taxonID	type	description	language	source
35368F2DFFB8FF9452DCFA14FBA1F980.taxon	type_taxon	Type species: Systenostoma tamlod Panha & Burch, 1999, by original designation.	en	Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László, Schilthuizen, Menno (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). Contributions to Zoology 91: 62-78, DOI: 10.1163/18759866-BjA10025
35368F2DFFB8FF9752DCF9C7FC58FEE3.taxon	materials_examined	Type material: Laos, Bolikhamsai Province, 15 km southeast + 4.5 km northeast (on a side road) towards centre of Lak Sao, Phu Phako, limestone gorge, 510 m a. s. l., 18 ° 06.546 ’ N, 105 ° 03.778 ’ E, (locality code: 2019 / 110), leg. A. Hunyadi, 2 October 2019, holotype (HNHM 104885); CUMZ 7435 / 2 paratypes; SMF 365053 / 2 paratypes; VNMN / 2 paratypes; ZRC. MOL. 23217 / 2 paratypes, HA / 697 paratypes.	en	Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László, Schilthuizen, Menno (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). Contributions to Zoology 91: 62-78, DOI: 10.1163/18759866-BjA10025
35368F2DFFB8FF9752DCF9C7FC58FEE3.taxon	diagnosis	Diagnosis: A strongly depressed-globular Angustopila species with a wide umbilicus, strong spiral sculpture consisting of a series of coarse elevations (flat-topped beads) in a chain-like pattern and four well-developed teeth (1 parietal, 2 palatal, 1 basal).	en	Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László, Schilthuizen, Menno (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). Contributions to Zoology 91: 62-78, DOI: 10.1163/18759866-BjA10025
35368F2DFFB8FF9752DCF9C7FC58FEE3.taxon	description	Description: Shell tiny, whitish, strongly depressed-globular, dorsal part domeshaped, body whorl widest in standard apertural view; protoconch with slightly less than 1.5 whorls and a minute and very uneven pattern of polygonal, rather sharpcrested ridges locally forming minute, conical peaks where ridges split (the structure also reminiscent of cross-sectional trabecular bone); this surface structure forms raised spiral striae on last half of protoconch whorl, and gradually becomes stronger towards first teleoconch whorl; teleoconch bears similar basal sculpture as that of protoconch, strongly ornamented by a pattern of elevated, equidistantly-arranged rows of elegantly linked flat-topped beads; some fine, irregular radial sculpture also visible, but none especially conspicuous due to prominent bead-like striations; whorls 3.25 – 3.5, separated by deep suture; whorls slightly shouldered; aperture oblique to shell axis in lateral view; umbilicus wide, ca. one third of shell width, in living specimens containing several mud granules; aperture narrow, semilunar, sinulus relatively wide, rather strongly separated due to strong parietal tooth; peristome expanded, not reflected; parietal callus strongly protruding, remarkably visible in lateral perspective as projecting beyond peristome and nearly in line with shell periphery, detached from penultimate whorl; parietal tooth strongly developed, long, curved, pointing towards upper palatal tooth; palatal wall with an upper, smaller tooth situated at peristome edge, and a slightly larger, lower tooth situated somewhat deeper, both blunt; columellar / basal, tooth blunt with wide base, positioned on lower part of columellar area, located at similar distance from peristome edge as lower palatal tooth. Soft anatomy unknown. Measurements (in mm): SW = 0.66 – 0.76; SH = 0.49 – 0.58 (n = 19).	en	Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László, Schilthuizen, Menno (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). Contributions to Zoology 91: 62-78, DOI: 10.1163/18759866-BjA10025
35368F2DFFB8FF9752DCF9C7FC58FEE3.taxon	diagnosis	Differential diagnosis: Angustopila coprologos n. sp. can be easily distinguished from all other Angustopila species by its depressed shell, the four apertural denticles and the pronounced sculpture.	en	Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László, Schilthuizen, Menno (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). Contributions to Zoology 91: 62-78, DOI: 10.1163/18759866-BjA10025
35368F2DFFB8FF9752DCF9C7FC58FEE3.taxon	etymology	Etymology: The specific epithet coprologos (Ancient Greek for dung gatherer) refers to the mud granules (here assumed to be faeces) placed on this species’ shell. Used as a noun in apposition.	en	Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László, Schilthuizen, Menno (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). Contributions to Zoology 91: 62-78, DOI: 10.1163/18759866-BjA10025
35368F2DFFB8FF9752DCF9C7FC58FEE3.taxon	distribution	Distribution: This species is known only from the type locality in Bolikhamsay Province, Laos: ca. 13 km southeast (in a straight line) from the town Lak Sao (see also supplementary figs S 1 – S 2).	en	Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László, Schilthuizen, Menno (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). Contributions to Zoology 91: 62-78, DOI: 10.1163/18759866-BjA10025
35368F2DFFB8FF9752DCF9C7FC58FEE3.taxon	biology_ecology	Habitat: Since no living individuals were found (only shells of freshly dead specimens), no observations about the specific habitat of Angustopila coprologos n. sp. could be made. The living snails presumably live deep in limestone crevices close to (or on) root systems. The summer of 2019 was very wet and rainy with extensive flooding throughout the flood plains of the Mekong River. This might have played a role in the vast accumulation of empty shells at the base of rock formations (collection in October 2019).	en	Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László, Schilthuizen, Menno (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). Contributions to Zoology 91: 62-78, DOI: 10.1163/18759866-BjA10025
35368F2DFFBBFF9952FDFE26FD8BFA34.taxon	materials_examined	Type material: Vietnam, Quang Ninh Province, Ha Long Bay, Cap La Cave, 20 ° 51.793 ’ N, 107 ° 13.541 ’ E, soil deposit fallen through roof in pristine cave, vegetation outside cave tall and woody, leg. J. J. Vermeulen & K. Anker, 07 March 2018, holotype (HNHM 104886); CUMZ 7436 / 2 paratypes; SMF 365052 / 2 paratypes; VNMN / 2 paratypes; ZRC. MOL. 23218 / 2 paratypes, jjV 17633 / 409 paratypes.	en	Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László, Schilthuizen, Menno (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). Contributions to Zoology 91: 62-78, DOI: 10.1163/18759866-BjA10025
35368F2DFFBBFF9952FDFE26FD8BFA34.taxon	diagnosis	Diagnosis: An Angustopila species with a depressed-globular shell with dome-shaped spire, thick spiral striae, kidney-shaped aperture with single parietal denticle not reaching parietal callus.	en	Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László, Schilthuizen, Menno (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). Contributions to Zoology 91: 62-78, DOI: 10.1163/18759866-BjA10025
35368F2DFFBBFF9952FDFE26FD8BFA34.taxon	description	Description: Shell tiny, off-white, depressed-globular form with domed spire; body whorl widest in standard apertural view; protoconch with 1.5 whorls and a minute and very uneven pattern of polygonal, rather sharp-crested ridges locally forming minute, conical peaks where ridges split (the structure also reminiscent of cross-sectional trabecular bone), no spiral striation discernible; teleoconch ornamented by some weak radial growth lines and much stronger, ropelike, equidistantly-arranged spiral striae (ca. 14 – 15 on body whorl from standard apertural view); the 3.25 – 3.5 whorls are separated by a deep suture; whorls slightly shouldered; aperture oblique to shell axis in lateral view; umbilicus relatively wide, less than one third of shell width; aperture kidney-shaped with strongly concave parietal side; sinulus wide, weakly separated due to weak parietal tooth; peristome expanded, not reflected; parietal callus strongly protruding, but in line with curvature of penultimate whorl and beyond aperture edge (profile) in lateral view (fig. 3 C), detached from penultimate whorl; parietal tooth weak, low, short, rounded, does not reach peristome. Soft anatomy unknown. Measurements (in mm): SW = 0.6 – 0.68, SH = 0.46 – 0.57 (n = 24).	en	Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László, Schilthuizen, Menno (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). Contributions to Zoology 91: 62-78, DOI: 10.1163/18759866-BjA10025
35368F2DFFBBFF9952FDFE26FD8BFA34.taxon	diagnosis	Differential diagnosis: Angustopila pallgergelyi Dumrongrojwattana, Chuenit & Wongkamhaeng, 2021 is similar in shell and aperture shape, but is larger, has a prominent palatal tooth (absent in Angustopila psammion n. sp.) and a stronger parietal tooth. Angustopila coprologos n. sp. is slightly larger, has a rough shell surface, and has an additional subcolumellar tooth and two palatal teeth.	en	Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László, Schilthuizen, Menno (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). Contributions to Zoology 91: 62-78, DOI: 10.1163/18759866-BjA10025
35368F2DFFBBFF9952FDFE26FD8BFA34.taxon	etymology	Etymology: The specific epithet (ΨαμμϯΟν) means a grain of sand in Greek and is used as a noun in apposition.	en	Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László, Schilthuizen, Menno (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). Contributions to Zoology 91: 62-78, DOI: 10.1163/18759866-BjA10025
35368F2DFFBBFF9952FDFE26FD8BFA34.taxon	distribution	Distribution: This new species is known only from the type locality, Cap La Cave, Ha Long Bay, Quang Ninh Province, Vietnam.	en	Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László, Schilthuizen, Menno (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). Contributions to Zoology 91: 62-78, DOI: 10.1163/18759866-BjA10025
35368F2DFFBBFF9952FDFE26FD8BFA34.taxon	biology_ecology	Habitat: Angustopila psammion n. sp. was found in large numbers in small sediment deposits along the walls of a dry cave (Cap La Cave), in complete darkness. We assume that the sediment had fallen in through crevices in the rock, because it contains a species-rich assemblage of bleached, opaque shells of surface-dwelling terrestrial gastropods. Shells of Angustopila psammion n. sp., however, look fresh and are probably autochthonous.	en	Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László, Schilthuizen, Menno (2022): The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae). Contributions to Zoology 91: 62-78, DOI: 10.1163/18759866-BjA10025
