identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
35508796FFF5FFBCFF79FB8DFDB7FB39.text	35508796FFF5FFBCFF79FB8DFDB7FB39.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apseudomorpha Sieg 1980	<div><p>Suborder Apseudomorpha Sieg, 1980</p><p>Superfamily Apseudoidea Leach, 1814</p></div>	https://treatment.plazi.org/id/35508796FFF5FFBCFF79FB8DFDB7FB39	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Morales-Núñez, Andrés G.;Heard, Richard W.;Bird, Graham J.	Morales-Núñez, Andrés G., Heard, Richard W., Bird, Graham J. (2019): Two new apseudomorphan species (Crustacea: Tanaidacea: Metapseudidae) from Mo‘orea Island (Society Islands, French Polynesia) with taxonomic keys. Zootaxa 4564 (1): 213-247, DOI: 10.11646/zootaxa.4564.1.8
35508796FFF5FFBFFF79FAA7FDA3FF7F.text	35508796FFF5FFBFFF79FAA7FDA3FF7F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apseudomorpha Miller 1940	<div><p>Genus Apseudomorpha Miller, 1940</p><p>Type-species. Apseudomorpha oahuensis Miller, 1940 .</p><p>Species. Apseudomorpha albida (Shiino, 1951); A. avicularia (Barnard, 1914); A. drummi sp. nov.; A. fontainei Guţu, 1987; A. glebosa (Menzies, 1953); A. hirsuta (Stebbing, 1910); A. magdalenensis (Menzies, 1953); A. martinicana Guţu, 2009; A. negoescuae Guţu, 2007; A. oahuensis Miller, 1940; A. ortizi Guţu, 2006; A. timaruvia (Chilton, 1882); A. veleronis (Menzies, 1953); A. vestafricana Guţu, 2006 .</p><p>Diagnosis. (modified from Guţu 2006). Body subcylindrical. Carapace usually with short and denticulate rostrum; ocular lobes well-developed containing visual elements. Pereonites wider than long. Pleon distinctly narrower than pereon, with five unfused, short, pleonites; pleotelson short, subcylindrical. Antennule biramous, usually shorter than cephalothorax; peduncle having one or more spiniform apophyses on inner margin of article-1; article-2 longer than article-3. Antenna with small squama. Mandible having thick palp with articles. Maxillule with palp having two articles. Cheliped and pereopod-1 lacking exopod. Cheliped sexually dimorphic; male strongly developed and subequal in size, basis appearing sub-ovoid, carpus stout and short, propodus and dactylus forming large chela; female having cheliped distinctly smaller and less robust than male, carpus narrow. Pereopod-1 subcylindrical, slightly large and generally similar to pereopods 2–3. Pleopods absent or reduced in size, when present (up to five pairs), uniramous or biramous. Uropods with short biramous rami</p><p>Remarks. Within the subfamily Metapseudinae, Apseudomorpha can be distinguished from the superficially similar genera Julmarichardia Guţu, 1989, Pseudoapseudomorpha Guţu, 1991, and Plectrocopus Guţu, 2006) by the absence of an exopod on the cheliped.</p></div>	https://treatment.plazi.org/id/35508796FFF5FFBFFF79FAA7FDA3FF7F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Morales-Núñez, Andrés G.;Heard, Richard W.;Bird, Graham J.	Morales-Núñez, Andrés G., Heard, Richard W., Bird, Graham J. (2019): Two new apseudomorphan species (Crustacea: Tanaidacea: Metapseudidae) from Mo‘orea Island (Society Islands, French Polynesia) with taxonomic keys. Zootaxa 4564 (1): 213-247, DOI: 10.11646/zootaxa.4564.1.8
35508796FFF7FFADFF79FF44FE37F9F7.text	35508796FFF7FFADFF79FF44FE37F9F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apseudomorpha drummi Morales-Núñez & Heard & Bird 2019	<div><p>Apseudomorpha drummi sp. nov.</p><p>(Figs 2 ̄9)</p><p>Material examined. Type material. Holotype— ovigerous ♀, TL 1.7 mm, (UF Arthropoda 49366), Stn BIZ-484 (Biocode No. MBIO40746; BMOO-14503: UF: Arthropoda: 28959), (-17.4824; -149.824), depth 1–2 m, lagoonal patch reef (in coral rubble/ Halimeda /green tufting algae), collector Charlotte Watson, November-01-2010, Mo‘orea Island, mouth of Cook's Bay.</p><p>Paratypes — Non-ovigerous ♀, TL 1.8 mm, (UF Arthropoda 49367), Stn BIZ-484 (Biocode No. MBIO40746; BMOO-14503: UF: Arthropoda: 28959), same collection data as holotype.— Two non-ovigerous ♀♀ (with oöstegites), TL 1.5 mm (GCRL 06599) and 1.7 mm (AF Arthropoda 49368), Stn BIZ-288 (Biocode No. MBIO30742; BMOO-10893; UF: Arthropoda: 24615), (-17.48968; -149.82587), depth 1–2 m, fringing reef (in rubble), collectors Kristine White, Sharein El-Tourky, and James Thomas, December-15-2009, Mo‘orea Island, Gump reef.—Non-ovigerous ♀ (with oöstegites), TL 1.8 mm, (USNM 1480633), Stn BIZ-288 (Biocode No. MBIO30681; BMOO-10837; UF: Arthropoda: 24601), (-17.48968; -149.82587), depth 1–2 m, fringing reef (in rubble), collectors Kristine White, Sharein El-Tourky, and James Thomas, December-15-2009, Mo‘orea Island, Gump reef.—Two non-ovigerous ♀♀ (with oöstegites), TL 2.2 mm (GCRL 06600) and TL 2.1 mm (USNM 1480634), and one adult ♂, TL 1.6 mm, (AF Arthropoda 49369), Stn BIZ-498 (BMOO-14237), (-17.4869; - 149.81824), depth 10–15 m, reef wall, collector Jenna Moore, November-05-2010, Mo‘orea Island, Cook's Bay Hotel, opposite Gump Station .—Ovigerous ♀, TL 1.4 mm, (USNM 1480635), Stn BIZ-288 (Biocode No. MBIO30683; BMOO-10839; UF: Arthropoda: 24602), (-17.48968; -149.82587), depth 1–2 m, fringing reef (in rubble), collectors Kristine White, Sharein El-Tourky, and James Thomas, December-15-2009, Mo‘orea Island, Gump reef.—Ovigerous ♀ (dissected), TL 1.6 mm, (AF Arthropoda 49370), Stn BIZ-237 (Biocode No. MBIO29351; BMOO-09640), (-17.59026; -149.83972), depth 2 m, fringing reef (under and on rocks), collectors Sarah McPherson, Kristine White, Sharein El-Tourky, Gustav Paulay, Seabird McKeon, and Chris Meyer, December-06-2009, Mo‘orea Island, Atiha Bay).—Adult ♂, TL 1.8 mm, (USNM 1480636), Stn BIZ-487 (Biocode No. MBIO40726; BMOO-13318; UF: Arthropoda: 28816), (-17.51012; -149.85129), depth 0.5–1 m, collectors Arthur Anker, Charlotte Watson, Gary Poore, and John Slapcinsky, October-31-2010, Mo‘orea Island, E side of Opunohu Bay, about 1 km N of Criobe, near white tower in water .</p><p>Additional material —adult ♂ (damaged), TL 1.8 mm, Stn BIZ-487 (Biocode No. MBIO40726; BMOO-13318; UF: Arthropoda: 28816), (-17.51012; -149.85129), depth 0.5–1 m, collectors Arthur Anker, Charlotte Watson, Gary Poore, and John Slapcinsky, October-31-2010, Mo‘orea Island, E side of Opunohu Bay, about 1 km N of Criobe, near white tower in water.—Non-ovigerous ♀ (with oöstegites), TL 1.5 mm,, Stn BIZ-288 (Biocode No. MBIO30676; BMOO-10833; UF: Arthropoda: 24599), (-17.489683; -149.825871), depth 1–2 m, fringing reef (in rubble), collectors Kristine White, Sharein El-Tourky, and James Thomas, December-15-2009, Mo‘orea Island, Gump reef.</p><p>Diagnosis. Female. Rostrum convex with broad base. Antennule main flagellum with three articles. Mandible palp article-2 with five setulate setae on inner margin. Labium palp with small spiniform setae on sub-proximal margin. Maxillipedal palp article-1 outer margin with spiniform seta. Pereopod-1 basis with dorso-proximal margin lacking blunt, spiniform process; carpus and propodus with ventral margin having two and four spiniform setae, respectively. Pleonites -2 and -3 with pleura having long distal seta. Pleopods biramous, exopod and endopod each with single long seta.</p><p>Male. Cheliped propodus with one simple and one bipinnate spiniform setae on mid-medial margin near articulation of dactylus.</p><p>Etymology. Named on honor of our friend and colleague David T. Drumm in recognition of his work on Tanaidacea and other invertebrate groups.</p><p>Type locality. Mouth of Cook's Bay (-17.4824; -149.824), Mo‘orea Island, Society Island, French Polynesia.</p><p>Distribution. Known from the shallow coastal waters of Mo‘orea Island at depths ranging from 1 to 15 m.</p><p>Description. Based on ovigerous ♀ (holotype (UF Arthropoda 49366) and paratype (UF Arthropoda 49370).</p><p>Body (Fig. 2). Length 1.7 mm, about 4.1 times ltb.</p><p>Cephalothorax (Fig. 2A ̄B). About 25% of TL, shorter than combined lengths of pereonites 1–3, just longer than broad; rostrum convex with broad base, with small denticles and median short trifurcate tip (Fig. 2B); eyelobes well defined, visual elements present. Carapace with simple seta near each ocular lobe and simple seta on each mid-proximal face; branchial region with dorsal red-pigmented area (retained in ethanol).</p><p>Pereon (Fig. 2A). About 60% of TL, all pereonites wider than long, all with two simple setae on sub-distal dorsal margin; pereonite-1 wider than others; pereonites 2̄6 longer than pereonite-1, with one small tuberculiform apophysis ending in a seta on each anterolateral margin; pereonite-6 shortest.</p><p>Pleon (Figs 2A, C, 8A). About 6% of TL, combined lengths of pleonites 1–5 sub-equal to pereonite-6; all pleonites sub-equal, wider than long, bearing pleopods (Fig. 8A); pleonite-1 laterally rounded with small simple seta (Fig. 2C); pleonites 2–5 epimera with bifid tip (Fig 2C); pleonites 2 and 5 with two (one long and one small) simple setae (Fig. 2C); pleonites 3–4 with one small simple seta (Fig. 2C).</p><p>Pleotelson (Figs 2A, C, 8A). About 9% of TL, longer than pereonite-6 (Fig. 2A); dorso-anteriorly with two tubercles, each with two simple setae (Figs. 2A, 8A); antero-laterally with apophysis with bifid tip and bearing a simple seta distally (Fig. 2C); apex with two sub-distal setae (Fig. 8A).</p><p>Antennule (Fig. 3A). Longer than cephalothorax. Peduncle with four articles. Article-1 almost four times ltb; inner margin with two (one long and one short) medial and one sub-distal pointed apophyses, with five simple setae; dorsal margin with two (one PSS and one simple) proximal setae, two medial simple setae, and row of three sub-distal setae; outer margin with one short medial apophysis and one long distal apophysis, with two cluster of three PSS each. Article-2 about 1.3 times ltb; inner margin with four simple setae, with dorsal sub-distal row of four simple setae and one PSS; outer distal margin with one PSS. Article-3 slightly wider than long; inner distal margin with simple seta; outer margin with two PSS. Article-4 shorter than broad; inner distal margin with two simple setae; dorsal margin with two simple setae. Outer flagellum with three articles. Article-1 just longer than broad, with outer distal simple seta. Article-2 about1.4 times ltb; inner distal margin with simple seta; outer distal margin with two simple setae. Article-3 about 2.7 times ltb, distally with one PSS, two aesthetascs, and five simple setae of various lengths. Inner flagellum uniarticulate, 3.2 times ltb, distally with two PSS and four simple setae of unequal lengths.</p><p>Antenna (Fig. 3B). With eight articles. Article-1 inner margin with mid simple seta and distal expansion; outer margin with distal denticles. Article-2 inner margin with distal apophysis; outer margin with two simple setae; squama present, just longer than article-3, with two distal simple setae of unequal lengths. Article-3 wider than long; inner sub-distal margin with simple seta. Article-4 inner distal margin with one PSS. Article-5 about 1.5 times ltb, longer than article-4, inner distal margin with two long simple setae of unequal lengths and two PSS; outer distal margin with two PSS. Article-6 half-length of article-5; outer distal margin with long simple long seta. Article-7 about 1.6 times ltb, inner distal margin with two simple setae of unequal lengths and one PSS; outer distal margin with simple seta. Article-8 shortest, with three distal setae of various lengths.</p><p>Mouthparts: Labrum (Fig. 3C). Shorter than broad, distally setulate. Mandibles (Fig. 3D ̄J). Left mandible incisor with five denticles (Fig. 3D, J); lacinia mobilis broad with three uneven denticles (Fig. 3D ̄E); setiferous lobe with four multi-furcate setae and one simple spiniform seta (Fig. 3D). Right mandible incisor with four uneven denticles (Fig. F̄G); lacinia mobilis bifurcate with several denticles on dorsal margin (Fig. 3F ̄H); setiferous lobe with four multi-furcate spiniform setae (Fig. 3F). Molar process of left and right mandible similar, with grinding surface with well-developed micro-denticles and simple small setae (Fig. 3I). Palp (Fig. 3J) article-1 shortest, inner margin with distal simple seta; article-2, 1.7 times ltb, longest, mid-inner margin with five setulate setae of unequal lengths; article-3 with seven setulate distal setae of different lengths.</p><p>Labium (Fig. 4A). Lobe with inner distal margin setose, with two (sub-proximal) and three (distal) denticles on outer margin. Palp oval, outer lobe setulose, with one small spiniform setae on sub-proximal margin and two distal spiniform setae.</p><p>Maxillule (Fig. 4B). Inner endite with five setulate distal setae, margins finely setose. Outer endite with eleven (one small) distal spiniform setae and two sub-distal setulose setae, margins finely setose; palp biarticulated with two sub-distal setae and one distal whip-like seta.</p><p>Maxilla (Fig. 4C ̄H). Inner and outer margins with three denticles, and outer margin finely setose; outer lobe of moveable endite with two simple setae and six inner pinnate spiniform setae (Fig. 4D); inner lobe of moveable endite with five simple spiniform setae; outer lobe of fixed endite with two bipinnate sub-distal spiniform setae (Fig. 4E) and three trifurcate-plumose spiniform setae (Fig 4F); inner lobe of fixed endite with one bipinnate spiniform seta (Fig. 4G), one inner trifurcate-plumose spiniform seta (Fig. 4H), one bipinnate sub-distal spiniform seta, and row of ~14 basally swollen setae.</p><p>Maxilliped (Fig. 4I ̄K). Coxa short and wide. Basis wider than long, with spiniform seta on distal inner margin, and four denticles on outer distal margin. Palp article-1 shortest, distal inner margin with long (passing article-3) simple seta, and distal outer margin expanded distally with spiniform seta; article-2 longest, inner margin finely setulose proximally, with 13 simple setae of various lengths, outer distal margin with strong spine; article-3, inner margin with five simple setae and three inner setulate spiniform setae; article-4, inner margin with seven setulate spiniform setae (Fig. 4J) and one simple seta. Endite (Fig. 4K) with inner margin having two coupling hooks, row of seven basally-swollen setulate setae, four apically bidentate or grooved spiniform setae with outer setulose margin, one sub-distal bipinnate spiniform setae, and four apically long bidentate or grooved spiniform setae; outer margin setulose with small denticles on mid margin.</p><p>Epignath. Not recovered.</p><p>Cheliped (Fig. 5A ̄D). Basis, 2.0 times ltb; ventral margin with two proximal simple setae, with one spine on mid-margin, and two (one short and one long) sub-distal simple setae; dorsal margin with three simple setae (Fig. 5B). Merus triangular; ventral margin with simple setae sub-distally; mid-outer margin with one simple seta; inner face with two sub-distal simple setae (Fig. 5B). Carpus 3.0 times ltb, as long as basis, widest distally; ventral margin with one small (medial) and two (sub-distal) simple setae; dorsal margin with one sub-distal simple seta. Propodus just longer than broad; outer margin with two (one near articulation of dactylus) simple setae of unequal lengths; dorsal margin with two (one distally) simple setae; inner face with one simple seta and one bipinnate spiniform seta on mid-medial margin near articulation of dactylus (Fig. 5C); fixed finger with three ventral simple setae, with five sub-marginal simple setae on outer incisive margin, claw short; inner face with sub-distal and middorsal (Fig. 5B) bipinnate spiniform setae (Fig. 5D). Dactylus and unguis longer than fixed finger, claw bifid distally; inner face with three bipinnate spiniform setae on sub-distal margin (Fig. 5B).</p><p>Pereopod -1 (Fig. 6A ̄C). Coxa with apophysis, with four simple setae. Basis 4.8 times ltb; distoventral margin with one spine and one long simple seta; dorsal margin with sub-proximal PSS and three simple setae. Ischium wider than long, with two simple setae of unequal lengths on distal ventral margin. Merus 2.5 times ltb, widest distally; ventral margin with three simple setae and one sub-distal bipinnate spiniform seta (Fig. 6B); outer margin with sub-distal simple seta; distodorsal margin with one simple seta and one bipinnate spiniform seta; inner face with sub-distal ventral simple seta (Fig. 6C). Carpus 1.3 times ltb, widest distally, ventral margin with two bipinnate spiniform seta; outer margin with two simple setae: distodorsal margin with three simple setae of varying lengths and bipinnate spiniform seta; inner face with two simple setae on ventral margin (Fig. 6C). Propodus twice as long as broad, ventral margin with four bipinnate spiniform seta and two simple setae; outer margin with two (one in the middle and one distal) simple setae; dorsal margin with mid PSS, two (one sub-distal and one distal) bipinnate spiniform seta, and one sub-distal simple seta; inner face with one setulate spiniform seta and one (longer as dactylus and unguis combined) simple seta (Fig. 6C). Dactylus curved, together with unguis shorter than propodus, dactylus longer than unguis; dactylus with two (one in the middle one sub-distal) ventral setae and small sub-distal denticle.</p><p>Pereopod-2 (Fig. 6D ̄E). Shorter and more gracile than pereopod-1. Coxa lacking apophysis, with two simple setae (Fig. 6D). Basis almost four times ltb; ventral margin with sub-proximal simple seta and two distal simple setae; dorsal margin with four simple setae. Ischium wider than long, with three simple setae of unequal lengths on distal ventral margin. Merus 1.8 times ltb, widest distally; ventral margin with three simple setae, with two subdistal bipinnate spiniform seta; outer margin with sub-distal medial simple seta; distodorsal margin with simple seta; inner face with sub-distal ventral simple seta (Fig. 6E). Carpus slightly ltb, wider distally; ventral margin with four bipinnate spiniform seta; sub-distal outer margin with two bipinnate spiniform setae; dorsal margin with two sub-distal setae and one distal simple seta. Propodus 2.2 times ltb; ventral margin with four bipinnate spiniform seta and four simple setae; dorsal margin with mid PSS, two (one sub-distal and one distal) bipinnate spiniform setae, and one sub-distal simple seta; inner face with one setulate spiniform seta (Fig. 6C). Dactylus curved, together with unguis shorter than propodus, dactylus longer than unguis; dactylus with small sub-distal denticle and one simple seta.</p><p>Pereopod-3 (Fig. 6F ̄G). Similar to pereopod-2 but shorter. Basis with four (one ventral and three dorsal) PSS. Merus shorter.</p><p>Pereopod-4 (Fig. 6H ̄J). Basis five times ltb; ventral margin with one PSS, one mid simple seta, and one subdistal simple seta; outer margin with one simple seta; dorsal margin with three sub-proximal PSS and two simple setae. Ischium wider than long, with three simple setae of unequal lengths on mid ventral margin. Merus 1.5 times ltb; sub-distally with three simple setae and two sub-distal bipinnate spiniform setae; outer margin with one medial simple seta. Carpus twice as long as broad; ventral margin with four bipinnate spiniform setae; distal mid-outer margin with simple setae; dorsal margin with one sub-distal simple seta, and one long simple seta and one bipinnate spiniform seta, distally; inner face with seven (six ventral and one mid sub-distally) bipinnate spiniform setae and one mid sub-distally simple seta (Fig. 6I). Propodus 3.4 times ltb; ventral margin with bipinnate spiniform seta; dorsal margin with mid PSS, and row of six sub-distal pectinate setae (Fig. 6J); inner face with row of five sub-distal pectinate setae (Fig. 6I). Dactylus curved, together with unguis shorter than propodus, dactylus longer than unguis; dactylus with two (one in the middle and one sub-distal) ventral setae and small sub-distal denticle.</p><p>Pereopod-5 (Fig. 7A ̄B). Similar to pereopod-4 but slightly smaller. Carpus with five bipinnate spiniform setae on ventral margin; sub-distal dorsal margin with a row of four simple setae. Propodus with two bipinnate spiniform setae on ventral margin; outer margin with two simple setae; distodorsal margin with two simple setae and one pectinate seta; inner face with row of five sub-distal pectinate setae (Fig. 7B). Dactylus with two sub-distal simple setae on ventral margin.</p><p>Pereopod-6 (Fig. 7C ̄D). Similar to pereopod-5 but slightly longer. Carpus with three bipinnate spiniform seta on ventral margin. Inner face of propodus with row of seven sub-distal pectinate setae (Fig. 7D).</p><p>Pleopods (Fig. 7E ̄F). Five similar, biramous pairs. Basal article 7.7 times ltb, longer than both rami, asetose. Exopod slightly longer than endopod, both uniarticulate, both with long plumose seta distally (Fig. 7F).</p><p>Uropod (Fig. 7G). Biramous. Basal article about twice as long as broad; inner margin with two distal simple setae of unequal lengths; outer margin with one mid and two sub-distal simple setae of various lengths. Exopod of three articles, longer than endopod articles 1̄2 combined; article-1 small, asetose, shorter than article-2 (the division between article-1 and 2 is often indistinct) with simple distal seta on both margins; article-3 with two simple setae of varying lengths distally. Endopod longer than pleon and pleotelson combined, with five articles; article-1 shorter, asetose; article-3 longest; articles 2̄4 bearing two to three simple setae on distal margins; terminal article with two PSS and four simple setae of unequal lengths distally.</p><p>Adult male. Length 1.6 mm. Similar to females except in the size of the chelipeds (paratype, UF Arthropoda 49369).</p><p>Cheliped (Fig. 8A ̄F). Robust. Basis slightly wider than long; ventral margin with three (two sub-proximal and one distal) simple setae, and one spiniform seta; dorsal margin with two simple setae. Merus triangular, mid margin with one simple seta; distoventral margin with three simple setae. Carpus broad and short; distoventral margin with a prolongation and one distal simple seta; outer distal margin with four denticles and one simple seta; inner face with series of ~12 ridges, with two simple setae on mid proximal margin, and two simple setae dorsally. Propodus 1.3 times as wide as long; ventral margin with proximal tridentate apophysis, (Fig. 8B), with two simple setae, and mid simple seta; outer margin with one simple seta near to articulation of dactylus; dorsal margin with two (one mid and one distal) simple setae; inner face with ventral simple seta, with one simple seta and two (one simple and one bipinnate (Fig. 8E)) spiniform setae on mid-medial margin near articulation of dactylus; fixed finger short, with three ventral simple setae, with nine sub-marginal simple setae on outer incisive margin; dorsal margin with two denticle and grasping edge with row of ~seven blade-like pectinate setae (Fig. 8C), claw short; inner face with three ventral simple setae, one sub-distal bipinnate spiniform seta, and three mid-dorsal simple setae (Fig. 8D). Dactylus curved, longer than fixed finger, ventral margin with mid-denticle and grasping edge with row of ~six blade-like pectinate setae having finely pectinate or serrate upper margins, claw with bidentate tip for insertion of tip of dactylus (Fig. 8C); inner face with three bipinnate spiniform setae (Fig. 8F) on sub-distal margin.</p><p>Intraspecific variation. Apseudomorpha drummi sp. nov. exhibits some degree of variation among the individuals examined during this study (Table 1), such as the number of articles on the antennular outer flagellum observed on non-ovigerous females that varied from 2–3 (Table 1), three being the most common. The presence, however, of two articles was only observed on the left antennular flagellum of a non-ovigerous, female paratype (USMN 01480634). Further, the number of articles on the antennular outer flagellum on males varied from 3–4 (four in paratype, USNM 1480636). The uropods also expressed considerable variation. The number of articles occurring in the exopods of a non-ovigerous females, ovigerous females, and males varied from 2–3 (Table 1). The total length of the exopod in relation to that of the endopod varied from being longer as endopodal article-1 to extending to the mid-length of endopod article-3 (Table 1). In some specimens the right endopod is shorter than left.</p><p>Size-distribution. The body sizes of individuals Apseudomorpha drummi measured are presented in Table 1. The non-ovigerous females ranged from 1.5 to 2.1 mm (n=7). Ovigerous females ranged from 1.4 to 1.7 mm (n=3). Adult males varied from 1.6 (Fig. 9) to 1.8 mm (n=3).</p><p>Remarks. Apseudomorpha drummi is the fourteenth member of the genus to be described (Table 2, Key). With A. fontainei, A. glebosa, and A. martinicana, it comprises a group of Apseudomorpha characterized by having exclusively long setae on the epimera of the second and fifth pleonites. The new species from Mo‘orea differs from A. fontainei by having (1) the mandibular palp with article-1 having an inner distal spiniform seta (two in fontainei), (2) biramous pleopods (uniramous in fontainei), (3) a uropodal exopod with three articles (two in fontainei), and (4) the uropodal endopod with five articles (seven in fontainei).</p><p>Apseudomorpha drummi is distinguished from A. glebosa by (1) presence of two and four spiniform setae on the ventral margin of carpus and propodus of pereopod-1, respectively (three and five in glebosa), (2) pereopod-6 with two ventral bipinnate spiniform setae (four in glebosa), (4) each branch of the pleopods with long setae (one branch with one long seta in glebosa), and (5) uropodal exopod article-1 shorter than endopodal article-1 (longer in glebosa).</p><p>* Indicates male known.</p><p>Based on the examination of specimens by G.J. Bird referable to Apseudomorpha timaruvia indicates that this species probably belongs in another genus as it differs from the other species of Apseudomorpha, including A. drummi, in having (1) a long apical seta only on the epimeron of pleonite-3, (2) more numerous antennular articles (ca. six inner, nine outer), and (3) a double row of spines on the carpus and propodus of pereopods 1–3. It is retained in the genus pending its complete redescription (G. J. Bird in prep.). Apseudomorpha drummi can be distinguished from the other species of Apseudomorpha in the following identification key.</p><p>The members of the genus Apseudomorpha are mostly confined to the Pacific and Indian Oceans (Figure 10); however, Heard et al. (2004) reported and illustrated an apparently undescribed species (“ Apseudomorpha sp. A”) with affinities to the Eastern Pacific species A. glebosa Menzies, 1953, from the Atlantic East Coast of Florida. Later, Guţu (2006, 2009) described A. ortizi from Cuba and A. martinica from Martinique Island. Further, Heard et al. (2009) reported A. glebosa or a very similar cryptic geminate species from coral reefs off the Southeast Caribbean Coast of Costa Rica.</p></div>	https://treatment.plazi.org/id/35508796FFF7FFADFF79FF44FE37F9F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Morales-Núñez, Andrés G.;Heard, Richard W.;Bird, Graham J.	Morales-Núñez, Andrés G., Heard, Richard W., Bird, Graham J. (2019): Two new apseudomorphan species (Crustacea: Tanaidacea: Metapseudidae) from Mo‘orea Island (Society Islands, French Polynesia) with taxonomic keys. Zootaxa 4564 (1): 213-247, DOI: 10.11646/zootaxa.4564.1.8
35508796FFE4FFACFF79F917FA53FCEB.text	35508796FFE4FFACFF79F917FA53FCEB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apseudomorpha Miller 1940	<div><p>Key to the known species (females) of Apseudomorpha, modified from (Guţu 2009)</p><p>1. Pereopod-1 dorsal proximal margin of basis with blunt, spiniform process..... A. negoescuae [Indian Ocean: Réunion Island] - Pereopod-1 dorsal proximal margin of basis lacking a blunt, spiniform process..................................... 2 2. Pleonites 1̅5 without a long seta on dorsolateral margins......... A. avicularia [Southeastern Atlantic Ocean: Cape Town] - At least the last pleonite with a long seta on dorsolateral margins................................................ 3 3. Pleopods absent.............................................. A. vestafricana [Eastern Atlantic Ocean: Mauritania] - Pleopods present (uni- or biramous)....................................................................... 4 4. Only the last pleonite with a long seta on distolateral margins................................................... 5 - More than the last pleonite with a long seta on distolateral margins.............................................. 6</p><p>5. First peduncular article of antennule with some dentiform processes on the proximal half of inner margin............................................................................................. A. ortizi [Caribbean Sea: Cuba]</p><p>- First peduncular article of antennule without dentiform processes on the proximal half of inner margin..................................................................................... A. albida [Northwest Pacific Ocean: Japan]</p><p>6. Last four pleonites with a long seta on distolateral margins..................................................... 7</p><p>- At most the last three pleonites with a long seta on distolateral margins........................................... 8</p><p>7. Pleopods biramous, with biarticulate exopodite................... A. magdalenensis [Northwest Atlantic Ocean: Mexico]</p><p>- Pleopods biramous, with uniarticulate exopodite (in male), or uniramous (in female).................................................... A. veleronis [Southeast Pacific Ocean and Northeast Atlantic Ocean: Ecuador, Colombia, and Panama]</p><p>8. Last three pleonites with a long seta on lateral margins......... A. oahuensis [Southern North Pacific Ocean: O‘ahu Island]</p><p>- Only two pleonites (second and fifth) with a long seta on distolateral margins...................................... 9</p><p>9. Pleopods uniramous........................................................ A. fontainei [Red Sea: Aqaba Gulf]</p><p>- Pleopods biramous................................................................................... 10</p><p>10. Pereopod-1 carpus and propodus ventral margin with three and five spiniform setae, respectively; one branch of pleopods with two long setae............................... A. glebosa [Northeast and Southeast Pacific Ocean: Mexico to Ecuador]</p><p>- Pereopod-1 carpus and propodus ventral margin with two and four spiniform setae, respectively; each branch of pleopods with one long setae....................................................................................... 11</p><p>11. Antennule main flagellum with four short, thick articles. Mandible palp article-2 with three setulate setae on inner margin. Labium palp outer lobe without spiniform setae on sub-proximal margin. Maxilliped palp article-1 outer margin with very small spiniform seta. Pereopod-2, ventral margin of propodus with three spiniform setae.......................................................................................... A. martinicana [Caribbean Sea: Martinique Island]</p><p>- Antennule main flagellum with three articles. Mandible palp article-2 with five setulate setae on inner margin. Labium palp outer lobe with small spiniform setae on sub-proximal margin. Maxilliped palp article-1 outer margin with spiniform seta. Pereopod-2, ventral margin of propodus with four spiniform setae....................................................................................... A. drummi sp. nov. [South Pacific Ocean: Mo‘orea Island (French Polynesia)]</p></div>	https://treatment.plazi.org/id/35508796FFE4FFACFF79F917FA53FCEB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Morales-Núñez, Andrés G.;Heard, Richard W.;Bird, Graham J.	Morales-Núñez, Andrés G., Heard, Richard W., Bird, Graham J. (2019): Two new apseudomorphan species (Crustacea: Tanaidacea: Metapseudidae) from Mo‘orea Island (Society Islands, French Polynesia) with taxonomic keys. Zootaxa 4564 (1): 213-247, DOI: 10.11646/zootaxa.4564.1.8
35508796FFE5FFACFF79FBD1FA89F8C1.text	35508796FFE5FFACFF79FBD1FA89F8C1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryptapseudes Bacescu 1976	<div><p>Genu s Cryptapseudes Băcescu, 1976</p><p>Type-species. Cryptapseudes sankarankuttyi Băcescu, 1976 Species. Cryptapseudes acutifrons Băcescu, 1976; C. leroyi David &amp; Heard, 2015; C. mamua sp. nov.; C. romanae Guţu, 1991; C. sankarankuttyi Băcescu, 1976 .</p><p>Diagnosis. “Body small, robust. Antennule having main flagellum with two to five articles; minor flagellum uniarticulate. Antenna with small squama. Maxilliped with palp article-2 having strongly developed, distolateral spiniform seta. Cheliped and pereopod-1 with exopods. Pereonites with long, strongly developed, stiff, dorsolateral setae. Pleonites compressed and completely fused with telson, strongly developed upturned, spiniform process dorsally; five pairs of reduced biramous pleopods present. Sexual dimorphism pronounced; adult males with massive chelipeds; chelipeds of female much smaller than those of male. Uropodal endopod with six or fewer articles, exopod small, uniarticulate” (David &amp; Heard 2015).</p><p>Remarks. The subfamilial status of metapseudid genus Cryptapseudes remains unsettled. Based on the presence of (1) an antennal squama, (2) exopods on the chela and pereopod-1, (3) five pairs of pleopods, and (4) the abdomen and telson being completely fused dorsally, David &amp; Heard (2015: p. 293) questioned the placement of this genus within the Synapseudinae Guţu, 1972, which, as then defined, shared none of these characters. Following the same reasoning, Heard et al. (2018: p. 7) removed Cryptapseudes from the Synapseudinae indicating that it shares more characters in common with the subfamilies Metapseudinae Lang, 1970 and Chondropodinae Guţu 2008 . The systematic status of Cryptapseudes and other genera within the Metapseudidae is being reevaluated and revised by Piotr Jóźwiak (pers. comm. 2016).</p><p>It should be noted that David &amp; Heard (2015: fig. 4D) inadvertently confused pereopod-4 of the female with that of the already described pereopod-3 (see their fig. 4C). Pereopod-4, which is morphologically similar to pereopods 5 and 6, is correctly indicated in their lateral views of the male and female (see their fig. 1A, C).</p></div>	https://treatment.plazi.org/id/35508796FFE5FFACFF79FBD1FA89F8C1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Morales-Núñez, Andrés G.;Heard, Richard W.;Bird, Graham J.	Morales-Núñez, Andrés G., Heard, Richard W., Bird, Graham J. (2019): Two new apseudomorphan species (Crustacea: Tanaidacea: Metapseudidae) from Mo‘orea Island (Society Islands, French Polynesia) with taxonomic keys. Zootaxa 4564 (1): 213-247, DOI: 10.11646/zootaxa.4564.1.8
35508796FFE6FF9DFF79FF44FE53FDC3.text	35508796FFE6FF9DFF79FF44FE53FDC3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryptapseudes mamua Morales-Núñez & Heard & Bird 2019	<div><p>Cryptapseudes mamua sp. nov.</p><p>(Figs 11 ̄21E, H, J)</p><p>Material examined. Type Material. Holotype— ovigerous ♀, TL 1.7 mm, (UF Arthropoda 49371), Stn BIZ-487 (Biocode No. MBIO40726; BMOO-13318; UF: Arthropoda: 28816), (-17.51012; -149.85129), depth 0.5–1 m, fore-reef, collectors Arthur Anker, Charlotte Watson, Gary Poore, and John Slapcinsky, October-31-2010, Mo‘orea Island, E side of Opunohu Bay, about 1 km N of Criobe, near white tower in water.</p><p>Paratypes —non-ovigerous ♀ (with oöstegites), TL 1.5 mm, (UF Arthropoda 49372) and one subadult(?) ♂, TL 1.2 mm, (UF Arthropoda 49373), Stn BIZ-508 (Biocode No. MBIO40637; BMOO-13955; UF: Arthropoda: 28838), (-17.47610; -149.83100), depth 10–11 m, fore-reef (in/on ruble), collector Kristine White, November-07- 2010, Mo‘orea Island, off channel MPA marker.— One non-ovigerous ♀, TL 1.0 mm, (USNM 1480637), Stn BIZ- 795.B (Biocode No. MBIO59835; BMOO-18252; UF: Arthropoda: 33940), (-17.47593; -149.84190), depth 13 m, outer reef slope (in ARMS 9), collectors BioCode IZ /ARMS team, February-04-2012, Mo‘orea Island, ARMS 9 site.—One ovigerous ♀, TL 1.2 mm, (USNM 1480638), Stn BIZ-485 (Biocode No. MBIO40747; BMOO-14504; UF: Arthropoda: 28960), (-17.49410; -149.86200), depth 10–20 m, small caves/hollows, collectors Jenna Moore, Mandy Bemis, and Seabird McKeon, November-02-2010, Mo‘orea Island, Opunohu Bay, wall near pylon .</p><p>Additional material. —Ovigerous ♀ (dissected), TL 1.7 mm, same collection data as holotype (remnants in collection of RWH). —Non-ovigerous ♀ (with oöstegites, damaged), TL 1.5 mm, same collection data as holotype. —Non-ovigerous ♀, TL 1.1 mm,, Stn BIZ-795.B (Biocode No. MBIO59835; BMOO-18252; UF: Arthropoda: 33940), (-17.47593; -149.84190), depth 13 m, outer reef slope (in ARMS 9), collectors BioCode IZ/ARMS team, February-04-2012, Mo‘orea Island, ARMS 9 site.</p><p>Diagnosis. Female. Rostrum expanded anteriorly into rosette cluster of short spines. Antennule with outer flagellum having two or three articles. Antenna with seven to nine articles. Mandible palp distal article with 5–6 setae. Maxilliped palp with article-1 having small spiniform seta on distolateral margin. Pereopods 2–3 with dorsal margin of basis having one or two blunt, spiniform process, on sub-proximal margin, and one blunt, spiniform process, on sub-distal margin. Pleotelson with two dorsolateral apophyses, each bearing two tubercles, long stiff seta and two simple setae; postero-dorsal surface with strong, apophysis bearing one tubercle and four simple setae. Subadult(?) male. Body similar to female. Cheliped more robust that female, propodus bearing two denticles on sub-distal ventral margin.</p><p>Etymology. After ‘ Mamua ’, the name used by the English poet Rupert Brooke for one of his Polynesian lovers (although probably not the Mo’orean, Taatamata—Tumatataata Topotofaerani) in the poem ‘Tiare Tahiti’ (Mamua, when our laughter ends, ….), during his stay in Papeete, Tahiti, February–April 1914; probably his rendition of Tahitian maimoa —‘favourite’, ‘chosen one’ (Delany 2015: 240); used as noun in apposition.</p><p>Type locality. East side of Opunohu Bay, about 1 km North of Criobe, near white tower in water (-17.5101; - 149.851), Mo‘orea Island, Society Island, French Polynesia.</p><p>Distribution. Known from the shallow coastal waters of the Mo‘orea Island at depths ranging from 0.5 to 20 m.</p><p>Description. Based on ovigerous ♀ (holotype (UF Arthropoda 49371), paratype (UF Arthropoda 49372), and additional topotypic material).</p><p>Body (Fig. 11). Sub-cylindrical, broader than deep. Holotype: TL 1.7 mm, about 3.2 ltb; widest at juncture of carapace and pereonite-1.</p><p>Cephalothorax (Fig. 11 A–B). About 30% of TL, slightly longer than combined lengths of pereonites 1–3; rostral base with anterior margin denticulate; rostrum rosette or mace-like, with five spines (Fig. 10B); eye-lobes well defined, visual elements present. Carapace with simple seta near each ocular lobe, and one dorsal protuberance with two long, stiff setae mid-dorsally; branchial region with dorsal circular red-pigmented area (retained in ethanol after).</p><p>Pereon (Fig. 11A, C ̄D). About 60% of TL, all pereonites wider than long, pereonite-1 widest; pereonites 2̄6 decreasing in width distally; pereonites 1–6 each with low, broadly rounded (lateral) protuberance, each with one to three strongly developed stiff setae (Fig. 11C); pereonites 1̄2 and 4̄6 with two large, stiff setae on anterior dorsal margin (Fig. 11A); pereonite-4 with anterior lateral denticulate projection (Fig. 110A); pereonite-6 with one small spine (Fig. 11C). Pereonites 3 and 5 with hyposphenia (Fig. 11D).</p><p>Pleotelson (Figs 11A, C ̄D, 17A). Reduced, about 10% of TL, shorter than pereonites 5–6 combined (Fig. 11A, C); abdomen not distinguishable from telson (all pleonites fused together and with telson), with five pairs of small, reduced pleopods indicating ventral remnants of pleonites at their attachment and with well-developed pair of uropods (Fig. 17A). Ventrally with five anterior keels (Fig. 11D); sub-proximal dorsal anterior region with pair of simple setae; mid-dorsal region with pair of apophyses each bearing two spiniform setae, long stiff seta, and two simple setae (Fig. 17A). Dorsally with posterior region having apophysis bearing one spiniform seta and four simple setae (Fig. 17A). Mid posterior margin region with three keels (Fig. 10D).</p><p>Antennule (Fig. 12A ̄B). Shorter than carapace. Peduncle with four articles. Article-1 three times ltb; inner margin with proximal row of setules, with medial and distal apophyses bearing two (medial) or three (distal) spiniform setae, and two simple setae; dorsal margin with one sub-proximal PSS and one mid simple seta; outer margin with four PSS and distal simple seta. Article-2 about 1.5 times ltb, distally widest; inner distal margin with one PSS, three simple setae of various lengths, and two spiniform setae; distodorsal margin with simple seta; outer distal margin with three simple setae of unequal lengths and two PSS. Article-3 just ltb, distinctly shorter and narrower than article-2, with distal simple setae at each margin. Article-4 shorter than broad; inner distal margin with simple seta. Outer flagellum (only right described in detail) with two (right; Fig. 12A) or three (left; Fig. 12B) articles. Article-1 about 1.8 times ltb; inner distal margin with one aesthetasc and two simple setae of unequal lengths; outer distal margin with simple seta. Article-2 three times ltb; distally with one aesthetasc and seven simple setae of unequal lengths. Left outer flagellum with one aesthetasc on inner distal margin at each article. Inner flagellum uniarticulate, five times ltb, distally with two simple setae. Left inner flagellum reduced, with two simple setae on outer distal margin.</p><p>Antenna (Fig. 12C) with nine articles. Article-1 having inner sub-distal process with distal denticles. Article-2 about 1.4 times ltb; inner distal margin with one PSS and two spiniform setae; outer mid margin with one PSS; squama present, small, about as long as broad, with two distal simple setae of unequal lengths. Article-3 wider than long; inner distal margin with one simple seta and one spiniform seta. Article-4 twice as long as broad; inner distal margin with one PSS. Article-5 longest, slightly less than three times ltb, distally widest; inner distal margin with three PSS; outer distal margin with two PSS. Article-6 shortest, asetose. Article-7 about 1.2 times ltb; inner distal margin with long simple seta. Article-8 about 2.3 times ltb; inner distal margin with one PSS and long simple seta; outer distal margin with one long simple seta. Article-9 nearly four times ltb, with three simple setae of varying lengths.</p><p>Mouthparts: Labrum not recovered. Mandibles (Fig. 13ĀD). Left mandible with incisor bearing three denticles (Fig. 13A); lacinia mobilis broad with three denticles (Fig. 13B); setiferous lobe with four multi-furcate spiniform setae (Fig. 13A). Right mandible with incisor having four (one acute) denticles (Fig. 13C); lacinia mobilis widely bidentate (Fig. 13C); setiferous lobe with three multi-furcate spiniform setae (Fig. 13C). Molar process of left and right (not illustrated) mandible similar, with grinding surface (Fig. 13A). Palp (13A, D) article-1 as long as article-2, inner margin with sub-distal simple seta; article-2 twice as long as broad, with two sub-distal simple setae on inner margin; article-3, with five (left; Fig. 13A) or six (right; Fig. 13D) setulate distal setae of various lengths.</p><p>Labium (Fig. 13ĒF). Lobe with mid-distal margin setulose, with outer margin serrate (Fig. 14E). Palp oval, outer margin setulose, with three distal setulate setae (Fig. 13F).</p><p>Maxillule (Fig. 13G). Inner endite with five setulate distal setae. Outer endite with ten (one small) distal spiniform setae and two sub-distal setulose setae; palp biarticulate with one sub-distal seta and two distal whip-like setae, distalmost seta longest.</p><p>Maxilla (Fig. 14A ̄E). Outer margin with two denticles and two simple setae, and inner margin with three denticles and one simple seta; outer lobe of movable endite with five inner pinnate spiniform setae (Fig. 14B); inner lobe of movable endite with five spiniform setae; outer lobe of fixed endite with two (one sub-distal and one distal) outer pinnate setae (Fig. 14C), three trifurcate spiniform (Fig. 14D) and two spiniform setae; inner lobe of fixed endite with five bifid spiniform setae (Fig. 14E), and row of ~17 basally swollen setae.</p><p>Maxilliped (Fig. 14F ̄G). Coxa short and wide. Basis bulbous, 1.2 times ltb, widest proximally; inner distal margin with long, stiff seta. Palp article-1 shortest, distal inner margin with long, stiff seta, and distal outer margin expanded distally with small spiniform seta; article-2 longest, inner margin finely setulose proximally, with four simple setae of varying lengths and row of nine distally attenuated spiniform setae, outer distal margin with stout spiniform seta; article-3 globose, smaller than article-2, inner margin with two distal spiniform seta and five welldeveloped, distally attenuated spiniform setae; article-4 narrower, inner margin with seven spiniform setae and two setulate spiniform setae. Endite (Fig. 14G) with inner margin having two coupling hooks (Fig. 14F), row of six basally-swollen setulate setae, four apically bidentate or grooved spiniform setae with outer setulose margin, three long apically bidentate or grooved spiniform setae, and one pinnate spiniform seta; outer margin expanded lateral, setulose, and with four denticles.</p><p>Epignath (Fig. 14H). Sub-oval, stout with distolateral margin having large, anteromedially directed seta.</p><p>Cheliped (Fig. 15A ̄C). Exopod with three articles, third article bearing four plumose setae. Basis 1.6 times ltb; ventral margin with two sub-proximal simple setae, one spiniform seta on mid-margin, and three sub-distal simple setae. Merus triangular, ventral margin with two sub-distal simple setae; mid-outer margin with one simple seta; mid-inner face with one simple seta (Fig. 15B). Carpus 2.5 times ltb, slightly longer than basis, ventral margin with two and one (distal) simple setae; inner distoventral face with one distal simple seta (Fig. 15B). Propodus 1.1 times as wide as long; outer margin with one (near to articulation of dactylus) simple seta; dorsal margin with two (one mid and one distal) simple setae; inner face with one simple seta and one bipinnate spiniform seta on midmedial margin near articulation of dactylus (Fig. 15B); fixed finger with three ventral simple setae, with four submarginal simple setae on outer incisive margin, grasping edge with ~six blade-like pectinate setae, claw well-</p><p>Pereopod -1 (Fig. 16A ̄C). Coxa asetose. Basis 3.2 times ltb; ventral margin with five (distal one longest) curved simple setae; dorsal margin with simple seta, five (left) or six (right) blunt, spiniform processes alternating with plumose setae (Fig. ĀB, respectively); inner face with proximal row of ~six simple setae (Fig. 16B). Ischium wider than long, with one short and one long simple seta on distal ventral margin. Merus 1.7 times ltb; ventral margin with two simple setae and one sub-distal spiniform seta; dorsal margin expanded distally, forming lobe bearing two curved, acutely tipped spiniform setae. Carpus 1.4 times ltb; ventral margin with two spiniform setae, and one simple seta; outer margin with mid-ventral simple seta; dorsal margin expanded distally, forming lobe bearing two curved, acutely tipped spiniform setae, and two simple setae. Propodus 2.7 times ltb; ventral margin with four spiniform setae; outer margin with sub-distal ventral simple seta; dorsal margin with mid PSS, and one spiniform seta and simple seta distally; inner face with bipinnate spiniform seta (Fig. 16C). Dactylus curved, together with unguis shorter than propodus; dactylus longer than unguis, with two ventral spines and one simple seta distally. Exopod with three articles, third article bearing five plumose setae (Fig. 16B).</p><p>Pereopod-2 (Fig. 16D). Shorter and more gracile than pereopod-1. Basis 3.8 times ltb; distoventral margin with simple seta; dorsal margin with one blunt, spiniform processes, on sub-proximal margin, with two simple setae. Ischium wider than long, with two (one short and one long) simple setae on distal ventral margin. Merus 1.8 times ltb; ventral margin with one simple seta and two sub-distal spiniform setae; dorsal margin expanded distally, forming lobe bearing one, acutely tipped simple seta. Carpus about as long as broad; ventral margin with two spiniform setae; outer margin with one mid-dorsal spiniform seta; dorsal margin expanded distally, forming lobe bearing two curved, acutely tipped spiniform setae, and one simple seta. Propodus 3.0 times ltb; ventral margin with four spiniform setae; dorsal margin with mid PSS, one spiniform seta, and simple seta distally. Dactylus curved, together with unguis shorter than propodus; dactylus longer than unguis, with one mid ventral spine, and two ventral simple setae distally.</p><p>Pereopod-3 (Fig. 16E). Similar to pereopod-2 but shorter. Basis ventral margin with three (two proximal and one sub-distal) simple setae; dorsal margin with one blunt, spiniform processes, on sub-proximal margin, and one mid simple seta. Propodus with four spiniform setae on ventral margin.</p><p>Pereopod-4 (Fig. 16F). Basis 5.3 times ltb; ventral margin with one simple seta. Ischium wider than long, with one short and one long simple setae on distal ventral margin. Merus 1.2 times ltb; ventral margin with two (one not shown) sub-distal spiniform setae; dorsal margin with acutely tipped simple seta. Carpus 1.7 times ltb; ventral margin with two spiniform setae; dorsal margin with one simple seta. Propodus almost three times ltb; ventral margin with two spiniform setae; dorsal margin with mid PSS, and three pectinate spiniform and one simple subdistal setae. Dactylus curved, together with unguis shorter than propodus; dactylus longer than unguis, with one mid ventral spine, and two ventral simple setae distally.</p><p>Pereopod-5 (Fig. 16G). Similar to pereopod-4 but slightly longer. Carpus asetose on dorsal margin.</p><p>Pereopod-6 (Fig. 16H). Similar to pereopod-5 but slightly shorter. Basis with three simple setae on ventral margin; mid-dorsal margin with one simple seta. Carpus with small simple seta on distodorsal margin. Propodus with four sub-distal pectinate setae.</p><p>Pleopods (Fig. 17A ̄B). Five similar, greatly reduced pairs, compressed into anterior half of fused pleon. Basal article sub-divided into two sub-articles, asetose, attached proximally to ventral protrusion. Exopod uniarticulate, shorter than endopodal article-1, with one simple seta and long plumose seta distally. Endopod biarticulate, article- 1 as long as article-2, with one long plumose seta on inner distal margin; article-2 with two long plumose setae of varying lengths distally.</p><p>Uropod (Fig. 17A, C). Biramous. Basal article, 2.1 times ltb; inner margin with two distal simple setae (Fig. 16A). Exopod small, uniarticulate, shorter than half length of endopodal article-1, with two long simple setae of unequal lengths distally. Endopod of five articles; articles 1̄2 bearing two PSS; article-2 shortest; articles 3̄5 bearing one PSS; articles 3̄4 with one long simple seta on distal outer margin; article-4 longest; article-5 with three simple setae of unequal lengths.</p><p>Subadult(?) male. Overall similar to adult female, except in having a longer antennule and cephalothorax, the size of the chelipeds, and the armament of the pleotelson (paratype, UF Arthropoda 49373).</p><p>Body (Fig. 18A). Small, ventrally flattened, TL 1.2 mm.</p><p>Cephalothorax (Fig. 18A). About 40% of TL.</p><p>Pereon (Fig. 18A ̄B). About 50% of TL. Pereonite-6 with one dorsal spiniform seta (Fig. 17B) and genital cone.</p><p>Pleotelson (Fig. 18A ̄C). Reduced, about 10% of TL (Fig. 18A), abdomen not distinguishable from telson (all pleonites fused together and with telson) (Fig.17B), with five pairs of small, reduced pleopods indicating ventral remnants of pleonites at their attachment (not illustrated) and with well-developed pair of uropods (Fig. 18B). Ventrally, with ~three anterior keels (Fig. 18B); sub-proximal dorsal anterior region with pair of simple setae and pair of spines, mid-dorsal region with pair of apophyses each bearing one spine, long stiff seta, and simple seta (Fig. 18B). Dorsally, with posterior region having apophysis bearing one spiniform seta and four simple setae (Fig. 18B). Mid posterior margin region with two keels (Fig.18C).</p><p>Antennule (Fig. 18A). Length about equal to that of cephalothorax. Article-1 with medial apophysis bearing two (left) or three (right) spiniform setae, and distal apophyses bearing two spiniform setae.</p><p>Cheliped (Fig. 19A ̄D). Shorter but robust. Exopod with three articles, third article bearing four plumose setae. Basis just longer than broad; ventral margin with two sub-proximal simple setae, one spiniform seta on midmargin, and three sub-distal simple setae; inner face with mid-dorsal small simple seta (Fig. 19B). Merus triangular, ventral margin with one sub-distal simple seta; mid-outer margin with one simple seta; inner face with two mid-dorsal simple setae (Fig. 19B). Carpus 1.6 times ltb, broad and short sub-distal ventral margin with two denticles (one of them apparently broken) and one simple seta between them, and two simple setae; dorsal margin with two sub-proximal simple setae and one distal simple seta; inner face with sub-proximal ventral seta and one small sub-distal simple seta (Fig. 19B). Propodus with palm about just wider than long; mid-ventral margin with two simple setae; mid-outer margin with two simple setae; mid distal margin with two simple setae near to articulation of dactylus; dorsal margin with four (two sub-proximal, one mid, and one distally) simple setae; inner face with simple seta on mid-medial margin (Fig. 19B), and one simple seta and one bipinnate spiniform seta on mid-medial margin near articulation of dactylus (Fig. 19B ̄C); fixed finger with two ventral simple setae, five submarginal simple setae on outer incisive margin, one long simple seta (near to articulation of dactylus), socket present at base of fixed finger for insertion of proximal tooth of dactylus, claw well-developed; inner face with one mid-inner bipinnate spiniform seta (Fig. 19B, D), and two (one bipinnate spiniform and one simple) setae on subproximal grasping edge (Fig. 19B, D). Dactylus and unguis longer than fixed finger and curving downward; inner face with three bipinnate spiniform setae on sub-distal margin (Fig. 19D).</p><p>Intraspecific variation. Cryptapseudes mamua sp. nov. exhibits a notable degree of variation among the eight individuals examined during this study (Table 3). The variation observed includes:</p><p>(1) Absence of a pair of large, stiff setae on anterior dorsal margin of pereonite-3 was observed on non-ovigerous females, ovigerous females, and males; however, this pair of setae was noted only on a female with marsupium (USNM 1480638).</p><p>(2) Presence of pair of small spiniform setae on the sub-anterior dorsal region of the paratype (UF Arthropoda 49372) (Fig. 11C), which were not observed on the holotype.</p><p>(3) Number of spiniform setae on the inner medial margin of the antennular article-1 varied from 2–3 in both sexes, and this variation can be observed between left and right antennal article-1 of the same specimen (e.g. subadult male) (Table 3).</p><p>(4) Number of articles on the antennular outer flagellum varied from 2–3 in non-ovigerous females, ovigerous females, and female with marsupium (Table 3), moreover, it was observed that the number of articles of the outer flagellum varied between two (right) or three (left) on the ovigerous females (dissected) (Table 3).</p><p>(5) Antenna with 7̄9 articles (Table 3).</p><p>(6) Number of sub-proximal, blunt, spiniform process on dorsal basis of pereopods 2–3 (varied from 1–2 (holotype: P-2 with two (Fig. 16I) and P-3 with one (Fig. 16J), Table 3), and the number varied between left and right (Table 3).</p><p>(7) Number of spiniform setae on ventral margin of pereopod-2 of ovigerous females differs from those of nonovigerous females/female with marsupium/subadult male (four versus three, respectively) (Table 3).</p><p>Size-distribution. The body sizes of individuals Cryptapseudes mamua measured during this study are presented in Table 3. The non-ovigerous females ranged from about 1.0 to 1.5 mm (n=4). Ovigerous females ranged from 1.2 to 1.7 mm (n=2). Subadult male TL 1.2 mm.</p><p>Remarks. Cryptapseudes mamua sp. nov. becomes the fifth member of the genus to be described (Table 4, Key). It appears to most closely resemble C. leroyi in the general body form. It can be distinguished from C. leroyi, however, by (1) the mandible palp article-1 as long as article-2 (article-1 shorter than article- 2 in leroyi), (2) maxilliped endite outer margin with four denticles (without denticles in leroyi), (3) antennal article-1 with inner apophysis extending laterally, reaching the half-length of article-2 (extending distally nearly reaching proximal margin of article- 3 in leroyi).</p><p>The new species differs from C. acutifrons in having the rostrum expanded into a rosette cluster of spines (acute tip in C. acutifrons). Cryptapseudes mamua is distinguished from C. sankarankuttyi by lacking a single pair of lateral tubercles adjacent to the rostrum (present in sankarankuttyi). It can be separated from C. romanae by having antennal articles 2̄3 with distal inner margin bearing two and one spines, respectively (without spines in romanae), a maxillipedal palp article-1 lacking a stout spiniform seta on the distolateral margin (present in romanae), maxilliped endite outer margin expanded lateral with four denticles (straight and without denticles in romanae). The following illustrated identification key may be used to further separate the species within the genus Cryptapseudes .</p><p>At present the members of Cryptapseudes are known from the mid and SW Pacific and Western Indian Oceans (Figure 20). The new species from Mo’orea is the second species described from Pacific Ocean; the first being C. leroyi, from the mid-Pacific (Hawai’i) and the only member of the genus reported from the Northern Hemisphere. The remaining three nominal species, C. acutifrons, C. romanae, C. sankarankuttyi, are known from the Western Indian Ocean off SE Africa.</p><p>The lack of records for Cryptapseudes in the coastal waters of the Eastern Pacific may be due to the lack of sampling in live-bottom habitats suitable for its small cryptic species. For the same reasons, its eventual discovery in the tropical, live-bottom habitats of the Eastern and Western Atlantic cannot be ruled out.</p><p>“?” indicates that it was not possible to determine the presence/absence of this character</p><p>information on distribution and depth range</p><p>* Indicates male known.</p></div>	https://treatment.plazi.org/id/35508796FFE6FF9DFF79FF44FE53FDC3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Morales-Núñez, Andrés G.;Heard, Richard W.;Bird, Graham J.	Morales-Núñez, Andrés G., Heard, Richard W., Bird, Graham J. (2019): Two new apseudomorphan species (Crustacea: Tanaidacea: Metapseudidae) from Mo‘orea Island (Society Islands, French Polynesia) with taxonomic keys. Zootaxa 4564 (1): 213-247, DOI: 10.11646/zootaxa.4564.1.8
35508796FFD5FF9CFF79FF44FA53FD9D.text	35508796FFD5FF9CFF79FF44FA53FD9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryptapseudes Bacescu 1976	<div><p>Key to the known species (females) of Cryptapseudes, modified from David &amp; Heard 2015</p><p>1. Rostrum with acute tip (Fig. 21A)................................... C. acutifrons [Western Indian Ocean: Tanzania]</p><p>- Rostrum expanded into rosette cluster of spines (Fig. 21B ̅E)................................................... 2</p><p>2. Frontal margin of carapace with single tubercle on each side of rostrum (Fig. 21B).............................................................................................. C. sankarankuttyi [Western Indian Ocean: Tanzania]</p><p>- Frontal margin of carapace crenulate, with low row of small tubercles on each side of rostrum (Fig. 21C ̅E)............. 3</p><p>3. Maxilliped endite outer margin straight (Fig. 21I)..................... C. romanae [Western Indian Ocean: Madagascar]</p><p>- Maxilliped endite outer margin expanded lateral (Fig. 21G ̅H).................................................. 4</p><p>4. Maxilliped endite outer margin without denticles (Fig. 21G). Antennal article-1 with inner apophysis extending distally, nearly reaching proximal margin of article-3 (Fig. 21I)............. C. leroyi [Mid-North Pacific Ocean: Ni‘ihau Island, Hawai‘i]</p><p>- Maxilliped endite outer margin with four denticles (Fig. 21H). Antennal article-1 with inner apophysis extending distally, reaching half-length of article-2 (Fig. 21J)........ C. mamua [Mid-South Pacific Ocean: Mo‘orea Island, French Polynesia]</p></div>	https://treatment.plazi.org/id/35508796FFD5FF9CFF79FF44FA53FD9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Morales-Núñez, Andrés G.;Heard, Richard W.;Bird, Graham J.	Morales-Núñez, Andrés G., Heard, Richard W., Bird, Graham J. (2019): Two new apseudomorphan species (Crustacea: Tanaidacea: Metapseudidae) from Mo‘orea Island (Society Islands, French Polynesia) with taxonomic keys. Zootaxa 4564 (1): 213-247, DOI: 10.11646/zootaxa.4564.1.8
