taxonID	type	description	language	source
3561FE47FFB3FFBE1588FA71FD41EA85.taxon	diagnosis	Diagnosis: Body elongate, widest near level of testes or slightly posterior; length / width ratio c. 2.8 – 8.7. Tegument with small to minute spines. Oral sucker funnel-shaped, with large oral spines, opens terminally. Ventral sucker unspecialised, embedded in ventrogenital sac. Ratio oral / ventral sucker width c. 1.6 – 2.6. Forebody occupies c. 1 / 2 – 2 / 3 body length. Prepharynx shorter than oesophagus. Oesophagus long. Intestinal bifurcation in approximately mid-forebody, well anterior to ventral sucker. Caeca blind, terminate close to posterior end of body. Testes two, opposite to oblique, in anterior hindbody. Seminal vesicle tubulosaccular. Common genital pore immediately anterior to ventral sucker. Gonotyl absent. Ovary deeply lobed, ventral to or immediately anterior to testes. Seminal receptacle tubulosaccular, posterior to ovary, may extend well posterior to testes. Vitelline follicles in two lateral groups, may extend from ovary to near intestinal bifurcation. Uterine coils restricted to hindbody, extend from posterior end of body to ventral sucker. Excretory vesicle Yshaped; arms reach to level of pharynx or between pharynx and oral sucker. In marine fishes (Haemulidae); tropical Indo-West Pacific.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB3FFBE1588FA71FD41EA85.taxon	etymology	Etymology: The name Beluesca is derived from the Latin belua, meaning beast or monster and esca, meaning bait. It is proposed in recognition of the original purpose for capturing the host individual found to harbour the type-species, which was as bait for a large serranid off Lizard Island. It is to be treated as feminine. Differential diagnosis: Beluesca n. gen. is distinguished from all other cryptogonimid genera by the combination of an elongate body, funnel-shaped oral sucker, the relatively small number of large oral spines, highly lobed ovary, opposite to slightly oblique testes, uterine loops that are restricted to the hindbody and extend well posterior to the testes, and vitelline follicles that may extend from the ovary into the forebody, but do not extend anterior to the intestinal bifurcation.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB4FFBE1588FD9BFB77ED53.taxon	materials_examined	Type host: Plectorhinchus gibbosus (Lacepède). Type locality: Lizard Island, Great Barrier Reef (14 ° 40 ’ S; 145 ° 27 ’ E), Queensland, Australia. Site: Intestine and pyloric caeca.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB4FFBE1588FD9BFB77ED53.taxon	description	Prevalence: 1 of 1 (100 %) at Lizard Island, 0 of 5 at Heron Island.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB4FFBE1588FD9BFB77ED53.taxon	materials_examined	Deposited specimens: holotype G 227654, eleven paratypes G 227655 – G 227665.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB4FFBE1588FD9BFB77ED53.taxon	etymology	Etymology: The epithet littlewoodi is in honour of Dr Tim Littlewood, in recognition of his contributions to platyhelminth systematics.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB4FFBE1588FD9BFB77ED53.taxon	description	Description: Based on 12 specimens. Body elongate, longer than wide, 2083 (1744 – 2472) long by 285 (240 – 368) wide; length / width ratio 7.4 (6.7 – 8.7). Oral sucker funnel-shaped, 234 (202 – 273) long by 207 (182 – 250) wide. Oral spines 21 (20 – 22), length 51 (40 – 64). Ventral sucker 91 (78 – 111) long by 108 (91 – 127) wide. Ratio oral / ventral sucker width 1.9 (1.6 – 2.2). Forebody occupying 52 (48 – 58) % of body length. Prepharynx 127 (59 – 182) long. Pharynx 68 (55 – 78) long by 67 (59 – 75) wide. Ventral sucker / pharynx width ratio 1.6 (1.4 – 1.8). Oesophagus 304 (241 – 384) long. Intestinal bifurcation at mid-forebody, well anterior to ventral sucker. Intestinal caeca blind, 1227 (1024 – 1456) long, terminate close to posterior end of body. Testes two, opposite to slightly oblique, in mid- to anterior hindbody, 181 (137 – 247) long by 116 (94 – 137) wide. Seminal vesicle tubulosaccular, between testes and ventral sucker. Genital pore immediately anterior to ventral sucker. Ovary deeply lobed 145 (114 – 189) long by 131 (104 – 182) wide, ventral to or immediately anterior to testes. Laurer’s canal present. Seminal receptacle tubulosaccular, posterior to ovary, may extend well posterior to testes. Vitelline follicles in two lateral groups, extend from level of ovary to midway between ventral sucker and intestinal bifurcation. Uterine coils restricted to hindbody, extend from posterior end of body to ventral sucker. Eggs small, darkly tanned, 17 (16 – 21) long by 9 (7 – 10) wide. Excretory vesicle Yshaped, bifurcates dorsal to ovary; arms extend anterior to pharynx, 1746 (1456 – 2064) long.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB4FFBD1588F956FDE5E82D.taxon	materials_examined	Type host: Plectorhinchus gibbosus (Lacepède). Type locality: Heron Island, Great Barrier Reef (23 ° 26 ’ S; 151 ° 54 ’ E), Queensland, Australia. Site: Intestine and pyloric caeca. Prevalence: 1 of 5 (20 %) at Heron Island, 0 of 1 at Lizard Island. Deposited specimens: holotype G 227666, ten paratypes G 227667 – 227676.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB4FFBD1588F956FDE5E82D.taxon	etymology	Etymology: The epithet longicolla is derived from the Latin longus, meaning long and collum, meaning neck, referring to the long forebody of this species.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB4FFBD1588F956FDE5E82D.taxon	description	Description: Based on 11 specimens. Body elongate, longer than wide, 2361 (2216 – 2536) long by 357 (320 – 384) wide; length / width ratio 6.7 (6 – 7.5). Oral sucker funnel-shaped, 226 (208 – 244) long by 186 (176 – 195) wide. Oral spines 19 (19 – 21), length 64 (51 – 82). Ventral sucker 94 (91 – 96) long by 98 (94 – 109) wide. Ratio oral / ventral sucker width 1.9 (1.8 – 2). Forebody occupying 66 (64 – 67) % of body length. Prepharynx much shorter than oesophagus, 56 (46 – 85) long. Pharynx 72 (68 – 78) long by 71 (68 – 75) wide. Ventral sucker / pharynx width ratio 1.4 (1.3 – 1.5). Oesophagus 687 (592 – 774) long. Intestinal bifurcation near midforebody, well anterior to ventral sucker. Intestinal caeca blind, 1099 (1053 – 1157) long, terminate close to posterior end of body. Testes two, opposite to oblique, in mid- to anterior hindbody, 259 (228 – 304) long by 152 (125 – 189) wide. Seminal vesicle tubulosaccular, between testes and ventral sucker. Genital pore immediately anterior to ventral sucker. Ovary deeply lobed 138 (99 – 163) long by 132 (109 – 144) wide, ventral to or immediately anterior to testes. Laurer’s canal present. Seminal receptacle tubulosaccular, posterior to ovary, may extend well posterior to testes. Vitelline follicles in two lateral groups, extend from level of ventral sucker to midway between ventral sucker and intestinal bifurcation, do not extend posterior to ventral sucker. Uterine coils restricted to hindbody, extend from posterior end of body to ventral sucker. Eggs small, darkly tanned, 16 (13 – 18) long by 7 (7 – 8) wide. Excretory vesicle Y-shaped, bifurcates dorsal to ovary; arms extend to pharynx, 2038 (1888 – 2224) long.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB7FFBD1588FC63FD23EC63.taxon	diagnosis	Diagnosis: Body lanceolate, widest near level of ovary; length / width ratio c. 3.7 – 6. Tegument with small to minute spines. Oral sucker funnel-shaped, with large oral spines, opens terminally. Ventral sucker unspecialised, embedded in ventrogenital sac. Ratio oral / ventral sucker width c. 1.6 – 2.2. Forebody occupies c. 1 / 2 body length. Prepharynx long, much longer than oesophagus. Oesophagus short. Intestinal bifurcation dorsal to or immediately anterior to ventral sucker. Caeca blind, terminate close to posterior end of body. Testes two, entire, tandem, adjacent, in mid-hindbody. Seminal vesical saccular, between ovary and ventral sucker. Common genital pore immediately anterior to ventral sucker. Gonotyl absent. Ovary entire, anterior and adjacent to or ventral to anterior testis. Laurer’s canal present. Seminal receptacle saccular, between ovary and ventral sucker. Vitelline follicles in two lateral groups, extend from ovary to pharynx. Uterine coils restricted to hindbody, extend from posterior end of body to ventral sucker. Excretory vesicle Y-shaped; arms reach to near pharynx. In marine fishes (Haemulidae); tropical Indo-West Pacific.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB7FFBD1588FC63FD23EC63.taxon	etymology	Etymology: The name Chelediadema is derived from the Greek chele, meaning nail or claw and diadema, meaning crown, referring to the crown of large oral spines that are analogous to claws. It is to be treated as neuter. Differential diagnosis: Chelediadema n. gen. is distinguished from all other cryptogonimid genera by the combination of a lanceolate body, the relatively small number of large oral spines, a prepharynx that is much longer than the oesophagus, caeca that terminate close to the posterior end of the body, tandem testes, uterine loops that are extensive in the hindbody and extend well posterior to the testes, and vitelline follicles that extend from the ovary to the pharynx.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB9FFB31588FF43FDDCEFA3.taxon	materials_examined	Type host: Diagramma labiosum (Macleay) Perciformes, Haemulidae. Additional hosts: Plectorhinchus albovittatus (Rüppell); P. gibbosus (Lacepède). Type locality: Heron Island, Great Barrier Reef (23 ° 26 ’ S; 151 ° 54 ’ E), Queensland, Australia. Additional localities: Lizard Island, Great Barrier Reef (14 ° 40 ’ S; 145 ° 27 ’ E), Queensland, Australia.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB9FFB31588FF43FDDCEFA3.taxon	description	Site: Intestine and pyloric caeca. Prevalence: Diagramma labiosum, 7 of 43 (16 %) at Heron Island, 0 of 7 at Lizard Island; P. albovittatus, 1 of 1 (100 %) at Lizard Island; P. gibbosus, 1 of 1 (100 %) at Lizard Island, 0 of 5 at Heron Island.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB9FFB31588FF43FDDCEFA3.taxon	materials_examined	Deposited specimens: holotype G 227677, nine paratypes G 227678 – G 227686.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB9FFB31588FF43FDDCEFA3.taxon	etymology	Etymology: The epithet marjoriae is in honour of Mrs Marjorie Neill in recognition of her support and encouragement of this work.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
3561FE47FFB9FFB31588FF43FDDCEFA3.taxon	description	Description: Based on 10 specimens. Body lanceolate, longer than wide, 1150 (930 – 1368) long by 240 (195 – 309) wide; length / width ratio 4.9 (3.7 – 6.0). Oral sucker funnel-shaped, 117 (102 – 134) long by 92 (83 – 106) wide. Oral spines 14 (14 – 15), length 32 (24 – 47). Ventral sucker 64 (56 – 69) long by 67 (54 – 75) wide. Ratio oral / ventral sucker width 1.37 (1.24 – 1.59). Forebody occupying 50 (46 – 53) % body length. Prepharynx 305 (227 – 381) long. Pharynx 67 (54 – 80) long by 66 (58 – 85) wide. Ventral sucker / pharynx width ratio 1.03 (0.87 – 1.11). Oesophagus 46 (26 – 70) long. Intestinal bifurcation at level of or immediately anterior to ventral sucker. Intestinal caeca blind, 591 (501 – 670) long, terminate close to posterior end of body. Testes two, tandem, adjacent, in mid-hindbody; anterior testis 106 (90 – 122) long by 112 (86 – 136) wide, posterior testis 122 (102 – 147) long by 111 (90 – 144) wide. Seminal vesicle saccular, between ovary and ventral sucker. Genital pore immediately anterior to ventral sucker. Ovary entire, anterior and adjacent to or ventral to anterior testis, 74 (67 – 80) long by 70 (58 – 90) wide. Laurer’s canal present. Seminal receptacle saccular, between ovary and ventral sucker. Vitelline follicles in two lateral groups, extend from level of ovary to pharynx. Uterine coils restricted to hindbody, extend from posterior end of body to ventral sucker. Eggs small, darkly tanned, 21 (18 – 24) long by 9 (7 – 11) wide. Excretory vesicle Y-shaped, bifurcates dorsal to ovary; arms extend to posterior end of pharynx, 661 (520 – 767) long.	en	Miller, Terrence L., Cribb, Thomas H. (2007): Two new cryptogonimid genera Beluesca n. gen. and Chelediadema n. gen. (Digenea: Cryptogonimidae) from tropical Indo-West Pacific Haemulidae (Perciformes). Zootaxa 1543: 45-60, DOI: 10.5281/zenodo.177912
