taxonID	type	description	language	source
367587E8FF90FFF7FF1B5461A477DB51.taxon	diagnosis	Diagnosis. Males of this genus can be recognised by the presence of a basal retrolateral apophysis on the tibia of the pedipalp (Fig. 8), armed with short, stout setae (Fig 3 a) except in the case of A. grismadoi sp. nov., which has stout setae but no apophyses (Figs 20 a, b). The embolic division has an sclerotised area divided in two branches, one large ventral (VB in Fig. 2 a – d; sclerotised ridge in A. grismadoi sp. nov., Figs 19 d, 20 c, e) and the other smaller and dorsal (DB in Fig. 2 a – d), which is sometimes not visible in the unexpanded bulb (i. e. in the case of A. velox, Fig. 10 d). Females resemble those of Pirata Sundevall, 1833 and Piratula Roewer, 1960, but can be differentiated by the lack of the posterior lobes of the epigyne (Figs 42 – 43 and fig. 70 in Omelko et al. 2011).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF90FFF7FF1B5461A477DB51.taxon	description	Description. Medium-sized wolf spiders (TL, 4.20 – 9.98 mm). Females larger than males. Carapace brown with a light brown median band and a Y-shaped mark that extends from fovea to the PLE (tenuous in A. grismadoi sp. nov. and A. punctata, absent in A. luteonigra). Eyes surrounded by black pigment. Dorsum of abdomen brown to dark olive-grey, with light yellow lanceolate mark in the cardiac area and a pattern of white spots laterally and posterior to the cardiac mark, composed by glistening setae (Fig. 3 b) (except in A. luteonigra, which have a black abdomen). Carapace longer than wide, dorsal line straight in lateral view (Figs 9 b, e; domed in A. grismadoi sp. nov. – Figs 18 b, e). Caput flanks in frontal view steep in males, gently sloping in females. AME larger than ALE, AER slightly procurved in anterior view (Fig. 24). Chelicerae with three promarginal and three retromarginal teeth. Abdomen oval. Colulus a fleshy triangular lobe, with several setae (fig. 5 a). The spinnerets have a similar conformation to what Townley and Tillinghast (2003) reported for Lycosidae, with six spinnerets, ALS and PLS two segmented, PMS with a single segment (Figs 5 a, 6 a). ALS with two major ampullate gland spigots on the mesal margin (the posterior one reduced to a nubbin in the male (Fig. 6 b )) and piriform gland spigots (Fig. 5 b). PMS with about 30 aciniform and two cylindrical gland spigots in the female (Fig. 5 c) and about 20 aciniform in the male and without cylindrical gland spigots (Fig. 6 c), with a few setae between them. Female with two minor ampullate gland spigot with a nubbin and a tartipore close to them on the PMS (Fig. 5 c) and only one minor ampullate gland spigot in the male (Fig. 6 c). PLS long, tubular, distal segment short, conical, with aciniform and cylindrical gland spigots, with setae among them (Fig. 5 d). Leg formula IV> I> II> III or IV> I> III> II, except in A. luteonigra where the leg formula is IV> III> I> II. Tibia I with two pairs of ventral spines (2 - 2 - 0), except in A. pirity sp. nov. (0 - 2 - 0) and A. grismadoi sp. nov. (2 - 0 - 0). Femur of male palp without modifications. Tibia with a basal retrolateral apophysis, covered with strong setae (Fig. 3 a), reduced in A. grismadoi sp. nov. (Fig. 8 f). Cymbium symmetric, without macrosetae in its tip. Tegulum with a prolateral furrow, deep in A. velox Keyserling, 1891 (Figs 2 a, 10 d) and A. tricuspidata Tullgren, 1905 (Fig. 12 d) and shallow in the rest of the species. Median apophysis membranous, composed of two elements, the ventral one is flattened and directed longitudinally, the dorsal is transverse and usually retrolaterally directed (Fig. 2 a, b). The embolic division possesses a grooved and complex terminal apophysis, with a pointed tip (Figs 2 a, b, 7) and two additional processes, one ventral and a dorsal (Fig. 2). Embolus as a curved spine, not visible in the unexpanded palp, originating prolaterally, below the ventral process of the embolic division and running retrolaterally hidden by the terminal apophysis (Fig. 7). Epigyne composed of a simple plate with two soft sclerotised areas; in A. velox, A. punctata, A. gentilis and A. grismadoi sp. nov. this area is well developed and surrounds the copulatory open, and in A. luteonigra, A. pirity sp nov., A. tricuspidata and A. gamas sp. nov. is reduced to a small area, posterior to the copulatory open. Vulva: spermathecae with a distal head connected to the base by a more or less defined stalk, vulval chambers rounded. Dictynoid pore conspicuous, located on the boundaries of the vulval chamber and the spermathecal stalk (Fig. 4 a).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF90FFF7FF1B5461A477DB51.taxon	distribution	Distribution. Most of the species are known from the eastern Argentina and neighbouring countries, with one species from subtropical montane forest of northwestern Argentina (Figs 25 – 27).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF90FFF7FF1B5461A477DB51.taxon	discussion	Remarks. The original illustrations of the epigyne of A. bryantae (Roewer, 1951) from Hispanola, A. denisi (Caporiacco, 1947) from Guyana, A. fallax (L. Koch, 1877) from Queensland, A. kolbei (Dahl, 1908) from Bismark Arch., A. pickeli (Mello-Leitão, 1937) from Brazil, A. yaucensis (Petrunkevitch, 1929) from Puerto Rico shows the typical shape of a Lycosinae epigyne, with an inverted T-shaped septum and anterior pockets. The type specimens of A. chacoensis (Mello-Leitão, 1942) and A. melanotaenia (Mello-Leitão, 1941) were examined; both belong to an undescribed Lycosinae genus (Alvares 2006). The authors of the remaining two species, A. helvola (C. L. Koch, 1847) from Mexico and Colombia, A. subinermis (Simon, 1897) from India not provided drawings of the copulatory organs, but the size, in the case of A. helvola, and the description of the epigyne, in A. subinermis suggest that those two species not belong to Agalenocosa. The ten species mentioned above where transferred to Agalenocosa by Roewer (1955) without any justification; none of these species fit in Agalenocosa and are here considered as incertae sedis; the final placement of these species is beyond the scope of this work. Agalenocosa fimbriata is considered here as species inquirenda because the epigyne of the holotype (from Arrecifes, Buenos Aires, deposited in the MLP number 15995, examined) was removed and is probably lost. There are no morphological characters that allow me to recognise which species it is.	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF96FFF6FF1B57CBA542DEB0.taxon	discussion	Remarks. The catalogue of the type material available at the Museu Nacional (Rio de Janeiro, Brazil) published by Silva Moreira et al. (2010) refers a vial of A. singularis from the type locality labelled as “ type ”, but in the original description of Mello-Leitão (1944) clearly indicates that the types are deposited in the MLP with the numbers 16010 and 16011, both examined in this work. The types of P. velox and A. singularis were compared, and show no significant morphological differences.	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF96FFF6FF1B57CBA542DEB0.taxon	materials_examined	Other material examined. ARGENTINA. Entre Ríos: Parque Nacional Predelta, bañado cerca de entrada, 6 km S Diamante, S 32.12194 ° W 60.62996 ° (GPS, + - 20 m), elev. 10 m, Ramírez, M. J., Piacentini, L. N., González, M. E., Laborda, A. & Aisen, S., 30. iv. 2013, hand collection in marshes, 1 ♀ (MACN-Ar 30310); Arroyo Gualeguaychú y ruta 14, Goloboff, P. A. & Ramírez, M. J., 10. xii. 1982, 1 ♀ (MACN-Ar 22713); Ruta 12 en el Empalme con ruta a Paranacito, Galiano, M. E. & Scioscia, C. L., 11. x. 1987, 1 ♀ (MACN-Ar 22775). San Luis: Villa Elena, Maury, E., 12. xi. 1982, 1 ♂ (MACN-Ar 22723); Cortaderas, Roig, A., 10. xi. 1982, 2 ♀ (MACN-Ar 22733). Buenos Aires: no precise locality, Viana, J. M., x. 1941, 1 ♂ (MACN-Ar 22768); Tigre, INTA Delta, xii. 1988, 1 ♂ (MACN-Ar 30625); Delta del Paraná, San Antonio, frente al Urion, Bachmann, A., 15. i. 1950, 6 ♀ 5 eggsac (MACN-Ar 3096); Campana, Reserva Natural Otamendi, Estación Río Luján, - 34,277352 º - 58,88054 º (WG 84), elev. 5 m, Ramírez, M. J., Grismado, C., Piacentini, L., Soto, E. González-Marquez, M. E., Guala, M., Andía Navarro, J. M., López, N., Ortega Insaurralde, I., & Zermoglio, P., 9. ix. 2010, in marsh and grassland, 1 ♀ (MACN-Ar 28440); same locality, Grismado, C. J., Damer, L., López, N., Trivero, S. & Grismado, A., 3 – 17. ii. 2007, pitfall on grassland of “ pelo de chancho ”, Distichlis sp, 2 ♀ (MACN-Ar 12122); same locality, Grismado, C. J., Damer, L., López, N., Trivero, S., Crudele, I. & Olejnik, N., 10. iii. 2007, hand collected on grassland, 1 ♀ (MACN-Ar 12215); same locality and collectors, 30. vi. 2007, 1 ♀ 1 immature (MACN-Ar 12663); same locality, Grismado, C. J., Damer, L., Crudele, I., Olejnik, N., López, N. & De Biase, S., 28. vii. 2007, hand collected on grassland, 1 ♂ 1 immature (MACN-Ar 12788); same locality, Zanetic, C. & Goloboff, P. A., 12. i. 1980, 1 ♀ (MACN-Ar 28988); Tigre, 23. xii. 1979, 1 ♀ (MACN-Ar 22744); Isla entre Zárate y Brazo Largo, del lado de Zárate, Zanetic, A. & Goloboff, P. A., 10. i. 1979, 1 ♀ (MACN-Ar 22750); Don Torcuato, Ropps, J. J., 1939, 1 ♀ (MACN-Ar 22726); Adolfo Sordeaux, Piacentini, L., i. 2002, 1 ♂ (MACN-Ar 22741); Punta Lara, Galiano, M. E. & Scioscia, C. L., 17. ix. 1986, 2 ♂ 1 ♀ (MACN-Ar 22716); same locality, 2. xi. 1946, 1 ♀ (MACN- Ar 22757); same locality, Fuentes, B, 27. x. 1997, hand collected on grassland, 1 ♂ (MACN-Ar 28493); same locality, Goloboff, P. A., 28. xii. 1989, 1 ♂ (MACN-Ar 30641); Mar del Plata, Sander, V., x. 2002, 1 ♂ (MACN-Ar 22747); Carmen de Patagones, Daguerre, J. B., 2 ♀ (MACN-Ar 22770). Ciudad Autónoma de Buenos Aires: Palermo, Frers, A. G., 11. x. 1914, under trunks in marshes, 1 ♂ (MACN-Ar 22715); Reserva Ecológica Costanera Sur: Laguna de los Macáes, S 34 º 36 ' 21,7 '' W 58 º 20 ' 55 '', Turienzo, P., 13. x. 2008, collected with pitfall traps, 2 ♂ (MACN-Ar 28446); same locality, Benedictto & Mamani, A., 5. i. 2009, collected with pitfall traps, 1 ♀ (MACN-Ar 28448); same locality, Mamani, A. & Zapata, L., 24. xi. 2008, 1 ♂ (MACN-Ar 28445); same locality, Mamani, A. & Turienzo, P., 2. ii. 2009, 1 ♀ (MACN-Ar 28480); same locality, Benedictto & Mamani, A., 5. i. 2009, 1 ♀ (MACN- Ar 28444); Reserva Ecológica Costanera Sur: Canal Viamonte, S 34 º 35 ' 47 '' W 58 º 21 ' 36,8 '', Turienzo, P., 13. x. 2008, collected with pitfall traps, 1 ♂ (MACN-Ar 28479); same locality, Mamani, A., 27. x. 2008, 1 ♂ (MACN-Ar 28449); same locality, Turienzo, P., 13. x. 2008, 1 ♀ 1 immature (MACN-Ar 28450); same locality, Mamani, A. & Turienzo, P., 29. ix. 2008, 1 ♀ (MACN-Ar 28478).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF96FFF6FF1B57CBA542DEB0.taxon	diagnosis	Diagnosis. Males and females of A. velox resemble those of A. punctata by the coloration pattern; males can be distinguished by the straight and larger tibial apophysis (Figs 8 a, c). Females can be distinguished by the large epigynal plate, that cover almost the anterior ventral side of the abdomen (Figs 10 a, 16 a). Agalenocosa velox vulvae also resemble those of A. punctata, but differs by the long twisted stalk of spermathecae (Fig. 10 b), instead of short and straight as in A. punctata (Fig. 16 b).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF96FFF6FF1B57CBA542DEB0.taxon	description	Description. Male (MACN-Ar 12788). Carapace brown with a light brown median band, with a conspicuous Y-shaped dark mark that extends from the fovea to the PLE and with marginal pale bands (Fig. 9). Sternum yellow, sparsely covered with brown bristles, more abundant on the margins (Fig. 9 f). Labium dark grey (Fig. 9 f). Chelicerae lighter than labium, light brown, covered with brown bristles. Dorsum of abdomen brown with longitudinal, irregular pale marks on the center and longitudinal lines of white pubescence on its lateral side, composed by bunches of glistening setae (Figs 3 b, 9). Venter pale brown (Fig. 9 f). Legs pale brown without annulations. Leg formula IV> I> II> III. Spination: femur I p 0 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, II p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, III p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, IV p d 1 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1; patella I d 1 - 1, II p 1 d 1 - 1, III p 1 d 1 - 1 r 1, IV p 1 d 0 - 1 r 1; tibia I v 2 - 2 - 0, II p d 1 - 1 v r 1 - 2 - 0, III p 1 - 1 d 1 - 1 r 1 - 1 v 2 - 2 - 2 ap, IV p 1 - 1 d 1 - 1 r 1 - 1 v 2 - 2 - 2 ap; metatarsus I v 2 - 2 - 2 ap, II p 0 - 1 - 2 ap r 0 - 0 - 2 ap v 2 - 2 - 0, III p d 1 - 1 - 2 ap r d 1 - 0 - 2 ap v 2 - 2 - 1 ap, IV p d 1 - 1 - 2 ap r d 1 - 1 - 2 ap v 2 - 2 - 1 ap. Palp. Basal apophysis of the palpal tibia longer than the diameter of the tibia (Figs 3 a, 8 a). Subtegulum in the mesal side of the alveolus (Fig. 10 d). Tegulum with deep prolateral furrow (Figs 2 a, 10 d). Ventral branch of the median apophysis triangular, the dorsal branch triangular, with the tip apicad (Figs 2, 10 c – e). Terminal apophysis with a triangular end, pointing apicad (Figs 2, 10 c – e). Female (MACN-Ar 22716). Color (Fig. 9), as in male. Leg formula IV> I> III> II. Spination: femur I p 0 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - 0, II p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, III p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, IV p d 1 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1; patella I d 1 - 0, II p 1 d 1 - 0, III p 1 d 1 - 1 r 1, IV p 1 d 1 - 1 r 1; tibia I v 2 - 2 - 0, II p d 1 - d 1 v r 1 - 2 - 0, III p d 1 - 1 d 1 - 1 r 1 - 1 v 2 - p 1 - 2 ap, IV p d 1 - 1 d 1 - 1 r d 1 - 1 v p 1 - p 1 - 2 ap; metatarsus I v 2 - 2 - 2 ap, II p 1 - 0 - 2 ap r 0 - 0 - 1 ap v 2 - 2 - 1 ap, III p 1 - 1 - 2 ap r 1 - 1 - 2 ap v 2 - 2 - 1 ap, IV p d 1 - 1 - 2 ap r d 1 - 1 - 2 ap v 2 - 2 - 1 ap. Epigynal plate without pockets or atria, enlarged anteriorly, covering almost all the ventral surface between the pedicel and the epigastric furrow, with two oval soft sclerotised parts that surround the copulatory opening (Figs 4 b, 10 a). Vulva: head of spermatheca simple, not wider than the U-shaped stalk (Fig. 10 b). Vulval chamber rounded, behind of the base of spermathecae in dorsal view (Figs 4 a, 10 b). Measurements. Female, MACN-Ar 22716 (Male, MACN-AR 12788): TL 6.33 (5.60), CL 3.00 (2.80), CW 2.07 (2.13), CH 0.93 (1.00), AL 3.33 (2.80). Eyes: AME 0.13 (0.17), ALE 0.10 (0.10), PME 0.27 (0.27), PLE 0.20 (0.20). Row of eyes: AER 0.60 (0.57), PME 0.63 (0. 60), PLE 0.83 (0.83). Sternum (length / width) 1.53 / 1.20 (1.33 / 1.07). Labium (length / width) 0.40 / 0.37 (0.33 / 0.33). Legs: length of segments (femur + patella / tibia + metatarsus + tarsus = total length): I 2.20 + 2.53 + 1.67 + 0.93 = 7.33, II 2.13 + 2.27 + 1.67 + 0.93 = 7.00, III 1.93 + 2.27 + 1.73 + 0.8 = 6.73, IV 2.67 + 3.40 + 2.80 + 1.07 = 9.94 (I 2.00 + 2.60 + 1.73 + 0.93 = 7.26, II 2.00 + 2.33 + 1.67 + 0.93 = 6.93, III 2.00 + 2.00 + 1.73 + 0.87 = 6.60, IV 2.60 + 3.20 + 2.67 + 1.07 = 9.54). Variation. Females (Males) (range, mean ± s. d.): TL 4.60 – 7.46, 5.82 ± 0.85; CL 2.53 – 3.33, 2.93 ± 0.20; CW 1.80 – 2.40, 2.14 ± 0.19 n = 10; (TL 4.80 – 7.34, 5.67 ± 0.68; CL 2.60 – 3.67, 2.89 ± 0.30; CW 1.87 – 2.67, 2.09 ± 0.23 n = 10). Variation in coloration was observed in males and females, there are some darker specimens, which have dark annulations on the legs and show a thin marginal dark band on the carapace.	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF96FFF6FF1B57CBA542DEB0.taxon	distribution	Distribution. Central and central-eastern Argentina (San Luis, Entre Ríos and Buenos Aires Province) and one record from southern Brazil (state of Rio Grande do Sul) (Fig. 25). Natural history. Representatives of this species were found under dead vegetation in flooded grasslands or marshes, between the water surface and the dead vegetation in the Otamendi National Reserve. Some specimens were collected with pitfall traps placed on the margins of a canal in the Costanera Sur Ecological Reserve.	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF94FFF9FF1B53F5A146DF6D.taxon	materials_examined	Other material examined. ARGENTINA: Jujuy: Parque Nacional Calilegua, Alisal, altura km. 21 de la ruta provincial Nº 83, 23 ° 41 ’ 08.2 ” S, 64 ° 53 ’ 02.1 ” W (WGS 84), elev. 1508 m (GPS), Andía, J. M., 12. viii. 2010, 1 ♀ (MACN-Ar 26941); same data, 1 ♂ (MACN-Ar 26946); same data, 1 ♀ (MACN-Ar 26952); same data, 1 ♀ (MACN-Ar 26940); same data, 1 ♂ (MACN-Ar 26942); same data, 2 ♂ 8 ♀ (MACN-Ar 26951); Monolito, S 23 º 40´52.3 ´´ W 064 º 54´05.1 ´´ (+ / - 13 m, WGS 84), elev. 1709 m (GPS), Grismado, C., Izquierdo, M., Labarque, F., Rubio, G., Burger, M., Michalik, P., Carrera, P., Ojanguren A. & Mattoni, C., 06 – 11. xii. 2008, 1 ♀ one eggsac (MACN-Ar 28489); same data, 1 ♂ 1 ♀ 1 immature (MACN-Ar 28487); same locality, Izquierdo, M., Zapata, L. & Akmentis, M., 27 – 31. i. 2009, night hand collecting, 1 ♀ one eggsac (MACN-Ar 27503); same data, 1 ♀ (MACN- Ar 27513); Monolito, Arroyo Tres Cruces, cerca de Mesada de las colmenas, Grismado, C., Izquierdo, M., Labarque, F., Rubio, G., Burger, M., Michalik, P., Carrera, P., Ojanguren, A. & Mattoni, C., 06 – 11. xii. 2008, 1 ♂ 4 ♀ 4 immatures 2 eggsacs (MACN-Ar 28488); Mesada Las Colmenas, Ramírez, M. J. & Compagnucci, L., 08. viii. 1997, 1 ♀ (MACN-Ar 22734); toma de agua, 2 km arriba de Mesada Las Colmenas, 09. viii. 1997, 2 ♀ (MACN-Ar 22735); same data, 2 ♀ (MACN-Ar 22738); cerca de Sevenguillar, 3 km arriba de Mesada Las Colmenas, Ramírez, M. J. & Compagnucci, L., 07. viii. 1997, 1 ♀ (MACN-Ar 22736); Cerca de Monolito, Ramírez, M. J. & Compagnucci, L., 09. viii. 1997, 1 ♀ (MACN-Ar 22737); 11,3 km arriba de entrada al parque, Ramírez, M. J., Goloboff, P. A. & Szumik, C., 23. ix. 1995, 2 ♀ (MACN-Ar 22739); Camino Valle Grande-Calilegua, Monolito, Galiano, M. E., xi. 1967, 1 ♀ (MACN-Ar 22755); Mesada de las Colmenas, ribera y lecho del arroyo Negrito S 23 º 42´38,9 ´´ W 064 º 51´49,9 ´´ (WGS 84), elev. 802 m (GPS), Andía, J. M., 10. viii. 2010, 1 ♀ (MACN-Ar 26947); same data, 5 ♂ 5 ♀ 4 immature (MACN-Ar 26953); P. N. Calilegua, Mesada de las Colmenas, sendero el Negrito (ambos márgenes) S 23 º 42´01,1 ´´ W 064 º 51´59,1 ´´ (WGS 84), elev. 1135 m (GPS), Andía, J. M., 11. viii. 2010, 1 ♂ 1 immature (MACN-Ar 26945); 19,7 km arriba de entrada al parque, Ramírez, M. J., Goloboff, P. A. & Szumik, C., 23. ix. 1995, 2 ♂ 4 ♀ (MACN-Ar 22710); San Antonio, Casal, G., xi. 1965, 3 ♀ (MACN-Ar 22748); Yala, Maury E., x. 1967, 1 ♀ (MACN-Ar 6022): same locality, Roig, A., 09. i. 1983, 2 ♀ (MACN-Ar 22780); same locality, Roig, A. 07. i. 1983, 7 ♂ 1 ♀ 1 eggsac (MACN-Ar 22751); Laguna de Yala, Goloboff, P. A., v. 1983, 1 ♂ 3 ♀ (MACN-Ar 22717). Salta: P. N. El Rey, Galiano, M. E., 14. xi. 1966, 1 ♂ 1 immature (MACN-Ar 22712); same data, 2 ♂ 2 ♀ 1 immature (MACN-Ar 22722); same data, 4 ♀ (MACN-Ar 22731); same locality, Goloboff, P. A., i. 1981, 1 ♀ (MACN-Ar 22771); Finca Jakulica, 25 NW Aguas Blancas, Ramírez, M. J. & Goloboff, P. A., 15. xi. 1994, 1 ♂ 2 ♀ (MACN-Ar 22740); San Lorenzo, camino a ciudad de Salta, Ramírez, M. J. & Goloboff, P. A., 18. vii. 1995, 3 ♀ (MACN-Ar 22742); Camino Calilegua a Valle Grande, Galiano, M. E, xi. 1966, 2 ♀ 2 eggsacs (MACN-Ar 22754); Ojo de Agua, ruta nacional 9 km 1640 – 1642. límite con Jujuy, Ramírez, M. J., Goloboff, P. A. & Szumik, C., 22. ix. 1995, 1 ♀ (MACN-Ar 22767); San Lorenzo, 22. ix. 1988, 1 ♀ (MACN-Ar 22789); 17 km N. La Caldera, 1550 m, Alto de la Sierra, 2 – 30. xii. 1987, S & J Peck, subtrop. humid for., malaise FIT 87 – 134, 1 ♀ (AMNH); same data, 1 ♀ (AMNH). Tucumán: Cuesta de San Javier, Birabén, M., xi. 1960, 2 ♀ (MACN-Ar 22761); Tafí del Valle, Galiano, M. E., xi. 1960, 1 ♀ (MACN-Ar 5342). Catamarca: Viana, J. M., vii. 1949, 1 ♀ (MACN-Ar 2946).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF94FFF9FF1B53F5A146DF6D.taxon	diagnosis	Diagnosis. Males and females of A. tricuspidata resemble these of A. pirity sp. nov. in the size and coloration pattern. The submarginal pale, undulating bands of the carapace (Fig. 11) help to recognize well-preserved specimens of A. tricuspidata as these bands are absent in all other species of Agalenocosa. Males of A. tricuspidata differ in the broad terminal apophysis (Fig. 12 d). Females have a large and oval head of spermathecae, as in A. pirity sp. nov., but can be differentiated by the position of the vulval chamber lateral to the head of spermathecae (Fig. 12 b), not ventral as in A. pirity sp. nov.	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF94FFF9FF1B53F5A146DF6D.taxon	description	Description. Male (MACN-Ar 22946). Carapace brown with a light brown median band with a conspicuous Y-shaped dark mark that extends from the fovea to the PLE and pale submarginal bands with irregular margins (Fig. 11). Sternum yellow, sparsely covered with brown bristles, more abundant on the margins. Labium dark grey. Chelicerae lighter than labium, light brown, covered with brown bristles. Dorsum of abdomen brown with small longitudinal irregular pairs of pale marks on the center, and two pale longitudinal margins at sides (Fig. 11). Venter pale brown with irregular darker marks. Legs pale brown with dark annulations. Leg formula IV> I> III> II. Spination: femur I p 0 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, II p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, III p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, IV p d 1 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1; patella II p 1, III p 1 r 1, IV p 1 r 1; tibia I p 0 - 0 - v 1 v 2 - 2 - 0, II p d 1 - 1 r 0 - 0 - 1 v r 1 - 2 - 0, III p 1 - 1 d 1 - 0 r 0 - 1 - 1 v p 2 - 2 - 2 ap, IV p 1 - 1 d 0 - 1 r 1 - 1 v 2 - 2 - 2 ap; metatarsus I v 2 - 2 - 2 ap, II p 1 - 0 - 2 ap r 1 - 0 - 2 ap v 2 - 2 - 1 ap, III p 1 - 0 - 2 ap r 1 - 0 - 2 ap v 2 - 2 - 1 ap, IV p 1 - 0 - 2 ap r 1 - 0 - 2 ap v 2 - 2 - 1 ap. Palp. Basal apophysis of the palpal tibia rounded, medium sized (Fig. 8 c). Subtegulum in the meso-prolateral side of the alveolus (Fig. 12 d). Tegulum with a deep prolateral furrow (Figs 12 c – e). Ventral branch of the median apophysis broad, with a semicircular distal margin, and the dorsal branch triangular, with the tip pointing to the retrolateral side (Fig. 12 d). Terminal apophysis broad and retrolaterally pointed (Fig. 12 d). Female (MACN-Ar 26940). Colour in alcohol (Fig. 11) as in male. Leg formula IV> I> III> II. Spination: femur I p 0 - 0 - d 1 d 1 - 1 - 1, II p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, III p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, IV p d 1 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1; patella II p 1 d 1 - 0, III p 1 d 1 - 1 r 1, IV p 1 d 1 - 1 r 1; tibia I p 0 - 0 - v 1 v 2 - 2 - 0, II p d 1 - 0 - 1 v 2 - 2 - 0 - 1 ap, III p 1 - 1 d 1 - 1 r 1 - 1 v 2 - 2 - 2 ap, IV p 1 - 1 d 1 - 1 r 1 - 1 v 2 - 2 - 2 ap; metatarsus I v 2 - 2 - 2 ap, II p 1 - 0 - 2 ap r 1 - 0 - 1 ap v 2 - 2 - 1 ap, III p d 1 - 1 - 2 ap r 1 - 1 - 2 ap v 2 - 2 - 1 ap, IV p 1 - 1 - 2 ap r 1 - 1 - 2 ap v 2 - 2 - 1 ap. Epigynal plate without pockets or atria, copulatory openings with strongly sclerotised internal margin, and a lightly sclerotised posterior area (Fig. 12 a). Vulva: head of spermatheca enlarged, wider than the short stalk. Vulval chamber rounded, next to the base of spermathecae in dorsal view (Fig. 12 b). Measurements. Female, MACN-Ar 26940 (male, MACN-AR 26946): TL 7.19 (7.32), CL 3.86 (3.86), CW 2.66 (2.66), CH 1.20 (1.46), AL 3.33 (3.46). Eyes: AME 0.13 (0.17), ALE 0.10 (0.13), PME 0.30 (0.30), PLE 0.23 (0.23). Row of eyes: AER 0.70 (0.67), PME 0.83 (0.73), PLE 1.17 (1.00). Sternum (length / width) 2.00 / 1.47 (2.00 / 1.40). Labium (length / width) 0.73 / 0.50 (0.63 / 0.47). Legs: length of segments (femur + patella / tibia + metatarsus + tarsus = total length): I 2.80 + 3.47 + 2.27 + 1.27 = 9.81, II 2.60 + 3.27 + 2.13 + 1.20 = 9.20, III 2.60 + 3.27 + 2.40 + 1.20 = 9.47, IV 3.33 + 4.40 + 3.67 + 1.40 = 12.80 (I 2.73 + 3.40 + 2.40 + 1.27 = 9.8, II 2.47 + 2.93 + 2.27 + 1.20 = 8.87, III 2.47 + 2.93 + 2.40 + 1.20 = 9, IV 3.20 + 4.00 + 3.67 + 1.47 = 12.34). Variation. Females (males) (range, mean ± s. d.): TL 4.20 – 8.60, 6.81 ± 1.27; CL 2.47 – 4.07, 3.44 ± 0.53; CW 1.73 – 3.00, 2.42 ± 0.46; n = 10; (TL 5.34 – 7.32, 6.29 ± 0.65; CL 2.67 – 3.86, 3.33 ± 0.36; CW 2.00 – 2.67, 2.32 ± 0.23; n = 10).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF94FFF9FF1B53F5A146DF6D.taxon	distribution	Distribution. Montane forest (Yungas) in northwestern Argentina (Jujuy, Salta, Tucumán and Catamarca) (Fig. 25). Natural history. Some specimens were found running on the bank of the El Negrito stream in the Calilegua National Park.	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF98FFFEFF1B530BA1E6DDA9.taxon	discussion	Remarks. Mello-Leitão (1945) indicated as “ types ” of this species one male and one female from Argentina, Misiones, Pindapoy deposited in MLP 16480. Pereira et al. (1999) and Capocasale (2001) reported that such number does not exist in the MLP collection, but there are one female and one male from the same locality labelled as “ cotypus ” in Birabén´s collection in the MLP. Pereira et al. (1999) suggested that the specimens of Birabén´s collections should be the types, but Capocasale (2001) proposed as holotype and paratype the specimens deposited in Museu Nacional (Rio de Janeiro, Brazil), labelled as types by Mello-Leitão. We agree with Pereira et al. (1999) because the MLP specimens fit the description that Mello-Leitão provided.	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF98FFFEFF1B530BA1E6DDA9.taxon	materials_examined	Other material examined. ARGENTINA. Misiones: Santa María, Viana, J. M., 1 ♀ (MACN-Ar 3520). URUGUAY. Maldonado: Arroyo de la Llama, sobre ruta 39 (Km 75), 131 m, S 34 º 19 ’ 03.6 ’’ W 54 ª 48 ’ 10.7 ’’, Labarque, F., Ojanguren, A. & Mattoni, C., 10. x. 2005, 1 ♂ 3 ♀ 1 immature (MACN-Ar 24206).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF98FFFEFF1B530BA1E6DDA9.taxon	diagnosis	Diagnosis. This species is easily recognised by the dark coloration of the body. Males of A. luteonigra resemble those of A. tricuspidata in the shape of the tibial apophysis but can be recognised by the long and retrolaterally pointed terminal apophysis and the reduced dorsal branch of the median apophysis (Fig. 14 d). Females resemble to those of A. punctata in the shape of the vulva, but can be differentiated by the elongated head of the spermathecae (Fig. 14 b).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF98FFFEFF1B530BA1E6DDA9.taxon	description	Description. Male (MACN-Ar 24206). Carapace uniform black (Figs 13 d – f). Sternum and labium black. Chelicerae lighter than labium, light brown, covered with brown bristles. Dorsum of abdomen uniformly black, lighter ventrally. Legs: Femora dark brown with paler distal parts and the other legs are pale brown without annulations. Leg formula IV> III> I> II. Spination: femur I p 0 - 0 - d 1 d 1 - 1 - 1, II p 0 - d 1 - d 1 d 1 - 1 - 1 r 0 - d 1 - d 1, III p 0 - d 1 - d 1 d 1 - 1 - 1 r 0 - d 1 - d 1, IV p 0 - d 1 - d 1 d 1 - 1 - 1; patella I d 0 - 1, II p 1 d 1 - 1, III p 1 d 1 - 1 r 1, IV p 1 d 1 - 1 r 1; tibia I p d 1 - 1 v r 1 - 2 - 0, II p d 1 - 1 v 2 - 2 - 0, III p d 1 - 1 d 1 - 1 r 1 - 1 v 2 - 2 - 2 ap, IV p 1 - 1 d 1 - 1 r 1 - 1 v 2 - 2 - 2 ap; metatarsus I v 2 - 2 - 2 ap, II p d 1 - d 1 - 2 ap r 0 - 0 - 2 ap v 2 - 2 - 1 ap, III p d 1 - 1 - 2 ap r d 1 - 1 - 2 ap v 2 - 2 - 1 ap, IV p d 1 - 1 - 2 ap r d 1 - 1 - 2 ap v 2 - 2 - 1 ap. Palp. Basal apophysis of the palpal tibia rounded, medium sized (Fig. 8 b). Subtegulum in the mesal side of the alveolus. Tegulum, with a shallow prolateral furrow (Figs 14 c – e). Ventral branch of the median apophysis triangular, dorsal branch reduced to a transverse pointing protuberance directed retrolaterally (Fig. 14 d). Terminal apophysis elongated and pointed retrolaterally (Fig. 14 d). Female (MACN-Ar 24206). Colour in ethanol (Figs 13 a – c), as in male. Leg formula IV> III> I> II. Spination: femur I p 0 - 0 - d 1 d 1 - 1 - 1, II p d 1 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1, III p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, IV p d 1 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1; patella I d 1 - 0, II p 1 d 1 - 1, III p 1 d 1 - 1 r 1, IV p 1 d 1 - 1 r 1; tibia I v 2 - 2 - 0 d 1 - 0 - 1, II p d 1 - 0 - d 1 d 1 - 0 - 1 v r 1 - 2 - 0, III p 0 - d 1 - 1 d 0 - 1 - 1 r 0 - 1 - 1 v p 1 - p 1 - 0, IV p 0 - d 1 - 1 d 0 - 1 - 1 r 0 - 1 - 1 v p 1 - 2 - 2 ap; metatarsus I v 2 - 2 - 2 ap, II p 1 - 0 - 2 ap r 0 - 0 - 1 ap v 2 - 2 - 1 ap, III p d 1 - d 1 - 2 ap r d 1 - d 1 - 2 ap v 2 - 2 - 1 ap, IV p d 1 - 1 - 1 d 0 - 0 - 2 r d 1 - 1 - 1 v 2 - 2 - 1 ap. Epigynal plate without pockets or atria, copulatory openings surrounded by a small and rounded lightly sclerotised area (Fig. 14 a). Vulva: head of spermatheca digitiform, shorter than the stalk. Vulval chamber rounded, behind to the base of spermathecae in dorsal view (Fig. 14 b). Measurements. Female, MACN-Ar 24206 (male, MACN-Ar 24206): TL 4.60 (4.80), CL 2.40 (2.60), CW 1.87 (1.80), CH 0.80 (0.70), AL 2.40 (2.10). Eyes: AME 0.08 (0.10), ALE 0.07 (0.07), PME 0.23 (0.20), PLE 0.20 (0.15). Row of eyes: AER 0.47 (0.50), PME 0.55 (0.57), PLE 0.80 (0.75). Sternum (length / width) 1.33 / 1.00 (1.37 / 1.07). Labium (length / width) 0.18 / 0.17 (0.38 / 0.33). Legs: length of segments (femur + patella / tibia + metatarsus + tarsus = total length): I 1.93 + 2.20 + 1.47 + 0,73 = 6.33, II 1.87 + 2.27 + 1.40 + 0.73 = 6.27, III 1.87 + 2.20 + 0.73 + 1.67 = 6.47, IV 2.33 + 3.20 + 2.80 + 0.87 = 9.20 (I 2.33 + 2.67 + 1.80 + 0.80 = 7.60, II 2.33 + 2.47 + 1.80 + 0.80 = 7.40, III 1.93 + 2.33 + 1.80 + 0.73 = 6.79, IV 2.33 + 2.47 + 1.80 + 0.73 = 7.33). Variation. Females (males) (range, mean ± s. d.): TL 4.60 – 5.60, 5.22 ± 0.43; CL 2.40 – 2.93, 2.65 ± 0.23; CW 1.87 – 2.13, 1.98 ± 0.12 n = 4.	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF98FFFEFF1B530BA1E6DDA9.taxon	distribution	Distribution. Known only from the northeastern Argentina (Misiones) and southeastern Uruguay (Maldonado) (Fig. 25).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF9FFFE2FF1B5053A1F8DB9C.taxon	materials_examined	Other material examined. ARGENTINA. Entre Ríos: Parque Nacional Predelta, bañado cerca de entrada, 6 km S Diamante, S 32.12194 ° W 60.62996 ° (GPS, + - 20 m), elev. 10 m, 30. iv. 2013, Ramírez, M. J., Piacentini, L. N., González, M. E., Laborda, A. & Aisen, S., 1 ♀ (MACN-Ar 30644). Buenos Aires: Tigre: Isla Tres Bocas, Olejnik, N. & López, N., 3. vi. 2007, in leaf litter, 1 ♂ (MACN-Ar 12519); Boca del Río Salado, canal T- 1 - F, Botto & Irigoyen, 30. vi. 1973, 2 ♂ 1 ♀ 3 immatures (MACN-Ar 22707); Río Salado, canal T- 1 - B, Botto & Irigoyen, 29. vi. 1973, 1 ♂ (MACN-Ar 22719); Río Salado, arroyo T- 1 - C; cangrejal, Botto & Irigoyen, 3. viii. 1973, 1 ♂ (MACN-Ar 22729). Ciudad Autónoma de Buenos Aires: Playa Núñez, Castillo, F., viii. 1936, 1 ♂ 1 ♀ (MACN- Ar 22725); same locality, 1 ♀ (MACN-Ar 22766); Museo Argentino de Cs. Naturales, Galiano, M. E., 7. v. 1963, 1 ♂ (MACN-Ar 22756); Bañados de Palermo, Frers, A. G., 6. ix. 1917, under trunks in marshes, 1 ♂ 1 ♀ (MACN-Ar 22782).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF9FFFE2FF1B5053A1F8DB9C.taxon	diagnosis	Diagnosis. Males of A. punctata resemble those of A. velox by the coloration pattern, but can by recognised by the broad and short terminal apophysis, longitudinally oriented (Figs 16 d, 10 d). The females also resembles those of A. velox in the shape of the epigyne, but have a smaller epigynal plate (Figs 16 a, 10 a) and a short and straight stalk of the spermathecae (Figs 16 b, 10 b).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF9FFFE2FF1B5053A1F8DB9C.taxon	description	Description. Male (MACN-Ar 22756). Carapace brown, with a light brown median band and a conspicuous Y-shaped dark mark that extends from the fovea to the PLE, submarginal bands pale (Figs 15 d – f). Sternum yellow, sparsely covered with brown bristles, more abundant on margins. Labium dark grey. Chelicerae lighter than labium, light brown, covered with brown bristles. Dorsum of abdomen brown with longitudinal irregular pale marks on the center and longitudinal lines of white pubescence on its lateral sides (Figs 15 d – f). Venter pale brown. Legs pale brown without annulations. Leg formula IV> I> II> III. Spination: femur I p 0 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1, II p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, III p 0 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1, IV p 0 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1; patella I d 1 - 1, II p 1 d 0 - 1, III p 1 d 1 - 1 r 1, IV p 1 d 0 - 1 r 1; tibia I p d 1 - 1 v 2 - 2 - 0, II p d 1 - 1 v 2 - 2 - 0, III p 1 - 1 d 1 - 1 r 1 - 1 v 2 - 2 - 2 ap, IV p 1 - 1 d 1 - 1 r 1 - 1 v 2 - 2 - 2 ap; metatarsus I p 1 - ap 1 r 0 - 1 ap v 2 - (p 1 r 1) - 1 ap, II p 1 - ap 1 r 0 - 1 ap v 2 - (p 1 r 1) - 1 ap, III p d 1 - 1 - 2 ap r d 1 - 1 - 2 ap v 2 - 2 - 1 ap, IV p d 1 - 1 - 1 d 0 - 0 - 2 r d 1 - 1 - 1 v 2 - 2 - 1 ap. Palp. Basal apophysis of the palpal tibia rounded, medium sized (Fig. 8 d). Subtegulum in the meso-prolateral side of the alveolus. Tegulum with a shallow prolateral furrow (Figs 16 c, d). Ventral branch of the median apophysis rounded, dorsal branch triangular, with the tip curved ventrad (Fig. 16 d). Terminal apophysis short and longitudinally oriented (Fig. 16 d). Female (MACN-Ar 22725). Color in ethanol (Figs 15 a – c) as in male. Leg formula IV> I> II> III. Spination: femur I p 0 - 0 - d 1 d 1 - 1 - 1, II p d 1 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1, III p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, IV p d 1 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1; patella I d 0 - 1, II p 1 d 0 - 1, III p 1 d 1 - 1, IV p 1 d 1 - 1 r 1; tibia I v 2 - 2 - 0, II p d 1 - d 1 v 2 - 2 - 2 ap, III p d 1 - 1 d 1 - 1 r 1 - 1 v 2 - 2 - 2 ap, IV p d 1 - 1 d 1 - 1 r d 1 - 1 v p 1 - p 1 - 2 ap; metatarsus I v 2 - 2 - 2 ap, II p 1 - 0 - 2 ap r 0 - 0 - 1 ap v 2 - 2 - 1 ap, III p 1 - 1 - 2 ap r 1 - 1 - 2 ap v 2 - 2 - 1 ap, IV p d 1 - 1 - 1 d 0 - 0 - 2 r d 1 - 1 - 1 v 2 - 2 - 1 ap. Epigynal plate without pockets or atria, enlarged anteriorly, copulatory openings surrounded by a rounded lightly sclerotised area (Fig. 16 a). Vulva: head of spermatheca rounded, almost of the same length of the stalk. Vulval chamber rounded, behind to the base of spermathecae in dorsal view (Fig. 16 b). Measurements. Female, MACN-Ar 22725 (male, MACN-Ar 22756): TL 5.53 (5.80), CL 2.87 (3.27), CW 2.27 (2.27), CH 0.93 (1.13), AL 3.00 (2.53). Eyes: AME 0.13 (0.13), ALE 0.10 (0.10), PME 0.23 (0.27), PLE 0.17 (0.20). Row of eyes: AER 0.60 (0.57), PME 0.67 (0.63), PLE 0.93 (0.83). Sternum (length / width) 1.67 / 1.33 (1.70 / 1.33). Labium (length / width) 0.50 / 0.40 (0.43 / 0.37). Legs: length of segments (femur + patella / tibia + metatarsus + tarsus = total length): I 2.13 + 2.60 + 1.87 + 0.93 = 7.53, II 1.93 + 2.13 + 1.60 + 0.87 = 6.53, III 1.73 + 2.07 + 1.73 + 0.87 = 6.40, IV 2,47 + 3,27 + 2,60 + 1,0 7 = 9,41 (I 2.47 + 3.07 + 2.07 + 1.00 = 8.61, II 2.33 + 2.73 + 1.93 + 0.93 = 7.92, III 2.27 + 2.60 + 2.20 + 0.93 = 8.00, IV 2.93 + 3.67 + 3.60 + 1.13 = 11.33). Variation. Females (males) (range, mean ± s. d.): TL 5.53 – 6.60, 6.08 ± 0.56; CL 2.87 – 3.33, 3.08 ± 0.22; CW 2.00 – 2.40, 2.20 ± 0.20 n = 4; (TL 5.07 – 6.27, 5.74 ± 0.42; CL 2.67 – 3.33, 3.07 ± 0.20; CW 2.00 – 2.47, 2.22 ± 0.17 n = 8).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF9FFFE2FF1B5053A1F8DB9C.taxon	distribution	Distribution. Eastern central Argentina (northeastern Buenos Aires Province, Buenos Aires city and southern Entre Rios) (Fig. 26).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF83FFE1FF1B5600A50FD82C.taxon	materials_examined	Other material examined. ARGENTINA. Buenos Aires: Paraná, Isla Catalina, Doello, Daguerre, J. B., 25. ii. 1934, 1 ♀ (MACN-Ar 22743); Paraná de las Palmas, Bachmann, A., ii. 1963, 2 ♀ 6 immatures (MACN-Ar 22788); Delta, Río Sarmiento, Bachmann, A., 4. xii. 1947, 1 ♀ 1 immature (MACN-Ar 22753); Tigre, Damico, xi. 1947, 4 ♀ 3 eggsac (MACN-Ar 22760); Delta, Paraná de las Palmas, Bachmann, A., 28. i. 1964, 1 ♀ (MACN-Ar 22765); Rincón de Milberg, xii. 1980, 1 ♀ (MACN-Ar 28987); San Fernando, Piacentini, L., xii. 2002, 1 ♀ 1 immature (MACN-Ar 22758); same data, 1 ♀ (MACN-Ar 22777); Punta Lara, Partridge, W., 1. xi. 1946, 1 ♀ (MACN-Ar 22776); same locality, Miranda & Ramírez, M., 4. xii. 1981, 1 ♀ (MACN-Ar 30642).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF83FFE1FF1B5600A50FD82C.taxon	diagnosis	Diagnosis. The females of A. gentilis resembles those of A. punctata and A. velox by the coloration pattern (Figs 15, 17), but can be differentiated by the acute shape on the anterior margin of the lightly sclerotised area that surrounds the copulatory opening, and by the vulval chamber dorsal to the spermathecae (Fig. 17 b).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF83FFE1FF1B5600A50FD82C.taxon	description	Description. Male. Unknown. Female (MACN-Ar 22765). Carapace brown with light brown median band with a conspicuous Y-shaped dark mark that extends from the fovea to the PLE, pale submarginal bands (Figs 17 a – c). Sternum yellow, sparsely covered with brown bristles, more abundant on the margins. Labium dark grey. Chelicerae lighter than labium, light brown, covered with brown bristles. Dorsum of abdomen brown with longitudinal irregular pale marks on the center and longitudinal lines of white pubescence on its lateral sides (Figs 17 a – c). Venter pale brown. Legs pale brown without annulations. Leg formula IV> I> II> III. Spination: femur I p 0 - 0 - d 1 d 1 - 1 - 1, II p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, III p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, IV p d 1 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1; patella II p 1, III p 1 d 1 - 1 r 1, IV p 1 d 1 - 1 r 1; tibia I v 2 - 2 - 0, II p d 1 - 0 - d 1 v 2 - 2 - 0, III p d 1 - 0 - 1 d 1 - 1 - 0 r 0 - 1 - 1 v 2 - 2 - 2 ap, IV p 0 - d 1 - d 1 d 1 - 0 - 1 r 0 - 1 - 1 v p 2 - 2 - 2 ap; metatarsus I v 2 - 2 - 2 ap, II p 1 - 0 - 2 ap r 0 - 0 - 1 ap v 2 - 2 - 1 ap, III p d 1 - d 1 - 2 ap r d 1 - d 1 - 2 ap v 2 - 2 - 1 ap, IV p d 1 - 1 - 1 d 0 - 0 - 2 r d 1 - 1 - 1 v 2 - 2 - 1 ap. Epigynal plate without pockets or atria, copulatory openings surrounded by a rounded lightly sclerotised area with an acute anterior margin (Fig. 17 d). Vulva: head of spermatheca rounded, almost of the same length of the stalk, which has a dorsal protrusion. Vulval chamber small and rounded, next to the base of spermathecae in dorsal view (Fig. 17 e). Measurements. MACN-Ar 22765: TL 5.47, PL 3.33, PW 2.27, PH 1.07, OL 2.27. Eyes: AME 0.13, ALE 0.08, PME 0.22, PLE 0.18. Row of eyes: AER 0.52, PME 0.55, PLE 0.78. Sternum (length / width) 1.42 / 1.22. Labium (length / width) 0.50 / 0.47. Legs: length of segments (femur + patella / tibia + metatarsus + tarsus = total length): I 2.47 + 3.07 + 2.00 + 1.00 = 8.54, II 2.33 + 2.73 + 1.93 + 1.00 = 7.99, III 2.27 + 2.40 + 2.00 + 0.87 = 7.54, IV 3.00 + 3.67 + 3.27 + 0.93 = 10.87. Variation (range, mean ± s. d.). TL 5.07 – 6.47, 5.82 ± 0.46; CL 3.00 – 3.67, 3.16 ± 0.20; CW 2.13 – 2.60, 2.33 ± 0.13 n = 10.	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF83FFE1FF1B5600A50FD82C.taxon	distribution	Distribution. Known only from eastern central Argentina at northeast of Buenos Aires Province (Fig. 26).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF81FFEAFF1B524DA1AADA80.taxon	materials_examined	Type material. Male holotype from Argentina, Santa Fe, Madrejón San Felipe, (probably “ Don Felipe ”), no collector or date (MACN-Ar 22790), and two female paratypes from Argentina, Buenos Aires province, Tigre, Arroyo Espera, Galiano, M. E., iv. 1983, collected on “ camalotes ” (MACN-Ar 22778).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF81FFEAFF1B524DA1AADA80.taxon	etymology	Etymology. The specific epithet is a patronym in honour of the arachnologist Cristian J. Grismado in recognition of his help in collecting and companionship during many productive field trips in Argentina.	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF81FFEAFF1B524DA1AADA80.taxon	materials_examined	Other material examined. PARAGUAY. Alto Paraguay: Pto. Vallemi, Bachmann, A., v. 1952, 2 ♀ (MACN- Ar 22711). ARGENTINA. Santa Fe: no further location or collector, viii. 1969, 1 ♂ 1 immature (MACN-Ar 22727); no further location, Daguerre, J. B., viii. 1969, 1 ♀ (MACN-Ar 22786); no further location or collector, 3. v. 1963, 1 ♀ (MACN-Ar 22791); Arroyo del Medio, Rosas, 2. i. 1983, 1 ♂ (MACN-Ar 22709). Buenos Aires: Entre río Luján y Paraná de las Palmas, Ferradás, B. R., xii. 1983, 1 ♀ (MACN-Ar 22718); Tigre, Viana, J. M., 12. xii. 1951, 3 ♂ 12 ♀ 15 immature (MACN-Ar 24011); Delta, Río Sarmiento, Oliva, A., 3. iv. 1983, 2 ♂ 1 ♀ (MACN-Ar 22721); Delta, INTA, Carbajal, M., 8. iv. 1982, 1 ♀ (MACN-Ar 22728); Berisso, Ferradás, B. R., 1. xii. 1982, 1 ♀ (MACN-Ar 22784).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF81FFEAFF1B524DA1AADA80.taxon	diagnosis	Diagnosis. This species can be differentiated from all other currently recognised Agalenocosa species by the small size, domed carapace, in lateral view, instead of straight as in other species of the genus (Figs 18 b, d, 9 b, d) and the presence of 2 - 0 - 0 ventral spines on the tibia I instead of 2 - 2 - 0 or 0 - 2 - 0. Males of this species can also be recognised by the absence of the tibial apophysis in the palp (Fig. 8 f), present in the rest of species, and by the ventral process of the embolic division as a sclerotised ridge, more developed in the rest of species (Figs 19 e, 10 c, e). Females can be differentiated by the rounded lightly sclerotised area with sclerotised borders (Fig. 19 a) and the vulvae with small spermathecae (Fig. 19 b).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF81FFEAFF1B524DA1AADA80.taxon	discussion	Remarks. The shape of the median and terminal apophyses, and the structure of the embolic division (Figs 20 c, e) suggests the tentative placement of this species in Agalenocosa, despite the lack of the tibial apophysis on the male palp and the shape of the ventral process of the embolic division. The retrolateral side of the male palpal tibia on A. grismadoi sp. nov. is covered with short, stout setae (Figs 20 a, b), more dense on the basal part (Fig. 8 f), this setae resemble to those present on the tibial apophysis of the rest Agalenocosa species (Fig. 3 a).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF81FFEAFF1B524DA1AADA80.taxon	description	Description. Male. Holotype (MACN-Ar 22790). Carapace with inconspicuous design (Figs 18 d – f). Sternum yellow, sparsely covered with brown bristles, more abundant on the margins. Labium dark grey. Chelicerae lighter than labium, light brown, covered with brown bristles. Dorsum of abdomen brown with a longitudinal wide, pale band with irregular margins, and with spots of whitish glistening setae in the front (Figs 18 d – f). Venter pale brown, slightly darkened at the spinnerets. Legs pale brown with darker annulations, except femora I and II, which are dark brown. Leg formula IV> I> II> III. Spination: femur I p 0 - 0 - d 1 d 1 - 1 - 1, II p 0 - 0 - d 1 d 1 - 1 - 1, III p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, IV p 0 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1; patella I d 1 - 0, II p 1 d 1 - 1, III p 1 d 1 - 1 r 1, IV p 1 d 1 - 1 r 1; tibia I v 2 - 0 - 0, II p d 1 - 0 - d 1 v 0 - r 1 - 0, III p 0 - d 1 - 1 d 1 - 0 - 1 r 0 - d 1 - 1 v p 1 - p 1 - 2 ap, IV p 0 - d 1 - 1 d 1 - 0 - 1 r 0 - 1 - 1 v 2 - 2 - 2 ap; metatarsus I p 0 - 0 - v 1 ap v 2 - 0 - 0 r 0 - 0 - v 1 ap, II p 1 - 1 - 2 ap v r 1 - 2 - 2 ap, III d 2 - 2 - 2 ap v 2 - 2 - 3 ap, IV p d 1 - d 1 - 2 ap r d 1 - 1 - 2 ap v 2 - 2 - 1 ap. Palp. Tibia without basal apophysis (Fig. 8 f), short, stout setae present (Figs 20 a, b). Subtegulum in the mesoprolateral part of the alveolus. Tegulum with a shallow prolateral furrow (Figs 19 c-d, 20 d). Ventral branch of the median apophysis spatula-shaped, dorsal branch beak-shaped, directed retrolaterally (Figs 19 d, 20 c, e). Terminal apophysis short, retrolaterally pointed (Fig. 19 d, 20 c – e). Female. Paratype (MACN-Ar 22778). Colour in ethanol (Figs 18 a – c) as in male. Leg formula IV> I> II> III Spination: femur I p 0 - 0 - d 1 d 1 - 1 - 1, II p 0 - 0 - d 1 d 1 - 1 - 1, III p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, IV p 0 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1; patella I d 1 - 0, II p 1 d 1 - 1, III p 1 d 1 - 1 r 1, IV p 1 d 1 - 1 r 1; tibia I v p 1 - 0 - 0, II p d 1 - 0 - d 1 v r 1 - r 1 - 0, III p 0 - d 1 - 1 d 1 - 0 - 1 r 0 - d 1 - 1 v p 1 - p 1 - 2 ap, IV p 0 - d 1 - 1 d 1 - 0 - 1 r 0 - d 1 - 1 v 2 - 2 - 2 ap; metatarsus I p 0 - 0 - v 1 ap v 2 - 0 - 0 r 0 - 0 - v 1 ap, II p 1 - 0 - 2 ap v r 1 - r 1 - 2 ap, III d 2 - 2 - 2 ap v 2 - 2 - 3 ap, IV p d 1 - d 1 - 2 ap r d 1 - 1 - 2 ap v 2 - 2 - 1 ap. Epigynal plate without pockets or atria, copulatory openings surrounded by a rounded lightly sclerotised area with sclerotised borders (Fig. 19 a). Vulva: head of spermatheca rounded, of the same length than the stalk, but little broader. Vulval chamber oval, behind to the base of spermathecae in dorsal view (Fig. 19 b). Measurements. Female, MACN-Ar 22778 (male, MACN-Ar 22790): TL 5.05 (7.58), CL 2.60 (4.00), CW 2.00 (3.00), CH 1.07 (1.40), AL 2.33 (3.00). Eyes: AME 0.10 (0.17), ALE 0.08 (0.13), PME 0.17 (0.25), PLE 0.13 (0.17). Row of eyes: AER 0.53 (0.68), PME 0.47 (0.68), PLE 0.68 (1.02). Sternum (length / width) 1.12 / 1.15 (2.00 / 1.67). Labium (length / width) 0.42 / 0.40 (0.62 / 0.58). Legs: length of segments (femur + patella / tibia + metatarsus + tarsus = total length): I 1.67 + 2.00 + 1.33 + 0.67 = 5.67, II 1.60 + 1.80 + 1.27 + 0.60 = 5,27, III 1.53 + 1.67 + 1.33 + 0.60 = 5.13, IV 2.33 + 2.60 + 2.00 + 0.80 = 7.73, (I 1.53 + 2.00 + 1.40 + 0.80 = 5.73, II 1.47 + 1.67 + 1.33 + 0.67 = 5.14, III 1.40 + 1.47 + 1.20 + 0.60 = 4.67, IV 1.93 + 2.33 + 1.93 + 0.73 = 6.92). Variation. Females (males) (range, mean ± s. d.): TL 4.26 – 5.32, 4.82 ± 0.40; CL 2.00 – 2.67, 2.31 ± 0.25; CW 1.53 – 2.07, 1.80 ± 0.22 n = 8 (TL 3.99 – 5.05, 4.32 ± 0.49; CL 2.00 – 2.33, 2.18 ± 0.14; CW 1.40 – 1.80, 1.61 ± 0.17 n = 4).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF81FFEAFF1B524DA1AADA80.taxon	distribution	Distribution. Known only from the eastern central Argentina (Santa Fe and northeastern Buenos Aires) and northern Paraguay (Fig. 27).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF8BFFE8FF1B517BA4AFDEB4.taxon	materials_examined	Type material. Male holotype from Argentina, Entre Ríos, Diamante, Parque Nacional Predelta, bañado al pie de bosque de barranca, S 32.12154 º W 60.62946 º (GPS, area + / - 10 m), elev. 4 m (GPS), 23 – 26. iii. 2011, Rubio, G., Izquierdo, M. & Piacentini, L., hand collected in rushes (MACN-Ar 28786) and three female paratypes of the same data (MACN-Ar 28784).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF8BFFE8FF1B517BA4AFDEB4.taxon	etymology	Etymology. The specific name is a noun in apposition meaning rushes in the Guarani language referring to the habitat where the type specimens was collected (Fig. 1 c).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF8BFFE8FF1B517BA4AFDEB4.taxon	materials_examined	Other material examined. ARGENTINA. Santa Fe: Isla Santa Fe, Zanetic, A. & Goloboff, P. A., 15. viii. 1981, 1 ♂ 4 ♀ (MACN-Ar 22708). Entre Ríos: Brazo Largo, Zanetic, A. & Goloboff, P. A., 12. vii. 1979, over a “ serrucheta ”, Eryngium sp. on the water, 1 ♀ (MACN-Ar 22745); same data, 1 ♀ (MACN-Ar 22752); Diamante, P. N. Predelta, bañado al pie de bosque de barranca, S 32.12154 º W 60.62946 º (GPS, area + / - 10 m), elev. 4 m (GPS), 23 – 26. iii. 2011, Rubio, G., Izquierdo, M. & Piacentini, L., hand collected in rushes, 1 ♂ (MACN-Ar 28787); same data, 1 ♂ (MACN-Ar 28788); same data, 3 ♂ 2 ♀ (MACN-Ar 28789); same data, 1 ♂ (MACN-Ar 28620); same data, 1 ♀ (MACN-Ar 28621); bañado en Laguna Las Piedras, 6 km S Diamante, S 32.12496 ° W 60.62878 ° (GPS, + - 150 m), elev. 12 m, 29. iv- 1. v. 2013, Ramírez, M. J., Piacentini, L. N., González, M. E., Laborda, A. & Aisen, S., collected on vegetation over the water, 1 ♂ (MACN-Ar 30648); same data, 4 ♂ 5 ♀ 2 immatures (MACN-Ar 30377); same data, 1 ♂ (MACN-Ar 30647); same data, 2 ♀ 1 immature (MACN-Ar 30649); same data, 1 ♂ 2 ♀ (MACN-Ar 30336); same data, 1 ♀ (MACN-Ar 30308); same data, 1 ♂ (MACN-Ar 30307); same data, 1 ♀ (MACN-Ar 30333); same data, 1 ♀ 1 eggsac (MACN-Ar 30646); same data, 1 ♂ (MACN-Ar 30645); Parque Nacional Predelta, bañado cerca de entrada, 6 km S Diamante, S 32.12194 ° W 60.62996 ° (GPS, + - 20 m), elev. 10 m, 30. iv. 2013, same collectors, 1 ♀ (MACN-Ar 30650); same data, 2 ♂ (MACN-Ar 30380); same data, collected in marshes, 5 ♀ 4 immature (MACN-Ar 30379). Buenos Aires: San Pedro, Vuelta de obligado, Olejnik, N. & López, N., 11. iv. 2009, 1 ♂ (MACN-Ar 25720); same locality and collectors, 21. iii. 2009, 1 ♀ (MACN-Ar 30640); Tigre, arroyo Espera, Galiano, M. E., iv. 1983, en camalotes, 1 ♂ 1 immature (MACN-Ar 22720); same data, 1 ♀ 1 eggsac (MACN-Ar 22781); same data, 3 ♀ (MACN-Ar 22783); Tigre, Bachmann, A., 30. xi. 1952, 1 ♀ (MACN-Ar 22746); Delta, Río Sarmiento, Oliva, A., over Pistia sp. and Sellenia sp. 1. iv. 1983, 1 ♀ (MACN-Ar 22759); Delta, en una isla, Bachmann, A., iv. 1977, 1 ♀ (MACN-Ar 22764); Delta, Estación experimental INTA, Bachmann, A., vii. 1968, 1 ♀ (MACN-Ar 22772); same locality and collector, 20 – 28. i. 1982, 1 ♀ 1 eggsac (MACN- Ar 22773); same locality and collector, 25. ii. 1973, 2 ♀ 1 immature (MACN-Ar 22779); Campana, Reserva Natural Otamendi, Estación Río Luján, - 34,277352 º - 58,88054 º (WG 84), elev. 5 m, Ramírez, M. J., Grismado, C., Piacentini, L., Soto, E., González-Marquez, M. E., Guala, M., Andía Navarro, J. M., López, N., Ortega Insaurralde, I. & Zermoglio, P., 9. ix. 2010, in marsh and grassland, 2 ♂ (MACN-Ar 28439); same data, 1 ♀ (MACN-Ar 28437); same data, 1 ♀ (MACN-Ar 28438); same data, 1 ♀ (MACN-Ar 28436); Playa Núñez, Castillo, F., viii. 1936, 3 ♂ 1 immature (MACN-Ar 22714). Ciudad Autónoma de Buenos Aires: Reserva Ecológica Costanera Sur, S 34 º 35 ' 47 '' W 58 º 21 ' 36,8 '', Canal Viamonte, Turienzo, P., 13. x. 2008, collected with pitfall traps, 1 ♀ (MACN-Ar 28472); same data, 2 ♀ (MACN-Ar 28447); same data, Mamani, A. & Turienzo, P., 10. xi. 2008, 1 ♀ (MACN-Ar 28443); same data, 29. ix. 2009, 1 ♀ (MACN-Ar 28474); same data 26. v. 2008, 1 ♀ (MACN-Ar 28475); same data, Mamani, A., 27. x. 2008, 1 ♀ (MACN-Ar 28442); same data, Mamani, A., 27. x. 2008, 3 ♀ 1 immature (MACN-Ar 28473); same data, Mamani, A., 27.10.2008, 1 ♀ 2 immatures (MACN-Ar 28476); same data, Mamani, A. & Turienzo, P., 1. ix. 2008, 1 ♀ (MACN-Ar 28477).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF8BFFE8FF1B517BA4AFDEB4.taxon	diagnosis	Diagnosis. This species can be differentiated from the other species of Agalenocosa by the combination of the large body size (TL 6.38 – 9.98), the presence of 0 - 2 - 0 ventral spines on the tibia I, whereas in the rest of the species has 2 - 0 - 0 or 2 - 2 - 0 and by the long setae covering almost all the body. Males resemble those of A. tricuspidata in the shape of the elements of the bulb, but can be differentiated by the shorter dorsal branch of the embolic division (Fig. 22 d).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF8BFFE8FF1B517BA4AFDEB4.taxon	description	Description. Male. Holotype (MACN-Ar 28786). Carapace light brown with pale brown median band and with a Y-shaped dark mark that extends from the fovea to the PLE, pale submarginal bands (as in Figs 21 d – f). Sternum yellow, sparsely covered with brown bristles, more abundant on the margins. Labium dark grey. Chelicerae lighter than labium, light brown, covered with brown bristles. Dorsum of abdomen brown with longitudinal irregular pale marks along the midline (as in Figs 21 d – f). Venter pale brown. Legs pale brown without annulations, densely covered with long setae. Leg formula IV> I> II> III. Spination: femur I p 0 - 0 - d 1 d 1 - 1 - 1, II p 0 - d 1 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1, III p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, IV p d 1 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1; patella II d 1 - 1 p 1, III p 1 d 1 - 1 r 1, IV p 1 d 1 - 1 r 1; tibia I v 0 - 2 - 0, II p d 1 - 0 - d 1 v 0 - 2 - 0, III p 0 - d 1 - 1 d 1 - 0 - 1 r 0 - d 1 - d 1 v p 1 - 2 - 2 ap, IV p 0 - d 1 - 1 d 1 - 0 - 1 r 0 - 1 - 1 v 2 - 2 - 2 ap; metatarsus I v 2 - 2 - 2 ap, II p d 1 - 0 - 2 ap r 0 - 0 - v 1 ap v 2 - 2 - 1 ap, III p d 1 - 1 - 2 ap r d 1 - 1 - 2 ap v p 1 - 2 - 1 ap, IV p d 1 - d 1 - 2 ap r d 1 - d 1 - 2 ap v 2 - 2 - 1 ap. Palp. Basal apophysis of the palpal tibia rounded, medium sized (as in Fig. 8 e). Tegulum with a shallow prolateral furrow (as in Figs 22 c – d). Subtegulum in the meso-prolateral part of the alveolus. Ventral branch of the median apophysis rounded, dorsal branch reduced to a transverse beak-like protuberance directed retrolaterally (as in Fig. 22 d). Terminal apophysis short, pointing retrolaterally (as in Fig. 22 d). Female (MACN-Ar 28437). Colouration in ethanol as in male (Figs 21 a – c). Leg formula IV> I> II> III. Spination: femur I p 0 - 0 - d 1 d 1 - 1 - 1, II p 0 - d 1 - d 1 d 1 - 1 - 1, III p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, IV p d 1 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1; patella I d 0 - 1, II d 1 - 1, III p 1 d 1 - 1 r 1, IV p 1 d 1 - 1 r 1; tibia I v 0 - 2 - 0 d 1 - 0 - 1, II p d 1 - 0 - d 1 v r 1 - r 1 - 0, III p 0 - d 1 - 1 d 1 - 0 - 1 r 0 - d 1 - d 1 v p 1 - p 1 - 2 ap, IV p 0 - d 1 - 1 d 1 - 0 - 1 r 0 - 1 - 1 v p 1 - 2 - 2 ap; metatarsus I v 2 - 2 - 2 ap, II p d 1 - 0 - 2 ap r 0 - 0 - v 1 ap v 2 - 2 - 1 ap, III p d 1 - 1 - 2 ap r d 1 - 1 - 2 ap v p 1 - 2 - 1 ap, IV p d 1 - d 1 - 2 ap r d 1 - d 1 - 2 ap v 2 - 2 - 1 ap. Epigynal plate without pockets or atria, copulatory openings with the anterior part of the internal margin sclerotised and a lightly sclerotised area posteriorly (Fig. 22 a). Epigynal plate simple, rounded atria with the anterior margin sclerotised, copulatory opening located in the middle of the external margin (Fig. 22 a). Vulva: head of spermatheca broad, longer than the stalk. Vulval chambers rounded, behind to the base of spermathecae in dorsal view (Fig. 22 b). Measurements. Female, MACN-Ar 28437 (male, Holotype): TL 9.04 (7.98), CL 3.93 (4.20), CW 3.07 (3.07), CH 1.20 (1.20), AL 4.33 (3.40). Eyes: AME 0.17 (0.17), ALE 0.10 (0.13), PME 0.27 (0.30), PLE 0.17 (0.23). Row of eyes: AER 0.73 (0.73), PME 0.75 (0.75), PLE 1.17 (1.08). Sternum (length / width) 2.13 / 1.67 (2.13 / 1.67). Labium (length / width) 0.65 / 0.75 (0.67 / 0.53). Legs: length of segments (femur + patella / tibia + metatarsus + tarsus = total length): I 3.07 + 3.73 + 2.40 + 1.13 = 10.33, II 2.67 + 3.27 + 2.27 + 1.07 = 9.28, III 2.67 + 3.00 + 2.33 + 1.07 = 9.07, IV 3.53 + 4.33 + 3.60 + 1.20 = 12.66, (I 3.00 + 4.00 + 2.33 + 1.27 = 10.60, II 2.73 + 3.40 + 2.53 + 1.07 = 9.73, III 2.67 + 3.20 + 2.53 + 1.07 = 9.47, IV 3.47 + 4.40 + 3.87 + 1.20 = 12.94). Variation. Females (males) (range, mean ± s. d.): TL 7.05 – 9.98, 8.16 ± 1.00; CL 3.27 – 5.13, 3.93 ± 0.60; CW 2.40 – 3.93, 3.02 ± 0.47 n = 10 (TL 6.38 – 8.51, 7.79 ± 0.58; CL 3.40 – 4.67, 4.14 ± 0.39; CW 2.53 – 3.40, 3.11 ± 0.24 n = 10).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF8BFFE8FF1B517BA4AFDEB4.taxon	distribution	Distribution. Known only from eastern central Argentina (Santa Fe, Entre Ríos and Buenos Aires Provinces) (Fig. 27). Natural history. This species was collected in a marsh, hidden at the base of reedmace (Typha angustifolia), in the Predelta National Park (Fig. 1 c) and in pitfall traps on the Costanera Sur Natural reservation.	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF8FFFEDFF1B503CA4D0DFA7.taxon	materials_examined	Type material. Female holotype from Argentina, Santa Fe, Las Gamas, 20 km W Vera, Ramírez, M. J. & Faivovich, J., 27. x. 1994 (MACN-Ar 22732) and two female paratypes with three eggsacs with the same location and data as the holotype (MACN-Ar 30548).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF8FFFEDFF1B503CA4D0DFA7.taxon	etymology	Etymology. The specific epithet is a noun in apposition taken from the type locality.	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF8FFFEDFF1B503CA4D0DFA7.taxon	diagnosis	Diagnosis. Females of A. gamas sp. nov. resemblesthose of A. gentilis by the coloration pattern (Figs 17, 23), but can be recognised by the small lightly sclerotised area that surrounds the copulatory openings and the relatively long, nearly parallel, anteriorly directed stalks of the spermathecae and the small, rounded heads (Fig. 23 d).	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF8FFFEDFF1B503CA4D0DFA7.taxon	description	Description. Male. Unknown. Female. Holotype (MACN-Ar 22732). Carapace brown with diffuse light brown median band and with a Yshaped dark mark that extends from the fovea to the PLE, pale submarginal bands (Figs 23 a – c). The border of the carapace clothed with white pubescence (Figs 23 a – c). Sternum yellow, sparsely covered with brown bristles, more abundant on the margins. Labium dark grey. Chelicerae lighter than labium, light brown, covered with brown bristles. Dorsum of abdomen brown with irregular pale dots, longitudinal lines of white pubescence on its lateral sides, and two lines of bunches of glistening setae on its posterior part (Figs 23 a – c). Venter pale brown. Legs pale brown without annulations. Leg formula IV> I> II> III. Spination: femur I p 0 - 0 - d 1 d 1 - 1 - 1, II p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, III p d 1 - 0 - d 1 d 1 - 1 - 1 r d 1 - 0 - d 1, IV p d 1 - 0 - d 1 d 1 - 1 - 1 r 0 - 0 - d 1; patella I d 0 - 1 II p 1 d 1 - 1, III p 1 d 1 - 1 r 1, IV p 1 d 1 - 1 r 1; tibia I v 2 - 2 - 0, II p d 1 - 0 - d 1 v 2 - 2 - 0, III p d 1 - 0 - 1 d 1 - 1 - 0 r 0 - 1 - 1 v p 1 - p 1 - 2 ap, IV p 0 - d 1 - d 1 d 1 - 0 - 1 r 0 - 1 - 1 v p 1 - p 1 - 2 ap; metatarsus I v 2 - 2 - 2 ap, II p 1 - 0 - 2 ap r 0 - 0 - 1 ap v 2 - 2 - 1 ap, III p d 1 - d 1 - 2 ap r d 1 - d 1 - 2 ap v 2 - 2 - 1 ap, IV p d 1 - 1 - 1 d 0 - 0 - 2 r d 1 - 1 - 1 v 2 - 2 - 1 ap. Epigynal plate without pockets or atria, copulatory openings with a lightly sclerotised area posteriorly (Fig. 23 b). Vulva: head of spermatheca rounded, stalk of spermathecae nearly parallel and longer than the head (Fig. 23 e). Vulval chamber small and rounded, next to the base of spermathecae in dorsal view (Fig. 23 e). Measurements. Female, MACN-Ar 22732: TL 5.32, CL 2.60, CW 1.87, CH 0.93, AL 2.53. Eyes: AME 0.12, ALE 0.08, PME 0.18, PLE 0.15. Row of eyes: AER 0.48, PME 0.50, PLE 0.72. Sternum (length / width) 1.42 / 1.08. Labium (length / width) 0.40 / 0.38. Legs: length of segments (femur + patella / tibia + metatarsus + tarsus = total length): I 2.07 + 2.47 + 1.67 + 0.93 = 7.14, II 1.87 + 2.13 + 1.33 + 0.80 = 6.13, III 1.47 + 2.07 + 1.47 + 0.80 = 5.81, IV 2.53 + 3.20 + 2.67 + 1.07 = 9.47. Variation. Females (range, mean ± s. d.): TL 4.39 – 5.32, 4.97 ± 0.50; CL 2.00 – 2.60, 2.40 ± 0.35; CW 1.40 – 1.87, 1.65 ± 0.24 n = 3.	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
367587E8FF8FFFEDFF1B503CA4D0DFA7.taxon	distribution	Distribution. Only known from the type locality, in northern Santa Fe Province (Fig. 27)	en	Piacentini, Luis N. (2014): A taxonomic review of the wolf spider genus Agalenocosa Mello-Leitão (Araneae, Lycosidae). Zootaxa 3790 (1): 1-35, DOI: 10.11646/zootaxa.3790.1.1
