identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
356287D9FF8CAF224C9BFE9AFC016E76.text	356287D9FF8CAF224C9BFE9AFC016E76.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Adolopus SHARP 1884	<div><p>GENUS ADOLOPUS SHARP, 1884</p> <p>(FIG. 12A–C)</p> <p>Material examined: Adolopus sp. 1: 4 males, 3 females (NMPC): New Zealand: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=168.07184&amp;materialsCitation.latitude=-46.898" title="Search Plazi for locations around (long 168.07184/lat -46.898)">Stewart Island</a>, Northwest Circuit Tk. at Kaipipi Bay, rotten longs/twigs in sparse hardwood forest with tree ferns, 46°53.88’S 168°4.31’E, 20 m a.s.l., 21.i.2016, M. Seidel &amp; M. Fikáček lgt. (2016- NZ 017).</p> <p>Adolopus sp. 2: 1 male (NMPC): New Zealand, Waikato (WO), Pirongia Forest Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=175.144&amp;materialsCitation.latitude=-37.966" title="Search Plazi for locations around (long 175.144/lat -37.966)">Ruapane Link Track</a> (lower part), 37.966°S 175.144°E, 235 m, 18– 21.xi.2016, M. Fikáček &amp; M. Seidel lgt. (MM02).</p> <p>Karyotype: 2 n = 22 + Xy (♂). In the Kaipipi Bay species (Fig. 12B), autosome pair 1 is the longest, and remaining autosomes gradually decrease in length so that pair 11 is about a quarter of the length of pair 1. All autosomes are metacentric to submetacentric. The X chromosome is metacentric, about as long as autosome pair 5, and the Y chromosome is dotlike. Adolopus from Pirongia (Fig. 12A) differs from that of Kaipipi Bay by subacrocentric pairs 7–9 and the acrocentric X chromosome slightly longer than autosome pair 2.</p> <p>Note: The identification of the examined specimens is impossible at the moment; however, the DNA data (Seidel, unpubl.) indicate that they represent two different species.</p></div> 	https://treatment.plazi.org/id/356287D9FF8CAF224C9BFE9AFC016E76	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF9EAF304E25F9B9FB366A0E.text	356287D9FF9EAF304E25F9B9FB366A0E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphiops Erichson 1843	<div><p>GENUS AMPHIOPS ERICHSON, 1843</p> <p>(FIG. 2A–E)</p> <p>Material examined: Amphiops mater Sharp, 1873: 2 females (BMNH); 1 female (NMPC): Taiwan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=121.06087&amp;materialsCitation.latitude=24.707466" title="Search Plazi for locations around (long 121.06087/lat 24.707466)">Beipu</a>, 24.707467°N 121.060870°E, 12.ix.2018, H.-C. Liu lgt.</p> <p>Karyotype: 2n = 18 (♀). The largest pair of chromosomes is submetacentric, the remaining pairs are metacentric.</p> <p>C-banding shows clear centromeric C-bands and terminal C-bands (heterozygous) on pairs 2 and 9.</p> <p>TRIBE BEROSINI</p></div> 	https://treatment.plazi.org/id/356287D9FF9EAF304E25F9B9FB366A0E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF96AF384E25FEE8FED66D7C.text	356287D9FF96AF384E25FEE8FED66D7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anacaena C. G. Thomson 1859	<div><p>GENUS ANACAENA THOMSON, 1859</p> <p>(FIG. 7A–D)</p> <p>Material examined: Anacaena gaetanae Bameul, 2001: 1 female (BMNH): France, Corsica, Corse du Sud, R. Stabiacciu, Porto Veccio, 12.vii.2009, R. B. &amp; E.M. Angus lgt. (BMNH).</p> <p>Anacaena lutescens (Stephens, 1829): 3 female specs. (BMNH): Germany, Hamburg district, 16.iv.1988, R. Angus lgt.; 1 spec. (BMNH): United Kingdom, Norfolk, R. Angus lgt.</p> <p>Karyotype: The karyotype of A. gaetanae (Fig. 7A) is similar to that of bisexual A. lutescens (2 n = 16 + Xy p (♂)) but the chromosome taken to be the X is noticeably larger.</p> <p>Comments on karyotypes: The karyotypes are discussed in detail by Shaarawi &amp; Angus (1991a). Parthenogenetic A. lutescens is heterozygous for loss of a small apical section of chromosome 8, distal to a secondary constriction (Fig. 7C). Anacaena globulus (Paykull, 1798) has a fusion of two chromosomes (autosomes) to give a reduction from eight to seven pairs (2 n = 14 + Xy p). Anacaena rufipes (Guillebeau, 1896) has two further fusions, one involving the X chromosome, to give five pairs of autosomes and neo-XY sex chromosomes (2 n = 10 + neo-XY). Shaarawi &amp; Angus (1991a) suggested that the original small Y chromosome had also fused with the neo-Y chromosome, but it is also possible that it has simply been lost. Triploidy (3 n = 27) was found in two widely separated populations of A. lutescens, from the United Kingdom (Armathwaite, Cumbria) and the Netherlands (Doetinchem, Gelderland). In both cases diploid females were also present. In the triploids there was one pair of intact chromosome 8 and one extra chromosome 8 with the deletion. In the Armathwaite population, one of three replicates of chromosome 8 has a pericentric inversion and is acrocentric.</p> </div>	https://treatment.plazi.org/id/356287D9FF96AF384E25FEE8FED66D7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF9EAF304CE1FE9FFB296F85.text	356287D9FF9EAF304CE1FE9FFB296F85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Berosus Leach 1817	<div><p>GENUS BEROSUS LEACH, 1817</p> <p>Comments on karyotypes: Shaarawi &amp; Angus (1991a) published the karyotype data of three species of the subgenus Berosus [Berosus affinis Brullé, 1835, Berosus luridus (Linnaeus, 1761) and Berosus signaticollis (Charpentier, 1825)] and two species of the subgenus Enoplurus (Berosus bispina Reiche &amp; Saulcy, 1856 and Berosus fulvus Kuwert, 1888) and Angus et al. (1994) added Berosus (s.s.) hispanicus Küster, 1847. The karyotype has 2 n = 16 + Xy p (♂), in all species except Berosus signaticollis which has an extra pair of autosomes 2 n = 18 + Xy p (♂). The karyotype of Berosus affinis may have an additional pair of B-chromosomes. The parachute configuration is shown by meiotic preparations of Berosus signaticollis. C-banding was not attempted; however, the chromosomes appear elongate and with small centromeric constrictions suggesting that the centromeric C-bands are small. Agarawal (1960b) presents data on mitosis and meiosis of Berosus (Enoplurus) indicus (Motschulsky, 1861), providing the meioformula 8 + Xy p which agrees with newer data; the species identification by Agarawal (1960b) may be easily incorrect since no reliable identification key for Asian Berosus existed at that time.</p> <p>TRIBE LACCOBIINI</p></div> 	https://treatment.plazi.org/id/356287D9FF9EAF304CE1FE9FFB296F85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FFB8AF164F82FD5BFD6B6811.text	356287D9FFB8AF164F82FD5BFD6B6811.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Berosus Leach 1817	<div><p>Berosus (s.s.) hispanicus Sexual Berosus (Enoplurus) indicus Sexual</p> <p>Hydrophilinae: Laccobiini</p> <p>Laccobius (s.s.) minutus Sexual Laccobius (s.s.) colon Sexual</p></div> 	https://treatment.plazi.org/id/356287D9FFB8AF164F82FD5BFD6B6811	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF84AF2B4CE1FE5DFDB56EBC.text	356287D9FF84AF2B4CE1FE5DFDB56EBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cercyon LEACH 1817	<div><p>GENUS CERCYON LEACH, 1817</p> <p>(FIGS 16-17)</p> <p>Material examined: Cercyon (Dicyrtocercyon) ustulatus (Preyssler, 1790): 1 male (BMNH): United Kingdom, Surrey, Egham, R. Angus lgt.</p> <p>Cercyon (Paracercyon) analis (Paykull, 1798): 1 female (NMPC): Czech Republic, Bohemia. Černíky, 3.vi.2019, collecting in compost, V. Sýkora lgt.</p> <p>Cercyon convexiusculus Stephens, 1829: 1 male, 1 female (BMNH): United Kingdom, Norfolk, East Walton, R. Angus lgt. 1 spec. (NMPC): Czech Republic, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.389573&amp;materialsCitation.latitude=49.079414" title="Search Plazi for locations around (long 14.389573/lat 49.079414)">Zliv</a>, 49.0794131°N 14.3895733°E, 24.iv.2019, V. Kolá ř lgt.</p> <p>Cercyon sternalis (Sharp, 1890): 2 females (BMNH): United Kingdom, Norfolk, East Walton, R. Angus lgt.</p> <p>Cercyon tristis (Illiger, 1801): 1 female (BMNH): United Kingdom, Norfolk, East Walton, R. Angus lgt.</p> <p>Cercyon marinus Thomson, 1853: 1 male (BMNH): United Kingdom, Norfolk, East Walton, R. Angus lgt.</p> <p>Cercyon impressus (Sturm, 1807): 2 males (BMNH): United Kingdom, Surrey, Egham, R. Angus lgt.</p> <p>Cercyon obsoletus (Gyllenhal, 1808): 1 male (BMNH): United Kingdom, Surrey, Bookham Common, R. Angus &amp; H. Gray lgt.</p> <p>Cercyon lateralis (Marsham, 1802): 1 male (BMNH): United Kingdom, Windsor, Boveney, R. Angus &amp; H. Gray lgt.; 1 male (BMNH): United Kingdom, Windsor, Windsor Deer Park, R. Angus &amp; H. Gray lgt.</p> <p>Cercyon haemorrhoidalis (Fabricius, 1775): 1 male (B M N H): U n i t e d K i n g d o m, Wi n d s o r, B o v e n e y, R. Angus &amp; H. Gray lgt.; 2 males (BMNH): United Kingdom, Windsor, Windsor Deer Park, R. Angus &amp; H. Gray lgt.</p> <p>Cercyon melanocephalus (Linnaeus, 1758): 1 male, 1 female (BMNH): United Kingdom, Windsor, Windsor Deer Park, R. Angus &amp; H. Gray lgt.</p> <p>Karyotypes: The examination of the eleven species shown here indicates that the subgenus Dicyrtocercyon (with the only species Cercyon ustulatus; Fig. 16A) and C. convexiusculus of the Cercyon convexiusculus group (Fig. 16F–I) have the karyotype of 2 n = 28 + Xy (♂). All nine remaining species (Cercyon sternalis and Cercyon tristis of the Cercyon convexiusculus group, Cercyon analis of the subgenus Paracercyon and other examined ‘usual’ European Cercyon) have the karyotype 2 n = 22 + Xy (♂).</p> <p>Cercyon ustulatus (Fig. 16A) has a large metacentric X chromosome, about the same size as autosome 6. The autosomes are a mixture of metacentric and subacrocentric ones, with pair 14 about a quarter of the length of pair 1.</p> <p>Cercyon convexiusculus (Fig. 16F–I) has a small metacentric X chromosome about the same size as autosome pair 9. Autosome pairs 4, 10 and 12–14 are acrocentric to subacrocentric whereas the remaining pairs are metacentric to submetacentric. Interestingly, pair 1 is polymorphic for a pericentric inversion— the East Walton specimens are either homozygous acrocentric (Fig. 16F) or heterozygous (Fig. 16G, H) whereas the Czech specimen (Fig. 16I) is homozygous submetacentric.</p> <p>The remaining species differ from each other in the relative length of the X chromosome: it is a long metacentric in Cercyon lateralis (Fig. 17B–E), Cercyon haemorrhoidalis (Fig. 17I–K) and Cercyon melanocephalus (Fig. 17L–N), and shorter but still metacentric in Cercyon marinus (Fig. 17A), Cercyon obsoletus (Fig. 17F, G) and Cercyon impressus (Fig. 17H). The X chromosome cannot be identified in Cercyon analis (Fig. 16B), Cercyon tristis (Fig. 16C) and Cercyon sternalis (Fig. 16D, E) because only females are available.</p> <p>Note: The yet unpublished DNA-based molecular phylogeny of the Megasternini (Arriaga-Varela &amp; Fikáček, unpubl.) revealed that the genus is polyphyletic. The groups used in Figures 16 and 17 correspond to these molecular findings, each of them represents a monophyletic DNA clade.</p> </div>	https://treatment.plazi.org/id/356287D9FF84AF2B4CE1FE5DFDB56EBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF96AF384CABFD33FAFA6EB0.text	356287D9FF96AF384CABFD33FAFA6EB0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chaetarthria STEPHENS 1835	<div><p>GENUS CHAETARTHRIA STEPHENS, 1835</p> <p>(FIG. 7L–M)</p> <p>M a t e r i a l e x a m i n e d: C h a e t a r t h r i a s i m i l l i m a Vorst &amp; Cuppen, 2003: 1 male (BMNH): United Kingdom: England, Oxfordshire, Cothill, R. Angus &amp; F. Shaarawi lgt.</p> <p>Karyotype: 2 n = 16 + Xy p (♂). There are distinct steps in the size decreases between chromosomes 1 and 2, 2 and 3, and 3 and 4, then a gradual decrease between chromosomes 4 and 6, a slightly larger one between 6 and 7 and a sharp decrease between 7 and 8. The X chromosome is about the same length as autosomes 4–6 and the y is dot like. Chromosomes 1–6 and the X chromosome are metacentric, chromosomes 7 and 8 are subacrocentric.</p> <p>Note: The karyotype was published by Angus &amp; Shaarawi (1997) under the name Chaetarthria seminulum (Herbst, 1797). It has been slightly rearranged and improved and it republished here.</p> <p>SUBFAMILY ENOCHRINAE</p></div> 	https://treatment.plazi.org/id/356287D9FF96AF384CABFD33FAFA6EB0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF81AF2F4CB1FE67FC77688A.text	356287D9FF81AF2F4CB1FE67FC77688A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coelostoma BRULLE 1835	<div><p>GENUS COELOSTOMA BRULLÉ, 1835</p> <p>(FIG. 15A–C, H)</p> <p>Material examined: Coelostoma orbiculare (Fabricius, 1775): 1 male, 2 females (BMNH): United Kingdom, Norfolk, East Walton, R. Angus &amp; F. Shaarawi lgt.</p> <p>Karyotype: 2 n = 26 + Xy (♂). The autosomes and the X chromosome are metacentric to submetacentric with strong centromeric C-bands. Autosome pair 6 has a distinct subterminal secondary constriction on its long arm. The Y chromosome is dot like, and there may be two small B-chromosomes in the examined specimens. All the nuclei are from eggs taken from a cocoon spun by one of the females. In total, 16 male and 6 female nuclei were photographed. The two small B-chromosomes were present in only one male nucleus.</p></div> 	https://treatment.plazi.org/id/356287D9FF81AF2F4CB1FE67FC77688A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF8CAF214CE7FA8FFDB46917.text	356287D9FF8CAF214CE7FA8FFDB46917.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyloma SHARP 1872	<div><p>GENUS CYLOMA SHARP, 1872</p> <p>(FIG. 12D–H)</p> <p>Material examined: Cyloma guttulatus Sharp, 1884: 1 female (NMPC): New Zealand: Fiordland (FD), Borland Road 24 km NWW of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=167.33817&amp;materialsCitation.latitude=-45.6855" title="Search Plazi for locations around (long 167.33817/lat -45.6855)">Monowai</a>, baited pitfall traps (rotten squid) in degraded remnants of Nothofagus forest with numerous rotten logs, 45°41.13’S 167°20.29’E, 320 m, 27.i.2016, M. Seidel, V. Sýkora &amp; M. Fikáček lgt. (2016- NZ 024).</p> <p>Cyloma sp.: (undescribed species illustrated in Fig. 12H). 1 male, 1 female (NMPC): New Zealand: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=167.33817&amp;materialsCitation.latitude=-45.6855" title="Search Plazi for locations around (long 167.33817/lat -45.6855)">Fiordland</a> (FD), Borland Road 24 km NWW of Monowai, on dead possum, 45°41.13’S 167°20.29’E,</p> <p>320 m, 27.i.2016, M. Seidel, V. Sýkora &amp; M. Fikáček lgt. (2016- NZ 024).</p> <p>Karyotype: 2 n = 22 + Xy (♂). In Cyloma guttulatus, chromosome pairs 1–3 and 5 are metacentric, with pair 2 half the size of pair 1, and the others decreasing in size from half the length of pair 2 to about quarter of the size of pair 2 (Fig. 12D). The X chromosomes cannot be identified. Cyloma sp. (Fig. 12E–G) has a similar karyotype to Cyloma guttulatus; however, autosome pair 2 is about two-thirds of the length of pair 1 and the smallest autosomes, and the X chromosome are about a quarter of the length of pair 1.</p> </div>	https://treatment.plazi.org/id/356287D9FF8CAF214CE7FA8FFDB46917	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF8FAF214FDEFD66FD646E71.text	356287D9FF8FAF214FDEFD66FD646E71.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cylomissus BROUN 1903	<div><p>GENUS CYLOMISSUS BROUN, 1903</p> <p>(FIG. 12I–N)</p> <p>Material examined: Cylomissus glabratus Broun, 1903: 1 male, 1 female (NMPC): New Zealand: Otago Lakes, Little Meg headwaters at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=168.96083&amp;materialsCitation.latitude=-44.876667" title="Search Plazi for locations around (long 168.96083/lat -44.876667)">Cardrona Skifield Rd.</a>, stream collecting in small exposed streamlets (stony and mossy), 44°52.60’S 168°57.65’E, 1280 m, 5.xii.2017, M. Fikáček, D. Sadílek &amp; V. Sýkora lgt. (2017-NZ63). 1 male (NMPC): New Zealand: Stewart Island, Fern Gully W of Oban, at bridge over <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=168.1&amp;materialsCitation.latitude=-46.892" title="Search Plazi for locations around (long 168.1/lat -46.892)">Mill Creek</a>, 17–21.i.2016, 46°53.52’S 168°6.00’E, 45 m, M. Seidel, V. Sýkora &amp; M. Fikáček lgt. (2016- NZ 004).</p> <p>Karyotype: 2 n = 22 + Xy (♂). Autosome pair 1 is 3–4 times the length of pair 2, whereas the remaining pairs have a gradual decrease in length along the karyotype, with pair 11 about half the length of pair 2, the X chromosome is only slightly longer than pair 11. The Y chromosome is dot-like. Examined specimens from both localities have a similar karyotype.</p></div> 	https://treatment.plazi.org/id/356287D9FF8FAF214FDEFD66FD646E71	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF96AF394C8AF9FDFC336C34.text	356287D9FF96AF394C8AF9FDFC336C34.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cymbiodyta BEDEL 1881	<div><p>GENUS CYMBIODYTA BEDEL, 1881</p> <p>(FIG. 8A, I)</p> <p>Material examined: Cymbiodyta marginella (Fabricius, 1792): 1 spec. (BMNH): Greece, Corfu, R. Angus lgt.</p> <p>K a r y o t y p e: 2 n = 1 6 + X y p (♂). E i g h t p a i r s o f metacentric or submetacentric autosomes showing a gradual decrease in size, pair 8 is about half the</p> <p>length of pair 1. The metacentric X chromosome is the smallest in the nucleus, apart from the dot-like y.</p> <p>Note: The monotypic genus Helocombus (LeConte, 1855) was synonymized with Cymbiodyta by Toussaint &amp; Short (2019). The karyotype of its only species, Cymbiodyta bifidus (LeConte, 1855), was examined by Smith (1960) and reported to be 2 n = 16 + Xy p (♂), corresponding to our findings in the European C. marginella.</p> </div>	https://treatment.plazi.org/id/356287D9FF96AF394C8AF9FDFC336C34	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF81AF2A4C7BFBF9FDA66B81.text	356287D9FF81AF2A4C7BFBF9FDA66B81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dactylosternum WOLLASTON 1854	<div><p>GENUS DACTYLOSTERNUM WOLLASTON, 1854</p> <p>(FIG. 15D–G, I)</p> <p>Material examined: Dactylosternum corbetti (Balfour- Browne, 1942): 1 male (NMPC): Taiwan, Nantou County, 4.8 km SSE of Tonglin, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.78434&amp;materialsCitation.latitude=24.04791" title="Search Plazi for locations around (long 120.78434/lat 24.04791)">Beikeng Creek Trail</a>, 24.04791°N 120.78434°E, 3.x.2018, M. Fikáček, F.-S. Hu, W.- R. Liang &amp; H.-C. Liu lgt.</p> <p>Dactylosternum flavicorne (Mulsant, 1844): 1 male, 1 female (BMNH): Dominican Republic, lowland rainforest, locality unknown, R. Angus lgt.</p> <p>Karyotype: 2 n = 26 + Xy (♂). In Dactylosternum flavicorne (Fig. 15D – F), the autosomes are metacentric to submetacentric, with the smallest pair about half of the length of the largest pair. The X chromosome is metacentric and the largest in the karyotype. All the chromosomes, except the dot-like Y, have large centromeric C-bands. The karyotype of Dactylosternum corbetti (Fig. 15G) has the autosomes and X chromosome metacentric to submetacentric and with smaller C-bands than in Dactylosternum flavicorne. Pair 1 is distinctly longer than the others, and the X chromosome is about as long as pair 5.</p> <p>TRIBE PROTOSTERNINI</p></div> 	https://treatment.plazi.org/id/356287D9FF81AF2A4C7BFBF9FDA66B81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF88AF224E2AFF39FE2E69D0.text	356287D9FF88AF224E2AFF39FE2E69D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Enochrus THOMSON 1859	<div><p>GENUS ENOCHRUS THOMSON, 1859</p> <p>(FIGS 8B–J, 9–10)</p> <p>Material examined: Enochrus (s.s.) melanocephalus (Olivier, 1792): 2 spec. (BMNH): United Kingdom, Surrey, Egham, R. Angus lgt.</p> <p>Enochrus (Methydrus) affinis (Thunberg, 1794): 2 spec. (BMNH): United Kingdom, Hampshire, New Forest, R. Angus lgt.</p> <p>Enochrus (Methydrus) coarctatus (Gredler, 1863): 1 spec. (BMNH): United Kingdom, Dorset, Studland Heath, R. Angus lgt.</p> <p>Enochrus (Methydrus) nigritus (Sharp, 1872): 1 spec. (BMNH): Spain, Peñalara, R. Angus lgt.; 1 spec. (BMNH): United Kingdom, Norfolk, East Walton, R. Angus lgt.; 1 spec. (BMNH): Greece, Corfu, R. Angus lgt.</p> <p>Enochrus (Methydrus) morenae (Heyden, 1870): 1 spec. (BMNH): Spain, Plasencia, R. Angus lgt.</p> <p>Enochrus (Methydrus) sauteri dʼ Orchymont, 1913: 2 spec. (NMPC): Taiwan, 4.8 km SEE of Tonglin, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.78434&amp;materialsCitation.latitude=24.04791" title="Search Plazi for locations around (long 120.78434/lat 24.04791)">Beikeng Creek Trail</a>, 24.04791°N 120.78434°E, 3.x.2018, Fikáček, Hu, Liang &amp; Liu lgt.</p> <p>Enochrus (Lumetus) bicolor (Fabricius, 1792): 1 spec. (BMNH): United Kingdom, Sussex, Cuckmere, R. Angus lgt.</p> <p>Enochrus (Lumetus) ochropterus (Marsham, 1802): 2 spec. (BMNH): United Kingdom, Dorset, Studland Heath, R. Angus lgt.; 2 spec. (BMNH): Germany, Niedersachsen, Staatsforest Gohrde, R. Angus lgt.</p> <p>Enochrus (Lumetus) testaceus (Fabricius, 1801): 2 spec. (BMNH): United Kingdom, Norfolk, East Walton, R. Angus lgt.; 2 spec. (BMNH): Sweden, Öland, R. Angus lgt.</p> <p>Enochrus (Lumetus) quadripunctatus (Herbst, 1797): 2 spec. (BMNH): United Kingdom, Norfolk, East Walton, R. Angus lgt.; 2 spec. (BMNH): Germany, Schleswig-Holstein, Ratzeburg, R. Angus lgt.</p> <p>Enochrus (Lumetus) fuscipennis (Thomson, 1884): 2 spec. (BMNH): United Kingdom, Cumbria, Drigg, R. Angus lgt.; 2 spec. (BMNH): United Kingdom, Scotland, Ayrshire, Lochton loch, R. Angus lgt.; 3 spec. (BMNH): Germany, Schleswig-Holstein, Ratzeburg, R. Angus lgt.; 2 spec. (BMNH): Spain, Provincia de Santander, Embalse de Ebro, R. Angus lgt.; 2 spec. (BMNH): Denmark, Rømø Island, R. Angus lgt.; 2 males (BMNH): Denmark, Rømø Island, R. Angus lgt. Enochrus (Lumetus) halophilus (Bedel, 1878): 2 spec. (BMNH): United Kingdom, Sheppey, Kent, R. Angus lgt.; 2 spec. (BMNH): United Kingdom, Bembridge, Isle of Wight, R. Angus lgt. 2 spec. (BMNH): Spain, Albacete, Pétrola, P. Arribas lgt.</p> <p>Karyotypes: 2 n = 16 + Xy p (♂). Enochrus s.s. (Fig. 8H) has autosomal pairs 3, 6 and 8 subacrocentric, pair 3 is heterozygous for an extra C-band. The other pairs, and the X chromosome, are metacentric, with well-developed centromeric C-bands. The subgenera Methydrus and Lumetus show no obvious features in their karyotypes, whereas the Taiwanese Enochrus sauteri (Fig. 8G) is unusual in having autosome pair 1 clearly twice the lengths of the others, which are all of the similar size.</p> <p>The karyotypes appear remarkably uniform with some apparent species differences. In European Methydrus, Enochrus coarctatus (Fig. 8C) differs from Enochrus affinis (Fig. 8B) and Enochrus nigritus (Fig. 8D–E) in having autosome pairs 7 and 8, and the X chromosome, subacrocentric, whereas in the others they are metacentric. No differences were found between Spanish (dark brown) and English pale coloured specimens of Enochrus nigritus (compare Fig. 8D and E). In Lumetus, English Enochrus halophilus (Fig. 9F–K) differs from Enochrus quadripunctatus and Enochrus fuscipennis (Fig. 10) in having autosome pairs 7 and 8, and the X chromosome acrocentric, with the centromeric C-bands only about half the size of those of the metacentric autosomes. However, in Spanish Enochrus halophilus, which are inseparable from English material by mitochondrial DNA (P. Arribas, unpubl.), all the chromosomes (apart from the dot-like Y) are metacentric with C-bands of the same size as in Enochrus halophilus (Fig. 9H–K). The separation of Enochrus fuscipennis from Enochrus quadripunctatus is also problematic as the mitochondrial DNA reveals Enochrus fuscipennis intermixed with Enochrus quadripunctatus (P. Arribas, unpubl.). The situation is made even more complicated by material from Ratzeburg (Germany). This material included clear Enochrus quadripunctatus whose karyotype matched that of English specimens (as in Fig. 10A–C) and also apparent Enochrus fuscipennis whose karyotype differed in the submetacentric autosome 1 and the clearly smaller X chromosome (Fig. 10D–G). The material from the Rømø Island (Denmark) (Fig. 10H–I) comprised two males whose karyotypes could only be understood if they were hybrids (but not F1) between western and Ratzeburg-type populations of Enochrus fuscipennis. Both males have small X chromosomes and one has autosome 1 heterozygous for western and Ratzeburg forms, whereas the other has both replicates of the western form. One male has a B-chromosome and both have normal first metaphase of meiosis.</p> <p>SUBFAMILY ACIDOCERINAE GENUS AGRAPHYDRUS RÉGIMBART, 1903</p> <p>(FIG. 11A–C, M)</p> <p>M a t e r i a l e x a m i n e d: A g r a p h y d r u s d e c i p i e n s Minoshima, Komarek &amp; Ohara, 2015: 1 male, 1 female (BMNH), 1 male (NMPC): Taiwan, 4.8 km</p> <p>SEE of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.78434&amp;materialsCitation.latitude=24.04791" title="Search Plazi for locations around (long 120.78434/lat 24.04791)">Tonglin</a>, Beikeng Creek Trail, 24.04791°N 120.78434°E, 3.x.2018, under leaves on a wet rock, lgt. M. Fikáček, H.-C. Liu, F.-S. Hu &amp; W.- R. Liang.</p> <p>Agraphydrus variabilis Komarek &amp; Hebauer, 2018: 1 female (BMNH): Taiwan, 4.8 km SEE of Tonglin, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.78434&amp;materialsCitation.latitude=24.04791" title="Search Plazi for locations around (long 120.78434/lat 24.04791)">Beikeng Creek Trail</a>, 24.04791°N 120.78434°E,</p> <p>3.x.2018, in gravel at side of a river, lgt. M. Fikáček, H.-C. Liu, F.-S. Hu &amp; W.-R. Liang.</p> <p>Karyotype: 2 n = 16 + Xy (♂). The autosomes show a gradual decrease in size along the karyotype so that pair 8 is about half the length of pair 1. In Agraphydrus decipiens (Fig. 11A–B), pairs 7 and 8, and the X chromosome, are acrocentric, whereas the rest (apart for the dot-like Y chromosome) are metacentric. The sex chromosomes of Agraphydrus variabilis (Fig. 11C) cannot be identified as only females are available. The smallest chromosome is acrocentric, the rest metacentric. C-bands are confined to the centromeres and are small, especially in Agraphydrus decipiens.</p> </div>	https://treatment.plazi.org/id/356287D9FF88AF224E2AFF39FE2E69D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF8FAF214E38FAA8FBFB6A88.text	356287D9FF8FAF214E38FAA8FBFB6A88.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Exydrus BROUN 1886	<div><p>GENUS EXYDRUS BROUN, 1886</p> <p>(FIG. 13A, D, I)</p> <p>Material examined: Exydrus gibbosus Broun, 1886: 1 female (NMPC): New Zealand: Wellington (WL), Tararua Range, 1 km W of Titurea Dam, start of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=175.66064&amp;materialsCitation.latitude=-40.4295" title="Search Plazi for locations around (long 175.66064/lat -40.4295)">Greens Rd.</a>, 40.4295°S 175.66064°E, 145 m, 26.xi.2016, fragment of broadleaf forest with sparse understory with ferns and Pandanus Parkinson: sifting, M. Fikáček &amp; M. Seidel lgt. (MM31).</p> <p>Karyotype: 2 n = 30 (♀). The only nucleus obtained has 29 chromosomes and the karyogram shows a serious size mismatch in pair 3 so there should be at least 30 chromosomes in the karyotype, presumably 14 pairs of autosomes plus the sex chromosomes. The smallest chromosome appears heavy and almost single-stranded; however, this is almost certainly because the two chromatids of a metacentric are lying on top of each other. This is frequent with small chromosomes. There is gradual decrease in length along the karyotype, with no pair strikingly longer than the others. The smallest pair is about a third of the length of the longest pair. Most of the chromosomes are metacentric or submetacentric; however, pairs 6–9 are subacrocentric.</p></div> 	https://treatment.plazi.org/id/356287D9FF8FAF214E38FAA8FBFB6A88	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF8CAF224FDFFD33FB096A0D.text	356287D9FF8CAF224FDFFD33FB096A0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helochares MULSANT 1844	<div><p>GENUS HELOCHARES MULSANT, 1844</p> <p>(FIG. 11D–L, N)</p> <p>Material examined: Helochares lividus (Forster, 1771): 2 spec. (BMNH): United Kingdom, Somerset, Shapwick, R. Angus lgt.; 1 spec. (BMNH): Spain, Provincia de Santander, Embalse de Ebro, R. Angus lgt.; 1 spec. (BMNH): Italy, Sardinia, Nuoro Province, R. Angus lgt.</p> <p>Helochares obscurus (Müller, 1776): 1 spec. (BMNH): Sweden, Öland, R. Angus lgt.</p> <p>Helochares punctatus Sharp, 1869: 3 spec. (BMNH): United Kingdom, Surrey, Chobham, R. Angus lgt.; 2 spec. (BMNH): United Kingdom, Hampshire, New Forest, R. Angus lgt.</p> <p>Helochares sauteri dʼ Orchymont, 1943: 3 spec. (NMPC): Taiwan, 2.5 km E of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.76323&amp;materialsCitation.latitude=24.06288" title="Search Plazi for locations around (long 120.76323/lat 24.06288)">Tonglin</a>, 24.06288°N 120.76323°E, 3.x.2018, M. Fikáček, F.-S. Hu, W.-R. Liang &amp; H.-C. Liu lgt.</p> <p>Karyotype: 2 n = 16 + Xy p (♂). In Helochares lividus (Fig. 11D–E), pairs 1–6 are metacentric, whereas pairs 7, 8 and the X are submetacentric; there are small centromeric C-bands on all the chromosomes except the Y. In Helochares obscurus (Fig. 11F), pairs 1–7 are metacentric, pair 8 is subacrocentric and the X chromosome submetacentric. In Helochares punctatus (Fig. 11G–H), the autosomes are all metacentric, the X chromosome is submetacentric and all chromosomes possess large centromeric C-bands. Some H. punctatus appear to have up to three dotlike Y chromosomes; however, the additional two dots are probably B-chromosomes. Helochares sauteri from Taiwan (Fig. 11L) differs from the other species in its small acrocentric X chromosome, only half the length of the smallest autosome.</p> <p>SUBFAMILY CYLOMINAE</p></div> 	https://treatment.plazi.org/id/356287D9FF8CAF224FDFFD33FB096A0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF96AF384FC7F99EFB09699A.text	356287D9FF96AF384FC7F99EFB09699A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horelophus d Orchymont 1913	<div><p>GENUS HORELOPHUS D’ ORCHYMONT, 1913</p> <p>(FIG. 7E–K)</p> <p>M a t e r i a l e x a m i n e d: H o r e l o p h u s w a l k e r i dʼ Orchymont, 1913: 2 males (NMPC): New Zealand, Nelson (NN), Kahurangi Nat. Park, AppleTree Flat at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=172.75761&amp;materialsCitation.latitude=-41.0744" title="Search Plazi for locations around (long 172.75761/lat -41.0744)">Cobb Dam Road</a>, 200 m, 41.0744°S 172.75761°E, 12.xii.2016, hygropetric habitat: exposed rocks with thin film of water, moss and algae, M. Fikáček &amp; M. Seidel lgt. (MM73). 1 male (NMPC): New Zealand: Marlborough, Pelorus Bridge Scenic Reserve, first waterfall at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=173.56738&amp;materialsCitation.latitude=-41.30526" title="Search Plazi for locations around (long 173.56738/lat -41.30526)">Tawa Walk</a>, 41.30526°S 173.56739°E, 55 m, floating moss from small waterfall in Nothofagus Blume forest, 11.xii.2016, M. Fikáček &amp; M. Seidel lgt. (MM69 b).</p> <p>Karyotype: 2 n = 16 + Xy (♂). Horelophus chromosomes seems to be acrocentric to subacrocentric and decrease in size along the karyotype. X chromosome is about as long as chromosomes 6–8.</p> <p>TRIBE CHAETARTHRIINI</p></div> 	https://treatment.plazi.org/id/356287D9FF96AF384FC7F99EFB09699A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FFB9AF174FB0FBFFFB386D9B.text	356287D9FFB9AF174FB0FBFFFB386D9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horelophus walkeri Sexual	<div><p>Horelophus walkeri Sexual 18 8 + Xy This paper</p> <p>Enochrinae Cymbiodyta marginella Sexual 18 8 + Xy This paper</p> <p>Cymbiodyta bifida Sexual 18 8 + Xy Smith (1960) (as</p> <p>p</p> <p>Helocombus bifidus)</p> <p>Enochrus s.s. melanocephalus Sexual 18 8 + Xy This paper</p> <p>Enochrus (Methydrus) affinis Sexual 18 8 + Xy This paper</p> <p>Enochrus (Methydrus) coarctatus Sexual 18 8 + Xy This paper</p> <p>Enochrus (Methydrus) nigritus Sexual 18 8 + Xy This paper</p> <p>Enochrus (Methydrus) morenae Sexual 18 8 + Xy This paper</p> <p>Enochrus (Methydrus) sauteri Sexual 18 8 + Xy This paper</p> <p>Enochrus (Lumetus) bicolor Sexual 18 8 + Xy This paper</p> <p>Enochrus (Lumetus) fuscipennis Sexual 18 8 + Xy</p> <p>p</p> <p>This paper</p> <p>Enochrus (Lumetus) halophilus Sexual 18 8 + Xy</p> <p>p</p> <p>This paper</p> <p>Enochrus (Lumetus) ochropterus Sexual 18 8 + Xy This paper</p> <p>Enochrus (Lumetus) Sexual 18 8 + Xy</p> <p>p</p> <p>This paper</p> <p>quadripunctatus Enochrus (Lumetus) testaceus Sexual 18 8 + Xy This paper</p> <p>Acidocerinae</p> <p>Agraphydrus decipiens Sexual 18 8 + Xy This paper</p> </div>	https://treatment.plazi.org/id/356287D9FFB9AF174FB0FBFFFB386D9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF9FAF3C4E28F98BFDB66A40.text	356287D9FF9FAF3C4E28F98BFDB66A40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrobius LEACH 1815	<div><p>GENUS HYDROBIUS LEACH, 1815</p> <p>(FIG. 4A–J)</p> <p>Material examined: Hydrobius fuscipes (Linnaeus, 1758): 2 spec. (BMNH): United Kingdom, England, Sheppey Island, R. Angus lgt.; 1 spec. (BMNH): Spain: Menorca, G. Foster lgt. [specimen barcoded</p> <p>by J. Bergsten and confirmed to belong to Hydrobius fuscipes sensu Fossen et al. (2016)].</p> <p>Hydrobius arcticus Kuwert, 1890: 1 spec. (BMNH): Sweden, Abisko National Park, G. N. Foster &amp; A. N. Nilsson lgt. [specimens from this locality barcoded by J. Bergsten and confirmed to belong to H. arcticus sensu Fossen et al. (2016)].</p> <p>Hydrobius rottenbergii Gerhardt, 1872: 1 spec. (BMNH): Sweden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=19.6597&amp;materialsCitation.latitude=63.4314" title="Search Plazi for locations around (long 19.6597/lat 63.4314)">Angermanland</a>, Nordmaling, Järnäsklubb 63.4314°N, 19.6597°E, rockpools by the sea, 13.x.2017, A.N. Nilsson lgt. [Material from this locality barcoded by J. Bergsten and confirmed to belong to H. rottenbergii sensu Fossen et al. (2016)].</p> <p>Hydrobius subrotundus Stephens, 1829: 3 spec. (BMNH): England, Westmorland, swamp near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-2.876&amp;materialsCitation.latitude=54.305" title="Search Plazi for locations around (long -2.876/lat 54.305)">Borderside</a>, 54.305°N 2.876°W. 34 m a.s.l., 18.iii.2016, G. Foster lgt.; 2 spec. (BMNH): France, Aubrac, R. Angus lgt.; 2 spec. (BMNH): Scotland, Kype Muir, S. Routledge lgt. [one specimen barcoded by J. Bergsten and confirmed to belong to H. subrotundus sensu Fossen et al. (2016)].</p> <p>Hydrobius pui Jia, 1995: 9 spec. (BMNH): China, Sichuan, Maba Prefecture, Songpan County, pools beside Chuanzhusi-Huanglong, road, by Mt Baoxueding, 3650 m a.s. l., 32°43’37.19”N 103°40’0.7”E, 8.vii.2016, R. Angus &amp; F-L Jia lgt. [These specimens were sequenced by J. Bergsten and indicated to belong to the same barcoding clade as specimens from Kangding County, Xinduqiao, Gaba, Liqi village, 29°56’35.80”N 101°35’21.46”E, 3359 m a.s.l., 28.vi.2016, R.B. Angus, F-L. Jia, Z-Q. Li &amp; K. Chen lgt. which were not karyotyped. Type locality of H. pui is Yushu in southern Qinghai, but the fact that our material from two widely separated localities belongs to the same DNA clade gives confidence that all these specimens are conspecific with H. pui.]</p> <p>Karyotype: 2 n = 16 + Xy (♂). All species with chromosomes 1 and 2 metacentric, 3–7 submetacentric, and 8 and X subacrocentric, X the smallest apart from dot-like y. In H. fuscipes, the English material (Fig. 4A–B) has a centromeric C-band on chromosome 8, Menorcan material (Fig. 4C) has additional bands on chromosomes 5 and 6. Hydrobius subrotundus (Fig. 4F–H) chromosomes have the size and form as H. fuscipes but often with apical satellite C-bands on chromosomes 3–7. Hydrobius arcticus (Fig. 4D) and H. rottenbergii (Fig. 4E) are characterized by weak C-bands on the X chromosome. Hydrobius pui (Fig. 4I) bears weak centromeric C-bands.</p> <p>Note: Smith (1960) mentions 2 n = 18 for H. fuscipes (sensu lato) without any additional information. This agrees with the data presented here.</p> </div>	https://treatment.plazi.org/id/356287D9FF9FAF3C4E28F98BFDB66A40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF92AF3C4FD6FA83FAA36BA1.text	356287D9FF92AF3C4FD6FA83FAA36BA1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrochara BERTHOLD 1827	<div><p>GENUS HYDROCHARA BERTHOLD, 1827</p> <p>(FIG. 6D–H, L)</p> <p>Material examined: Hydrochara caraboides (Linnaeus, 1758): 2 spec. (BMNH): Greece, Corfu, R. Angus lgt.; 1 spec. (BMNH): Germany, Hamburg, R. Angus lgt. (BMNH).</p> <p>Hydrochara flavipes (Boheman, 1851): 2 spec. (BMNH): Spain, Cáceres, Abadia, 12.v.1990, R. Angus lgt. (BMNH).</p> <p>Karyotype: 2 n = 28 + Xy (♂). The chromosomes showing a gradual decrease in length, the longer</p> <p>chromosomes about 8 µm long, the shorter about 3µm. Centromeric C-bands small but well-developed.</p></div> 	https://treatment.plazi.org/id/356287D9FF92AF3C4FD6FA83FAA36BA1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF92AF3C4CADFECBFB5F68B0.text	356287D9FF92AF3C4CADFECBFB5F68B0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrophilus Geoffroy 1762	<div><p>GENUS HYDROPHILUS GEOFFROY, 1762</p> <p>(FIG. 6A–C, M)</p> <p>Material examined: Hydrophilus piceus: 2 spec. (BMNH): Greece, Corfu, R. Angus lgt.</p> <p>Hydrophilus pistaceus Castelnau, 1840: 1 spec. (BMNH): Spain, Caceres, Abadia, R. Angus lgt.</p> <p>Karyotype: 2 n = 28 + Xy p (♂). The chromosomes are short. C-banding shows all the autosomes with heavy centromeric C-bands accounting for most of their length.</p> <p>Comments on karyotypes: Mitosis and meiosis were studied in four species of Hydrophilus [Hydrophilus acuminatus Motschulsky, 1853, Hydrophilus indicus (Bedel, 1891), Hydrophilus piceus and Hydrophilus triangularis Say, 1823] by previous authors (Arnold, 1909; Asana et al., 1942; Smith, 1953; Agarawal, 1960a), all revealing the karyotype 2 n = 28 + Xy p (♂), which is in agreement with our findings.</p> </div>	https://treatment.plazi.org/id/356287D9FF92AF3C4CADFECBFB5F68B0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FFB8AF164F82FC88FD6B6D9D.text	356287D9FFB8AF164F82FC88FD6B6D9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laccobius (Dimorpholaccobius) Sexual	<div><p>Laccobius (Dimorpholaccobius) Sexual</p> <p>bipunctatus</p> <p>Laccobius (Dimorpholaccobius) Sexual</p> <p>ytenensis</p> <p>Laccobius (Dimorpholaccobius) Sexual</p> <p>striatulus</p> <p>Laccobius (Dimorpholaccobius) Sexual</p> <p>sinuatus</p> <p>Laccobius (Compsolaccobius) Sexual</p> <p>decorus Paracymus aeneus Sexual Paracymus scutellaris Sexual Tormus helmsi Sexual Tormus posticalis Sexual</p> <p>Hydrophilinae: Hydrobiusini</p> <p>Hydrobius fuscipes Sexual</p> <p>Hydrobius arcticus Sexual Hydrobius pui Sexual Hydrobius rottenbergii Sexual Hydrobius subrotundus Sexual Limnohydrobius convexus Sexual Limnoxenus niger Sexual</p> <p>Hydrophilinae: Hydrophilini Sternolophus rufipes Sexual</p></div> 	https://treatment.plazi.org/id/356287D9FFB8AF164F82FC88FD6B6D9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF9EAF314C81FB12FE906A0D.text	356287D9FF9EAF314C81FB12FE906A0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laccobius Erichson 1837	<div><p>GENUS LACCOBIUS ERICHSON, 1837</p> <p>(FIG. 2F–L)</p> <p>Material examined: Laccobius (Compsolaccobius) decorus (Gyllenhal, 1827). 2 males. (BMNH): Sweden, Öland Island, R. Angus lgt.</p> <p>Karyotype: 2 n = 16 + Xy p (♂). Centromeric C-bands strong. All autosomes, and X chromosome, metacentric. Autosomes similarly sized, X chromosome the largest in the nucleus.</p> <p>Comments on karyotypes of Laccobius: Angus &amp; Shaarawi (1997) published the karyotype data of two species of the subgenus Laccobius [Laccobius colon (Stephens, 1829) and Laccobius minutus (Linnaeus, 1758)] and four species of the subgenus Dimorpholaccobius [Laccobius bipunctatus (Fabricius, 1775), Laccobius sinuatus Motschulsky, 1849, Laccobius striatulus (Fabricius, 1801) and Laccobius ytenensis Sharp, 1910]. Laccobius karyotypes all show 2 n = 16 + Xy p (♂). All autosomes and the X chromosome have strong centromeric C-bands, which are particularly strong in Laccobius striatulus and Laccobius sinuatus.</p> </div>	https://treatment.plazi.org/id/356287D9FF9EAF314C81FB12FE906A0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF92AF3C4FCAFEB7FD216839.text	356287D9FF92AF3C4FCAFEB7FD216839.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limnohydrobius REITTER 1909	<div><p>GENUS LIMNOHYDROBIUS REITTER, 1909</p> <p>(FIG. 5A–E)</p> <p>Material examined: Limnohydrobius convexus (Brullé, 1835): 2 spec. (BMNH): France, Corsica, R. Angus lgt.; 2 spec. (BMNH): Spain, Menorca, G. Foster lgt. (BMNH).</p> <p>Karyotype: 2 n = 16 + Xy(♂).Chromosomes1,3–5,8and X showing weak centromeric C-bands. Chromosomes 1–6 metacentric, 7, 8 and X submetacentric with X large, about as long as chromosome 5.</p> <p>Note: Limnohydrobius was recently separated from Hydrobius based on DNA-based phylogenetic data (Short et al., 2017); previously Limnohydrobius species were classified under the latter genus (e.g. Hansen, 1999).</p> </div>	https://treatment.plazi.org/id/356287D9FF92AF3C4FCAFEB7FD216839	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF92AF3C4FC5FC63FDB06E2A.text	356287D9FF92AF3C4FC5FC63FDB06E2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limnoxenus MOTSCHULSKY 1853	<div><p>GENUS LIMNOXENUS MOTSCHULSKY, 1853</p> <p>(FIG. 5F–K)</p> <p>Material examined: Limnoxenus niger (Gmelin, 1790): 2 spec. (BMNH): France, Indre, Brenne, R. Angus lgt.; 1 spec. (BMNH): Greece, Corfu, R. Angus lgt. (BMNH).</p> <p>Karyotype: 2 n = 16 + Xy p (♂). All chromosomes, except the dot-like y, with strong centromeric C-bands. Chromosomes 1, 3, 4 and 7 metacentric, 2 and X subacrocentric, 5–7 acrocentric.</p> <p>TRIBE HYDROPHILINI</p></div> 	https://treatment.plazi.org/id/356287D9FF92AF3C4FC5FC63FDB06E2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF81AF2F4FEBFEEEFDE16873.text	356287D9FF81AF2F4FEBFEEEFDE16873.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Noteropagus d'Orchymont 1919	<div><p>GENUS NOTEROPAGUS DʼORCHYMONT, 1919</p> <p>(FIG. 14A–B, G)</p> <p>Material examined: Noteropagus sp.: (unidentified species illustrated in Fig. 14G). 1 male (NMPC): Taiwan: Nantou County, Yushan National Park, Dongbu Scenic Area, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.93044&amp;materialsCitation.latitude=23.561" title="Search Plazi for locations around (long 120.93044/lat 23.561)">Dong Bu</a> 5.4 km SE of Heshe, rotten banana trunks on the margin of a village, 23.5610°N 120.93044°E, 15.v.2018, M. Fikáček, W.-R. Liang, H.-C. Liu &amp; Y. Minoshima lgt. (2018-TW33).</p> <p>Karyotype: 2 n = 22. Two nuclei from testis show probably the meiotic metaphase I with 11 bivalents. The sex bivalent cannot be recognized, although the smallest element in Figure 14A appears asymmetrical. The remaining bivalents suggest that the autosomes are all more or less similar in size.</p></div> 	https://treatment.plazi.org/id/356287D9FF81AF2F4FEBFEEEFDE16873	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF81AF2F4FECFC99FE166E07.text	356287D9FF81AF2F4FECFC99FE166E07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Omicrogiton D'ORCHYMONT 1919	<div><p>GENUS OMICROGITON DʼORCHYMONT, 1919</p> <p>(FIG. 14C–D, H)</p> <p>Material examined: Omicrogiton insularis dʼ Orchymont, 1919: 1 male, 1 female (BMNH): Taiwan, 4.8 km SEE of Tonglin, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.78434&amp;materialsCitation.latitude=24.04791" title="Search Plazi for locations around (long 120.78434/lat 24.04791)">Beikeng Creek Trail</a>, 24.04791°N 120.78434°E, 3.x.2018, rotten banana stem, M. Fikáček, H.-C. Liu, F.-S. Hu &amp; W.-R. Liang lgt.</p> <p>Karyotype: 2 n = 24 + Xy (♂). Autosome pairs 1–3 and the X chromosome are metacentric and clearly larger than the other pairs. Pairs 4 and 7–12 are subacrocentric to acrocentric, and the smallest autosome is about a quarter the length of the largest. The Y chromosome is dot like.</p></div> 	https://treatment.plazi.org/id/356287D9FF81AF2F4FECFC99FE166E07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF9FAF314FD4FE6FFF166834.text	356287D9FF9FAF314FD4FE6FFF166834.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paracymus THOMSON 1867	<div><p>GENUS PARACYMUS THOMSON, 1867</p> <p>(Fig. 3A–H)</p> <p>Material examined: Paracymus aeneus (Germar, 1824): 2 spec. (BMNH): United Kingdom, England, Isle of Wight, R. Angus lgt.</p> <p>Paracymus scutellaris (Rosenhauer, 1856): 2 spec. (BMNH): United Kingdom, England, Hampshire, New Forest, R. Angus lgt.</p> <p>Karyotype: 2 n = 16 + Xy p (♂). No C-banding was attempted. Autosome 1 is distinctly longer than the others and X is the shortest (apart from y) in both species.</p></div> 	https://treatment.plazi.org/id/356287D9FF9FAF314FD4FE6FFF166834	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF81AF2F4E23FA65FB186AD6.text	356287D9FF81AF2F4E23FA65FB186AD6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paromicrus SCOTT 1913	<div><p>GENUS PAROMICRUS SCOTT, 1913</p> <p>(FIG. 14E–F, I)</p> <p>Material examined: Paromicrus sp.: [unidentified species likely close to Paromicrus affinis (Fig. 14I)]: 2 females (BMNH): Taiwan, 4.8 km SEE of Tonglin, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.78434&amp;materialsCitation.latitude=24.04791" title="Search Plazi for locations around (long 120.78434/lat 24.04791)">Beikeng Creek Trail</a>, 24.04791°N 120.78434°E, 3.x.2018, rotten banana stem, M. Fikáček, H.-C. Liu, F.-S. Hu &amp; W.-R. Liang lgt.</p> <p>K a r y o t y p e: 2 n = 2 6 (♀). A l l t h e c h r o m o s o m e s are subacrocentric to acrocentric and there is a gradual decrease in size along the karyotype, with the smallest chromosomes about half of the length of the largest. The X chromosomes cannot be identified.</p> <p>TRIBE COELOSTOMATINI</p></div> 	https://treatment.plazi.org/id/356287D9FF81AF2F4E23FA65FB186AD6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF84AF2A4E20FF16FDB36800.text	356287D9FF84AF2A4E20FF16FDB36800.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protosternum Sharp 1890	<div><p>GENUS PROTOSTERNUM SHARP, 1890</p> <p>(FIG. 15J–L)</p> <p>Material examined: Protosternum abnormale (dʼ Orchymont, 1913): 1 male (NMPC): Taiwan: Nantou County, Yushan National Park, Dongbu Scenic Area, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.93044&amp;materialsCitation.latitude=23.561" title="Search Plazi for locations around (long 120.93044/lat 23.561)">Dong Bu</a>, 5.4 km SE of Heshe, rotten banana trunks on the margin of a village, 23.5610°N 120.93044°E, 15.v.2018, M. Fikáček, W.- R. Liang, H.-C. Liu &amp; Y. Minoshima lgt. (2018-TW33).</p> <p>Karyotype: 2 n = 32. The material to hand is from the testis and some nuclei at least, particularly that in Figure 15J, appear to be first of meiotic metaphase I and exhibit 16 bivalents. The karyotype hence likely comprise 15 pairs of autosomes plus Xy sex chromosomes.</p> <p>TRIBE SPHAERIDIINI</p></div> 	https://treatment.plazi.org/id/356287D9FF84AF2A4E20FF16FDB36800	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF8FAF214C83FDFAFB4B6FEF.text	356287D9FF8FAF214C83FDFAFB4B6FEF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rygmodus WHITE 1846	<div><p>GENUS RYGMODUS WHITE, 1846</p> <p>(FIG. 13F–H, J)</p> <p>Material examined: Rygmodus modestus White, 1846: 1 male (NMPC): New Zealand: Rangitikei (RI), Ruahine Forest Park, Kashmir Road, sweeping of flowering Gaultheria L. and Brachyglottis repanda J.R.Forster &amp; G.Forster, 39.94°S 176.16859°E, 655 m, 25.xi.2016, M. Fikáček &amp; M. Seidel lgt. (MM25). 1 female (NMPC): New Zealand: Taranaki, 0.2 km S of Pukeiti Garden, 9 km E of Okato, lowland Nothofagus forest, flight interception trap, 370 m, 173.98°S 39.20°E, 10.xii.2016, lgt. M. Fikáček &amp; M. Seidel (MM09).</p> <p>Karyotype: 2 n = 13 + Xy p (♂). There is a gradual decrease in length along the karyotype, with pair 13 about a quarter of the length of pair 1. Most of the chromosomes, including the X, are acrocentric or subacrocentric. The Y chromosome is small, though not dot like, and first metaphase of meiosis shows the typical parachute association, Xy</p> <p>p</p> <p>.</p></div> 	https://treatment.plazi.org/id/356287D9FF8FAF214C83FDFAFB4B6FEF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF84AF2A4FD8FC95FB146AC8.text	356287D9FF84AF2A4FD8FC95FB146AC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sphaeridium Fabricius 1775	<div><p>GENUS SPHAERIDIUM FABRICIUS, 1775</p> <p>(FIG. 14J–Q)</p> <p>Material examined: Sphaeridium scarabaeoides (Linnaeus, 1758): 2 spec. (BMNH): United Kingdom, Surrey, Egham, R. Angus &amp; F. Shaarawi lgt.</p> <p>Sphaeridium lunatum Fabricius, 1792: 2 spec. (BMNH): United Kingdom, Surrey, Tilford, R. Angus &amp; F. Shaarawi lgt.</p> <p>Sphaeridium bipustulatum Fabricius, 1781: 1 spec. (BMNH): United Kingdom, Surrey, Tilford, R. Angus &amp; F. Shaarawi lgt.</p> <p>Comments: Angus et al. (2000) studied the mitotic and meiotic karyotypes of four European species, Sphaeridium bipustulatum, Sphaeridium lunatum, Sphaeridium marginatum and Sphaeridium scarabaeoides. The published figures of karyotypes were of low quality, and we hence provide those of three species again here and add the meiotic figures as well.</p> <p>Karyotype: 2 n = 22 + Xy (♂). In the larger species, Sphaeridium lunatum (Fig. 14J–K) and Sphaeridium scarabaeoides (Fig. 14L), autosome pairs 4 and 5 are subacrocentric to acrocentric and pair 8 subacrocentric, with pair 11 also subacrocentric in Sphaeridium scarabaeoides. In the smaller species, Sphaeridium bipustulatum (Fig. 14M) and Sphaeridium marginatum, autosome pair 5 is metacentric and pairs 4 and 10 are subacrocentric. The X chromosome is medium sized metacentric and the Y is dot like in all species.</p> <p>TRIBE MEGASTERNINI</p></div> 	https://treatment.plazi.org/id/356287D9FF84AF2A4FD8FC95FB146AC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF92AF3C4CB9FBDAFA866EBB.text	356287D9FF92AF3C4CB9FBDAFA866EBB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sternolophus SOLIER 1834	<div><p>GENUS STERNOLOPHUS SOLIER, 1834</p> <p>(FIG. 6I–K)</p> <p>Material examined: Sternolophus solieri Castelnau, 1840: 1 male (BMNH): Egypt, surroundings of Cairo, 10 th Ramadan, R. Angus lgt.</p> <p>Karyotype: 2 n = 16 + Xy (♂). Chromosomes 1–7 metacentric, 8 subacrocentric, with an even decrease in length along the karyotype. X chromosome is metacentric, as large as chromosome 1.</p> <p>Note: Agarwal (1960a) provides a meioformula n = 8 + Xy p (♂) for Sternolophus rufipes (Fabricius, 1792), which agrees with the data presented here.</p> </div>	https://treatment.plazi.org/id/356287D9FF92AF3C4CB9FBDAFA866EBB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FFB8AF174F82F925FB3868AA.text	356287D9FFB8AF174F82F925FB3868AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sternolophus solieri Sexual	<div><p>Sternolophus solieri Sexual Tropisternus lateralis Sexual</p> <p>Tropisternus mutatus sapucay Sexual Hydrochara caraboides Sexual</p> <p>18 – This paper</p> <p>18 8 + Xy p Shaarawi &amp; Angus (1991a), Angus et al. (1994)</p> <p>18 8 + Xy p Shaarawi &amp; Angus (1991a) 18 8 + Xy p Shaarawi &amp; Angus (1991a) 18 8 + Xy p Shaarawi &amp; Angus (1991a) 20 9 + Xy p Shaarawi &amp; Angus (1991a) 18 8 + Xy</p> <p>p</p> <p>Angus et al. (1994)</p> <p>18 8 + Xy</p> <p>p</p> <p>Agarawal (1960b)</p> <p>18 – Angus &amp; Shaarawi (1997)</p> <p>18 8 + Xy</p> <p>p</p> <p>Angus &amp; Shaarawi (1997)</p> <p>(as L. biguttatus)</p> <p>18 8 + Xy</p> <p>p</p> <p>Angus &amp; Shaarawi (1997)</p> <p>18 8 + Xy p Angus &amp; Shaarawi (1997) (as L. atratus)</p> <p>18 – Angus &amp; Shaarawi (1997)</p> <p>18 – Angus &amp; Shaarawi (1997)</p> <p>18 8 + Xy This paper</p> <p>18 8 + Xy</p> <p>p</p> <p>This paper</p> <p>18 8 + Xy</p> <p>p</p> <p>This paper</p> <p>18 – This paper</p> <p>18 8 + Xy This paper</p> <p>18 8 + Xy</p> <p>p</p> <p>Smith (1960), this paper</p> <p>18 8 + Xy This paper</p> <p>18 8 + Xy This paper</p> <p>18 8 + Xy This paper</p> <p>18 8 + Xy This paper</p> <p>18 8 + Xy This paper</p> <p>18 8 + Xy</p> <p>p</p> <p>This paper</p> <p>18 8 + Xy</p> <p>p</p> <p>Agarwal (1960a)</p> <p>18 8 + Xy This paper</p> <p>18 8 + Xy</p> <p>p</p> <p>Smith (1953, 1960)</p> <p>18 8 + Xy p Pine et al. (2013)</p> <p>30 14 + Xy This paper</p> <p>Species Reproduction 2 n Meioformula Reference mode</p> <p>Hydrochara flavipes Sexual 30 14 + Xy p This paper Hydrophilus acuminatus Sexual 30 14 + Xy Asana et al. (1942) Hydrophilus indicus Sexual 30 14 + Xy p Agarawal (1960a) Hydrophilus piceus Sexual 30 14 + Xy p Arnold (1909), this paper Hydrophilus pistaceus Sexual 30 14 + Xy This paper Hydrophilus triangularis Sexual 30 14 + Xy p Smith (1953) Chaetarthriinae: Chaetarthriini Chaetarthria simillima Sexual 18 8 + Xy Angus &amp; Shaarawi (1997)</p> <p>p</p> <p>(as C. seminulum), this</p> <p>paper</p> <p>Chaetarthriinae: Anacaenini Anacaena bipustulata Sexual 18 8 + Xy p Shaarawi &amp; Angus (1991 b) Anacaena globulus Sexual 16 7 + Xy p Shaarawi &amp; Angus (1991 b) Anacaena limbata Sexual 18 8 + Xy p Shaarawi &amp; Angus (1991 b) Anacaena lutescens Sexual 18 8 + Xy</p> <p>p</p> <p>Shaarawi &amp; Angus (1991 b)</p> <p>Parthenogenetic 18 – Shaarawi &amp; Angus (1991 b)</p> <p>Parthenogenetic 3 n = 27 – Shaarawi &amp; Angus (1991 b)</p> <p>Anacaena rufipes Sexual 12 5 + NeoXY Shaarawi &amp; Angus (1991 b) Anacaena gaetanae Sexual 18 8 + Xy This paper</p> </div>	https://treatment.plazi.org/id/356287D9FFB8AF174F82F925FB3868AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF8FAF2F4CBFFB11FDDD6A59.text	356287D9FF8FAF2F4CBFFB11FDDD6A59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tormissus BROUN 1893	<div><p>GENUS TORMISSUS BROUN, 1893</p> <p>(FIG. 13B–C, E)</p> <p>Material examined: Tormissus magnulus Broun, 1893: 1 male (NMPC): New Zealand, Wellington (WL), Tararua Range, 1 km W of Titurea Dam, start of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=175.66064&amp;materialsCitation.latitude=-40.4295" title="Search Plazi for locations around (long 175.66064/lat -40.4295)">Greens Rd.</a>, fragment of broadleaf forest with sparse understory with ferns and Pandanus: pitfall trap, 40.4295°S 175.66064°E, 145 m, 28.xi.2016, M. Fikáček &amp; M. Seidel lgt. (MM31). 1 male (NMPC): New Zealand: Wellington, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=174.73285&amp;materialsCitation.latitude=-41.29683" title="Search Plazi for locations around (long 174.73285/lat -41.29683)">Wright Hill Reserve</a>, along unnamed stream along Deliverance Tk., 41.29683°S 174.73285°E, 190 m, 18.xi.2017, D. Sadílek lgt. (2017- NZ 07; DNA extraction NZ 718).</p> <p>Karyotype: 2 n = 26 + Xy p (♂). Based on the specimen from Titurea Dam (Fig. 13B, C, E), the karyotype comprises 13 pairs of autosomes plus Xy sex chromosomes. The specimen from Wellington provided two suitable mitotic nuclei and one meiotic nucleus, all showing one chromosome pair less (not illustrated here), despite the Titurea and Wellington specimens being conspecific as confirmed by their DNA barcodes. The Wellington specimen clearly shows the Xy</p> <p>p</p> <p>configuration of sex chromosomes. We suppose that the lower number of chromosomes in the Wellington specimen is caused by a loss of a chromosome pair during preparation.</p> <p>SUBFAMILY SPHAERIDIINAE</p> <p>TRIBE OMICRINI</p></div> 	https://treatment.plazi.org/id/356287D9FF8FAF2F4CBFFB11FDDD6A59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF9FAF314E3AFC4EFDC26D32.text	356287D9FF9FAF314E3AFC4EFDC26D32.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tormus SHARP 1884	<div><p>GENUS TORMUS SHARP, 1884</p> <p>(FIG. 3I–M)</p> <p>Material examined: Tormus helmsi Sharp, 1884: 1 female (NMPC): New Zealand: Buller (BR) <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=172.05333&amp;materialsCitation.latitude=-42.29202" title="Search Plazi for locations around (long 172.05333/lat -42.29202)">Tobins Bridge</a> at Hwy. 7 (11.6 km NWW of Springs Junction), 42.29202°S 172.05333°E, 527 m, 3.xii.2016, M. Fikáček &amp; M. Seidel lgt. (MM51).</p> <p>Tormus posticalis (Broun, 1917): 1 male (NMPC): New Zealand, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=168.54784&amp;materialsCitation.latitude=-44.1235" title="Search Plazi for locations around (long 168.54784/lat -44.1235)">Westland</a> (WD), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=168.54784&amp;materialsCitation.latitude=-44.1235" title="Search Plazi for locations around (long 168.54784/lat -44.1235)">Stream</a> at Jackson River Rd., Red Hill area, 44°07.41’S 168°32.87’E, 150 m, 7.xii.2017, sifting dry moss, M. Fikáček, D. Sadílek &amp; V. Sýkora lgt. (2017- NZ 73).</p> <p>Karyotype: 2 n = 16 + Xy p (♂). All autosomes and the X chromosome are metacentric. Autosomal pairs 1 and 2 are distinctly longer than the others which decrease in length gradually.</p> <p>TRIBE HYDROBIUSINI</p></div> 	https://treatment.plazi.org/id/356287D9FF9FAF314E3AFC4EFDC26D32	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
356287D9FF92AF384CBAF9D9FD7A6B81.text	356287D9FF92AF384CBAF9D9FD7A6B81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tropisternus Solier 1834	<div><p>GENUS TROPISTERNUS SOLIER, 1834</p> <p>Comments on karyotypes: The meioformula of Tropisternus lateralis (Fabricius, 1775) was published by Smith (1953, 1960), the karyotype and meiosis of Tropisternus mutatus d’Orchymont, 1921 was studied by Pine et al. (2013). Published data indicate that Tropisternus, like Sternolophus, has 2 n = 16 + Xy</p> <p>p</p> <p>(♂).</p> <p>SUBFAMILY CHAETARTHRIINAE</p></div> 	https://treatment.plazi.org/id/356287D9FF92AF384CBAF9D9FD7A6B81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Angus, Robert B;Sadílek, David;Shaarawi, Fatma;Dollimore, Hayley;Liu, Hsing-Che;Seidel, Matthias;Sýkora, Vít;Fikáček, Martin	Angus, Robert B, Sadílek, David, Shaarawi, Fatma, Dollimore, Hayley, Liu, Hsing-Che, Seidel, Matthias, Sýkora, Vít, Fikáček, Martin (2021): Karyotypes of water scavenger beetles (Coleoptera: Hydrophilidae): new data and review of published records. Zoological Journal of the Linnean Society 192 (3): 958, DOI: 10.1093/zoolinnean/zlaa105, URL: https://academic.oup.com/zoolinnean/article/192/3/958/5930869
