taxonID	type	description	language	source
35378770FFC90D73C8EAFDA401D73031.taxon	materials_examined	Type species: Ochlerotatus macfarlanei Edwards, 1914. Females Head: Vertex with decumbent scales narrow, curved, pale and dark; erect forked scales numerous on occiput and vertex; ocular line narrow, with narrow, curved, pale scales; normally 4 or 5 interocular setae (occasionally 6 in Co. elsiae elsiae); eyes contiguous or separated by less than width of 1 eye facet; antennal pedicel with small, broad, pale scales on mesal surface; clypeus bare; maxillary palpus dark-scaled with apex pale-scaled; proboscis longer than forefemur, dark-scaled with patch of pale scales on ventral and lateral surfaces near midlength (Assemblage A) or entirely dark-scaled (Assemblage B). Thorax: Scutum covered with narrow, curved scales except for bare median prescutellar area, scales sparse on scutal fossa, scales dark except for narrow, pale-scaled stripes on acrostichal (anterior and posterior) area, dorsocentral (anterior and posterior) area, outer and posterior margins of scutal fossa, outer margins of prescutellar area mesad of setae, and longitudinal stripe on supraalar area; setae well developed on anterior promontory, acrostichal (anterior and posterior) area, antedorsocentral area, dorsocentral (anterior and posterior) area, scutal fossa, antealar and supraalar areas, and lateral margins of prescutellar area; scutellum with narrow or broad scales on midlobe, narrow scales on lateral lobes; antepronota widely separated, with broad, pale scales, several setae; postpronotum with scales, few posterior setae; prespiracular setae absent; hypostigmal area bare; postspiracular area with scales, several setae; subspiracular area with pale scales (Co. banksi without or with few scales, Co. pseudotaeniatus without scales); upper proepisternum with pale scales, several setae, lower area bare; paratergite narrow, with pale scales; mesokatepisternum with broad, pale scales in upper and lower patches, several upper and posterior setae; prealar area with broad, pale scales on upper (absent in Co. banksi) and lower areas, numerous upper setae; mesepimeron with 1 patch of broad, pale scales on upper area and extending onto middle, several upper setae, lower setae absent. Wing: Dark-scaled with elongate, narrow, pale-scaled patch at or near base of anterior margin of costa; upper calypter with several setae on margin; alula with row of narrow, dark scales on margin; remigium with 2 or 3 setae distally on dorsal surface. Legs: Midfemur with narrow, longitudinal stripe of white-scales from base to or near apex on middle of anterior surface; hindfemur with subapical white scales but apex fringed with dark scales; hindtarsomeres 1 and 2 with narrow, white-scaled, basal and apical bands; fore- and midtarsi with 2 equal ungues, each with 1 tooth; hindtarsi with 2 equal ungues, simple. Abdomen: Tergum I with pale-scaled patch on lateral tergite; terga II – V with narrow, pale-scaled, basal bands. Genitalia: Tergum VIII with apex broadly rounded, numerous broad scales on approximately distal 0.30 – 0.40; sternum VIII sclerotized, width greater than length, apical margin with moderate, median emargination separating broadly rounded lobes (Co. hatorii and Assemblage B with apex nearly straight), numerous short, lanceolate scales and occasionally 1 – 3 broad scales (Co. hatorii and Assemblage B without lanceolate scales, but with several to numerous broad scales); seta 2 - S inserted posterior to 1 - S; tergum IX comprising single, moderately to heavily pigmented, elongate plate with median emargination apically in Assemblage A or with plate short and wide in Assemblage B, several setae on each side of midline distally; postgenital lobe relatively long, narrow, apex normally flat or with very shallow, median emargination (Co. hatorii and Assemblage B with postgenital lobe somewhat shorter and wider), several setae distally; upper vaginal sclerite small to moderate size; lower vaginal sclerite absent; insula liplike, with several setae in lateral patches; cercus moderately long, moderately wide throughout length, apex broadly rounded, scales absent (Assemblage B with cercus somewhat shorter and with scales); 3 spermathecal capsules. Males Head: Maxillary palpus with 5 palpomeres, slightly shorter than proboscis, palpomeres 4 and 5 slightly down-turned, palpomeres 3 (distal part) and 4 with moderate number of long setae ventrally; antennal flagellar whorls with numerous long setae directed dorsally and ventrally. Legs: Fore- and midtarsi with 2 unequal ungues, larger unguis with 2 teeth, smaller unguis with 1 tooth; hindtarsus with 2 equal ungues, simple. Genitalia: Tergum IX with poorly sclerotized, moderate lobe bearing several moderately long, slender setae on each side of midline of posterior margin; gonocoxite moderately long, relatively narrow throughout length, dorsal surface without apical and basal lobes on mesal margin, with few moderately long and long, relatively slender setae scattered over dorsal and ventral surfaces, mesal surface membranous, lateral, outer part of dorsal and most of ventral surfaces with broad scales; gonostylus attached at apex of gonocoxite, long, narrow, slightly curved, gonostylar claw short, narrow, attached at apex, 1 or more shorter, stout setae subapically (these similar to gonostylar claw but somewhat shorter and thinner, especially in Co. hatorii) (Assemblage A) or subapical, stout setae absent (Assemblage B); aedeagus simple, tubelike, distal part wider than middle (Assemblage A) or approximately same width as middle (Assemblage B); proctiger with few tiny, cercal setae; claspette with single, moderately long, columnar stem with small subapical, thumblike lobe (Assemblage A) or subapical lobe absent (Assemblage B), claspette filament attached apically, long, relatively narrow, strongly curved, apex pointed; sternum IX with several setae on posteromesal area. Pupae Cephalothorax: Setae 1,3 - CT similarly developed; 11 - CT normally single, rarely 2 - branched. Abdomen: Seta 3 - I, II branched (Assemblage A) or normally single (Assemblage B), longer than 6 - I, II; 6 - I shorter than 7 - I; 1 - II with several branches; 5 - II inserted lateral to 4 - II; 3 - III single (Co. banksi single or 2 - branched); 5 - V long, stout, single, longer than tergum VI median length; 2 - VI inserted mesal to 1,3 - VI (Assemblage A) or 2 - VI inserted lateral to 1 - VI and mesal to 3 - VI (Assemblage B); 6 - VII inserted posterior and mesal to 9 - VII; 9 - VIII inserted slightly anterior to posterolateral corner (Assemblage A) or on corner (Assemblage B), with multiple branches. Paddle: Apical margin rounded; midrib extends to apex of paddle; without hairlike spicules on margins; seta 1 - Pa short, single. Fourth-instar larvae Head: Seta 1 - C slender, single, apex attenuate; 4 - C short, slender, with few branches, inserted at approximately same level or slightly anterior to 5,6 - C; 5,6 - C relatively short, moderately stout, with several branches, 6 - C lateral to 5 - C, 4 – 6 - C all at approximately same level; 7 - C moderately long, relatively slender, with several branches, inserted lateral and posterior to 4 – 6 - C; 12 - C short, slender, with several branches, inserted mesal to 13 - C; 13 - C single, longer than 12 - C; 14 - C short, stout, normally single or 2 - branched, occasionally 3 - branched; 19 - C absent; antenna moderately long, narrow, with several to numerous spicules, seta 1 - A moderately long, normally with 2 or 3 branches (rarely single). Thorax: Setae 1 – 3 - P on common setal support plate; 4 - P single; 8 - P branched, shorter than 4 - P; 1 - M, T stout, single or 2 - branched, inserted on large, pigmented tubercle (Assemblage A) or 1 - M, T slender, with few branches, not on tubercle (Assemblage B); 5 - M longer than 7 - M; 2,6 - T single (Lu & Ji, 1997: fig. 44, of Co. ningheensis shows 2 - T with 3 branches). Abdomen: Seta 6 - I, II long, with 2 branches, 6 - II longer than 6 - III; 7 - I, II long, single (rarely 2 - branched); 12 - I present; 8 - II short, single or 2 - branched; 1 - VII long; 12 - VII single; 2,4 - VIII single; comb comprising numerous scales in patch; segment X with moderate saddle, incomplete ventrally, with numerous stout spicules on posterior margin dorsal to seta 1 - X, acus absent, 1 - X inserted on saddle, ventral brush with several long, branched setae with short stems, attached to grid with both transverse and lateral bars, some species with 1 shorter seta anterior to grid. Siphon: With attached acus; pecten with numerous closely set spines; seta 1 - S single or branched, inserted within distal part of pecten or slightly distal to pecten. Included species Assemblage A: Collessius elsiae elsiae (Barraud), Co. elsiae vicarius (Lien), Co. hatorii (Yamada), Co. macdougalli (Edwards), Co. macfarlanei (Edwards), Co. ningheensis (Lei) and Co. shortti (Barraud). Assemblage B: Collessius banksi (Edwards), Co. pseudotaeniatus (Giles) and Co. tonkinensis (Galliard & Ngu). Distribution Assemblage A: China, India, Indonesia, Japan, Malaysia, Pakistan, Sri Lanka, South Korea, Taiwan, Thailand and Vietnam. Assemblage B: Bangladesh, India, Myanmar, Nepal, Philippine Islands, Sri Lanka and Vietnam. Bionomics Assemblage A: Immatures are normally found in rock pools but also in artificial containers (e. g. concrete water tanks). Females of some species bite humans indoors and outdoors. Assemblage B: Immatures have been collected normally from rock pools but also from tree holes, and artificial containers (e. g. cement sinks and gutters, drains, iron cisterns, tires, etc.). Females of Co. pseudotaeniatus bite humans indoors and outdoors. Discussion Species of Collessius can be grouped into Assemblages based on morphological characters (see above). Barraud (1934: figs 37, 39) illustrated the development of the gonostylus and fourth-instar larva of Co. elsiae and Co. shortti. See Tanaka et al. (1979: figs 97, 98, 219) for illustrations of the male genitalia, fourthinstar larva and adults of Co. hatorii, Tanaka (2002: fig. 1) for the pupa and Moriya et al. (1973) for the egg of this species. Etymology Collessius is named in honour of Dr Donald H. Colless in recognition of his contributions to the taxonomy of Oriental mosquitoes. The generic name is masculine, formed from his surname and the masculine Latin suffix ‘ - ius ’. Recommended abbreviation = Co.	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFB70D71C8E6FB7A0286364F.taxon	materials_examined	Type species: Culex geniculatus Olivier, 1791. Females Head: Median area of vertex with narrow, curved, pale scales; occiput and vertex with numerous erect forked scales; ocular line narrow, with narrow, white scales; eyes above antennal pedicels separated by width of 2 – 4 ocular facets; antennal pedicel with few short, fine setae, few small, broad scales sometimes present on mesal surface; clypeus bare; maxillary palpus and proboscis dark-scaled, proboscis longer than forefemur. Thorax: Scutum with narrow, curved scales; pale scales forming narrow, longitudinal stripe on anterior and posterior acrostichal area and extending posteriorly to or near prescutellar area, posterior dorsocentral area extending posteriorly laterad of prescutellar setae to posterior margin of scutum, and on anterior and lateral margins of prescutellar bare area; white scales on antealar area and extending posteriorly as narrow, longitudinal stripe on supraalar area; scutal fossa covered with pale scales; acrostichal (anterior and posterior) and dorsocentral (anterior and posterior) areas with setae; scutellum with narrow, curved scales on lateral lobes; paratergite with broad, white scales; antepronota widely separated, with broad and few moderately broad, pale scales; postpronotum with broad, pale and dark scales, Da. echinus normally also with few narrow, curved scales dorsally; broad, white scales on postspiracular area, subspiracular area, upper proepisternum, mesokatepisternum (upper and lower posterior patches), upper and lower prealar areas, mesepimeron (1 large patch); mesepimeron without lower setae. Wing: Dark-scaled; upper calypter with several setae on margin; alula with row of narrow scales on margin; remigium with 2 or 3 setae on dorsal surface distally. Legs: Ante- and postprocoxal membranes bare; hindfemur with well-developed apical, white-scaled band; hindtibia and tarsus dark-scaled, ungues equal, simple; fore- and midungues equal, each with 1 tooth. Abdomen: Tergum I with pale-scaled patch on laterotergite; segment VII laterally compressed. Genitalia: Tergum VIII heavily pigmented, basolateral corners expanded into short, proximally curved, thumblike projections, apex broadly rounded, numerous broad scales on distal 0.56 – 0.58; sternum VIII heavily pigmented, base straight or very slightly concave mesally, apex noticeably wider than base, nearly straight but with shallow, median emargination, broad scales absent or very few (Da. geniculata) or several scales forming lateral patches (Da. echinus), 2 moderately long setae slightly subapical on lateral part of apical margin, width greater than length; tergum IX heavily pigmented with median area slightly lighter, apex with moderately deep, V-shaped, median emargination, about 6 moderately long, stout setae apically on each side of midline; postgenital lobe moderately long, relatively narrow, apex rounded, setae on distal part; upper vaginal sclerite moderately large to large, heavily pigmented; lower vaginal sclerite absent; insula liplike, with few setae in lateral patches; cercus relatively long, moderately wide, apex broadly rounded, without scales; 3 spermathecal capsules. Males Head: Maxillary palpus with 5 palpomeres, slightly shorter than proboscis, dark-scaled, numerous long and moderately long setae ventrolaterally on distal part of palpomere 3 and ventrolaterally on palpomeres 4 and 5; antennal flagellomeres with numerous long setae directed dorsally and ventrally. Legs: Foretarsus with 2 unequal ungues, larger unguis with 2 teeth, smaller with 1 tooth; midtarsus with 2 unequal ungues, each with 1 tooth; hindtarsus with 2 equal ungues, simple. Genitalia: Tergum IX comprising 2 relatively broad, lateral plates each with short, narrow, strongly sclerotized lobe on posteromesal area bearing 2 – 6 short, stout, flattened, dark setae; gonocoxite moderately long, relatively narrow, dorsal surface without apical and basal lobes on mesal margin, ventral surface without specialized setae or scales, mesal surface membranous, numerous scales on dorsal, lateral and ventral surfaces; gonostylus attached at apex of gonocoxite, relatively long, narrow, distal part gently curved mesally, single gonostylar claw, long, narrow, attached at apex; aedeagus simple, tubelike, lateral margins gently curved to apex, distal 0.50 narrowest; proctiger with few to several cercal setae; claspette with single moderately long to long, narrow, columnar stem, claspette filament attached apically, long, somewhat flattened, slightly curved to narrow, pointed apex; sternum IX with several setae on posteromedian area. Pupae (based on Da. geniculata) Cephalothorax: Setae 1,3 - CT similarly developed, normally with 2 branches; 11 - CT single. Abdomen: Seta 3 - I single, shorter than 6 - I; 6 - I longer than 7 - I; 2 - II inserted lateral to 1,3 - II; 3 - II moderately long, stout, normally single (occasionally 2 - forked), shorter than 6 - II; 5 - II long, stout, single, noticeably longer than 3 - II, slightly shorter than 6 - II, inserted lateral to 4 - II; 3,6 - III long, stout, single; 5 - V long, stout, single, longer than tergum VI median length; 2 - VI inserted anterior and slightly lateral to 1 - VI and mesal to 3 - VI; 6 - VII short, slender, inserted posterior and slightly mesal to 9 - VII; 9 - VII moderately long, stout, aciculate, usually with 2 branches; 9 - VIII moderately long, stout, aciculate, with 2 – 5 (usually 2 or 3) branches. Paddle: Length noticeably greater than width; midrib well developed, reaching apex of paddle; without hairlike spicules on margins; seta 1 - Pa short, slender, single. Fourth-instar larvae Head: Seta 1 - C single, moderately stout; 4 - C moderately long, slender, multiple-branched, inserted mesal and slightly anterior to 6 - C; 5 - C long, stout, single, inserted posterior to 4,6,7 - C; 6 - C moderately long to long, stout, single to 3 - branched, inserted anterior and approximately same lateral level to 5 - C and mesal and slightly anterior to 7 - C; 7 - C moderately long, moderately stout, with 2 or 3 branches; 12 - C branched, inserted mesal to 13 - C; 13 - C single, longer than 12 - C; 14 - C short, branched; 19 - C absent; antenna moderately long, narrow, without spicules, seta 1 - A moderately long, single or 2 - branched. Thorax: Setae 1 – 3 - P inserted on common setal support plate; 5 - P with 2 branches, longer than 6 - P; 8 - P short, multiple-branched; 4 - M single; 5 - M longer than 7 - M; 2,6 - T single. Abdomen: Seta 6 - I, II with 2 long, stout, aciculate branches; 7 - I, II long, stout, single; 12 - I present; 8 - II single; 1 - VII moderately long, stout, stellate, multiplebranched; 12 - VII single; 2,4 - VIII single; segment VIII with comb comprising several scales in single curved row; segment X with large saddle, incomplete ventrally, acus absent, seta 1 - X long, stout, branched, inserted on saddle, ventral brush with several long, normally 2 – 4 - branched setae with long stems, inserted on grid with both transverse and lateral bars, 2 precratal setae. Siphon: With attached acus; pecten with several, closely set spines; seta 1 - S moderately long, multiplebranched, inserted distal to pecten. Included species Dahliana echinus (Edwards), Da. geniculata (Olivier) and Da. gilcolladoi (Sanchez-Covisa Villa, Rodriguez Rodriguez & Guillen Llera). Distribution Albania, Algeria, Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, France, European Russia, Finland, Germany, Greece, Hungary, Iran, Italy, Lithuania, Macedonia, Morocco, Netherlands, Norway, Poland, Portugal, Romania, Slovenia, Slovakia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom and Yugoslavia. Bionomics Immature stages normally inhabit treeholes but have been found in open tree stumps, root holes of olive trees and pools in rocks. Discussion See discussion under Classification of Evaluated Taxa. Marshall (1938: figs 12, 57 (g), 69, 94 and pl. IV) provided illustrations of the egg, first- and fourth-instar larvae, male genitalia and female of Da. geniculata, and Reinert (2002 e: fig. 3) illustrated the female genitalia of this species. Villa, Rodriguez & Llera (1985) provide photographs of the fourth-instar larvae of Da. echinus, Da. geniculata and Da. gilcolladoi. Etymology Dahliana is named in honour of Dr Christine Dahl in recognition of her many important contributions to the taxonomy and bionomics of Palaearctic mosquitoes. The generic name is feminine, formed from her surname and the feminine Latin suffix ‘ - iana ’. Recommended abbreviation = Da.	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFB00D74CB65FB8D00343361.taxon	description	Females Thorax: Antealar scales pale; paratergite without scales; postpronotum with broad scales; postspiracular area with patch of broad, white scales. Legs: Postprocoxal membrane without scales; hindtarsomere 2 with narrow, apical, pale-scaled band. Abdomen: Tergum I without scales on laterotergite. Males Head: Maxillary palpus with few short setae ventrally on distal part of palpomere 3 and ventrolaterally on palpomere 4, palpomeres 3 and 4 not down-turned. Pupae	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFB10D7AC8FBF9DB01443679.taxon	materials_examined	Type species: Culex fluviatilis Lutz, 1904. Females Thorax: Antealar scales dark; paratergite with broad, pale scales (rarely absent?); postpronotum with narrow, curved scales dorsally; postspiracular area without scales. Legs: Postprocoxal membrane with pale scales; hindtarsomere 2 without apical, pale-scaled band. Abdomen: Tergum I with broad, pale scales on laterotergite. Males Head: Maxillary palpus with several to numerous moderate to long setae ventrally on distal part of palpomere 3 and ventrolaterally on palpomere 4, palpomeres 3 and 4 down-turned. Pupae Abdomen: Seta 5 - II inserted lateral to 4 - II; 6 - II longer than 7 - II; 6 - III-V relatively long; 5 - IV-VI very long, stout, each noticeably longer than following tergum; 2 - VI, VII inserted mesal to 1 - VI, VII; 8 - VI, VII inserted dorsally. Fourth-instar larvae Head: Seta 5 - C moderately long, multiple-branched. Thorax: Setae 1 – 3 - P and 5,6 - P inserted on common setal support plates; 4 - M branched. Abdomen: Seta 7 - II short, developed differently than 7 - I; 13 - III – V long, single; 1 - VII long, stout, aciculate, with 2 branches. Siphon: Pecten with spines evenly spaced; seta 1 - S inserted slightly distal to pecten. Included species Georgecraigius fluviatilis. Distribution Argentina (Misiones), Bolivia (Cochabamba), Brazil (Bahia, Goias, Minas Gerais, Para, Rio de Janeiro, Sao Paulo, Sergipe), Colombia (Boyaca, Cundinamarca, Meta), Costa Rica (Alajuela, Cartago, San Jose), Ecuador (Zamora), French Guiana (Guyane, Inini), Guyana (Essequibo), Honduras (Colon), Mexico (Veracruz), Nicaragua (Zelaya), Panama (Canal Zone, Chiriqui, Cocle, Colon, Darien, Panama), Surinam (Suriname) and Venezuela (Aragua, Distrito Federal, Territorio Amazonas). Bionomics Immature stages usually are collected from rockholes, rock pools and stream pools. They are found occasionally in artificial containers, ground pools and treeholes. Etymology Subgenus Horsfallius is named in honour of Dr William Robert Horsfall in recognition of his outstanding contributions to the biology of New World mosquitoes and descriptions of the eggs of aedine species of the Nearctic Region. The name is masculine, formed from his surname and the masculine Latin suffix ‘ - ius ’. Recommended abbreviation = Hor.	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFBE0D7BC893FDBD000232AD.taxon	materials_examined	Type species: Culex pulchriventer Giles, 1901. Females Head: Vertex with decumbent scales narrow and curved, several broad scales present or absent on anterolateral area; erect forked scales numerous on occiput and vertex; ocular line narrow, with narrow, curved, pale scales; 4 or 5 interocular setae; eyes separated by approximately width of 3 ocular facets; antennal pedicel with small, broad, pale scales on mesal surface; clypeus bare; maxillary palpus dark-scaled, with or without pale scales on subbasal area; proboscis longer than forefemur, dark-scaled. Thorax: Scutum covered with narrow, curved scales except bare median prescutellar area; scales pale on following: combined anterior acrostichal and anterior dorsocentral areas, moderate stripe on posterior acrostichal area, narrow stripe on posterior dorsocentral area, narrow stripe on outer margins mesad of setae on prescutellar area, and moderate, longitudinal stripe on supraalar area; acrostichal (anterior and posterior) and dorsocentral (anterior and posterior) setae short; prescutellar area with several setae laterally; scutellum with narrow, curved scales on all lobes, median lobe also with or without few broad scales; antepronota widely separated, with broad, pale scales, several setae; postpronotum with scales, few posterior setae; prespiracular setae absent; hypostigmal area with broad, pale scales; postspiracular area with pale scales, several setae; subspiracular area with broad, pale scales; upper proepisternum with broad, pale scales, several setae, lower area bare; paratergite with pale scales; mesokatepisternum with upper and lower patches of broad, pale scales, upper and posterior setae; prealar area with broad, pale scales on upper and lower areas, numerous upper setae; mesepimeron with 2 patches of broad, pale scales, 1 on upper and 1 on middle areas, several upper setae, lower setae absent. Wing: Dark-scaled, with or without small, pale-scaled patch on anterobasal area of costa; upper calypter with several setae on margin; alula with row of narrow, dark scales on margin; remigium with 2 or 3 setae distally on dorsal surface. Legs: Femora without subapical pale-scaled bands; midfemur without median, longitudinal white-scaled stripe on anterior surface; hindfemur with apex with white scales; tibiae and tarsi dark-scaled; fore- and midtarsi with 2 equal ungues, each with 1 tooth; hindtarsi with 2 equal ungues, simple. Abdomen: Tergum I with pale-scaled patch on laterotergite; terga dark-scaled with basal pale-scaled areas; segment VII dorsoventrally flattened. Genitalia: Tergum VIII with apex broadly rounded, nearly covered with broad scales; sternum VIII sclerotized, width slightly greater than length, apical margin with small, median emargination, several broad scales on lateral areas, seta 2 - S inserted posterior to 1 - S; tergum IX comprising 2 moderately pigmented, lateral plates connected mesally by lightly pigmented area, several short setae distally on each plate; postgenital lobe relatively long, moderately wide, apex flat, several setae distally; upper vaginal sclerite small to moderate size; lower vaginal sclerite absent; insula liplike, few setae in lateral patches; cercus moderately long, moderately wide throughout length, apex broadly rounded, numerous broad scales on dorsal surface; 3 spermathecal capsules. Males Head: Maxillary palpus with 5 palpomeres, shorter than proboscis, palpomeres 4 and 5 slightly downturned; antennal flagellar whorls with numerous long setae directed dorsally and ventrally. Legs: Fore- and midtarsi with 2 unequal ungues, larger unguis with 2 teeth, smaller unguis with 1 tooth; hindtarsus with 2 equal ungues, simple. Genitalia: Tergum IX with median part relatively narrow with bilobed, pincushion-like structure bearing numerous, short to moderately long, very slen- der, curved setae, lateral area developed as large, oblong lobe projecting cephalad; gonocoxite moderately long, dorsal surface without apical and basal lobes on mesal surface, middle part of dorsal surface wide with mesal area broad, curved, projecting ventrally with row of short, stout, closely set, comblike setae on mesal margin, much of dorsal surface covered with short to moderately long, very slender setae, ventral surface with elongate patch of numerous, long, moderately wide, fusiform scales on most of distal 0.50 of mesal margin, mesal surface membranous, lateral, outer part of dorsal and most of ventral surfaces with broad scales; gonostylus attached at apex of gonocoxite, long, narrow but proximal part somewhat thicker, somewhat sigmoidshaped, single gonostylar claw short, curved, spiniform; aedeagus simple, tubelike; proctiger with few tiny, cercal setae; claspette with single, short to moderately long, narrow, columnar stem with small subapical lobe, claspette filament attached apically, long, curved, relatively slender throughout length, apex pointed; sternum IX with few setae (Gi. pulchriventer also with 2 or 3 broad scales) on posteromesal area. Pupae Cephalothorax: Setae 1,3 - CT similarly developed; 11 - CT single, longer than 10 - CT. Abdomen: Seta 3 - I, II single, longer than 6 - I, II; 6 - I shorter than 7 - I; 1 - II long, with 3 branches; 5 - II inserted lateral to 4 - II; 3 - III relatively short, single, approximately equal to 5 - III length; 5 - V moderately long, stout, single, shorter than tergum VI median length; 2 - VI inserted mesal to 1,3 - VI; 6 - VII single, inserted slightly posterior and mesal to 9 - VII; 9 - VII with 3 branches; 9 - VIII inserted on posterolateral corner, with multiple branches. Paddle: Apical margin rounded; midrib extends to apex of paddle; without hairlike spicules on margin; seta 1 - Pa short, single. Fourth-instar larvae Head: Seta 1 - C slender, single, apex attenuate; 4 - C short, with few slender branches, inserted mesal and slightly posterior to 6 - C; 5 - C moderately long, stout, lightly aciculate, with several branches, inserted mesal to 6 - C; 6 - C moderately long, stout, with 3 or 4 aciculate branches, inserted lateral and approximately same level to 5 - C; 4 – 6 - C inserted noticeably anterior to 7 - C; 7 - C moderately long, moderately stout, lightly aciculate, with 4 or 5 branches; 12 - C short, with 2 or 3 slender branches, inserted mesal to 13 - C; 13 - C single, longer than 12 - C; 14 - C short, moderately stout, single or 2 - branched; 19 - C absent; antenna moderately long, narrow, with several small spicules, seta 1 - A moderately long, with 3 or 4 (rarely 2) slender branches. Thorax: Setae 1 – 3 - P on common setal support plate, 3 - P multiple-branched; 4 - P relatively long; 5 - P long, stout, with 2 or 3 aciculate branches; 8 - P short, stellate, with 5 or 6 branches, shorter than 4 - P; 4 - M single; 5 - M longer than 7 - M; 2,6 - T single. Abdomen: Seta 6 - I, II long, stout, aciculate, with 2 branches, 6 - II longer than 6 - III; 7 - I, II long, stout, aciculate, single; 12 - I present; 8 - II short, with 2 branches; 1 - VII long, stout, aciculate, single; 12 - VII single; 2,4 - VIII single; segment VIII with comb comprising numerous scales in patch; segment X with moderate saddle, incomplete ventrally, with numerous stout spicules on posterior margin dorsal to seta 1 - X, acus absent, 1 - X moderately long to long, stout, aciculate, single, inserted on saddle, ventral brush with several long, 2 – 5 - branched setae with long stems, attached to grid with both transverse and lateral bars, anterior few setae somewhat shorter. Siphon: With attached acus; pecten with numerous closely set spines; seta 1 - S relatively long with several aciculate branches, inserted very slightly distal to pecten. Included species Gilesius alius (Lien) and Gi. pulchriventer (Giles). Distribution China, India, Nepal, Pakistan and Taiwan. Bionomics Immature stages inhabit rock pools and treeholes but also have been collected in fresh water ground pools, a seepage pool, mud pools, barrels, tree stump holes, and holes in a fallen tree. Females of Gi. pulchriventer have been collected from humans at dusk. Discussion See Lien (1968 b: figs 1, 6) for illustrations of the male genitalia, pupa and fourth-instar larva of Gi. alius. Etymology Gilesius is named in honour of Dr (Lt-Col) George Michael James Giles in recognition of his contributions to the taxonomy of mosquitoes, primarily of India and surrounding areas. The generic name is masculine, formed from his surname and the masculine Latin suffix ‘ - ius ’. Recommended abbreviation = Gi.	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFBF0D79CB72F8F907333242.taxon	materials_examined	Type species: Finlaya gilli Barraud, 1924. Females Head: Vertex with decumbent, narrow, curved, goldenwhite to white scales mesally, broad, golden-white to white scales laterally; occiput with numerous erect forked scales; clypeus bare; maxillary palpus and proboscis dark brown-scaled, proboscis approximately equal to forefemur length. Thorax: Scutum with narrow, curved scales, goldenwhite scales covering approximately anterior 0.60 of scutum, similar scales forming narrow stripe on posterior acrostichal area extending from anterior patch to anterior part of prescutellar area where it forks and extends along lateral margins of prescutellar bare area, similar narrow stripe on posterior dorsocentral area extending posteriorly along lateral margin of prescutellar setae, longitudinal stripe on supraalar area, scutal fossa with small, posteromedian patch of white scales, prealar area with small patch of somewhat broader, narrow, white scales; setae on posterior dorsocentral area; scutellum with narrow, curved, golden-white scales on all lobes; paratergite without scales; antepronota widely separated, with numerous moderately broad to broad, golden-white scales; postpronotum with golden-white scales, broad scales on most of area except narrow, curved scales dorsally; broad, white scales on upper proepisternum, mesokatepisternum (upper and lower posterior patches), lower prealar area and mesepimeron (single large patch), other areas without scales; mesepimeron without lower setae. Wing: Dark-scaled; upper calypter with several setae on margin; alula with row of narrow, dark scales on margin; remigium with 2 or 3 setae on dorsal surface distally. Legs: Ante- and postprocoxal membranes without scales; hindfemur with apex dark-scaled; hindtibia dark-scaled with basal band of white scales, band narrow dorsally, broader on anterior, posterior and ventral surfaces; hindtarsomeres 1 – 3 with broad, basal bands of white scales, hindungues equal, simple; fore- and midungues equal, each with 1 tooth. Abdomen: Tergum I with white-scaled patch on laterotergite; segment VII laterally compressed. Genitalia: Tergum VIII with broad scales covering all but narrow basal strip; sternum VIII sclerotized, slightly wider than long, several broad scales on proximal area, apical margin nearly straight with several moderately stout setae decreasing in length mesally, seta 2 - S inserted posterior to 1 - S; tergum IX comprising lightly to moderately pigmented sclerite with relatively deep, median, apical emargination separating pair of narrow lobes each bearing 3 or 4 short, slender setae; postgenital lobe relatively long and narrow, apex rounded, several setae distally; upper vaginal sclerite small, lightly pigmented; lower vaginal sclerite absent; insula liplike, with few setae in lateral patches; cercus relatively moderately long, moderately wide, apex rounded, mesal margin gently concave, without scales; 3 spermathecal capsules. Males Head: Maxillary palpus with 5 palpomeres, 0.86 – 0.95 length of proboscis, dark-scaled, distal part of palpomere 3 bearing several long setae ventrolaterally, palpomeres 4 and 5 short, slightly down-turned, with several short and moderately long setae primarily on ventral and distal areas. Legs: Fore- and midtarsi with 2 unequal ungues, larger unguis with 2 teeth, smaller unguis with 1 tooth; hindungues equal, simple. Genitalia: Tergum IX comprising 2 moderately pigmented, lateral plates, each with small, short, narrow, heavily sclerotized lobe on posterior margin bearing 5 or 6 short, stout setae; gonocoxite moderately long, relatively narrow, dorsal surface without apical and basal lobes on mesal margin, several scales present, ventral surface without specialized setae or scales, mesal surface membranous; gonostylus long, attached at apex of gonocoxite, narrow but distal part narrower, nearly straight, with single relatively short, narrow, gonostylar claw at apex; aedeagus simple, tubelike, moderately long, relatively narrow; proctiger with distal part heavily pigmented, apex flattened with 2 or 3 short, stout teeth, cercal setae present; claspette with single long, narrow columnar stem, claspette filament attached at apex of stem, long, relatively narrow, slightly flattened, distal part terminating in long, slender, pointed apex; sternum IX relatively short, with several setae on posteromedian area. Pupae Cephalothorax: Seta 1 - CT very long, stout, with 2 branches; 3 - CT very long, with 3 – 5 branches; 4 - CT shorter than 5 - CT; 7 - CT long, with 3 – 5 branches; 11 - CT relatively long, normally with 2 branches (single on 1 side of 1 specimen). Abdomen: Seta 3 - I long, moderately stout, with 3 or 4 branches; 6 - I long, slender, single or 2 - branched, longer than 7 - I; 2 - II inserted lateral to 1 - II and mesal to 3 - II; 3 - II long, stout, aciculate, with 2 or 3 branches; 4 - II inserted mesal to 5 - II; 6 - II long, slen- der, single to 3 - branched, longer than 7 - II; 3 - III long, stout, with 2 branches; 5 - IV, V exceptionally long; 2 - IV inserted mesal to 1,3 - IV; 6 - VII short, slender, inserted mesal and slightly anterior to 9 - VII; 9 - VII moderately long, stout, aciculate, with 3 – 5 branches; 9 - VIII moderately long, stout, aciculate, with 10 or 11 branches, inserted noticeably far anteriorly from posterior margin. Paddle: Ovoid; midrib well developed, reaching apex of paddle; without hairlike spicules on lateral margins; seta 1 - Pa very long, 0.65 – 1.02 length of paddle, single. Fourth-instar larvae Head: With scooplike pouch anteroventral to antennal base; seta 1 - C relatively short, moderately thickened throughout length but distal part slightly wider, apex bluntly pointed, single; 4 - C very short, slender, with 6 – 9 branches, inserted near, mesal and slightly posterior to 5 - C; 5,6 - C very long, stout, dark, single, 5 - C inserted near, mesal and slightly posterior to 6 - C; 7 - C long, stout, lightly aciculate, with 2 branches (3 - branched on 1 side of 1 specimen), inserted anterior and lateral to 4 – 6 - C and posterior to antennal base; 12 - C short, slender, with 6 – 9 branches, inserted mesal to 13 - C; 13 - C moderately long, slender, with 3 – 5 branches; 19 - C absent; antenna long, narrow, with few small spicules, seta 1 - A single, extending beyond apex of antennal shaft. Thorax: Seta 2 - P longer than 1,3 - P; 4 - P moderately long, slender, with 3 or 4 branches; 6 - P longer than 7 - P; 6,7 - P longer than 5 - P; 8 - P relatively short, slender, with 2 or 3 branches; 1 - M moderately long, slender, with 3 – 6 branches, approximately 2.9 times length of 2 - M; 4 - M with 3 – 6 branches; 5 - M longer than 7 - M; 2 - T with 8 – 12 branches; 6 - T with 3 – 5 branches. Abdomen: Seta 12 - I present; 6,7 - I moderately long, stout, 6 - I with 4 – 6 branches, 7 - I with 4 or 5 branches, only slightly shorter than 6 - I; 6 - II moderately long, stout, with 3 or 4 branches; 7 - II relatively moderately long, slender, with 4 – 6 branches; 8 - II single; 1 - VII very long, moderately stout, with 2 or 3 branches; 3 - VII short, slender with 3 – 8 branches, inserted lateral and slightly anterior to 1 - VII; 7,10,12 - VII branched; 1 - VIII with 5 or 6 branches; 2,4 - VIII branched; segment VIII with comb comprising 80 – 94 scales in patch; segment X with moderately large saddle, incomplete ventrally, acus absent, 1 - X moderately long, slender, with 3 branches, inserted on saddle, 3 - X very long, stout, single, ventral brush inserted on grid with both transverse and lateral bars, 2 precratal setae. Siphon: Acus attached; pecten comprising 24 – 36 evenly spaced spines; seta 1 - S short, slender, with 6 – 8 branches, inserted distal to pecten. Included species Himalaius gilli (Barraud) and Hi. simlensis (Edwards). The latter species is provisionally placed in Himalaius. Distribution Himalaius gilli: India (Western Himalayas, Himachal Pradesh State, Krol Mountains near Solan, Naini Tal District, Uttar Pradesh State, Himachal Pradesh State, Uttaranchal State) and Nepal (Okaldunga District, Ramechhap District). Himalaius simlensis: India (Western Himalayas, Bhowali, Kumaon, Lansdowne, Muktesar, Simla) (Barraud, 1934), and Himachal Pradesh and Uttaranchal (Kaur, 2003). Bionomics Barraud (1924, 1934) reported that the immature stages of Hi. gilli (including those of the type series) were collected from water in treeholes. However, Kaur (1992) states that Barraud’s paratypes deposited in the National Institute of Communicable Diseases, New Delhi, India, were reared from larvae collected in bamboo. Darsie, Pradhan & Vaidya (1992) collected pupae from turbid, chocolate-coloured water in treeholes. The immatures of Hi. simlensis are not known. Discussion Few specimens of Hi. gilli were available for examination but these bear distinct characters in all life stages that readily distinguish the genus (see above description based on Hi. gilli). Several abdominal segments of the larval specimens are in poor condition and could not be described. The species simlensis is provisionally included in Himalaius based on the characters mentioned in the brief and incomplete descriptions of Edwards (1922) and Barraud (1924). Etymology The generic name Himalaius is formed from ‘ Himalaya’, the mountains, and the masculine Latin suffix ‘ - ius ’, meaning ‘ pertaining to the Himalayas’. Recommended abbreviation = Hi.	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFBD0D7FCB7DF9940733315F.taxon	materials_examined	Type species: Culex varipalpus Coquillett, 1902. Females Head: Vertex with decumbent scales narrow (few broad scales occasionally present laterally in some species); occiput and vertex with erect forked scales; eyes moderately separated above antennal pedicels; antennal pedicel with few to several, often partially overlapping, small, broad scales on mesal surface; clypeus bare; maxillary palpus dark with pale-scaled areas (all dark-scaled in Ja. muelleri); proboscis without median, pale-scaled band. Thorax: Scutum with moderately wide to wide, median, pale-scaled stripe or patch extending over anterior acrostichal area laterally to inner dorsocentral area or wider and extending laterally to mesal margin of scutal fossa, pale-scaled, median area normally continuing posteriorly to prescutellar area (except in Ja. varipalpus); narrow, curved, white scales on scutal fossa (lateral and posterior margins), supraalar area (elongate longitudinal line) and prescutellar space (lateral and anterior margins of median, bare area); acrostichal and dorsocentral areas with both anterior and posterior setae; paratergite with broad, pale scales; postpronotum with narrow, curved scales dorsally and broad scales ventrally (except all narrow scales in Ja. muelleri); postspiracular, upper proepisternal, lower prealar, mesokatepisternal (large upper patch and moderate lower posterior patch), mesepimeral (upper and lower patches, except 1 large patch in Ja. muelleri) and metameral (except Ja. deserticola) areas with broad, pale scales. Wing: Dark-scaled with small white-scaled patch at base of costa (pale-scaled patch reduced to 2 or 3 scales or occasionally absent in Ja. muelleri); upper calypter with several setae on margin; alula with row of narrow scales on margin; remigium without setae ventrally and normally absent dorsally (1 or 2 setae distally in Ja. muelleri and Ja. varipalpus). Legs: Anteprocoxal membrane bare, postprocoxal membrane normally with broad, pale scales (absent in Ja. deserticola and Ja. sierrensis); hindtarsomere 1 with broad, basal and apical, white-scaled bands, hindtarsomeres 2 and 3 with broad, apical, whitescaled bands (hindtarsus entirely dark-scaled in Ja. muelleri), ungues simple. Abdomen: Tergum I with white-scaled patch on laterotergite; segment VIII dorsoventrally flattened. Genitalia: Tergum VIII sclerotized, width greater than length, setae on distal 0.70 or more of tergum; sternum VIII sclerotized, width greater than length, seta 2 - S inserted posterolateral to 1 - S, scales normally present (not seen in only available specimen of Ja. monticola); tergum IX comprising single sclerite with moderately deep, median, apical emargination, 3 – 12 setae distally on each side of midline; postgenital lobe moderately long, relatively broad, apex broad with very small, median emargination, 16 – 25 setae distally; upper and lower vaginal sclerites absent; insula liplike, with setae in lateral patches; cercus moderately long, relatively narrow with apex sharply rounded, without scales (moderately wide with apex broadly rounded and 1 – 3 scales / cercus in Ja. muelleri); 3 spermathecal capsules. Males Head: Maxillary palpus with 5 palpomeres, longer than proboscis, moderate number of moderately long to long setae ventrolaterally on distal part of palpomere 3 and ventrolaterally on palpomere 4; antennal flagellar whorls with numerous long setae directed dorsally and ventrally. Legs: Fore- and midtarsi with 2 unequal ungues, each with 1 tooth (smaller ungues simple in Ja. muelleri); hindungues equal, both simple. Genitalia: Tergum IX with posterior margin with pair of short, relatively narrow, heavily sclerotized lobes bearing several short, stout setae; gonocoxite moderately long, relatively narrow, mesal margin of dorsal surface without apical lobe but with small basal lobe bearing several long setae, 1 or more of these setae stout and somewhat flattened, ventral surface without specialized scales or setae, mesal surface membranous; gonostylus attached at apex of gonocoxite, moderately long, narrow, distal part slender and slightly curved mesally, with single moderately long, narrow gonostylar claw at apex; aedeagus simple, tubelike, moderately long, apex broadly rounded; proctiger with cercal setae; claspette with single moderately long, slender, columnar stem, claspette filament attached apically, relatively long, relatively narrow, somewhat flattened with distal part narrowing to pointed, slightly recurved apex; sternum IX with setae on posteromedian area. Pupae Cephalothorax: Seta 1 - CT longer than 3 - CT, single or 2 - branched; 11 - CT single. Abdomen: Setae 3,6 - I single, moderately stout; 2 - II shorter and inserted mesal to 1 - II; 3 - II single, moderately long, stout, inserted mesal to 2 - II; 5 - II less than 3 - II length, inserted mesal to 4 - II (lateral to 4 - II in Ja. muelleri); 3 - III single, stout, longer than 5 - III; 5 - V single, less than tergum VI median length (slightly greater than tergum length in Ja. muelleri); 2,3 - VI inserted lateral to 1 - VI (? in Ja. laguna); 6 - VII inserted posterior and mesal to 9 - VII (at same level in Ja. muelleri); 9 - VII longer and stouter than 6 - VII; 9 - VIII stout, with 2 or 3 branches. Paddle: Length greater than width; apex broadly rounded or with minute, broad, median emargination; midrib extending 0.60 – 0.70 from base; without fringe of hairlike spicules; seta 1 - Pa short, single. Fourth-instar larvae Head: Seta 1 - C single, spiniform; 4 - C short (barely moderately long in Ja. muelleri), multiple-branched, inserted mesal and slightly anterior to 6 - C; 5 - C moderately long (approaching long), single, inserted mesal and slightly anterior to 7 - C and posterior to 4,6 - C; 6 - C moderately long, with 2 branches (single in Ja. muelleri); 7 - C moderately long, multiple-branched; 12 - C branched, inserted mesal to 13 - C; 13 - C single; 19 - C very short; antenna without spicules (with few spicules in Ja. muelleri), seta 1 - A single, 2,3 - A inserted apically. Thorax: Setae 1 – 3 - P inserted on common setal support plate; 4 - P branched, longer than 8 - P; 5 - P branched; 4 - M single (rarely 2 - branched in Ja. sierrensis); 5 - M longer than 7 - M; 2,6 - T single. Abdomen: Setae 3 - I – VI and 2,4 - VIII single; 6 - I – III long, aciculate, branched; 7 - I moderately long to long, stout, aciculate, single; 12 - I absent (present in Ja. deserticola and Ja. sierrensis); 7 - II moderately long, single, except relatively short and branched in Ja. muelleri; 1 - VII longer than dorsal length of segment X (see method of measurement for character 49 in Character Descriptions); 2 - VII inserted near 1 - VII; 3 - VII moderately long, inserted anterolaterad of 1 - VII; segment VIII with comb comprising several scales in 1 or 2 irregular rows; segment X with saddle incomplete ventrally, without acus, seta 1 - X single or 2 - branched, inserted on saddle in small unsclerotized area (reduced in Ja. muelleri), 2 - X branched, 3 - X single, ventral brush attached to grid with only transverse bars (Ja. muelleri with both transverse and lateral bars), seta 4 a-X long, except relatively short in Ja. muelleri, 4 relatively long, broad anal papillae with apices broadly rounded. Siphon: Acus present; pecten comprising several evenly spaced spines, on proximal 0.50 of siphon; seta 1 - S branched, aciculate, inserted distal to pecten; 6 - S short, single. Included species Jarnellius deserticola (Zavortink), Ja. laguna (Arnell & Nielsen), Ja. monticola (Belkin & McDonald), Ja. muelleri (Dyar), Ja. sierrensis (Ludlow) and Ja. varipalpus (Coquillett). Distribution Canada (British Columbia), Mexico (Baja California Sur, Distrito Federal, Durango, Hidalgo, Mexico, Sonora, Zacatecas) and United States (Arizona, California, Idaho, Montana, Nevada, New Mexico, Oregon, Texas, Utah, Washington). Bionomics Immature stages are usually found in treeholes. Jarnellius muelleri are found occasionally in artificial containers and were once collected from leaf axils of maguey (Zavortink, 1972). Insight into the ecology of treeholes in which most species of Jarnellius are found is given in Zavortink (1985). Discussion The above generic description of Jarnellius is additional to the description (Varipalpus Group) given by Arnell & Nielsen (1972), who should be referred to for the systematic background, descriptions and illustrations of the included species. Zavortink (1972) provided a description and illustrations of Ja. muelleri (as Ae. (Och). muelleri), the type species of the new subgenus Lewnielsenius. The genus includes two subgenera (see below). Myers (1967) described and illustrated eggs of Ja. sierrensis and Ja. varipalpus. Etymology Jarnellius is named in honour of Dr J. Hal Arnell in recognition of his important work on species of this genus and other New World aedines. The generic name is masculine, formed from his first initial ‘ J’, surname and the masculine Latin suffix ‘ - ius ’. Recommended abbreviation = Ja.	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFBB0D7CCB48FA7301493152.taxon	description	Females Head: Maxillary palpus dark-scaled with pale-scaled areas. Thorax: Scutal fossa with white-scaled lines on lateral and posterior margins that are not expanded mesally; postpronotum with narrow, curved scales dorsally and broad scales ventrally; hypostigmal area without scales; mesepimeron with upper and lower patches of white scales. Legs: Hindtibia dark-scaled with white scales basally; hindtarsomere 1 with broad basal and apical, whitescaled bands, hindtarsomeres 2 and 3 with broad, apical, white-scaled band; fore- and midungues each with 1 tooth. Genitalia: Cercus moderately long, moderately wide, apex sharply rounded, without scales. Males Legs: Fore- and midungues each with 1 tooth. Pupae Abdomen: Seta 5 - II inserted lateral to 4 - II; 5 - V shorter than median length of tergum VI; 6 - VII inserted posterior and mesal to 9 - VII. Fourth-instar larvae Head: Seta 6 - C with 2 branches; antenna without spicules. Abdomen: Seta 7 - II developed similarly to 7 - I, single (occasionally 2 - branched in Ja. sierrensis), moderately long to long; ventral brush attached to grid with only transverse grid bars, seta 4 a-X long. Distribution Canada (British Columbia), Mexico (Baja California Sur, Sonora) and United States (Arizona, California, Idaho, Montana, Nevada, New Mexico, Oregon, Utah, Washington). Included species Jarnellius deserticola, Ja. laguna, Ja. monticola, Ja. sierrensis and Ja. varipalpus. Recommended abbreviation = Jar.	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFB80D7CC8B9FA85062D3260.taxon	materials_examined	Type species: Aedes (Heteronycha) muelleri Dyar, 1920. Females Head: Maxillary palpus dark-scaled. Thorax: Scutal fossa with white-scaled areas on lateral and posterior margins that are expanded mesally; postpronotum with narrow scales; hypostigmal area with pale scales; mesepimeron with single, large patch of white scales extending over upper and lower areas. Legs: Hindtibia and hindtarsus entirely dark-scaled; fore- and midungues simple. Genitalia: Cercus moderately long, moderately wide, apex broadly rounded, with 1 – 3 scales. Males Head: Maxillary palpus dark-scaled. Legs: Fore- and midtarsi with larger ungues with 1 tooth, smaller ungues simple. Pupae Abdomen: Seta 5 - II inserted mesal to 4 - II; 5 - V longer than median length of tergum VI; 6 - VII inserted mesal and at approximately same level as 9 - VII. Fourth-instar larvae Head: Seta 6 - C single; antenna with few spicules. Abdomen: Seta 7 - II developed differently from 7 - I, relatively short, branched; ventral brush attached to grid with both transverse and lateral grid bars, seta 4 a-X relatively short. Distribution Mexico (Distrito Federal, Durango, Hidalgo, Mexico, Zacatecas) and United States (Arizona, New Mexico, Texas). Included species Jarnellius muelleri (Dyar). Etymology Subgenus Lewnielsenius is named in honour of Dr Lewis T. Nielsen in recognition of his many contributions to the taxonomy, distribution and bionomics of North American mosquitoes, especially aedine species. The subgenus is masculine, formed from the combination of his first name, surname and the Latin masculine suffix ‘ - ius ’. Recommended abbreviation = Lew.	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFB80D62CB6CF9B2038C3588.taxon	materials_examined	Type species: Aedes (Finlaya) chungi Lien, 1968. Females Head: Vertex with decumbent scales broad, pale on median area and dark on lateral areas; erect forked scales numerous on occiput, few semierect scales on vertex caudal to ocular setae; ocular line narrow, covered with broad, pale scales; 2 long interocular setae; eyes above antennal pedicels contiguous or separated by less than width of 1 eye facet; antennal pedicels with few small, broad, dark scales on mesal surface; clypeus bare; maxillary palpus and proboscis dark-scaled; proboscis longer than forefemur. Thorax: Covered with narrow, relatively straight scales except bare, median, prescutellar area and elongate strip between supraalar and prescutellar setae, scales dark except for following pale-scaled areas: narrow stripe on anterior promontory and extending caudally on acrostichal area to approximately level of scutal angle where it becomes less distinct for short distance and then distinct on caudal part of posterior acrostichal area, narrow line on caudal part of anterior dorsocentral area and extending over posterior dorsocentral area to caudal margin of scutum, stripe broadened at juncture of prescutal suture into small triangular patch which connects with narrow stripe extending outward along scutal suture and cephalad along margin of scutal fossa onto antedorsocentral area, more or less elongate stripe on supraalar area, and indistinct stripe on lateral margins of prescutellar area mesad of setae; setae on following: few on anterior promontory, antedorsocentral area, dorsocentral (anterior and posterior) area, scutal fossa and antealar area, numerous on supraalar and prescutellar areas; scutellum with broad or broad and narrow scales on lobes; antepronota widely separated, with broad, pale scales, several setae; postpronotum with pale scales, few posterior setae; prespiracular setae absent; hypostigmal and subspiracular areas bare; postspiracular area with few setae, scales absent; upper proepisternum with broad, pale scales, several setae on upper area, lower area bare; paratergite narrow, with broad, pale scales; mesokatepisternum with broad, pale scales in upper and lower patches, 3 or 4 upper and several posterior setae; prealar area with broad, pale scales in patch on lower area and extending onto lower part of knob, several upper setae; mesepimeron with 1 patch of broad, pale scales on upper area and extending onto middle, several upper setae, lower setae absent. Wing: Dark-scaled with small pale-scaled patch at base of costa; upper calypter with several setae on margin; alula with row of narrow, dark scales on margin (Ji. chungi); remigium with 2 or 3 setae distally on dorsal surface. Legs: Hindfemur with narrow basal and moderate preapical bands dark-scaled, remainder pale-scaled; hindtibia and hindtarsus dark-scaled except tarsomere 1 with broad, pale-scaled, basal band; fore- and midtarsi with 2 equal ungues, each with 1 tooth; hindtarsus with 2 equal ungues, simple. Abdomen: Tergum I with pale-scaled patch on lateral tergite; terga dark-scaled with small, basolateral, pale-scaled patch; segment VII laterally compressed. Genitalia: Tergum VIII with apex broadly rounded, numerous broad scales on most of surface; sternum VIII sclerotized, width greater than length, apical margin with median emargination, scales absent, seta 2 - S inserted posterior to 1 - S; tergum IX comprising 2 moderately pigmented, moderately long, lateral plates connected mesally by basal pigmented strip, base wider than apex, few short setae distally on each plate; postgenital lobe long, relatively narrow, apex rounded, several setae distally; upper vaginal sclerite small; lower vaginal sclerite absent; insula liplike, with few setae in lateral patches; cercus moderately long, relatively wide, outer distal margin gently oblique, without scales; 3 spermathecal capsules. Males Head: Maxillary palpus with 5 slender palpomeres in nearly straight line, shorter than proboscis, few moderately long setae on distal part of palpomere 3 ventrally and palpomere 4 ventrolaterally; antennal flagellar whorls with numerous long setae directed dorsally and ventrally. Thorax: Scaled areas of scutum with narrow, relatively long, pale scales except small area of scutal fossa with narrow, dark scales. Legs: Fore- and midtarsi with 2 unequal ungues, larger unguis with 2 teeth, smaller unguis with 1 tooth; hindtarsus with 2 equal ungues, simple. Genitalia: Tergum IX with median area narrow, lateral area developed as broad, rounded lobe projecting cephalad, posterior margin with very small, sclerotized lobe on each side of midline bearing few short, slender, curved setae; gonocoxite moderately long, relatively narrow, dorsal surface without apical and basal lobes on mesal margin, however, with small, dark, oval area basomesally bearing straight setae, few of these setae on most proximal part long and somewhat flattened, middle part with several moderately long setae and distal part with several short setae, ventral surface with narrow, elongate patch of moderately long, slender, lanceolate setae on distal area of mesal margin (Lu & Ji, 1997: fig. 34, shows narrow fusiform scales in this area of Ji. gonguoensis), mesal surface membranous, lateral and outer parts of dorsal and ventral surfaces with broad scales; gonostylus attached at apex of gonocoxite, relatively short, narrow, single gonostylar claw, long, narrow, attached at apex; aedeagus simple, tubelike, apex concave; proctiger with few tiny, cercal setae; claspette with single, moderately long, narrow, columnar stem, claspette filament attached apically, long, somewhat flattened beyond midlength, with transverse striations on much of median area, apex curved into narrow point; sternum IX with few setae on posteromedian area. Pupae	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFA60D60CB17F9F7000C35CA.taxon	materials_examined	Type species: Leucomyia australiensis variety papuensis Taylor, 1914. Females Head: Vertex with median area covered with narrow, curved, pale scales; occiput and vertex with erect forked scales; interocular space with patch of narrow, curved, pale scales projecting ventrad over mesal margins of eyes; eyes contiguous or nearly so; antennal pedicel with few short, fine setae and small, broad, nonoverlapping scales on mesal surface; clypeus bare; maxillary palpus and proboscis dark-scaled. Thorax: Scutum with narrow, curved, dark scales except for moderate to broad median, longitudinal patch of white scales on approximately anterior 0.55 – 0.70 (white-scaled patch on acrostichal area and extending laterally over mesal part of dorsocentral area or to near lateral part of scutal fossa, but not on lateral part of scutal fossa), other scutal areas normally dark-scaled including supraalar and prescutellar areas (but not median, prescutellar bare area) (few species with white scales along lateral margin of prescutellar bare area); scutellum with narrow, curved scales on all lobes; acrostichal (both anterior and posterior) areas without setae, anterior dorsocentral area normally without setae (occasionally 1 seta on caudal part), posterior dorsocentral area with several setae; paratergite with broad, pale scales; postpronotum normally with patch of broad, scales ventrally and narrow, curved scales dorsally (no narrow scales seen in Pm. dobodura but this area rubbed in specimens examined; only narrow, curved, pale scales in Pm. anggiensis); mesokatepisternum with upper and lower posterior patches of broad, pale scales; metameron bare. Wing: Costa dark-scaled; upper calypter with several setae on margin; alula with row of narrow scales on margin; remigium with dorsal setae distally. Legs: Ante- and postprocoxal membranes bare; hindfemur with dorsoapical patch of pale scales; tibiae dark-scaled (except narrow, pale-scaled, basal band in Pm. anggiensis); hindtarsus with broad, basal, palescaled bands on tarsomeres 1 – 4. Genitalia: Tergum VIII sclerotized, with numerous broad scales on distal part; sternum VIII sclerotized, length greater than width, normally moderate to large number of broad scales in lateral patches, apex with small to moderately deep, median emargination separating pair of small to broad lobes, apical margin with numerous short, curved setae, seta 2 - S inserted posterior to seta 1 - S; tergum IX comprising 2 moderately pigmented, narrow, lateral plates connected by basal or subbasal pigmented or membranous band, distal part of mesal surfaces of plates darkly pigmented and strongly fused to basolateral margins of cerci, without setae; postgenital lobe with several setae distally; upper vaginal sclerite small, moderately pigmented; lower vaginal sclerite absent; insula liplike, V-shaped medially (depressed), with setae in lateral patches; cercus moderately long, moderately wide, apex broadly rounded, without scales (rarely 1 or 2 scales present); 3 spermathecal capsules. Males Head: Maxillary palpus with 5 palpomeres, slightly shorter to slightly longer than proboscis, numerous long setae ventrolaterally on distal part of palpomere 3 and ventrolaterally on palpomere 4; antennal flagellar whorls with numerous long setae directed dorsally and ventrally. Legs: Fore- and midtarsi with 2 ungues, both toothed. Genitalia: Tergum IX comprising 2 narrow, lateral plates narrowly joined mesally, without setae; gonocoxite moderately long, moderately wide, mesal margin of dorsal surface without apical lobe but with small basal lobe bearing several short and moderately long, slender setae, ventral surface without specialized setae or scales on mesal margin, mesal surface membranous; gonostylus attached at apex of gonocoxite, moderately long, narrow, with single, short, narrow, gonostylar claw at apex; aedeagus simple, tubelike, moderately long, distal 0.50 widest; proctiger with cercal setae; claspette with single moderately long, slender, columnar stem, claspette filament attached apically, moderately long to long, somewhat flattened, relatively narrow with distal part tapering to slender apex, middle part with transparent sheaf with transverse striations encircling spiniform; sternum IX with setae on posteromedian area. Pupae Cephalothorax: Seta 11 - CT single (rarely 2 - branched in Pm. argyronotum). Abdomen: Seta 3 - I long, stout, single, longer than 6 - I; 6 - I single, longer than 7 - I; 2 - II short, inserted lateral to 1 - II; 3 - II long, stout, single, longer than 6 - II; 3 - III long, stout, single, longer than 5 - III; 5 - IV, V long, stout, longer than following tergum; 6,9 - VII both relatively stout, aciculate, branched; 9 - VIII stout, aciculate, multiple-branched. Paddle: Broad; apex broadly rounded or flattened; midrib nearly reaching apex; without hairlike spicules on margin; seta 1 - Pa single. Fourth-instar larvae Head: Seta 1 - C relatively stout, single; 4 - C short, branched, inserted slightly posterior and mesal to 5 - C; 5,6 - C moderately long, multiple-branched, displaced far anteriorly, 6 - C inserted lateral and slightly anterior to 5 - C; 7 - C moderately long, multiplebranched, inserted lateral and posterior to 4 – 6 - C; 12 - C short, branched, inserted mesal to 13 - C; 13 - C single; 14 - C short, normally single; 19 - C absent; antenna with few to numerous spicules, 1 - A normally branched. Thorax: Setae 1 – 3 - P inserted on common setal support plate; 8 - P short, normally branched; 1 - M, T not on tubercle, 1 - M moderately long, 1 - T short; 6 - T single. Abdomen: Seta 7 - I long, stout, normally single; 12 - I present; 7 - II short, branched; 7,10,12 - VII single; 2,4 - VIII single; segment VIII with comb normally with numerous scales in patch (some species with several scales in 1 or 2 irregular rows); segment X with saddle incomplete ventrally, moderately developed spicules on posterior margin dorsal to seta 1 - X (occasionally spicules smaller), ventral brush with setae attached to grid with both transverse and lateral bars, seta 1 - X single (except 2 or 3 branches in Pm. argenteitarsis and Pm. dobodura), inserted on saddle. Siphon: With attached acus; pecten with spines evenly spaced; seta 1 - S branched, inserted distal to pecten. Included species Patmarksia anggiensis (Bonne-Wepster), Pm. argenteitarsis (Brug), Pm. argyronotum (Belkin), Pm. buxtoni (Belkin), Pm. clintoni (Taylor), Pm. derooki (Brug), Pm. dobodura (King & Hoogstraal), Pm. hollandia (King & Hoogstraal), Pm. mackerrasi (Taylor), Pm. novalbitarsis (King & Hoogstraal), Pm. palmarum (Edwards), Pm. papuensis (Taylor) and Pm. subalbitarsis (King & Hoogstraal). Distribution Australia (Queensland), Indonesia (Irian Jaya, Moluccas), Papua New Guinea and Solomon Islands (Guadalcanal, New Georgia). Bionomics Immature stages of Patmarksia species are found in plant containers (e. g. fallen palm fronds, coconut husks and leaves, treeholes, holes in fallen logs, cut bamboo, funnel-shaped fungi), artificial containers (e. g. metal cans, drum heads, flower bowls), and small puddles, ruts, borrow pits and rock pools in stream beds. Discussion See King & Hoogstraal (1946: pls 20, 21, 23) for illustrations of the male genitalia, fourth-instar larva and adults of Pm. papuensis, Penn (1949: fig. 35) for the pupa and Reinert (2002 e: fig. 9) for the female genitalia. Etymology Patmarksia is named in honour of Dr Elizabeth Nesta Marks in recognition of her pioneering and outstanding contributions to the taxonomy of Australasian mosquitoes. The generic name is feminine, formed from her nickname ‘ Pat ’, her surname and the feminine Latin suffix ‘ - ia ’. Recommended abbreviation = Pm.	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFA40D61CB6AFE1903B8323C.taxon	materials_examined	Type species: Culex notoscriptus Skuse, 1889. Females Head: Vertex with decumbent scales both broad and narrow; occiput and vertex with erect forked scales; eyes contiguous or separated by less than or equal to 1 eye facet; antennal pedicel with patch of overlapping, broad, silvery scales on mesal surface; clypeus bare; maxillary palpus with apex pale-scaled; proboscis with well-developed pale-scaled band at about midlength. Thorax: Acrostichal area with narrow stripe of pale scales on anterior and posterior areas; posterior dorsocentral area with narrow stripe of pale scales; scutal fossa with narrow stripe of pale scales on lateral and usually posterior margins; supraalar area with narrow, longitudinal patch of pale scales; prescutellar median, bare area with pale scales laterally mesal to setae; acrostichal and dorsocentral areas with both anterior and posterior setae; scutellum with broad scales on all lobes; scutal fossa with 1 or 2 posterior setae; paratergite with broad, pale scales; broad scales on antepronotum, postpronotum, subspiracular area, upper proepisternum, lower prealar area, upper and lower posterior mesokatepisternum, and upper and lower mesepimeron, other pleural areas without scales; mesepimeron without lower setae. Wing: Dark-scaled; upper calypter with several setae on margin; alula with row of narrow scales on margin; remigium without dorsal and ventral setae. Legs: Ante- and postprocoxal membranes bare; mid- and hindfemora with median stripe of pale scales on anterior surface; hindtarsus with wide, basal, palescaled bands on tarsomeres 1 – 4; fore- and midungues equal, each with 1 tooth; hindungues equal, simple. Abdomen: Tergum I with silvery-scaled patch on lateral tergite; segment VII laterally compressed. Genitalia: Tergum VIII sclerotized, width greater than length, without (rarely with 1 or 2 adventitious) scales, setae on distal 0.30 or less of tergum; sternum VIII sclerotized, width greater than length, without (rarely with 1 or 2 adventitious) scales, apex broad, each side nearly flat and very gently sloping to midline, seta 2 - S inserted posterior to seta 1 - S; tergum IX comprising single sclerite with moderately deep, median, apical emargination, 2 – 7 setae distally on each side of midline; postgenital lobe relatively long, narrow, with spicules well developed on lateral margins, few setae distally; upper vaginal sclerite well developed, moderately to heavily pigmented; lower vaginal sclerite absent; insula liplike, with 5 – 7 setae in lateral patches; cercus moderately long, moderately wide, apex broadly rounded, without scales; 3 spermathecal capsules. Males Head: Maxillary palpus with 5 palpomeres, longer than proboscis, numerous long setae ventrolaterally on distal part of palpomere 3 and ventrolaterally on palpomere 4; antennal flagellar whorls with numerous long setae directed dorsally and ventrally. Legs: Fore- and midtarsi with 2 unequal ungues, both toothed; hindtarsus with 2 equal ungues, simple. Genitalia: Posterior margin of tergum IX with pair of short, narrow, acute lobes bearing few slender to moderately stout setae; gonocoxite relatively long, relatively narrow, mesal margin of dorsal surface without apical and basal lobes, ventral surface with patch of long, narrow, lanceolate setae on distal part of mesal margin, mesal surface membranous; gonostylus attached at apex of gonocoxite, relatively short, less than or equal to 0.25 length of gonocoxite, narrow, distal part slender and slightly curved mesally, with single long, narrow gonostylar claw at apex; aedeagus simple, tubelike, noticeably long and narrow; paramere relatively long and narrow; proctiger with cercal setae; claspette with single, long, slender, columnar stem, claspette filament attached apically, flattened, curved with distal part tapering to bluntly pointed apex; sternum IX with setae on posteromedian area. Pupae	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFA20D67C8BCFD330601326A.taxon	materials_examined	Type species: Aedes (Finlaya) pulchrithorax Edwards, 1939. Females Head: Vertex with median, pale-scaled patch; vertex and occiput with erect forked scales; ocular line narrow, with broad, silvery scales; eyes contiguous; clypeus bare; maxillary palpus and proboscis darkscaled. Thorax: Scutum with narrow, curved scales; pale scales forming relatively narrow stripe on anterior and posterior acrostichal areas and extending to prescutellar area, similar stripe on posterior dorsocentral area extending caudally laterad of prescutellar setae to posterior margin of scutum, elongate, longitudinal patch on supraalar area, prescutellar area with narrow patches on lateral and anterior parts of median, bare area; scutal fossa covered with narrow, curved, dark scales; dorsocentral area with anterior and posterior setae; acrostichal area without setae; scutellum with narrow, curved scales on all lobes; mesopostnotum bare; paratergite with broad, silvery scales; broad, silvery scales on antepronotum, postpronotum (large patch), upper proepisternum, mesokatepisternum (small upper and lower posterior patches), lower prealar area and mesepimeron (single large patch extending over upper and lower areas); subspiracular and postspiracular areas without scales; upper proepisternum with 3 – 5 setae; mesepimeron without lower setae. Wing: Dark-scaled; upper calypter with several setae on margin; alula with row of narrow scales on margin; remigium with few dorsal setae distally. Legs: Ante- and postprocoxal membranes bare; hindfemur without pale scales dorsoapically; hindtibia with broad, basal, white-scaled patch; hindtarsus with white-scaled, basal bands on tarsomeres 1 and 2, narrow on tarsomere 1, broad on tarsomere 2; fore-, mid- and hindungues equal, each with 1 tooth. Abdomen: Tergum I with silvery-scaled patch on lateral tergite; segment VII laterally compressed. Genitalia: Tergum VIII with numerous broad scales on much of surface; sternum VIII sclerotized, width greater than length, apical margin nearly flat or with very shallow, broad, median emargination, with several stout setae decreasing in length mesally, few to several broad scales, seta 2 - S inserted posterior to 1 - S; tergum IX comprising 2 long, very narrow, darkly pigmented sclerites connected on approximately mesal 0.70 by moderately pigmented band, 2 or 3 short setae apically on each side of midline; postgenital lobe long, relatively narrow, with several setae distally; upper and lower vaginal sclerites absent; insula liplike, with few setae in lateral patches; cercus moderately long, relatively narrow, apex sharply rounded, without scales; 3 spermathecal capsules. Males Head: Maxillary palpus with 5 palpomeres, all relatively slender and in approximately straight line, dark-scaled, slightly longer to slightly shorter than proboscis, palpomeres 4 and 5 with few setae; antennal flagellar whorls with numerous long setae directed dorsally and ventrally. Legs: Fore-, mid- and hindtarsi each with 2 ungues, each with 1 tooth. Genitalia: Tergum VIII with small, median lobe on posterior margin; tergum IX comprising 2 very small sclerites connected mesally by tonguelike, median, cephalad-directed lobe of tergum X, each sclerite with small, strongly sclerotized lobe on posterior margin bearing few short, stout setae, lateral part of lobe connected by narrow band with sternum IX; gonocoxite moderately long, moderately wide, dorsal surface with scales all approximately same length, mesal margin without apical lobe, with moderate-size, basal lobe, distal part of lobe narrow and bearing 2 or 3 moderately long to long, moderately stout setae whereas proximal part of lobe is expanded with series of wrinkles at base and bearing several short to moderately long setae (some of which may be narrowly lanceolate), ventral surface without specialized scales or setae, mesal surface membranous; gonostylus attached at apex of gonocoxite, relatively short, slen- der, single gonostylar claw moderately long, narrow, attached at apex; aedeagus simple, tubelike, moderately long; claspette with single moderately long, slen- der, columnar stem, claspette filament long, proximal part narrow, distal part expanded, flattened but terminating in pointed apex; proctiger with few cercal setae; sternum IX with few setae on posteromedian area. Pupae Cephalothorax: Setae 1,3 - CT branched; 7 - CT very long, greater than 6.0 times length of 6 - CT; 8 - CT moderately long, moderately stout, aciculate, with 5 – 14 branches; 9 - CT moderately long, moderately stout, with 2 – 6 branches; 10 - CT relatively short, very slen- der, with 6 – 19 branches; 11 - CT long, stout, single. Abdomen: Seta 3 - I long, with 2 or 3 branches; 6 - I very long, stout, aciculate, single, noticeably longer than 7 - I; 7 - I long, stout, aciculate, normally with 2 branches (rarely single or 3 - branched); 2 - II shorter and inserted lateral to 1 - II and mesal to 3 - II; 3 - II, III long, stout, single; 5 - II shorter than 3 - II; 6 - II long, stout, aciculate, single, longer than 7 - II; 7 - II long, branched; 5 - IV – VI long, stout, aciculate, longer than following tergum; 2 - VI inserted mesal to 1,3 - VI; 6 - VII short, slender, with 4 – 10 branches, inserted posterior and mesal to 9 - VII; 9 - VII stout, aciculate, branched, longer than 6 - VII; 9 - VIII stout, aciculate, multiple-branched, inserted near posterolateral corner of segment VIII. Paddle: Length greater than width, index 1.20 – 1.42; without hairlike spicules on margins; midrib reaching apex of paddle; seta 1 - Pa single. Fourth-instar larvae Head: Seta 1 - C single, spiniform; 4 - C short, very slen- der, branched, inserted mesal and slightly posterior to 6 - C; 5 - C long, stout, aciculate, often single (range single to 3 - branched), inserted posterior and mesal to 6 - C and posterior and lateral to 4 - C; 6 - C long, stout, aciculate, longer than 5 - C, inserted mesal and very slightly anterior to 7 - C; 7 - C moderately long, relatively stout, aciculate, with 3 – 6 branches; 12 - C branched, inserted mesal to and shorter than 13 - C; 13 - C branched; 19 - C absent; antenna relatively long, slender, slightly curved mesally, with few scattered spicules, seta 1 - A moderately long, single or 2 - branched (occasionally 3 - branched). Thorax: Seta 5 - P branched, shorter than 6 - P; 8 - P short; 4 - M branched; 6 - T single. Abdomen: Setae 6 - I, II long, branched (6 - II rarely single in Va. luteostriata); 6 - III branched, shorter than 6 - II; 7 - I long, with 2 or 3 branches; 7 - II short, with 4 – 15 branches, differently developed than 7 - I; segment VIII with comb comprising numerous scales in patch; 2 - VIII branched; segment X with saddle incomplete ventrally, acus absent, setae 1 - X short, branched, inserted on saddle, ventral brush with several long, branched setae with short stems, attached to grid with well developed transverse grid bars, lateral bars absent but represented by faint depression, 2 precratal setae. Siphon: With acus attached; pecten with number of evenly spaced spines; seta 1 - S branched, inserted distal to pecten. Included species Vansomerenis hancocki (van Someren), Va. luteostriata (Robinson) and Va. pulchrithorax (Edwards). Distribution Angola, Kenya, Tanzania, Uganda, Zaire and Zambia. Bionomics Immature stages have been found in treeholes, a rot hole in the fork of a tree, a tree buttress and bored bamboo. Discussion See Edwards (1941: figs 6 m, 176 and pl. III- 14) for illustrations of the female genitalia, fourth-instar larva and female of Va. pulchrithorax. Hopkins (1952: figs 62, 63) also illustrated the fourth-instar larva of this species. Etymology Vansomerenis is named in honour of Mrs E. C. C. van Someren in recognition of her many important contributions to the taxonomy of Afrotropical mosquitoes, especially those in tribe Aedini. The generic name is feminine, formed from her surname and the feminine Greek suffix ‘ - is ’ denoting ‘ daughter of ’, i. e. daughter of van Someren. Recommended abbreviation = Va.	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFAE0D6AC8A7FBCC02883239.taxon	description	annulirostris (Theobald, 1905) chionodes (Belkin, 1962) gombakensis (Mattingly, 1959) ibis (Barraud, 1931) thomsoni (Theobald, 1905)	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFAE0D6ACB0FFEAC065831A6.taxon	description	Diceromyia Theobald, 1911 adersi (Edwards, 1917) agastyai (Tewari & Hiriyan, 1992) bananea (Wolfs, 1958) cordellieri (Huang, 1986) coulangesi (Rodhain & Boutonnier, 1983) fascipalpis (Edwards, 1912) flavicollis (Edwards, 1928) franciscoi (Mattingly, 1959) furcifer (Edwards, 1913) grassei (Doucet, 1951) iyengari (Edwards, 1923) kanarensis (Edwards, 1934) madagascarensis (van Someren, 1949) mefouensis (Ferrara, 1974) meronephada (Dyar & Shannon, 1925) micropterus (Giles, 1901) nummata (Edwards, 1923) periskelata (Giles, 1902) pseudonummata (Reinert, 1973) punctipes (Edwards, 1921) ramachandrai (Reuben, 1967) reginae (Edwards, 1922) reubenae (Tewari & Hiriyan, 1992) scanloni (Reinert, 1970) sylvatica (Brunhes, 1983) taylori (Edwards, 1936) tiptoni (Grjebine, 1953) whartoni (Mattingly, 1965) zethus (de Meillon & Lavoipierre, 1944)	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFAE0D68CB0FFA1A02C636C0.taxon	description	Downsiomyia Vargas, 1950 albolateralis (Theobald, 1908) albonivea (Barraud, 1934) axitiosa (Kulasekera, Knight & Harbach, 1990) dorseyi (Knight, 1946) ganapathi (Colless, 1958) harinasutai (Knight, 1978) idjenensis (Brug, 1934) inermis (Colless, 1958) lactea (Knight, 1946) laoagensis (Knight, 1946) leonis (Colless, 1958) litorea (Colless, 1958) mikrokopion (Knight & Harrison, 1988) mjobergi (Edwards, 1926), comb. nov. mohani (Knight, 1969) nipponica (LaCasse & Yamaguti, 1948) nipoononivea (Sasa & Nakahashi, 1952) nishikawai (Tanaka, Mizusawa & Saugstad, 1979) nivea (Ludlow, 1903) niveoides (Barraud, 1934) novonivea (Barraud, 1934) omorii (Lien, 1968) pexa (Colless, 1958) pseudonivea (Theobald, 1905) saperoi (Knight, 1946) shehzadae (Qutubuddin, 1972), comb. nov. sinensis (Chow, 1950) subnivea (Edwards, 1922) vana (Colless, 1958) watteni (Lien, 1968) Edwardsaedes Belkin, 1962 bekkui (Mogi, 1977) imprimens (Walker, 1860) pingpaensis (Chang, 1965) Eretmapodites Theobald, 1901 adami Ferrara & Eouzan, 1974 angolensis da Cunha Ramos & Ribeiro, 1992 argyrurus Edwards, 1936 brenguesi Rickenbach & Lombrici, 1975 brottesi Rickenbach, 1967 caillardi Rickenbach, Ferrara & Eouzan, 1968 chrysogaster Graham, 1909 corbeti Hamon, 1962 dracaenae Edwards, 1916 dundo da Cunha Ramos & Ribeiro, 1992 eouzani Rickenbach & Lombrici, 1974 ferrarai Rickenbach & Eouzan, 1970 forcipulatus Edwards, 1936 germaini Rickenbach & Eouzan, 1970 gilletti van Someren, 1949 grahami Edwards, 1911 grenieri Hamon & van Someren, 1961 haddowi van Someren, 1949 hamoni Grjebine, 1972 harperi van Someren, 1949 hightoni van Someren, 1947 inornatus Newstead, 1907 intermedius Edwards, 1936 jani Rickenbach & Lombrici, 1976 lacani Rickenbach & Eouzan, 1970 leucopous Graham, 1909 mahaffyi van Someren, 1949 marcellei Adam & Hamon, 1959 mattinglyi Hamon & van Someren, 1961 melanopous Graham, 1909 mortiauxi Cunha Ramos & Ribeiro, 1990 oedipodeios Graham, 1909 parvipluma Edwards, 1941 pauliani Grjebine, 1950 penicillatus Edwards, 1941 plioleucus Edwards, 1941 ssp. brevis Edwards, 1941 productus Edwards, 1941 quinquevittatus Theobald, 1901 ravissei Rickenbach & Eouzan, 1970 rickenbachi Ferrara & Eouzan, 1974 salauni Rickenbach, Ferrara & Eouzan, 1968 semisimplicipes Edwards, 1914 silvestris Ingram & de Meillon, 1927 ssp. conchobius Edwards, 1941 subsimplicipes Edwards, 1914 tendeiroi da Cunha Ramos, Ribeiro & Machado, 1992 tonsus Edwards, 1941 vansomereni Hamon, 1962 wansoni Edwards, 1941 ssp. douceti Adam & Hamon, 1959 Finlaya Theobald, 1903 alocasicola (Marks, 1947) ananae (Knight & Laffoon, 1946) avistylus (Brug, 1939) bougainvillensis (Marks, 1947) burnetti (Belkin, 1962) crocea (Knight & Laffoon, 1946) dobrotworskyi (Marks, 1958) fijiensis (Marks, 1947) flavipennis Giles, 1904 franclemonti (Belkin, 1962) freycinetiae (Laird, 1957) fuscipalpis (Belkin, 1962) fuscitarsis (Belkin, 1962) gahnicola (Marks, 1947) gani (Bonne-Wepster, 1940) * gressitti (Bohart, 1947) hollingsheadi (Belkin, 1962) horotoi (Taylor, 1972) hui (Bohart, 1957) josephinae (Marks, 1958) knighti (Stone & Bohart, 1944) kochi (Dönitz, 1901) lewelleni (Starkey & Webb, 1946) lutea (Ludlow, 1905) maffii (Taylor & Tenorio, 1974) medleri (Knight & Laffoon, 1946) neogeorgiana (Belkin, 1962) oceanica (Belkin, 1962) poicilia Theobald, 1903 samoana Grünberg, 1913 schlosseri (Belkin, 1962) solomonis (Stone & Bohart, 1944) sorsogonensis (Banez & Jueco, 1966) stonei (Knight & Laffoon, 1946) tutuilae (Ramalingam & Belkin, 1965) wallacei (Edwards, 1926) Fredwardsius Reinert, 2000 vittatus (Bigot, 1861)	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFAC0D69C8A7FD3D015B362C.taxon	description	Subgenus Georgecraigius atropalpus (Coquillett, 1902), comb. nov. epactius (Dyar & Knab, 1908), comb. nov. Subgenus Horsfallius Reinert, Harbach & Kitching, subgen. nov. fluviatilis (Lutz, 1904), comb. nov. Geoskusea Edwards, 1929 baisasi (Knight & Hull, 1951) becki (Belkin, 1962) daggyi (Stone & Bohart, 1944) fimbripes (Edwards, 1924) kabaenensis (Brug, 1939) longiforceps (Edwards, 1929) lunulata (King & Hoogstraal, 1946) perryi (Belkin, 1962) timorensis (Miyagi, Toma & Lien, 2004), comb. nov. tonsa (Edwards, 1924) Gilesius Reinert, Harbach & Kitching, gen. nov. alius (Lien, 1968), comb. nov. pulchriventer (Giles, 1901), comb. nov. Gymnometopa Coquillett, 1905 mediovittata (Coquillett, 1906) Haemagogus Williston, 1896 Subgenus Conopostegus Dyar, 1925 clarki (Galindo, Carpenter & Trapido, 1953) leucocelaenus (Dyar & Shannon, 1924) leucophoebus (Galindo, Carpenter & Trapido, 1953) leucotaeniatus (Komp, 1938) Subgenus Haemagogus acutisentis Arnell, 1973 aeritinctus Galindo & Trapido, 1967 albomaculatus Theobald, 1903 anastasionis Dyar, 1921 andinus Osorno-Mesa, 1944 argyromeris Dyar & Ludlow, 1921 baresi Cerqueira, 1960 boshelli Osorno-Mesa, 1944 capricornii Lutz, 1904 celeste Dyar & Nunez Tovar, 1927 chalcospilans Dyar, 1921 chrysochlorus Arnell, 1973 equinus Theobald, 1903 iridicolor Dyar, 1921 janthinomys Dyar, 1921 lucifer (Howard, Dyar & Knab, 1913) mesodentatus Komp & Kumm, 1938 nebulosus Arnell, 1973 panarchys Dyar, 1921 regalis Dyar & Knab, 1906 soperi Levi-Castillo, 1955 spegazzinii Brethes, 1912 splendens Williston, 1896 tropicalis Cerqueira & Antunes, 1938 Halaedes Belkin, 1962 ashworthi (Edwards, 1921) australis (Erichson, 1842) wardangensis (Brust, Ballard, Driver, Hartley, Galway & Curran, 1998) Heizmannia Ludlow, 1905 Subgenus Heizmannia aurea Brug, 1932 aureochaeta (Leicester, 1908) carteri Amerasinghe, 1993 chandi Edwards, 1922 chengi Lien, 1968 communis (Leicester, 1908) complex (Theobald, 1910) covelli Barraud, 1929 demeilloni Mattingly, 1970 funerea (Leicester, 1908) greenii (Theobald, 1905) heterospina Gong & Lu, 1986 himalayensis Edwards, 1922 indica (Theobald, 1905) kana Tanaka, Mizusawa & Saugstad, 1979 kanhsienensis Tung, 1955 lii Wu, 1936 macdonaldi Mattingly, 1957 mattinglyi Thurman, 1959 menglianensis Lu & Gong, 1986 menglianeroides Dong, Dong & Zhou, 2003 persimilis Mattingly, 1970 propinqua Mattingly, 1970 proxima Mattingly, 1970 reidi Mattingly, 1957 ruiliensis Dong, Zhou & Wang, 1997 scanloni Mattingly, 1970 scintillans Ludlow, 1905 taiwanensis Lien, 1968 tengchongensis Dong, Wang & Zhou, 2002 thelmae Mattingly, 1970 viridis Barraud, 1929 Subgenus Mattinglyia Lien, 1968 achaetae (Leicester, 1908) catesi (Lien, 1968) discrepans (Edwards, 1922) occidentayunnana (Gong & Lu, 1991) tripunctata (Theobald, 1908)	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFAD0D69C88BFD7F072E30CA.taxon	description	Howardina Theobald, 1903 albonotata (Coquillett, 1906) allotecnon (Kumm, Komp & Ruiz, 1940) arborealis (Bonne-Wepster & Bonne, 1920) argyriti (Dyar & Nunez Tovar, 1927) aureolineata (Berlin, 1969) auriti Theobald, 1907 aurivittata (Cerqueira, 1943) bahamensis (Berlin, 1969) brevis (Berlin, 1969) brevivittata (Berlin, 1969) busckii (Coquillett, 1906) cozumelensis (Diaz Najera, 1966) ecuadoriensis (Berlin, 1969) eleanorae (Berlin, 1969) fulvithorax (Lutz, 1904) grabhami (Berlin, 1969) guatemala (Berlin, 1969) guerrero (Berlin, 1969) inaequalis Grabham, 1907 ioliota (Dyar & Knab, 1913) leei (Berlin, 1969) lorraineae (Berlin, 1969) marinkellei (Berlin, 1969) martinezi (Berlin, 1969) osornoi (Berlin, 1969) pseudodminicii (Komp, 1936) quadrivittata (Coquillett, 1902) septemstriata (Dyar & Knab, 1907) sexlineata (Theobald, 1901) spinosa (Berlin, 1969) stenei (Thompson, 1956) vanemdeni (Martini, 1931) walkeri (Theobald, 1901) whitmorei (Dunn, 1918) Huaedes Huang, 1968 medialis (Brug, 1932) variepictus (King & Hoogstraal, 1946) wauensis (Huang, 1968) Hulecoeteomyia Theobald, 1904, stat. nov., restored to genus rank chrysolineata (Theobald, 1907), comb. nov. formosensis (Yamada, 1921), comb. nov. harveyi (Barraud, 1923), comb. nov. japonica (Theobald, 1901), comb. nov. ssp. shintienensis (Tsai & Lien, 1950), comb. nov. ssp. amamiensis (Tanaka, Mizusawa & Saugstad, 1979), comb. nov. ssp. yaeyamensis (Tanaka, Mizusawa & Saugstad, 1979), comb. nov. jugraensis Leicester, 1908, comb. nov. koreica (Edwards, 1917), comb. nov. nigrorhynchus (Brug, 1931), comb. nov. pallirostris (Edwards, 1922), comb. nov. reinerti (Rattanarithikul & Harrison, 1988), comb. nov. rizali (Banks, 1906), comb. nov. saxicola (Edwards, 1922), comb. nov. sherki (Knight, 1948), comb. nov. yunnanensis (Gaschen, 1934) *, comb. nov. Indusius Edwards, 1934 pulverulentus (Edwards, 1922) Isoaedes Reinert, 1979 cavaticus (Reinert, 1979)	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFAD0D69CB13FB2607AA3258.taxon	description	Subgenus Jarnellius deserticola (Zavortink, 1969), comb. nov. laguna (Arnell & Nielsen, 1972), comb. nov. monticola (Belkin & McDonald, 1957), comb. nov. sierrensis (Ludlow, 1905), comb. nov. varipalpus (Coquillett, 1902), comb. nov. Subgenus Lewnielsenius Reinert, Harbach & Kitching, subgen. nov. muelleri (Dyar, 1920), comb. nov.	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFAD0D6ECB13F99401143070.taxon	description	Kenknightia Reinert, 1990 dissimilierodes (Dong, Zhou & Dong, 2002) dissimilis (Leicester, 1908) gaffigani (Reinert, 1990) harbachi (Reinert, 1990) karwari (Barraud, 1924) lerozeboomi (Reinert, 1990) leucomeres (Giles, 1904) litwakae (Reinert, 1990) luzonensis (Rozeboom, 1946) paradissimilis (Rozeboom, 1946) pecori (Reinert, 1990) wilkersoni (Reinert, 1990) Kompia Aitken, 1941 purpureipes (Aitken, 1941) Leptosomatomyia Theobald, 1905 aurimargo (Edwards, 1922) Levua Stone & Bohart, 1944 dufouri (Hamon, 1953) fryeri (Theobald, 1912) geoskusea (Amos, 1944) Lorrainea Belkin, 1962 amesii (Ludlow, 1903) celebica (Mattingly, 1959) dasyorrhus (King & Hoogstraal, 1946) fumida (Edwards, 1928) lamellifera (Bohart & Ingram, 1946)	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFAA0D6EC8A7FBCC0135327B.taxon	description	Subgenus Macleaya, transferred from ‘ Ochlerotatus ’ littlechildi (Taylor, 1933), comb. nov. stoneorum (Marks, 1977), comb. nov. tremula Theobald, 1903, comb. nov.	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFAA0D6DC8A7F9B600FF3184.taxon	description	alternans (Westwood, 1835) aurantius (Theobald, 1907) ssp. chrysogaster Taylor, 1927 ferinus (Knight, 1947) grahamii Theobald, 1909 laniger (Wiedemann, 1820) lucianus (Muspratt, 1959) mucidus (Karsch, 1887) nigerrimus Theobald, 1913 painei (Knight, 1948) quadripunctis (Ludlow, 1910) quasiferinus (Mattingly, 1961) scatophagoides Theobald, 1901 sudanensis Theobald, 1908 tonkingi (Gebert, 1948) Neomelaniconion Newstead, 1907 albicosta (Edwards, 1913) albothorax (Theobald, 1907) aurovenatum (Worth, 1960) bequaerti (Wolfs, 1947) bergerardi (Pajot & Geoffroy, 1971) bolense (Edwards, 1936) carteri (Edwards, 1936) circumluteolum (Theobald, 1908) crassiforceps (Edwards, 1927) ellinorae (Edwards, 1941) flavimargo (Edwards, 1941) fuscinerve (Edwards, 1914) jamoti (Hamon & Rickenbach, 1954) lineatopenne (Ludlow, 1905) ssp. aureum (Gutsevich, 1955) luridum (McIntosh, 1971) luteolaterale (Theobald, 1901) mcintoshi (Huang, 1985) monotrichus (Edwards, 1936) palpale Newstead, 1907 pogonurum (Edwards, 1936) punctocostale (Theobald, 1909) taeniarostre (Theobald, 1909) unidentatum (McIntosh, 1971) Ochlerotatus Lynch Arribalzaga, 1891 Subgenus Rusticoidus Shevchenko & Prudkina, 1973 albescens (Edwards, 1921) bicristatus (Thurman & Winkler, 1950) krymmontanus (Alekseev, 1989) lepidonotus (Edwards, 1920) provocans (Walker, 1848) quasirusticus (Torres Canamares, 1951) refiki (Medschid, 1928) rusticus (Rossi, 1790) var. subtrichurus (Martini, 1927) subdiversus (Martini, 1926) Unassigned to subgenus atlanticus (Dyar & Knab, 1906) communis (de Geer, 1776) dorsalis (Meigen, 1830) fulvus (Wiedemann, 1828) ssp. pallens (Ross, 1943) infirmatus (Dyar & Knab, 1906) scapularis (Rondani, 1848) sollicitans (Walker, 1856) ‘ Ochlerotatus ’ sensu auctorum Subgenus ‘ Finlaya ’ Theobald, 1903 sensu auctorum albocinctus (Barraud, 1924) albotaeniata (Leicester, 1904) alticola (Bonne-Wepster, 1948) auronitens (Edwards, 1922) australiensis (Theobald, 1910) barnardi (Edwards, 1924) biocellatus (Taylor, 1915) britteni (Marks & Hodgkin, 1958) candidoscutellum (Marks, 1947) crossi (Lien, 1967) eatoni Edwards, 1916 embuensis (Edwards, 1930) fengi (Edwards, 1935) gracilelineatus (Bonne-Wepster, 1937) harperi (Knight, 1948) ingrami (Edwards, 1930) keefei (King & Hoogstraal, 1946) koreicoides (Sasa, Kano & Hayashi, 1950) lauriei Carter, 1920 lepchana (Barraud, 1923) loi (Lien, 1968) mallochi (Taylor, 1944) monocellatus (Marks, 1948) nyasae (Edwards, 1930) oreophilus Edwards, 1916 peipingensis (Feng, 1938) plagosus (Marks, 1959) quasirubithorax (Theobald, 1910) roai (Belkin, 1962) rubiginosus (Belkin, 1962) seoulensis (Yamada, 1921) sintoni (Barraud, 1924) stanleyi (Peters, 1963) subauridorsum (Marks, 1948) suffusus (Edwards, 1922) toxopeusi (Bonne-Wepster, 1948) tsiliensis (King & Hoogstraal, 1946) versicolor (Barraud, 1924) wasselli (Marks, 1947) wellmanii (Theobald, 1905) Subgenus ‘ Ochlerotatus ’ Lynch Arribalzaga, 1891 sensu auctorum aboriginis (Dyar, 1917) abserratus (Felt & Young, 1904) aculeatus (Theobald, 1903) aenigmaticus (Cerqueira & Costa, 1946) akkeshiensis (Tanaka, 1998) albifasciatus (Macquart, 1838) albineus (Seguy, 1923) aloponotum (Dyar, 1917) ambreensis (Rodhain & Boutonnier, 1983) andersoni (Edwards, 1926) angustivittatus (Dyar & Knab, 1907) annulipes (Meigen, 1830) antipodeus Edwards, 1920 atactavittatus (Arnell, 1976) auratus (Grabham, 1906) aurifer (Coquillett, 1903) behningi (Martini, 1926) bejaranoi (Martinez, Carcavallo & Prosen, 1960) berlandi (Seguy, 1921) bimaculatus (Coquillett, 1902) biskraensis (Brunes, 1999) bogotanus (Arnell, 1976) breedensis (Muspratt, 1953) burjaticus (Kuchartshuk, 1973) burpengaryensis (Theobald, 1905) caballus (Theobald, 1912) cacozelus (Marks, 1963) calcariae (Marks, 1957) calumnior (Belkin, Heinemann & Page, 1970) campestris (Dyar & Knab, 1907) camptorhynchus (Thomson, 1869) canadensis (Theobald, 1901) ssp. mathesoni (Middlekauff, 1944) cantans (Meigen, 1818) var. subvexans (Martini, 1922) cantator (Coquillett, 1903) caspius (Pallas, 1771) ssp. meirai Ribeiro, da Cunha Ramos, Capela & Pires, 1980 var. hargreavesi Edwards, 1920 cataphylla (Dyar, 1916) var. rostochiensis (Martini, 1920) chelli Edwards, 1915 churchillensis (Ellis & Brust, 1973) clelandi (Taylor, 1914) clivis (Lanzaro & Eldridge, 1992) coluzzii (Rioux, Guilvard & Pasteur, 1998) comitatus (Arnell, 1976) condolescens (Dyar & Knab, 1907) continentalis (Dobrotworsky, 1960) crinifer (Theobald, 1903) cunabulanus (Edwards, 1924) cyprioides (Danilov & Stupin, 1982) cyprius (Ludlow, 1920) decticus (Howard, Dyar & Knab, 1917) deficiens (Arnell, 1976) detritus (Haliday, 1833) diantaeus (Howard, Dyar & Knab, 1913) duplex (Martini, 1926) dupreei (Coquillett, 1904) dzeta (Seguy, 1924) edgari (Stone & Rosen, 1952) eidsvoldensis (Mackerras, 1927) eucephalaeus (Dyar, 1918) euedes (Howard, Dyar & Knab, 1913) euiris (Dyar, 1922) euplocamus (Dyar & Knab, 1906) excrucians (Walker, 1856) explorator (Marks, 1964) fitchii (Felt & Young, 1904) flavescens (Müller, 1764) flavifrons (Skuse, 1889) grossbecki (Dyar & Knab, 1906) gutzevichi (Dubitsky & Deshevykh, 1978) hakusanensis (Yamaguti & Tamaboko, 1954) harrisoni (Muspratt, 1953) hastatus (Dyar, 1922) hesperonotius (Marks, 1959) hexodontus (Dyar, 1916) hodgkini (Marks, 1959) hokkaidensis (Tanaka, Mizusawa & Saugstad, 1979) hortator (Dyar & Knab, 1907) hungaricus (Mihalyi, 1955) imperfectus (Dobrotworsky, 1962) impiger (Walker, 1848) ssp. daisetsuzanus (Tanaka, Mizusawa & Saugstad, 1979) implicatus (Vockeroth, 1954) incomptus (Arnell, 1976) increpitus (Dyar, 1916) inexpectatus (Bonne-Wepster, 1948) intermedius (Danilov & Gornostaeva, 1987) intrudens (Dyar, 1919) jacobinae (Serafim & Davis, 1933) jorgi (Carpintero & Leguizamón, 2000) juppi (McIntosh, 1973) kasachstanicus (Gutsevich, 1962) lasaensis (Meng, 1962) ssp. gyirongensis (Ma, 1982) lepidus (Cerqueira & Paraense, 1945) leucomelas (Meigen, 1804) linesi (Marks, 1964) longifilamentus (Su & Zhang, 1988) luteifemur (Edwards, 1926) macintoshi (Marks, 1959) mariae (Sergent & Sergent, 1903) martineti (Senevet, 1937) mcdonaldi (Belkin, 1962) melanimon (Dyar, 1924) meprai (Martinez & Prosen, 1953) mercurator (Dyar, 1920) milleri (Dyar, 1922) mitchellae (Dyar, 1905) montchadskyi (Dubitsky, 1968) nevadensis (Chapman & Barr, 1964) nigrinus (Eckstein, 1918) nigripes (Zetterstedt, 1838) nigrithorax (Macquart, 1847) nigrocanus (Martini, 1927) nigromaculis (Ludlow, 1906) niphadopsis (Dyar & Knab, 1918) nivalis (Edwards, 1926) normanensis (Taylor, 1915) nubilis (Theobald, 1903) obturbator (Dyar & Knab, 1907) oligopistus (Dyar, 1918) patersoni (Shannon & del Ponte, 1928) pectinatus (Arnell, 1976) pennai (Antunes & Lane, 1938) perkinsi (Marks, 1949) pertinax (Grabham, 1906) perventor (Cerqueira & Costa, 1946) phaecasiatus (Marks, 1964) phaeonotus (Arnell, 1976) phoeniciae (Coluzzi & Sabatini, 1968) pionips (Dyar, 1919) procax Skuse, 1889 pseudonormanensis (Marks, 1949) pulcritarsis (Rondani, 1872) ssp. asiaticus (Edwards, 1926) pullatus (Coquillett, 1904) punctodes (Dyar, 1922) punctor (Kirby, 1837) purpuraceus (Brug, 1932) purpureifemur (Marks, 1959) purpuriventris (Edwards, 1926) ratcliffei (Marks, 1959) raymondi (del Ponte, Castro & Garcia, 1951) rempeli (Vockeroth, 1954) rhyacophilus (da Costa Lima, 1933) riparioides (Su & Zhang, 1987) riparius (Dyar & Knab, 1907) sagax (Skuse, 1889) sapiens (Marks, 1964) schizopinax (Dyar, 1929) schtakelbergi (Shingarev, 1928) scutellalbum (Boshell-Manrique, 1939) sedaensis (Lei, 1989) sergievi (Danilov, Markovich & Proskuryakova, 1978) serratus (Theobald, 1901) shannoni (Vargas & Downs, 1950) silvestris (Dobrotworsky, 1961) simanini (Gutsevich, 1966) sinkiangensis (Hsiao, 1977) spencerii (Theobald, 1901) ssp. idahoensis (Theobald, 1903) spilotus (Marks, 1963) squamiger (Coquillett, 1902) sticticus (Meigen, 1838) stigmaticus (Edwards, 1922) stimulans (Walker, 1848) stramineus (Dubitzky, 1970) stricklandi Edwards, 1912 subalbirostris (Klein & Marks, 1960) surcoufi (Theobald, 1912) synchytus (Arnell, 1976) taeniorhynchus (Wiedemann, 1821) tahoensis (Dyar, 1916) thelcter (Dyar, 1918) theobaldi (Taylor, 1914) thibaulti (Dyar & Knab, 1910) tormentor (Dyar & Knab, 1906) tortilis (Theobald, 1903) trivittatus (Coquillett, 1902) turneri (Marks, 1963) upatensis (Anduze & Hecht, 1943) ventrovittis (Dyar, 1916) vigilax (Skuse, 1889) ssp. ludlowae (R. Blanchard, 1905) ssp. vansomerenae (Mattingly, 1955) vittiger (Skuse, 1889) washinoi (Lanzaro & Eldridge, 1992) zammitii (Theobald, 1903) Subgenus ‘ Protomacleaya ’ Theobald, 1907 sensu auctorum aitkeni (Schick, 1970) alboapicus (Schick, 1970) amabilis (Schick, 1970) argyrothorax (Bonne-Wepster & Bonne, 1920) berlini (Schick, 1970) bertrami (Schick, 1970) braziliensis (Gordon & Evans, 1922) brelandi (Zavortink, 1972) buenaventura (Schick, 1970) burgeri (Zavortink, 1972) campana (Schick, 1970) casali (Schick, 1970) chionotum (Zavortink, 1972) daryi (Schick, 1970) diazi (Schick, 1970) gabriel (Schick, 1970) galindoi (Schick, 1970) hendersoni (Cockerell, 1918) heteropus (Dyar, 1921) homoeopus (Dyar, 1922) idanus (Schick, 1970) impostor (Schick, 1970) insolitus (Coquillett, 1906) knabi (Coquillett, 1906) kompi (Vargas & Downs, 1950) metoecopus (Dyar, 1925) niveoscutum (Zavortink, 1972) podographicus (Dyar & Knab, 1906) sandrae (Zavortink, 1972) schicki (Zavortink, 1972) schroederi (Schick, 1970) sumidero (Schick, 1970) tehuantepec (Schick, 1970) terrens (Walker, 1856) thorntoni (Dyar & Knab, 1907) triseriatus (Say, 1823) vargasi (Schick, 1970) zavortinki (Schick, 1970) zoosophus (Dyar & Knab, 1918) Subgenus uncertain daliensis (Taylor, 1916) Opifex Hutton, 1902 Subgenus Nothoskusea Dumbleton, 1962 chathamicus (Dumbleton, 1962) Subgenus Opifex fuscus Hutton, 1902 Paraedes Edwards, 1934 barraudi Edwards, 1934 bonneae (Mattingly, 1958) chrysoscuta (Theobald, 1910) collessi (Mattingly, 1958) menoni (Mattingly, 1958) ostentatio (Leicester, 1908) pagei (Ludlow, 1911) thailandensis (Reinert, 1976) Patmarksia Reinert, Harbach & Kitching, gen. nov. anggiensis (Bonne-Wepster, 1937), comb. nov. argenteitarsis (Brug, 1932), comb. nov. argyronotum (Belkin, 1962), comb. nov. buxtoni (Belkin, 1962), comb. nov. clintoni (Taylor, 1946), comb. nov. derooki (Brug, 1932), comb. nov. dobodura (King & Hoogstraal, 1946), comb. nov. hollandia (King & Hoogstraal, 1946), comb. nov. mackerrasi (Taylor, 1927), comb. nov. novalbitarsis (King & Hoogstraal, 1946), comb. nov. palmarum (Edwards, 1924), comb. nov. papuensis (Taylor, 1914), comb. nov. subalbitarsis (King & Hoogstraal, 1946), comb. nov.	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FFA90D52CB13F9F8028136D9.taxon	description	Pseudoarmigeres Stone & Knight, 1956 albomarginatus (Newstead, 1907) argenteoventralis (Theobald, 1909) ssp. dunni (Evans, 1928) kummi (Edwards, 1930) michaelikati (van Someren, 1946) ssp. gurneri (van Someren, 1946) natalensis (Edwards, 1930)	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FF960D52C8A7FD2C070037DC.taxon	description	Subgenus Grabhamia Theobald, 1903 cingulata (Fabricius, 1805) columbiae (Dyar & Knab, 1906) confinnis (Lynch Arribalzaga, 1891) dimidiata Cerqueira, 1943 discolor (Coquillett, 1903) infinis (Dyar & Knab, 1906) insularia (Dyar & Knab, 1906) jamaicensis Theobald, 1901 leucocnemis Martini, 1931 paulli Paterson & Shannon, 1927 pruinosa Martini, 1935 pygmaea (Theobald, 1903) santamarinai Broche, 2000 signipennis (Coquillett, 1904) varinervis Edwards, 1922 Subgenus Janthinosoma Lynch Arribalzaga, 1891 albigenu (Peryassu, 1908) albipes (Theobald, 1907) amazonica Cerqueira, 1960 champerico (Dyar & Knab, 1906) circumflava Cerqueira, 1943 cyanescens (Coquillett, 1902) discrucians (Walker, 1856) ferox (von Humboldt, 1819) fiebrigi Edwards, 1922 forceps Cerqueira, 1939 horrida (Dyar & Knab, 1908) johnstonii (Grabham, 1905) lanei Shannon & Cerqueira, 1943 longipalpus Randolph & O’Neill, 1944 lutzii (Theobald, 1901) mathesoni Belkin & Heinemann, 1975 melanota Cerqueira, 1943 mexicana (Bellardi, 1859) pilosa Duret, 1971 pseudoalbipes Duret, 1971 pseudomelanota Barata & Cotrim, 1971 totonaci Lassmann, 1951 varipes (Coquillett, 1904) Subgenus Psorophora ciliata (Fabricius, 1794) cilipes (Fabricius, 1805) holmbergii Lynch Arribalzaga, 1891 howardii Coquillett, 1901 lineata (von Humboldt, 1819) ochripes (Macquart, 1850) pallescens Edwards, 1922 pilipes (Macquart, 1834) saeva Dyar & Knab, 1906 stonei Vargas, 1956 Nomina dubia goeldii (Giles, 1904) marmorata (Philippi, 1865) perterrens (Walker, 1856)	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FF960D50CB0FFC180285312D.taxon	description	Rhinoskusea Edwards, 1929 longirostris (Leicester, 1908) pillaii (Mattingly, 1958) portonovoensis (Tewari & Hiriyan, 1992) wardi (Reinert, 1976) Scutomyia Theobald, 1904 albolineata Theobald, 1904 arboricola (Knight & Rozeboom, 1946) bambusicola Knight & Rozeboom, 1946 boharti (Knight & Rozeboom, 1946) hoogstraali (Knight & Rozeboom, 1946) impatibilis (Walker, 1859) laffooni (Knight & Rozeboom, 1946) platylepida (Knight & Hull, 1951) pseudalbolineata (Brug, 1939) Skusea Theobald, 1903 cartroni (Ventrillon, 1906) lambrechti (van Someren, 1971) moucheti (Ravaonjanahary & Brunhes, 1977) pembaensis (Theobald, 1901) Stegomyia Theobald, 1901 aegypti (Linnaeus, 1762) ssp. formosus (Walker, 1848) africana Theobald, 1901 agrihanensis (Bohart, 1957) albopicta (Skuse, 1895) alcasidi (Huang, 1972) alorensis (Bonne-Wepster & Brug, 1932) amalthea (de Meillon & Lavoipierre, 1944) andrewsi (Edwards, 1926) angusta (Edwards, 1935) annandalei Theobald, 1910 var. quadricincta (Barraud, 1923) aobae (Belkin, 1962) apicoargentea Theobald, 1909 bambusae (Edwards, 1935) blacklocki (Evans, 1925), comb. nov. bromeliae Theobald, 1911 burnsi (Basio & Reisen, 1971) calceata (Edwards, 1924) chaussieri (Edwards, 1923) chemulpoensis (Yamada, 1921) contigua (Edwards, 1936) cooki (Belkin, 1962) corneti (Huang, 1986) cozi (Cornet, 1973) craggi Barraud, 1923 cretina (Edwards, 1921) daitensis (Miyagi & Toma, 1981) deboeri (Edwards, 1926) demeilloni (Edwards, 1936) denderensis (Wolfs, 1949) dendrophila (Edwards, 1921) desmotes Giles, 1904 dybasi (Bohart, 1957) ealaensis (Huang, 2004), comb. nov. edwardsi Barraud, 1923 ethiopiensis (Huang, 2004), comb. nov. flavopicta (Yamada, 1921) ssp. downsi (Bohart & Ingram, 1946) ssp. miyarai (Tanaka, Mizusawa & Saugstad, 1979) fraseri Edwards, 1912 futunae (Belkin, 1962) galloisi (Yamada, 1921) galloisiodes (Liu & Lu, 1984) gandaensis (Huang, 2004), comb. nov. gardnerii Ludlow, 1905 ssp. imitator Leicester, 1908 grantii Theobald, 1901 guamensis (Farner & Bohart, 1944) gurneyi (Stone & Bohart, 1944) hakanssoni (Knight & Hurlbut, 1949) hansfordi (Huang, 1997) hebridea (Edwards, 1926) heischi (van Someren, 1951) hensilli (Farner, 1945) hogsbackensis (Huang, 2004), comb. nov. hoguei (Belkin, 1962) horrescens (Edwards, 1935) josiahae (Huang, 1988) katherinensis (Woodhill, 1949) keniensis (van Someren, 1946) kenyae (van Someren, 1946) kesseli (Huang & Hitchcock, 1980) kivuensis (Edwards, 1941) krombeini (Huang, 1975) lamberti (Ventrillon, 1904) langata (van Someren, 1946) ledgeri (Huang, 1981) lilii Theobald, 1910 luteocephala Newstead, 1907 maehleri (Bohart, 1957) malayensis (Colless, 1962) malikuli (Huang, 1973) marshallensis (Stone & Bohart, 1944) mascarensis (MacGregor, 1924) masseyi (Edwards, 1923) mattinglyorum (Huang, 1994) maxgermaini (Huang, 1990) mediopunctata Theobald, 1905 var. sureilensis (Barraud, 1934) metallica (Edwards, 1912) mickevichae (Huang, 1988) mpusiensis (Huang, 2004), comb. nov. muroafcete (Huang, 1997) neoafricana (Cornet, Valade & Dieng, 1978) neogalloisi (Chen & Chen, 2000) neopandani (Bohart, 1957) njombiensis (Huang, 1997) novalbopicta (Barraud, 1931) opok (Corbet & van Someren, 1962) palauensis (Bohart, 1957) pandani (Stone, 1939) patriciae (Mattingly, 1954) paullusi (Stone & Farner, 1945) pernotata (Farner & Bohart, 1944) perplexa Leicester, 1908 polynesiensis (Marks, 1951) poweri Theobald, 1905 pseudalbopicta Borel, 1928 pseudoafricana (Chwatt, 1949) pseudonigeria Theobald, 1910 pseudoscutellaris Theobald, 1910 quasiscutellaris (Farner & Bohart, 1944) rhungkiangensis (Chang & Chang, 1974) riversi (Bohart & Ingram, 1946) robinsoni (Belkin, 1962) rotana (Bohart & Ingram, 1946) rotumae (Belkin, 1962) ruwenzori (Haddow & van Someren, 1950) saimedres (Huang, 1988) saipanensis (Stone, 1945) sampi (Huang, 2004), comb. nov. schwetzi (Edwards, 1926) scutellaris (Walker, 1858) scutoscripta (Bohart & Ingram, 1946) seampi (Huang, 1974) seatoi (Huang, 1969) segermanae (Huang, 1997) sibirica (Danilov & Filippova, 1978) simpsoni Theobald, 1905 soleata (Edwards, 1924) strelitziae (Muspratt, 1950) subalbopicta (Barraud, 1931) subargentea (Edwards, 1925) tabu (Ramalingam & Belkin, 1965) tongae (Edwards, 1926) tulagiensis (Edwards, 1926) unilineata (Theobald, 1906) upolensis (Marks, 1957) usambara (Mattingly, 1953) varuae (Belkin, 1962) vinsoni (Mattingly, 1953) wadai (Tanaka, Mizusawa & Saugstad, 1979) w-albus Theobald, 1905 woodi (Edwards, 1922) Nomina dubia ciliaris (Linnaeus, 1767) niveus (von Eichwald, 1837) rimandoi (Basio, 1971) Tanakaius Reinert, Harbach & Kitching, 2004 savoryi (Bohart, 1957) togoi (Theobald, 1907) Udaya Thurman, 1954 argyrurus (Edwards, 1934) lucaris Macdonald & Mattingly, 1960 subsimilis (Barraud, 1927)	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
35378770FF940D51C8A7FA8F02F53315.taxon	description	Verrallina Theobald, 1903 Subgenus Harbachius Reinert, 1999 abdita (Barraud, 1931) consonensis (Reinert, 1973) fragilis Leicester, 1908 hamistylus (Laffoon, 1946) indecorabilis Leicester, 1908 nobukonis (Yamada, 1932) pahangi (Delfinado, 1968) ramalingami (Reinert, 1974) robertsi (Laffoon, 1946) srilankensis (Reinert, 1977) stunga (Klein, 1973) uniformis (Theobald, 1910) yusafi (Barraud, 1931) Subgenus Neomacleaya Theobald, 1907 adusta (Laffoon, 1946) agrestis (Barraud, 1931) andamanensis (Edwards, 1922) assamensis Bhattacharyya, Tewari, Prakash, Mohapatra & Mahanta, 2004 atriisimilis (Tanaka & Mizusawa, 1973) atria (Barraud, 1928) campylostylus (Laffoon, 1946) cauta (Barraud, 1928) clavata (Barraud, 1931) comata (Barraud, 1931) comosa (Reinert, 1974) cretata (Delfinado, 1967 cyrtolabis (Edwards, 1928) gibbosa (Delfinado, 1967) harrisonica (Reinert, 1974) hispida (Delfinado, 1967) incerta (Edwards, 1922) indica (Theobald, 1907) johnsoni (Laffoon, 1946) johorensis (Reinert, 1974) komponga (Klein, 1973) lankaensis (Stone & Knight, 1958) latipennis (Delfinado, 1967) leicesteri (Edwards, 1917) macrodixoa (Dyar & Shannon, 1925) margarsen (Dyar & Shannon, 1925) neomacrodixoa (King & Hoogstraal, 1947) nigrotarsis (Ludlow, 1908) notabilis (Delfinado, 1967) nubicola (Laffoon, 1946) panayensis (Ludlow, 1914) petroelephantus (Wijesundara, 1951) philippinensis (Delfinado, 1968) phnoma (Klein, 1973) prioekanensis (Brug, 1931) protuberans (Delfinado, 1967) pseudodiurna (Theobald, 1910) pseudomediofasciata (Theobald, 1910) pseudovarietas (Reinert, 1974) rami (Barraud, 1928) rara (Delfinado, 1968) sabahensis (Reinert, 1974) seculata (Menon, 1950) singularis (Leicester, 1908) sohni (Reinert, 1974) spermathecus (Wijesundara, 1951) torosa (Delfinado, 1967) uncus (Theobald, 190 l) vallistris (Barraud, 1928) varietas (Leicester, 1908) virilis Leicester, 1908 yerburyi (Edwards, 1917) Subgenus Verrallina azureosquamata (Bonne-Wepster, 1948) bifoliata (King & Hoogstraal, 1947) butleri (Theobald, 1901) carmenti (Edwards, 1924) cuccioi (Belkin, 1962) cunninghami (Taylor, 1944) dux (Dyar & Shannon, 1925) embiensis (Huang, 1968) foliformis (King & Hoogstraal, 1947) funerea (Theobald, 1903) iriomotensis (Tanaka & Mizusawa, 1973) killertonis (Huang, 1968) leilae (King & Hoogstraal, 1947) lineata (Taylor, 1914) lugubris (Barraud, 1928) mccormicki (Belkin, 1962) milnensis (King & Hoogstraal, 1947) multifolium (King & Hoogstraal, 1947) obsoleta (Huang, 1968) parasimilis (King & Hoogstraal, 1947) pipkini (Bohart, 1957) quadrifolium (Brug, 1934) quadrispinata (King & Hoogstraal, 1947) reesi (King & Hoogstraal, 1947) sentania (King & Hoogstraal, 1947) similis (Theobald, 1910) simpla (King & Hoogstraal, 1947) trispinata (King & Hoogstraal, 1947) vanapa (Huang, 1968) variabilis (Huang, 1968) Nomen dubium perdita Leicester, 1908 Zavortinkius Reinert, 1999 brunhesi (Reinert, 1999) brygooi (Brunhes, 1971) fulgens (Edwards, 1917) geoffroyi (Reinert, 1999) huangae (Reinert, 1999) interruptus (Reinert, 1999) longipalpis (Grünberg, 1905) monetus (Edwards, 1935) mzooi (van Someren, 1962) phillipi (van Someren, 1949) pollinctor (Graham, 1910) Zeugnomyia Leicester, 1908 aguilari Baisas & Feliciano, 1953 fajardoi Baisas & Feliciano, 1953 gracilis Leicester, 1908 lawtoni Baisas, 1946	en	Reinert, John F., Harbach, Ralph E., Kitching, Ian J. (2006): Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society 148 (1): 1-101, DOI: 10.1111/j.1096-3642.2006.00254.x, URL: https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2006.00254.x
