convex, partly bulgy, forming conical to cyrtoconoid, gradate spire; suture deeply incised, not channelled. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Last whorl with weakly angulated shoulder, position of maximum diameter slightly below shoulder. Inflated to cylindrical below shoulder and rapidly contracting in lower half of shell. Strongly constricted at base separating a comparatively narrow and moderately long, weakly recurved siphonal canal. Few spiral cords on base; faint spiral cords may appear on entire last whorl. Siphonal fasciole nearly absent. Colour pattern very strong in UV light, consisting of four broad bands of partly amalgamated and partly well separated blotches; first one closely below shoulder, second one slightly above midwhorl, third one below mid-whorl above constriction, fourth one on base and siphonal channel. Blotches of the uppermost band continue as narrower flammulae on the last spire whorl; additional wide-spaced flammulae on spire whorls. Shell measurements and ratios. n = 11: largest specimen: SL: 20.3 mm, MD: 12.9 mm, mean SL: 18.6 mm (σ = 1.4), mean MD: 11.4 mm (σ = 0.9), spire angle: µ = 102 ° (σ = 8.2), last whorl angle: µ = 42.3 ° (σ = 3.0), LW: µ = 1.64 (σ = 0.09), RD: µ = 0.74 (σ = 0.06), PMD: µ = 0.80 (σ = 0.03), RSH: µ = 0.17 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD3FFC0FF5FADFBFE3E47EC.taxon	discussion	Discussion. This is a very peculiar small cone, which is characterised by its globose, columbellid shape unknown from any other Paratethyan species. It is reminiscent of some species of the Indo-West Pacific genus Pseudolilliconus Tucker & Tenorio, 2009, such as Pseudolilliconus traillii (Adams, 1855) and Pseudolilliconus wallacei (Lorenz & Morrison, 2004), but the much larger size of the Paratethyan species makes a direct relation with this tiny cone genus unlikely. This strange species was identified by Kovács & Vicián (2013) as Varioconus taurinensis (Bellardi & Michelotti, 1840), which develops also a weakly gradate spire but differs in its elongate and straight-sided last whorl (see Landau et al. 2013 and references therein). Artemidiconus dertogranularis (Sacco, 1893), from the Tortonian of Stazzano in Italy, is slightly similar in shape and size but differs in its typical midwhorl groove on the spire whorls. This species is known so far only from Letkés. A non-Paratethyan occurrence, however, might be represented by the specimen illustrated in Peyrot (1931, pl. 2, fig. 11) as Conus mucronatolaevis globospira (Sacco) from the Langhian of Salies-de-Béarn in the Aquitaine Basin. Conus globospira Sacco, 1893, from the Burdigalian of the Turin Hills, differs clearly in its much more elongate last whorl. Paleoenvironment. The locality Letkés comprises a mixed assemblage of partly reworked taxa (Kovács & Vicián 2013); the frequent occurrence of corals suggests a shallow marine depositional environment.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD3FFC0FF5FADFBFE3E47EC.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Pannonian Basin: Letkés (Hungary). Proto-Mediterranean Sea and north eastern Atlantic.? Langhian (middle Miocene): Aquitaine Basin: Salies-de-Béarn (France).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD1FFC1FF5FAEABFB7545A4.taxon	materials_examined	Type material. Holotype NHMW 2016 / 0001 / 0001, Lăpugiu de Sus (Romania), illustrated in Hoernes & Auinger (1879, pl. 6, fig. 5). Studied material. Holotype and 1 spec. NHMW 2016 / 0001 / 0002, illustrated in Hoernes & Auinger (1879, pl. 1, fig. 6), 1 spec. NHMW 1870 / 0033 / 0002, all Lăpugiu de Sus (Romania); middle Miocene, Badenian (Langhian). Illustrated material. Figs 18 A 1 – A 3: Lăpugiu de Sus (Romania): SL: 65.5 mm, MD: 45.1 mm, NHMW 2016 / 0001 / 0002, illustrated in Hoernes & Auinger (1879, pl. 1, fig. 7); Figs 18 B 1 – B 3: holotype, Lăpugiu de Sus (Romania): SL: 68.2 mm, MD: 44.0 mm, NHMW 2016 / 0001 / 0001, illustrated in Hoernes & Auinger (1879, pl. 1, fig. 6); Figs 18 C 1 – C 2, 17 A: Lăpugiu de Sus (Romania), SL: 59.3, MD: 38.4 mm, NHMW 1870 / 0033 / 0002. Revised description. Moderately large, solid shells with low spire and stout, ventricose, quickly contracting last whorl. Early spire whorls eroded in all specimens; later spire whorls concave with bulgy abapical convexity, faintly striate; suture narrow, shallow. Subsutural flexure deep, strongly curved, moderately asymmetrical. Last whorl with prominent, rounded shoulder; position of maximum diameter slightly below; weakly constricted at base. Faint spiral sculpture on base. Siphonal fasciole narrow, slightly swollen, twisted; siphonal canal short and wide. Aperture moderately wide, broadening towards siphonal canal. Colour pattern consisting of strong flammulae on spire whorls and shoulder and densely spaced delicate spirally arranged dashes on last whorl.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD1FFC1FF5FAEABFB7545A4.taxon	description	Shell measurements and ratios. Only three specimens are available: SL: 68.2 / 65.5 / 59.3 mm, MD: 44.0 / 45.1 / 38.4 mm, spire angle: 132 / 141 / 130 °, last whorl angle: 42 / 45 / 44 °, LW: 1.55 / 1.45 / 1.54, RD: 0.71 / 0.74 / 0.77, PMD: 0.86 / 0.87 / 0.97, RSH: 0.10 / 0.07 / 0.16.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD1FFC1FF5FAEABFB7545A4.taxon	discussion	Discussion. This species is characterised by its stout ventricose outline and dense colour pattern and cannot be confused with any other Paratethyan species. Sacco (1893 b) introduced this taxon with a question mark as variety of Conus tauroventricosus Sacco, 1893 from the Burdigalian of the Turin Hills. Hall (1966) treated this species and the varietal name magnolapugyensis as synonym of Lautoconus bitorosus (Fontannes, 1880). Conus tauroventricosus, which is most probably unrelated to L. bitorosus, differs from the Paratethyan species in its figshaped outline and more rounded shoulder. Lautoconus bitorosus is similar concerning its ventricose last whorl but has a higher spire and the last whorl is more cylindrical-conical (see Muñiz-Solís 1999). Paleoenvironment. The sandy sediment preserved in the shells suggests a shallow marine environment.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD1FFC1FF5FAEABFB7545A4.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Transylvanian Basin: Lăpugiu de Sus (Romania). The record from the Burdigalian of the Turin Hills mentioned by Sacco (1893 b) needs confirmation.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD0FFDDFF5FAD53FCFA4532.taxon	description	Lautoconus bitorosus (Fontannes, 1880) — Landau et al. 2013: 239, pl. 38, figs 2 – 4, pl. 41, fig. 9, pl. 42, fig. 3, pl. 81, fig. 3. non Conus (Chelyconus) miovoeslauensis Sacco, 1893 — Strausz 1966: 457, text-fig. 204.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD0FFDDFF5FAD53FCFA4532.taxon	materials_examined	Type material. Holotype NHMW 1849 / 0023 / 0003, Bad Vöslau (Austria), illustrated in Hoernes & Auinger (1879, pl. 6, fig. 6); middle Miocene, Badenian (Langhian). Studied material. Holotype and 10 spec. NHMW 1855 / 0045 / 0857, Bad Vöslau (Austria); 1 spec. NHMW 1937 / 0002 / 0272, Bad Vöslau (Austria), illustrated in Schaffer (1908, pl. 10, fig. 15); 4 spec. NHMW 2010 / 0004 / 1459 a-b Bad Vöslau (Austria); 7 spec. NHMW 1846 / 0037 / 0060, Steinebrunn (Austria); 17 spec. NHMW 1846 / 0037 / 0062, Steinebrunn (Austria); 7 spec. NHMW 1863 / 0015 / 0398, Forchtenau (Austria); 9 spec. NHMW 1870 / 0033 / 0003, Lăpugiu de Sus (Romania). Illustrated material. Figs 18 D 1 – D 3: holotype, Bad Vöslau (Austria): SL: 51.3 mm, MD: 34.5 mm, NHMW 1849 / 0023 / 0003, illustrated in Hoernes & Auinger (1879, pl. 6, fig. 6); Figs 18 E 1 – E 3: Bad Vöslau (Austria): SL: 65.7 mm, MD: 40.9 mm, NHMW 2010 / 0004 / 1459 a; Figs 18 F 1 – F 3: Bad Vöslau (Austria): SL: 50.3 mm, MD: 30.8 mm, NHMW 2010 / 0004 / 1459 b; Fig. 17 B: Bad Vöslau (Austria): SL: 42.6 mm, MD: 25.6 mm, NHMW 1855 / 0045 / 0855. Revised description. Medium-sized shells, spire coeloconoid, low to medium in height; very variable in height; spire whorls channelled, weakly striate to striate; more or less deep groove in mid-whorl. Subsutural flexure of medium depth, moderately curved, strongly asymmetrical; last whorl elongate and fig-shaped, rarely strongly ventricose. Shoulder high, rounded, sides convex, position of maximum diameter about ¼ of the way below the shoulder; very weakly constricted at base; bearing weak spiral grooves on the abapical third; aperture straight, widening and slightly flared abapically; siphonal canal short; siphonal fasciole weakly developed; colour pattern seen under UV light consisting of spirally arranged small irregular dashes, partly underlain by very thin spirals. Broad, wide-spaced flammulae on shoulder and last spire whorl.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD0FFDDFF5FAD53FCFA4532.taxon	description	Shell measurements and ratios. n = 17 adult specimens (only large morphs): largest specimen: SL: 65.7 mm, MD: 40.9 mm, mean SL: 51.2 mm (σ = 5.1), mean MD: 30.4 mm (σ = 3.6), spire angle: µ = 121.9 ° (σ = 8.6 °), last whorl angle: µ = 38.7 ° (σ = 1.5 °), LW: µ = 1.69 (σ = 0.08), RD: µ = 0.68 (σ = 0.02), PMD: µ = 0.86 (σ = 0.02), RSH: µ = 0.12 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD0FFDDFF5FAD53FCFA4532.taxon	discussion	Discussion. Sacco (1893 b) considered the Miocene specimen illustrated by Hörnes (1851, pl. 3, fig. 6) as Conus ventricosus to be conspecific with the Pliocene species Conus exventricosus Sacco, 1893. He referred to the specimen as a typical representative, which could be read as designation as type specimen. Unfortunately, this specimen is lost. Later, Hoernes & Auinger (1879, pl. 6, fig. 6) illustrated another specimen from Bad Vöslau as Conus ventricosus. Their selection was somewhat unfortunate as Hoernes & Auinger (1879) illustrated an atypical low spired and broad specimen. Therefore, Sacco (1893 b) introduced the varietal name miovoeslauensis for this specimen. The middle Miocene specimens from the Paratethys, like the Serravallian specimens from the Turkish Karman Basin described by Landau et al. (2013), differ from the Pliocene Lautoconus bitorosus in their lower spires and more elongate shapes (see also Landau et al. 2013). Therefore, it might be justified to separate the Miocene specimens as Lautoconus miovoeslauensis (Sacco, 1893). This name was overlooked by Sieber (1958 a) in his synthesis on Miocene gastropods of the Vienna Basin but was used by authors such as Strausz (1966) and Kovács & Vicián (2013). Unfortunately, the specimen illustrated by Strausz (1966, text-fig. 204) has a very broad last whorl and is not conspecific with L. miovoeslauensis. Similarly, the ovoid shells from Letkés in Hungary illustrated by Kovács & Vicián (2013) seem to represent a different species. The Italian specimen described by Sacco (1893 b) as Conus exventricosus differs from the Paratethyan species in its rounded shoulder, the lower position of the maximum diameter and the more pronounced constriction of the base. Hall (1966) treated Conus miovoeslauensis as conspecific with Conus magnolapugyensis and both as synonyms of Lautoconus bitorosus (Fontannes, 1880). Lautoconus miovoeslauensis differs from L. magnolapugyensis (Sacco, 1893) as redescribed herein in its smaller size, narrower and striate spire whorls, the distinct spiral threads on the base and the overall distinctly more slender outline. Paleoenvironment. The pelitic sediment infill suggests offshore environments at least for specimens from Bad Vöslau; other localities, such as Steinebrunn, represent nearshore environments.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFD0FFDDFF5FAD53FCFA4532.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Bad Vöslau, Baden, Steinebrunn (Austria); Eisenstadt-Sopron Basin: Mattersburg, Forchtenau (Austria); Alpine-Carpathian Foredeep: Grund (Austria) (Hoernes & Auinger 1879; Sieber 1949, 1956, 1958 b), Korytnica (Poland) (Friedberg 1911), Caransebeş-Mehadia Basin: Valea Bela Reca (Romania) (Hinculov 1968); Transylvanian Basin: Lăpugiu de Sus (Romania) (Hoernes & Auinger 1879). The record of an early Miocene occurrence from Mörtersdorf (Austria) of Lower Austria mentioned by Schaffer (1910) and Mandic & Steiniger (2003) is based on poorly preserved specimens, which are almost certainly misidentified; note that the specimen illustrated by Schaffer (1912, pl. 49, fig. 4) derives from Bad Vöslau and is of middle Miocene age. Proto-Mediterranean Sea and northeastern Atlantic. Serravallian (middle Miocene): Karaman Basin (Turkey) (Landau et al. 2013). See Landau et al. (2013) for a detailed list of early Miocene to early Pleistocene occurrences of the closely related L. bitorosus (Fontannes, 1880).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFCCFFDBFF5FACDAFA844237.taxon	description	Conus pelagicus nob. — Brocchi 1814: 289, pl. 2, fig. 9. [Chelyconus] [Conus] Mariae n. f. — Hoernes 1878 a: 195 (nomen nudum). C. [onus] Mariae — Hoernes 1878 b: 207 (nomen nudum). Conus (Chelyconus) Mariae nov. form. — Hoernes & Auinger 1879: 49, pl. 6, fig. 7.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFCCFFDBFF5FACDAFA844237.taxon	materials_examined	Type material. Lectotype (following ICZN Article 74.6) stored in the Museo Civico di Storia Naturale di Milano, Catalogue number 4668, Piacentino, Italy, Pliocene; illustrated in Pinna & Spezia (1978; pl. 18, fig. 4). The type locality of Conus mariae Hoernes & Auinger, 1879, which is considered to be a junior synonym of C. pelagicus, is the middle Miocene locality Bujtur in Romania. Studied material. 1 spec. NHMW 1861 / 0035 / 0004, Bujtur (Romania), illustrated in Hoernes & Auinger (1879, pl. 6, fig. 7); 2 spec. NHMW 1868 / 0001 / 0387, Lăpugiu de Sus (Romania). Illustrated material. Figs 19 A 1 – A 4, 17 J: Bujtur (Romania): SL: 53.6 mm, MD: 24.9 mm, NHMW 1861 / 0035 / 0 0 0 4, illustrated in Hoernes & Auinger (1879, pl. 6, fig. 7). Revised description. Medium-sized, biconical, solid, glossy shells with high conical spire; early spire whorls probably tuberculate; spire whorls flat to weakly convex, not striate; suture narrow but deeply impressed, almost canaliculated. Subsutural flexure shallow, moderately curved, moderately asymmetrical; last spire whorl weakly concave; indistinct angulation on shoulder; last whorl slightly ventricose, constricted at base. No spiral sculpture except for several spiral cords on base. Aperture moderately narrow. Siphonal fasciole strongly swollen, convex, twisted; inner lip narrow, straight; siphonal canal moderately long, reflected. Colour pattern under UV light consisting of flammulae on shoulder and spire and irregularly spaced short dashes on the entire last whorl, forming three broad bands separated by two fluorescing bands.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFCCFFDBFF5FACDAFA844237.taxon	description	Shell measurements and ratios. Three specimens are available: largest specimen SL: 57.6 mm, MD: 28.2 mm, spire angle: 82 / 85 / 83 °, last whorl angle: 33 / 35 / 32, LW: 2.04 / 2.09 / 2.15, RD: 0.59 / 0.59 / 0.56, PMD: 0.86 / 0.88 / 0.91, RSH: 0.17 / 0.18 / 0.17; spire angle: 95 °, last whorl angle: 37 °; second specimen: SL: 50.4 mm, MD: 25.2 mm, LW: 2.0, RD: 0.64, PMD: 0.85, RSH: 0.22.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFCCFFDBFF5FACDAFA844237.taxon	discussion	Discussion. Differs from the somewhat similar Conus s. l. posticestriatus (Kojumdgieva in Kojumdgieva & Strachimirov 1960) by the absence of any spiral grooves on the spire whorls, the constricted base and the reflected siphonal canal. Hall (1894) and Landau et al. (2013) synonymized Conus mariae Hoernes & Auinger, 1879 with Lautoconus pelagicus (Brocchi, 1814), which is followed herein.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFCCFFDBFF5FACDAFA844237.taxon	description	Paleoenvironment. The occurrences in the Turkish Karman Basin suggest shallow marine environments.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFCCFFDBFF5FACDAFA844237.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn (Austria), Mikulov- Kienberk, Hrušovany (Czech Republic) (Hoernes & Auinger 1879); Eisenstadt-Sopron Basin: Forchtenau (Austria) (Sieber 1956); Pannonian Basin: Letkés (Hungary) (Kovács & Vicián 2013); Transylvanian Basin: Lăpugiu de Sus, Bujtur (Romania) (Hoernes & Auinger 1879); Buzău Basin: Valea Muscel (Romania) (Ionesi & Nicorici 1994); Caransebeş-Mehadia Basin: Calvei (Romania) (Hinculov 1968); Dacian Basin: Opanec (Bulgaria) (Kojumdgieva & Strachimirov 1960). Proto-Mediterranean Sea and north eastern Atlantic. Burdigalian (early Miocene): Turin Hills (Italy); Langhian (middle Miocene): Aquitaine Basin (France); Serravallian (middle Miocene): Karaman Basin (Turkey); Tortonian (late Miocene): Po Basin (Italy); early Pliocene: western Mediterranean, Estepona Basin (Spain), central Mediterranean (Italy); late Pliocene: central Mediterranean (Italy) (see Landau et al. 2013 for details).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFCAFFD8FF5FABD3FD7740E2.taxon	materials_examined	Holotype: Figs 20 A 1 – A 3: SL: 25.2 mm, MD: 15.2 mm, NHMW 2016 / 0008 / 0001. Paratype: Figs 20 B 1 – B 3, 34 C: SL: 20.4 mm, MD: 11.8 mm, NHMW 2016 / 0008 / 0002.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFCAFFD8FF5FABD3FD7740E2.taxon	description	Paratype: Figs 20 C 1 – C 2: SL: 19.2 mm, MD: 10.7 mm, NHMW 2016 / 0008 / 0002. Paratype: Figs 20 D 1 – D 3: SL: 23.6 mm, MD: 13.1 mm, NHMW 2016 / 0008 / 0004.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFCAFFD8FF5FABD3FD7740E2.taxon	materials_examined	Additional material: 12 specimens, NHMW 2016 / 0008 / 0004, all specimens from Letkés, Hungary. Type stratum: fossil-rich marly sand with coral blocks and andesite boulders of the Sámsonháza Formation (Császár 1997). Type locality: Letkés at the western part of the Börzsöny Mts. (Hungary); see Kovács & Vicián (2013) for a map and description. Age: Middle Miocene, early Badenian (= Langhian).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFCAFFD8FF5FABD3FD7740E2.taxon	etymology	Etymology: Referring to Pest, the county of the type locality Letkés in Hungary.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFCAFFD8FF5FABD3FD7740E2.taxon	description	Description. Small stout-olivoid shells with low mammillate to weakly cyrtoconoid spire; spire whorls smooth, weakly convex; suture impressed. Last spire whorl relatively broad, passing via rounded shoulder in ventricose last whorl. Subsutural flexure shallow, weakly curved, strongly asymmetrical. Position of maximum diameter in posterior quarter of last whorl; faint constriction at base. Siphonal canal very short, slightly recurved; siphonal fasciole short, twisted, weakly swollen. About 5 deep spiral grooves on base delimitate broad flat spiral cords. Aperture narrow, strongly narrowing posteriorly, slightly widening towards siphonal canal. Very intense colour pattern in UV light consisting of broad spirals of densely spaced subquadratic to weakly rectangular spots; not axially arranged. Slightly axially elongated, prosocline dots on shoulder; additional dots and narrow flammulae on spire whorls. Shell measurements and ratios. n = 12: largest specimen: SL: 28.5 mm, MD: 16.4 mm, mean SL: 22.3 mm (σ = 2.8), mean MD: 12.9 mm (σ = 1.7), spire angle: µ = 116.6 ° (σ = 6.0 °), last whorl angle: µ = 37.3 ° (σ = 1.2 °), LW: µ = 1.7 (σ = 0.04), RD: µ = 0.67 (σ = 0.02), PMD: µ = 0.83 (σ = 0.02), RSH: µ = 0.13 (σ = 0.00).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFCAFFD8FF5FABD3FD7740E2.taxon	discussion	Discussion. Among the Paratethyan Conidae, only Lautoconus olivaeformis (Hoernes & Auinger, 1879) has a comparable olivoid outline but differs in its higher spire, narrower shape and the colour pattern of broad bands. The specimen from the Badenian of Bosnia and Herzegovina illustrated by Atanacković (1985, pl. 39, fig. 9) might represent this species as well, although it is more barrel-shaped. Paleoenvironment. The locality Letkés comprises a mixed assemblage of partly reworked taxa (Kovács & Vicián 2013); the frequent occurrence of corals suggests a shallow marine depositional environment.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFCAFFD8FF5FABD3FD7740E2.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Pannonian Basin: Letkés (Hungary);? southern Pannonian Basin: Hrvaćani (Bosnia and Herzegovina).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC9FFD9FF5FA9AAFB334076.taxon	description	Conus ponderosus nob. — Brocchi 1814: 293, pl. 3, fig. 1. Conus ponderosus Brocc. — Hörnes 1851: 26, pl. 2, figs 6 a – b. Conus (Rhizoconus) ponderosus Brocc. — Hoernes & Auinger 1879: 38 (partim, non figs.). Conus ponderosus Brocc. F. elmenus De Greg. — De Gregorio 1885: 369 [nov. nom. pro Conus ponderosus in Hörnes 1851, pl. 2, fig. 6].	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC9FFD9FF5FA9AAFB334076.taxon	materials_examined	Type material. Lectotype (following ICZN Article 74.6), Museo Civico di Storia Naturale di Milano, catalogue number 4674, illustrated in Brocchi (1814, p. 293, pl. 3, fig. 1) and Pinna & Spezia (pl. 18, figs 1 - 1 a), Parlascio, Colle, San Geminiano, Sogliano or Piacentino (Italy), Pliocene. The type locality of Conus elmenus De Gregorio, 1885, considered to be a junior synonym of C. ponderosus, is the middle Miocene locality Steinebrunn in Austria. Studied material. 1 spec. GBA 1856 / 004 / 0001 / 01, Steinebrunn (Austria), illustrated in Hörnes (1851, pl. 2, fig. 6, = holotype of Conus elmenus De Gregorio, 1885); 2 spec. NHMW 1846 / 0037 / 0064, Steinebrunn (Austria); 1 spec. NHMW 1860 / 0001 / 0051, Grund, (Austria). Illustrated material. Figs 19 B 1 – B 3: Steinebrunn (Austria), SL: 70.0 mm, MD: 36.3 mm: NHMW 1846 / 0037 / 0064; Figs 19 C 1 – C 3: Steinebrunn (Austria): SL: 88 mm, MD: 49 mm, GBA 1856 / 004 / 0001 / 01, illustrated in Hörnes (1851, pl. 2, fig. 6). Revised description. Moderately large shells with low-medium conical to slightly cyrtoconoid spire; incised suture; spire whorls nearly flat or weakly convex, adapically concave; with faint spiral threads. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Last spire whorl forming a nearly flat sutural ramp passing via a weakly angulated shoulder on the elongate conical last whorl; position of maximum diameter slightly below angulation. Base slightly constricted with weak spiral grooves; indistinct siphonal fasciole well demarcated from narrow inner lip. Siphonal canal moderately long, somewhat reflected. No colour pattern preserved.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC9FFD9FF5FA9AAFB334076.taxon	description	Shell measurements and ratios. 3 specimens: SL: 88 / 71.0 / 70.0 mm, MD: 49 / 41.5 / 36.3 mm, spire angle: 107 / 93 / 100 °, last whorl angle: 35 / 36 / 36 °, LW: 1.80 / 1.71 / 1.93, RD: 0.65 / 0.74 / 0.62, PMD: 0.90 / 0.91 / 0.91, RSH: 0.15 / 0.21 / 0.16.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC9FFD9FF5FA9AAFB334076.taxon	discussion	Discussion. This species is rather rare in the Paratethyan basins. Only few specimens from Steinebrunn (Austria) agree with Miocene and Pliocene specimens as described by Hall (1966) and Davoli (1972). Several additional specimens from Poland and Hungary described by Krach (1981) and Kovács & Vicián (2013) might need confirmation. Herein we follow Hall (1966) in treating the Paratethyan shells described by Hörnes (1851) as conspecific with Conus ponderosus. Consequently, we consider Conus elmenus, which was introduced by De Gregorio (1885) as new name for the specimen illustrated by Hörnes (1851, pl. 2, fig. 6), as subjective junior synonym of Conus ponderosus Brocchi, 1814. Nevertheless, the species concept of Hall (1966) was too wide and we reject his decision to treat Conus olivaeformis Hoernes & Auinger, 1879 and C. transsylvanicus Hoernes & Auinger, 1879 as synonyms of C. ponderosus (see discussions of respective species). Paleoenvironment. Shallow marine environments (e. g. Letkés, Kovács & Vicián 2013).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC9FFD9FF5FA9AAFB334076.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Steinebrunn (Austria) (Hörnes 1851); Pannonian Basin: Letkés (Kovács & Vicián 2013); Carpathian Foredeep: Korytnica, Węglinek, Łychów (Poland) (Bałuk 1997). Proto-Mediterranean Sea and north eastern Atlantic. Tortonian (late Miocene): Po Basin: Montegibbio (Italy) (Davoli 1972); Pliocene: Po Basin, Toscana (Italy) (Pinna & Spezia 1978; Chirli 1997).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC8FFD9FF5FA919FD764770.taxon	description	Conus (Rhizoconus) ponderosus grinzingensis Sacco — Csepreghy-Meznerics 1956: 436, pl. 10, figs 3 – 4 [non Conus grinzingensis Sacco, 1893 = Lautoconus steinabrunnensis (Sacco, 1893)]. Conus (Chelyconus) pseudoponderosus Dollfus & Dautzenberg (in Glibert, 1952) — Strausz 1966: 463, pl. 68, figs 6 – 7.? Conus (Chelyconus) cfr. ponderosus Brocchi — Strausz 1966: 463, pl. 69, fig. 11, pl. 70, fig. 1. Conus (Chelyconus) pseudoponderosus Glibert 1952 — Bohn-Havas 1973: 1122, pl. 7, fig. 4, pl. 9, fig. 12.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC8FFD9FF5FA919FD764770.taxon	description	Conus pseudoponderosus Glibert, 1952 — Vaessen 2010: 12, figs 10 C, 15 – 18 [cum syn.].	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC8FFD9FF5FA919FD764770.taxon	materials_examined	Type material. Holotype: Royal Belgian Institute of Natural Sciences, Nr. 10591, Cat. Types Invert. tert. I. R. Sc. N. B. nr. 2816, Manthelan, France; middle Miocene, Langhian; illustrated in Glibert (1952 a, pl. 13, fig. 4); paratypes: Royal Belgian Institute of Natural Sciences, Nr. 10591, Cat. Types Invert. tert. I. R. Sc. N. B. nr. 2817, Bossée, Manthelan, France; middle Miocene, Langhian. Revised description. Moderately large to large, solid shells with broad conical to weakly cyrtoconoid spire and broad conical last whorl. Spire whorls weakly convex with deep suture, not striate. Subsutural flexure asymmetrically curved, rather shallow. Last whorl with broadly rounded shoulder, rarely slightly angulate. Siphonal fasciole broad, weakly swollen; siphonal canal short, wide, straight. Colour pattern consisting of about 15 spirals of widely-spaced subquadratic and rectangular dots (see Bohn-Havas 1973, pl. 9, fig. 12).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC8FFD9FF5FA919FD764770.taxon	discussion	Discussion. This species was described by Glibert (1952 a) from the Langhian of the Loire Basin. Later Strausz (1966) and Bohn-Havas (1973) identified this species also in the coeval deposits of the Paratethys. The broad conical outline, rounded shoulder and slightly convex spire whorls allow a clear separation from Lautoconus ponderosus. Therefore, we consider it a distinct species as proposed also by Vaessen (2010). The internal cast illustrated by Strausz (1966, pl. 69, fig. 11, pl. 70, fig. 1) might also represent this species. In our opinion, the Paratethyan records of Lautoconus conoponderosus (Sacco, 1893) also represent L. pseudoponderosus (e. g. Kovács & Vicián 2013, fig. 113, which is a hardly distinguishable from the holotype of L. pseudoponderosus). Despite the similarity in shape, L. conoponderosus as defined by Davoli (1972) is smaller and more slender. Paleoenvironment. Shallow marine environments (e. g. Letkés, Kovács & Vicián 2013).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC8FFD9FF5FA919FD764770.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Pannonian Basin: Várpalota, Bánd, Diósd, Letkés, Szob, Budapest: Illés street, Kerepesi street (Hungary). Proto-Mediterranean Sea and northeastern Atlantic. Langhian: Loire Basin (Glibert 1952 a); Tortonian: Cacela Velha, Cacela Basin (Portugal) (Glibert 1952 a).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC8FFD7FF5FAE1BFB7F471D.taxon	materials_examined	Holotype: Figs 20 E 1 – E 3, 17 D: SL: 21.5 mm, MD: 12.9 mm, NHMW 2016 / 0009 / 0001. Paratype: Figs 20 F 1 – F 3: SL: 22.5 mm, MD: 13.2 mm, NHMW 2016 / 0009 / 0002. Paratype: Figs 20 G 1 – G 2: SL: 24.1 mm, MD: 14.1 mm, NHMW 2016 / 0009 / 0003. Paratype: Figs 20 I 1 – I 3: SL: 24.3 mm, MD: 15.5 mm, NHMW 2016 / 0009 / 0004 a. Paratype: Fig. 20 J: SL: 23.0 mm, MD: 13.4 mm, NHMW 2016 / 0009 / 0004 b. Additional material: 5 specimens, NHMW 2016 / 0009 / 0004, 2 spec. private collection Anton Breitenberger, Figs 20 H 1 – H 3: SL: 31.5 mm, MD: 19.4 mm. Type stratum: fossil-rich marly sand with coral blocks and andesite boulders of the Sámsonháza Formation (Császár 1997). Type locality: Letkés at the western part of the Börzsöny Mts. (Hungary); see Kovács & Vicián (2013) for a map and description. Age: Middle Miocene, early Badenian (= Langhian).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC8FFD7FF5FAE1BFB7F471D.taxon	etymology	Etymology: Referring to the zebra-like colour pattern of subadult specimens.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC8FFD7FF5FAE1BFB7F471D.taxon	description	Description. Small shells; early spire whorls pointed; later spire whorls nearly flat with moderately incised suture, forming low conical to slightly gradate spire; spire whorls narrowly coiled, last spire whorl relatively broader and faintly striate (only visible in grazing light). Subsutural flexure of medium depth, moderately curved, moderately asymmetrical. Narrowly rounded to weakly subangulate shoulder; position of maximum diameter slightly below shoulder. Last whorl stout and conical to weakly ventricose with faint constriction on base; weak spiral cords on lower third of last whorl. Aperture narrow, angulated at shoulder, nearly straight, only weakly widening towards short and straight siphonal canal; siphonal fasciole nearly absent. Colour pattern very intense under UV light, consisting of frequently interrupted broader fluorescing and narrower dark spirals crossed by broad axial bands, which may be nearly straight to strongly zig-zag shaped within the same specimen. This pattern is dominant in subadult specimens. Axial elements become subordinate in adult shells, which display mainly broad, spirally arranged light and dark dashes. Spire whorls with flammulae. Shell measurements and ratios. n = 12: largest specimen: SL: 31.5 mm, MD: 19.4 mm, mean SL: 24.2 mm (σ = 3.7), mean MD: 14.6 mm (σ = 2.4), spire angle: µ = 122.8 ° (σ = 5.6 °), last whorl angle: µ = 41.0 ° (σ = 1.5 °), LW: µ = 1.66 (σ = 0.05), RD: µ = 0.69 (σ = 0.02), PMD: µ = 0.84 (σ = 0.01), RSH: µ = 0.12 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC8FFD7FF5FAE1BFB7F471D.taxon	discussion	Discussion. This shell probably belongs to the same group as the West African genus Africonus Petuch, 1975 as defined by Tucker & Tenorio (2009). These are all small shelled species, with a short siphonal canal, shallow anal notch, weak siphonal fasciole and rather narrow aperture. Moreover, zig-zag colour patterns are frequently developed within Africonus (e. g. A. decoratus Röckel, Rolán & Monteiro, 1980, A. verdensis (Trovão, 1979), A. derrubado Rolán & Fernandes in Rolán, 1990). According to Tucker & Tenorio (2009), Africonus (type species: Conus cuneolus Reeve, 1843, by original designation, present day, West Africa) is characterised by having a paucispiral protoconch, the whorl tops may be concave when viewed in cross section, with cords on the whorl tops that may be lost in middle spire whorls or persist thereafter. The shell has a shallow to moderately deep subsutural flexure. The molecular phylogeny presented by Puillandre et al. (2014 a, b) does not support the separation of Africonus and Lautoconus Monterosato, 1923. Shells of the genus Lautoconus are characterised by their turgid shape with convex sides. Like Africonus, the protoconch is paucispiral. The whorl tops are ornamented with cords that reach the middle spire whorls and often persist and the subsutural flexure is shallow to moderate in depth (Tucker & Tenorio 2009). On shell characters, alone we cannot convincingly distinguish the two genera and therefore follow Puillandre et al. (2014 a) in considering Africonus a junior synonym of Lautoconus. Aside from its very narrow aperture, this cone snail has few shell features catching the eye at first sight. Therefore, it might have been overlooked so far. Its spectacular colouring under UV light, however, distinguishes it distinctly from all other Paratethyan species and allows an easy identification. Paleoenvironment. The locality Letkés comprises a mixed assemblage of partly reworked taxa (Kovács & Vicián 2013); the frequent occurrence of corals suggests a shallow marine depositional environment.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC8FFD7FF5FAE1BFB7F471D.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Pannonian Basin: Letkés (Hungary).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC6FFD5FF5FAEB6FDF24285.taxon	description	[Chelyconus] [Conus] rotundus n. f. — Hoernes 1878 a: 195 (nomen nudum). Conus (Chelyconus) rotundus nov. form. — Hoernes & Auinger 1879: 50, pl. 6, fig. 8. Conus (Chelyconus) rotundus Hoernes et Auinger — Csepreghy-Meznerics 1956: 419, pl. 10, figs 5 – 6.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC6FFD5FF5FAEB6FDF24285.taxon	materials_examined	Type material. Syntypes: 1 spec. NHMW 1858 / 0015 / 0072, Steinebrunn (Austria), illustrated in Hoernes & Auinger (1879, pl. 6, fig. 8), 1 spec. NHMW 1853 / 0003 / 0003, Gainfarn (Austria), 3 spec. NHMW 1867 / 0019 / 0 0 0 3, Coşteiu de Sus (Romania); middle Miocene, Badenian (late Langhian). Studied material. Syntypes. Illustrated material. Figs 17 E, 21 A 1 – A 3: syntype, Steinebrunn (Austria): SL: 42.2 mm, MD: 25.0 mm, NHMW 1858 / 0015 / 0072, illustrated in Hoernes & Auinger (1879, pl. 6, fig. 8). Revised description. Medium-sized shells with low conical to weakly cyrtoconoid spire and faintly ventricose last whorl. Early spire pointed, later whorls weakly convex with more or less prominent striae, which may grade into delicate cords on the last whorl. Most prominent spiral cord coincides with angulate shoulder. Deep suture; subsutural flexure of medium depth, strongly curved, moderately asymmetrical. Last whorl with spiral cords on lower third; weakly constricted at base with short canal; siphonal fasciole indistinct. No colour pattern preserved.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC6FFD5FF5FAEB6FDF24285.taxon	description	Shell measurements and ratios. Only two adult specimens are available: SL: 42.2 / 34.1 mm, MD: 25.0 / 20.5 mm, spire angle: 129 / 125 °, last whorl angle: 36 / 38 °, LW: 1.69 / 1.66, RD: 0.65 / 0.69, PMD: 0.90 / 0.90, RSH: 0.08 / 0.13.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC6FFD5FF5FAEB6FDF24285.taxon	discussion	Discussion. Hall (1966) discussed a close relationship between Lautoconus rotundus and L. bitorosus (Fontannes, 1880), which is larger and lacks a distinct angulation along the shoulder. Some of the specimens identified as Conus rotundus by Bałuk (1997, pl. 22, figs 7 – 8) differ in their elongate outline, the twisted siphonal fasciole and long siphonal canal and represent another species. Paleoenvironment. Probably shallow coastal environments based on its occurrence at Gainfarn and Steinebrunn (Zuschin et al. 2007).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC6FFD5FF5FAEB6FDF24285.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn, Steinebrunn (Austria) (Hoernes & Auinger 1879); Carpathian Foredeep: Korytnica (Poland) (Bałuk 1997); Pannonian Basin: Pécsszabolcs, Szob, Letkés (Hungary) (Kovács & Vicián 2013); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus (Romania) (Hoernes & Auinger 1879).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC4FFD2FF5FAC34FDF24783.taxon	description	non Conus (Rhizoconus) ponderosus steinabrunnensis — Csepreghy-Meznerics 1956: plate captions, pl. 11, figs 11 – 12. non Conus (Rhizoconus) ponderosus grinzingensis Sacco — Csepreghy-Meznerics 1956: 436, pl. 10, figs 1 – 2. non Conus (Chelyconus) ponderosus steinabrunnensis (Sacco, 1893) — Atanacković 1985: 177, pl. 39, figs 12 – 13. non Conus ponderosus steinabrunnensis (Sacco) — Ionesi & Nicorici 1994: 62, pl. 5, fig. 6. non Conus (Chelyconus) ponderosus var. steinabrunnensis (Sacco) — Kojumdgieva in Kojumdgieva & Strachimirov 1960: 213, pl. 50, fig. 8 [? Kalloconus ponderoaustriacus (Sacco, 1893)]. non Conus (Rhizoconus) steinabrunnensis (Sacco, 1893) — Bałuk 1997: 61, pl. 22, figs 1 – 3. non Chelyconus steinabrunnensis (Sacco, 1893) — Kovács & Vicián 2013: 61, figs 22 – 27.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC4FFD2FF5FAC34FDF24783.taxon	materials_examined	Type material. Holotype NHMW 1855 / 0045 / 0005, Steinebrunn (Austria), illustrated in Hoernes & Auinger (1879, pl. 5, fig. 4), middle Miocene (Badenian). The type locality of Conus grinzingensis Sacco, 1893, considered to be a junior synonym of L. steinabrunnensis, is the middle Miocene (Badenian) locality Grinzing in Austria. Studied material. Holotype and 3 spec. NHMW 1855 / 0045 / 0005, Steinebrunn (Austria); 3 spec. NHMW 1857 / 0014 / 0007, Steinebrunn (Austria); 7 spec. NHMW 1846 / 0037 / 0049, Gainfarn (Austria); 1 spec. NHMW 1855 / 0045 / 0356, Gainfarn (Austria); 4 spec. NHMW 1847 / 0046 / 0003, Szob (Hungary); 8 spec. NHMW 1865 / 0001 / 0764, Grinzing (Austria), including specimen illustrated in Hoernes & Auinger (1879, pl. 5, fig. 5) (= holotype of Conus grinzingensis Sacco, 1893); 2 spec. NHMW 1846 / 0037 / 0045, Grinzing (Austria); 2 spec. NHMW 1854 / 0035 / 0037, Lăpugiu de Sus (Romania); 3 spec. NHMW 1870 / 0054 / 0038, Enzesfeld (Austria); 2 spec. NHMW 1860 / 0001 / 0067, Mikulov-Kienberk (Czech Republic); middle Miocene, Badenian (Langhian, Serravallian?). Illustrated material. Figs 21 B 1 – B 3: Grinzing (Austria): SL: 48.8 mm, MD: 26.4 mm, NHMW 1865 / 0001 / 0 764, specimen illustrated in Hoernes & Auinger (1879, pl. 5, fig. 5) (= holotype of Conus grinzingensis Sacco, 1893); Figs 21 C 1 – C 3: holotype, Steinebrunn (Austria): SL: 54.6 mm, MD: 30.2 mm, NHMW 1855 / 0045 / 0005; Figs 21 D 1 – D 3, 17 H: Lăpugiu de Sus (Romania): SL: 54.2 mm, MD: 28.3 mm, NHMW 1854 / 0035 / 0037. Revised description. Medium-sized shells with medium-low conical to slightly coeloconoid spire; spire whorls flat, weakly striate, slowly increasing in width. Last spire whorl often smooth; suture channelled. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Last whorl elongate conical, not constricted, usually distinctly angulated, with weak convexity below shoulder, coinciding with position of maximum diameter. Weak spiral grooves may be present on lower third of last whorl. Siphonal canal moderately long, straight or weakly recurved. Aperture moderately narrow, anteriorly widening; siphonal fasciole indistinct, weakly swollen, twisted, well demarcated from narrow inner lip. Colour pattern under UV light consists of large and irregular blotches on the last whorl and flammulae on spire whorls.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC4FFD2FF5FAC34FDF24783.taxon	description	Shell measurements and ratios. n = 24 adult specimens: largest specimen: SL: 61.5 mm, MD: 32.5 mm, mean SL: 51.8 mm (σ = 5.5), mean MD: 28.2 mm (σ = 2.8), spire angle: µ = 101.1 ° (σ = 8.6 °), last whorl angle: µ = 35.3 ° (σ = 2.1 °), LW: µ = 1.8 (σ = 0.08), RD: µ = 0.65 (σ = 0.03), PMD: µ = 0.88 (σ = 0.03), RSH: µ = 0.17 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC4FFD2FF5FAC34FDF24783.taxon	discussion	Discussion. Hoernes & Auinger (18979) separated several shells from the Vienna Basin as variety II based on the slightly more elongate last whorl and the less pronounced angulation. Later, Sacco (1893 b) introduced the variety name grinzingensis for these shells, based on the illustration in Hoernes & Auinger (1879, pl. 5, fig. 5). In the lots assigned to variety II by Hoernes & Auinger (1879), there is only a single pathologic specimen with a more elongate last whorl, whereas all others do not differ at all from the specimens assigned by them to variety I. Consequently, the morphometric analysis of the measurements and shell ratios as given in Table 1 did not support any separation. Therefore, we consider Conus grinzingensis Sacco, 1893 as a subjective junior synonym of Lautoconus steinabrunnensis. This species was established by Sacco (1893 b) as variety of his Conus conoponderosus from the Tortonian of Italy. The Paratethyan species differs from the rather stout Lautoconus conoponderosus (Sacco, 1893) in the larger size, more slender outline and the distinct angulation at the shoulder (see syntype of L. conoponderosus in Ferrero- Mortara et al. 1984, pl. 18, fig. 10).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC4FFD2FF5FAC34FDF24783.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Steinebrunn, Enzesfeld, Gainfarn, Grinzing (Austria), Mikulov-Kienberk (Czech Republic) (Hoernes & Auinger 1879; Sieber 1958 b); Eisenstadt- Sopron Basin: Forchtenau (Austria) (Sieber 1958 b); Carpathian Foredeep: Bořitov (Czech Republic) (Hoernes & Auinger 1879), Pannonian Basin: Szob (Hungary) (Hoernes & Auinger 1879); Transylvanian Basin: Lăpugiu de Sus (Romania) (Hoernes & Auinger 1879).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC3FFD0FF5FAF42FA0840E2.taxon	materials_examined	Type material. Lectotype NHMW 1949 / 0005 / 0004, Bad Vöslau (Austria); illustrated in Hoernes & Auinger (1879, pl. 3, fig. 3) (designated herein); middle Miocene, Badenian (late Langhian). Studied material. 1 spec. NHMW 1949 / 0005 / 0004, Bad Vöslau (Austria); illustrated in Hoernes & Auinger (1879, pl. 3, fig. 3); 15 spec. NHMW 1855 / 0045 / 0858, Bad Vöslau (Austria); 5 spec. NHMW 1997 z 0178 / 1179, Bad Vöslau (Austria); 3 spec. NHMW A 1631, Lăpugiu de Sus (Romania); 4 spec. NHMW 1856 / 0004 / 0413, Lăpugiu de Sus (Romania). Illustrated material. Figs 21 E 1 – E 3: Bad Vöslau (Austria), SL: 51.2 mm, MD: 32.1 mm, NHMW 1949 / 0005 / 0 0 0 4, illustrated in Hoernes & Auinger (1879, pl. 3, fig. 3); Figs 21 F 1 – F 3: Bad Vöslau (Austria), SL: 47.0 mm, MD: 30.5 mm, NHMW 1997 z 0178 / 1179; Fig. 17 F: Bad Vöslau (Austria), SL: 41.0 mm, MD: 25.2 mm, NHMW 1855 / 0045 / 0858. Revised description. Medium-sized, club-shaped solid shells with low conical spire. Pointed early spire; later spire whorls weakly convex, faintly striate; deep, narrowly canaliculated suture. Subsutural flexure shallow, weakly curved, strongly asymmetrical. Last spire whorl rapidly broadening; position of maximum diameter coinciding with periphery of prominent subangular to rounded shoulder; last whorl rapidly contracting, straightsided except for a very faint convexity mid-whorl. Prominent, strongly twisted siphonal fasciole; siphonal canal moderately wide, long, strongly recurved. Surface glossy with delicate growth lines and rather weak spiral grooves on lower third of last whorl. Colour pattern very weak (or weakly preserved) consisting of numerous thin spirals, which are continuous or composed of nearly continuous spirals of long dashes.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC3FFD0FF5FAF42FA0840E2.taxon	description	Shell ratios and measurement. n = 10 adult specimens: largest specimen: SL: 51.2 mm, MD: 33.4 mm, mean SL: 46.4 mm (σ = 3.7), mean MD: 29.4 mm (σ = 2.5), spire angle: µ = 116.9 ° (σ = 6.5 °), last whorl angle: µ = 37.5 ° (σ = 1.0 °), LW: µ = 1.58 (σ = 0.04), RD: µ = 0.72 (σ = 0.03), PMD: µ = 0.89 (σ = 0.03), RSH: µ = 0.12 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC3FFD0FF5FAF42FA0840E2.taxon	discussion	Discussion. Hoernes & Auinger (1879) described this species as Conus hochstetteri in the text of their monograph. This name was already preoccupied by Conus hochstetteri Martin, 1879, a Miocene species from Indonesia. The monograph of Martin was already available to R. Hoernes before publication of the monograph of Hoernes & Auinger (1879) but it was too late for changes in type setting. When recognizing the homonymy Hoernes (1879) introduced Conus steindachneri as new name for the Viennese species. This name was subsequently also added in the plate captions of Hoernes & Auinger (1879). Nevertheless, the authorship has to be passed to Hoernes (1879) and not to Hoernes & Auinger (1879) as frequently done in the literature. When introducing this taxon Hoernes & Auinger (1879) referred to early Miocene specimens from Niederkreuzstetten (Austria) and to middle Miocene specimens from Bad Vöslau (Austria), thus mixing two species. The early Miocene specimens, including the specimen illustrated by Hörnes (1851, pl. 3, fig. 8), represent Kalloconus ponderovagus (Sacco, 1893) (see discussion of K. ponderovagus). The description of Conus steindachneri by Hoernes & Auinger (1879) is mainly based on the middle Miocene specimens but the early Miocene specimens are syntypes as well. To settle this problem, we designate the specimen from Bad Vöslau, illustrated by Hoernes & Auinger (1879, pl. 3, fig. 3) as lectotype to clarify the status of this species.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC3FFD0FF5FAF42FA0840E2.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Alpine-Carpathian Foredeep: Grund (Austria); Vienna Basin: Bad Vöslau (Austria), Mikulov-Kienberk (Czech Republic) (Hoernes & Auinger 1879); Pannonian Basin: Szob, Letkés (Hungary) (Strausz 1966; Kovács & Vicián 2013); Dacian Basin: Târnene, Orehovica (Bulgaria) (Kojumdgieva & Strachimirov 1960); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus (Romania) (Hoernes & Auinger 1879; Boettger 1906); Krka Basin: Golobinjek pri Šentjerneju (Slovenia) (Mikuž 2009). The occurrence of this species in the Tortonian of Barcelona, mentioned by Faura I Sans (1908), needs confirmation.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC1FFD1FF5FA9AAFBA4430A.taxon	description	Conus (Chelyconus) Enzesfeldensis nov. form. — Hoernes & Auinger 1879: 46 [nov. nom. pro Conus raristriatus in Hörnes 1851, pl. 3, fig. 2].? Conus (Dendroconus) subraristriatus Da Costa — Csepreghy-Meznerics 1956: 422, pl. 11, figs 1 – 2. Conus (Chelyconus) enzesfeldensis Hoernes et Auinger — Csepreghy-Meznerics 1956: 420, pl. 3, fig. 9.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC1FFD1FF5FA9AAFBA4430A.taxon	materials_examined	Type material. Lectotype designated by Landau et al. (2013): specimen illustrated in Pereira da Costa (1866, pl. 4, fig. 7); Cacela Basin (Portugal); late Miocene, Tortonian. The type locality of Conus enzesfeldensis Hoernes & Auinger, 1879, considered to be a junior synonym of L. subraristriatus, is the middle Miocene (Badenian) locality Enzesfeld in Austria. Studied material. 4 spec NHMW 1853 / 0010 / 0004, 4 spec. NHMW 1853 / 0010 / 0006, 6 spec. NHMW A 1610, including specimen illustrated in Schultz (1998, pl. 29, fig. 8.), all Enzesfeld (Austria), 1 spec NHMW 1857 / 0014 / 0 0 0 5, Steinebrunn (Austria), 2 spec. NHMW A 992, Grund (Austria). Illustrated material. Figs 21 G 1 – G 3: Enzesfeld (Austria): SL: 57.9 mm, MD: 29.8 mm, NHMW 1853 / 0010 / 0006 a; Figs. 21 H: Enzesfeld (Austria), SL: 52.6 mm, MD: 27.9 mm, NHMW A 1610; Fig. 17 I: Enzesfeld (Austria): SL: 48.7 mm, MD: 25.3 mm, NHMW 1853 / 0010 / 0006 b. Revised description. Medium-sized, solid shells; spire of medium height, cyrtoconoid; spire whorls very weakly convex, smooth; subsutural flexure shallow, weakly curved, moderately asymmetrical; last whorl elongate, roundly shouldered, straight-sided, position of maximum diameter at shoulder; hardly constricted at base; bearing few weak, finely beaded spiral cords at the anterior end in some specimens; aperture straight, relatively narrow, widening slightly abapically; siphonal canal short, straight; siphonal fasciole rounded, relatively well-developed; colour pattern under UV light of widely spaced, narrow brownish dots and dashes, coalescing into straight spiral lines forming three continuous fluorescing bands, separating two narrow regions lacking pigmentation.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC1FFD1FF5FA9AAFBA4430A.taxon	description	Shell measurements and ratios. n = 13: largest specimen: SL: 80.4 mm, MD: 39.4 mm, mean SL: 54.3 mm (σ = 9.2), mean MD: 28.1 mm (σ = 4.2), spire angle: µ = 92.6 ° (σ = 2.9 °), last whorl angle: µ = 34.6 ° (σ = 1.0 °), LW: µ = 1.9 (σ = 0.06), RD: µ = 0.63 (σ = 0.02), PMD: µ = 0.84 (σ = 0.03), RSH: µ = 0.18 (σ = 0.02).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC1FFD1FF5FA9AAFBA4430A.taxon	discussion	Discussion. The status of this species was discussed in detail by Landau et al. (2013), who designated the specimen illustrated by Pereira da Costa (1866, pl. 4, fig. 7) as lectotype of L. subraristriatus, which is conspecific with Conus enzesfeldensis Hoernes & Auinger, 1879. Paleoenvironment. Typically found in shallow marine, sandy nearshore environments.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFC1FFD1FF5FA9AAFBA4430A.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn, Enzesfeld, Bad Vöslau, Steinebrunn, Pötzleinsdorf (Austria) (Hoernes & Auinger 1879; Sieber 1958 b); Alpine-Carpathian Foredeep: Grund (Austria) (Hoernes & Auinger 1879),? Bükk Mountains: Borsodbóta (Hungary) (Csepreghy-Meznerics 1972); Pannonian Basin: Szob, Letkés (Hungary) (Kovács & Vicián 2013); southern Pannonian Basin: Miljevići (Bosnia and Herzegovina) (Atanacković 1985); Transylvanian Basin: Nemeşeşti, Lăpugiu de Sus, Coşteiu de Sus, Bujtur (Romania) (Hoernes & Auinger 1879);? Buzău Basin: Valea Muscel (Romania) (Ionesi & Nicorici 1994). Proto-Mediterranean Sea and northeastern Atlantic. Serravallian: Karman Basin (Turkey) (Landau e t al. 2013); Tortonian: Cacela Velha, Cacela Basin (Portugal) (Pereira da Costa, 1866).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFFFFEEFF5FA8CAFB7945D4.taxon	materials_examined	Illustrated material. Figs 22 A 1 – A 3, 17 G: Letkés (Hungary): SL: 55.3 mm, MD: 31.9 mm, private collection Anton Breitenberger (Bad Vöslau, Austria).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFFFFEEFF5FA8CAFB7945D4.taxon	description	Description. Medium-sized shells of subcylindrical outline. Spire very low; suture impressed. Early spire whorls smooth, weakly convex; last whorl smooth with faint central concavity. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Distinctly angulated shoulder, position of maximum diameter somewhat below shoulder. Last whorl slightly ventricose, subcylindrical; weakly constricted at base. Surface smooth except for prominent spiral cords on base. Siphonal fasciole weakly swollen, twisted, separated from short inner lip by slight notch. Siphonal canal short, broad, slightly recurved. Aperture moderately narrow, widening anteriorly. Shell measurements and ratios. SL: 55.3 mm, MD: 31.9 mm, spire angle: 130 °, last whorl angle: 37 °, LW: 1.73, RD: 0.65, PMD: 81, RSH: 0.11.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFFFFEEFF5FA8CAFB7945D4.taxon	discussion	Discussion. Only a single specimen is available to us. However, a second specimen from the same locality, stored in the Hungarian Natural History Museum (HNHM 2013.213), was illustrated by Kovács & Vicián (2013). Both specimens are identical in outline and therefore, it is unlikely that they represent aberrant specimens. This species is reminiscent of Lautoconus bitorosus miovoeslauensis (Sacco, 1893) but differs in its subcylindrical outline, the smooth spire and the pronounced shoulder. Moreover, both species seem also to be ecologically separated. Lautoconus b. miovoeslauensis is found in offshore clays whereas the occurrence at Letkés indicates shallow marine environments with corals. Despite the numerous “ varieties ” described by Sacco (1893 a, b), we are not aware of any comparably cylindrical morphotype. Nevertheless, we refrain from formally introducing a new species name without having seen more material. Paleoenvironment. The locality Letkés comprises a mixed assemblage of partly reworked taxa (Kovács & Vicián 2013); the frequent occurrence of corals suggests a shallow marine depositional environment.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFFFFEEFF5FA8CAFB7945D4.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Pannonian Basin: Letkés (Hungary).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFFFFEFFF5FAD67FB6D409F.taxon	materials_examined	Type species (by original designation): Conus glans Hwass in Bruguière, 1792. Recent, Indian Ocean. Note. According to Tucker & Tenorio (2009) the genus Leporiconus Iredale, 1930 is characterised by subcylindrical to pyriform, sometimes torpedo shaped shells, with beaded early spire whorls, two or more cords on the spire whorls and ridges on the body whorl, which are usually well developed and pustulose. The shoulder is rounded to indistinct. The subsutural flexures of the Paratethyan species are shallow, weakly to moderately curved and moderately asymmetrical and the protoconch is multispiral. The generic description given by Tucker & Tenorio (2009) was based on the species considered by these authors to be included in the genus: C. glans Hwass in Bruguière, 1792, C. caffeae Gmelin, 1791, C. corallinus Kiener, 1845, C. cylindraceus Broderip & Sowerby I, 1830, C. granum Röckel & Fischöder, 1985, C. luteus Sowerby I, 1833, C. mitratus Hwass in Bruguière, 1792, C. nucleus Reeve, 1848 and C. tenuistriatus Sowerby II, 1858. However, Puillandre et al. (2014 b) defined Leporiconus as a monophyletic clade containing five of the species included by Tucker & Tenorio: C. coffeae, C. glans, C. granum, C. luteus, and C. tenuistriatus, but excluded C. nucleus and C. corallinus, which they placed in the closely related genus Splinoconu s da Motta, 1991. Conus cylindraceus and C. mitratus they placed in the genus Turriconus Shikama & Habe, 1968, which is not closely related to Leporiconus. Therefore, the generic description should be modified to include only the species shown to be monophyletic by Puillandre et al. (2014 b). The wording given above still applies to the genus, as recognised by Puillandre et al. (2014 b), but torpedo-shaped shells should be removed, as the two species now included in Turriconus are more slender torpedo shaped, with far higher spires than species in Leporiconus. We cannot see any reliable shell characters by which to separate the present-day species of Leporiconus and Splinoconus, except that the spiral sculpture seems to be less prominent in Splinoconus. This not surprising as the two genera are closely related (Puillandre et al. 2014 b; fig 2). We have attributed to this genus a small number of Paratethyan species of which one, Leporiconus transsylvanicus (Hoernes & Auinger, 1879) is remarkably similar to the type species L. glans (Hwass in Bruguière, 1792). The other species are less obviously torpedo-shaped, but all have beaded early spire whorls. Conus transsylvanicus and C. suessi Hoernes & Auinger, 1879 were placed in the genus Varioconus da Motta 1991 in a unpublished generic list made available to us by J. K. Tucker (personal comm. 2016), however, Varioconus (considered as synonym of Lautoconus by Puillandre et al. 2014 a) lacks beads on spire whorls. The presence of Leporiconus in the Miocene Paratethys is interesting, as it documents the wide post-Tethyan distribution of a taxon during middle Miocene times nowadays restricted to the Indo-West Pacific.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFEFFEFFF5FAA31FB7F468B.taxon	materials_examined	Holotype: Figs 23 A 1 – A 3, 17 K: SL: 34.6 mm, MD: 17.8 mm, NHMW 2016 / 0005 / 0001. Paratype: Figs 23 B 1 – B 3: SL: 32.6 mm, MD: 16.3 mm, NHMW 2016 / 0005 / 0002.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFEFFEFFF5FAA31FB7F468B.taxon	description	Paratype: Figs 23 C 1 – C 2: SL: 30.8 mm, MD: 16.9 mm, NHMW 2016 / 0005 / 0003 a.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFEFFEFFF5FAA31FB7F468B.taxon	materials_examined	Additional material: 1 specimen NHMW 2016 / 0005 / 0003, all specimens from Letkés, Hungary. Type stratum: fossil-rich marly sand with coral blocks and andesite boulders of the Sámsonháza Formation (Császár 1997). Type locality: Letkés at the western part of the Börzsöny Mts. (Hungary); see Kovács & Vicián (2013) for a map and description. Age: Middle Miocene, early Badenian (= Langhian).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFEFFEFFF5FAA31FB7F468B.taxon	etymology	Etymology: Referring to the Paratethys Sea.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFEFFEFFF5FAA31FB7F468B.taxon	description	Description. Moderately small biconical to stout torpedo-shaped shell with moderately high conical to weakly cyrtoconoid spire and conical last whorl. Early spire whorls gradate, tuberculate along lower suture, striate; later spire whorls high, moderately convex, distinctly striate; suture impressed, narrowly canaliculated, undulating. Subsutural flexure shallow, weakly curved, moderately asymmetrical. Last whorl conical, squat, with broadly rounded, sometimes weakly angulated rounded shoulder; not constricted at base. Narrow aperture strongly narrowing at posterior canal, weakly broadening towards short, straight siphonal canal. Siphonal fasciole indistinct. Sculpture of last whorl consisting of densely spaced spirals of tiny beads, which are also roughly axially arranged. Lower half of last whorl with prominent, beaded spiral cords. Colour pattern under UV light consisting of narrow, axially elongate flammulae on spire whorls and very dense, regular pattern of spirally arranged tiny dots, coinciding with beads on last whorl. Shell measurements and ratios. Only 4 specimens are available, of these three are adult: SL: 34.6 / 32.6 / 30.8 mm, MD: 17,8 / 16.3 / 16.9 mm, spire angle: 85 / 83 / 98 °, last whorl angle: 39 / 37 / 36 °, LW: 1.94 / 2.00 / 1.82, RD: 0.70 / 0.68 / 0.66, PMD: 0.86 / 0.83 / 0.79, RSH: 0.26 / 0.26 / 0.17.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFEFFEFFF5FAA31FB7F468B.taxon	discussion	Discussion. We place this species in Leporiconus based on the beaded spiral cords, the tuberculate early spire whorls and the striate spire whorls. The tiny dotted colour pattern coinciding with the beads is also seen in the type species L. glans. This is a very peculiar species, which is characterised by its stout torpedo-shaped outline and the densely beaded sculpture. In their paper on the Conidae from Letkés, Kovács & Vicián (2013) might have partly referred to this species as Lautoconus pelagicus (Brocchi, 1814) as they mention beads and small tubercles in their description. Lautoconus pelagicus differs in its larger size, smooth shell surface, slightly recurved siphonal canal and completely different colour pattern (se Pinna & Spezia 1978; Landau et. al. 2013). Leporiconus transsylvanicus (Hoernes & Auinger, 1879) is congeneric, but differs in its more slender torpedo-like shell with a longer last whorl, higher position of the shoulder, lower spire and less prominent sculpture. Paleoenvironment. The locality Letkés comprises a mixed assemblage of partly reworked taxa (Kovács & Vicián 2013); the frequent occurrence of corals suggests a shallow marine depositional environment.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFEFFEFFF5FAA31FB7F468B.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Pannonian Basin: Letkés (Hungary).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFCFFEAFF5FA8E1FEFF40C7.taxon	description	[Chelyconus] [Conus] Suessi n. f. — Hoernes 1878 a: 195 (nomen nudum). C. [onus] Suessi — Hoernes 1878 b: 207 (nomen nudum).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFCFFEAFF5FA8E1FEFF40C7.taxon	materials_examined	Type material. The syntype illustrated in Hoernes & Auinger (1879, pl. 1, fig. 1) was stored in the collections of the Geological Survey of Austria (GBA), but seems to be lost. Other syntypes: NHMW 1858 / 0043 / 0001 a, illustrated in Hoernes & Auinger (1879, pl. 6, fig. 1), syntype NHMW 1854 / 0035 / 0033, illustrated in Hoernes & Auinger (1879, pl. 6, fig. 2), syntype NHMW 1858 / 0043 / 0001 b, illustrated in Hoernes & Auinger (1879, pl. 1, fig. 15), 2 syntypes NHMW 1858 / 0043 / 0001 a, 4 syntypes NHMW A 457, 5 syntypes NHMW 1868 / 0001 / 0386, all Lăpugiu de Sus (Romania); middle Miocene, Badenian (Langhian). Studied material. Syntypes and 6 spec. NHMW 1865 / 0001 / 0158, Lăpugiu de Sus (Romania). Illustrated material. Figs 23 D 1 – D 2: syntype, SL: 44.1 mm, MD: 19.8 mm, NHMW 1858 / 0043 / 0001 b, illustrated in Hoernes & Auinger (1879, pl. 1, fig. 15); Figs 23 E 1 – E 2: syntype, SL: 64.6 mm, MD: 29.0 mm, NHMW A 457; Figs 23 F 1 – F 3: syntype, SL: 63.6 mm, MD: 29.4 mm, NHMW 1858 / 0043 / 0001 a, illustrated in Hoernes & Auinger (1879, pl. 6, fig. 1); Fig. 23 H: SL: 55.6, MD: 23.4 mm, NHMW 1865 / 0001 / 0158; Figs 23 G 1 – G 2: SL: 72.3, MD: 32.3 mm, NHMW 1865 / 0001 / 0158; Fig. 17 L: syntype, SL: 51.2 mm, MD: 23.3 mm, NHMW A 457 a; all Lăpugiu de Sus (Romania). Revised description. Moderately large, not very solid, biconical shells; straight-sided conical spire; early spire whorls tuberculate; later spire whorls flat, striate; suture moderately incised, undulating; last spire whorl forming flat to weakly concave sutural ramp passing via distinct, slightly angulated shoulder into elongate, feebly convex last whorl. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Aperture narrow with straight outer lip, slightly widening towards short, straight, weakly recurved siphonal canal; fasciole indistinct, slightly swollen. Sculpture consisting of delicate, finely pustulose spiral threads on the entire whorl in subadult shells; beads are successively reduced during ontogeny; fully-grown specimens develop only weak spiral cords on base. Colour pattern under UV light consisting of densely spaced thin, continuous spiral lines and broad flammulae on spire whorls, becoming axially elongate opisthocline on shoulder. Last whorl with moderately broad discontinuous bands with axial streaks aligned with the axis of coiling.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFCFFEAFF5FA8E1FEFF40C7.taxon	description	Shell measurements and ratios. n = 7: largest specimen: SL: 68.9 mm, MD: 30.5 mm, mean SL: 56.8 mm (σ = 9.7), mean MD: 25.7 mm (σ = 4.1), spire angle: µ = 88.6 ° (σ = 6.0 °), last whorl angle: µ = 31.3 ° (σ = 1.8 °), LW: µ = 2.2 (σ = 0.07), RD: µ = 0.54 (σ = 0.01), PMD: µ = 0.89 (σ = 0.02), RSH: µ = 0.16 (σ = 0.02).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFCFFEAFF5FA8E1FEFF40C7.taxon	discussion	Discussion. We place this species in Leporiconus based on the pustulose spiral cords, the tuberculate early spire whorls and the striate spire whorls, which contradict a placement in Varioconus (= Lautoconus) as proposed by Kovács & Vicián (2013). A modern look-alike of Leporiconus suessi is the Indo-West Pacific Leporiconus corallinus (Kiener, 1847). This species was synonymized with Lautoconus pelagicus (Brocchi, 1814) by Kovács & Vicián (2013), but the Miocene to Pliocene L. pelagicus, as described by Hall (1966), Pinna & Spezia (1978), Muñiz Solís (1999) and Landau et al. (2013), differs in its larger size, the more convex spire whorls, the broader last whorl, which lacks the slight angulation and tends to be slightly ventricose, and its colour pattern of spirally arranged blotches (see Landau et al. 2013).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFCFFEAFF5FA8E1FEFF40C7.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Eisenstadt-Sopron Basin: Forchtenau (Hoernes & Auinger 1879); Pannonian Basin: Borsodbóta, Diósd, Letkés, Szob, Budapest: Illés street, Örs vezér square (Hungary) (Kovács & Vicián 2013); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus (Romania) (Hoernes & Auinger 1879). The occurrence of this species in the Tortonian of Barcelona, mentioned by Faura I Sans (1908), needs confirmation.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFBFFEAFF5FA989FE444791.taxon	materials_examined	Type material. Syntype NHMW 1854 / 0035 / 0034 a, Lăpugiu de Sus (Romania), illustrated in Hoernes & Auinger (1879, pl. 1, fig. 14); 4 syntypes NHMW 1854 / 0035 / 0034, Lăpugiu de Sus (Romania); middle Miocene, Badenian (Langhian). Studied material. Syntypes. Illustrated material. Figs 23 I 1 – I 5, 17 M: syntype, Lăpugiu de Sus (Romania): SL: 49.8 mm, MD: 21.5 mm, NHMW 1854 / 0035 / 0034 a, illustrated in Hoernes & Auinger (1879, pl. 1, fig. 14); Figs 23 J 1 – J 3: syntype, Lăpugiu de Sus (Romania): SL: 52.5 mm, MD: 21.2 mm, NHMW 1854 / 0035 / 0034. Revised description. Medium-sized torpedo-shaped shells; broad mammillate spire with pointed apex and strongly tuberculate above suture on early whorls; later spire whorls nearly flat, striate; suture impressed; subsutural flexure shallow, weakly curved, moderately asymmetrical. Indistinct shoulder coinciding with position of maximum diameter. Last whorl narrowly elongate, faintly convex, weakly constricted close to moderately long, slightly recurved siphonal canal. Aperture narrow, only slightly widening towards siphonal canal. Siphonal fasciole indistinct, twisted, poorly demarcated from thin, narrow inner lip. Sculpture of last whorl consisting of pustulose cords on lower half of whorl, becoming very weak and smooth in upper part. Colour pattern under UV light consisting of weak flammulae on shoulder and narrow spirals of long dashes and speckles forming nearly continuous spiral lines on last whorl.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFBFFEAFF5FA989FE444791.taxon	description	Shell measurements and ratios. n = 5: largest specimen: SL: 52.5 mm, MD: 21.2 mm, mean SL: 47.1 mm (σ = 5.8), mean MD: 19.9 mm (σ = 2.1), spire angle: µ = 84.6 ° (σ = 4.5 °), last whorl angle: µ = 29.2 ° (σ = 1.5 °), LW: µ = 2.4 (σ = 0.08), RD: µ = 0.51 (σ = 0.02), PMD: µ = 0.87 (σ = 0.04), RSH: µ = 0.17 (σ = 0.02).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFBFFEAFF5FA989FE444791.taxon	discussion	Discussion. This rare species agrees in all aspects with the definition of Leporiconus Iredale, 1930 by Tucker & Tenorio (2009). Aside from the general shell shape with the typical dome-shaped spire, especially the pustulose spiral cords, the tuberculate early spire whorls, the striate spire whorls and the shallow subsutural flexure support the placement in this genus. It differs from its Paratethyan congeners in being more elongated and obviously torpedo-shaped and in having a domed spire with a pointed tip. Among the modern representatives of this genus, the Indo-West Pacific species Leporiconus tenuistriatus (Sowerby II, 1858), Leporiconus granum (Röckel & Fischöder, 1985) and Leporiconus nucleus (Reeve, 1848) are highly reminiscent of L. transsylvanicus. Paleoenvironment. No information.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFBFFEAFF5FA989FE444791.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Transylvanian Basin: Lăpugiu de Sus (Romania) (Hoernes & Auinger 1879).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFBFFEBFF5FAF39FC4443C2.taxon	materials_examined	Type species (by original designation): Conus ambiguus Reeve, 1844. Recent, West Africa. Note. According to Tucker & Tenorio (2009), shells of the genus Monteiroconus da Motta, 1991 are characterised by having concave spire whorls with cords present or absent; the spire may be convex in profile; the subsutural sinus is shallow to moderately deep, and the protoconch is multispiral. The genus has a rich European fossil record and today has a West African distribution. The molecular phylogeny presented by Puillandre et al. (2014 a) confirmed the group to be monophyletic, although the authors recognise these groups at subgenus level. However, they also included two species (C. gladiator Broderip, 1833 and C. mus Hwass in Brugière, 1792), which one would not have expected based on shell characters, as both of these species have coarse tubercles on the spire whorls. Moreover, they both have a Caribbean distribution, whereas Monteiroconus as defined by Tucker & Tenorio (2009) is West African. We provisionally restrict the usage of the genus to that suggested by Tucker & Tenorio (2009).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFBFFEBFF5FAF39FC4443C2.taxon	description	Monteiroconus is a speciose conid genus in the Paratethyan assemblages. Based on these species we add that the genus is characterised by moderately large to large-sized, relatively solid shells. The spire is low to almost flat and in most species the spire whorls are striate. Often, spire whorls are bulgy along the lower suture. The subsutural flexures in all the species treated here are medium deep to deep, moderately to strongly curved and moderately to strongly asymmetrical. The relative diameter of the last whorl and length of the siphonal canal are variable but generally species have a wide to moderately wide last whorl.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFAFFE9FF5FAA8AFC0045AF.taxon	description	Conus (Lithoconus) subacuminatus d’Orbigny, 1852 — Strausz 1966: 454, pl. 67, fig. 13.? Conus (Lithoconus) subacuminatus d’Orbigny — Bohn-Havas 1973: 1069, pl. 7, fig. 10.? Conus antiqus [sic] Lamarck – Chira & Voia 2001: 156, pl. 1, figs 3 a – b.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFAFFE9FF5FAA8AFC0045AF.taxon	materials_examined	Type material. Lectotype catalogue number MNHN B 35774, stored in the Muséum national d'histoire naturelle, Paris; no detailed locality given; Piedmont Italy, probably Burdigalian (early Miocene). This specimen was found again in the Paris collection in the 1980 s (Ferrero-Mortara et al. 1984). Therefore, Hall (1966) considered the type specimen as lost and designated a neotype based on a Burdigalian specimen from the Turin Hills in the Saccocollection of the Museo Regionale di Scienze Naturali, Torino (BS. 038.02.021). Studied material. 1 spec. NHMW 1853 / 0003 / 0007, 1 spec. NHMW 1846 / 0037 / 0035, 2 spec. NHMW 1855 / 0014 / 0364, 3 spec. NHMW 1856 / 0001 / 0364, 2 spec. NHMW 1856 / 0004 / 0034, 1 spec. NHMW 1856 / 0050 / 0112, 1 spec. NHMW 1863 / 0015 / 1283, 2 spec. NHMW 1997 z 0178 / 1480, 1 spec. NHMW 1970 / 1396 / 1486, 9 spec. NHMW 2013 / 0479 / 2041, 2 spec. NHMW 2013 / 0479 / 1623, GBA 1856 / 004 / 0003, illustrated in Hörnes (1851, pl. 4, fig. 1), all Gainfarn (Austria); 4 spec. NHMW 1858 / 0015 / 0067, Steinebrunn (Austria); 2 spec. NHMW 1860 / 0001 / 0063, 1 spec. NHMW 1851 / 0010 / 0007, illustrated in Hörnes (1851, pl. 4, fig. 2), 1 spec. NHMW 1850 / 0001 / 0 0 54, illustrated in Hoernes & Auinger (1879, pl. 4, fig. 1), all Mikulov-Kienberk (Czech Republic). Illustrated material. Figs 24 A 1 – A 3: SL: 137.3 mm, MD: 68.7 mm, NHMW 2016 / 0048 / 0001; all Gainfarn (Austria); Figs 24 B 1 – B 3, 17 N: SL: 109.8 mm, MD: 49.1 mm, NHMW 1970 / 1396 / 1486; Figs 24 C 1 – C 3: SL: 128.4 mm, MD: 59.9 mm, NHMW 1856 / 0050 / 0112.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFAFFE9FF5FAA8AFC0045AF.taxon	description	Description: Large shells with low, coeloconoid spire. Early spire whorls angulated, later distinctly concave with spiral striae, which are most prominent on the adapical half of the whorls. Subsutural flexure moderately deep, moderately curved, strongly asymmetrical. Last whorl elongate, straight-sided; position of maximum diameter close below shoulder; weakly constricted at base. Weak spiral grooves on abapical third of last whorl are usually developed and become stronger and more closely spaced abapically. Aperture narrow, straight, slightly narrowed in middle part. Siphonal canal long, weakly recurved posteriorly with broad, rounded fasciole. Colour pattern under UV light consisting of numerous, thin, continuous and regular brown spiral lines covering the entire last whorl. Shell measurements and ratios. n = 10 adult and subadult specimens: largest specimen: SL: 128.4 mm, MD: 59.9 mm, mean SL: 107.9 mm (σ = 14.2), mean MD: 51.12 mm (σ = 6.5), spire angle: µ = 107.9 ° (σ = 8.8 °), last whorl angle: µ = 26 ° (σ = 1.9 °), LW: µ = 2.1 (σ = 0.1), RD: µ = 0.56 (σ = 0.03), PMD: µ = 0.94 (σ = 0.03), RSH: µ = 0.16 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFAFFE9FF5FAA8AFC0045AF.taxon	discussion	Discussion. This species is an outlier within Monteiroconus concerning the strongly elongate last whorl. Nevertheless, we follow Tucker & Tenorio (2009) and place it in Monteiroconus based on its genus-typical spire and sculpture. The colour pattern of Serravallian specimens of M. antiquus from the Karaman Basin in Turkey consists of narrow, axially arranged flammules (Landau et al. 2013), thus differing from the Paratethyan shell with densely spaced, continuous spirals. Comparable patterns, however, are also documented from Tortonian shells from Montegibbio (Italy) (see Davoli 1972). This is among the largest cone shells in the Paratethys and is recognized easily by its slender shell with short coeloconoid spire and concave spire whorls. Hoernes & Auinger (1879) illustrated a rather bulky subadult shell, which they identified with the smaller and shorter Conus sharpeanus Pereira da Costa, 1866. De Gregorio (1885) introduced epellus as new name for the specimens illustrated in Hörnes (1851, pl. 4, figs 1 – 3), which is considered herein to be a subjective junior synonym of Conus antiquus Lamarck, 1810. The juvenile specimen illustrated by Hörnes (1851, pl. 4, fig 3.) was excluded by Hall (1966) from C. antiquus and placed it in Plagioconus elatus (Michelotti, 1847). Unfortunately, this specimen is lost and its status remains unclear. Paleoenvironment. Monteiroconus antiquus occurs only in shallow sublittoral settings. At localities, such as Gainfarn and Mikulov it was inhabiting sea grass meadows (Zuschin et al. 2007).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFFAFFE9FF5FAA8AFC0045AF.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn, Steinebrunn, Pötzleinsdorf (Austria) (Hoernes & Auinger 1879; Sieber 1958 b), Mikulov-Kienberk (Czech Republic) (Hoernes & Auinger 1879); Pannonian Basin: Pécsszabolcs, Szob (Hungary) (Strausz 1966); Transylvanian Basin: Lăpugiu de Sus (Romania) (Chira & Voia 2001). Proto-Mediterranean Sea and northeastern Atlantic. Widespread during the early Miocene to late Miocene in the northeastern Atlantic and the Proto-Mediterranean Sea; the species persists into the Pliocene in the Mediterranean Sea (see Landau et al. 2013 for detailed references).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF8FFE6FF5FAD61FC6F4516.taxon	materials_examined	Type material. The holotype was stored in the collection of the Geological and Geophysical Institute of Hungary in Budapest but seems to be lost (pers. comm. Klára Palotás); middle Miocene, Badenian (Langhian). The type locality of Conus pseudaldrovandi Sacco, 1893, considered to be a junior synonym of M. boeckhi, is the middle Miocene (Badenian) locality Ritzing in Austria. Studied material. 1 spec. NHMW 1870 / 0037 / 00021, 1 spec. NHMW 1870 / 0037 / 0002, 6 spec. NHMW 1930 / 0006 / 0037, Ritzing (Austria). Illustrated material. Figs 24 D 1 – D 6, 17 O: Ritzing (Austria): SL: 72.6 mm, MD: 45.5 mm, NHMW 1930 / 0006 / 0037; Figs 24 E 1 – E 3: Ritzing (Austria): SL: 81.5 mm, MD: 51.2 mm, NHMW 1870 / 0037 / 0002, illustrated in Hoernes & Auinger (1879, pl. 4, fig. 2), holotype of Conus pseudaldrovandi Sacco, 1893; Figs 24 F 1 – F 3: Ritzing (Austria), SL: 54.5 mm, MD: 31.8 mm, NHMW 1930 / 0006 / 0037. Revised description. Moderately large shells, up to 81 mm in height; about 10 teleoconch whorls; early spire conical with nearly flat, weakly striate to nearly smooth spire whorls; late spire strongly depressed with slightly concave, faintly striate whorls and somewhat irregular, deeply impressed, irregular suture; subsutural flexure deep, strongly curved, strongly asymmetrical. Last whorl conical with distinct shoulder, position of maximum diameter directly below angulation; very weak and somewhat irregular constriction below shoulder within the adapical third and distinct constriction at base. Fasciole short, strongly raised; siphonal canal short, recurved, thickened; inner lip broad, twisted. Colour pattern in UV light comprising axially elongated blotches parallel to the subsutural flexure on spire, slightly narrower than the interspaces; blotches continue via shoulder on last whorl. Last whorl covered by 3 – 4 broad bands and small dashes and blotches in the interspaces; position of bands variable but typically in some distance from shoulder, at mid-whorl and in lower third of last whorl.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF8FFE6FF5FAD61FC6F4516.taxon	description	Shell measurements and ratios. Only four adult shells are available: largest specimen: SL: 81.5 mm, MD: 51.2 mm, mean SL: 73.5 mm (σ = 5.7), mean MD: 45.5 mm (σ = 4.2), spire angle (without high conical initial part): µ = 142, 3 ° (σ = 9.5), last whorl angle: µ = 38.8 ° (σ = 0.96), LW: µ = 1.6 (σ = 0.07), RD: µ = 0.70 (σ = 0.03), PMD: µ = 0.95 (σ = 0.02), RSH: µ = 0.11 (σ = 0.02); the holotype is a subadult specimen: SL: 47 mm, MD: 28 mm (according to Halaváts 1884).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF8FFE6FF5FAD61FC6F4516.taxon	discussion	Discussion. The illustration of the holotype of Monteiroconus boeckhi from Hidas shows the characteristic colour pattern, which is also preserved in the specimens from Ritzing. Therefore, Conus pseudaldrovandi Sacco, 1893, which was established by Sacco (1893 a) for the specimen from Ritzing illustrated in Hoernes & Auinger (1879, pl. 4, fig. 2), is a subjective junior synonym of C. boeckhi. In their discussion of Conus aldrovandi, Hoernes & Auinger (1979) also referred to the specimen from Grund discussed by Hörnes (1851, pl. 1, fig. 2). Doderlein (1863) proposed Conus hornesi as new name for this shell from Grund in Austria, which is not conspecific with either M. boeckhi or M. aldrovandi. Despite some similarities, the Italian Pliocene C. aldrovandi differs from both Paratethyan species in its convex spire whorls and the position of the aperture distinctly below the shoulder of the preceding whorl. The lack of spiral striae on the spire allows a clear separation from M. mojsvari. Moreover, the ice cream cone shaped last whorl, the conspicuous, swollen fasciole and the colour pattern of broad bands allow a separation from all other Paratethyan species. Paleoenvironment. The mollusc assemblage and geological situation of Ritzing suggests sandy fore shore settings in an embayment of the Pannonian Basin (Harzhauser et al. 2014 and references therein).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF8FFE6FF5FAD61FC6F4516.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Pannonian Basin: Hidas (Hungary) (Halaváts 1884); Oberpullendorf Basin: Ritzing (Austria) (Hoernes & Auinger 1879).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF7FFE7FF5FACB9FEC8400F.taxon	description	Conus Mercati Brocc. — Hörnes 1851: 23 (partim), pl. 2, figs 3 a – c. Conus (Lithoconus) Mercati Brocc. — Hoernes & Auinger 1879: 27 (partim). [Conus (Lithoconus)] conicomaculata Sacc. — Sacco 1893 a: 20 [nov. nom. pro Conus mercati in Hörnes 1851, pl. 2, fig. 3].	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF7FFE7FF5FACB9FEC8400F.taxon	materials_examined	Type material. Holotype NHMW 1860 / 0001 / 0054 a, illustrated in Hörnes (1851, pl. 2, fig. 3), Mikulov-Kienberk (Czech Republic); middle Miocene, Badenian (late Langhian). Studied material. Holotype and 2 spec. NHMW 1860 / 0001 / 00054, Mikulov-Kienberk (Czech Republic). Illustrated material. Figs 17 P, 24 G 1 – G 3: Mikulov-Kienberk (Czech Republic): SL: 60.1 mm, MD: 37.8 mm NHMW 1860 / 0001 / 00054. Revised description. Medium-sized shells with low conical spire consisting of broad, flat to slightly concave spire whorls with weak striae. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Distinct, subangulate shoulder; position of maximum diameter slightly below shoulder. Last whorl broadly conical to weakly ventricose, not constricted; few weak spiral cords on base. Aperture wide with straight outer lip. Siphonal canal wide, short, straight; siphonal fasciole weakly swollen. Colour pattern under UV light consisting of about 12 spirals of wide-spaced subquadratic to elongate rectangular blotches.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF7FFE7FF5FACB9FEC8400F.taxon	description	Shell measurements and ratios. Only 3 specimens are available: SL: 60.1 / 53.3 / 49.5 mm, MD: 37.8 / 32.9 / 31.3 mm, spire angle: 118 / 110 / 119 °, last whorl angle: 41 / 37 / 38 °, LW: 1.59 / 1.62 / 1.58, RD: 0.71 / 0.72 / 0.73, PMD: 0.85 / 0.91 / 0.85, RSH: 0.12 / 0.14 / 0.14.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF7FFE7FF5FACB9FEC8400F.taxon	discussion	Discussion. This species was introduced by Sacco (1893 a) as variety name of Conus mercati Brocchi, 1814. Morphologically, it is very similar to Pliocene specimens of Monteiroconus mercati (Brocchi, 1814) as shown by Pinna & Spezia (1978) and Hall (1966). Nevertheless, it differs in its striate spire whorls, which are smooth in M. mercati (see discussion in Landau et al. 2013). The smaller size, higher spire and shorter last whorl allow a separation from Monteiroconus mojsvari (Hoernes & Auinger, 1879). Paleoenvironment. Shallow marine seagrass environments.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF7FFE7FF5FACB9FEC8400F.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Mikulov-Kienberk (Czech Republic) (Hörnes 1851).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF6FFE5FF5FA9C1FDA84286.taxon	description	Conus (Dendroconus) Daciae nov. form. — Hoernes & Auinger 1879: 21, pl. 3, fig. 1.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF6FFE5FF5FA9C1FDA84286.taxon	description	Conus (Lithoconus) mercati miocaenicus [sic] Sacco — Nicorici 1972: 70, pl. 17, figs 1 – 2 [non Conus miocenicus Sacco, 1893].	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF6FFE5FF5FA9C1FDA84286.taxon	materials_examined	Type material. Syntype NHMW 1858 / 0043 / 0002, Lăpugiu de Sus (Romania), illustrated in Hoernes & Auinger (1879, pl. 3, fig. 1); syntype NHMW 1949 / 0005 / 0002, Nemeşeşti (Romania), illustrated in Hoernes & Auinger (1879, pl. 3, fig. 5), holotype of Conus exloroisi Sacco, 1893. Studied material. Syntypes and 2 spec. NHMW 1856 / 0007 / 0001, 2 spec. NHMW 1856 / 0007 / 0001, Lăpugiu de Sus (Romania); 2 spec. NHMW 1848 / 003 / 003, 6 spec. NHMW 1930 / 0006 / 0034, 1 NHMW 1970 / 1396 / 0834, Ritzing (Austria); 2 spec. NHMW 1856 / 0002 / 0002, Grund (Austria); 1 spec. NHMW 1862 / 0032 / 0001, Pöls (Austria); all middle Miocene, Badenian (Langhian). Illustrated material. Figs 25 A 1 – A 3: syntype, Lăpugiu de Sus (Romania), SL: 85.9 mm, MD: 64.5 mm, NHMW 1858 / 0043 / 0002, illustrated in Hoernes & Auinger (1879, pl. 3, fig. 1); Figs 25 B 1 – B 3, 17 Q: Ritzing (Austria): SL: 81.9 mm, MD: 56.1 mm, NHMW 1930 / 0006 / 0034. Revised description. Moderately large to large shells with a maximum height of 87 mm. Spire depressed to flat; adapical part of spire whorls flat, striate; abapical part weakly convex, often elevated, resulting in weakly concave cross section. Subsutural flexure of medium depth, moderately curved, strongly asymmetrical. Last whorl broad, straight-sided, rapidly contracting, not constricted at base. Position of maximum diameter slightly below rounded shoulder. Aperture wide, broadening abapically, protruding above shoulder adapically in well preserved, fully grown specimens. Siphonal canal short, straight, wide; siphonal fasciole broad, rounded. Spiral grooves on base, rarely extending to mid-whorl.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF6FFE5FF5FA9C1FDA84286.taxon	description	Shell measurements and ratios. n = 14 adult and subadult specimens: largest specimen: SL: 86.8 mm, MD: 64.5 mm, mean SL: 67.8 mm (σ = 13.3), mean MD: 48.2 mm (σ = 9.1), spire angle: µ = 154 ° (σ = 12.3 °), last whorl angle: µ = 42.2 ° (σ = 2.3 °), LW: µ = 1.41 (σ = 0.07), RD: µ = 0.73 (σ = 0.03), PMD: µ = 0.9 (σ = 0.02), RSH: µ = 0.03 (σ = 0.02).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF6FFE5FF5FA9C1FDA84286.taxon	discussion	Discussion. The larger syntype of Monteiroconus daciae is a slightly worn specimen with abraded base, fasciole and adapical part of the aperture. Therefore, it lacks the characteristic protruding adapical aperture. This specimen is also extraordinarily large and broad compared to other specimens. However, the specimens from the Karaman Basin of Turkey attain an even greater maximum size (height 104.3 mm; Landau et al. 2013). We were not able to see a colour pattern in the Paratethyan specimens under UV light, however, a pattern is present in the Karaman specimens, consisting of axial comma-shaped reddish stripes on the spire and irregular, elongate, oblong blotches arranged in three bands, coalescent into broad axial flammules on some specimens (Landau et al. 2013).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF6FFE5FF5FA9C1FDA84286.taxon	description	Paleoenvironment. The locality Ritzing represents a shallow sublittoral environment (Harzhauser et al. 2014). The deeper water occurrences from Lăpugiu de Sus may represent allochthonous specimens transported by storms into basinal settings (own observations M. H.).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF6FFE5FF5FA9C1FDA84286.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Bad Vöslau (Austria), Oberpullendorf Basin: Ritzing (Austria) (Hörnes 1851; Hoernes & Auinger 1879); North Alpine Foreland Basin: Grund (Austria); Styrian Basin: Pöls (Austria) (Hoernes & Auinger 1879); Pannonian Basin: Letkés (Hungary) (Kovács & Vicián 2013); Transylvanian Basin: Nemeșești, Timiș, Lăpugiu de Sus, Coşteiu de Sus (Romania) (Hoernes & Auinger 1879; Boettger 1906); Caransebeş-Mehadia Basin: Valea Bela Reca (Romania) (Hinculov 1968); Dacian Basin: Opanec (Bulgaria) (Kojumdgieva & Strachimirov 1960), southern Pannonian Basin: Relievac at Prnjavor (Bosnia and Herzegovina) (Eremija 1971). Proto-Mediterranean Sea and northeastern Atlantic: Serravallian (middle Miocene): Karaman Basin, Turkey (Landau et al. 2013). Tortonian (late Miocene): Cacela Basin, Portugal (Pereira da Costa 1866); Montegibbio (Italy) (Davoli 1972).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF4FFE2FF5FAC49FF5643A6.taxon	description	Conus (Dendroconus) betulinoides Lamarck mut. girondicus nov. mut — Peyrot 1931: 98, nr. 1170, pl. 3, figs 4 – 6.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF4FFE2FF5FAC49FF5643A6.taxon	materials_examined	Type material. Holotype illustrated in Peyrot (1931, pl. 3, fig. 5), Aquitaine Basin, Salles (Debat), France; middle Miocene, Serravallian. Studied material. Bad Vöslau (Austria): 1 spec. NHMW 2010 / 0004 / 1569 a, 1 spec. NHMW 2010 / 0004 / 1569 b 3 spec. NHMW 2010 / 0004 / 1569 c, 4 spec. NHMW 2010 / 0004 / 1568 a – 2010 / 0004 / 1568 d, 2 spec. NHMW 2010 / 0004 / 1571 a – 2010 / 0004 / 1571 b, 1 spec. NHMW 2010 / 0004 / 1572. Illustrated material. Figs 25 C 1 – C 3, 17 R: Bad Vöslau (Austria): SL: 127 mm, MD: 79.5 mm, NHMW 2010 / 0004 / 1569 a; Figs 25 D 1 – D 3: Bad Vöslau (Austria): SL: 108.6 mm, MD: 62 mm, NHMW 2010 / 0004 / 1569 b.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF4FFE2FF5FAC49FF5643A6.taxon	description	Description: Very large and solid species; strongly depressed, nearly flat spire; spire whorls striate with deep concavity in adapical half, becoming more prominent during ontogeny; abapical half of spire whorls bulging; spiral whorls increase regularly in width, last spire whorl not exceptionally broad. Suture impressed. Strongly shouldered with rounded periphery; position of maximum diameter just below shoulder. Subsutural flexure deep, strongly curved, strongly asymmetrical. Last whorl moderately elongate, regularly conical below periphery, not constricted. Slightly allometric growth, fully grown specimens being more elongate than subadults. Weak spiral cords on base and prominent prosocline, straight growth lines on last whorl; fasciole raised, rather narrow, covered by prominent growth lines. Inner lip broad, twisted, glossy well demarcated from fasciole. Aperture moderately wide, only weakly widening abapically; siphonal canal wide, short and straight. No colour pattern preserved. Shell measurements and ratios. n = 12: largest specimen: SL: 127.0 mm, MD: 79.5 mm, mean SL: 85.9 mm (σ = 5.3), mean MD: 54.4 mm (σ = 10.7), spire angle: µ = 154. 9 ° (σ = 11.1), last whorl angle: µ = 38.6 ° (σ = 1.98), LW: µ = 1.6 (σ = 0.08), RD: µ = 0.67 (σ = 0.03), PMD: µ = 0.86 (σ = 0.03), RSH: µ = 0.06 (σ = 0.02).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF4FFE2FF5FAC49FF5643A6.taxon	discussion	Discussion. This species is among the largest cone shells of the Paratethys and it is morphologically quite clearly defined. Nevertheless, all specimens were labelled in the NHMW collection as Conus betulinoides Lamarck, 1810 and Conus mercati miocaenicus Sacco, 1893 (= Monteiroconus mojsvari). In respect to the inventory history, none of the specimens was probably known to Hörnes (1851) and Hoernes & Auinger (1879) and later curators followed a rather broad species concept. A separation from Kalloconus betulinoides (Lamarck, 1810) is based on the nearly flat spire with deeply concave spire whorls and the comparatively narrow last spire whorl. Monteiroconus mojsvari (Hoernes & Auinger, 1879) differs in its more conical spire, the shallower concavity on spire whorls and especially in the less prominent shoulder; its fasciole is weaker and the inner lip much narrower. Monteiroconus antiquus (Lamarck, 1810), which is of comparable size, differs in its elongate slender outline and basal constriction. Monteiroconus daciae (Hoernes & Auinger, 1879) can attain a comparable size and also has an almost flat striate spire, but the spire whorls do not have the deep concavity in the adapical half seen in M. girondicus (Peyrot, 1931) and M. daciae has a shorter, wider last whorl (RD 0.73 vs. 0.67). Paleoenvironment. All specimens derive from offshore clays in Bad Vöslau, whereas Monteiroconus mojsvari and M. antiquus are typically found in nearshore deposits. This suggests also an ecological separation of these species.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF4FFE2FF5FAC49FF5643A6.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Bad Vöslau (Austria). Northeastern Atlantic: Serravallian (middle Miocene): Aquitaine Basin: Salles (Debat, Minoy) (Peyrot 1931).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF3FFE3FF5FAB69FD4B448F.taxon	description	C. [onus] Karreri — Hoernes 1878 b: 205 (nomen nudum). Conus (Lithoconus) Karreri nov. form. — Hoernes & Auinger 1879: 26, pl. 4, fig. 7. Conus (Lithoconus) Fuchsi nov. form. — Hoernes & Auinger 1879: 26, pl. 4, figs 4 – 5. Lithoconus parvicaudatus (Sacco, 1893) — Kovács & Vicián 2013: 74, figs 77 – 78 [non Conus parvicaudatus Sacco, 1893].	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF3FFE3FF5FAB69FD4B448F.taxon	materials_examined	Type material. Lectotype (designated herein), NHMW 1851 / 0002 / 0004, Grund (Austria), specimen illustrated in Hörnes (1851, pl. 1, fig. 2); middle Miocene, Badenian (Langhian). The type locality of Conus karreri Hoernes & Auinger, 1879, considered to be junior synonym of M. hoernesi, are the middle Miocene (Badenian) localities Bad Vöslau and Gainfarn in Austria. The type locality of Conus fuchsi Hoernes & Auinger, 1879, considered to be junior synonym of M. hoernesi, is the middle Miocene (Badenian) locality Lăpugiu de Sus in Romania. Studied material. Lectotype and 2 spec. NHMW 1868 / 0001 / 0203, Grund (Austria); 1 spec. NHMW 1874 / 0024 / 0058 Ritzing (Austria); 1 spec. NHMW 1853 / 0002 / 0045, Bad Vöslau (Austria); 1 spec. NHMW 1849 / 0023 / 0 0 0 4, Bad Vöslau (Austria), syntype of Conus fuchsi illustrated in Hoernes & Auinger (1879, pl. 4, fig. 5); 1 spec. NHMW 1856 / 0050 / 0110, Gainfarn (Austria), syntype of Conus fuchsi illustrated in Hoernes & Auinger (1879, pl. 4, fig. 4); 1 spec. NHMW 1870 / 0033 / 0001, Lăpugiu de Sus (Romania), syntype of Conus karreri illustrated in Hoernes & Auinger (1879, pl. 4, fig. 7). Illustrated material. Figs 26 A 1 – A 3: lectotype, Grund (Austria): SL: 76.0 mm, MD: 49.1 mm, NHMW 1851 / 0002 / 0004, illustrated in Hörnes (1851, pl. 1, fig. 2); Figs 26 B 1 – B 3: Lăpugiu de Sus (Romania): SL: 78.2 mm, MD: 51.2 mm, NHMW 1870 / 0033 / 0001, illustrated in Hoernes & Auinger (1879, pl. 4, fig. 7); Figs 26 C 1 – C 3, 17 S: Gainfarn (Austria): SL: 75.5 mm, MD: 47.5 mm, NHMW 1856 / 0050 / 0110, illustrated in Hoernes & Auinger (1879, pl. 4, fig. 4). Revised description. Large robust shells; spire low and conical with weakly convex, blunt, smooth to faintly striate whorls; rarely a shallow concavity may be developed (“ fuchsi ” - morph). Late spire whorls slightly gradate; suture impressed. Subsutural flexure of medium depth, moderately curved, moderately asymmetrical. Prominent, rounded shoulder, with position of maximum diameter slightly below shoulder, whorl profile straight to slightly constricted below, resulting in a depressed weakly club-shaped profile. Moderately narrow aperture with shallow, narrow posterior canal; anteriorly widening, terminating in wide, very short, straight canal. Fasciole weakly swollen, indistinct, demarcated from twisted inner lip by deep groove; base somewhat constricted below siphonal fasciole. Faint spiral cords on last whorl, visible only in grazing light. No colour pattern preserved.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF3FFE3FF5FAB69FD4B448F.taxon	description	Shell measurements and ratios. n = 8: largest specimen: SL: 81.1 mm, MD: 53.9 mm, mean SL: 74.8 mm (σ = 2.1), mean MD: 48.7 mm (σ = 1.6), spire angle: µ = 123.6 ° (σ = 7.4 °), last whorl angle: µ = 43.5 ° (σ = 2.8 °), LW: µ = 1.5 (σ = 0.1), RD: µ = 0.75 (σ = 0.03), PMD: µ = 0.84 (σ = 0.01), RSH: µ = 0.13 (σ = 0.06).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF3FFE3FF5FAB69FD4B448F.taxon	discussion	Discussion. Doderlein (1863) introduced Conus Hornesi as new name for Conus Aldrovandi sensu Hörnes, 1851 (non Brocchi, 1814) including a very brief description. He obviously referred to the Viennese palaeontologist Moritz Hörnes and “ hornesi ” is a type setting error for “ hörnesi ”, which should be corrected to “ hoernesi ” according to ICZN Article 32.5. Therefore, we propose to emend the name to Conus hoernesi. Doderlein (1863) used the name for Tortonian specimens from Montegibbio and S. Agata in Italy but also as replacement name for the specimen of Hörnes (1851). Therefore, the Badenian shell from Grund in Austria is a syntype. For the sake of nomenclatural stability we select the specimen NHMW 1851 / 0002 / 0004 from Grund (Austria) as lectotype. Hoernes & Auinger (1879) stated that the specimen of Hörnes (1851, pl. 1, fig. 2) is conspecific with their Conus karreri, which is confirmed herein, but obviously they overlooked the paper by Doderlein (1863). Therefore, Conus karreri is a subjective junior synonym of C. hoernesi. Conus fuchsi was separated by Hoernes & Auinger (1879) based on the slight mid-whorl constriction of the last whorl and the higher position of the fasciole. We consider these differences to reflect only intraspecific variability and treat the illustrated syntypes of Conus fuchsi as subadult specimens of Monteiroconus hoernesi. The seemingly concave and incised siphonal canal of the specimen illustrated by Hoernes & Auinger (1879, pl. 4, fig. 7) is an artefact due to fragmentation. All other specimens show a wide, straight and convex margin of the siphonal canal.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF3FFE3FF5FAB69FD4B448F.taxon	description	The specimens described by Kovács & Vicián (2013) as Lithoconus parvicaudatus (Sacco, 1893) might rather represent a low spired M. hoernesi, representing the “ fuchsi ” - morph with concave last whorl. Paleoenvironment. Probably shallow marine coastal habitats based on the co-occurring mollusc assemblages (e. g. Letkés, Kovács & Vicián 2013, Ritzing, Harzhauser et al. 2014).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF3FFE3FF5FAB69FD4B448F.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn, Bad Vöslau (Austria); Alpine Carpathian Foredeep: Grund (Austria) (Hoernes & Auinger 1879); Oberpullendorf Basin: Ritzing (Austria) (own data); Pannonian Basin: Letkés (Hungary) (Kovács & Vicián 2013); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus (Romania) (Boettger 1902, 1906).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF2FFFEFF5FAE41FC32402A.taxon	description	[Dendroconus] [Conus] Gainfahrensis n. f. — Hoernes 1878 a: 195 (nomen nudum). Conus (Dendroconus) Mojsvari nov. form. — Hoernes & Auinger 1879: 18, pl. 3, fig. 2. Conus (Dendroconus) Gainfahrenensis nov. form. — Hoernes & Auinger 1879: 18, pl. 2, fig. 4. Conus (Lithoconus) Mercati Brocc. — Hoernes & Auinger 1879: 27 (partim). [Conus] L. [ithoconus] Mercatii var. miocaenica Sacc. — Sacco 1893 a: 20 [nov. nom. pro Conus mercati in Hörnes 1851, pl. 2, fig. 1]. Conus (Lithoconus) mercatii [sic] miocaenica [sic] Sacco — Csepreghy-Meznerics 1956: 421, pl. 10, figs 7 – 8.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF2FFFEFF5FAE41FC32402A.taxon	materials_examined	Type material. Syntype NHMW 1853 / 0010 / 0001, Gainfarn (Austria), illustrated in Hoernes & Auinger (1879, pl. 3, fig. 2), 2 spec. syntypes NHMW 1867 / 9919 / 0001, Coşteiu de Sus (Romania); middle Miocene, Badenian (late Langhian). The type locality of Conus gainfahrenensis Hoernes & Auinger, 1879, considered to be junior synonym of M. mojsvari, is the middle Miocene locality Gainfarn in Austria. The type locality of Conus miocaenicus Sacco, 1893, considered to be junior synonym of M. mojsvari, is the middle Miocene locality Pötzleinsdorf in Austria. Studied material. Syntypes and 1 spec. NHMW 1847 / 0037 / 0024, Pötzleinsdorf (Austria), illustrated specimen in Hörnes (1851, pl. 2, fig. 1), syntype of Conus miocaenicus Sacco, 1893; 1 spec. NHMW 1855 / 0045 / 0 0 0 1, Steinebrunn (Austria); 1 NHMW 1853 / 0003 / 0001, 1 spec. NHMW 1997 z 0178 / 1484, 1 spec. NHMW 1855 / 0045 / 0354, 4 spec. NHMW 1846 / 0037 / 0046, Gainfarn (Austria); 1 spec. NHMW 1848 / 0021 / 0023, Grinzing (Austria). Illustrated material. Figs 25 E 1 – E 2,26 I, 17 T: Pötzleinsdorf (Austria): SL: 103.6 mm, MD: 60.3 mm, NHMW 1847 / 0037 / 0024, illustrated in Hörnes (1851, pl. 2, fig. 1); Figs 25 F 1 – F 2,26 J: syntype, Gainfarn (Austria): SL: 87.2 mm, MD: 54.3 mm, NHMW 1853 / 0010 / 0001, illustrated in Hoernes & Auinger (1879, pl. 3, fig. 2). Revised description. Moderately large to large shell up to 105 mm in height; about 10 teleoconch whorls. Depressed spire with initially moderately coeloconoid and later low conical outline; adapical half of spire whorls weakly concave, distinctly striate; abapical part convex, sometimes bulging. Subsutural flexure of medium depth, strongly curved, moderately asymmetrical; suture usually wavy and irregular. Shoulder weakly angulated; position of maximum diameter slightly below shoulder; last whorl regularly conical below periphery, not constricted. Surface smooth except for weak growth lines and indistinct, low spiral cords on base. Aperture moderately wide, anteriorly broadening; siphonal canal very short, wide, nearly straight; fasciole not well demarcated, weakly swollen with prominent growth lines; inner lip twisted. No colour pattern preserved.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF2FFFEFF5FAE41FC32402A.taxon	description	Shell measurements and ratios. n = 8 adult and subadult specimens: largest specimen: SL: 105 mm, MD: 63.8 mm, mean SL: 89.3 mm (σ = 11.2), mean MD: 53.7 mm (σ = 6.7), spire angle: µ = 132.3 ° (σ = 8.5 °), last whorl angle: µ = 37.3 ° (σ = 2.1 °), LW: µ = 1.67 (σ = 0.06), RD: µ = 0.66 (σ = 0.02), PMD: µ = 0.88 (σ = 0.02), RSH: µ = 0.09 (σ = 0.01).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF2FFFEFF5FAE41FC32402A.taxon	discussion	Discussion. The very broad species concept of Hörnes (1851) united several species as Conus mercati, which were later recognized as distinct taxa by Hoernes & Auinger (1879) and Sacco (1893 a). Monteiroconus mercati has smooth spire whorls, whereas the Paratethyan shells variously placed on M. mercati in the literature all have striate spire whorls (see Landau et al. 2013). The first available name for this mercati- like shells with conical spire (in contrast to M. daciae) and moderately elongate last whorl is Conus mojsvari Hoernes & Auinger, 1879. Later, Sacco (1893 a) introduced Conus mercatii miocaenica as variation name for the specimen illustrated by Hörnes (1851, pl. 2, fig. 1) without discussing its relation with C. mojsvari. The only difference between the syntypes of Monteiroconus mojsvari and the syntype of M. miocaenicus is the larger size, more elongate last whorl and the slightly gradate last two spire whorls of M. miocaenicus. As there are also intermediate specimens between both morphotypes, we consider M. miocaenicus a very large specimen of M. mojsvari and treat it as subjective junior synonym. The single specimen, described by Hoernes & Auinger (1879) as Conus gainfahrenensis (NHMW 1855 / 0045 / 0351), is just an aberrant specimen of M. mojsvari with a slight constriction in the middle of the last whorl. Paleoenvironment. All occurrences reflect shallow water environments, partly with sea grass meadows and coral patches (Zuschin et al. 2007; Kovács & Vicián 2013).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFF2FFFEFF5FAE41FC32402A.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn Pötzleinsdorf, Steinebrunn, Grinzing (Austria) (Hoernes & Auinger 1879; Sieber 1958 b; NHMW collection); Pannonian Basin: Várpalota, Letkés, Szob, Biatorbágy, Devecser, Diósd, Kemence, Márkháza, Mátraverebély, Sámsonháza, Budapest: Adria street, Illés street, Rákos (Hungary) (Kovács & Vicián 2013); Caransebeş-Mehadia Basin: Zorlenţu-Mare (Romania) (Florei 1961); Transylvanian Basin: Coşteiu de Sus, Lăpugiu de Sus (Romania) (NHMW collection); Zârand Basin: Minişul de Sus (Romania) (Nicorici & Sagatovici 1973).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFEFFFFEFF5FA9E2FE4444B4.taxon	description	[Dendroconus] [Conus] Neugeboreni n. f. — Hoernes 1878 a: 195 (nomen nudum). Conus (Dendroconus) Neugeboreni nov. form. — Hoernes & Auinger 1879: 19, pl. 1, fig. 5, pl. 2, fig. 5.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFEFFFFEFF5FA9E2FE4444B4.taxon	materials_examined	Type material. Holotype NHMW 1949 / 0005 / 0001, Lăpugiu de Sus (Romania). Studied material. Holotype. Illustrated material. Figs 26 D 1 – D 3, 17 U: holotype, Lăpugiu de Sus (Romania), SL: 66 mm, MD: 38 mm, NHMW 1949 / 0005 / 0001; middle Miocene, Badenian (Langhian).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFEFFFFEFF5FA9E2FE4444B4.taxon	description	Description: Moderately large shell of pyriform shape. Low conical spire with pointed apex; spire whorls deeply concave in adapical half, forming a convex bulge in abapical part; smooth, suture narrow and regular. Shoulder rounded; last whorl moderately convex above maximum diameter, strongly constricted below; subsutural flexure shallow, moderately curved, moderately asymmetrical. Aperture narrow adapically, widening towards moderately narrow and long canal. Base with narrow, widely-spaced, convex spiral cords; siphonal fasciole very weak; inner lip narrow, slightly twisted. Colour pattern under UV light consisting of numerous, densely spaced spirals of dashes covering the last whorl; the blotches on the shoulder indicated by Hoernes & Auinger (1879, pl. 1 fig. 5) are a misinterpretation or are not visible any more. Shell measurements and ratios. Holotype: SL: 66 mm, MD: 38 mm, spire angle = 128 °, last whorl angle = 39 °, LW = 1.74, RD = 0.64, PMD = 0.8, RSH = 0.1.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFEFFFFEFF5FA9E2FE4444B4.taxon	discussion	Discussion. At first sight, this species might be considered synonym with Pseudonoduloconus austriacus (Hoernes & Auinger, 1879), due to its pyriform shape. It differs, however, in the narrower, deeply concave spire whorls, the regular suture, the rounded shoulder and the even more pronounced constriction of the base. The aperture is adapically narrower, as is the siphonal canal. The position of maximum diameter is placed lower, resulting in a Ficus - like outline and lastly M. neugeboreni (Hoernes & Auinger, 1879) lacks the subobsolete tubercles on the spire whorls seen in P. austriacus. As the fig-shaped outline is also atypical for Monteiroconus, this generic placement is tentative. Paleoenvironment. No information is available; shallow water elements are frequently found transported into deep marine settings at Lăpugiu de Sus.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFEFFFFEFF5FA9E2FE4444B4.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Transylvanian Basin: Lăpugiu de Sus (Romania) (Hoernes & Auinger 1879).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFEFFFFFFF5FAE47FB3545FF.taxon	materials_examined	Type material. The holotype illustrated in Hörnes (1851, pl. 1, fig. 1) is lost. Studied material. 1 spec. NHMW 1869 / 0001 / 0020, 1 spec. NHMW 2010 / 0004 / 1574, 2 spec. NHMW 2010 / 0004 / 1575 Bad Vöslau (Austria); 1 spec NHMW A 989 Immendorf (Austria); 2 spec. NHMW 1852 / 0012 / 0010, 2 NHMW 1851 / 0026 / 0004 Grund (Austria). Illustrated material. Figs 26 E 1 – E 2: Bad Vöslau (Austria): SL: 124.5, mm, MD: 74.0 mm, 1 spec. NHMW 1869 / 0001 / 0020. Revised description. Very large shells with low spire; spire whorls striate, weakly convex with shallow central concavity in some specimens. Subsutural flexure shallow and asymmetrically curved. Shoulder rounded or faintly subangulate, position of maximum diameter close below shoulder. Last whorl elongate, regular conical in dorsal view, ventricose in apertural view; not constricted. Aperture wide, slightly excavated at junction from fasciole and inner lip. Fasciole broad, swollen, demarcated from glossy, inner lip by distinct groove. Growth lines on last whorl below shoulder prosocline and straight. Siphonal canal wide, slightly deflected. No colour pattern preserved.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFEFFFFFFF5FAE47FB3545FF.taxon	description	Shell measurements and ratios. n = 5: largest specimen: SL: 124.5 mm, MD: 74.00 mm, mean SL: 101.3 mm (σ = 13.3), mean MD: 62.8 mm (σ = 6.5), spire angle: µ = 135.6 ° (σ = 7.3 °), last whorl angle: µ = 39.2 ° (σ = 2.6 °), LW: µ = 1.61 (σ = 0.08), RD: µ = 0.69 (σ = 0.02), PMD: µ = 0.9 (σ = 0.02), RSH: µ = 0.1 (σ = 0.02), Hörnes (1851, pl. 1, fig. 1) reports a height of 128 mm and a width of 73 mm for the illustrated but lost specimen.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFEFFFFFFF5FAE47FB3545FF.taxon	discussion	Discussion. The specimen of Hörnes (1851, pl. 1, fig. 1) from Steinebrunn was already lost in 1879 (Hoernes & Auinger, 1879) and only a few additional specimens in the NHMW collection are available. Unfortunately, Hörnes (1851) presented only the dorsal view of his specimen. Therefore, the conspecificity with the Pliocene Kalloconus betulinoides (Lamarck, 1810) remained unchallenged. Only Sacco (1893 a) proposed the variety name pervindobonensis for this specimen, without discussing specific differences from the type. A re-investigation of the few additional Paratethyan specimens seems to support Sacco’s position. The concave and striate spire whorls, the excavated columella and the deflected siphonal canal of the Vienna Basin specimens differ from the Italian Kalloconus betulinoides as illustrated by Davoli (1972), Pinna & Spezia (1978) and Chirli (1997). The shell is broader and club-shaped, the last whorl less elongate and the last spire whorl is narrower. Therefore, we propose to separate the middle Miocene Paratethyan shells as Monteiroconus pervindobonensis (Sacco, 1893). The generic placement is based on the concave and striate spire whorls. Paleoenvironment. The specimens from Bad Vöslau, Grund and Immendorf were found in clays indicating an offshore environment but transport from shallow habitats cannot be excluded (Zuschin et al. 2005, 2006).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFEFFFFFFF5FAE47FB3545FF.taxon	distribution	Distribution in Paratethys.? Karpatian (early Miocene): Alpine-Carpathian Foredeep: Niederkreuzstetten (Austria) (Harzhauser 2002). Badenian (middle Miocene): Vienna Basin: Gainfarn, Steinebrunn, Bad Vöslau, (Austria); Alpine-Carpathian Foredeep: Grund, Immendorf (Austria) (Hoernes & Auinger 1879);? Korytnica (Poland) (Bałuk 1997);? Pannonian Basin: Letkés, Márkháza, Mátraverebély, Sámsonháza, Budapest: Rákos (Hungary) (Kovács & Vicián 2013);? Krka Basin: Gorenje Vrhpolje (Slovenia) (Mikuž 2009).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFEEFFFCFF5FACABFE5B4741.taxon	materials_examined	Type material. Holotype NHMW 1851 / 0010 / 0003, Mikulov-Kienberk (Czech Republic), illustrated in Hörnes (1851, pl. 2, fig. 2); middle Miocene, Badenian (late Langhian). Studied material. Holotype and 1 spec. NHMW 1851 / 0010 / 0004, Mikulov-Kienberk (Czech Republic); 3 spec. NHMW 1847 / 0037 / 0025, 2 spec. NHMW 1850 / 0009 / 0018, 2 spec. NHMW 1851 / 0026 / 0074, 2 spec. NHMW 1878 / 0041 / 0001, Pötzleinsdorf (Austria). Illustrated material. Figs 26 F 1 – F 3: holotype, Mikulov (Czech Republic): SL: 62.9 mm, 43.2 mm, NHMW 1851 / 0010 / 0004, illustrated in Hörnes (1851, pl. 2, fig. 2); Figs 26 G 1 – G 3: Mikulov (Czech Republic): SL: 62.8 mm, 38.7 mm, NHMW 2016 / 0034 / 0001; Fig. 26 H, 17 V: Mikulov (Czech Republic), SL: 53.9 mm, MD: 33.0 mm, NHMW 2016 / 0034 / 0001. Revised description. Moderately large shells; spire strongly depressed to flat with slightly protruding early spire. Adapical part of spire whorls flat to weakly concave, abapical part convex, often bulgy; faintly striate. Irregular, deeply incised suture. Subsutural flexure deep, moderately curved, strongly asymmetrical. Last whorl moderately broad, straight-sided to weakly ventricose, not constricted at base. Position of maximum diameter slightly below rounded shoulder. Aperture moderately wide, broadening abapically; adapically slightly expanding, extending to or slightly above apex. Siphonal canal very short, straight, wide; siphonal fasciole broad but weak, rounded. Spiral grooves on base. Colour pattern under UV light consisting of narrow flammulae on shoulder and broad, somewhat blotchy dark bands separated by three narrow fluorescing stripes.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFEEFFFCFF5FACABFE5B4741.taxon	description	Shell measurements and ratios. n = 13 adult and subadult specimens: largest specimen: SL: 82.8 mm, MD: 49.5 mm, mean SL: 64.0 mm (σ = 8.4), mean MD: 40.9 mm (σ = 4.9), spire angle: µ = 153 ° (σ = 9.7 °), last whorl angle: µ = 37.7 ° (σ = 1.7 °), LW: µ = 1.57 (σ = 0.07), RD: µ = 0.65 (σ = 0.03), PMD: µ = 0.93 (σ = 0.03), RSH: µ = 0.02 (σ = 0.02).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFEEFFFCFF5FACABFE5B4741.taxon	discussion	Discussion. This species is most reminiscent of Monteiroconus daciae (Hoernes & Auinger, 1879) but is distinctly more slender (RD 0.65 vs. 0.73). The differences in shape allow also a clear separation of both species in a Principal Component Analysis based on shell ratios (Fig. 27). Moreover, the broad bands on the last whorl differ from the dashes and blotches of M. daciae as documented by Landau et al. (2013). This species is known so far only from the mid-Badenian locality Mikulov-Kienberk and the late Badenian locality Pötzleinsdorf, both in the Vienna Basin. It might represent a northern offshoot of Monteiroconus daciae, which is mainly found in southern basins of the Paratethys. Paleoenvironment. The localities Pötzleinsdorf and Mikulov-Kienberk represent shallow sublittoral environments within the photic zone based on the co-occurring mollusc assemblages (Sieber 1953; own data, M. H.).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFEEFFFCFF5FACABFE5B4741.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Pötzleinsdorf (Austria), Mikulov- Kienberk (Czech Republic).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFEDFFF4FF5FAE02FE2240C6.taxon	materials_examined	Type species (subsequent designation by Cotton, 1945): Conus radiatus Gmelin, 1791. Recent, Indo-West Pacific. Note. Tucker & Tenorio (2009) gave only a brief description of Phasmoconus, stating that the shell was elongated and subcylindrical in shape with angulate shoulders. The protoconch is usually multispiral, but rarely paucispiral. The anal notch is shallow, and an anterior notch is either slight or absent. In addition, we notice that tuberculate early whorls are not rare in the genus. Puillandre et al. (2014 a, b) accepted Phasmoconus as a subgenus of Conus and listed Fulgiconus da Motta, 1991, Graphiconus da Motta, 1991, Thoraconus da Motta, 1991 and Nimboconus Tucker & Tenorio, 2013 as subjective synonyms. corđs, SCES = spiral corđs only on early spire whorls, FS = faint striae, US = unđulating suture, CW = concave spire whorls, T = tuberculate, BLW = beađs on last whorl; subsutural flexure (SSF): SSFD = đepth, SSFđ = relative đepth, PV = position of the vertex; MH = međium height, MW = međium wiđth, SA = spire angle, LWA = angle of last whorl, LW = length wiđth ratio, RD = relative điameter, PMD = position of maximum điameter, RSH = relative height of spire).. Genus Species SC SCES FS US CW T BLW SSFD SSFd PV Artemidiconus granularis 0 0 0 0 0 1 1 4.70 5.45 0.81 Conasprella berwerthi 0 0 0 0 1 1 1 2.28 4.73 0.41 Conasprella minutissima 1 1 0 0 0 1 1 2.22 4.07 0.44 Conilithes allioni 0 0 0 0 1 1 0 1.75 3.56 0.43 Conilithes antidiluvianus 0 0 0 0 1 1 0 1.52 2.36 0.44 Conilithes brezinae 0 0 1 0 1 1 0 1.45 2.95 0.53 Conilithes eichwaldi 0 0 1 0 0 1 0 ??? Conilithes exaltatus 0 0 1 0 1 1 0 1.68 3.04 0.53 Conilithes sceptophorus 0? 0 0 1 1 0 ???	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFEDFFF4FF5FAE02FE2240C6.taxon	description	Conus s. l. vindobonensis 1 1 0 0 0 1 0 1.55 5.17 0.43 Conus s. l. johannae 1 1 0 0 0 1 0 3.05 6.18 0.54 Conus s. l. posticestriatus 1 1 0 0 0 1 0 3.55 4.95 0.71 Conus s. l. sturi 1 1 0 0 0 1 0 2.04 4.98 0.36 Conus s. l. mucronatolaevis 1 1 0 0 0 1 0 2.75 6.85 0.53 Conus s. l. olivaeformis 0 0 0 0 0 1 0 2.73 6.25 0.55 Conus s. l. praelongus 1 1 0 0 0 1 0 2.91 6.74 0.55 Conus s. l. argillicola 1 1 0 0 0 1 0 ??? Kalloconus berghausi 0 1 0 1 0 0 0 3.24 5.48 0.47 Kalloconus hendricksi 0 1 1 1 0 0 0 2.76 5.59 0.50 Kalloconus cacellensis 0 0 1 0 0 0 0 4.02 13.75 0.41 Kalloconus tietzei 0 0 0 0 1 0 0 1.85 5.80 0.32 Kalloconus gallicus 0 0 1 0 0 0 0 1.96 5.03 0.50 Kalloconus hungaricus 0 0 0 0 0 0 0 1.94 5.67 0.42 Kalloconus letkesensis 1 1 1 1 0 0 0 2.01 4.53 0.33 …… continued on the next page. Genus Species SC SCES FS US CW T BLW SSFD SSFd PV …… continued on the next page. Genus Species SC SCES FS US CW T BLW SSFD SSFd PV …… continued on the next page. Genus Species MH MW SA LWA LW RD PMD RSH Artemidiconus granularis 13.60 7.20 68.00 41.00 1.90 0.74 0.90 0.29 Conasprella berwerthi 21.10 10.20 78.00 32.30 2.10 0.60 0.90 0.20 Conasprella minutissima 10.40 5.10 63.20 32.40 2.04 0.68 0.81 0.28 Conilithes allioni 23.90 12.70 100.10 33.70 1.89 0.66 0.90 0.21 Conilithes antidiluvianus 59.20 21.80 61.40 29.40 2.69 0.52 0.88 0.27 Conilithes brezinae 32.80 14.30 58.40 32.70 2.30 0.63 0.90 0.31 Conilithes eichwaldi 15.00 7.00 65.00 36.50 ???? Conilithes exaltatus 40.80 16.40 59.70 30.90 2.50 0.56 0.90 0.29 Conilithes sceptophorus 13.50 6.60 68.00 44.00 2.05 0.70 0.89 0.30 Conus s. l. vindobonensis 51.9 27.20 98.30 35.10 1.91 0.63 0.89 0.17 Conus s. l. johannae 74.25 38.75 94.00 32.50 1.92 0.64 0.90 0.19 Conus s. l. posticestriatus 50.10 26.10 80.00 37.00 1.90 0.65 0.86 0.20 Conus s. l. sturi 41.30 18.30 89.80 30.50 2.30 0.55 0.89 0.19 Conus s. l. mucronatolaevis 50.90 26.80 98.80 36.00 1.89 0.64 0.85 0.18 Conus s. l. olivaeformis 22.30 11.20 87.70 36.60 2.00 0.60 0.87 0.16 Conus s. l. praelongus 33.70 14.90 71.50 33.80 2.30 0.58 0.87 0.23 Conus s. l. argillicola ???????? Kalloconus berghausi 35.00 24.30 128.00 40.10 1.40 0.75 0.87 0.08 Kalloconus hendricksi 22.60 14.40 128.30 39.90 1.60 0.70 0.90 0.09 Kalloconus cacellensis 70.30 43.60 130.00 30.00 1.61 0.69 0.93 0.10 Kalloconus tietzei 62.10 41.20 141.00 41.00 1.51 0.72 0.85 0.08 Kalloconus gallicus 50.10 31.90 123.00 37.00 1.57 0.73 0.90 0.12 Kalloconus hungaricus 62.00 44.20 141.60 43.30 1.41 0.76 0.89 0.07 Kalloconus letkesensis 33.80 22.30 138.50 41.50 1.52 0.73 0.87 0.10 Kalloconus moravicus 39.40 26.20 132.10 37.90 1.50 0.72 0.91 0.08 Kalloconus neumayri 28.80 18.50 128.10 37.90 1.60 0.71 0.91 0.09 …… continued on the next page. Genus Species MH MW SA LWA LW RD PMD RSH Kalloconus ponderoaustriacus 49.80 27.60 115.20 34.60 1.80 0.64 0.90 0.10 Kalloconus ponderovagus 48.60 31.40 106.10 38.20 1.55 0.76 0.89 0.14 Kalloconus pseudohungaricus 52.40 35.60 134.70 40.50 1.47 0.74 0.90 0.08 Kalloconus tschermaki 62.70 36.80 108.30 38.50 1.70 0.70 0.93 0.16 Kalloconus voeslauensis 50.20 31.20 118.80 38.90 1.60 0.71 0.87 0.12 Lautoconus andreei? 39.50 ?????? Lautoconus eschewegi 37.40 20.80 113.30 37.00 1.80 0.63 0.86 0.12 Lautoconus ex. gr. bitorosus 55.30 31.90 130.00 37.00 1.73 0.65 0.81 0.11 Lautoconus kovacsi 18.60 11.40 102.00 42.30 1.64 0.74 0.80 0.17 Lautoconus magnolapugyensis 64.30 42.50 134.30 43.70 1.51 0.74 0.90 0.11 Lautoconus miovoeslauensis 51.20 30.40 121.90 38.70 1.69 0.68 0.86 0.12 Lautoconus pelagicus 57.60 28.20 83.30 33.30 2.09 0.58 0.88 0.17 Lautoconus pestensis 22.30 12.90 116.60 37.30 1.70 0.67 0.83 0.13 Lautoconus ponderosus 76.30 42.30 100.00 36.00 1.81 0.67 0.91 0.17 Lautoconus pseudoponderosus ???????? Lautoconus quaggaoides 24.20 14.60 122.8 41.00 1.66 0.69 0.84 0.12 Lautoconus rotundus 38.15 22.75 127.00 37.00 1.68 0.67 0.90 0.11 Lautoconus steinabrunnensis 51.80 28.20 101.10 35.30 1.80 0.65 0.88 0.17 Lautoconus steindachneri 46.40 29.40 116.90 37.50 1.58 0.72 0.89 0.12 Lautoconus subraristriatus 54.30 28.10 92.60 34.60 1.90 0.63 0.84 0.18 Leporiconus paratethyianus 32.67 17.00 88.67 37.30 1.92 0.68 0.83 0.23 Leporiconus suessi 56.80 25.70 88.60 31.30 2.20 0.54 0.89 0.16 Leporiconus transsylvanicus 47.10 19.90 84.60 29.20 2.40 0.51 0.87 0.17 Monteiroconus antiquus 107.90 51.12 107.90 26.00 2.10 0.56 0.94 0.16 Monteiroconus boeckhi 73.50 45.50 142.30 38.80 1.60 0.70 0.95 0.11 Monteiroconus conicomaculatus 54.30 34.00 115.70 38.70 1.60 0.72 0.87 0.13 …… continued on the next page Within the Paratethyan cones, Conus fuscocingulatus Hörnes, 1851 and C. schroeckingeri Hoernes & Auinger, 1879 are characterised by very shallow and weakly curved subsutural flexures and are reminiscent of some Lautoconus species, such L. eschewegi. These species plot also within the Lautoconus - field in the PCA based on shell ratios (Fig. 28). Both species, however, differ from Lautoconus s. s. in the presence of tuberculate early whorls. Thus, we base the placement of both species in the Indo-West Pacific genus Phasmoconus on the similarities in shell outline, the morphology of the subsutural flexure and the tuberculate spire whorls. In addition, we place Conus ottiliae Hoernes & Auinger, 1879 in Phasmoconus, based on the results of the PCA (Fig. 28) (see discussion in P. ottiliae).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE5FFF2FF5FA983FB7F461C.taxon	description	Conus (Lithoconus) friedbergi nomen nov. — Moisescu 1955 a: 165, pl. 14, figs 5 – 6. Conus fuscocingulatus Bronn. — Eremija 1959: 187, pl. 1, figs 7 – 7 a. Conus (Dendroconus) fuscocingulatus Bronn 1848 — Kojumdgieva in Kojumdgieva & Strachimirov 1960: 214, pl. 51, fig. 2.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE5FFF2FF5FA983FB7F461C.taxon	description	Conus (Chelyconus) friedbergi Moisescu, 1955 — Hinculov 1968: 149, pl. 37, figs 14 a – b. Conus (Dendroconus) berghausi exfuscocingulatus (Sacco, 1893) — Hinculov 1968: 152, pl. 38, fig. 9 [non Conus exfuscocingulatus Sacco, 1893]. Conus (Dendroconu s) fuscocingulatus Bronn — Atanacković 1963: 78, pl. 15, figs 5 – 5 a.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE5FFF2FF5FA983FB7F461C.taxon	materials_examined	Type material. Syntype NHMW 1846 / 0037 / 0055, Pötzleinsdorf (Austria), illustrated in Hörnes (1851, pl. 1, fig. 5); middle Miocene, Badenian (Serravallian). Studied material. Syntypes and 6 spec. NHMW 1861 / 0033 / 0001, Bujtur (Romania), 7 spec. NHMW 1861 / 0033 / 0002, Bujtur (Romania) including specimen illustrated in Hoernes & Auinger (1879, pl. 1, figs 10 – 11), including syntypes of Conus ochreocingulatus Sacco, 1893; 32 spec. NHMW 1846 / 0037 / 0054 a, Pötzleinsdorf (Austria); 8 spec. NHMW A 1608, Bujtur (Romania), 41 spec. NHMW 1836 / 0012 / 0108, Bujtur (Romania). Illustrated material. Figs 29 A 1 – A 3: Bujtur (Romania), SL: 36.5 mm, MD: 18.2 mm, NHMW 1861 / 0033 / 0 0 0 1, Bujtur (Romania); Figs 29 B 1 – B 3: Bujtur (Romania), SL: 34.1 mm, MD: 17.5 mm, NHMW 1861 / 0033 / 0001, Bujtur (Romania); Figs 29 C 1 – C 3: Bujtur (Romania), SL: 31.7 mm, MD: 18.7 mm, NHMW A 1608 (poetzleinsdorfensis- morph); Figs 29 D 1 – D 3, 17 W: Bujtur (Romania), SL: 34.4 mm, MD: 17.6 mm, NHMW 1861 / 0033 / 0002, specimen in Hoernes & Auinger (1879, pl. 1, fig. 11); Figs 29 E 1 – E 2: Bujtur (Romania), SL: 30.8 mm, MD: 15.6 mm, NHMW 1836 / 0012 / 0108 a; Figs 29 F 1 – F 3, 17 X: Bujtur (Romania), SL: 24.7 mm, MD: 14.7 mm, NHMW 1836 / 0012 / 0108 b (poetzleinsdorfensis- morph). Revised description. Moderately small to medium-sized shells; paucispiral protoconch; high conical initial spire; later spire low, with beaded keel just above suture; beads move towards suture within 3 rd – 5 th spire whorls and fade out as indistinct swellings within 6 th whorl. Later spire of variable height, usually elevated and slightly coeloconoid; spire whorl tops weakly convex, faintly striate (only visible in high magnification); impressed suture. Subsutural flexure variable, very shallow, weakly curved, moderately asymmetrical. Last spire whorl slightly broadening with indistinct shoulder. Last whorl moderately elongate, weakly ventricose, slightly constricted; aperture regularly widening towards long and reflected anterior canal. Fasciole swollen, long, twisted, often with weak axial threads; inner lip narrow, straight. Deep irregularly spaced spiral grooves with broad cords in lower third of last whorl. Colour pattern often preserved as brown lines and very prominent in UV light, consisting of about 12 – 15 rather regularly spaced, well defined, continuous spirals on last whorl (rarely discontinuous spirals occur), with broad interspaces; broadest interspace usually two spirals below shoulder. Spire with axially elongate flammulae and blotches especially on shoulder and last whorl; some specimens display a broad dark band on spire whorl tops (in UV) and a light spiral at suture.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE5FFF2FF5FA983FB7F461C.taxon	description	Shell measurements and ratios. n = 14: largest specimen: SL: 39.7 mm, MD: 21.9 mm, mean SL: 34.4 mm (σ = 2.3), mean MD: 18.5 (σ = 1.6), spire angle: µ = 104.6 ° (σ = 11.2 °), last whorl angle: µ = 35.2 ° (σ = 2.1 °), LW: µ = 1.87 (σ = 0.14), RD: µ = 0.63 (σ = 0.04), PMD: µ = 0.89 (σ = 0.03), RSH: µ = 0.14 (σ = 0.02).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE5FFF2FF5FA983FB7F461C.taxon	discussion	Discussion. The authorship is often attributed to Bronn, who used the name in his correspondence and listed it in Bronn (1848) as nomen nudum (considering it as junior synonym of Conus raristriatus Bellardi & Michelotti, 1840). Hörnes (1851) made the name available and based his description on specimens from numerous localities of the Austro-Hungarian Empire, clearly mixing several different species. Consequently, the status of this species is highly confused. Hörnes (1851) illustrated specimens as Conus fuscocingulatus, which represent two different species. He illustrated a specimen from Mikulov (Czech Republic) as figure 4 on plate 1 and a second one as variety from Pötzleinsdorf (Austria) as figure 5. Later, Hoernes & Auinger (1879) separated the specimen from Mikulov as Conus moravicus and kept the “ variety ” of Hörnes (1851) in the synonymy of Conus fuscocingulatus. The confusion was completed by Hoernes & Auinger (1879) by illustrating additional specimens as C. fuscocingulatus, which represent at least three species. The specimens illustrated by Hoernes & Auinger as fig. 12 is a stout shell with very low spire and distinct shoulder; under UV light it displays an irregular pattern of densely spaced, spirally arranged dashes and differs completely from Phasmoconus fuscocingulatus [the illustration of the colour pattern in Hoernes & Auinger (1879) is incorrect]. It is most probably a Kalloconus moravicus (Hoernes & Auinger, 1879). Their fig. 13 is also a very stout shell with nearly flat spire. For this specimen Sacco (1893 a) introduced Conus poetzleinsdorfensis as new name. Unfortunately, the specimens of Hörnes (1851, pl. 1, fig. 5) and Hoernes & Auinger (1879, pl. 1, fig. 10) are rather untypical for Phasmoconus fuscocingulatus due to the broad last whorl, whereas all other specimens are more slender. Consequently, Sacco (1893 a) proposed Conus ochreocingulata as new name for the shells o Hoernes & Auinger (1879, pl. 1, figs 10 – 11), which are treated as typical P. fuscocingulatus herein. This confusion is reflected in a very broad range of morphologies and lumped species identified in Paratethyan literature as this species. A Principal Component Analysis of the available specimens did not allow any separation of the “ poetzleinsdorfensis ” - morphs from Phasmoconus fuscocingulatus (Fig. 12); moreover, both morphs co-occur at Pötzleinsdorf and Bujtur. Therefore, we treat Conus poetzleinsdorfensis Sacco, 1893 and Conus ochreocingulata Sacco, 1893 as subjective junior synonyms of P. fuscocingulatus. Moisescu (1955) introduced Conus friedbergi for specimens from Bujtur (Romania), which do not differ from P. fuscocingulatus. In her discussion she referred to the specimen from Białogon (Poland) illustrated by Friedberg (1911, pl. 2, fig. 19) as Conus cf. avellana Lamarck. This specimen is lost and the illustration does not allow a clear identification. As Moisescu (1955) did not clearly state whether she considered the Polish or her Romanian specimen as type specimen, the status of Conus friedbergi remains vague. It is either a nomen dubium (or species inquirenda) or a subjective junior synonym of P. fuscocingulatus. Paleoenvironment. Most probably shallow marine environments based on the co-occurring mollusc assemblage and the geological setting at the very margin of the Vienna Basin (Sieber 1954).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE5FFF2FF5FA983FB7F461C.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): widespread in all Paratethyan basins.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE2FFF0FF5FA8CAFAC14192.taxon	description	[Chelyconus] [Conus] Ottiliae n. f. — Hoernes 1878 a: 195 (nomen nudum). Conus (Chelyconus) Ottiliae nov. form. — Hoernes & Auinger 1879: 42, pl. 6, figs 12 – 13. [Conus] Stephanoconus? Ottiliae (H. A.) — Sacco 1893 b: 119.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE2FFF0FF5FA8CAFAC14192.taxon	materials_examined	Type material. 2 syntypes NHMW 1870 / 0033 / 0009, Lăpugiu de Sus (Romania), illustrated in Hoernes & Auinger (1879, pl. 6, figs 12 – 13); middle Miocene, Badenian (Langhian). Studied material. Syntypes and 1 spec. NHMW 1847 / 0061 / 0001, Szob (Hungary). Illustrated material. Figs 29 G 1 – G 3: syntype, Lăpugiu de Sus (Romania): SL: 24.9 mm, MD: 12.1 mm, NHMW 1870 / 0033 / 0009, illustrated in Hoernes & Auinger (1879, pl. 6, fig. 13) (= holotype of Conus ottiliae miolapugyensis Sacco, 1893); Figs 29 H 1 – H 3, 30 A: Szob (Hungary): SL: 23.1 mm, MD: 11.4 mm, NHMW 1847 / 0061 / 0001. Revised description. Small biconical shells with low to moderately elevated spire and elongate conical last whorl. Protoconch multispiral. Pointed early spire with beaded whorls; beads become weak during ontogeny; later spire whorls with 5 – 6 prominent spiral cords; spire weakly gradate; nearly flat sutural ramp; subsutural flexure shallow, weakly curved, moderately asymmetrical. Last whorl elongate, with subangular shoulder, position of maximum diameter slightly below shoulder, faintly constricted. Shell surface glossy, sculpture consisting of about 20 raised spiral cords with spirally elongated beads, being most prominent on lower third of whorl but weak and smooth in upper half of last whorl. Aperture narrow with subparallel margins; siphonal canal short, nearly straight; fasciole very weak. Colour pattern in UV light consisting of two light (fluorescing) spiral lines just below shoulder and at midshell and dots coinciding with sculpture on spire and last whorl.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE2FFF0FF5FA8CAFAC14192.taxon	description	Shell measurements and ratios. 2 adult specimens: SL: 24.9 / 23.1 mm, MD: 12.1 / 11.4 mm, spire angle: 103 ° / 105 °), last whorl angle: 34 ° / 33 °, LW: 2.06 / 2.03, RD: 0.58 / 0.57, PMD: 0.95 / 0.96, RSH: µ = 0.16 / 0.14.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE2FFF0FF5FA8CAFAC14192.taxon	discussion	Discussion. This species was placed by Tucker & Tenorio (2009) in Conilithes Swainson, 1840 but the shallow subsutural flexure, the elongate conical shape with low spire and the beaded spiral cords on the last whorl do not support this placement. Despite an overall similarity with Conasprella Thiele 1929, the weakly curved subsutural flexure does not correspond to the other species placed herein in Conasprella. Moreover, it does not plot close to other Conasprella species in the PCA based on shell ratios (fig. 28). Instead, this species plots close to Phasmoconus fuscocingulatus and P. schroeckingeri and agrees with these species in overall outline and subsutural flexure. The only marked difference is the beads on the last whorl. This feature, however, does not contradict a placement in Phasmoconus, as extant species of this genus may form beaded cords as well [e. g. Phasmoconus alabaster (Reeve, 1849)]. Sacco (1893 b) mentioned this species as rare from the Tortonian of Stazzano and Montegibbio and introduced several names for the late Miocene Italian specimens. He even proposed to separate the specimen illustrated by Hoernes & Auinger (1879, pl. 6, fig. 13) as distinct variety Conus ottiliae miolapugyensis, which is unjustified. Therefore, we consider Conus miolapugyensis Sacco, 1893 to be a subjective junior synonym of Conus ottiliae Hoernes & Auinger, 1879. Conus ottiliae asperula Sacco, 1893 and C. o. ovulatina Sacco, 1893 might be conspecific with the Paratethyan species but we did not study the types in Turin. In any case it is surely incorrect to treat these specimens as Lautoconus pelagicus (Brocchi, 1814) as proposed by Hall (1966). Paleoenvironment. The mollusc fauna and lithofacies of Szob suggest a sandy coastal environment with sea grass (Dulai 1996); there is no information for the other occurrences.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE2FFF0FF5FA8CAFAC14192.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Carpathian-Foredeep: Ostrava (Czech Republic) (Kittl, 1887);? Bükk Mountains: Borsodbóta (Hungary) (Csepreghy-Meznerics 1972); Pannonian Basin: Szob, Pécsszaolcs, Ófalu (Hungary) (NHMW collection; Bohn-Havas 1973); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus (Hoernes & Auinger 1879; Boettger 1902);? Caransebeş-Mehadia Basin: Valea Bela Reca, Valea Calvei, (Romania) (Hinculov 1968). The illustrations in Hinculov (1968) and Csepreghy-Meznerics (1972) do not allow a clear identification and might represent another species as well. Proto-Mediterranean Sea. Tortonian (late Miocene): Po Basin: Stazzano, Montegibbio (Saco 1893 b).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE1FFF0FF5FA93AFD834418.taxon	description	[Chelyconus] [Conus] Schroeckingeri n. f. — Hoernes 1878 a: 195 (nomen nudum). Conus (Chelyconus) Schroeckingeri nov. form. — Hoernes & Auinger 1879: 51, pl. 1, fig. 19.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE1FFF0FF5FA93AFD834418.taxon	materials_examined	Type material. Holotype, NHMW 1854 / 0035 / 0040, Lăpugiu de Sus (Romania). Illustrated material. Figs 29 I 1 – I 4, 30 B: Holotype, NHMW 1854 / 0035 / 0040, height 23.7 mm, MD: 13.8 mm, Lăpugiu de Sus (Romania). Studied material. Holotype and 1 spec. NHMW A 1607, Lăpugiu de Sus (Romania). Revised description. Small pyriform shell with low conical to coeloconoid spire; early spire whorls angulated, tuberculate; later whorls flat, striate. Subsutural flexure very shallow, weakly curved, moderately asymmetrical. Suture impressed. Last whorl faintly angulated and convex below; constricted. Lower third of last whorl and base covered by wide-spaced, raised spiral cords. Moderately long, weakly reflected siphonal canal. Siphonal fasciole very weak. Aperture narrowing adapically and widening towards siphonal canal. Colour pattern under UV light consisting of regularly spaced, thin spirals on entire last whorl, coinciding with spiral cords on base. Spirals overlain by large, roughly trigonal blotches amalgamating into two irregular bands below shoulder and below midwhorl.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE1FFF0FF5FA93AFD834418.taxon	description	Shell ratios and measurements. n = 2 specimens: SL: 23.7 / 23.8 mm, MD: 13.8 / 13.4 mm, spire angle: 110 ° / 104 °), last whorl angle: 40 ° / 40 °, LW: 1.72 / 1.78, RD: 0.67 / 0.66, PMD: 0.83 / 0.80, RSH: µ = 0.14 / 0.15.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE1FFF0FF5FA93AFD834418.taxon	discussion	Discussion. Although known by just two specimens, this seems to be a distinct species. None of the other Paratethyan cone species develops such a stout pyriform shape with striate spire whorls and raised spiral cords on the base. This sculpture of wide-spaced, prominent spiral cords on the last whorl is reminiscent of Conus s. l. sturi (Hoernes & Auinger, 1879) but the strongly constricted base and the low conical coeloconoid spire exclude that P. schroeckingeri is just a juvenile C. sturi. Further, the colour pattern of delicate spirals overlain by large blotches is unknown from any other Paratethyan species. Paleoenvironment. No information.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE1FFF0FF5FA93AFD834418.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Transylvanian Basin: Lăpugiu de Sus (Romania) (Hoernes & Auinger 1879). The occurrence of this species in the Tortonian of Barcelona, mentioned by Faura I Sans (1908), needs confirmation.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FFE1FFF0FF5FADB3FEC4460E.taxon	materials_examined	Type species (by original designation): Conus elatus Michelotti, 1847. Miocene, Europe. Note. Tucker & Tenorio (2009) erected the genus Plagioconus, which they characterised as including species with an elongated last whorl, spire with a convex profile; the spire whorls devoid of sculpture and a shallow subsutural flexure. This is an exclusively European Miocene genus. If our interpretation of the genus is correct, the generic description must be emended. The shells have indeed an elongated last whorl, but the spire, which can be mediumhigh is always beaded on the earliest whorls and spiral sculpture on the ramp may range from finely to distinctly striate. Tucker & Tenorio (2009) described the anal notch as shallow, however, most species, including the type species, have a deep to very deep, moderately to strongly asymmetrical subsutural flexure, which is moderately to strongly curved.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1FFF0CFF5FA8E1FCD643C2.taxon	description	Conus (Leptoconus) Puschi Michti. — Hoernes & Auinger 1879: 34 (partim), pl. 5, fig. 7. [Conus] Chelyconus austriaconoe Sacc. — Sacco 1893 b: 85 [nov. nom. pro. Conus noe in Hörnes 1851, pl. 3, fig. 1].	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1FFF0CFF5FA8E1FCD643C2.taxon	materials_examined	Type material. Holotype: NHMW 1843 / 0032 / 0026, Baden (Austria), illustrated in Hörnes (1851, pl. 3, fig. 1). Studied material. Holotype and 2 spec. NHMW A 1612, 4 spec. NHMW 2013 / 0300 / 0526, Baden-Sooß (Austria), 1 spec. NHMW 1869 / 0001 / 0423, 1 spec. NHMW 1863 / 0015 / 0399, Forchtenau (Austria), 2 spec. NHMW 1868 / 0001 / 0382, Lăpugiu de Sus (Romania); middle Miocene, Badenian (Langhian). Illustrated material. Figs 30 F, 31 A 1 – A 3: Baden-Sooß (Austria): SL: 92.7 mm, MD: 35.7 mm, NHMW 2013 / 0300 / 0526; Figs 31 B 1 – B 3: Lăpugiu de Sus (Romania): SL: 76.6 mm, MD: 32.5 mm, NHMW 1868 / 0001 / 0382. Revised description. Large club-shaped shell, up to 94 mm in height with tall mammillate-cyrtoconoid spire. Earliest spire whorls strongly pointed; spire whorls broad, weakly convex but of bulgy appearance due to the deeply incised suture. Very faint spiral threads on early whorls; weak beads above suture, rarely preserved. Last whorl with broadly rounded shoulder; subsutural flexure deep, strongly curved, strongly asymmetrical. Position of maximum diameter some distance below upper suture; straight sided to weakly constricted in middle part, moderately constricted at base. Fasciole long, twisted and demarcated by deep and broad groove from narrow plait. Siphonal canal long, slightly recurved. Aperture narrow, widening anteriorly. Sculpture consisting of spiral threads on last whorl, strengthening on lower half; spiral threads interrupted by growth lines causing a wavy pattern. Colour pattern under UV light consisting of weak spirals of speckles on last whorl.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1FFF0CFF5FA8E1FCD643C2.taxon	description	Shell measurements and ratios. Only two shells are complete, all others lack the siphonal canal; for these species the ratios are: largest specimen: SL: 92.7 mm, MD: 35.7 mm, last whorl angle: µ = 23 / 28 °, LW: µ = 2.6 / 2.4, RD: µ = 0.47 / 0.52, PMD: µ = 0.91 / 0.89, RSH: µ = 0.19 / 0.18.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1FFF0CFF5FA8E1FCD643C2.taxon	discussion	Discussion. The specimens display some variability concerning spire height, but the conspicuous club-shaped outline and large size allow this species to be easily separated from other Paratethyan cones shells. Plagioconus austriaconoe is reminiscent of P. clavatulus (d’Orbigny, 1852), from the European early and middle Miocene, and is probably closely related (see Peyrot 1931; Hall 1966; Janssen 1984). Both develop convex spires with markedly pointed early spire and have broadly rounded shoulders. A separation is based on the higher spire, the bulgy spire whorls, the more elongate last whorl and the lower position of the maximum diameter of P. austriaconoe. Conus pseudopuschi Sacco, 1893 was based on a single strongly fragmented and slightly deformed specimen from offshore clays of the Vienna Basin. The spire is largely destroyed except for parts of the last three whorls. We consider this specimen to represent a P. austriaconoe with somewhat deeper concavity below the adsutural swelling. Therefore, Conus pseudopuschi is a subjective junior synonym of Plagioconus austriaconoe. Paleoenvironment. Plagioconus austriaconoe is a rare species, which was found so far only in offshore clays.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1FFF0CFF5FA8E1FCD643C2.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Baden (Austria) (Hoernes & Auinger 1879); Eisenstadt-Sopron Basin: Forchtenau (Austria) (Sieber 1956); Alpine-Carpathian Foredeep: Grund (Austria); Pannonian Basin: Letkés (Hungary) (Kovács & Vicián 2013); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus (Romania) (Boettger 1906; Chira & Voia 2001).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1DFF0DFF5FAA8AFDB040BA.taxon	description	Conus (Chelyconus) sp. — Caze et al. 2010: 35, fig. 5 / M 1 - M 2.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1DFF0DFF5FAA8AFDB040BA.taxon	materials_examined	Holotype: Figs 31 C 1 – C 3: SL: 70.8, MD: 27.3 mm, NHMW 1874 / 0025 / 0002 a, Lăpugiu de Sus (Romania). Paratype: Figs 31 D 1 – D 2: SL: 74.4, MD: 27.9 mm, NHMW 1874 / 0025 / 0002 b, Lăpugiu de Sus (Romania).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1DFF0DFF5FAA8AFDB040BA.taxon	description	Paratype: Figs 31 E 1 – E 2: SL: 66.9, MD: 25.0 mm, NHMW 1890 / 0001 / 0012 a, Lăpugiu de Sus (Romania). Paratype: Fig. 30 C: SL: 66.9, MD: 24.7 mm, NHMW 1890 / 0001 / 0012 b, Lăpugiu de Sus (Romania). Fig. Figs 31 F 1 – F 3: SL: 58.7, MD: 24.4 mm, Letkés (Hungary), private collection Anton Breitenberger (Bad Vöslau, Austria).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1DFF0DFF5FAA8AFDB040BA.taxon	materials_examined	Additional material: 2 spec. NHMW 1874 / 0025 / 0002; 2 spec. NHMW 1890 / 0001 / 0012; 1 spec., NHMW 1973 / 1615 / 0069; all Lăpugiu de Sus (Romania). Type stratum: Badenian marly-clayey deposits with thin interlayers of sand and corallinacean limestones. Type locality: Lăpugiu de Sus (Romania). Age: Middle Miocene, early Badenian (= Langhian).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1DFF0DFF5FAA8AFDB040BA.taxon	etymology	Etymology: Referring to the aesthetic appearance of the species, which was overlooked so far in the collections.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1DFF0DFF5FAA8AFDB040BA.taxon	description	Description. Moderately large, very elongate shells; spire medium-high conical, somewhat variable in height; early spire pointed, angulated just above suture; suture canaliculated. Later spire whorls feebly convex, glossy with faint spiral threads, not striate; suture deeply impressed. Subsutural flexure deep, moderately curved, strongly asymmetrical. Early spire whorls coarsely tuberculate at suture; tubercles usually covered by subsequent whorl. Last whorl convex, with rounded, sometimes bulgy shoulder coinciding with position of maximum diameter. Last whorl very elongate, conical, constricted at base. Siphonal canal very long, narrow, twisted, slightly reflected. Siphonal fasciole distinctly swollen, marked by densely spaced and prominent growth lines. Aperture narrow with subparallel margins, only weakly widening anteriorly. Wavy spiral grooves on base and lower quarter of last whorl may be developed. Colour pattern under UV light consisting of dense flammulae on spire and shoulder; last whorl with irregular pattern of densely spaced spirally arranged and amalgamating blotches, partly with vague axial arrangement. Two light (fluorescing) bands in the middle and lower third of the last whorl. Shell measurements and ratios. n = 8: largest specimen: SL: 74.4 mm, MD: 27.9 mm, mean SL: 68.1 mm (σ = 4.4), mean MD: 25.2 mm (σ = 2.2), spire angle: µ = 86 ° (σ = 8.23), last whorl angle: µ = 24.3 ° (σ = 0.9 °), LW: µ = 2.7 (σ = 0.07), RD: µ = 0.44 (σ = 0.01), PMD: µ = 0.95 (σ = 0.02), RSH: µ = 0.16 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1DFF0DFF5FAA8AFDB040BA.taxon	discussion	Discussion. This species was identified as Conus puschi in NHMW-collection lots and may also be mixed with Plagioconus puschi (Michelotti, 1847) in other museum collections. Both species, however, are readily distinguished by the higher spire, the broader and shorter siphonal canal and the less constricted base of P. puschi, which is also larger. In addition, P. puschi seems to have preferred shallow water environments, whereas P. bellissimus might rather represent an offshore species. Subadult specimens of Plagioconus extensus (Hörnes, 1851) are reminiscent of P. bellissimus due to the elongate shape and long siphonal canal but are distinguished by the sutural concavity and the prominent spiral cords. Plagioconus lapugyensis (Hoernes & Auinger, 1879) differs in its striate spire whorls, the less elongate last whorl, the broader shoulder and the shorter siphonal canal. Paleoenvironment. All specimens from Lăpugiu de Sus are filled with clay, probably suggesting offshore environments as preferred habitat.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1DFF0DFF5FAA8AFDB040BA.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Transylvanian Basin: Lăpugiu de Sus (Romania); Pannonian Basin: Letkés (own data).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1CFF0DFF5FAA52FB794664.taxon	materials_examined	Holotype: Figs 31 G 1 – G 5: SL: 43.6 mm, MD: 17.6 mm, NHMW 2016 / 0041 / 0001. Paratype: Figs 31 H 1 – H 2: SL: 43.9 mm, MD: 16.1 mm, NHMW 2016 / 0041 / 0002.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1CFF0DFF5FAA52FB794664.taxon	description	Paratype: SL: 34.7 mm, MD: 14.3 mm, NHMW 2016 / 0041 / 0003.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1CFF0DFF5FAA52FB794664.taxon	materials_examined	Additional material: 2 spec. NHMW 2016 / 0041 / 0004, Fig. 30 G: SL: 31.0 mm, MD: 12.3 mm, 1 spec. private collection Anton Breitenberger; all specimens from Letkés, Hungary. Type stratum: fossil-rich marly sand with coral blocks and andesite boulders of the Sámsonháza Formation (Császár 1997). Type locality: Letkés at the western part of the Börzsöny Mts. (Hungary); see Kovács & Vicián (2013) for a map and description. Age: Middle Miocene, early Badenian (= Langhian).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1CFF0DFF5FAA52FB794664.taxon	etymology	Etymology: Referring to Anton Breitenberger, who collected and donated the holotype.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1CFF0DFF5FAA52FB794664.taxon	description	Description. Medium-sized, slender elongate shells; spire mammillate, dome-shaped, high. Early spire whorls angulate just above suture, tuberculate. Tubercles become obsolete within 3 rd – 4 th teleoconch whorl; angulation migrates to suture during ontogeny. Spire whorls weakly convex, faintly striate; suture narrowly impressed. Subsutural flexure very deep, moderately curved, strongly asymmetrical. Last whorl angulated; very slender, elongate, not constricted. Aperture narrow and only weakly widening anteriorly; siphonal canal long, straight to faintly recurved. Siphonal fasciole indistinct, narrow. Weak spiral cords on lower third of last whorl. Colour pattern in UV light consisting of broad axial stripes and flammulae on spire and shoulder. A light band below shoulder angulation follows in upper quarter of last whorl. Below appear broad, stretched zig-zag shaped axial streaks crossed mid-whorl by a spiral band; base and siphonal canal dark. Shell measurements and ratios. n = 4: largest specimen: SL: 43.9 mm, MD: 17.6 mm, mean SL: 38.3 mm (σ = 6.5), mean MD: 15.1 mm (σ = 2.3), spire angle: µ = 84 ° (σ = 2.7 °), last whorl angle: µ = 26 ° (σ = 1.4 °), LW: µ = 2.54 (σ = 0.14), RD: µ = 0.50 (σ = 0.02), PMD: µ = 0.89 (σ = 0.02), RSH: µ = 0.21 (σ = 0.01).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1CFF0DFF5FAA52FB794664.taxon	discussion	Discussion. This species is not rare at Letkés but was probably mistaken as subadult Plagioconus marii (Sacco, 1893) by Kovács & Vicián (2013). It differs from Plagioconus marii in its dome-shaped spire, the lower height of the spire whorls, the less impressed suture and the smaller size. Conus s. l. mucronatolaevis Sacco (1893) and especially its globospira - morphotype develop a comparable spire but are broader and the last whorl is less elongate. The much deeper subsutural flexure allows a clear separation from Plagioconus puschi (Michelotti, 1847). Paleoenvironment. The locality Letkés comprises a mixed assemblage of partly reworked taxa (Kovács & Vicián 2013); the frequent occurrence of corals suggests a shallow marine depositional environment.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1CFF0DFF5FAA52FB794664.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Pannonian Basin: Letkés (Hungary).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1CFF0AFF5FAF17FE76462B.taxon	materials_examined	Type material. Syntype or holotype illustrated by Michelotti (1847, pl. 13, fig. 16), Tortona, Italy; the specimen might have been stored in the collections of the Dipartimento di Scienze della Terra, Sapienza Università di Roma but parts of the Michelotti collection were destroyed during WWII (Manni 2005); type specimen is lost according to Hall (1966); late Miocene, Tortonian. Studied material. 1 spec. NHMW 1997 z 0178 / 1471, 1 spec. NHMW 1856 / 0050 / 0114, 4 spec. NHMW 1855 / 0045 / 0367, 1 spec. NHMW 1997 z 0178 / 1473, Gainfarn (Austria); 1 spec. NHMW 1846 / 0037 / 0023, Grinzing (Austria). Illustrated material. Figs 32 A 1 – A 3: Gainfarn (Austria): SL: 91.6 mm, MD: 38.4 mm, NHMW 1855 / 0045 / 0367; Figs 30 H, 32 B 1 – B 2: Grinzing (Austria): SL: 103.8 mm, MD: 37.3 mm, NHMW 1846 / 0037 / 0023. Revised description. Large shells; spire short with coeloconoid outline. Teleoconch comprising at least 12 whorls; early spire whorls beaded, rounded or with weak angulation in the middle, migrating towards lower suture on last whorls. Suture deeply incised, emphasized by weak subsutural inflation; delicate spiral threads appear on sutural ramp. Subsutural flexure deep, moderately curved, strongly asymmetrical. Last whorl slightly allometric in growth due to rapid widening of last two whorls. Sutural ramp relatively shallow and wide, distinctly shouldered at periphery, whorl straight below, hardly constricted at base. Aperture narrow; siphonal canal of moderate length and width; not recurved or twisted. Narrow, flattened inner lip demarcated from short fasciole by distinct notch. No spiral grooves on base. No colour pattern is preserved.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1CFF0AFF5FAF17FE76462B.taxon	description	Shell measurements and ratios. n = 8 adult and subadult specimens: largest specimen: SL: 103.8 mm, MD: 39 mm, mean SL: 87.9 mm (σ = 8.1), mean MD: 35.4 mm (σ = 2.6), spire angle: µ = 82.4 ° (σ = 3.3 °), last whorl angle: µ = 24.5 ° (σ = 1.2 °), LW: µ = 2.48 (σ = 0.15), RD: µ = 0.50 (σ = 0.03), PMD: µ = 0.96 (σ = 0.01), RSH: µ = 0.19 (σ = 0.02).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1CFF0AFF5FAF17FE76462B.taxon	discussion	Discussion. This species was frequently treated as Conus elongatus Borson, 1820 (e. g. Hall 1966; Davoli 1972; Bałuk 2006). This name, however, is preoccupied by C. elongatus Holten, 1802 and consequently C. elatus Michelotti, 1847 is the next available name. In the Paratethys, it was identified as Conus haueri Hörnes, 1851, which is a problematic species. The syntype illustrated by Hörnes (1851, pl. 4, fig. 4) is lost and the spire whorls of the illustration display a weak concavity as typical for Monteiroconus antiquus. The syntype illustrated by Hörnes (1851, pl. 4, fig. 5) is still preserved in the NHMW collection and is a specimen of Plagioconus puschi. The syntype of Conus haueri from Grinzing mentioned by Hörnes (1851, not illustrated) and all other shells identified as C. haueri in the NHMW collection agree fully with Plagioconus elatus (Michelotti, 1847) as understood by Hall (1966). Therefore, we consider Conus haueri Hörnes, 1851 a nomen dubium. Paleoenvironment. All Austrian specimens were found in assemblages from shallow sublittoral settings (e. g. Zuschin et al. 2007).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF1CFF0AFF5FAF17FE76462B.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn, Grinzing (Austria); Pannonian Basin: Szob (Hungary) (Hoernes & Auinger 1879); Carpathian Foredeep: Korytnica (Poland) (Bałuk 2006); Dacian Basin: Opanec (Bulgaria) (Kojumdgieva & Strachimirov 1960). Proto-Mediterranean Sea and northeastern Atlantic. Burdigalian (Italy: Colli Torinesi), Tortonian (Italy: Sant'Agata Fossili, Stazzano Montegibbio). A Pliocene occurrence from Borzoli (Italy) was mentioned by Sacco (1893 a) but needs confirmation.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF19FF09FF5FA8E1FC75400F.taxon	description	[Conus] extensus n. sp. — Hauer 1837: 416 (nomen nudum).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF19FF09FF5FA8E1FC75400F.taxon	materials_examined	Type material. 2 syntypes NHMW 1847 / 0037 / 0031, Baden (Austria), including specimen illustrated in Hörnes (1851, pl. 5, fig. 1), middle Miocene, Badenian (late Langhian). Studied material. Syntypes and 1 spec. NHMW 1869 / 0001 / 0159, 1 spec. NHMW 1997 z 0178 / 0871, Badenian Baden (Austria); 2 spec. NHMW 2015 / 0391 / 0001 including specimen illustrated in Schultz (1998, pl. 29, fig. 13), 3 spec. NHMW 1872 / 0030 / 0017, 10 spec. NHMW 2013 / 0300 / 0525, all Baden-Sooß (Austria); 1 spec. NHMW 1855 / 0045 / 0370, Gainfarn (Austria), 3 spec. NHMW 1854 / 00035 / 0053, 3 spec. NHMW 1854 / 0035 / 0052, 2 spec. NHMW 1855 / 0043 / 0003, 2 spec. NHMW 1859 / 0037 / 0001, 1 spec. NHMW 1868 / 0001 / 0384, 2 spec. NHMW 1866 / 0040 / 0247, 4 spec. NHMW 1875 / 0009 / 0005, 1 spec. NHMW 1875 / 0009 / 0004, 2 spec. NHMW 1890 / 0001 / 0 0 11, 2 spec. NHMW 1890 / 0001 / 0013, 2 spec. NHMW 1973 / 1615 / 0062, 3 spec. NHMW 2015 / 0392 / 0001, 8 spec. NHMW 2015 / 0392 / 0002, 2 spec. NHMW 1858 / 0043 / 0004, all Lăpugiu de Sus (Romania), 1 spec. NHMW 1867 / 0019 / 0010, Coşteiu de Sus (Romania): 1 spec. NHMW 2015 / 0393 / 0001, Nemeşeşti (Romania). Illustrated material. Figs 32 C 1 – C 3: Lăpugiu de Sus (Romania): SL: 89.3 mm, MD: 35.3 mm, NHMW 1890 / 0001 / 0011; Figs 32 D: Lăpugiu de Sus (Romania), SL: 89.0, MD: 36.0 mm, NHMW 1859 / 0037 / 0001; Figs 30 K, 32 E 1 – E 3: Baden-Sooß (Austria): SL: 96.0 mm, MD: 33.2 mm, NHMW 1872 / 0030 / 0017. Revised description. Large shells comprising at least 10 tall teleoconch whorls; spire short, conical to weakly coeloconoid. Early whorls angular above suture with coarse beads, central angulation becoming mid-whorl inflation abapically. Sutural ramp bearing subsutural collar / band and spiral striae developed to a variable degree; ramp becoming increasingly concave abapically. Suture deeply impressed. Subsutural flexure very deep, moderately curved, strongly asymmetrical. Last whorl very elongate, weakly constricted at base; moderately round shoulder with position of maximum diameter a short distance below shoulder. Aperture narrow, widening abapically. Siphonal canal long, weakly recurved; fasciole elongate, slightly twisted and demarcated from narrow glossy inner lip by a narrow notch. Base covered by very delicate spiral grooves on adapical half; grooves become stronger and wider abapically; usually somewhat irregular and pitted. Colour pattern in UV light consisting of light flammulae on spire whorls and shoulder of last whorl parallel to subsutural flexure; indistinct bands on last whorl.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF19FF09FF5FA8E1FC75400F.taxon	description	Shell measurements and ratios. n = 12 adult and subadult specimens: largest specimen: SL: 107 mm, MD: 39 mm, mean SL: 88.2 mm (σ = 10.4), mean MD: 32 mm (σ = 3.7), spire angle: µ = 82.7 ° (σ = 4.3 °), last whorl angle: µ = 22.3 ° (σ = 1.7 °), LW: µ = 2.76 (σ = 0.19), RD: µ = 0.44 (σ = 0.03), PMD: µ = 0.95 (σ = 0.02), RSH: µ = 0.17 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF19FF09FF5FA8E1FC75400F.taxon	discussion	Discussion. This species is characterized by the concave sutural ramp bearing spiral striae, the distinct but rounded shoulder and rather moderate variability in outline. The largest variability is represented by the more or less prominent spiral sculpture. Hörnes (1851) had only a single complete specimen at hand, which has been largely destroyed thereafter; only the spire is preserved. Since then, numerous shells were collected at Baden, Baden-Sooß and Lăpugiu de Sus. Paleoenvironment. The majority of shells were collected in basinal clays. In contrast, only a single specimen was found in the seagrass environment of Gainfarn (Austria). This suggests that Plagioconus extensus was probably an offshore species.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF19FF09FF5FA8E1FC75400F.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Alpine-Carpathian Foredeep: Grund (Austria) (Sieber 1947); Vienna Basin: Baden, Baden-Sooß, Gainfarn, Steinebrunn (Austria) (Hoernes & Auinger 1879; Sieber 1958 b); Lavanttal Basin: Mühldorf (Austria) (Wank 1981); Pannonian Basin: Hont, Letkés, Szob (Hungary) (Kovács & Vicián 2013); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus, Nemeşeşti (Romania) (Boettger 1902, 1906; Chira & Voia 2001); Dacian Basin: Opanec (Bulgaria) (Kojumdgieva & Strachimirov 1960). No reliable occurrences from the proto-Mediterranean Sea are known to date.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF18FF09FF5FA9C1FEB947DC.taxon	description	2013 Leptoconus hirmetzli sp. n. — Kovács & Vicián, 62, figs 30 – 37.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF18FF09FF5FA9C1FEB947DC.taxon	materials_examined	Type material. Holotype HNHM PAL 2013.3.1 (Hungarian Natural History Museum), paratypes HNHM PAL 2013.4. 1, HNHM PAL 2013.5.1 – PAL 2013.9. 1 (Hungarian Natural History Museum) and private collection of Tamás Hirmetzl (Hungary), all from Letkés (Hungary); middle Miocene, Badenian (Langhian). Illustrated material. Fig. 32 H: paratype, SL: 130 mm, MD: 34 mm, private collection of T. Hirmetzl; Fig. 32 I: paratype, SL: 67 mm, MD: 17 mm, private collection of T. Hirmetzl, Fig. 32 J: holotype, SL: 62 mm, MD: 21 mm, HNHM, PAL 2013.3.1; Fig. 30 E: SL: 114 mm, MD: 32.3 mm, private collection Anton Breitenberger (Bad Vöslau, Austria); all Letkés (Hungary). Revised description. Large, elongate, slender, biconical shell with very high spire and strongly elongate last whorl; protoconch paucispiral. Early spire whorls weakly angulate, tuberculate; later spire whorls high and convex. Suture deeply incised; subsutural flexure very deep, moderately curved, strongly asymmetrical. Last whorl with rounded to subangulate shoulder; position of maximum diameter slightly below shoulder; straight sided to faintly concave, not constricted at base; smooth surface. Aperture very narrow with subparallel margins. Fasciole very weak, twisted; siphonal canal slightly reflected and long. No colour pattern preserved.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF18FF09FF5FA9C1FEB947DC.taxon	description	Shell measurements and ratios. Holotype: height (fragmentary): 62 mm, MD: 21 mm, spire angle: 31 °, last whorl angle: 19 °; paratype 1: height (fragmentary): 132 mm, MD: 42 mm, spire angle: 38 °, last whorl angle: 18 °; paratype 2: height (fragmentary): 130 mm, MD: 34 mm, spire angle: 35 °, last whorl angle: 21 °.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF18FF09FF5FA9C1FEB947DC.taxon	discussion	Discussion. This is a very rare species, which is only known so far from its early Badenian type locality Letkés in Hungary. Due to the peculiar morphology, Kovács & Vicián (2013) placed the species in Leptoconus Swainson, 1840. They referred to the comparably elongate and slender Leptoconus milneedwardsi (Jousseaume, 1894), which is an extant species in the Indo-West Pacific Region, and to L. aratispira (Pilsbry, 1905), from Pleistocene of Japan. Aside from these two extremely elongate species, Leptoconus species, like the extant type species Leptoconus amadis (Gmelin, 1791), have low conical spires. All develop angulated whorls, tend to have concave adapical parts of the spire whorls and many species have sculptured last whorls. The high and convex spire whorls of Plagioconus hirmetzli, the deep suture and smooth shell surface do not support the placement in Leptoconus. Therefore, we consider the similarity between the Miocene species with the extant L. milneedwardsi to represent a striking case of convergent evolution and consider it an extraordinarily high spired Plagioconus species comparable with Plagioconus marii (Sacco, 1893). Paleoenvironment. The locality Letkés comprises a mixed assemblage of partly reworked taxa (Kovács & Vicián 2013); the frequent occurrence of corals suggests a shallow marine depositional environment.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF18FF09FF5FA9C1FEB947DC.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene). Pannonian Basin: Letkés (Hungary) (Kovács & Vicián 2013).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF18FF06FF5FAF7FFCA0445C.taxon	description	Conus (Chelyconus) Lapugyensis nov. form. — Hoernes & Auinger 1879: 42, pl. 1, fig. 9, pl. 5, fig. 8.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF18FF06FF5FAF7FFCA0445C.taxon	materials_examined	Type material. Syntype NHMW 1870 / 0033 / 0006 a, Lăpugiu de Sus (Romania), illustrated in Hoernes & Auinger (1879, pl. 1, fig. 9), syntype NHMW 1867 / 0019 / 0004, Coşteiu de Sus (Romania), illustrated in Hoernes & Auinger (1879, pl. 5, fig. 8). 7 syntypes NHMW 1867 / 0019 / 0005, Coşteiu de Sus (Romania); 4 syntypes NHMW 1870 / 0033 / 0006, Lăpugiu de Sus (Romania); middle Miocene, Badenian (Langhian). Studied material. Syntypes. Illustrated material. Figs 30 D, 33 D 1 – D 3: syntype, Lăpugiu de Sus (Romania), SL: 60.6 mm, MD: 28.2 mm, NHMW 1870 / 0033 / 0006 a, illustrated in Hoernes & Auinger (1879, pl. 1, fig. 9); Figs 33 E 1 – E 3: syntype, Coşteiu de Sus (Romania), SL: 44.6 mm, MD: 22.0 mm, NHMW 1867 / 0019 / 0005. Revised description. Medium-sized to moderately large, not very solid, elongate pyriform shells with mammillate spire and pointed apex with weakly coeloconoid initial part. Early spire whorls carinate, coarsely beaded; suture deeply canaliculated, undulating; later spire whorls convex with deep suture; weakly striate. Subsutural flexure of medium depth, moderately curved, strongly asymmetrical. Last spire whorl slightly broader and inflated, forming a strongly convex shoulder coinciding with position of maximum diameter. Last whorl constricted mid-whorl passing into moderately narrow, long, weakly recurved canal. Distinct spiral grooves on lower half of last whorl demarcating flat to weakly convex cords. Aperture narrow; fasciole weak, slightly twisted, usually with 2 – 3 strongly raised growth lines from previous siphonal canal margins. Colour pattern under UV light consisting of flammulae on shoulder reaching down on upper part of last whorl; below on last whorl follow spirally arranged short dashes and speckles, arranged into discontinuous bands and into irregular axial stripes.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF18FF06FF5FAF7FFCA0445C.taxon	description	Shell measurements and ratios. n = 10: largest specimen: SL: 60.6 mm, MD: 28.2 mm, mean SL: 44.8 mm (σ = 8.2), mean MD: 21.4 mm (σ = 3.2), spire angle: µ = 98.2 ° (σ = 5.1 °), last whorl angle: µ = 34.3 ° (σ = 1.9 °), LW: µ = 2.1 (σ = 0.15), RD: µ = 0.6 (σ = 0.03), PMD: µ = 0.89 (σ = 0.03), RSH: µ = 0.17 (σ = 0.05).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF18FF06FF5FAF7FFCA0445C.taxon	discussion	Discussion. Despite some superficial similarities, this rare species differs from Leporiconus suessi (Hoernes & Auinger, 1879) in its shorter and more dome-shaped spire, the canaliculated suture, the broad, inflated last spire whorl, the constricted base, the pyriform outline and the prominent spiral cords on the base. It can generally be distinguished from Paratethyan Leporiconus species by its strongly asymmetrical subsutural flexure. This species was synonymized with Lautoconus pelagicus (Brocchi, 1814) by Hall (1966) from which it is distinguished by its spire shape, the canaliculated suture and elongate pyriform outline. In addition, the colour pattern of L. pelagicus, as described by Hall (1966) and Landau et al. (2013) differs from Plagioconus lapugyensis. Conus sturi Hoernes & Auinger, 1879 and Leporiconus transsylvanicus (Hoernes & Auinger, 1879) differ in their well defined tubercles on early spire whorls. Paleoenvironment. Unknown; the fine grained sediment infill might indicate offshore habitats.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF18FF06FF5FAF7FFCA0445C.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus (Romania) (Hoernes & Auinger 1879; Boettger 1902).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF17FF04FF5FADFFFEAD4478.taxon	description	[Conus] Chelyconus Marii Sacc. — Sacco 1893 b: 62, pl. 6, fig. 1. Conus marii (Sacco), 1893 — Hall 1966: 148, pl. 26, figs 21 – 22 [? figs 8, 15, 16]. Chelyconus marii Sacco, 1893 — Ferrero-Mortara 1984: 114, pl. 17, figs 8 a – b.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF17FF04FF5FADFFFEAD4478.taxon	materials_examined	Type material. Lectotype BS. 038.05.020 in Museo Regionale di Scienze Naturali, Torino; Albugnano (Italy), middle Miocene, illustrated in Sacco (1893 b, pl. 6, fig. 1); designated herein. Studied material. 2 spec. NHMW 1846 / 0037 / 0025, 1 spec. NHMW 1851 / 0013 / 0019, Vöslau (Austria); 1 spec. NHMW 1874 / 0024 / 0001, 1 spec. NHMW 1869 / 0001 / 0160, Baden (Austria); 5 spec NHMW 2013 / 0300 / 0 529 (4 spire fragments) Baden-Soo (Austria). Illustrated material. Figs 30 I, 32 F 1 – F 3: Baden-Soo (Austria): SL: 67.0 mm, MD: 21.0 mm, NHMW 2013 / 0300 / 0529; Figs 32 G 1 – G 3: Vöslau (Austria): SL: 67.8 mm, MD: 23.4 mm, NHMW 1846 / 0037 / 0025. Revised description. Moderately large shell with maximum height of nearly 70 mm; elongate outline with elevated conical spire comprising at least 13 whorls; spire height varies considerably. Early spire whorls angulated and beaded. Later spire whorls subangular without beads, with faint spiral grooves strongest along the angulation and below. Deeply incised suture; subsutural flexure very deep, moderately curved, strongly asymmetrical. Last whorl with strongly rounded shoulder and position of maximum diameter close below shoulder; straight sided with very slight constriction at base. Aperture narrow, narrowing adapically. Inner lip with callous notch and weak parietal callus. Siphonal canal moderately long, straight; fasciole indistinct, slightly twisted. Last whorl with narrow, finely wavy spiral grooves over lower half of last whorl.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF17FF04FF5FADFFFEAD4478.taxon	description	Shell measurements and ratios. n = 4 adult and subadult specimens: largest specimen: SL: 67.8 mm, MD: 23.4 mm, mean SL: 64 mm (σ = 5.4), mean MD: 21.4 mm (σ = 1.5), spire angle: µ = 55.3 ° (σ = 3.8 °), last whorl angle: µ = 24.8 ° (σ = 1.0 °), LW: µ = 2.99 (σ = 0.16), RD: µ = 0.47 (σ = 0.01), PMD: µ = 0.93 (σ = 0.01), RSH: µ = 0.29 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF17FF04FF5FADFFFEAD4478.taxon	discussion	Discussion. The Paratethyan shells are very well preserved and show even the delicate beads on the early spire and the spiral sculpture on spire whorls and last whorl. These features are not preserved in the Italian type specimens but the size and general shape agree well. Unfortunately, Sacco (1893 b) based this species on occurrences from different early and middle Miocene localities in the Turin Hills. These specimens represent syntypes and therefore, the designation of a holotype as proposed by Hall (1966) is inappropriate. To clarify this situation. the specimen from the middle Miocene of Albugnano illustrated by Sacco (1893 b, pl. 6, fig. 1) is designated herein as the lectotype (inventory number BS. 038.05.020 according to Ferrero-Mortara et al. 1984). In the NHMW collections this species was mixed with P. puschi due to the superficial similarity in outline. Its rareness in the Paratethys might thus rather be an artefact. It differs from P. puschi and P. elatus in the shorter last whorl, the comparatively higher spire and higher spire whorls, the deep subsutural flexure and the smaller size. Moreover, the occurrences in coastal shallow water settings (P. puschi, P. elatus) versus offshore clays (P. marii) suggest an ecological and bathymetric separation. Paleoenvironment. The species is only known from offshore clays of the inner to outer shelf.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF17FF04FF5FADFFFEAD4478.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Bad Vöslau, Baden, Baden-Sooß (Austria) (own data); Pannonian Basin: Letkés (Hungary) (Kovács & Vicián 2013); Transylvanian Basin: Lăpugiu de Sus (Romania) (Chira & Voia 2001). Proto-Mediterranean Sea and northeastern Atlantic. Burdigalian (Italy: Colli Torinesi), Langhian (Italy: Albugnano).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF15FF02FF5FAD13FC00402A.taxon	description	Conus Puschi mihi — Michelotti 1847: 340, pl. 14, fig. 6. Conus Puschi Micht. — Hörnes 1851: 35, pl. 4, figs 6 – 7.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF15FF02FF5FAD13FC00402A.taxon	description	[Conus (Chelyconus)] Puschi] var. postica Sacc. — Sacco 1893 b: 60 [nov. nom. pro Conus Puschi in Hörnes 1851, pl. 4, fig. 6]. [Conus (Chelyconus)] Puschi] var. sulcopostica Sacc. — Sacco 1893 b: 60 [nov. nom. pro Conus Puschi in Hörnes 1851, pl. 4, fig. 7, erroneously given as fig. 5 by Sacco 1893]. Conus (Chelyconus) puschi Micht. — Csepreghy-Meznerics 1956: 419, pl. 10, figs 3 – 4.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF15FF02FF5FAD13FC00402A.taxon	materials_examined	Type material. Syntype illustrated by Michelotti (1847, pl. 14, fig. 6), Tortona, Italy; the specimen might have been stored in the collections of the Dipartimento di Scienze della Terra, Sapienza Università di Roma but parts of the Michelotti collection were destroyed during WWII (Manni 2005); type specimen is lost according to Hall (1966); late Miocene, Tortonian. The type locality Tortona was questioned by Sacco (1893 b). In addition to the Italian specimen (s), Michelotti (1847) referred also to material from the Burdigalian of Bordeaux (France). Studied material. 1 spec. NHMW 2013 / 0479 / 1615, 4 spec. NHMW 2013 / 0479 / 1616, 4 spec. NHMW 2013 / 0479 / 2042, 5 spec. NHMW 1855 / 0045 / 0368, all Gainfarn (Austria); 2 spec. NHMW 1851 / 0013 / 0062, illustrated in Hörnes (1851, pl. 4, fig. 7), 3 spec. NHMW 1860 / 0001 / 0062; 4 spec. NHMW 1846 / 0037 / 0026, 2 spec. NHMW 1884 / 2869, 1 spec. GBA 1856 / 004 / 0004 / 01 illustrated in Hörnes (1851, pl. 4, fig. 6), all Steinebrunn (Austria); 1 spec. NHMW 1846 / 0037 / 0024, Grinzing (Austria), illustrated in Hörnes (1851, pl. 4, fig. 5). Illustrated material. Fig. 30 J: Steinebrunn (Austria), SL: 60.6 mm, MD: 22.1 mm, NHMW 1846 / 0037 / 0026; Figs 33 A 1 – A 3: Gainfarn (Austria): SL: 98.7 mm, MD: 37.8 mm, NHMW 2013 / 0479 / 1615; Figs 33 B 1 – B 3: Gainfarn (Austria): SL: 78.6 mm, MD: 27.9 mm, NHMW 1860 / 0001 / 0062; Figs 33 C 1 – C 3: Gainfarn (Austria): SL: 66.20 mm, MD: 24.7 mm, NHMW 1851 / 0013 / 0062, illustrated in Hörnes (1851, pl. 4, fig. 7). Revised description. Moderately large to large shell, up to ca. 100 mm in height; elongate slender outline with moderately elevated conical spire. Early spire whorls weakly angulated and beaded, later whorls convex with deep suture. Subsutural flexure of medium depth, strongly curved, strongly asymmetrical. Shoulder broadly rounded, position of maximum diameter slightly below shoulder; last whorl straight sided, not constricted at base. Spiral threads on late spire whorls and along shoulder of last whorl; spiral grooves may appear on lower third of last whorl, becoming stronger and closely spaced abapically. Aperture narrow, slightly narrowing adapically; siphonal canal almost straight, fasciole weak. No colour pattern preserved.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF15FF02FF5FAD13FC00402A.taxon	description	Shell measurements and ratios. n = 17 adult and subadult specimens: largest specimen: SL: 98.7 mm, MD: 37.8 mm, mean SL: 76.0 mm (σ = 9.4), mean MD: 28.8 mm (σ = 3.8), spire angle: µ = 75.1 ° (σ = 5.4 °), last whorl angle: µ = 22.2 ° (σ = 1.8 °), LW: µ = 2.64 (σ = 0.15), RD: µ = 0.48 (σ = 0.02), PMD: µ = 0.95 (σ = 0.02), RSH: µ = 0.21 (σ = 0.02).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF15FF02FF5FAD13FC00402A.taxon	discussion	Discussion. Plagioconus puschi is reminiscent of subadult P. elatus (Michelotti, 1847), which lack the prominent shoulder of adult specimens. Both species can be distinguished based on the nearly diagonal and shallow subsutural flexure of P. puschi and the angulation of the spire whorls of P. elatus. Moreover, fully grown specimens of P. elatus are always distinctly larger. Sacco (1893 b) doubted that the specimens from the Vienna Basin are conspecific with the Italian P. puschi and proposed new names for each of the two shells illustrated by Hörnes (1851): Conus postica and C. sulcopostica. Both shells derive from Steinebrunn in the northern Vienna Basin and represent a subadult and a fully grown specimen. Therefore, Conus postica Sacco, 1893 and Conus sulcopostica Sacco, 1893 are considered subjective junior synonyms of Conus puschi Michelotti, 1847. The syntype of Conus haueri Hörnes, 1851 (pl. 4, fig. 5) is also Plagioconus puschi — a fact that was already recognized by Hall (1966). Therefore, Conus haueriana Sacco, 1893, which was introduced as new name for that specimen, is also a subjective junior synonym of Conus puschi Michelotti, 1847. The specimen from Bosnia, illustrated by Atanacković (1963, 1985, same specimen in both papers) has a marked angulation of the last whorl and might rather be an aberrant Plagioconus extensus (Hörnes, 1851). The shells illustrated by Hall (1966) as Conus puschi differ in their dome-shaped spire and represent a different species. Similarly, the specimens illustrated by Davoli (1972) differ in their high conical spire and might belong to another species. Paleoenvironment. Typically found in shallow water assemblages; for some localities sea grass meadows are documented (e. g. Gainfarn, Zuschin et al. 2007).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF15FF02FF5FAD13FC00402A.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn, Steinebrunn, Bad Vöslau, Baden, Niederleis (Austria), Mikulov-Kienberk, Hrušovany (Czech Republic) (Hörnes 1851); Alpine-Carpathian Foredeep: Grund (Austria), Lysice (Czech Republic) (Hörnes 1851; Sieber 1956); Eisenstadt-Sopron Basin: Forchtenau (Austria); Pannonian Basin: Devecser, Diósd, Letkés, Magyaregregy, Szob, Zebegény (Hungary) (Kovács & Vicián 2013); southern Pannonian Basin: Zapešić-Brijeg at Samobor (Croatia); Banja Luka Basin: Miljevići (Bosnia and Herzegovina) (Pavlovsky 1957), Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus (Romania) (Boettger 1902, 1906); Caransebeş-Mehadia Basin: Valea Satului (Romania) (Hinculov 1968). Proto-Mediterranean Sea and northeastern Atlantic. Widespread during the early Miocene to late Miocene in the northeastern Atlantic and the Proto-Mediterranean Sea; the species persists into the Pliocene in the Mediterranean Sea (see Landau et al. 2013 for detailed references).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF13FF02FF5FA9E2FF7C4258.taxon	materials_examined	Type species (by original designation): Conus carnalis Sowerby III, 1879. Recent. Western Africa. According to Tucker & Tenorio (2009) this genus is characterised by shells with false nodules, formed by elevations at the junction of the shoulder and major growth lines. Subsutural flexure shallow, moderately curved, strongly asymmetrical. Based on the species included in the genus by Tucker & Tenorio (2009) the shells have spiral cords or faint spiral striae on the spire whorls, which are more or less concave. Aside from the spire sculpture, the species are reminiscent of Monteiroconus species. The genus is restricted to Western Africa; Miocene and Pliocene species are documented from the proto-Mediterranean Sea and the Eastern Atlantic including the Azores. This genus was not treated by Puillandre et al. (2014 b), but accepted by Puillandre et al. (2014 a).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF13FF03FF5FABF3FAA84236.taxon	description	[Dendroconus] [Conus] austriacus n. f. — Hoernes 1878 a: 195 (nomen nudum). [Dendroconus] [Conus] Reussii n. f. — Hoernes 1878 a: 195 (nomen nudum). Conus (Dendroconus) austriacus nov. form. — Hoernes & Auinger 1879: 19, pl. 2, figs 2 – 3. Conus Dendroconus Reussi nov. form. — Hoernes & Auinger 1879: 20, pl. 2, fig. 1.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF13FF03FF5FABF3FAA84236.taxon	materials_examined	Type material. Syntype NHMW 1875 / 0009 / 0001, Lăpugiu de Sus (Romania), specimen illustrated in Hoernes & Auinger (1879, pl. 2, fig. 2); syntype NHMW 1866 / 0011 / 0106, Steinebrunn (Austria), specimen illustrated in Hoernes & Auinger (1879, pl. 2, fig. 3); syntype NHMW 1853 / 0003 / 0002, Gainfarn (Austria); middle Miocene, Badenian (Langhian). Studied material. Syntypes and 1 spec. NHMW 1847 / 0037 / 0030, Steinebrunn (Austria), holotype of Conus reussi Hoernes & Auinger, 1879. Illustrated material. Figs 33 G 1 – G 2: syntype, Lăpugiu de Sus (Romania): SL: 73.8 mm, MD: 46.3 mm, NHMW 1875 / 0009 / 0001, specimen illustrated in Hoernes & Auinger (1879, pl. 2, fig. 2); Figs 30 P, 33 H 1 – H 4: syntype, Steinebrunn (Austria): SL: 112.0 mm, MD: 62.8 mm, NHMW 1866 / 0011 / 0106, illustrated in Hoernes & Auinger (1879, pl. 2, fig. 3). Revised description. Moderately large to large pyriform shells of up to 112 mm height. Spire ranging from flat to very low conical; early spire whorls with granulose spiral threads on adapical half; tuberculate along lower suture. Later whorls striate with weak concavity in adapical half and convex abapical part developing large, wide, blurred nodes; suture undulose. Subsutural flexure shallow, moderately curved, strongly asymmetrical. Shoulder angulated; position of maximum diameter distinctly below shoulder passing into relatively elongate last whorl; markedly constricted at base with long, moderately wide, nearly straight siphonal canal. Siphonal fasciole indistinct with prominent growth lines; very weak spiral cords on base; faint spiral cords on last whorl, visible only in tangential light. Colour pattern under UV light only seen on spire whorls, consisting of broad flammulae.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF13FF03FF5FABF3FAA84236.taxon	description	Shell measurements and ratios. Only three shells are more or less completely preserved: largest specimen: SL: 112.0 mm, MD: 62.8 mm, mean SL: 92.6 mm (σ = 19.1), mean MD: 54.0 mm (σ = 8.3), spire angle: µ = 142 ° (σ = 4.4), last whorl angle: µ = 39.3 ° (σ = 2.1), LW: µ = 1.7 (σ = 0.1), RD: µ = 0.64 (σ = 0.04), PMD: µ = 0.85 (σ = 0.02), RSH: µ = 0.09 (σ = 0.04).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF13FF03FF5FABF3FAA84236.taxon	discussion	Discussion. This rare species is recognized easily by its pyriform shape and the undulose suture around the broad nodes of the spire whorls and the tuberculate early spire whorls. These features separate it well from Monteiroconus. We place this species in Pseudonoduloconus based on the low spire, striate spire whorls, tuberculate early spire whorls and the undulose suture. Hoernes & Auinger (1879) already discussed if their Conus reussi might simply represent a very large C. austriacus but separated it based on its larger size and because of the less distinct nodes on the spire. Both morphs were collected at the same locality (Steinebrunn); the single specimen of C. reussi has a worn spire and therefore the sculpture is partly abraded. Aside from the size, there is little reason to separate this specimen and we consider C. reussi a subjective junior synonym of Pseudonoduloconus austriacus (as already proposed by Bałuk 1997). A specimen from the Tortonian of Stazzano in Italy, described by Sacco (1893 a) as Conus mercati subaustriacus, might also represent this species, but is too fragmentary for a clear identification. Paleoenvironment. The Austrian localities Gainfarn and Steinebrunn represent shallow water environments with sea grass (e. g. Zuschin et al. 2007).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF13FF03FF5FABF3FAA84236.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn, Steinebrunn (Austria) (Hoernes & Auinger 1879; Sieber 1958 b); Carpathian Foredeep: Korytnica (Poland); Transylvanian Basin: Lăpugiu de Sus (Romania) (NHMW collection). Proto-Mediterranean Sea and northeastern Atlantic: Langhian (middle Miocene): Loire Basin: Ferrière- Larçon (Glibert 1952 a); Tortonian (late Miocene):? Stazzano (Italy) (as subaustriacus in Sacco 1893 a).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF12FF00FF5FABD9FA1543C2.taxon	description	Conus Gastaldii mihi — Michelotti 1847: 344.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF12FF00FF5FABD9FA1543C2.taxon	description	[Conus] S. [tephanoconus] Gastaldii var. superneasulcata Sacc. — Sacco 1893 b: 117, pl. 11, fig. 12. [Conus] S. [tephanoconus] Gastaldii var. supraproducta Sacc. — Sacco 1893 b: 117, pl. 11, fig. 13.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF12FF00FF5FABD9FA1543C2.taxon	materials_examined	Type material. Syntype, illustrated by Sacco (1893 b, pl. 11, fig. 10), Museo Regionale di Scienze Naturali, Torino (BS. 038.08.003), Turin Hills, early Miocene, Burdigalian. The type locality of Conus subcoronatus Boettger, 1887, considered to be a junior synonym of P. gastaldii, is the middle Miocene (Badenian) locality Lăpugiu de Sus in Romania. Studied material. 1 spec. SMF XII. 2246 a, Lăpugiu de Sus (Romania). Illustrated material. Figs 33 F 1 – F 3: SMF XII. 2246 a coll. O. Boettger ex G. Schmidt 1885, SL: 50.1 mm, MD: 26.8 mm, Lăpugiu de Sus (Romania); holotype of Conus subcoronatus Boettger, 1887; picture Sigrid Hof; courtesy Ronald Janssen, section Malacology, Senckenberg Forschungsinstitut Frankfurt / Main. Revised description. Moderately large and robust shell with low spire; early spire whorls tuberculate just above suture. Spire whorls striate, weakly convex with wide, blurred nodes (= pseudo-nodes sensu Tucker & Tenorio 2009); suture undulose. Subsutural flexure shallow, moderately curved, strongly asymmetrical. Rounded shoulder with blurred nodes; position of maximum diameter slightly below shoulder; last whorl elongate, faintly ventricose, weakly constricted. Aperture and siphonal canal largely destroyed; siphonal fasciole indistinctly swollen. Few prominent spiral cords on base. No colour pattern preserved.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF12FF00FF5FABD9FA1543C2.taxon	description	Shell measurements and ratios. Holotype: SL: 50.1 mm, MD: 26.8 mm, spire angle: 112 °, last whorl angle: 32 °, LW: 1.87, RD: 0.63, PMD: 0.95, RSH: 0.15.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF12FF00FF5FABD9FA1543C2.taxon	discussion	Discussion. Tucker & Tenorio (2009) placed this rare species in Pseudonoduloconus based on the conspicuous spire sculpture; this was followed by Kovács & Balász (2016), who treated the Paratethyan Pseudonoduloconus subcoronatus as distinct species. The illustration in Boettger (1887) is misguiding and suggests a broader shell with slightly opisthocline and very prominent nodes. In fact, the holotype is a rather slender shell with low spire, which does not differ from the Italian Burdigalian morphotype described by Sacco (1893 b, pl. 11, fig. 13) as Conus gastaldii supraproducta. Therefore, we consider Conus subcoronatus Boettger, 1887 to be a subjective junior synonym of Conus gastaldii Michelotti, 1847. It is not surprising that Boettger (1887) did not recognise the synonymy because the first figures of Conus gastaldii were published much later by Sacco (1893 b). This species is distinguished from P. austriacus (Hoernes & Auinger, 1879) by its distinctly more slender outline, the less constricted base, the narrower and higher spire and the more swollen “ pseudo-nodes ”. Paleoenvironment. Unknown.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF12FF00FF5FABD9FA1543C2.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Transylvanian Basin: Lăpugiu de Sus (Romania) (Boettger 1887). Proto-Mediterranean Sea and northeastern Atlantic: Burdigalian (early Miocene): Turin Hills (Sacco 1893 b); Aquitaine Basin: Léognan, Mérignac, Saint-Paul-lès-Dax (France) (Peyrot 1931); middle Miocene (Langhian): Langhian: Aquitaine Basin: Manciet (France) (Peyrot 1931, as Conus subnocturnus, d’Orbigny, 1852).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF11FF00FF5FAA8AFED946BC.taxon	description	Conus (Stephanoconus) wagneri n. sp. — Boettger 1902: 7.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF11FF00FF5FAA8AFED946BC.taxon	materials_examined	Type material. Holotype, illustrated by Zilch (1934, pl. 22, fig. 9), Senckenberg Museum, Frankfurt / Main, German, SMF XII. 2204 a, Coşteiu de Sus (Romania); middle Miocene, Badenian. Studied material. Holotype. Illustrated material. Figs 33 I 1 – I 3: Holotype, SMF XII. 2204 a; coll. O. Boettger 1899, SL: 15.0 mm, diameter: 8.5 mm, Coşteiu de Sus (Romania); picture Sigrid Hof, courtesy Ronald Janssen, section Malacology, Senckenberg Forschungsinstitut Frankfurt / Main. Revised description. Stout, biconical shell with moderately high, coeloconoid to conical spire; spire whorls with bulgy rim along lower suture and shallow mid-whorl concavity; striate. Subsutural flexure moderately deep, asymmetrically curved. Blurred nodes on shoulder, causing strongly undulating suture. Distinctly angulated shoulder; last whorl slightly ventricose, weakly constricted at base; entirely covered by very weak spiral cords. Siphonal fasciole weakly swollen and twisted. Aperture and siphonal canal largely destroyed. No colour pattern preserved.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF11FF00FF5FAA8AFED946BC.taxon	description	Shell measurements and ratios. SL: 15.0 mm, MD:. 8.5 mm, spire angle: 85 °, last whorl angle: 50 °, LW: 1.76, RD: 0.79, PMD: 1.09, RSH: 0.28.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF11FF00FF5FAA8AFED946BC.taxon	discussion	Discussion. This is a poorly known species, documented only by its holotype, which might represent a subadult specimen. Despite the small size, the sculpture of the spire whorls suggests a placement in Pseudonoduloconus Tucker & Tenorio, 2009. The lack of tubercles on early spire whorls does not contradict this allocation as the extant type species P. carnalis (Sowerby III, 1879) lacks tubercles as well. Nevertheless, this feature allows a separation from the two other Paratethyan species of this genus, which both have strongly tuberculate early spire whorls. Hence, we can exclude that the shell is just a juvenile of P. austriacus (Hoernes & Auinger, 1879) or P. gastaldii (Michelotti, 1847). Kovács & Balász (2016) treated Conus wagneri Boettger, 1902 as a junior synonym of Conilithes granularis (Borson) [herein described as Artemidiconus granularis (Borson, 1820)]. In our opinion, both taxa differ considerably in outline and spire sculpture. Especially the nodulose shoulder and the moderately deep and asymmetrical subsutural flexure of P. wagneri differ considerably from A. granularis. Moreover, it lacks the characteristic beads on the last whorl of A. granularis. Paleoenvironment. Unknown.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF11FF00FF5FAA8AFED946BC.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Transylvanian Basin: Coşteiu de Sus (Romania) (Boettger 1902).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF10FF01FF5FAB68FE2545AB.taxon	description	The shells are reminiscent of Lautoconus species in general shell shape. The striate spire whorls would not contradict a placement in this genus as well. The prominently tuberculate early spire whorls, however, are unknown in Lautoconus.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF10FF1EFF5FAD63FEFF43C2.taxon	description	[Chelyconus] [Conus] Johannae n. f. — Hoernes 1878 a: 195 (nomen nudum). Conus (Chelyconus) Johannae nov. form. — Hoernes & Auinger 1879: 40, pl. 1, fig. 4.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF10FF1EFF5FAD63FEFF43C2.taxon	materials_examined	Type material. Syntype NHMW 1949 / 0005 / 0003, illustrated in Hoernes & Auinger (1879, pl. 1, fig. 4), syntype NHMW 1869 / 0001 / 0332, all Steinebrunn (Austria); middle Miocene, Badenian (late Langhian). Studied material. Syntypes and 4 subadult spec. NHMW A 458, all Steinebrunn (Austria). Illustrated material. Figs 30 O, 34 A 1 – A 4: Steinebrunn (Austria): SL: 75.5 mm, MD: 38.9 mm, NHMW 1949 / 0005 / 0003, illustrated in Hoernes & Auinger (1879, pl. 1, fig. 4); Figs 34 I: Steinebrunn (Austria), SL: 73.0 mm, MD: 38.6 mm, NHMW 1869 / 0001 / 0332. Revised description. Moderately large, robust shells; mammillate spire regularly conical, elevated. Early spire whorls with tuberculate keel and distinct striae; later spire whorls convex, bulgy; only weakly striate with deep suture. Subsutural flexure shallow, moderately curved, moderately asymmetrical; subangulated middle spire whorls, rounded penultimate whorl; dense pattern of growth lines on spire. Last whorl with rounded shoulder, position of maximum diameter slightly below; last whorl slender, regularly conical, not constricted; aperture moderately narrow, broadening towards short, weakly recurved canal; fasciole weak, slightly twisted and not well demarcated from base. Last whorl covered by broad, indistinct spiral cords. Colour pattern under UV light consisting of flammulae on shoulder and spire whorls; last whorl with about 15 broad spirals of subquadratic blotches, coinciding with spiral cords.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF10FF1EFF5FAD63FEFF43C2.taxon	description	Shell measurements and ratios. Only two adult specimens are available: SL: 75.5 / 73.0 mm, MD: 38.9 / 38.6 mm, spire angle: 91 / 97 °, last whorl angle: 32 / 33 °, LW: 1.94 / 1.89, RD: 0.63 / 0.65, PMD: 0.88 / 0.93, RSH: 0.18 / 0.19.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF10FF1EFF5FAD63FEFF43C2.taxon	discussion	Discussion. This rare species is highly reminiscent of Lautoconus subraristriatus (Pereira da Costa, 1866) in overall shape and size and could be easily confused with the widespread species. Nevertheless, the tuberculate keel and prominent striae on the early spire of C. johannae and its conspicuous colour pattern allow a clear separation. The colour pattern is unique within Paratethyan Conidae. The record of this species from the Croatian Karlovac- Glina Basin by Eremija (1959) is based on a misidentification. Paleoenvironment. The co-occurring mollusc assemblage is indicative for shallow sublittoral enbvironments (Sieber, 1958 b).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF10FF1EFF5FAD63FEFF43C2.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Steinebrunn (Austria) (Hoernes & Auinger 1879). The occurrence of this species in the Tortonian of Barcelona, mentioned by Faura I Sans (1908), needs confirmation.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0FFF1FFF5FAA8AFD4A4372.taxon	description	Conus (Chelyconus) Suessi Varietät III — Hoernes & Auinger 1879: 44, pl. 6. figs 3 – 4.? Conus an Suessi R. Hoern. i Auinger — Friedberg 1911: 61, text-fig. 15.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0FFF1FFF5FAA8AFD4A4372.taxon	materials_examined	Type material. Holotype: NHMW 1999 z 0077 / 0024, Lăpugiu de Sus (Romania), illustrated in Hoernes & Auinger (1879, pl. 6, fig. 3); middle Miocene, Badenian (Langhian). Studied material. Holotype and 1 spec. NHMW 1854 / 0035 / 0038, Lăpugiu de Sus (Romania), illustrated in Hoernes & Auinger (1879, pl. 6, fig. 4). Illustrated material. Figs 34 E 1 – E 3: Lăpugiu de Sus (Romania): SL: 49.8 mm, MD: 27 mm, NHMW 1999 z 0077 / 0024, illustrated in Hoernes & Auinger (1879, pl. 6, fig. 3); 30 M, Figs 34 F 1 – F 3: holotype, Lăpugiu de Sus (Romania): SL: 50.4 mm, MD: 25.2 mm, NHMW 1854 / 0035 / 0038, illustrated in Hoernes & Auinger (1879, pl. 6, fig. 4). Revised description. Medium-sized, biconical, moderately robust shells; tall, mammillate spire, conical with impressed suture; early spire whorls flat, striate with broad tubercles along carina, which is placed at the lower suture on earliest whorls, migrating slightly adapically on later whorls, to be placed a short distance above lower suture, making later whorls weakly angular. Deeply incised, narrow, slightly pitted spiral grooves separate 6 regular, narrow spiral cords on sutural ramp; surface smooth below angulation. Last spire whorl with concave sutural ramp. Shoulder slightly angulated; last whorl weakly convex; subsutural flexure shallow, moderately curved, nearly symmetrical. Aperture moderately narrow, slightly widening anteriorly; very narrow anal notch. Siphonal fasciole and inner lip indistinct; siphonal canal short, feebly reflected; faint spiral threads on base. Colour pattern under UV light consisting of wide-spaced flammulae on shoulder and numerous spirals of close-spaced dashes on last whorl.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0FFF1FFF5FAA8AFD4A4372.taxon	description	Shell measurements and ratios. Holotype: SL: 49.8 mm, MD: 27 mm, spire angle: 80 °, last whorl angle: 37, LW: 1.8, RD: 0.66, PMD: 0.86, RSH: 0.18; spire angle: 95 °, last whorl angle: 37 °; second specimen: SL: 50.4 mm, MD: 25.2 mm, LW: 2.0, RD: 0.64, PMD: 0.85, RSH: 0.22.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0FFF1FFF5FAA8AFD4A4372.taxon	discussion	Discussion. When introducing Conus suessi var. posticestriatus Kojumdgieva in Kojumdgieva & Strachimirov (1960) designated the specimen illustrated by Hoernes & Auinger (1879, pl. 6, fig 3) as the holotype. As already pointed out by Bałuk (1997), the Bulgarian specimen identified by Kojumdgieva in Kojumdgieva & Strachimirov (1960) is not conspecific with the type from Lăpugiu de Sus. The illustrated Bulgarian specimen is fragmentary and seems to represent Conus s. l. sturi (Hoernes & Auinger, 1879). Bałuk (1997) applied posticestriatus to shells from the Badenian of Poland, which correspond fairly well with Conus posticestriatus in outline but are rather poorly preserved and thus, the identification remains doubtful. Moreover, the spiral threads on the last whorl are not seen in the Polish specimens. The stout biconical outline and especially the characteristic deeply incised narrow spiral grooves on the spire of Plagioconus posticestriatus separate it distinctly from Leporiconus suessi. The mode of sculpture formation on the spire whorls differs fundamentally from the striae and cords of other Paratethyan species and characterises this species. Paleoenvironment. The specimens contained sand with elphidiid foraminifers suggesting a shallow water environment.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0FFF1FFF5FAA8AFD4A4372.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene):? Carpathian Foredeep: Korytnica (Poland); Transylvanian Basin: Lăpugiu de Sus (Romania).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0EFF1DFF5FAA1AFCB54192.taxon	materials_examined	Type material. Syntype NHMW 1854 / 0035 / 0034 a, Lăpugiu de Sus (Romania), illustrated in Hoernes & Auinger (1879, pl. 1, fig. 16), 10 syntypes NHMW 1854 / 0035 / 0034, Lăpugiu de Sus (Romania); middle Miocene, Badenian (Langhian). Studied material. Syntypes. Illustrated material. Figs 34 B 1 – B 3: syntype, Lăpugiu de Sus (Romania): SL: 44.9 mm, MD: 19.5 mm, NHMW 1854 / 0035 / 0034; Figs 30 L, 34 C 1 – C 3: syntype, Lăpugiu de Sus (Romania): SL: 40.8 mm, MD: 17.5 mm, NHMW 1854 / 0035 / 0034 a, illustrated in Hoernes & Auinger (1879, pl. 1, fig. 16); Figs 34 D 1 – D 2: Lăpugiu de Sus (Romania): SL: 32.3 mm, MD: 14.9 mm, NHMW 1854 / 0035 / 0034. Revised description. Medium-sized olivoid shells; pointed apex with tuberculate early spire whorls; later spire whorls weakly convex to nearly flat, striate, forming a high, nearly straight-sided conical to weakly cyrtoconoid spire. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Very indistinct shoulder coinciding with slight angulation and position of maximum diameter; last whorl moderately elongate, weakly ventricose, not constricted. Siphonal canal very short, moderately wide, straight; fasciole very weak and indistinct; few thin spiral threads on base. Colour pattern under UV light consisting of wide-spaced spirals of long dashes, intercalated by 1 – 2 weaker spirals of shorter dashes. If uppermost shell layer is eroded, these spirals are underlain by a dense pattern of spirals covering the entire shell.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0EFF1DFF5FAA1AFCB54192.taxon	description	Shell measurements and ratios. n = 11: largest specimen: SL: 44.9 mm, MD: 19.5 mm, mean SL: 33.7 mm (σ = 1.8), mean MD: 14.9 mm (σ = 2.6), spire angle: µ = 71.5 ° (σ = 3.1 °), last whorl angle: µ = 33.8 ° (σ = 1.1 °), LW: µ = 2.3 (σ = 0.1), RD: µ = 0.58 (σ = 0.03), PMD: µ = 0.87 (σ = 0.03), RSH: µ = 0.23 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0EFF1DFF5FAA1AFCB54192.taxon	discussion	Discussion. The strong spiral pattern on the syntype illustrated by Hoernes & Auinger (1879) is somewhat misleading, as it represents a specimen with corroded uppermost shell layer. These spirals are just a colour pattern and do not represent a spiral sculpture as suggested by Halaváts (1884) and Strausz (1966). The stout, robust specimen with somewhat ventricose last whorl, illustrated by Atanacković (1969, 1985) as Conus praelongus, might represent another species. Lautoconus praelongus differs from the slightly similar Leporiconus suessi (Hoernes & Auinger, 1879), Leporiconus transsylvanicus (Hoernes & Auinger, 1879) and Conus s. l. sturi Hoernes & Auinger, 1879 in its higher conical spire, shorter last whorl and lower position of the shoulder. Paleoenvironment. No information.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0EFF1DFF5FAA1AFCB54192.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Eisenstadt-Sopron Basin: Forchtenau (Austria) (Sieber 1956); Transylvanian Basin: Lăpugiu de Sus (Romania) (Hoernes & Auinger 1879); Dacian Basin: Staropatica (Bulgaria) (Kojumdgieva & Strachimirov 1960).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0CFF1DFF5FA93AFDB146CA.taxon	description	non Conus cf. Sturi R. Hoern. i. Auinger — Friedberg 1911: 53, pl. 2, fig. 15 [= Conus s. l. mucronatolaevis (Sacco, 1893)].	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0CFF1DFF5FA93AFDB146CA.taxon	materials_examined	Type material. Syntype NHMW 1854 / 0035 / 0043 a, Lăpugiu de Sus (Romania), illustrated in Hoernes & Auinger (1879, pl. 5, fig. 9); syntype NHMW 1854 / 0035 / 0043 b, Lăpugiu de Sus (Romania), illustrated in Hoernes & Auinger (1879, pl. 5, fig. 10); 11 syntypes NHMW 1854 / 0035 / 0045, NHMW 1854 / 0035 / 0047, NHMW 1858 / 0043 / 0003, Lăpugiu de Sus (Romania); middle Miocene, Badenian (Langhian). Studied material. Syntypes. Illustrated material. Fig. 30 N: syntype, Lăpugiu de Sus (Romania): SL: 38.1 mm, MD: 17.1 mm, NHMW 1854 / 0035 / 0043 a, illustrated in Hoernes & Auinger (1879, pl. 5, fig. 9); Figs 34 G 1 – G 3: syntype, Lăpugiu de Sus (Romania): SL: 43.7 mm, MD: 20.5 mm, NHMW 1854 / 0035 / 0043 b, illustrated in Hoernes & Auinger (1879, pl. 5, fig. 10); Figs 34 H 1 – H 3: syntype, Lăpugiu de Sus (Romania): SL: 49.1 mm, MD: 22.1 mm, NHMW 1854 / 0035 / 0 0 47.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0CFF1DFF5FA93AFDB146CA.taxon	description	Description: Medium-sized, moderately solid shells with medium high mammillate-conical spires and elongate conical last whorls. First 5 – 6 spire whorls high and conical, forming pointed apex, tuberculate along lower suture; striate. Later spire whorls distinctly striate, convex, forming low dome-shaped spire; suture impressed but narrow. Subsutural flexure shallow, moderately curved, strongly asymmetrical. Rounded shoulder coinciding with position of maximum diameter. Last whorl elongate conical, not constricted. Narrow aperture, narrowing adapically, only weakly widening towards short, weakly recurved siphonal canal. Siphonal fasciole narrow, indistinct. Sculpture of last whorl consisting of wide-spaced, convex spiral cords, which may be largely reduced on adapical half. Colour pattern not well preserved; spiral cords appear as light spiral bands in UV light; spirally arranged small speckles seem to be present on upper half of last whorl. Shell measurements and ratios. n = 8: largest specimen: SL: 49.1 mm, MD: 22.1 mm, mean SL: 41.3 mm (σ = 4.5), mean MD: 18.3 mm (σ = 2.0), spire angle: µ = 89.8 ° (σ = 5.6 °), last whorl angle: µ = 30.5 ° (σ = 1.1 °), LW: µ = 2.3 (σ = 0.11), RD: µ = 0.55 (σ = 0.02), PMD: µ = 0.89 (σ = 0.01), RSH: µ = 0.19 (σ = 0.02).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0CFF1DFF5FA93AFDB146CA.taxon	discussion	Discussion. A characteristic feature of this species is the spiral cords on the last whorl, which are true cords on the shell surface and not just intervening shell surface between spiral groves as seen in Plagioconus lapugyensis. The Italian Burdigalian Conus mucronatolaevis Sacco, 1893 is quite similar but differs in the lower position of the shoulder and lacks the raised spiral cords on the base. Indirectly, Hall (1966) considered Conus sturi as a synonym of Conus pyrula Brocchi, 1814 (by referring to a figure in Friedberg 1911 and not to the original description of Hoernes & Auinger, 1879), which is incorrect in respect to the much broader and slightly ventricose shell of Lautoconus pyrula (see holotype in Pinna & Spezia 1978).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0CFF1DFF5FA93AFDB146CA.taxon	distribution	Distribution in Paratethys. Transylvanian Basin: Lăpugiu de Sus (Romania) (Hoernes & Auinger 1879);? Dacian Basin: Radomirci (Bulgaria).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0BFF1AFF5FAA30FDB144C4.taxon	materials_examined	Type material. Syntypes described by Eichwald (1830) from Salisze about 50 km NE of Ternopil (Ukraine) (see map in Dubois de Montpéreux 1831), syntype illustrated in Eichwald (1852, pl. 9, fig. 2). The specimens are most probably stored in the Zoological Museum of St. Petersburg (Russia) but could not be detected; middle Miocene, Badenian (Langhian).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0BFF1AFF5FAA30FDB144C4.taxon	discussion	Discussion. Eichwald (1830, 1852, 1853) described and figured a medium-sized species (SL: c. 38 mm, MD: c. 18 mm) with a conical spire of weakly convex whorls with a deep suture and prominent spiral cords. Its last whorl develops a rounded shoulder and an elongate, weakly ventricose last whorl with a long and reflected siphonal canal. In the description Eichwald (1853) mentioned nodulose tubercles on the early spire whorls, which are not visible on the illustration. Although Hall (1966) synonymized this species with Conus vindobonensis Hoernes & Auinger, 1879, the two taxa are not very similar. Especially the elongate last whorl and the long siphonal canal exclude conspecificity with Conus s. l. vindobonensis. Moreover, the spiral cords on the spire whorls of C. s. l. vindobonensis are always much weaker. Bałuk (1997) treated Conus argillicola as a subjective junior synonym of Conus pyrula Brocchi, 1814. The Pliocene Lautoconus pyrula, however, lacks spiral cords, is broader, and has rounded shoulders and very prominent and broad spiral cords on the base. The illustration of Eichwald (1852) does not correspond unambiguously to any other Paratethyan and Mediterranean species and the status of Conus argillicola remains unclear without observation of type specimens or new material.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0BFF1AFF5FAA30FDB144C4.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Carpathian Foredeep: Salisze at Tarnów (Ukraine), Korytnica (Poland) (Eichwald 1830).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0BFF18FF5FAE77FE7944AC.taxon	description	[Chelyconus] [Conus] olivaeformis n. f. — Hoernes 1878 a: 195 (nomen nudum). Conus (Chelyconus) olivaeformis nov. form. — Hoernes & Auinger 1879: 52, pl. 1, fig. 23.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0BFF18FF5FAE77FE7944AC.taxon	description	Conus (Chelyconus) praelongus fusiformis Halaváts, 1884 — Strausz 1966: 461, text-fig. 208.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0BFF18FF5FAE77FE7944AC.taxon	materials_examined	Type material. Syntype NHMW 1854 / 0035 / 0046, Lăpugiu de Sus (Romania), illustrated in Hoernes & Auinger (1879, pl. 1, fig. 23); 10 syntypes NHMW 1858 / 0035 / 0008, Lăpugiu de Sus (Romania); middle Miocene, Badenian (Langhian). The type locality of Conus fusiformis Halaváts, 1884, considered to be a junior synonym of C. olivaeformis, is the middle Miocene (Badenian) locality Hidas in Hungary. Studied material. Syntypes. Illustrated material. Figs 35 A 1 – A 3: syntype, Lăpugiu de Sus (Romania): SL: 24.5 mm, MD: 12.6 mm, NHMW 1858 / 0035 / 0008; Figs 30 Q, 35 B 1 – B 3: syntype, Lăpugiu de Sus (Romania): SL: 21.9 mm, MD: 10.4 mm, NHMW 1854 / 0035 / 0046, illustrated in Hoernes & Auinger (1879, pl. 1, fig. 23). Revised description. Small subcylindrical shells; high conical protoconch comprising c. 3 whorls; mammillate spire of medium height, with weakly tuberculate whorls; later spire whorls flat to weakly convex, forming conical to cyrtoconoid spire; suture narrow but impressed, very irregular. Surface glossy, smooth except for delicate growth lines, not striate. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Shoulder indistinct to rarely subangulate. Narrow spiral grooves on base, delimiting broader cords. Siphonal fasciole very short, moderately swollen, poorly demarcated from thin and narrow inner lip. Moderately narrow aperture, adapically distinctly narrowing; siphonal canal moderately wide, short. Colour pattern under UV light consisting of three indistinct spiral bands of blotches, which are composed of densely spaced spiral dashes; additional dashes appear between these bands.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0BFF18FF5FAE77FE7944AC.taxon	description	Shell measurements and ratios. n = 7: largest specimen: SL: 24.5 mm, MD: 12.6 mm, mean SL: 22.3 mm (σ = 2.0), mean MD: 11.2 mm (σ = 1.0), spire angle: µ = 87.7 ° (σ = 6.1 °), last whorl angle: µ = 36.6 ° (σ = 2.7 °), LW: µ = 2.0 (σ = 0.08), RD: µ = 0.6 (σ = 0.02), PMD: µ = 0.87 (σ = 0.03), RSH: µ = 0.16 (σ = 0.05).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0BFF18FF5FAE77FE7944AC.taxon	discussion	Discussion. The spire height and shape are variable and the pointed apex and the tuberculate early spire are usually eroded. Hall (1966) listed this well-defined species as a synonym of Lautoconus ponderosus (Brocchi, 1814), which was probably a mistake. Lautoconus ponderosus is much larger, broader and has a distinctly more conical last whorl. Halaváts (1884) described this species as Conus fusiformis. This name is preoccupied by Conus fusiformis Lamarck, 1810 and Conus fusiformis Pease, 1861. Halaváts (1884) discussed similarities with Lautoconus praelongus (Hoernes & Auinger, 1879) and Strausz (1966) considered this taxon a subspecies of L. praelongus. Both species differ in the much lower spire, the more ventricose and cylindrical last whorl, and the incised suture of L. olivaeformis. The holotype of Conus fusiformis Halaváts, 1884 was stored in the collection of the Geological and Geophysical Institute of Hungary in Budapest but seems to be lost (pers. comm. Klára Palotás). The specimen illustrated by Halaváts (1884) does not differ from L. olivaeformis and we consider it to be a subjective junior synonym of Conus olivaeformis. Therefore, no replacement name for the preoccupied C. fusiformis is necessary. Paleoenvironment. No information.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF0BFF18FF5FAE77FE7944AC.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Pannonian Basin: Hidas, Letkés, Pécsszabolcs (Hungary) (Halaváts 1884; Kovács & Vicián 2013);? southern Pannonian Basin: Miljevići, Hrvaćani (Bosnia and Herzegovina) (Atanacković 1985); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus (Romania) (Hoernes & Auinger 1879; Boettger 1902).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF09FF16FF5FAE6FFC334252.taxon	description	[Conus] vindobonensis P. [artsch] — Hörnes 1848: 16 (nomen nudum).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF09FF16FF5FAE6FFC334252.taxon	materials_examined	Type material. Syntypes: 3 spec. NHMW 1846 / 0037 / 0043, Gainfarn (Austria), including specimen illustrated in Hörnes (1851, pl. 3, fig. 5); 5 spec. NHMW 1846 / 0037 / 0049, Enzesfeld (Austria), including specimen illustrated in Hörnes (1851, pl. 3, fig. 7). Studied material. 3 spec. NHMW 1846 / 0037 / 0043, Gainfarn (Austria), including specimen illustrated in Hörnes (1851, pl. 3, fig. 5); 5 spec. NHMW 1846 / 0037 / 0049, Enzesfeld (Austria), including specimen illustrated in Hörnes (1851, pl. 3, fig. 7); 10 spec. NHMW 1855 / 0045 / 0359, 9 spec. NHMW 1853 / 0003 / 0005, 9 spec. NHMW 1856 / 0050 / 0115, all Gainfarn (Austria); 4 spec. NHMW 1846 / 0037 / 0052, Gainfarn and Steinebrunn (Austria); 7 spec. NHMW 1846 / 00027 / 0028, Steinebrunn (Austria); 16 spec. NHMW 1860 / 0001 / 0067, Mikulov-Kienberk (Czech Republic); 10 spec. NHMW 1849 / 0023 / 0002, Bad Vöslau, including specimen illustrated in Hoernes & Auinger (1879, pl. 6, fig. 9); 1 spec. NHMW 1849 / 0023 / 0002 a, Bad Vöslau, specimen illustrated in Hoernes & Auinger (1879, pl. 6, fig. 10); 3 spec. NHMW 1846 / 0037 / 0048, Enzesfeld (Austria), including specimen illustrated in Hörnes (1851, pl. 2, fig. 4. Illustrated material. Figs 35 C 1 – C 2: Gainfarn (Austria): SL: 53.4 mm, MD: 26.5 mm, NHMW 1856 / 0050 / 0115; Figs 35 D 1 – D 3: syntype, Enzesfeld (Austria): SL: 48.8 mm, MD: 26.7 mm, NHMW 1846 / 0037 / 0049, illustrated in Hörnes (1851, pl. 3, fig. 7); Figs 35 E 1 – E 3: Mikulov-Kienberk (Czech Republic): SL: 47.5 mm, MD: 23.8 mm, NHMW 1860 / 0001 / 0067; Figs 35 F: Gainfarn (Austria): SL: 58.4 mm, MD: 29.2 mm, NHMW 1856 / 0050 / 0115; Figs 35 G 1 – G 2: Gainfarn (Austria): SL: 30.6 mm, MD: 14.3 mm, NHMW 1846 / 0037 / 0043 a; Fig. 30 R: Gainfarn (Austria): SL: 30.1 mm, MD: 15.2 mm, NHMW 1846 / 0037 / 0043 a. Revised description. Medium-sized moderately slender shells; spire coeloconoid to conical and of variable height. Spire whorls channelled, moderately convex to nearly flat; early spire whorls scalariform, strongly tuberculate and striate (but rarely preserved); later whorls with more or less prominent striae, which may fade out completely on last whorls. Subsutural flexure deep, moderately curved, moderately asymmetrical. Last whorl elongate, slightly ventricose, very weakly constricted with weak spiral grooves on abapical third; shoulder high, more or less prominent, rounded to subangular; position of maximum diameter slightly below shoulder. Siphonal canal short, weakly recurved; siphonal fasciole indistinct. Aperture straight, moderately narrow. Colour pattern under UV light only poorly preserved, consisting of a dense pattern of thin, roughly spirally arranged short dashes and slightly spirally extended dots.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF09FF16FF5FAE6FFC334252.taxon	description	Shell measurements and ratios. n = 20 adult specimens (only large morphs): largest specimen: SL: 58.4 mm, MD: 29.2 mm, mean SL: 51.9 mm (σ = 3.3), mean MD: 27.2 mm (σ = 1.5), spire angle: µ = 98.3 ° (σ = 10.4 °), last whorl angle: µ = 35.1 ° (σ = 1.9 °), LW: µ = 1.91 (σ = 0.1), RD: µ = 0.63 (σ = 0.03), PMD: µ = 0.89 (σ = 0.03), RSH: µ = 0.17 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF09FF16FF5FAE6FFC334252.taxon	discussion	Discussion. Hall (1966: 137) discussed a close relation of Conus vindobonensis with Conus ventricosus Gmelin, 1791, which would result in a placement in Lautoconus. The tuberculate, depressed scalariform early spire, however, contradicts a relation with Lautoconus. In any case, Hall (1966) was correct to interpret this species as a very polymorphic species comparable to the extant Lautoconus ventricosus. Especially the height of the spire and the convexity of the spire whorls are very variable. Similarly, the outline of the last whorl ranges from slightly ventricose to nearly straight-sided. Like the recent L. ventricosus, the spire of the Miocene C. vindobonensis tends to be more or less gradate in some specimens. Consequently, the high spired, gradate specimens illustrated by Hoernes & Auinger (1879, pl. 6, figs 10 – 11) as Conus mediterraneus are most probably aberrant specimens of Conilithes vindobonensis. Therefore, Conus mediterraneus var. pupoidemiocenica and permiocenica, which were introduced by Sacco (1893 b) as new names for these specimens, are subjective junior synonyms of Conus vindobonensis. The same holds true for the aberrant specimen illustrated by Hörnes (1851, pl. 2, fig. 4) as Conus clavatus, for which Sacco (1893 b) introduced the superfluous varietal name mioexclavata, although already Hoernes & Auinger (1879) identified this specimen as their Conus vindobonensis. Hall (1966: 137) considered this species to be a subjective junior synonym of Conus argillicola Eichwald, 1830, from the Badenian of Ukraine. The specimen illustrated by Eichwald (1852, pl. 9, fig. 2) as C. argillicola differs from C. vindobonensis in its constricted base and the long and reflected siphonal canal. Moreover, the spiral striae on the spire whorls of C. argillicola are much stronger. Therefore, we doubt that both taxa represent the same species. Unfortunately, we were not able to find the type specimen (s) of C. argillicola to solve this question definitely. Paleoenvironment. Typically found in shallow marine nearshore environments; at Gainfarn, where it is among the most frequent cones, it was associated with seagrass.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF09FF16FF5FAE6FFC334252.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn, Enzesfeld, Steinebrunn, Baden, Baden-Sooß, Bad Vöslau, Möllersdorf, Pötzleinsdorf, Grinzing, (Austria), Mikulov-Muschelberg, Mikulov-Kienberk, Hrušovany (Czech Republic), Eisenstadt-Sopron Basin: Marz, Forchtenau (Austria); Styrian Basin: Pöls; Alpine-Carpathian Foredeep: Grund, Guntersdorf (Austria) (Hoernes & Auinger 1879; Sieber 1958 a, b), Korytnica, Węglinek, Łychów (Poland) (Krach 1981; Bałuk 1997); Pannonian Basin: Hidas (Hungary) (Strausz 1966); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus (Romania) (Boettger 1902); Buzău Basin: Crivineni, Valea Muscel (Romania); Zârand Basin: Minişul de Sus (Romania) (Nicorici & Sagatovici 1973); Caransebeş-Mehadia Basin: Valea Satului (Romania) (Hinculov 1968).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF07FF17FF5FAC84FB54461C.taxon	description	[Conus] Chelyconus globoponderosus Sacc. — Sacco 1893 b: 85 (nov. nom. pro Conus avellana in Hörnes 1851, pl. 3, fig. 3).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF07FF17FF5FAC84FB54461C.taxon	description	Conus mucronatolaevis Sacco 1893 — Krach 1981: 76, pl. 21, figs 3 – 4, 11 – 13, 21.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF07FF17FF5FAC84FB54461C.taxon	materials_examined	Type material. Syntype illustrated in Sacco (1893 b, pl. 6, fig. 26) and Ferrero-Mortara et al. (1984, pl. 18, fig. 8), stored in the Museo Regionale di Scienze Naturali, Torino (BS. 038.05.043); Baldissero (Italy), Burdigalian (early Miocene). Studied material. 4 spec. NHMW 1847 / 0037 / 0027, Mikulov-Muschelberg (Czech Republic), including specimen illustrated in Hörnes (1851, pl. 3, figs 3 a – c) (= syntype of Conus globoponderosus Sacco, 1893); 1 spec. NHMW 1853 / 0003 / 0006, Gainfarn (Austria); 1 spec. NHMW 1846 / 0037 / 0064, Steinebrunn (Austria). Illustrated material. Figs 35 H 1 – H 3: Mikulov-Muschelberg: SL: 59.2 mm, MD: 32.3 mm, NHMW 1847 / 0037 / 0 0 27, illustrated in Hörnes (1851, pl. 3, figs 3 a – c); Figs 35 I 1 – I 3: Mikulov-Muschelberg (Czech Republic), SL: 45.4 mm, MD: 23.8 mm, NHMW 1847 / 0037 / 0027; Figs 30 S, 35 J: Gainfarn (Austria), SL: 42.6 mm, MD: 23.3 mm, private collection Anton Breitenberger (Bad Vöslau, Austria). Revised description. Moderately large, solid shells; early spire mammillate, coeloconoid with prominent beads along lower suture; later spire globose, cyrtoconoid with nearly flat and striate spire whorls; impressed but not channelled suture. Subsutural flexure shallow, weakly curved, moderately asymmetrical. Last whorl conical, contracting quickly below broadly rounded shoulder; not constricted. Siphonal fasciole nearly absent; weak spiral cords on base. Aperture narrow, slightly flaring towards short siphonal canal. Colour pattern under regular and UV light consisting of densely spaced delicate spirals of tiny dots. In slightly eroded specimens, these dots form tiny beads.	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF07FF17FF5FAC84FB54461C.taxon	description	Shell measurements and ratios. n = 5: largest specimen: SL: 66.6 mm, MD: 32.3 mm, mean SL: 50.9 mm (σ = 11.3), mean MD: 26.8 (σ = 5.0), spire angle: µ = 98.8 ° (σ = 8.0 °), last whorl angle: µ = 36 ° (σ = 1.4 °), LW: µ = 1.89 (σ = 0.1), RD: µ = 0.64 (σ = 0.02), PMD: µ = 0.85 (σ = 0.02), RSH: µ = 0.18 (σ = 0.03).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF07FF17FF5FAC84FB54461C.taxon	discussion	Discussion. Hall (1966) revised Conus mucronatolaevis and synonymized all varieties named by Sacco (1893 b). Typical specimens of Conus s. l. mucronatolaevis are rather slender and have a more or less conical spire. Broader specimens with cyrtoconoid spires were described by Sacco (1893 b) as Conus m. globospira and C. m. glandispira. The Paratethyan specimens are closer to these morphotypes. In addition, to the numerous variety names for C. mucronatolaevis, Sacco (1893 b) introduced globoponderosus as new name for the specimens referred to as Conus avellana by Hörnes (1851). The two specimens illustrated by Hörnes (1851) are almost certainly not conspecific and his fig 3 d is most probably a poorly preserved Plagioconus lapugyensis (Hoernes & Auinger, 1879). Therefore, Conus globoponderosus Sacco, 1893 should be restricted to the specimen illustrated by Hörnes (1851, pl. 3) as figs 3 a – c. Within the broad species concept of Hall (1966), Conus globoponderosus Sacco, 1893 is a subjective junior synonym of C. mucronatolaevis; within a narrower frame, it would still be a subjective junior synonym of Conus globospira Sacco, 1893. Curiously, Hall (1966) included the specimen illustrated as figs 3 a – c in Hörnes (1851, pl. 3) in the synonymy of Conus belus d’Orbigny, 1852. In our opinion, this decision is unjustified as Lautoconus belus, as redefined by Hall (1966), differs in its smaller size, shorter last whorl, the slightly constricted base and the conspicuous spiral sculpture on the last whorl (see also Muñiz-Solís 1999). The specimen described by Friedberg (1911, pl. 2, fig. 16) as Conus argillicola Eichwald is stored in the Geological Survey of Austria (GBA 1911 / 2 / 2) and represents a subadult C. mucronatolaevis. Mayer (1864: 77) introduced Conus borsoni Mayer, 1864 for a species from the Pliocene or Pleistocene of the Azores without illustration. In the discussion he referred to specimens from the Burdigalian of France (Grateloup, 1846: pl. 44, fig. 5) and the Badenian of Austria (Hörnes, 1851, pl. 3, fig. 3), both previously described as Conus avellana. As discussed above, the illustrations in Hörnes (1851, pl. 3, figs 3 a – d) represent two different species. Moreover, it is doubtful if the much younger specimens from the Azores are conspecific with any of the Miocene species at all. It is beyond the scope of this paper to revise the fossil Conidae of the Azores but to solve the confusion we designate the well preserved syntype from the tuff deposits of Ilhéu de Baixo, stored in the geological-paleontological collections of the ETH Zurich (Switzerland), as lectotype of Conus borsoni. Paleoenvironment. Shallow marine environments, partly with seagrass (e. g. Zuschin et al. 2007).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
373F87D7FF07FF17FF5FAC84FB54461C.taxon	distribution	Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn, Steinebrunn (Austria), Mikulov-Muschelberg (Czech Republic) (Hörnes 1851); Pannonian Basin: Letkés (Hungary) (Kovács & Vicián 2013); Carpathian Foredeep: Czepiele, Ternopil (Ukraine) (Friedberg 1911); Węglinek, Łychów (Poland) (Krach 1981). Proto-Mediterranean Sea and northeastern Atlantic. Burdigalian (early Miocene): Turin Hills: Baldissero (Italy) (Sacco 1893 b), middle Miocene: Aquitaine Basin, Salies-de-Béarn (France) (Peyrot 1931).	en	Harzhauser, Mathias, Landau, Bernard (2016): A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1-178, DOI: 10.11646/zootaxa.4210.1.1
