identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
6D48946A8FA5C527097A26353C04EFC8.text	6D48946A8FA5C527097A26353C04EFC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glyphidrilus Horst 1889	<div><p>Genus Glyphidrilus Horst, 1889</p><p>Diagnosis.</p><p>Prostomium zygolobous. Body shape nearly circular in cross section in anterior part, and becoming quadrangular in posterior part or after clitellum. Anus dorsal or dorso-terminal. A longitudinal lamellar ridge at maturity from body wall on each side in bc, through several of the clitellar segments, which are called wings. Dorsal pores absent. Setae four pairs per segment. Clitellum annular. Genital apertures, all minute and superficial. Male pores inter-or intraclitellar. Spermathecal pores usually all behind the testis segments. Gizzard in VII or VIII sometimes extending into an adjacent segment. Calciferous glands absent. Seminal vesicles usually short, usually four pairs in IX–XII . Holonephridia. Nephrostomes single ducts avesiculate and without sphincters or caeca. Testis and funnels free in X and XI. Male ducts intramural. Ovaries fan shaped and with several egg strings. Ovisacs present or absent. Prostate glands absent and spermathecae without diverticulum (Chanabun et al. 2013, Gates 1972).</p><p>Type species.</p><p>Glyphidrilus weberi Horst, 1889. Type species by original designation in Horst (1889: 77).</p></div>	https://treatment.plazi.org/id/6D48946A8FA5C527097A26353C04EFC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Chanabun, Ratmanee;Inkavilay, Khamla;Panha, Somsak	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
830715AC146E9FADBBD0CED922F826CD.text	830715AC146E9FADBBD0CED922F826CD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glyphidrilus nanensis Chanabun & Panha	<div><p>Glyphidrilus nanensis Chanabun &amp; Panha sp. n. Figs 1, 3, 4, Table 1</p><p>Type material.</p><p>Holotype: One adult (CUMZ 3403) in a rice field near Nan River at Saklek, Phichit, north Thailand (16°30'28.4"N, 100°31'15.0"E), 49 meters elevation on 16 June 2012. Paratypes: 42 adults and 10 juveniles (CUMZ 3404), 2 adults (ZMH 14579), 2 adults (NHMUK), and 2 adults (ZRC), all same collection data as holotype.</p><p>Diagnosis.</p><p>A small sized earthworm with the clitellar wings on the lateral side of the body in XXIV, XXV, XXVI– XXVII, XXVIII, XXIX. Clitellum in XVII, XVIII–XXXIII, XXXIV. Female pores, male pores, and spermathecal pores not visible. Genital markings: paired or asymmetrical on aa in X, XI, XII, XIII, XIV and XXVIII, XXIX, XXX; paired or asymmetrical on bc in XV, XVI, XVII–XXIII, XXIV, XXV. Four pairs of seminal vesicles in IX–XII . Intestinal origin in XV. Ovaries in XIII–XIV . Spermathecae between 13/14-17/18.</p><p>Description of holotype.</p><p>Dimensions: body length 78 mm, diameter 2.6 mm in segment VIII, 4.7 mm before the clitellar wing in segment XXIII, 4.2 mm after wing in segment XXX in clitellar region; body cylindrical in anterior part, quadrangular in transverse section behind clitellum. 241 segments. Body color pale brown with variations from red to pink at adjacent tissues of wing portion in different individuals of newly collected specimens. At posterior end dorsal surface considerably broader than the ventral. Clitellar wing on ventro-lateral part of clitellum in XXVI–XXIX, 3.6 mm in height, and 0.3 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in XVII–XXXIII . Four pairs of setae per segment from II, setal formula aa:ab:bc:cd:dd = 1.0:0.6:1.3:0.6:1.4 in segment VIII and 1.0:0.6:1.5:0.6:1.5 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: paired on aa in XII–XIII, laterally paired or asymmetrical on bc in XVII–XXIV .</p><p>Septa 5/6-8/9 thicker than between other segments, 9/10-14/15 thick and 15/16 to the last segment thin. Gizzard small, globular in 7/8. Intestine enlarged from XV. Hearts, five pairs in VII–XI, first in VII and last in XI. A pair of holonephridia in each segment from segment XIII onwards. Seminal vesicles, four pairs in IX, X, XI, XII. Ovaries, two pairs in XIII–XIV . Testes free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, elongated oval or globular between 13/14-17/18, about 0.2-0.3 mm in diameter, two to ten on each side per segment.</p><p>Variations.</p><p>Body lengths of adult (n = 49) and juvenile paratypes (n = 10). Adults ranged from 72-99 mm (85.2 ± 19.0), with 186-257 segments. Juveniles ranged from 73-93 mm (82.3 ± 7.5), with 186-251 segments. Wings in XXIV, XXV, XXVI–XXVII, XXVIII, XXIX, clitellum in XVII, XVIII -XXXIII, XXXIV. Genital markings: paired or asymmetrical on aa in X, XI, XII, XIII, XIV and XXVIII, XXIX, XXX; paired or asymmetrical on bc in XV, XVI, XVII–XXIII, XXIV, XXV.</p><p>Distribution.</p><p>The new species was found in the river banks of Nan River at Saklek, Phichit, north Thailand, which was covered with worm casts.</p><p>Etymology.</p><p>The species was named after the locality on the banks of the Nan River. This was the first time that the worm genus was ever recorded from near this river.</p><p>Remarks.</p><p>For a summary of the following comparisons please see Table 1. The new species differs from G. yunnanensis Chen &amp; Xu, 1977 reported from China by the latter having longer wings in XXII–XXXII, the clitellum in XVIII–XXXVIII, paired genital markings on bc in XVII–XXI and XXXII–XXXIV, lacking a spermathecae. It differs from G. vangviengensis Chanabun &amp; Panha, 2011 from Laos by G. vangviengensis having longer wings in XXIV, XXV–XXXI, XXXII, the clitellum in XIX, XX–XXXV, XXXVI, XXXVII, and lacking spermathecae. It differs from the species recorded as G. mekongensis Panha &amp; Chanabun, 2012 from the Mekong River, Ubon Ratchathani, northeast Thailand by the latter species having longer wings in XXIV–½ XXXIII, XXXIII, XXXIV, ½ XXXV, the clitellum in XIX–XXXVII, XXXVIII, and lacking spermathecae. It differs from G. borealis Chanabun &amp; Panha, 2013 from Chiangmai, north Thailand by the latter having longer wings in XXI, XXII–XXVII, XXVIII, XXIX, a longer clitellum in XIV, XVI, XVII–XXXI, XXXII, XXXIII, XXXIV, XXXV, XXXVI, and the intestine enlarged from XIII. It differs from G. vangthongensis Chanabun &amp; Panha, 2013 reported from Phitsanulok, north Thailand by G. vangthongensis having longer wings in XXIV, XXV, XXVI–XXXI, XXXII, a longer clitellum in XII, XIII, XIV, XV, XVI–XL, XLI, XLII, an intestine enlarged from XIV, and spermathecae between 12/13-18/19. It differs from G. chaophraya Chanabun &amp; Panha, 2013 from Chaophraya River, Nakhonsawan, central Thailand by the latter having longer wings in XXIV, XXV–XXXII, XXXIII, a longer clitellum in XX–XLIII, XLIV, XLV, and spermathecae between 16/17-22/23. It differs from G. chiensis Chanabun &amp; Panha, 2013 from Mahasarakham, northeast Thailand by G. chiensis having wings in XXIII, XXIV, XXV, XXVI–XXIX, XXX, XXXI, XXXII, the clitellum in XVII, XVIII–XXXIII, XXXIV, XXXV, XXXVI, XXXVII, XXXVIII, and spermathecae between 12/13-18/19. It differs from G. quadratus Chanabun &amp; Panha, 2013 reported from the Mun River drainage area, northeast Thailand by G. quadratus having wings in XXIII, XXIV–XXVIII, XXIX, XXX, XXXI, the clitellum in XV, XVI, XVII, XVIII–XXXI, XXXII, XXXIII, XXXIV, XXXV, XXXVI, and spermathecae between 12/13-17/18. It differs from G. huailuangensis Chanabun &amp; Panha, 2013 recorded from Najahlauy National Park, Ubon Ratchathani, northeast Thailand which has wings in XXV, XXVI–XXX, XXXI, a clitellum in XII, XIII, XVI–XXXII, XXXIII, and lacking spermathecae. It differs from G. sekongensis sp. n. by G. sekongensis sp. n. having longer wings in XXV–XXXI, a longer clitellum in XVI, XVII–XXXVI, XXXVII, and spermathecae between 12/13-15/16 (see Table 1).</p></div>	https://treatment.plazi.org/id/830715AC146E9FADBBD0CED922F826CD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Chanabun, Ratmanee;Inkavilay, Khamla;Panha, Somsak	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
9CEA78A4F959CFD6127E63BFC52A5437.text	9CEA78A4F959CFD6127E63BFC52A5437.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glyphidrilus satunensis Chanabun & Panha	<div><p>Glyphidrilus satunensis Chanabun &amp; Panha sp. n. Figs 2, 5, Table 1</p><p>Material examined.</p><p>Holotype: One adult (CUMZ 3405), in Nong Prakpraya, Mueang, Satun, Thailand (06°44'34.0"N, 100°02'23.0"E), 27 meters elevation on 16 January 2014. Paratypes: 15 adults and 21 juveniles (CUMZ 3406), 2 adults (ZMH 14580), 2 adults (NHMUK), and 2 adults (ZRC), all same collection data as holotype.</p><p>Diagnosis.</p><p>Glyphidrilus satunensis sp. n. has the clitellar wings on the lateral side of the body in XXIV, XXV, XXVI–XXIX, XXX, XXXI. Clitellum in XVII, XVIII–XXXII, XXXIII, XXXIV, XXXV. Female pores, male pores and spermathecal pores not visible. Genital markings: unpaired on aa in XVI, XVII, XVIII–XXII, XXIII, XXIV; paired or asymmetrical on bc in XVII, XVIII, XIX. Four pairs of seminal vesicles in IX–XII . Intestinal origin in XVI. Ovaries in XIII–XIV . Spermathecae between 13/14-15/16.</p><p>Description of holotype.</p><p>Dimensions: body length 131 mm, diameter 2.2 mm in segment VIII, 2.3 mm before the clitellar wing in segment XXIII, 2.4 mm after wing in segment XXX in clitellar region; body cylindrical in anterior part, quadrangular in transverse section behind clitellum. 326 segments. Body color pale brown with variations from red to pink at adjacent tissues of wing portion in different individuals of newly collected specimens. At posterior end dorsal surface considerably broader than the ventral. Clitellar wing on ventro-lateral part of clitellum in XXIV–XXIX, 2.5 mm in height, and 0.5 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in XVII–XXXIII . Four pairs of setae per segment from II, setal formula aa:ab:bc:cd:dd = 1.0:0.5:1.5:1.0:1.5 in segment VIII and 1.0:0.5:1.5:0.5:2.0 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: unpaired on aa in XVII–XXII .</p><p>Septa 5/6-7/8 thicker than between other segments, 8/9-9/10 thick and 10/11 to the last segment thin. Gizzard small, globular in VII. Intestine enlarged from XVI. Hearts, four pairs in VIII–XI . A pair of holonephridia in each segment from segment XIII onwards. Seminal vesicles, four pairs in IX–XII . Ovaries, two pairs in XIII–XIV . Testes free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, elongated oval or globular between 13/14-15/16, about 0.2-0.3 mm in diameter, four to five on each side per segment.</p><p>Variations.</p><p>Body lengths of adult (n = 22) and juvenile paratypes (n = 21). Adults ranged from 60+-131 mm (95.8 ± 24.5), with 156+-326 segments. Juveniles ranged from 62-129 mm (82.5 ± 22.1), with 166-323 segments. Wings in XXIV, XXV, XXVI–XXIX, XXX, XXXI, clitellum in XVII, XVIII–XXXII, XXXIII, XXXIV, XXXV. Genital markings: unpaired on aa in XVI, XVII, XVIII–XXII, XXIII, XXIV; paired or asymmetrical on bc in XVII, XVIII, XIX.</p><p>Distribution.</p><p>The new species was found at a pond in Satun, south Thailand.</p><p>Etymology.</p><p>The name " satunensis " is given in reference to the type locality, Satun.</p><p>Remarks.</p><p>See Table 1 for a summary of these comparisons. Glyphidrilus satunensis sp. n. is quite similar to G. kratuensis Chanabun &amp; Panha, 2013 from Kratu waterfall, Kratu, Phuket, south Thailand in the location of wings but G. kratuensis has a smaller size, a shorter clitellum in XVIII–XXX, XXXI, XXXII, the intestine begins from XIV, and spermathecae are between 14/15-17/18. It differs from G. trangensis Chanabun &amp; Panha, 2013 from Trang River, Nayong, Trang, south Thailand by the latter having wings in XXII, XXIII–XXVII, XXVIII, the clitellum in XVII, XVIII–XXX, and spermathecae in XVIII–XXI . It differs from G. wararamensis Chanabun &amp; Panha, 2013 from stream near Wattham Wararam, Phanom, Suratthani, south Thailand by G. wararamensis having wings in XX, XXI–XXVI, XXVII, a clitellum in XI, XII, XIII–XXXIII, XXXIV, XXXV, and spermathecae between 13/14-17/18. It differs from G. nanensis sp. n. by G. nanensis having a smaller size, slightly shorter wings in XXIV, XXV, XXVI–XXVII, XXVIII, XXIX, unpaired genital markings absent, five pairs of hearts in VII–XI, and spermathecae between 13/14-17/18 (see Table 1).</p></div>	https://treatment.plazi.org/id/9CEA78A4F959CFD6127E63BFC52A5437	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Chanabun, Ratmanee;Inkavilay, Khamla;Panha, Somsak	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
375DC007151F758BA5E7BF3FDA373DD8.text	375DC007151F758BA5E7BF3FDA373DD8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glyphidrilus chiangraiensis Chanabun & Panha	<div><p>Glyphidrilus chiangraiensis Chanabun &amp; Panha sp. n. Figs 1, 6, 7, Table 1</p><p>Type material.</p><p>Holotype: One adult (CUMZ 3407) in the river banks of Mekong River at Wat Hatkai, Chiangkhong, Chiangrai, north Thailand (20°15'8.5"N, 100°24'46.8"E), 384 meters elevation on 14 March 2014. Paratypes: 9 adults (CUMZ 3408), 2 adults (ZMH 14581), 2 adults (NHMUK), and 2 adults (ZRC), all specimens collected only from the type locality.</p><p>Other material examined.</p><p>43 adults (CUMZ 3409), in the river banks of Mekong River at Mueang, Buengkan, northeast Thailand (18°22'2.4"N, 103°38'58.0"E), 144 meters elevation on 6 December 2013. 33 adults (CUMZ 3410), in the river banks of Mekong River at Kang Kudku, Chiangkhan, Loei, northeast Thailand (17°54'24.5"N, 101°42'7.5"E), 195 meters elevation on 8 December 2013. 25 adults (CUMZ 3411), in the river banks of Mekong River at Wat Srisomsanook, Chiangkhan, Loei, northeast Thailand (17°59'25.4"N, 101°44'51.3"E), 212 meters elevation on 7 December 2013 . 11 adults (CUMZ 3412), in the river banks of Mekong River at Wat Hadphatum, Srichiangmai, Nongkhai, northeast Thailand (17°57'32.2"N, 102°35'26.8"E), 174 meters elevation on 7 December 2013. 38 adults (CUMZ 3413), in the river banks of Mekong River at Wat Jomnang, Phonphisai, Nongkhai, northeast Thailand (18°01'53.6"N, 103°4'47.4"E), 165 meters elevation on 6 December 2013. 30 adults (CUMZ 3414), in the river banks of Mekong River at Wat Prayanakmai, Wiangkaen, Chiangrai, north Thailand (20°11'45.2"N, 100°27'32.0"E), 359 meters elevation on 15 March 2014. 19 adults (CUMZ 3415), in the river banks of Mekong River at Wat Bansaw, Chiangsan, Chiangrai, north Thailand (20°15'19.4"N, 100°10'44.9"E), 385 meters elevation on 14 March 2014. 26 adults (CUMZ 3416), in river banks of Mekong River at Ban Rimkhong, Pakchom, Loei, northeast Thailand (18°12'48.9"N, 102°04'52.2"E), 181 meters elevation on 8 December 2013. 3 adults (CUMZ 3417), in river banks of Mekong River, Mueng Paksay, Chaiyaburi, Laos (18°12'40.0"N, 101°24'28.1"E), 214 meters elevation on 15 April 2014. 15 adults (CUMZ 3418) in river banks of Mekong River, Bandon, Luangprabang, Laos (19°55'27.6"N, 102°10'49.7"E), 304 meters elevation on 14 April 2014. 24 adults (CUMZ 3419) in river banks of Mekong River, between Sanakham to Vientiane, Laos (17°57'39.7"N, 101°43'53.8"E), 224 meters elevation on 15 April 2014.</p><p>Diagnosis.</p><p>Glyphidrilus chiangraiensis sp. n. has clitellar wings on the lateral side of the body in XXIII, XXIV–XXVI, ½ XXVII, XXVII, ½ XXVIII, XXVIII. Clitellum in XVII, XVIII, XIX, XX, XXI–XXXVI, XXXVII, XXXVIII, XXXIX. Female pores, male pores and spermathecal pores not visible. Genital markings: paired or asymmetrical on aa in XII–XIV, XV, XVI, paired or asymmetrical on bc in XXI, XXII, XXIII, XXIV and XXVII, XXVIII, XXIX–XXXV . Four pairs of seminal vesicles in IX–XII . Intestinal origin in XVI. Ovaries in XIII–XIV . Spermathecae between 15/16-20/21.</p><p>Description of holotype.</p><p>Dimensions: body length 158 mm, diameter 3.0 mm in segment VIII, 4.0 mm before the clitellar wing in segment XXIII, 4.0 mm after wing in segment XXIX in clitellar region; body cylindrical in anterior part, quadrangular in transverse section behind clitellum. 282 segments. Body color pale brown with variations from red to pink at adjacent tissues of wing portion in different individuals of newly collected specimens. Clitellar wing on ventro-lateral part of clitellum in XXIV–XXVII, ½ XXVIII, 3.7 mm and 4.0 mm in height, and 0.5 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in XVII–XXXVIII . Four pairs of setae per segment from II, setal formula aa:ab:bc:cd:dd=2.0:1.0:2.0:1.0:2.0 in segment VIII and 2.0:1.0:2.0:1.0:2.5 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: paired on aa in XIV, laterally paired or asymmetrical on bc in XXIII and XXIX–XXXV .</p><p>Septa 5/6-8/9 thicker than between other segments, 9/10-13/14 thick and 14/15 to the last segment thin. Gizzard small, globular in VIII. Intestine enlarged from XVI. Hearts, five pairs in VII–XI, first in VII and last in XI. A pair of holonephridia in each segment from segment XIII onwards. Seminal vesicles, four pairs in IX, X, XI, XII. Ovaries, two pairs in XIII and XIV. Testis free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, small elongated oval or globular between 15/16-20/21, about 0.1-0.2 mm in diameter, three to twelve per segment on each side per segment.</p><p>Variations.</p><p>Body lengths of adults (n = 283) ranged from 94-340 mm (155.42 ± 54.93), with 89-394 segments. Wings in XXIII, XXXIV–XXVI, ½ XXVII, XXVII, ½ XXVIII, XXVIII, clitellum in XVII, XVIII, XIX, XX, XXI–XXXVI, XXXVII, XXXVIII, XXXIX. Genital markings: paired or asymmetrical on aa in XII–XIV, XV, XVI, paired or asymmetrical on bc in XXI, XXII, XXIII, XXIV and XXVII, XXVIII, XXIX–XXXV .</p><p>Distribution.</p><p>The new species is known from the type locality in the river banks of Mekong River at Wat Hatkai, Chiangkhong, Chiangrai, north Thailand, and was found in several locations along the Mekong River and its tributaries in the northeast and north of Thailand at Buengkan, Loei, Nongkhai, and Chiangrai and in Chaiyaburi, Luangprabang, and Sanakham to Vientiane of Laos.</p><p>Etymology.</p><p>The species was named after Chiangrai, the locality name.</p><p>Remarks.</p><p>(see Table 1) Glyphidrilus chiangraiensis sp. n. differs from G. vangviengensis Chanabun &amp; Panha, 2011 from Song River, Vieintiane, Laos by the latter having longer wings in XXIV, XXV–XXXI, XXXII, the genital markings widely paired in bc XVIII, XIX, XX, XXI–XXIV and XXXIII, XXXIV, paired on aa in XII–XIV, XV and lacking spermathecae. It differs from G. yunnanensis Chen &amp; Xu, 1977 reported from China by G. yunnanensis having longer wings in XXII–XXXII, clitellum in XVIII–XXXVIII, and lacking spermathecae. It differs from the species recorded as G. mekongensis Panha &amp; Chanabun, 2012 from Mekong River, Thailand by G. mekongensis having longer wings in XXIV–½ XXXIII, XXXIII, XXXIV, ½ XXXV, and lacking spermathecae. It differs from G. chiensis Chanabun &amp; Panha, 2013 from Chi River, Mahasarakham, northeast Thailand by G. chiensis having longer wings in XXIII, XXIV, XXV, XXVI–XXIX, XXX, XXXI, XXXII, and spermathecae between 12/13-18/19. It differs from G. quadratus Chanabun &amp; Panha, 2013 reported from the Mun River by G. quadratus having longer wings in XXIII, XXIV–XXVIII, XXIX, XXX, XXXI, a bit longer clitellum in XV, XVI, XVII, XVIII–XXXI, XXXII, XXXIII, XXXIV, XXXV, XXXVI, and spermathecae between 12/13-17/18. It differs from G. huailuangensis Chanabun &amp; Panha, 2013 recorded from Najahlauy National Park, Ubon Ratchathani, northeast Thailand by the latter having longer wings in XXV, XXVI–XXX, XXXI, clitellum in XII, XIII, XVI–XXXII, XXXIII, and lacks spermathecae. Glyphidrilus chiangraiensis sp. n. differs from G. namphao sp. n. by the latter having wings in XVIII–XXIV, clitellum in XVII–XXVI, XXIX, genital markings: paired on aa in XVII; paired or asymmetrical on bc in XII–XV, XXV, XXVI, and spermathecae between 13/14-16/17. It differs from G. sekongensis sp. n. by the latter having wings in XXV–XXXI, clitellum in XVI, XVII–XXXVI, XXXVII, and spermathecae between 12/13-15/16. It differs from G. champasakensis sp. n. from Mekong River at Ban Khonkhen, Champasak, Laos by the latter having longer wings in XXIII, XXIV–½ XXXII, XXXII, XXXIII, longer clitellum in XIX, XX–XLIX, L, LI, LII, and spermathecae between 14/15-19/20.</p></div>	https://treatment.plazi.org/id/375DC007151F758BA5E7BF3FDA373DD8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Chanabun, Ratmanee;Inkavilay, Khamla;Panha, Somsak	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
1ED54A23B5607D1B3F79877C70544E99.text	1ED54A23B5607D1B3F79877C70544E99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glyphidrilus namphao Chanabun & Panha	<div><p>Glyphidrilus namphao Chanabun &amp; Panha sp. n. Figs 1, 8, Table 1</p><p>Type material.</p><p>Holotype: One adult (CUMZ 3420) in a river banks of Phao River between Ban Lak 7 and Ban Lak 5, Kamkerd, Bolikhamxai, Laos (18°16'27.7"N, 105°2'44.0"E), 525 meters elevation on 7 December 2013. Paratypes: 2 adults and 8 juveniles (CUMZ 3421), 1 adult (ZMH 14582), all specimens collected from only the type locality.</p><p>Diagnosis.</p><p>Glyphidrilus namphao sp. n. is a small worm with the clitellar wings on the lateral side of the body in XVIII–XXIV . Clitellum in XVII–XXVI, XXIX. Female pores, male pores and spermathecal pores not visible. Genital markings: paired or asymmetrical on aa in XVII; paired or asymmetrical on bc in XII–XV and XXV, XXVI. Four pairs of seminal vesicles in IX–XII . Intestinal origin in XVI. Ovaries in XIII–XIV . Spermathecae between 13/14-16/17.</p><p>Description of holotype.</p><p>Dimensions: body length 92 mm, diameter 3.1 mm in segment VIII, 4.2 mm before the clitellar wing in segment XVII, 4.2 mm after wing in segment XXV in clitellar region; body cylindrical in anterior part, quadrangular in transverse section behind clitellum. 269 segments. Body color pale brown with variations from red to pink at adjacent tissues of wing portion in different individuals of newly collected specimens. At posterior end dorsal surface considerably broader than the ventral. Clitellar wing on ventro-lateral part of clitellum in XVIII–XXIV, 3.5 mm in height, and 0.25 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in XVII–XXIX . Four pairs of setae per segment from II, setal formula aa:ab:bc:cd:dd =1.5:0.5:1.5:0.5:2.0 in segment VIII and 1.5:0.5:1.5:0.5:2.5 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: paired on aa in XVII, laterally paired or asymmetrical on bc in XII–XV and XXV.</p><p>Septa 4/5-6/7 thicker than between other segments, 7/8-11/12 thick and 12/13 to the last segment thin. Gizzard small, globular in VIII. Intestine enlarged from XVI. Hearts, five pairs in VII–XI, first in VII and last in XI. A pair of holonephridia in each segment from segment XIII onwards. Seminal vesicles, four pairs in IX, X, XI, XII. Ovaries, two pairs in XIII and XIV. Testis free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, elongated oval or globular between 13/14-16/17, about 0.3-0.4 mm in diameter, two to seven on each side per segment.</p><p>Variations.</p><p>Body lengths of adult (n = 4) and juvenile paratypes (n = 8). Adults ranged from 64-122 mm (89.7 ± 20.8), with 190-320 segments. Juveniles ranged from 64-120 mm (98.3 ± 17.6), with 193-311 segments. Wings in XVIII–XXIV, clitellum in XVII–XXVI, XXIX. Genital markings: paired or asymmetrical on aa in XVII; paired or asymmetrical on bc in XII–XV and XXV, XXVI.</p><p>Distribution.</p><p>The new species was found in the river banks of Phao River which was covered with worm casts.</p><p>Etymology.</p><p>The species was named after the Phao River, the type locality.</p><p>Remarks .</p><p>Glyphidrilus namphao sp. n. is different from G. vangviengensis Chanabun &amp; Panha, 2011 reported from Song River Veintiane, Laos in different locations of wings in XXIV, XXV–XXXI, XXXII, longer clitellum in XIX, XX–XXXV, XXXVI, XXXVII and lacking spermathecae. It differs from G. nanensis sp. n. from Nan River at Saklek, Phichit, north Thailand by G. nanensis sp. n. having wings in XXIV, XXV, XXVI -XXVII, XXVIII, XXIX, a longer clitellum in XVII, XVIII–XXXIII, XXXIV, and spermathecae between 13/14-17/18. It is different from G. sekongensis sp. n. from Ban Kiangkong, Lamarm, Sekong, Laos in the locations of wings in XXV–XXXI, a longer clitellum in XVI, XVII–XXXVI, XXXVII, the genital markings paired or asymmetrical on bc in XVI, XVII–XXIV, XXXII, and spermathecae between 12/13-15/16. Glyphidrilus namphao sp. n. differs from G. chiangraiensis sp. n. from Mekong River at Wat Hatkai, Chiangkhong, Chiangrai, north Thailand in the locations of wings in XXIII, XXIV–XXVI, ½ XXVII, XXVII, ½ XXVIII, XXVIII, a longer clitellum in XVII, XVIII, XIX, XX, XXI–XXXVI, XXXVII, XXXVIII, XXXIX, and spermathecae between 15/16-20/21 (see Table 1).</p></div>	https://treatment.plazi.org/id/1ED54A23B5607D1B3F79877C70544E99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Chanabun, Ratmanee;Inkavilay, Khamla;Panha, Somsak	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
D19F4D5966A6F9EF119FBC1730D3B83E.text	D19F4D5966A6F9EF119FBC1730D3B83E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glyphidrilus sekongensis Chanabun & Panha	<div><p>Glyphidrilus sekongensis Chanabun &amp; Panha sp. n. Figs 1, 9, Table 1</p><p>Type material.</p><p>Holotype: One adult (CUMZ 3422) in a stream at Ban Kiangkong, Lamarm, Sekong, Laos (15°33'30.6"N, 106°19'19.4"E), 472 meters elevation on 17 October 2013. Paratypes: 2 adults and 11 juveniles (CUMZ 3423) all same collection data as holotype.</p><p>Diagnosis.</p><p>Glyphidrilus sekongensis sp. n. is a small sized earthworm with the clitellar wings on the lateral side of the body in XXV–XXXI . Clitellum in XVI, XVII–XXXVI, XXXVII. Female pores, male pores and spermathecal pores not visible. Genital markings: paired or asymmetrical on bc in XVI, XVII–XXIV and XXXII. Four pairs of seminal vesicles in IX–XII . Intestinal origin in XVI. Ovaries in XIII–XIV . Spermathecae between 12/13-15/16.</p><p>Description of holotype.</p><p>Dimensions: body length 134 mm, diameter 2.4 mm in segment VIII, 2.3 mm before the clitellar wing in segment XXIV, 2.4 mm after wing in segment XXXII in clitellar region; body cylindrical in anterior part, quadrangular in transverse section behind clitellum. 329 segments. Body color pale brown with variations from red to pink on adjacent tissues of wing portions in different individuals of newly collected specimens. The posterior end on the dorsal surface is considerably broader than the ventral. Clitellar wing on ventro-lateral part of clitellum in XXV–XXXI, 4.0 mm in height, and 0.5 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in XVI–XXXVII . Four pairs of setae per segment from II, setal formula aa:ab:bc:cd:dd = 1.0:0.3:1.0:0.5:1.3 in segment VIII and 1.8:0.8:1.3:0.6:1.5 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: laterally paired or asymmetrical on bc in XVI–XXIV, XXXII.</p><p>Septa 5/6-8/9 thicker than between other segments, 9/10-14/15 thick and 15/16 to the last others segment, 9/10-14/15 thick and 15/16 to the last segment thin. Gizzard small, globular in VIII. Intestine enlarged from XVI. Hearts, four pairs in VIII–XI, first in VIII and last in XI. A pair of holonephridia in each segment from segment XIII onwards. Seminal vesicles, four pairs in IX, X, XI, XII. Ovaries, two pairs in XIII and XIV. Testis free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, small elongated oval or globular between 12/13-15/16, about 0.1-0.3 mm in diameter, two to seven on each side per segment.</p><p>Variations.</p><p>Body lengths of adult (n = 3) and juvenile paratypes (n = 11). Adults ranged from 90-134 mm (109.6 ± 12.2), with 237-337 segments. Juveniles ranged from 86+-130 mm (104.2 ± 14.0), with 209+-329 segments. Wings in XXV–XXXI, clitellum in XVI, XVII–XXXVI, XXXVII. Genital markings: laterally paired or asymmetrical on bc in XVI, XVII–XXIV, XXXII.</p><p>Distribution.</p><p>The new species was found in a stream at Ban Kiangkong, Lamarm, Sekong, Laos. Soils are slightly sandy mixed with black organic matter.</p><p>Etymology.</p><p>The species was named after Sekong, the type locality in Laos.</p><p>Remarks.</p><p>(see Table 1). Glyphidrilus sekongensis sp. n. differs from other species by the locations of external and internal characteristics such as wings, clitellum, the arrangement of genital markings, and spermathecae. The new species differs from G. vangviengensis Chanabun &amp; Panha, 2011 from Song River, Vientiane, Laos by the latter having a shorter clitellum in XIX, XX–XXXV, XXXVI, XXXVII, widely paired genital markings in bc XVIII, XIX, XX, XXI–XXIV, XXXIII, XXXIV, paired on aa in XII, XIII, XIV, XV, and lacks spermathecae. Differences from G. yunnanensis Chen &amp; Xu, 1977 reported from China are that G. yunnanensis has longer wings in XXII–XXXII, clitellum in XVIII–XXXVIII, and lacks spermathecae. Differences from G. mekongensis Panha &amp; Chanabun, 2012 are that G. mekongensis has longer wings in XXIV–½ XXXIII, XXXIII, XXXIV, ½ XXXV, and lacks spermathecae. It differs from G. huailuangensis Chanabun &amp; Panha, 2013 recorded from Najahlauy National Park, Ubon Ratchathani, northeast Thailand by the latter having a larger body size, clitellum in XII, XIII, XVI–XXXII, XXXIII, and lacks spermathecae. It differs from G. champasakensis sp. n. from Mekong River at Ban Khonkhen, Champasak, Laos by the latter having longer wings in XXIII, XXIV–½ XXXII, XXXII, XXXIII, longer clitellum in XIX, XX–XLIX, L, LI, LII, and spermathecae between 14/15-19/20.</p></div>	https://treatment.plazi.org/id/D19F4D5966A6F9EF119FBC1730D3B83E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Chanabun, Ratmanee;Inkavilay, Khamla;Panha, Somsak	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
84DAC461D84A960851E34F34C2436402.text	84DAC461D84A960851E34F34C2436402.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glyphidrilus namdonensis Chanabun & Panha	<div><p>Glyphidrilus namdonensis Chanabun &amp; Panha sp. n. Figs 1, 10, Table 1</p><p>Type material.</p><p>Holotype: One adult (CUMZ 3424) in the banks of Done River at Ban Namdone, Thakhek, Khammouan, Laos (17°28'39.9"N, 104°45'35.1"E), 161 meters elevation on 6 December 2013. Paratypes: 16 adults and 6 juveniles (CUMZ 3425), 2 adults (ZMH 14583), 2 adults (NHMUK), and 2 adults (RMBR), all same collection data as holotype.</p><p>Diagnosis.</p><p>Glyphidrilus namdonensis sp. n. is small sized earthworm with distinctly expanded tissues, the clitellar wings on the lateral side of the body in XXIV, XXV–XXVIII, XXX. Clitellum in XIX, XX–XXXVI, XXXVII, XXXVIII. Female pores, male pores and spermathecal pores not visible. Genital markings: medium paired or asymmetrical on aa in XIII, XIV; paired or asymmetrical on bc in XXII, XXIII, XXIV and XXVII, XXVIII–XXXIII . Four pairs of seminal vesicles in IX–XII . Intestinal origin in XVI. Ovaries in XIII–XIV . Spermathecae between 14/15-18/19.</p><p>Description of holotype.</p><p>Dimensions: body length 92 mm, diameter 3.5 mm in segment VIII, 4.0 mm before the clitellar wing in segment XXIII, 4.0 mm after the wing in segment XXXI in the clitellar region; body cylindrical in the anterior part, quadrangular in transverse section behind clitellum. 216 segments. Body color pale brown with variations from red to pink in adjacent tissues of the wing portions in different individuals of newly collected specimens. Clitellar wing on ventro-lateral part of clitellum in XXIV–XXX, 5.0 mm in height, and 0.2 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in XX–XXXVII . Four pairs of setae per segment from II, setal formula aa:ab:bc:cd:dd = 1.3:0.5:1.0:0.5:2.0 in segment VIII and 1.5:0.5:1.0:0.5:2.3 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: medium paired on aa in XIII, XIV, laterally paired or asymmetrical on bc in XXII, XXIII.</p><p>Septa 5/6-8/9 thicker than between other segments, 9/10-14/15 thick, and 15/16 to the last segment thin. Gizzard small, globular in VIII. Intestine enlarged from XVI. Hearts, five pairs in VII–XI, first in VII and last in XI. A pair of holonephridia in each segment from segment XII onwards. Seminal vesicles, four pairs in IX, X, XI, XII. Ovaries two pairs in XIII and XIV. Testis free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, small elongated oval or globular between 14/15-18/19, about 0.1-0.2 mm in diameter, two to thirteen on each side per segment.</p><p>Variations.</p><p>Body lengths of adult (n = 23) and juvenile paratypes (n = 6). Adults ranged from 90-139 mm (121.9 ± 25.9), with 183-259 segments. Juveniles ranged from 87-136 mm (103.3 ± 17.4), with 185-223 segments. Wings in XXIV, XXV–XXVIII, XXX, clitellum in XIX, XX–XXXVI, XXXVII, XXXVIII. Genital markings: medium paired or asymmetrical on aa in XIII, XIV; paired or asymmetrical on bc in XXII, XXIII, XXIV and XXVII, XXVIII–XXXIII .</p><p>Distribution.</p><p>The new species was found only from the banks of Done River at Ban Namdone, Thakhek, Khammouan, Laos.</p><p>Etymology.</p><p>The species was named after the Done River.</p><p>Remarks.</p><p>Glyphidrilus namdonensis sp. n. is compared with other Glyphidrilus having spermathecae and a clitellum starting feom segment XIX. Glyphidrilus namdonensis differs from G. champasakensis sp. n. by the latter having longer wings in XXIII, XXIV–½ XXXII, XXXII, XXXIII, longer clitellum in XIX, XX–XLIX, L, LI, LII, and spermathecae between 14/15-19/20 (see Table 1).</p></div>	https://treatment.plazi.org/id/84DAC461D84A960851E34F34C2436402	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Chanabun, Ratmanee;Inkavilay, Khamla;Panha, Somsak	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
BEBC94D9DC8FF68804DC4BE7B52753FF.text	BEBC94D9DC8FF68804DC4BE7B52753FF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glyphidrilus champasakensis Chanabun & Panha	<div><p>Glyphidrilus champasakensis Chanabun &amp; Panha sp. n. Figs 1, 11, 12, Table 1</p><p>Type material.</p><p>Holotype: One adult (CUMZ 3426) in the banks of Mekong River at Ban Khonkhen, Champasak, Laos (15°02'21.2"N, 105°51'20.4"E), 106 meters elevation on 17 April 2014. Paratypes: 17 adults (CUMZ 3427), 2 adults (ZMH 14584), 2 adults (NHMUK), and 2 adults (RMBR), all specimens collected from type locality.</p><p>Diagnosis .</p><p>Glyphidrilus champasakensis sp. n. is a large sized earthworm with distinct expanded tissues of the clitellar wings on the sides of the body in XXIII, XXIV–½ XXXII, XXXII, XXXIII. Clitellum in XIX, XX– XLIX, L, LI, LII. Female pores, male pores and spermathecal pores not visible. Genital markings: medium paired or asymmetrical on aa in XII, XIII, XXXIII, XXXIV; paired or asymmetrical on bc in XVII, XVIII, XIX–XXII, XXIII, XXIV and XXXII, XXXIII, XXXIV, XXXV. Four pairs of seminal vesicles in IX–XII . Intestinal origin in XVI. Large ovaries in XIII–XIV . Spermathecae between 14/15-19/20.</p><p>Description of holotype.</p><p>Dimensions: body length 301 mm, diameter 5.0 mm in segment VIII, 5.0 mm before the clitellar wing in segment XXII, 5.0 mm after wing in segment XXXIII in clitellar region; body cylindrical in anterior part, quadrangular in transverse section behind clitellum. 424 segments. Body color pale brown with variations from red to pink at adjacent tissues of wing portion in different individuals of newly collected specimens. Clitellar wing on ventro-lateral part of clitellum in XXIII–XXXII, 10.0 mm in height, and 0.2 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in XIX– XLIX. Four pairs of setae per segment from II, setal formula aa:ab:bc:cd:dd = 2.0:1.0:2.0:1.0:3.0 in segment VIII and 2.0:1.0:2.0:1.0:2.5 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: medium paired or asymmetrical on aa in XIII, XXXIV, laterally paired or asymmetrical on bc in XIX–XXII and XXXIII, XXXIV.</p><p>Septa 4/5-8/9 thicker than between other segments others segment, 9/10-11/12 thick and 12/13 to the last segment thin. Gizzard small, globular in VIII. Intestine enlarged from XVI. Hearts, five pairs in VII–XI, first in VII and last in XI. A pair of holonephridia in each segment from segment XIV onwards. Seminal vesicles, four pairs in IX, X, XI, XII. Ovaries, two pairs in XIII and XIV. Testis free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, small elongated oval or globular between 14/15-19/20, about 0.1-0.3 mm in diameter, five to thirteen on each side per segment.</p><p>Variations.</p><p>Body lengths of adult (n = 24) ranged from 167-301 mm (212.75 ± 42.15), with 248-424 segments. Wings in XXIII, XXIV–½ XXXII, XXXII, XXXIII, clitellum in XIX, XX– XLIX, L, LI, LII. Genital markings: paired or asymmetrical on aa in XII, XIII, XXXIII, XXXIV; paired or asymmetrical on bc in XVII, XVIII, XIX–XXII, XXIII, XXIV and XXXII, XXXIII, XXXIV, XXXV.</p><p>Distribution.</p><p>The new species was found only on the banks of the Mekong River at Ban Khonkhen, Champasak, Laos.</p><p>Etymology.</p><p>The new species was named after Champasak, Laos, the type locality.</p><p>Remarks.</p><p>Here we compare Glyphidrilus champasakensis sp. n. to the species with spermathecae and a clitellum beginning from segment XIX. It differs from G. namdonensis sp. n. by G. namdonensis sp. n. having shorter wings in XXIV, XXV–XXVIII, XXX, and spermathecae between 14/15-18/19 (see Table 1).</p></div>	https://treatment.plazi.org/id/BEBC94D9DC8FF68804DC4BE7B52753FF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Chanabun, Ratmanee;Inkavilay, Khamla;Panha, Somsak	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
