taxonID	type	description	language	source
6D48946A8FA5C527097A26353C04EFC8.taxon	diagnosis	Diagnosis. Prostomium zygolobous. Body shape nearly circular in cross section in anterior part, and becoming quadrangular in posterior part or after clitellum. Anus dorsal or dorso-terminal. A longitudinal lamellar ridge at maturity from body wall on each side in bc, through several of the clitellar segments, which are called wings. Dorsal pores absent. Setae four pairs per segment. Clitellum annular. Genital apertures, all minute and superficial. Male pores inter-or intraclitellar. Spermathecal pores usually all behind the testis segments. Gizzard in VII or VIII sometimes extending into an adjacent segment. Calciferous glands absent. Seminal vesicles usually short, usually four pairs in IX-XII. Holonephridia. Nephrostomes single ducts avesiculate and without sphincters or caeca. Testis and funnels free in X and XI. Male ducts intramural. Ovaries fan shaped and with several egg strings. Ovisacs present or absent. Prostate glands absent and spermathecae without diverticulum (Chanabun et al. 2013, Gates 1972).	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
830715AC146E9FADBBD0CED922F826CD.taxon	diagnosis	Diagnosis. A small sized earthworm with the clitellar wings on the lateral side of the body in XXIV, XXV, XXVI- XXVII, XXVIII, XXIX. Clitellum in XVII, XVIII-XXXIII, XXXIV. Female pores, male pores, and spermathecal pores not visible. Genital markings: paired or asymmetrical on aa in X, XI, XII, XIII, XIV and XXVIII, XXIX, XXX; paired or asymmetrical on bc in XV, XVI, XVII-XXIII, XXIV, XXV. Four pairs of seminal vesicles in IX-XII. Intestinal origin in XV. Ovaries in XIII-XIV. Spermathecae between 13 / 14 - 17 / 18.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
830715AC146E9FADBBD0CED922F826CD.taxon	description	Description of holotype. Dimensions: body length 78 mm, diameter 2.6 mm in segment VIII, 4.7 mm before the clitellar wing in segment XXIII, 4.2 mm after wing in segment XXX in clitellar region; body cylindrical in anterior part, quadrangular in transverse section behind clitellum. 241 segments. Body color pale brown with variations from red to pink at adjacent tissues of wing portion in different individuals of newly collected specimens. At posterior end dorsal surface considerably broader than the ventral. Clitellar wing on ventro-lateral part of clitellum in XXVI-XXIX, 3.6 mm in height, and 0.3 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in XVII-XXXIII. Four pairs of setae per segment from II, setal formula aa: ab: bc: cd: dd = 1.0: 0.6: 1.3: 0.6: 1.4 in segment VIII and 1.0: 0.6: 1.5: 0.6: 1.5 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: paired on aa in XII-XIII, laterally paired or asymmetrical on bc in XVII-XXIV. Septa 5 / 6 - 8 / 9 thicker than between other segments, 9 / 10 - 14 / 15 thick and 15 / 16 to the last segment thin. Gizzard small, globular in 7 / 8. Intestine enlarged from XV. Hearts, five pairs in VII-XI, first in VII and last in XI. A pair of holonephridia in each segment from segment XIII onwards. Seminal vesicles, four pairs in IX, X, XI, XII. Ovaries, two pairs in XIII-XIV. Testes free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, elongated oval or globular between 13 / 14 - 17 / 18, about 0.2 - 0.3 mm in diameter, two to ten on each side per segment.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
830715AC146E9FADBBD0CED922F826CD.taxon	distribution	Distribution. The new species was found in the river banks of Nan River at Saklek, Phichit, north Thailand, which was covered with worm casts.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
830715AC146E9FADBBD0CED922F826CD.taxon	etymology	Etymology. The species was named after the locality on the banks of the Nan River. This was the first time that the worm genus was ever recorded from near this river.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
9CEA78A4F959CFD6127E63BFC52A5437.taxon	materials_examined	Material examined. Holotype: One adult (CUMZ 3405), in Nong Prakpraya, Mueang, Satun, Thailand (06 ° 44 ' 34.0 " N, 100 ° 02 ' 23.0 " E), 27 meters elevation on 16 January 2014. Paratypes: 15 adults and 21 juveniles (CUMZ 3406), 2 adults (ZMH 14580), 2 adults (NHMUK), and 2 adults (ZRC), all same collection data as holotype.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
9CEA78A4F959CFD6127E63BFC52A5437.taxon	diagnosis	Diagnosis. Glyphidrilus satunensis sp. n. has the clitellar wings on the lateral side of the body in XXIV, XXV, XXVI-XXIX, XXX, XXXI. Clitellum in XVII, XVIII-XXXII, XXXIII, XXXIV, XXXV. Female pores, male pores and spermathecal pores not visible. Genital markings: unpaired on aa in XVI, XVII, XVIII-XXII, XXIII, XXIV; paired or asymmetrical on bc in XVII, XVIII, XIX. Four pairs of seminal vesicles in IX-XII. Intestinal origin in XVI. Ovaries in XIII-XIV. Spermathecae between 13 / 14 - 15 / 16.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
9CEA78A4F959CFD6127E63BFC52A5437.taxon	description	Description of holotype. Dimensions: body length 131 mm, diameter 2.2 mm in segment VIII, 2.3 mm before the clitellar wing in segment XXIII, 2.4 mm after wing in segment XXX in clitellar region; body cylindrical in anterior part, quadrangular in transverse section behind clitellum. 326 segments. Body color pale brown with variations from red to pink at adjacent tissues of wing portion in different individuals of newly collected specimens. At posterior end dorsal surface considerably broader than the ventral. Clitellar wing on ventro-lateral part of clitellum in XXIV-XXIX, 2.5 mm in height, and 0.5 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in XVII-XXXIII. Four pairs of setae per segment from II, setal formula aa: ab: bc: cd: dd = 1.0: 0.5: 1.5: 1.0: 1.5 in segment VIII and 1.0: 0.5: 1.5: 0.5: 2.0 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: unpaired on aa in XVII-XXII. Septa 5 / 6 - 7 / 8 thicker than between other segments, 8 / 9 - 9 / 10 thick and 10 / 11 to the last segment thin. Gizzard small, globular in VII. Intestine enlarged from XVI. Hearts, four pairs in VIII-XI. A pair of holonephridia in each segment from segment XIII onwards. Seminal vesicles, four pairs in IX-XII. Ovaries, two pairs in XIII-XIV. Testes free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, elongated oval or globular between 13 / 14 - 15 / 16, about 0.2 - 0.3 mm in diameter, four to five on each side per segment.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
9CEA78A4F959CFD6127E63BFC52A5437.taxon	distribution	Distribution. The new species was found at a pond in Satun, south Thailand.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
9CEA78A4F959CFD6127E63BFC52A5437.taxon	etymology	Etymology. The name " satunensis " is given in reference to the type locality, Satun.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
375DC007151F758BA5E7BF3FDA373DD8.taxon	materials_examined	Other material examined. 43 adults (CUMZ 3409), in the river banks of Mekong River at Mueang, Buengkan, northeast Thailand (18 ° 22 ' 2.4 " N, 103 ° 38 ' 58.0 " E), 144 meters elevation on 6 December 2013. 33 adults (CUMZ 3410), in the river banks of Mekong River at Kang Kudku, Chiangkhan, Loei, northeast Thailand (17 ° 54 ' 24.5 " N, 101 ° 42 ' 7.5 " E), 195 meters elevation on 8 December 2013. 25 adults (CUMZ 3411), in the river banks of Mekong River at Wat Srisomsanook, Chiangkhan, Loei, northeast Thailand (17 ° 59 ' 25.4 " N, 101 ° 44 ' 51.3 " E), 212 meters elevation on 7 December 2013. 11 adults (CUMZ 3412), in the river banks of Mekong River at Wat Hadphatum, Srichiangmai, Nongkhai, northeast Thailand (17 ° 57 ' 32.2 " N, 102 ° 35 ' 26.8 " E), 174 meters elevation on 7 December 2013. 38 adults (CUMZ 3413), in the river banks of Mekong River at Wat Jomnang, Phonphisai, Nongkhai, northeast Thailand (18 ° 01 ' 53.6 " N, 103 ° 4 ' 47.4 " E), 165 meters elevation on 6 December 2013. 30 adults (CUMZ 3414), in the river banks of Mekong River at Wat Prayanakmai, Wiangkaen, Chiangrai, north Thailand (20 ° 11 ' 45.2 " N, 100 ° 27 ' 32.0 " E), 359 meters elevation on 15 March 2014. 19 adults (CUMZ 3415), in the river banks of Mekong River at Wat Bansaw, Chiangsan, Chiangrai, north Thailand (20 ° 15 ' 19.4 " N, 100 ° 10 ' 44.9 " E), 385 meters elevation on 14 March 2014. 26 adults (CUMZ 3416), in river banks of Mekong River at Ban Rimkhong, Pakchom, Loei, northeast Thailand (18 ° 12 ' 48.9 " N, 102 ° 04 ' 52.2 " E), 181 meters elevation on 8 December 2013. 3 adults (CUMZ 3417), in river banks of Mekong River, Mueng Paksay, Chaiyaburi, Laos (18 ° 12 ' 40.0 " N, 101 ° 24 ' 28.1 " E), 214 meters elevation on 15 April 2014. 15 adults (CUMZ 3418) in river banks of Mekong River, Bandon, Luangprabang, Laos (19 ° 55 ' 27.6 " N, 102 ° 10 ' 49.7 " E), 304 meters elevation on 14 April 2014. 24 adults (CUMZ 3419) in river banks of Mekong River, between Sanakham to Vientiane, Laos (17 ° 57 ' 39.7 " N, 101 ° 43 ' 53.8 " E), 224 meters elevation on 15 April 2014.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
375DC007151F758BA5E7BF3FDA373DD8.taxon	diagnosis	Diagnosis. Glyphidrilus chiangraiensis sp. n. has clitellar wings on the lateral side of the body in XXIII, XXIV-XXVI, 1 / 2 XXVII, XXVII, 1 / 2 XXVIII, XXVIII. Clitellum in XVII, XVIII, XIX, XX, XXI-XXXVI, XXXVII, XXXVIII, XXXIX. Female pores, male pores and spermathecal pores not visible. Genital markings: paired or asymmetrical on aa in XII-XIV, XV, XVI, paired or asymmetrical on bc in XXI, XXII, XXIII, XXIV and XXVII, XXVIII, XXIX-XXXV. Four pairs of seminal vesicles in IX-XII. Intestinal origin in XVI. Ovaries in XIII-XIV. Spermathecae between 15 / 16 - 20 / 21.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
375DC007151F758BA5E7BF3FDA373DD8.taxon	description	Description of holotype. Dimensions: body length 158 mm, diameter 3.0 mm in segment VIII, 4.0 mm before the clitellar wing in segment XXIII, 4.0 mm after wing in segment XXIX in clitellar region; body cylindrical in anterior part, quadrangular in transverse section behind clitellum. 282 segments. Body color pale brown with variations from red to pink at adjacent tissues of wing portion in different individuals of newly collected specimens. Clitellar wing on ventro-lateral part of clitellum in XXIV-XXVII, 1 / 2 XXVIII, 3.7 mm and 4.0 mm in height, and 0.5 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in XVII-XXXVIII. Four pairs of setae per segment from II, setal formula aa: ab: bc: cd: dd = 2.0: 1.0: 2.0: 1.0: 2.0 in segment VIII and 2.0: 1.0: 2.0: 1.0: 2.5 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: paired on aa in XIV, laterally paired or asymmetrical on bc in XXIII and XXIX-XXXV. Septa 5 / 6 - 8 / 9 thicker than between other segments, 9 / 10 - 13 / 14 thick and 14 / 15 to the last segment thin. Gizzard small, globular in VIII. Intestine enlarged from XVI. Hearts, five pairs in VII-XI, first in VII and last in XI. A pair of holonephridia in each segment from segment XIII onwards. Seminal vesicles, four pairs in IX, X, XI, XII. Ovaries, two pairs in XIII and XIV. Testis free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, small elongated oval or globular between 15 / 16 - 20 / 21, about 0.1 - 0.2 mm in diameter, three to twelve per segment on each side per segment.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
375DC007151F758BA5E7BF3FDA373DD8.taxon	distribution	Distribution. The new species is known from the type locality in the river banks of Mekong River at Wat Hatkai, Chiangkhong, Chiangrai, north Thailand, and was found in several locations along the Mekong River and its tributaries in the northeast and north of Thailand at Buengkan, Loei, Nongkhai, and Chiangrai and in Chaiyaburi, Luangprabang, and Sanakham to Vientiane of Laos.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
375DC007151F758BA5E7BF3FDA373DD8.taxon	etymology	Etymology. The species was named after Chiangrai, the locality name.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
1ED54A23B5607D1B3F79877C70544E99.taxon	diagnosis	Diagnosis. Glyphidrilus namphao sp. n. is a small worm with the clitellar wings on the lateral side of the body in XVIII-XXIV. Clitellum in XVII-XXVI, XXIX. Female pores, male pores and spermathecal pores not visible. Genital markings: paired or asymmetrical on aa in XVII; paired or asymmetrical on bc in XII-XV and XXV, XXVI. Four pairs of seminal vesicles in IX-XII. Intestinal origin in XVI. Ovaries in XIII-XIV. Spermathecae between 13 / 14 - 16 / 17.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
1ED54A23B5607D1B3F79877C70544E99.taxon	description	Description of holotype. Dimensions: body length 92 mm, diameter 3.1 mm in segment VIII, 4.2 mm before the clitellar wing in segment XVII, 4.2 mm after wing in segment XXV in clitellar region; body cylindrical in anterior part, quadrangular in transverse section behind clitellum. 269 segments. Body color pale brown with variations from red to pink at adjacent tissues of wing portion in different individuals of newly collected specimens. At posterior end dorsal surface considerably broader than the ventral. Clitellar wing on ventro-lateral part of clitellum in XVIII-XXIV, 3.5 mm in height, and 0.25 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in XVII-XXIX. Four pairs of setae per segment from II, setal formula aa: ab: bc: cd: dd = 1.5: 0.5: 1.5: 0.5: 2.0 in segment VIII and 1.5: 0.5: 1.5: 0.5: 2.5 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: paired on aa in XVII, laterally paired or asymmetrical on bc in XII-XV and XXV. Septa 4 / 5 - 6 / 7 thicker than between other segments, 7 / 8 - 11 / 12 thick and 12 / 13 to the last segment thin. Gizzard small, globular in VIII. Intestine enlarged from XVI. Hearts, five pairs in VII-XI, first in VII and last in XI. A pair of holonephridia in each segment from segment XIII onwards. Seminal vesicles, four pairs in IX, X, XI, XII. Ovaries, two pairs in XIII and XIV. Testis free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, elongated oval or globular between 13 / 14 - 16 / 17, about 0.3 - 0.4 mm in diameter, two to seven on each side per segment.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
1ED54A23B5607D1B3F79877C70544E99.taxon	distribution	Distribution. The new species was found in the river banks of Phao River which was covered with worm casts.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
1ED54A23B5607D1B3F79877C70544E99.taxon	etymology	Etymology. The species was named after the Phao River, the type locality.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
D19F4D5966A6F9EF119FBC1730D3B83E.taxon	diagnosis	Diagnosis. Glyphidrilus sekongensis sp. n. is a small sized earthworm with the clitellar wings on the lateral side of the body in XXV-XXXI. Clitellum in XVI, XVII-XXXVI, XXXVII. Female pores, male pores and spermathecal pores not visible. Genital markings: paired or asymmetrical on bc in XVI, XVII-XXIV and XXXII. Four pairs of seminal vesicles in IX-XII. Intestinal origin in XVI. Ovaries in XIII-XIV. Spermathecae between 12 / 13 - 15 / 16.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
D19F4D5966A6F9EF119FBC1730D3B83E.taxon	description	Description of holotype. Dimensions: body length 134 mm, diameter 2.4 mm in segment VIII, 2.3 mm before the clitellar wing in segment XXIV, 2.4 mm after wing in segment XXXII in clitellar region; body cylindrical in anterior part, quadrangular in transverse section behind clitellum. 329 segments. Body color pale brown with variations from red to pink on adjacent tissues of wing portions in different individuals of newly collected specimens. The posterior end on the dorsal surface is considerably broader than the ventral. Clitellar wing on ventro-lateral part of clitellum in XXV-XXXI, 4.0 mm in height, and 0.5 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in XVI-XXXVII. Four pairs of setae per segment from II, setal formula aa: ab: bc: cd: dd = 1.0: 0.3: 1.0: 0.5: 1.3 in segment VIII and 1.8: 0.8: 1.3: 0.6: 1.5 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: laterally paired or asymmetrical on bc in XVI-XXIV, XXXII. Septa 5 / 6 - 8 / 9 thicker than between other segments, 9 / 10 - 14 / 15 thick and 15 / 16 to the last others segment, 9 / 10 - 14 / 15 thick and 15 / 16 to the last segment thin. Gizzard small, globular in VIII. Intestine enlarged from XVI. Hearts, four pairs in VIII-XI, first in VIII and last in XI. A pair of holonephridia in each segment from segment XIII onwards. Seminal vesicles, four pairs in IX, X, XI, XII. Ovaries, two pairs in XIII and XIV. Testis free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, small elongated oval or globular between 12 / 13 - 15 / 16, about 0.1 - 0.3 mm in diameter, two to seven on each side per segment.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
D19F4D5966A6F9EF119FBC1730D3B83E.taxon	distribution	Distribution. The new species was found in a stream at Ban Kiangkong, Lamarm, Sekong, Laos. Soils are slightly sandy mixed with black organic matter.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
D19F4D5966A6F9EF119FBC1730D3B83E.taxon	etymology	Etymology. The species was named after Sekong, the type locality in Laos.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
84DAC461D84A960851E34F34C2436402.taxon	diagnosis	Diagnosis. Glyphidrilus namdonensis sp. n. is small sized earthworm with distinctly expanded tissues, the clitellar wings on the lateral side of the body in XXIV, XXV-XXVIII, XXX. Clitellum in XIX, XX-XXXVI, XXXVII, XXXVIII. Female pores, male pores and spermathecal pores not visible. Genital markings: medium paired or asymmetrical on aa in XIII, XIV; paired or asymmetrical on bc in XXII, XXIII, XXIV and XXVII, XXVIII-XXXIII. Four pairs of seminal vesicles in IX-XII. Intestinal origin in XVI. Ovaries in XIII-XIV. Spermathecae between 14 / 15 - 18 / 19.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
84DAC461D84A960851E34F34C2436402.taxon	description	Description of holotype. Dimensions: body length 92 mm, diameter 3.5 mm in segment VIII, 4.0 mm before the clitellar wing in segment XXIII, 4.0 mm after the wing in segment XXXI in the clitellar region; body cylindrical in the anterior part, quadrangular in transverse section behind clitellum. 216 segments. Body color pale brown with variations from red to pink in adjacent tissues of the wing portions in different individuals of newly collected specimens. Clitellar wing on ventro-lateral part of clitellum in XXIV-XXX, 5.0 mm in height, and 0.2 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in XX-XXXVII. Four pairs of setae per segment from II, setal formula aa: ab: bc: cd: dd = 1.3: 0.5: 1.0: 0.5: 2.0 in segment VIII and 1.5: 0.5: 1.0: 0.5: 2.3 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: medium paired on aa in XIII, XIV, laterally paired or asymmetrical on bc in XXII, XXIII. Septa 5 / 6 - 8 / 9 thicker than between other segments, 9 / 10 - 14 / 15 thick, and 15 / 16 to the last segment thin. Gizzard small, globular in VIII. Intestine enlarged from XVI. Hearts, five pairs in VII-XI, first in VII and last in XI. A pair of holonephridia in each segment from segment XII onwards. Seminal vesicles, four pairs in IX, X, XI, XII. Ovaries two pairs in XIII and XIV. Testis free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, small elongated oval or globular between 14 / 15 - 18 / 19, about 0.1 - 0.2 mm in diameter, two to thirteen on each side per segment.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
84DAC461D84A960851E34F34C2436402.taxon	distribution	Distribution. The new species was found only from the banks of Done River at Ban Namdone, Thakhek, Khammouan, Laos.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
84DAC461D84A960851E34F34C2436402.taxon	etymology	Etymology. The species was named after the Done River.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
BEBC94D9DC8FF68804DC4BE7B52753FF.taxon	diagnosis	Diagnosis. Glyphidrilus champasakensis sp. n. is a large sized earthworm with distinct expanded tissues of the clitellar wings on the sides of the body in XXIII, XXIV- 1 / 2 XXXII, XXXII, XXXIII. Clitellum in XIX, XX- XLIX, L, LI, LII. Female pores, male pores and spermathecal pores not visible. Genital markings: medium paired or asymmetrical on aa in XII, XIII, XXXIII, XXXIV; paired or asymmetrical on bc in XVII, XVIII, XIX-XXII, XXIII, XXIV and XXXII, XXXIII, XXXIV, XXXV. Four pairs of seminal vesicles in IX-XII. Intestinal origin in XVI. Large ovaries in XIII-XIV. Spermathecae between 14 / 15 - 19 / 20.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
BEBC94D9DC8FF68804DC4BE7B52753FF.taxon	description	Description of holotype. Dimensions: body length 301 mm, diameter 5.0 mm in segment VIII, 5.0 mm before the clitellar wing in segment XXII, 5.0 mm after wing in segment XXXIII in clitellar region; body cylindrical in anterior part, quadrangular in transverse section behind clitellum. 424 segments. Body color pale brown with variations from red to pink at adjacent tissues of wing portion in different individuals of newly collected specimens. Clitellar wing on ventro-lateral part of clitellum in XXIII-XXXII, 10.0 mm in height, and 0.2 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in XIX- XLIX. Four pairs of setae per segment from II, setal formula aa: ab: bc: cd: dd = 2.0: 1.0: 2.0: 1.0: 3.0 in segment VIII and 2.0: 1.0: 2.0: 1.0: 2.5 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: medium paired or asymmetrical on aa in XIII, XXXIV, laterally paired or asymmetrical on bc in XIX-XXII and XXXIII, XXXIV. Septa 4 / 5 - 8 / 9 thicker than between other segments others segment, 9 / 10 - 11 / 12 thick and 12 / 13 to the last segment thin. Gizzard small, globular in VIII. Intestine enlarged from XVI. Hearts, five pairs in VII-XI, first in VII and last in XI. A pair of holonephridia in each segment from segment XIV onwards. Seminal vesicles, four pairs in IX, X, XI, XII. Ovaries, two pairs in XIII and XIV. Testis free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, small elongated oval or globular between 14 / 15 - 19 / 20, about 0.1 - 0.3 mm in diameter, five to thirteen on each side per segment.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
BEBC94D9DC8FF68804DC4BE7B52753FF.taxon	distribution	Distribution. The new species was found only on the banks of the Mekong River at Ban Khonkhen, Champasak, Laos.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
BEBC94D9DC8FF68804DC4BE7B52753FF.taxon	etymology	Etymology. The new species was named after Champasak, Laos, the type locality.	en	Chanabun, Ratmanee, Inkavilay, Khamla, Panha, Somsak (2017): New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae). ZooKeys 672: 1-34, DOI: http://dx.doi.org/10.3897/zookeys.672.10212, URL: http://dx.doi.org/10.3897/zookeys.672.10212
