taxonID	type	description	language	source
E298CAFD44CB561FBDB4CFB02D697F45.taxon	materials_examined	Additional material. PNG: Sandaun: 2 males " Papua New Guinea: Sandaun, Ofektaman, 820 m, 17. x. 2008, 5.04.113 S 141.35.841 E, Ibalim (PNG 190) ", one with an additional green label " M. Balke 3727 " (ZSM).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
E298CAFD44CB561FBDB4CFB02D697F45.taxon	diagnosis	Diagnosis. For complete description, see Balke (1998: 322). Beetle medium-sized (TL-H 4.0 - 4.35 mm), oblong-oval; piceous, sometimes with paler pronotal sides; dorsally shiny, with extremely fine, inconspicuous punctation and weakly impressed microreticulation; pronotum with distinct lateral bead (Fig. 1); male protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta; male protarsomere 5 ventrally with anterior band of more than 60 and posterior row of 14 relatively long setae (Fig. 5 D); median lobe simple, in lateral view, evenly tapering to broadly pointed, somehow elongate and gently curved downwards apex, in ventral view, apex more or less rounded; paramere with distinct dorsal notch and large, long subdistal part, dorsal setae numerous and strong, subdistal slightly denser and longer than proximal ones, the latter more or less distinct (Fig. 5 A-C).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
E298CAFD44CB561FBDB4CFB02D697F45.taxon	distribution	Distribution. Indonesia: Papua Province: Pegunungan Bintang Regency and Papua New Guinea: Sandaun Province (Fig. 11).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
941A80ECFECC53C9A730A5A8FB08B04E.taxon	description	Description. Body size and form: Beetle small: TL-H 3.35 mm, TL 3.8 mm, MW 1.8 mm, with oblong-oval habitus. Colouration: Dark brown, with reddish pronotal sides and head anteriorly. Head reddish brown, paler anteriorly; pronotum dark brown on disc, with reddish sides; elytra dark brown, with weakly indicated reddish sutural lines; head appendages and legs proximally yellowish, legs distally darker, reddish brown (Fig. 13). Surface sculpture: Shiny dorsally, with weak and sparse punctation and weakly impressed microreticulation. Head with fine and sparse punctation (spaces between punctures 2 - 3 times size of punctures); diameter of punctures equal to or smaller than diameter of cells of microreticulation. Pronotum with much finer and sparser punctation than on head, very inconspicuous. Punctation on elytra invisible. Pronotum and elytra with weakly impressed microreticulation; head with microreticulation slightly stronger. Metaventrite, metacoxae, and abdominal ventrites distinctly microreticulate. Metacoxal plates with longitudinal strioles and weak transverse wrinkles; abdominal ventrites with strioles. Punctation on venter invisible; inconspicuous on two last abdominal ventrites. Structures: Pronotum with narrow lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, very slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 slightly truncate. Male: Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior band of more than 30 and posterior row of 7 relatively long setae (Fig. 19 D). Abdominal ventrite 6 with 7 - 8 lateral striae on each side. Median lobe short, robust, evenly tapering to slightly pointed apex in lateral and ventral views; apex slightly sinuate in lateral view (Fig. 19 A, B). Paramere as in Fig. 19 C. Female: Unknown.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
941A80ECFECC53C9A730A5A8FB08B04E.taxon	distribution	Distribution. Papua New Guinea: Madang Province, Bismarck Range (Fig. 25).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
941A80ECFECC53C9A730A5A8FB08B04E.taxon	etymology	Etymology. The species is named after Akameku Village. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
D998CB1A157159F89BA0272DABEE75AF.taxon	description	Description. Body size and form: Beetle small: TL-H 3.4 mm, TL 3.8 mm, MW 1.8 mm, with oblong-oval habitus. Colouration: Reddish brown. Head reddish brown. Pronotum reddish brown, dark brown on disc and almost yellowish on lateral sides. Elytra brown, with reddish sutural lines. Head appendages and legs proximally yellowish, legs distally darker, reddish (Fig. 26). Surface sculpture: Submatt dorsally, with strong and dense punctation and strongly impressed microreticulation. Head with dense and coarse punctation (spaces between punctures 0 - 1 times size of punctures); diameter of punctures equal to or larger than diameter of cells of microreticulation. Pronotum and elytra with finer and sparser punctation than on head, very distinct, more even on elytra. Pronotum and elytra with strongly impressed microreticulation; head with microreticulation stronger. Metaventrite, metacoxae, and abdominal ventrites distinctly microreticulate, but shiny. Metacoxal plates with longitudinal strioles and weak transverse wrinkles; abdominal ventrites with strioles. Punctation on venter weak; more distinct on two last abdominal ventrites. Structures: Pronotum with lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 broadly rounded. Male: Protarsomere 4 with anterolateral seta very long and thin, evenly curved, in size equal to more laterally situated large seta. Protarsomere 5 ventrally with anterior band of ca. 80 and posterior row of ca. 16 relatively long setae, which mixed up medially (Fig. 30 D). Abdominal ventrite 6 with 4 - 6 lateral striae on each side. Median lobe in lateral view with apical lobes distinct but shallow, slightly rounded, " nose " elongate, large (Fig. 30 A, B). Paramere as in Fig. 30 C. Female: Unknown.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
D998CB1A157159F89BA0272DABEE75AF.taxon	distribution	Distribution. Papua New Guinea: Morobe Province (Fig. 34).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
D998CB1A157159F89BA0272DABEE75AF.taxon	etymology	Etymology. The species is named after Aseki Village. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
5D8909D7AF755338AA64C5DC20DBF7A0.taxon	materials_examined	Additional material. Madang: 2 males " Papua New Guinea: Madang, Simbai area, 1200 m, 11. iii. 2007, 05.13.333 S 144.37.611 E, Kinibel (PNG 153) (ZSM). 15 males, 6 females " Papua New Guinea: Madang, Simbai-Mombeen, 1100 m, 11. iii. 2007, 05.12.876 S 144.41.759 E, Kinibel (PNG 154), one male with an additional green label " DNA M. Balke 3318 " (NHMW, ZSM). 1 male, 14 females " Papua New Guinea: Madang, Keki-Sewan, Adalbert [sic!] Mts., 700 m, 30. xi. 2006, nr 04.41.802 S 145.25.460 E, Binatang Boys (PNG 120) " (ZSM). 4 males, 4 females " Papua New Guinea: Madang, Adalbert [sic!] Mts., creek nr Keki, 790 m, 28. xi. 1994, 04.42.300 S 145.25.089 E, Binatang Boys leg. (PNG 53 a) " (NHMW, ZSM). 2 males, 5 females " Papua New Guinea: Madang, Adalbert [sic!] Mts., Keki, 850 m, 4. v. 2006, nr 04.42.300 S 145.25.089 E, Manaono leg. (PNG 52) " (ZSM). 83 males, 84 females " Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 1200 m, 24. xi. 2006, nr 05.52.754 S 145.23.209 E, Balke & Kinibel (PNG 110) ", one of them with an additional green label " DNA M. Balke 1521 " (NHMW, ZSM). 7 males, 3 femlaes " Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 1500 m, 24. xi. 2006, 05.51.964 S 145.23.604 E, Balke & Kinibel (PNG 111) " (ZSM). 3 females " Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 800 m, 24. xi. 2006, 05.50.021 S 145.24.664 E, Balke & Kinibel (PNG 112) " (ZSM). 1 male, 1 female " Papua New Guinea: Madang, Akameku - Brahmin, Bismarck Range, 750 m, 25. xi. 2006, 05.49.892 S 145.24.491 E, Balke & Kinibel (PNG 113) " (ZSM). 1 male " Ibisca Niugini, PNG 26 - 28. x. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - P- 1 / 8 - d 02 / Plot 16 / P 1608 Vial 18767 " (ZSM). 1 male " Ibisca Niugini, PNG 26 - 28. x. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - T- 1 / 8 - d 02 / Plot 20 / P 1640 Vial 18781 " (ZSM). 3 females " Ibisca Niugini, PNG 26 - 28. x. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - P- 1 / 8 - d 02 / Plot 16 / P 1608 Vial 18767 " (ZSM). 1 female " Ibisca Niugini, PNG 26 - 28. x. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - O- 1 / 8 - d 02 / Plot 15 / P 1600 Vial 18763 " (ZSM). 1 female " Ibisca Niugini, PNG 26 - 28. x. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - K- 1 / 8 - d 02 / Plot 11 / P 1568 Vial 17122 " (ZSM). 1 female " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - A- 2 / 8 - d 03 / Plot 1 / P 1489 Vial 17237 " (ZSM). 1 female " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - C- 2 / 8 - d 03 / Plot 3 / P 1505 Vial 17179 " (ZSM). 6 females " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1529 Vial 16853 " (ZSM). 1 male, 4 females " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 1200 m - 5,720873833 145,269470 2 MW 1200 / P 1553 Vial 09007 " (ZSM). 1 male, 4 females " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1545 Vial 16863 " (ZSM). 1 male, 7 females " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1561 Vial 16873 " (ZSM). 1 male " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - A- 2 / 8 - d 03 / Plot 1 / P 1489 Vial 17237 " (ZSM). 7 females " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - E- 2 / 8 - d 03 / Plot 5 / P 1521 Vial 17210 " (ZSM). 1 female " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1569 Vial 17302 " (ZSM). 3 females " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1601 Vial 17313 " (ZSM). 11 females " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1595 Vial 18799 " (ZSM). 1 female " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1593 Vial 17462 " (ZSM). 2 males " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1577 Vial 18802 " (ZSM). 1 male " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1625 Vial 18821 " (ZSM). 1 female " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1633 Vial 18848 " (ZSM). 1 female " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1617 Vial 18813 " (ZSM). 1 female " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1601 Vial 17313 " (ZSM). 2 females " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1585 Vial 18825 " (ZSM). 1 male, 13 females " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1609 Vial 18855 " (ZSM). 1 female " Ibisca Niugini, PNG 29 - 31. x. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - D- 3 / 8 - d 05 / Plot 4 / P 1514 Vial 16946 " (ZSM). 1 female " Ibisca Niugini, PNG 29 - 31. x. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - F- 3 / 8 - d 05 / Plot 6 / P 1530 Vial 16936 " (ZSM). 1 female " Ibisca Niugini, PNG 29 - 31. x. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - A- 3 / 8 - d 05 / Plot 1 / P 1490 Vial 16931 " (ZSM). 2 females " Ibisca Niugini, PNG 29 - 31. x. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - H- 3 / 8 - d 05 / Plot 8 / P 1546 Vial 17249 " (ZSM). 1 female " Ibisca Niugini, PNG 29 - 31. x. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - C- 3 / 8 - d 05 / Plot 3 / P 1506 Vial 17235 " (ZSM). 1 female " Ibisca Niugini, PNG 29 - 31. x. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - J- 3 / 8 - d 05 / Plot 10 / P 1562 Vial 16881 " (ZSM). 2 females " Ibisca Niugini, PNG 30. x. - 1. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1578 Vial 17625 " (ZSM). 4 females " Ibisca Niugini, PNG 30. x. - 1. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1570 Vial 17585 " (ZSM). 1 female " Ibisca Niugini, PNG 30. x. - 1. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1634 Vial 17605 " (ZSM). 1 female " Ibisca Niugini, PNG 31. x. - 2. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1547 Vial 17303 " (ZSM). 1 female " Ibisca Niugini, PNG 31. x. - 2. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1515 Vial 17326 " (ZSM). 1 female " Ibisca Niugini, PNG 31. x. - 2. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1491 Vial 17331 " (ZSM). 1 female " Ibisca Niugini, PNG 31. x. - 2. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1531 Vial 17347 " (ZSM). 1 female " Ibisca Niugini, PNG 31. x. - 2. xi. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - E- 4 / 8 - d 07 / Plot 5 / P 1523 Vial 17348 " (ZSM). 2 females " Ibisca Niugini, PNG 1 - 3. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1579 Vial 18787 " (ZSM). 8 females " Ibisca Niugini, PNG 1 - 3. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1643 Vial 18794 " (ZSM). 1 female " Ibisca Niugini, PNG 1 - 3. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1595 Vial 18799 " (ZSM). 3 females " Ibisca Niugini, PNG 1 - 3. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1571 Vial 16947 " (ZSM). 1 female " Ibisca Niugini, PNG 1 - 3. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1635 Vial 16968 " (ZSM). 1 female " Ibisca Niugini, PNG 1 - 3. xi. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - S- 4 / 8 - d 08 / Plot 19 / P 1635 Vial 16968 - CODYTI " (ZSM). 1 male, 10 females " Ibisca Niugini, PNG 1 - 3. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1611 Vial 16950 " (ZSM). 2 females " Ibisca Niugini, PNG 1 - 3. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1611 Vial 16950 " (ZSM). 2 females " Ibisca Niugini, PNG 1 - 3. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1635 Vial 16968 " (ZSM). 1 female " Ibisca Niugini, PNG 2 - 4. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1516 Vial 17287 " (ZSM). 2 females " Ibisca Niugini, PNG 2 - 4. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1548 Vial 17297 " (ZSM). 2 females " Ibisca Niugini, PNG 2 - 4. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1532 Vial 17314 " (ZSM). 1 female " Ibisca Niugini, PNG 2 - 4. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1492 Vial 17355 " (ZSM). 1 male, 2 females " Ibisca Niugini, PNG 2 - 4. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1564 Vial 16857 " (ZSM). 1 female " Ibisca Niugini, PNG 2 - 4. xi. 2012 Mount Wilhelm 1200 m ", " - 5,72090292 145,2714691 FIT-MW 1200 - C- 5 / 8 - d 09 / Plot 3 / P 1508 Vial 14052 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 2 - 4. xi. 2012 Mount Wilhelm 1200 m ", " - 5,72090292 145,2714691 FIT-MW 1200 - I- 5 / 8 - d 09 / Plot 9 / P 1556 Vial 17374 " (ZSM). 1 female " Ibisca Niugini, PNG 2 - 4. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 ", " FIT-MW 1200 - E- 5 / 8 - d 09 / Plot 5 / P 1524 Vial 16861 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 3 - 5. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1636 Vial 17324 " (ZSM). 2 males " Ibisca Niugini, PNG 3 - 5. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1612 Vial 17292 " (ZSM). 1 male " Ibisca Niugini, PNG 3 - 5. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1572 Vial 18837 " (ZSM). 1 female " Ibisca Niugini, PNG 3 - 5. xi. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - O- 5 / 8 - d 10 / Plot 15 / P 1604 Vial 17325 " (ZSM). 1 male " Ibisca Niugini, PNG 4 - 6. xi. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - F- 6 / 8 - d 11 / Plot 6 / P 1533 Vial 17257 " (ZSM). 1 male " Ibisca Niugini, PNG 5 - 7. xi. 2012 Mount Wilhelm 1200 m ", " - 5,720873833 145,2694702 FIT-MW 1200 - K- 6 / 8 - d 12 / Plot 11 / P 1573 Vial 17082 " (ZSM). 1 female " Ibisca Niugini, PNG 8 - 10. xi. 2012 Mount Wilhelm 1200 m - 5,721022129 145,2703094 MW 1200 / P 1503 Vial 16995 " (ZSM). 1 female " Ibisca Niugini, PNG 8 - 10. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1527 Vial 16879 " (ZSM). 1 female " Ibisca Niugini, PNG 8 - 10. xi. 2012 Mount Wilhelm 1200 m - 5,720873833 145,2694702 MW 1200 / P 1503 Vial 16995 " (ZSM). 1 male, 1 female " Ibisca Niugini, PNG 9 - 11. xi. 2012 Mount Wilhelm 1200 m, " " - 5,720873833 145,2694702 FIT-MW 1200 - T- 8 / 8 - d 16 / Plot 20 / P 1647 Vial 17039 - CODYTI " (ZSM). 4 females " Ibisca Niugini, PNG 9 - 11. xi. 2012 Mount Wilhelm 1200 m, " " - 5,720873833 145,2694702 FIT-MW 1200 - P- 8 / 8 - d 16 / Plot 16 / P 1615 Vial 17012 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 9 - 11. xi. 2012 Mount Wilhelm 1200 m, " " - 5,720873833 145,2694702 FIT-MW 1200 - M- 8 / 8 - d 16 / Plot 13 / P 1591 Vial 17035 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 1 - 3. xi. 2012 Mount Wilhelm 1700 m - 5,79269238 145,235611 MW 1700 / P 1961 Vial 06629 " (ZSM). Eastern Highlands: 3 male, 3 females " Papua New Guinea: Eastern Highlands, Bena Bridge, 1400 m, 8. xii. 2007, 06.10.781 S 145.26.034 E, Balke & Sagata (PNG 164) " (ZSM). Simbu / Eastern Highlands: 2 males " Papua New Guinea: Simbu / EHPr. Crater Mountain, Sera - Herowana, Wara Pima, 900 m, 15 IX 2002, Balke & Sagata (PNG 011) " (ZSM). 16 males " Papua New Guinea: Crater Mountain, Sera - Herowana, upper Oh River, 1200 m, 15 IX 2002, Balke & Sagata (PNG 012) " (NHMW, ZSM). 2 males, 4 females " Papua New Guinea: Crater Mountain, Sera - Herowana, Jau river, 1100 m, 15 IX 2002, Balke & Sagata (PNG 013) " (ZSM). 12 males " Papua New Guinea: Simbu / EHPr. Crater Mountain, Sera - Herowana, Jau river, 1000 m, 15 IX 2002, Balke & Sagata (PNG 015) " (NHMW, ZSM). 1 male, 3 females " Papua New Guinea: Simbu / EHP, Crater Mountain, Sera - Herowana, Sima river, 1250 m, 15 IX 2002, Balke & Sagata (PNG 016) " (ZSM). Simbu: 5 males " Papua New Guinea: Supa Haia, 1023 m, 10. ix. 2002, K. Sagata (WB 1) " (NHMW, ZSM). 1 male " Papua New Guinea: Crater Mountain, trek Haia - Wara Sera, 600 m, 12 IX 2002, Balke & Sagata, (PNG 003) " (ZSM). 3 males " Papua New Guinea: Crater Mountain, trek Haia - Wara Sera, 500 m, 12 IX 2002, Balke & Sagata, (PNG 005) " (ZSM). 1 male " Papua New Guinea: Crater Mountain, trek Haia - Wara Sera, 500 m, 12 IX 2002, Balke & Sagata, (PNG 006) " (ZSM). 11 males, 14 females " Papua New Guinea: Simbu / EHPr. Crater Mountain, Wara Sera Station, 800 m, 14 IX 2002, Balke & Sagata (PNG 009) " (NHMW, ZSM). 12 males, 7 females " Papua New Guinea: Simbu / EHPr. Crater Mountain, Wara Sera Station, 800 m, 14 IX 2002, Balke & Sagata (PNG 10) ", one male with additional labels " 257 DNA M Balke " [green], " sp. 17 SEM 19 " (ZSM). 27 males, 31 females " Papua New Guinea: Crater Mountain, Wara Sera Station, 800 m, 14 IX 2002, Balke & Sagata (PNG 010) " (NHMW, ZSM). Morobe: 62 males, 27 females " PAPUA N. G.: Morobe Prov. E Pindiu, Kobau 24.4.1998, 1400 m, leg. A. Riedel " (NHMW, ZSM). 1 male " Papua New Guinea: Morobe, Pindiu, Sulemana, 850 m, 15. x. 2009, 06.25.169 S 147.32.11 E, Inaho (08) (PNG 208) ", " DNA M. Balke 3825 " [green] (ZSM). 2 males " PNG: Huon Peninsula, Morobe Prov., Yus conservation area [5 ° 53 ' 54 " S, 146 ° 48 ' 15 " E], 1398 m, 24. viii. 2010, Bega ", " DNA M. Balke 6531 " [green text], " DNA M. Balke 6532 " [green text] (ZSM). 6 females " PNG: Huon Peninsula, Morobe Prov., Yus conservation area (Y 7), 1398 m, 24. viii. 2010, Huon, Bega " (ZSM). Gulf: 9 males, 8 females " Papua New Guinea: Gulf, Marawaka, nr Ande, 1000 m, 10. xi. 2006, 07.03.598 S 145.44.375 E, Balke & Kinibel (PNG 89) " (NHMW, ZSM).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
5D8909D7AF755338AA64C5DC20DBF7A0.taxon	diagnosis	Diagnosis. For complete description, see Balke (1998: 326). Beetle small to medium-sized: TL-H 3.05 - 3.9 mm, oblong-oval; dorsally uniformly reddish to dark brown or with paler head and sides of pronotum; shiny, with very fine to distinct punctation and usually weakly impressed microreticulation; pronotum with distinct lateral bead (Fig. 16); male protarsomere 4 with anterolateral seta very long and thin, evenly curved, smaller than more laterally situated large seta; male protarsomere 5 ventrally with anterior band of more than 50 and posterior row of 8 relatively long setae (Fig. 22 D); median lobe simple, evenly attenuated to broadly pointed apex in lateral and ventral views; paramere very slightly concave on dorsal side and with long, dense, thin setae, situated along dorsal margin; proximal setae longer that subdistal, more distinct (Fig. 22 A-C).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
5D8909D7AF755338AA64C5DC20DBF7A0.taxon	distribution	Distribution. Papua New Guinea: Madang, Simbu, Eastern Highlands, Morobe and Gulf Provinces (Fig. 25). This is one of the most abundant species in the region.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
BB967394B81D56C7A13525136A9E8C64.taxon	description	Description. Body size and form: Beetle small: TL-H 3.4 - 3.6 mm, TL 3.8 - 4.0 mm, MW 1.85 - 2.0 mm (holotype: TL-H 3.6 mm, TL 4.0 mm, MW 2.0 mm), with oblong-oval habitus. Colouration: Yellow reddish to brown. Head reddish brown to brown, dark brown posterior to eyes. Pronotum yellowish reddish, with small dark area on disc or brown, with paler sides. Elytra yellow reddish to brown. Head appendages and legs proximally yellowish, legs distally darker, reddish brown (Fig. 17). Teneral specimen yellowish. Surface sculpture: Shiny dorsally, with very fine punctation and weakly impressed microreticulation. Elytral punctation and microreticulation finer then in nominotypical subspecies. Elytral punctation usually invisible. Structures: Pronotum with lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 slightly truncate. Male: Protarsomere 4 with anterolateral seta rather long and thing, evenly curved, smaller than more laterally situated large seta. Protarsomere 5 ventrally with anterior band of more than 60 and posterior row of ten relatively long setae (Fig. 23 D). Abdominal ventrite 6 with 6 - 8 lateral striae on each side. Median lobe simple, evenly tapering towards apex in lateral and ventral views; in lateral view, apex elongate, thin, with slightly enlarged, rounded tip (Fig. 23 A, B). Paramere as in Fig. 23 C. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
BB967394B81D56C7A13525136A9E8C64.taxon	distribution	Distribution. Papua New Guinea: Morobe and Central Provinces (Fig. 25).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
BB967394B81D56C7A13525136A9E8C64.taxon	etymology	Etymology. The subspecies is named after Herzog Mts., where the subspecies was the first time discovered. The name is an adjective in the nominative singular.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
807019FED654537594C76E6B28173514.taxon	materials_examined	Additional material. 1 male " IRIAN JAYA Zentralmassive 140 ° 25 ' E 04 ° 24 ' S ", " Borme, 1800 m 16.8.1992 leg. Balke (12, 12 A) ", " Paratypus Copelatus fume Balke des. 1997 " [red] (NHMW).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
807019FED654537594C76E6B28173514.taxon	diagnosis	Diagnosis. For complete description, see Balke (1998: 330). Beetle small (TL-H 3.45 - 3.75 mm), oblong-oval; brown to piceous, usually with paler pronotal sides; dorsally more or less shiny, with fine but conspicuous punctation and weakly impressed microreticulation; pronotum with distinct lateral bead (Fig. 14); male protarsomere 4 with anterolateral seta thin, weakly curved, smaller than more laterally situated large seta; male protarsomere 5 ventrally with anterior band of ca. 70 and posterior row of 6 relatively long setae (Fig. 20 D); median lobe in lateral view evenly attenuated to elongate, thin apex, which slightly pointed in ventral view; paramere slightly concave on dorsal side and with distinct, long, dense, uniform setae, situated along dorsal margin (Fig. 20 A-C).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
807019FED654537594C76E6B28173514.taxon	distribution	Distribution. Indonesia: Papua Province: Pegunungan Bintang Regency, Borme (Fig. 25). The species is known only from the type material.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
4DF0A0EF01DF5152AF146ABE6886DD34.taxon	description	Description. Body size and form: Beetle small: TL-H 3.4 - 3.45 mm, TL 3.7 - 3.75 mm, MW 1.8 mm (holotype: TL-H 3.4 mm, TL 3.7 mm, MW 1.8 mm), with oblong-oval habitus. Colouration: Dark brown, with paler sides of pronotum and head anteriorly. As in E. warasera sp. nov. (Fig. 49). Surface sculpture: Shiny dorsally, with extremely fine and sparse punctation and weakly impressed microreticulation. As in E. warasera sp. nov. Structures: Pronotum with lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 very slightly concave. Male: Protarsomere 4 with anterolateral seta thin and evenly curved, smaller than more laterally situated large seta. Protarsomere 5 ventrally with anterior band of ca. 80 and posterior row of two relatively long setae (Fig. 53 D). Abdominal ventrite 6 with 6 - 8 lateral striae on each side. Median lobe short, curved, with broadly pointed apex in lateral view, and evenly tapering to pointed apex in ventral view (Fig. 53 A, B). Paramere concave on dorsal side, with dorsal setae distinct: subdistal setae only slightly stronger and denser than proximal (Fig. 53 C). Female: Unknown.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
4DF0A0EF01DF5152AF146ABE6886DD34.taxon	distribution	Distribution. Papua New Guinea: Simbu Province, Crater Mountain area (Fig. 54). The species is named after Haia Village. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
757E69DBAB555E01AA5EDD3AA356177C.taxon	materials_examined	Additional material. Morobe: 1 male " Stn. No. 149 A ", " NEW GUINEA: Morobe Dist., Herzog Mts., Vagau, C. 4,000 ft. 4 - 17. i. 1965 ", " M. E. Bacchus. B. M. 1965 - 120 ", " Paratypus Copelatus monae sp. n. Balke des. 1997 " [red], " Exocelina jaseminae (Balke) det. H. Shaverdo 2014 " (BMNH). 2 males " Papua New Guinea: Morobe, Wagau, Herzog Mts., 1150 m, 19. xi. 2006, 06.51.067 S 146.48.068 E, Balke & Kinibel (PNG 102) ", one male with an additional green label " DNA M. Balke 1382 " (ZSM). 1 male, 2 females " Papua New Guinea: Morobe, Wagau, Herzog Mts., 1150 m, 19. xi. 2006, 06.51.067 S 146.48.068 E, Balke & Kinibel (PNG 103) " (ZSM). 2 males " Papua New Guinea: Gulf [sic!], Menyamya, Mt Inji 1700 m, 14. xi. 2006 nr 07.14.813 S 146.01.330 E Balke & Kinibel (PNG 96) " (ZSM). 9 males " PAPUA N. G.: Morobe Prov. Aseki, Oiwa, 1600 - 1700 m, 11. - 12.3.1998 leg. A. Riedel " (NHMW). 6 males, 2 females " Papua New Guinea: Gulf [Morobe], Marawaka, Andakombe towards Morobe, 1100 m, 12. xi. 200, 07.09.766 S, 145.46.333 E, Balke & Kinibel (PNG 92) " (NHMW, ZSM). Eastern Highlands: 38 males, 38 females " Papua New Guinea: Eastern Highlands, Marawaka, Ande, 1700 m, 8. xi. 2005, 07.01.697 S 145.49.807 E, Balke & Kinibel (PNG 86) ", one male with an additional green label " DNA M. Balke 1365 " (NHMW, ZSM). 49 males, 40 females " Papua New Guinea: Eastern Highlands, Marawaka, Ande, 1700 - 1800 m, 9. xi. 2006, 07.01.697 S 145.49.807 E, Balke & Kinibel (PNG 87) " (NHMW, ZSM).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
757E69DBAB555E01AA5EDD3AA356177C.taxon	diagnosis	Diagnosis. For complete description, see Balke (1998: 331). Beetle medium-sized: TL-H 3.55 - 4.1 mm; oblong-oval; brown to dark brown, with reddish to reddish brown pronotal sides or pronotum and often head; shiny, with very fine, on elytra often almost invisible punctation and weakly impressed microreticulation; pronotum with lateral bead (Fig. 27); male protarsomere 4 with anterolateral seta thin, evenly curved, smaller than or equal to more laterally situated large seta; male protarsomere 5 ventrally with anterior band of more than 60 and posterior row of 13 - 15 relatively long setae (Fig. 31 D); median lobe in lateral view with apical lobes distinctly developed and rounded, " nose " small but distinct; paramere slightly concave on dorsal side and with long, dense, thin setae, situated along dorsal margin distinctly divided to dense and strong subdistal setae and sparser proximal ones, setae in middle short and fine (Fig. 31 A-C).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
757E69DBAB555E01AA5EDD3AA356177C.taxon	distribution	Distribution. Papua New Guinea: Eastern Highlands and Morobe Provinces (Fig. 34).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
208EA2242D04579F99A6A6530E9A407F.taxon	description	Description. Body size and form: Beetle small: TL-H 3.1 - 3.85 mm, TL 3.45 - 4.35 mm, MW 1.7 - 2.05 mm (holotype: TL-H 3.4 mm, TL 3.75 mm, MW 1.85 mm), with oblong-oval habitus. Colouration: Piceous, with paler sides of pronotum and head anteriorly. Head reddish brown to dark brown, paler anteriorly. Pronotum dark brown, to piceous on disc and to reddish on sides. Elytra uniformly dark brown to piceous. Head appendages and legs proximally yellowish to reddish, legs distally darker, reddish brown (Fig. 28). Teneral specimen paler, brown to reddish brown with to yellowish pronotum and head. Surface sculpture: Shiny dorsally, with extremely fine, sparse punctation and weakly impressed microreticulation. Head with fine and sparse punctation (spaces between punctures 2 - 3 times size of punctures); diameter of punctures smaller than diameter of cells of microreticulation. Pronotum with much finer and sparser punctation than on head, very inconspicuous. Punctation on elytra invisible. Pronotum and elytra with weakly impressed microreticulation; head with microreticulation slightly stronger. Metaventrite, metacoxae, and abdominal ventrites distinctly microreticulate. Metacoxal plates with longitudinal strioles and weak transverse wrinkles. Punctation on venter invisible; inconspicuous on two last abdominal ventrites. Structures: Pronotum with narrow lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 truncate. Male: Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior narrow band of 26 setae and posterior row of six relatively long setae (Fig. 32 D). Abdominal ventrites 1 - 3 with long strioles, abdominal ventrites 4 - 6 without strioles or with 1 - 2 small lateral strioles on each side. Median lobe with apical lobes weakly developed, not rounded, truncate in lateral view, " nose " usually indistinct (Fig. 32 A, B). Paramere as in Fig. 32 C. Female: Without evident differences in external morphology from males, except for not modified protarsi. Abdominal ventrites 1 - 2 with strioles, abdominal ventrites 3 - 6 without strioles.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
208EA2242D04579F99A6A6530E9A407F.taxon	distribution	Distribution. Papua New Guinea: Central Province (Fig. 34).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
208EA2242D04579F99A6A6530E9A407F.taxon	etymology	Etymology. The species is named after Kailaki Village. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
110D3B4B1C1C5EC99F6E781AEF4783D0.taxon	description	Description. Body size and form: Beetle medium-sized: TL-H 4.25 mm, TL 4.75 mm, MW 2.2 mm, with oblong-oval habitus. Colouration: Piceous, with paler sides of pronotum and head anteriorly. Head dark brown, paler anteriorly. Pronotum dark brown, with brown sides. Elytra piceous, with weakly indicated reddish sutural lines. Head appendages and legs proximally reddish, legs distally darker, reddish brown (Fig. 46). Surface sculpture: Shiny dorsally, with extremely fine and sparse punctation and weakly impressed microreticulation. Head with fine and sparse punctation (spaces between punctures 2 - 3 times size of punctures); diameter of punctures equal to or smaller than diameter of cells of microreticulation. Pronotum with much finer and sparser punctation than on head, very inconspicuous. Punctation on elytra invisible. Pronotum and elytra with weakly impressed microreticulation; head with microreticulation slightly stronger. Metaventrite, metacoxae, and abdominal ventrites distinctly microreticulate. Metacoxal plates with longitudinal strioles and weak transverse wrinkles; abdominal ventrites with strioles. Punctation on venter invisible; inconspicuous on two last abdominal ventrites. Structures: Pronotum with lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 truncate. Male: Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior band of 25 and posterior row of five relatively long setae (Fig. 50 D). Abdominal ventrite 6 with 3 - 4 lateral striae on each side. Median lobe short, robust, evenly tapering to broadly pointed apex in lateral and ventral views (Fig. 50 A, B). Paramere slightly concave on dorsal side, subdistal setae strong and dense, proximal setae inconspicuous (Fig. 50 C). Female: Unknown.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
110D3B4B1C1C5EC99F6E781AEF4783D0.taxon	distribution	Distribution. Papua New Guinea: Morobe Province (Fig. 54).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
110D3B4B1C1C5EC99F6E781AEF4783D0.taxon	etymology	Etymology. The species is named after Kobau Village. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
341A511B7DCC5FFE9F173732B4E29950.taxon	description	Description. Body size and form: Beetle medium-sized: TL-H 3.95 - 4.4 mm, TL 4.4 - 4.55 mm, MW 2.2 - 2.35 mm (holotype: TL-H 3.95 mm, TL 4.4 mm, MW 2.2 mm), with oblong-oval habitus. Colouration: Piceous, with paler sides of pronotum. Head piceous, paler anteriorly; pronotum piceous, with brown sides; elytra piceous, with reddish sutural lines; head appendages and legs proximally reddish, legs distally darker, reddish brown (Fig. 2). Teneral specimen paler, brown. Surface sculpture: Submatt dorsally, with relatively dense and coarse punctation and evident microreticulation. Head with relatively dense and coarse punctation (spaces between punctures 1 - 2 times size of punctures); diameter of punctures almost equal to diameter of cells of microreticulation. Pronotum with finer, sparser punctation, and more evenly distributed punctation than on head. Elytra with coarser punctation than on pronotum. Pronotum and elytra with distinct microreticulation. Head with microreticulation slightly stronger. Metaventrite, metacoxae, and abdominal ventrites distinctly microreticulate. Metacoxal plates with longitudinal strioles and weak transverse wrinkles; abdominal ventrites with strioles. Venter with inconspicuous punctation, more evident on metacoxal plates and two last abdominal ventrites. Structures: Pronotum with distinct lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded and with few transverse strioles anteriorly, without anterolateral extensions. Blade of prosternal process lanceolate, relatively broad, convex, with distinct bead and few setae laterally. Abdominal ventrite 6 slightly truncate. Male: Protarsomere 4 with rather small, slightly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of 23 short setae and posterior row of 6 setae (Fig. 8 D). Abdominal ventrite 6 with 12 - 15 lateral striae on each side. Median lobe slightly curved, with apex thick, short, pointed and strongly curved downwards in lateral view (Fig. 8 A-B). Paramere with distinct notch on dorsal side, subdistal part relatively large, rounded, with dense and strong setae, proximal setae thin and sparse, inconspicuous. (Fig. 8 C). Female: Unknown.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
341A511B7DCC5FFE9F173732B4E29950.taxon	distribution	Distribution. Papua New Guinea: Hela Province, Koroba area (Fig. 11).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
341A511B7DCC5FFE9F173732B4E29950.taxon	etymology	Etymology. The species is named after Koroba Village. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
92803D30F9395308812E99856A8F6091.taxon	materials_examined	Additional material. Madang: 7 males, 2 females " Papua New Guinea: Madang Province, Wanang village, ca 110 m, 20. ix. 2013, 05.15.458 S 145.02.389 E, David Boukal (PNG 2013 - 13) " (NHMW, ZSM). 4 males, 5 females " Papua New Guinea: Madang Province, Wanang village env., Wanang conservation area, 250 m, 21. ix. 2013, 05.15.458 S 145.02.389 E, David Boukal (PNG 2013 - 20) " (NHMW, ZSM). 1 male " Papua New Guinea: Madang Province, Wanang village env., Wanang conservation area, 250 m, 22. ix. 2013, 05.15.458 S 145.02.389 E, David Boukal (PNG 2013 - 24) " (NHMW). 1 female " Papua New Guinea: Madang Province, Wanang village env., Wanang conservation area, 250 m, 22. ix. 2013, 05.15.458 S 145.02.389 E, David Boukal (PNG 2013 - 31) " (NHMW). 16 males, 4 females " Papua New Guinea: Madang Province, Wanang village env., Wanang conservation area, 250 m, 23. ix. 2013, 05.15.458 S 145.02.389 E, David Boukal (PNG 2013 - 33) " (NHMW, ZSM). 8 males, 11 females " Papua New Guinea: Madang Province, Wanang village env., Wanang conservation area, 250 m, 23. ix. 2013, 05.15.458 S 145.02.389 E, David Boukal (PNG 2013 - 37) " (NHMW, ZSM). 2 males, 1 female " Papua New Guinea: Madang Province, Wanang village env., Wanang conservation area, 250 m, 24. ix. 2013, 05.15.458 S 145.02.389 E, David Boukal (PNG 2013 - 39) " (NHMW). 33 males, 3 females " Papua New Guinea: Madang Province, Wanang village env., Wanang conservation area, 250 m, 25. ix. 2013, 05.15.458 S 145.02.389 E, David Boukal (PNG 2013 - 44) " (NHMW, ZSM). 4 males, 1 female " V. Kolar Lgt. Papua New Guinea Wanang III 4 - 20.7.2013 " (NHMW). 1 female " Ibisca Niugini, PNG 18 - 20. xi. 2012 Wanang - 5,227670193 145,0797424 ", " FIT-WAN-G- 1 / 8 - d 01 / Plot 7 / P 0596 Vial 22305 - CODYTI " (ZSM). 1 male " Ibisca Niugini, PNG 20 - 22. xi. 2012 Wanang - 5,227670193 145,0797424 ", " FIT-WAN-P- 2 / 8 - d 04 / Plot 16 / P 0669 Vial 22273 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 24 - 26. xi. 2012 Wanang - 5,227670193 145,0797424 ", " FIT-WAN-H- 4 / 8 - d 07 / Plot 8 / P 0607 Vial 17640 - CODYTI " (ZSM). 1 male " Ibisca Niugini, PNG 30. x. - 2. xii. 2012 Wanang - 5,227670193 145,0797424 ", " FIT-WAN-L- 7 / 8 - d 14 / Plot 12 / P 0642 Vial 17744 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 2 - 4. xii. 2012 Wanang FIT-WAN-D- 8 / 8 - d 15 / Plot 4 / P 0579 Vial 17568 - CODYTI " (ZSM). 3 males " Papua New Guinea: Madang, below Bundi, 500 m, 26. IX. 2002 Balke & Sagata (PNG 023) ", one of them with an additional green label " 269 DNA M Balke " and " sp 24 SEM 19 " (ZSM). 5 males, 3 females " Papua New Guinea: Madang, Brahmin, 150 m, 26. IX. 2002 Balke & Sagata (PNG 024) ", two of males with additional green labels " 274 DNA M Balke ", " 275 DNA M Balke " (NHMW, ZSM). 1 male " Papua New Guinea: Madang, Adalbert [sic!] Mts., Keki, 850 m, 4. v. 2006, nr 04.42.300 S 145.25.089 E, Manaono leg. (PNG 52) " (ZSM). 2 males, 5 females " Papua New Guinea: Madang, Adalbert [sic!] Mts., creek nr Keki, 790 m, 28. xi. 1994, 04.42.300 S 145.25.089 E, Binatang Boys leg. (PNG 53 a) " (ZSM). 1 male, 1 female " Papua New Guinea: Madang, Adalbert Mts., Keki to Sewan, 650 m, 7. v. 1994, 04.41.802 S 145.25.460 E, Balke (PNG 54) ", male with an additional green label " DNA M. Balke 1299 " (ZSM). 16 males, 8 females " Papua New Guinea: Madang, Usino, 260 m, 15. iii. 2007, 05.31.125 S 145.25.316 E, Kinibel (PNG 158) ", one male with an additional green label " DNA M. Balke 3307 " (NHMW, ZSM). 1 male, 1 female " Papua New Guinea: Madang, Mt. Tapo, 180 m, ii. 2008 5 24.11.00 S 145 36 17 16 E, BRC leg. (PNG 178) " (ZSM). 4 males, 8 females " Papua New Guinea: Madang, Wannang, 270 m 31. x. 2008, 05.15.458 S 145.02.389 E, Posman, (PNG 187) " (ZSM). 17 males, 25 females " Papua New Guinea: Madang, Wannang, 230 m 3. x. 2008, 05.17.235 S 145.06.160 E, Posman (PNG 188) ", three males additionally with green labels " DNA M. Balke 3763 ", " DNA M. Balke 3767 ", " DNA M. Balke 3768 " (NHMW, ZSM). 2 males " Papua New Guinea: Madang, Akameku - Brahmin, Bismarck Range, 250 - 500 m, 25. xi. 2006, nr 05.47.026 S 145.24.131 E, Balke & Kinibel (PNG 115) ", one of them with an additional green abel " DNA M. Balke 1363 " (ZSM). 13 males, 22 females " Papua New Guinea: Madang, Keki, Adalbert Mts., 500 m, 29. xi. 1994, 04.43.058 S 145.24.437 E, Balke & Kinibel (PNG 118) " (NHMW, ZSM). 38 males, 56 females " Papua New Guinea: Madang, Keki, Adalbert Mts., 400 m, 29. xi. 1994, 04.43.058 S 145.24.437 E, Binatang Boys, (PNG 119) " (NHMW, ZSM). 60 males, 97 females " Papua New Guinea: Madang, Keki-Sewan, Adalbert Mts., 700 m, 30. xi. 1994 nr 04.41.802 S 145.25.460 E Binatang Boys (PNG 120) " (NHMW, ZSM). 7 males, 8 females " Papua New Guinea: Madang, Keki-Sewan, Adalbert Mts., 300 m 30. xi. 1994, 04.40.558 S 145.27.187 E, Binatang Boys, (PNG 121) " (ZSM). 1 female " Ibisca Niugini, PNG 25 - 27. x. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 ", " FIT-MW 700 - C- 1 / 8 - d 01 / Plot 3 / P 1114 Vial 16039 - CODYTI " (ZSM). 1 male " Ibisca Niugini, PNG 26 - 28. x. 2012 Mount Wilhelm 700 m FIT-MW 700 - O- 1 / 8 - d 02 / Plot 1 / P 1210 Vial 16172 - CODYTI " (ZSM). 4 females " Ibisca Niugini, PNG 26 - 28. x. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 MW 0700 / P 1210 Vial 16172 " (ZSM). 1 male, 1 female " Ibisca Niugini, PNG 26 - 28. x. 2012 Mount Wilhelm 700 m ", " - 5,731960773 145,2521667 FIT-MW 700 - S- 1 / 8 - d 02 / Plot 19 / P 1242 Vial 16118 - CODYTI " (ZSM). 2 males, 3 females " Ibisca Niugini, PNG 26 - 28. x. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 MW 700 / P 1234 Vial 16270 " (ZSM). 1 female " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 700 m ", " - 5,731960773 145,2521667 FIT-MW 700 - A- 2 / 8 - d 03 / Plot 1 / P 1099 Vial 15960 - CODYTI " (ZSM). 2 females " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 700 m ", " - 5,731960773 145,2521667 FIT-MW 700 - C- 2 / 8 - d 03 / Plot 3 / P 1115 Vial 15923 - CODYTI " (ZSM). 1 male, 2 females " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 ", " FIT-MW 700 - A- 2 / 8 - d 03 / Plot 1 / P 1099 Vial 15958 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 ", " FIT-MW 700 - G- 2 / 8 - d 03 / Plot 7 / P 1147 Vial 16042 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 ", " FIT-MW 700 - H- 2 / 8 - d 03 / Plot 8 / P 1155 Vial 15976 - CODYTI " (ZSM). 2 females " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 ", " FIT-MW 700 - E- 2 / 8 - d 03 / Plot 5 / P 1131 Vial 15937 - CODYTI " (ZSM). 2 females " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 MW 0700 / P 1243 Vial 16156 " (ZSM). 2 males, 5 females " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 MW 700 / P 1235 Vial 16164 " (ZSM). 3 males, 5 females " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 700 m FIT-MW 700 - O- 2 / 8 - d 0 / Plot 15 / P 1211 Vial 16189 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 30. x. - 1. xi. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 MW 0700 / P 1252 Vial 15993 " (ZSM). 1 female " Ibisca Niugini, PNG 1 - 3. xi. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 MW 0700 / P 1213 Vial 16236 " (ZSM). 1 female " Ibisca Niugini, PNG 1 - 3. xi. 2012 Mount Wilhelm 700 m FIT-MW 700 - M- 4 / 8 - d 08 / Plot 13 / P 1197 Vial 15762 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 2 - 4. xi. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 ", " FIT-MW 700 - A- 5 / 8 - d 09 / Plot 1 / P 1102 Vial 16027 - CODYTI " (ZSM). 2 females " Ibisca Niugini, PNG 2 - 4. xi. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 ", " FIT-MW 700 - E- 5 / 8 - d 09 / Plot 5 / P 1134 Vial P 1134 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 3 - 5. xi. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 MW 0700 / P 1222 Vial 16098 " (ZSM). 1 female " Ibisca Niugini, PNG 3 - 5. xi. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 MW 0700 / P 1254 Vial 16105 " (ZSM). 1 female " Ibisca Niugini, PNG 3 - 5. xi. 2012 Mount Wilhelm 700 m FIT-MW 700 - M- 5 / 8 - d 10 / Plot 13 / P 1198 Vial 16087 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 3 - 5. xi. 2012 Mount Wilhelm 700 m FIT-MW 700 - O- 5 / 8 - d 10 / Plot 15 / P 1214 Vial 16088 - CODYTI " (ZSM). 1 male, 2 females " Ibisca Niugini, PNG 3 - 5. xi. 2012 Mount Wilhelm 700 m ", " - 5,731960773 145,2521667 FIT-MW 700 - R- 5 / 8 - d 10 / Plot 18 / P 1238 Vial 15969 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 3 - 5. xi. 2012 Mount Wilhelm 700 m ", " - 5,731960773 145,2521667 FIT-MW 700 - K- 5 / 8 - d 10 / Plot 11 / P 1182 Vial 16083 - CODYTI " (ZSM). 4 females " Ibisca Niugini, PNG 4 - 6. xi. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 ", " FIT-MW 700 - E- 6 / 8 - d 11 / Plot 5 / P 1135 Vial 07232 - CODYTI " (ZSM). 2 males, 1 female " Ibisca Niugini, PNG 4 - 6. xi. 2012 Mount Wilhelm 700 m ", " - 5,731960773 145,2521667 FIT-MW 700 - A- 6 / 8 - d 11 / Plot 1 / P 1103 Vial 07195 - CODYTI " (ZSM). 4 females " Ibisca Niugini, PNG 4 - 6. xi. 2012 Mount Wilhelm 700 m ", " - 5,731960773 145,2521667 FIT-MW 700 - C- 6 / 8 - d 11 / Plot 3 / P 1119 Vial 07081 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 4 - 6. xi. 2012 Mount Wilhelm 700 m FIT-MW 700 - D- 6 / 8 - d 11 / Plot 4 / P 1127 Vial 07061 " (ZSM). 1 male, 1 female " Ibisca Niugini, PNG 5 - 7. xi. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 MW 0700 / P 1239 Vial 16263 " (ZSM). 1 male, 1 female " Ibisca Niugini, PNG 6 - 8. xi. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 ", " FIT-MW 700 - E- 7 / 8 - d 13 / Plot 5 / P 1136 Vial 15886 - CODYTI " (ZSM). 2 females " Ibisca Niugini, PNG 9 - 11. xi. 2012 Mount Wilhelm 700 m - 5,731960773 145,2521667 MW 0700 / P 1217 Vial 16106 " (ZSM). 1 male, 3 females " Ibisca Niugini, PNG 3 - 5. xi. 2012 Mount Wilhelm 200 m - 5,739897251 145,329742 MW 0200 / P 0856 Vial 09605 " (ZSM). 1 female " Ibisca Niugini, PNG 3 - 5. xi. 2012 Mount Wilhelm 200 m - 5,739897251 145,329742 MW 0200 / P 0865 Vial 09576 " (ZSM). Eastern Highlands: 6 males, 7 females " Papua New Guinea: Eastern Highlands, Bena Bridge, 1400 m, 8. xii. 2007, 06.10.781 S 145.26.034 E, Balke & Sagata (PNG 164) " (NHMW, ZSM).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
92803D30F9395308812E99856A8F6091.taxon	diagnosis	Diagnosis. For complete description, see Balke (1998: 332). Beetle small or medium-sized: TL-H 3.45 - 3.9 mm; oblong-oval, sometimes slightly more attenuated posteriorly; with characteristic dorsal colouration: dark (brown to piceous) elytra and pale (reddish to reddish brown pronotum, except for darker disc, and head; shiny, with very fine, on elytra often almost invisible punctation and weakly impressed microreticulation; pronotum with very narrow lateral bead (Fig. 35); male protarsomere 4 with medium-sized, thick, strongly curved anterolateral hook-like seta; male protarsomere 5 with anterior band of ca. 30 and posterior row of nine relatively long setae (Fig. 40 D); median lobe robust, without lateral setae apically; in lateral view, with broadened, triangle apex; in ventral view, apex deeply concave; paramere very slightly concave on dorsal side and with long and dense subdistal and inconspicuous proximal setae (Fig. 40 A-C).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
92803D30F9395308812E99856A8F6091.taxon	distribution	Distribution. Papua New Guinea: Madang and Eastern Highlands Provinces. The species is known from numerous specimens from the central and wertern part of Madang and from northern part of Eastern Highlands (Fig. 45).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
A9608F8EFD8A5A1D8717FBE6E87EDF24.taxon	description	Description. Body size and form: Beetle medium-sized: TL-H 3.85 - 4.4 mm, TL 3.45 - 3.95 mm, MW 1.8 - 2.05 mm (holotype: TL-H 3.8 mm, TL 4.2 mm, MW 2.0 mm), with oblong-oval habitus. Colouration: Dark brown, with paler sides of pronotum and head anteriorly. Head dark brown, piceous posteriorly; pronotum dark brown, with brown sides; elytra uniformly dark brown; head appendages and legs proximally reddish, legs distally darker, reddish brown (Fig. 3). Teneral specimen paler, brown to reddish brown with yellowish pronotal sides. Surface sculpture: Shiny dorsally, with fine, sparse punctation and weakly impressed microreticulation. Head with relatively fine and sparse punctation (spaces between punctures 2 - 3 times size of punctures); diameter of punctures almost equal to or smaller than diameter of cells of microreticulation. Pronotum and elytra with much finer and sparser punctation than on head, inconspicuous. Pronotum and elytra with weakly impressed microreticulation. Head with microreticulation slightly stronger. Metaventrite, metacoxae, and abdominal ventrites distinctly microreticulate. Metacoxal plates with longitudinal strioles and weak transverse wrinkles; abdominal ventrites with strioles. Venter with extremely inconspicuous punctation, more evident on metacoxal plates and two last abdominal ventrites. Structures: Pronotum without lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 slightly truncate. Male: Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior band of ca. 60 and posterior row of ten relatively long setae (Fig. 9 D). Abdominal ventrite 6 with 6 - 10 lateral striae on each side. Median lobe simple, slightly curved, in lateral view, apex thick, short and slightly curved downwards, its minuscule tip curved upwards (Fig. 9 A, B). Paramere without notch on dorsal side, evenly tapering to distal part, with numerous small spines and without long setae (Fig. 9 C). Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
A9608F8EFD8A5A1D8717FBE6E87EDF24.taxon	distribution	Distribution. Papua New Guinea: Sandaun Province (Fig. 11).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
A9608F8EFD8A5A1D8717FBE6E87EDF24.taxon	etymology	Etymology. The species is named after Mekil Village where most specimens of the species were found. The name is an adjective in the nominative singular.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
D5CB43EB7D465FAB8B2DE29C5ED73336.taxon	description	Description. Body size and form: Beetle medium-sized: TL-H 3.75 - 4.25 mm, TL 4.15 - 4.6 mm, MW 1.95 - 2.3 mm (holotype: TL-H 3.75 mm, TL 4.15 mm, MW 1.95 mm), with oblong habitus. Colouration: Piceuos. Head piceous, with reddish brown anterior margin. Pronotum dark brown to piceous, with reddish brown to brown sides. Elytra uniformly piceous. Head appendages and legs proximally yellowish, legs distally darker, reddish brown (Fig. 38). Teneral specimen paler. Surface sculpture: Submatt dorsally, with dense and coarse punctation and weakly impressed microreticulation. Head with very dense and coarse punctation (no spaces between punctures or spaces 1 - 2 times size of punctures); diameter of punctures equal to diameter of cells of microreticulation. Pronotum with distinct punctation, finer than on head. Punctation on elytra distinct, finer and sparser than on head. Elytra with weakly impressed microreticulation; head and pronotum with microreticulation stronger than on elytra. Metaventrite, metacoxae, and abdominal ventrites distinctly microreticulate. Metacoxal plates with longitudinal strioles and weak transverse wrinkles; abdominal ventrites 2 - 4 with few strioles, two last one without strioles but with very weak wrinkles. Punctation on venter fine but distinct. Structures: Pronotum with narrow lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, narrow, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 truncate or very slightly concave. Male: Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior band of ca. 60 and posterior row of 17 relatively long setae (Fig. 43 D). Abdominal ventrite 6 without lateral striae on each side, except one with setae. Median lobe slender, lateral sides slightly thickened; in lateral view, apex short, pointed, and curved downwards; in ventral view, lateral sides evenly expanded subdistally and apex slightly concave (Fig. 43 A, B). Paramere with distinct dorsal notch and subdistal part well developed, with long and dense subdistal and inconspicuous proximal setae (Fig. 43 C). Female: Without evident differences in external morphology from males, except for not modified protarsi.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
D5CB43EB7D465FAB8B2DE29C5ED73336.taxon	distribution	Distribution. Papua New Guinea: Sandaun Province (Fig. 45).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
D5CB43EB7D465FAB8B2DE29C5ED73336.taxon	etymology	Etymology. The species is named after Mianmin Village. The name is an adjective in the nominative singular.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
B4CF65541BFD5C3A9470C9317EA6D974.taxon	description	Description. Body size and form: Beetle medium-sized, rarely small: TL-H 3.3 - 4.1 mm, TL 3.6 - 4.5 mm, MW 1.75 - 2.1 mm (holotype: TL-H 3.9 mm, TL 4.35 mm, MW 2.05 mm), with oblong-oval habitus. Colouration: Brown to dark brown, usually with reddish pronotum and head. Head reddish to brown, sometimes darker posterior eyes; pronotum reddish to brown, often broader or narrower darker area on disc; elytra brown to dark brown, sometimes with reddish sutural lines; head appendages and legs proximally reddish, legs distally darker, reddish brown to brown (Fig. 4). Teneral specimen paler. Surface sculpture: Shiny dorsally, with fine, sparse punctation and weakly impressed microreticulation. Head with relatively fine and sparse punctation (spaces between punctures 2 - 3 times size of punctures); diameter of punctures almost equal to or smaller than diameter of cells of microreticulation. Pronotum and elytra with much finer and sparser punctation than on head, often inconspicuous on elytra. Pronotum and elytra with weakly impressed microreticulation. Head with microreticulation slightly stronger. Metaventrite, metacoxae, and abdominal ventrites distinctly microreticulate. Metacoxal plates with longitudinal strioles and weak transverse wrinkles; abdominal ventrites with strioles. Venter with extremely inconspicuous punctation, more evident on metacoxal plates and two last abdominal ventrites. Structures: Pronotum with narrow lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 very slightly truncate. Male: Protarsomere 4 with with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior band of ca. 50 and posterior row of seven relatively long setae (Fig. 10 D). Abdominal ventrite 6 with 5 - 7 lateral striae on each side. Median lobe evenly curved, rather thin, lateral margins thickened proximally; in lateral view, apex elongate, slightly thickened and rounded; in ventral view, median lobe broad proximally and distinctly narrowed in distal half, apex bluntly pointed (Fig. 10 A, B). Paramere slightly concave on dorsal side, its subdistal part with dense, strong setae, proximal setae weaker, less distinct (Fig. 10 C). Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
B4CF65541BFD5C3A9470C9317EA6D974.taxon	distribution	Distribution. Papua New Guinea: Morobe Province (Fig. 11).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
B4CF65541BFD5C3A9470C9317EA6D974.taxon	etymology	Etymology. The species is named after Morobe Province, the only province of PNG where the species has been found. The name is an adjective in the nominative singular.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
AC52001898CC5FFBA6B8FEAC44FACAC6.taxon	materials_examined	Additional material. Central: 23 males, 32 females " Papua New Guinea: Central, Kokoda Trek, 320 m, i. 2008 [09 °] 19.236 S 147.31.791 E, Posman (PNG 168) ", one male with an additional green label " DNA M. Balke 3405 " (NHMW, ZSM). 4 males " Papua New Guinea: Central, Kokoda Trek, 980 m, i. 2008, [09 °] 15.933 S 147.36.590 E, Posman (PNG 169) ", one of them with an additional green label " DNA M. Balke 3411 " (ZSM). 1 female " Papua New Guinea: Central, Kokoda Trek, 980 m, i. 2008, [09 °] 15.933 S 147.36.590 E, Posman (PNG 169) ", " DNA M. Balke 4117 " [green] (ZSM). 10 males, 2 females " Papua New Guinea: Central, Kokoda Trek, 590 m, i. 2008, [09 °] 14.339 S 147.36.920 E, Posman (PNG 170) " (NHMW, ZSM). 19 males, 31 females " Papua New Guinea: Central, Tapini, 1200 m, 31. x. 2007, 08.21.557 S 146.58.712 E, Kinibel (PNG 162) ", one of the males with an additional green label " DNA M. Balke 3306 " (NHMW, ZSM). 2 males, 3 females " Papua New Guinea: Central, Moreguina [10 ° ' 57 " S, 148 ° 28 ' 27 " E], 16. viii. 2008, Posman (PNG 183) ", males and one female with additional green labels " DNA M. Balke 3745 ", " DNA M. Balke 3816 ", " DNA M. Balke 3746 " respectively (ZSM). 5 males, 9 females " Papua New Guinea: Central, Moroka area, Kailaki, 827 m, 26. x. 2009, 9.24.134 S 147.33.521 E, Sagata (PNG 225) " (NHMW, ZSM). 14 males, 8 females " Papua New Guinea: Central, Moroka, Kailaki, 827 m, 26. x. 2009, 9.24.113 S 147.33.524 E, Sagata (PNG 226) " (NHMW, ZSM). 1 female " Papua New Guinea Central, Moroka, Kailaki Wareaga, 760 m, 27 x 2009 9.25.424 S 147.31.068 E Sagata (PNG 227) " (ZSM). 27 males, 39 females " Papua New Guinea: Central, 755 m, 28. x. 2009 S 9 25 47 5 E 147 32 59.1, Sagata (PNG 229) " (NHMW, ZSM). National Capital District: 2 males, 3 females " Papua New Guinea: National Capital District, Varirata NP, 600 m, 16. xii. 2007, 09.26.13 S 147.22.09 E, Balke & Sagata (PNG 159) ", males with additional green labels " DNA M. Balke 3304 " and " DNA M. Balke 3305 " (ZSM).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
AC52001898CC5FFBA6B8FEAC44FACAC6.taxon	distribution	Distribution. Papua New Guinea: Central and National Capital District Provinces (Fig. 45). The species is known from numerous specimens in the Central Province and a small population in the National Capital District.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
EEE0CC5E6EB0541ABDE7DE47B43DD574.taxon	description	Description. Body size and form: Beetle small: TL-H 3.3 - 3.5 mm, TL 3.7 - 3.95 mm, MW 1.85 - 1.95 mm (holotype: TL-H 3.5 mm, TL 3.95 mm, MW 1.95 mm), with oblong-oval habitus. Colouration: Fast uniformly reddish brown. Head reddish brown, darker posterior eyes. Pronotum reddish brown, slightly darker on disc. Elytra reddish brown, sometimes slightly darker than pronotum. Head appendages and legs proximally yellowish, legs distally darker, reddish (Fig. 12). Surface sculpture: Submatt dorsally, with strong and dense punctation and strongly impressed microreticulation. Head with dense and coarse punctation (spaces between punctures 0 - 1 times size of punctures); diameter of punctures equal to or larger than diameter of cells of microreticulation. Pronotum and elytra with finer and sparser punctation than on head, very distinct, more even on elytra. Pronotum and elytra with strongly impressed microreticulation; head with microreticulation stronger. Metaventrite, metacoxae, and abdominal ventrites distinctly microreticulate, but shiny. Metacoxal plates with longitudinal strioles and weak transverse wrinkles; abdominal ventrites with strioles. Punctation on venter weak; more distinct on two last abdominal ventrites. Structures: Pronotum with lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 broadly rounded or slightly truncate. Male: Protarsomere 4 with anterolateral seta rather long and thin, evenly curved, equal to laterally situated large seta. Protarsomere 5 ventrally with anterior band of ca. 60 and posterior row of five relatively long setae (Fig. 18 D). Abdominal ventrite 6 with 8 - 9 lateral striae on each side. Median lobe short, evenly tapering to apex in lateral and ventral views, very tip of apex thickened dorsally (Fig. 18 A, B). Paramere as in Fig. 18 D. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
EEE0CC5E6EB0541ABDE7DE47B43DD574.taxon	distribution	Distribution. Papua New Guinea: Morobe Province (Fig. 25).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
EEE0CC5E6EB0541ABDE7DE47B43DD574.taxon	etymology	Etymology. The species is named after Oiwa Village. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
6731F430FEC65C59880946C59359CEE5.taxon	description	Description. Body size and form: Beetle small: TL-H 3.05 - 3.35 mm, TL 3.5 - 3.7 mm, MW 1.7 - 1.85 mm (holotype: TL-H 3.35 mm, TL 3.7 mm, MW 1.85 mm), usually with oval, egg-shaped habitus. Colouration: Reddish brown to brown. Head reddish brown to dark brown, paler anteriorly. Pronotum dark brown on disc and narrower or broader reddish on sides. Elytra reddish brown to dark brown, with reddish sutural lines. Head appendages and legs proximally yellowish, legs distally darker, reddish brown (Fig. 15). Surface sculpture: Shiny dorsally, with fine punctation and weakly impressed microreticulation. Head with fine and sparse punctation (spaces between punctures 2 - 3 times size of punctures); diameter of punctures equal to or smaller than diameter of cells of microreticulation. Pronotum and elytra with much finer and sparser punctation than on head, sometimes inconspicuous. Pronotum and elytra with weakly impressed microreticulation; head with microreticulation slightly stronger. Metaventrite, metacoxae, and abdominal ventrites distinctly microreticulate. Metacoxal plates with longitudinal strioles and weak transverse wrinkles; abdominal ventrites with strioles. Punctation on venter invisible; inconspicuous on two last abdominal ventrites. Structures: Pronotum with narrow lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 slightly truncate. Male: Protarsomere 4 with anterolateral seta rather long and thing, evenly curved, smaller than more laterally situated large seta. Protarsomere 5 ventrally with anterior band of more than 60 and posterior row of 4 relatively long setae (Fig. 21 D). Abdominal ventrite 6 with 3 - 6 lateral striae on each side. Median lobe simple, evenly tapering to broadly pointed apex in lateral and ventral views (Fig. 21 A, B). Paramere as in Fig. 21 C. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
6731F430FEC65C59880946C59359CEE5.taxon	distribution	Distribution. Indonesia: Papua Province, Pegunungan Bintang Regency, Ok Sibil area (Fig. 25).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
6731F430FEC65C59880946C59359CEE5.taxon	etymology	Etymology. The species is named after Ok Sibil River. The name is an adjective in the nominative singular.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
E9FA166BF0D25934B1D36D25D0D8C619.taxon	description	Description. Body size and form: Beetle medium-sized: TL-H 3.4 - 3.85 mm, TL 3.8 - 4.25 mm, MW 1.8 - 2.1 mm (holotype: TL-H 3.65 mm, TL 4.0 mm, MW 1.95 mm), with oblong-oval habitus. Colouration: Brown to dark brown, with paler sides of pronotum and head. Head reddish brown, dark brown posterior to eyes. Pronotum reddish brown to brown, with reddish sides. Elytra brown to dark brown, sometimes with weak reddish sutural lines. Head appendages and legs proximally yellowish to reddish, legs distally darker, reddish brown (Fig. 29). Surface sculpture: More or less shiny dorsally, with fine but distinct punctation and distinctly impressed microreticulation. Head with coarse and dense punctation (no spaces between punctures or spaces 1 - 2 times size of punctures); diameter of punctures equal to diameter of cells of microreticulation. Pronotum with much finer and sparser punctation than on head. Elytra with distinct punctation, slightly finer and sparser than on pronotum. Pronotum and elytra with weakly or more strongly impressed microreticulation; head with microreticulation stronger. Metaventrite, metacoxae, and abdominal ventrites distinctly microreticulate. Metacoxal plates with longitudinal strioles and weak transverse wrinkles. Abdominal ventrites with strioles and very fine, sparse punctation. Structures: Pronotum with lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 slightly truncate. Male: Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior band of more than 60 and posterior band of ca. 30 relatively long setae, which connected approximately in middle (Fig. 33 D). Abdominal ventrite 6 with 1 - 4 lateral strioles on each side. Median lobe with apical lobes very strongly developed, rounded in lateral view, " nose " small but distinct (Fig. 33 A, B). Paramere as in Fig. 33 C. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrites 5 and 6 without strioles.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
E9FA166BF0D25934B1D36D25D0D8C619.taxon	distribution	Distribution. Papua New Guinea: Central Province (Fig. 34).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
E9FA166BF0D25934B1D36D25D0D8C619.taxon	etymology	Etymology. The species was mistaken for E. jaseminae due to their similarity in general appearance and shape of the median lobe. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
FA0E68D4F22A587F9675ABD615E5B6FB.taxon	description	Description. Body size and form: Beetle small: TL-H 2.85 - 3.3 mm, TL 3.15 - 3.7 mm, MW 1.6 - 1.8 mm (holotype: TL-H 3.05 mm, TL 3.4 mm, MW 1.75 mm), with oblong-oval habitus. Colouration: Reddish head and bicoloured elytra: yellowish at shoulders and brownish distally. Head reddish, reddish brown posterior eyes. Pronotum reddish brown to brown on disc (broader or narrower) and yellowish to yellowish reddish on sides. Elytra bicoloured: yellowish in proximal 1 / 4 to 1 / 3 (rarely to 1 / 2) and yellowish brown to brown distally, proximal yellowish colouration sometimes more distinctly boarded as shoulder spots slightly elongated along sutural lines, but mostly fuzzy, not boarded. Head appendages and legs proximally yellowish, legs distally darker, reddish to reddish brown (Fig. 47). Teneral specimens paler. Surface sculpture: Shiny dorsally, with extremely fine and sparse punctation and weakly impressed microreticulation. Head with fine and sparse punctation (spaces between punctures 2 - 3 times size of punctures); diameter of punctures smaller than diameter of cells of microreticulation. Pronotum with much finer and sparser punctation than on head, very inconspicuous. Punctation on elytra invisible. Pronotum and elytra with weakly impressed microreticulation; head with microreticulation slightly stronger. Metaventrite, metacoxae, and abdominal ventrites distinctly microreticulate. Metacoxal plates with longitudinal strioles and weak transverse wrinkles; abdominal ventrites with strioles. Punctation on venter invisible; inconspicuous on two last abdominal ventrites. Structures: Pronotum with lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 broadly rounded or slightly truncate. Male: Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior band ca. 60 and posterior row of eight relatively long setae (Fig. 51 D). Abdominal ventrite 6 with 4 - 7 lateral striae on each side. Median lobe simple, short, slightly curved, with broadly pointed apex in lateral view, and evenly tapering to broadly pointed apex in ventral view (Fig. 51 A, B). Paramere slightly concave on dorsal side, with strong, long, dense subdistal setae, proximal setae inconspicuous (Fig. 51 C). Female: Without evident differences in external morphology from males, except for not modified pro- and mesotarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
FA0E68D4F22A587F9675ABD615E5B6FB.taxon	distribution	Distribution. Papua New Guinea: Central Province (Fig. 54).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
FA0E68D4F22A587F9675ABD615E5B6FB.taxon	etymology	Etymology. The species name derives from Latin pulchellus, a diminutive of pulcher (beautiful), to express the small size and nice colouration of the beetles; this species is the most colourful of all known New Guinea Exocelina. The species name is an adjective in the nominative singular.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
733BAC0C78D75CBFBC8D78BA8FBAC482.taxon	materials_examined	Additional material. 2 females " IRIAN JAYA: 11.9.1993 Bime - Calab Gebiet, Bime, 1400 m ", " leg. M. Balke (12) ca. 140 ° 12 ' E 04 ° 20 ' S " (NHMW).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
733BAC0C78D75CBFBC8D78BA8FBAC482.taxon	diagnosis	Diagnosis. For complete description, see Balke (1998: 336). Beetle medium-sized: TL-H 4.1 - 4.5 mm; oblong-oval; dark brown to piceous, with brownish pronotal sides and head anteriorly; shiny, but with distinct dorsal punctation and weakly impressed microreticulation; pronotum with narrow lateral bead (Fig. 39); male protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta; male protarsomere 5 with anterior band of ca. 40 and posterior row of eight relatively long setae (Fig. 44 D); median lobe robust, in lateral view, evenly curved to narrowly rounded, not curved downwards apex; in ventral view, with more strongly thickened lateral sides, distinctly expanded in middle and narrowing to broadly pointed apex; paramere with distinct dorsal notch and subdistal part well developed, with long and dense subdistal and inconspicuous proximal setae (Fig. 44 A-C).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
733BAC0C78D75CBFBC8D78BA8FBAC482.taxon	distribution	Distribution. Indonesia: Papua Province: Pegunungan Bintang Regency. The species is known only from the type material, i. e., Borme - Bime - Emdoman area (Fig. 45).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
1F73681E510A532A98880E378BB11F16.taxon	description	Description. Body size and form: Beetle small to medium-sized: TL-H 3.4 - 3.9 mm, TL 3.7 - 4.3 mm, MW 1.8 - 2.2 mm (holotype: TL-H 3.9 mm, TL 4.3 mm, MW 2.1 mm), with oblong-oval habitus. Colouration: Piceous, with reddish sides of pronotum and head. Head reddish to reddish brown, darker posterior to eyes. Pronotum brown to piceous on disc and reddish to reddish brown on sides. Elytra dark brown to piceous. Head appendages and legs proximally yellowish, legs distally darker, reddish brown (Fig. 37). Teneral specimen paler. Surface sculpture: Shiny dorsally, with inconspicuous punctation and weakly impressed microreticulation. As in E. nomax. Structures: Pronotum with narrow lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 truncate or slightly concave. Male: Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of 20 and posterior row of seven relatively long setae (Fig. 42 D). Abdominal ventrite 6 with 9 - 12 lateral striae on each side. Median lobe slender, with lateral setae apically; in lateral view, evenly curved to broadly pointed, short apex; in ventral view, apex slightly concave (Fig. 42 A, B). Paramere very slightly concave on dorsal side, with long and dense subdistal and inconspicuous proximal setae (Fig. 42 C). Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
1F73681E510A532A98880E378BB11F16.taxon	distribution	Distribution. Papua New Guinea: Simbu and Eastern Highlands Provinces (Fig. 45). The species is known only from Crater Mountain area.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
1F73681E510A532A98880E378BB11F16.taxon	etymology	Etymology. The species is named after Wara Hulene Village where one of the paratype was collected. The name is an adjective in the nominative singular.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
FE62B5E8DBFA5779BE0D26BE7C8E920A.taxon	description	Description. Body size and form: Beetle small: TL-H 3.15 - 3.8 mm, TL 3.5 - 4.15 mm, MW 1.65 - 2.05 mm (holotype: TL-H 3.4 mm, TL 3.8 mm, MW 1.8 mm), with oblong-oval habitus. Colouration: Dark brown, with paler sides of pronotum and head anteriorly. Head dark brown, paler anteriorly. Pronotum dark brown, with brown sides. Elytra uniformly dark brown. Head appendages and legs proximally reddish, legs distally darker, reddish brown (Fig. 48). Teneral specimen paler, brown to reddish brown with yellowish pronotal sides and head anteriorly. Surface sculpture: Shiny dorsally, with extremely fine and sparse punctation and weakly impressed microreticulation. Head with fine and sparse punctation (spaces between punctures 2 - 3 times size of punctures); diameter of punctures smaller than diameter of cells of microreticulation. Pronotum with much finer and sparser punctation than on head, very inconspicuous. Punctation on elytra invisible. Pronotum and elytra with weakly impressed microreticulation; head with microreticulation slightly stronger. Metaventrite, metacoxae, and abdominal ventrites distinctly microreticulate. Metacoxal plates with longitudinal strioles and weak transverse wrinkles; abdominal ventrites with strioles. Punctation on venter invisible; inconspicuous on two last abdominal ventrites. Structures: Pronotum with lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 truncate or very slightly concave. Male: Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior band of ca. 50 and posterior row of eight relatively long setae (Fig. 52 D). Abdominal ventrite 6 with 2 - 4 lateral striae on each side. Median lobe short, curved, with broadly pointed apex in lateral view, and evenly tapering to pointed apex in ventral view; its right lateral margin slightly concave at apex (Fig. 52 A, B). Paramere slightly concave on dorsal side, its subdistal part with numerous, dense, very strong setae, proximal setae long, but weaker, less distinct (Fig. 52 C). Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
FE62B5E8DBFA5779BE0D26BE7C8E920A.taxon	distribution	Distribution. Papua New Guinea: Simbu and Eastern Highlands Provinces, Crater Mountain area (Fig. 54).	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
FE62B5E8DBFA5779BE0D26BE7C8E920A.taxon	etymology	Etymology. The species is named after Haia Village. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo, Balke, Michael (2019): Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 878: 73-143, DOI: http://dx.doi.org/10.3897/zookeys.878.37403, URL: http://dx.doi.org/10.3897/zookeys.878.37403
