taxonID	type	description	language	source
3B0087DD8C78043CD3BAB99AEA1FD488.taxon	diagnosis	Diagnosis. Anthidulum is characterized by the presence of a short juxtantennal carina; margin of pronotal lobe with a continuous lamina on anterior and lateral edges; omaular carina short; arolia present in the male and reduced in the female; male with seventh tergum bilobed and third sternum somewhat enlarged. The male genitalia have the gonocoxite wider than long and the gonostylus curved in the middle (Fig. 25).	en	Parizotto, Daniele R., Urban, Danúncia (2013): Contribution to the knowledge of the Anthidulum Michener and Ctenanthidium Urban (Hymenoptera, Apidae) with new species from Argentina and Peru. Zootaxa 3609 (3): 311-318, DOI: 10.11646/zootaxa.3609.3.5
3B0087DD8C79043DD3BABFA3EDD1D4D8.taxon	discussion	Comments and diagnosis. Head black with small yellow spot behind eyes (Figs. 3 – 6). Tegula with semicircular light yellow macula; all terga with light yellow bands (Figs. 4 – 6), medially interrupted on T 1 - T 3 (Fig. 4). Male: clypeus, supraclypeal and lower paraocular area light yellow (Fig. 1); T 7 with large lateral lobe (Fig. 7). The semicircular macula of tegula and the T 7 with a large lobe are unique characteristics of this species.	en	Parizotto, Daniele R., Urban, Danúncia (2013): Contribution to the knowledge of the Anthidulum Michener and Ctenanthidium Urban (Hymenoptera, Apidae) with new species from Argentina and Peru. Zootaxa 3609 (3): 311-318, DOI: 10.11646/zootaxa.3609.3.5
3B0087DD8C79043DD3BABFA3EDD1D4D8.taxon	description	Description. Holotype male. Approximate body length 6.1 mm; forewing length 5.4 mm; head width 2.3 mm; eye length 1.5 mm. Color: Integument predominantly black except with yellow as follows: clypeus, lower paraocular area to just above antennal socket, supraclypeal area superiorly concave (Fig. 1) and a small spot on vertex, just behind eyes (Figs. 3, 5). Mandible with yellowish brown dorsal macula (Fig. 1). Antenna brown. Mesoscutum with laterobasal light yellow band; scutellum with narrow distal yellow band, medially interrupted. Tegula brown with semicircular light yellow macula; wing membrane brown, darker at base and at costal margin (Fig. 3). Legs blackened, fore and middle with irregular yellow maculae on base and apex of tibiae and yellowish ferruginous tinged on internal surface; middle leg with light yellow basal macula on external surface of basitarsus; hind leg with femoral apex and tibial base yellowish brown; basitarsus light yellow with brown apex. Terga black; T 1 with lateral light yellow bands widely separated; T 2 with narrow light yellow band medially interrupted; T 3 band little narrower than T 2 and weakly interrupted medially; T 4 - T 5 with narrow, complete bands, irregular on sides; T 6 with large band, anteriorly and posteriorly sinuous, and distal margin translucent ferruginous; T 7 light yellow with narrow black base and brownish distal margin (Fig. 7). Sterna blackish basally, brown at middle and with translucent ferruginous margin. Pilosity: Mostly white and sparse. Hairs up to longer than twice median ocellar diameter; pilosity plumose, conspicuous on sides of mesepisternum. Second sternum with apical fringe of long and plumose hairs; hairs shorter at middle; S 3 with sparse fringe laterally, denser at middle. Punctures: Head and mesosoma densely punctate; punctures contiguous and slightly larger on mesosoma. Metasoma densely punctate, punctures shallower than on mesosoma. Structure: T 7 with wide and flat lateral lobes, space between lobes as wide as lobe width (Fig. 7). Description. Paratype female. Differs from male in the following characteristics: Approximate body length 6.2 mm; forewing length 4.8 mm; head width 2.2 mm; eye length 1.5 mm. Color: Head almost completely black, except from small yellow spot on vertex, just behind eyes (Fig. 2). Pronotal lobe with light yellow spot on apex (Fig. 4). Fore and middle legs with yellowish brown area on tibial base (Fig. 6). First tergum with large light yellow lateral band; T 2 - T 3 with narrower bands, closer than T 1; T 4 - T 5 with complete light yellow bands and T 6 black. Pilosity and Punctures: As in male except by more conspicuous pilosity in paraocular area. Scopa light yellow.	en	Parizotto, Daniele R., Urban, Danúncia (2013): Contribution to the knowledge of the Anthidulum Michener and Ctenanthidium Urban (Hymenoptera, Apidae) with new species from Argentina and Peru. Zootaxa 3609 (3): 311-318, DOI: 10.11646/zootaxa.3609.3.5
3B0087DD8C79043DD3BABFA3EDD1D4D8.taxon	materials_examined	Type material. Holotype male. PERU, Lima: “ PERU: Lima Dept. / Sta. Rosa de Quives, / near Yangas. 1100 m. / v- 28 - 29 - 96 / J. C. Rozen, A. Ugarte ” (AMNH). Paratypes: one male and one female of with the same data of holotype (AMNH); one male with the same data of holotype (DZUP); one male with the same data, except by date ‘ v- 28 - 96 ’ (AMNH); one male without metasoma: “ PERU: Lima Dept. / Ricardo Palma, V- 9 - 96 / J. C. Rozen, A. Ugarte ” (AMNH) and one female with the same data (DZUP); one male: “ PERU: Lambayeque / Dept. 20 km E. Olmos / v- 21 - 96 500 – 1000 m. / J. C. Rozen, A. Ugarte ” (AMNH).	en	Parizotto, Daniele R., Urban, Danúncia (2013): Contribution to the knowledge of the Anthidulum Michener and Ctenanthidium Urban (Hymenoptera, Apidae) with new species from Argentina and Peru. Zootaxa 3609 (3): 311-318, DOI: 10.11646/zootaxa.3609.3.5
3B0087DD8C79043DD3BABFA3EDD1D4D8.taxon	etymology	Etymology. The specific epithet honors Dr. Jerome G. Rozen, American Museum of Natural History, New York.	en	Parizotto, Daniele R., Urban, Danúncia (2013): Contribution to the knowledge of the Anthidulum Michener and Ctenanthidium Urban (Hymenoptera, Apidae) with new species from Argentina and Peru. Zootaxa 3609 (3): 311-318, DOI: 10.11646/zootaxa.3609.3.5
3B0087DD8C79043DD3BABFA3EDD1D4D8.taxon	description	Variation. All male paratypes have a yellow spot on the pronotal lobe. In the paratype male collected from Lambayeque, the yellow maculae of mesosoma and metasoma are larger than the remaining specimens.	en	Parizotto, Daniele R., Urban, Danúncia (2013): Contribution to the knowledge of the Anthidulum Michener and Ctenanthidium Urban (Hymenoptera, Apidae) with new species from Argentina and Peru. Zootaxa 3609 (3): 311-318, DOI: 10.11646/zootaxa.3609.3.5
3B0087DD8C79043DD3BABFA3EDD1D4D8.taxon	discussion	Comments. Anthidulum rozeni differs from all other species in the shape of T 7, by its wide lateral lobes, separated by narrow emargination. All other species have narrower lateral lobes, widely separated by more than three times the base of lobe width. Moreover, the new species has a small yellow spot behind the eyes while all other species have a yellow or ferruginous postocellar band extending across the vertex. The two other species of Anthidulum known from Peru, Anthidulum lamasi Urban, 2003 and Anthidulum clausi Urban, 2003, have the vertex with a yellow postocellar band and the mesoscutum with a band along the anterior and lateral margins. Anthidulum lamasi has the supraclypeal area black, T 4 - T 5 almost totally yellow and A. clausi has a large yellow spot on the mesepisternum laterally and T 4 - T 7 are predominantly yellow.	en	Parizotto, Daniele R., Urban, Danúncia (2013): Contribution to the knowledge of the Anthidulum Michener and Ctenanthidium Urban (Hymenoptera, Apidae) with new species from Argentina and Peru. Zootaxa 3609 (3): 311-318, DOI: 10.11646/zootaxa.3609.3.5
3B0087DD8C7A043ED3BAB99DECA8D4C0.taxon	diagnosis	Diagnosis. Ctenanthidium can be defined by the juxtantennal carina strong and laminated; female with apex of labrum with hairs; margin of pronotal lobe with lamina interrupted on lateral edge; arolia well developed in both sexes; S 4 - S 5 of male with lateral projections; sixth sternum with a transverse integumentary fold. The genitalia is characterized by having a gonocoxite longer than wide and gonostylus curved on the apical half (Fig. 26). Additionally, an important character indicated by its generic name, is the third sternum of the male with a comb of coarse black setae on the distal margin medially.	en	Parizotto, Daniele R., Urban, Danúncia (2013): Contribution to the knowledge of the Anthidulum Michener and Ctenanthidium Urban (Hymenoptera, Apidae) with new species from Argentina and Peru. Zootaxa 3609 (3): 311-318, DOI: 10.11646/zootaxa.3609.3.5
3B0087DD8C7B0439D3BABFA3EFA4D020.taxon	discussion	Comments and diagnosis. Head predominantly tinged by yellow and ferruginous; mesoscutum with a reverse Lshaped macula, yellow along anterior edges and ferruginous along lateral edges. Integument of mesoscutum smooth between punctures (Figs. 13 – 14). Labrum with subapical hairs, plumose in the male; thickened and simple in the female (Fig. 10). The color pattern of head and mesoscutum does not occur in other species and the features of labrum pilosity are clearly unique within the genus.	en	Parizotto, Daniele R., Urban, Danúncia (2013): Contribution to the knowledge of the Anthidulum Michener and Ctenanthidium Urban (Hymenoptera, Apidae) with new species from Argentina and Peru. Zootaxa 3609 (3): 311-318, DOI: 10.11646/zootaxa.3609.3.5
3B0087DD8C7B0439D3BABFA3EFA4D020.taxon	description	Description. Holotype male. Approximate body length 5.5 mm; forewing length 4.5 mm; head width 2.0 mm; eye length 1.3 mm. Color: Integument predominantly black. Head widely tinged by yellow and ferruginous; yellow colored in mandible (except teeth and margins), clypeus, supraclypeal area and lower paraocular area; tinged by yellow and ferruginous in interalveolar area including juxtantennal carina; ferruginous in upper paraocular area extending to vertex and all of gena. Antenna with four basal articles amber, remaining flagellomeres ferruginous (Fig. 8). Mesosoma and metasoma predominantly black except as follows: pronotal lobe with apical ferruginous spot; mesoscutum with reverse L-shaped macula, yellow along anterior edge and ferruginous along lateral edge; axilla and scutellum with yellow subapical band and translucent margin. Tegula amber with pale yellow anterior area; wing membrane mostly black infumated, darker at base and at costal margin (Fig. 13). Legs predominantly amber; middle leg with coxa and trochanter blackened; hind leg with nearly entire coxa, trochanter, femur and two distal tarsomeres blackened (Fig. 17). First tergum with yellow band, narrower at middle, and area posterior to band ferruginous; T 2 with lateral yellow macula and very narrow, obsolete, ferruginous dorsal band; T 3 - T 5 with yellow complete band, wider at middle and with area posterior to band ferruginous, apically translucent (Fig. 24); T 6 - T 7 with apical yellow band. Sterna brown; S 1 with yellowish ferruginous medial area; S 2 with three apical yellow spots; S 3 with apical yellow margin, disc yellowish brown. Pilosity. Predominantly white; labrum with subapical plumose hairs; lower paraocular area, lateral and ventral surfaces of mesepisternum, coxae, trochanters, femora and tibiae with plumose hairs. Terga with translucent, short (<1 PD) yellow setae. Sterna with longer (> 1 PD) plumose hairs, denser and longer at sides. Punctures. Head densely punctate, sparser and more shallow on clypeus and supraclypeal area. Mesoscutum and mesepisternum with punctures slightly larger and deeper than on head, integument smooth between punctures. Scutellum with punctures slightly larger than mesoscutum. Tegula with shallow punctures at sides, dorsally smooth (Fig. 16). Terga with punctures sparser and shallow than on mesoscutum. Distance between punctures at least a puncture diameter. T 2 with smaller, sparser and shallower punctures than T 1. T 3 - T 6 with larger, denser and deeper punctures. T 7 with smaller punctures and even denser than T 6 (Fig. 24). Description. Paratype female. Differs from male in the following characteristics: Approximate body length 6.6 mm; forewing length 4.7 mm; head width 2.1 mm; eye length 1.4 mm. Color: mandibles blackened; clypeus with large basal ferruginous macula, supraclypeal area and juxtantennal carina blackened (Fig. 9). Fore and middle legs with coxae and trochanters darkish brown; remaining areas of fore leg ferruginous; middle leg with brown femur, except apex ferruginous, tibia ferruginous and tarsomeres brown; hind leg dark brown with ferruginous areas on apex of femora and base of tibiae. Terga black; T 1 and T 3 with yellow complete band; T 2 with lateral yellow maculae; T 4 - T 5 with traces of band, leaving a ferruginous area (Fig. 14, 19). Sterna black. Pilosity whitish, including scopa. Subapical setae on labrum thickened. (Fig. 10). Punctures as in male.	en	Parizotto, Daniele R., Urban, Danúncia (2013): Contribution to the knowledge of the Anthidulum Michener and Ctenanthidium Urban (Hymenoptera, Apidae) with new species from Argentina and Peru. Zootaxa 3609 (3): 311-318, DOI: 10.11646/zootaxa.3609.3.5
3B0087DD8C7B0439D3BABFA3EFA4D020.taxon	materials_examined	Type material. Holotype male. Argentina, Tucumán. “ ARGENTINA, Tucumán: / 12 km N El Cadillal / December 3, 1989 / J. G. Rozen & A. Roig ” (AMNH). Paratypes males: one specimen with the same date of holotype (DZUP); “ ARGENTINA / Tucumán: 11 Km NW / Cadillal XII- 4 - 1993 / J. G. Rozen ” (AMNH); “ ARGENTINA: Tucumán / 11 Km NW Cadillal / XII- 1 - 93 J. G. Rozen ” (AMNH); “ ARG. – SALTA / Rosario Lerma / Fritz - 11.84 ” (AMNH); “ ARG. – SALTA / Alemania / Peña - 2.92 ” (AMNH); “ ARG. – SALTA / Cnel. [Coronel] Moldes / Fritz - 2.95 ” (AMNH); “ ARG. – Córdoba / Quilpo / Fritz - 2.80 (AMNH). Paratypes females: “ ARG. – SALTA / Rosario Lerma / Fritz – X. 85 ” (DZUP); one specimen with the same date above except by “ Fritz – XII. 86 ” (AMNH); “ ARG. – SALTA / Sumalao / Fritz – II. 96 ” (AMNH).	en	Parizotto, Daniele R., Urban, Danúncia (2013): Contribution to the knowledge of the Anthidulum Michener and Ctenanthidium Urban (Hymenoptera, Apidae) with new species from Argentina and Peru. Zootaxa 3609 (3): 311-318, DOI: 10.11646/zootaxa.3609.3.5
3B0087DD8C7B0439D3BABFA3EFA4D020.taxon	etymology	Etymology. From the Latin versicolor (= various colors) in reference to the coloration from head and mesosoma of this species.	en	Parizotto, Daniele R., Urban, Danúncia (2013): Contribution to the knowledge of the Anthidulum Michener and Ctenanthidium Urban (Hymenoptera, Apidae) with new species from Argentina and Peru. Zootaxa 3609 (3): 311-318, DOI: 10.11646/zootaxa.3609.3.5
3B0087DD8C7D043AD3BABC56EB85D096.taxon	discussion	Comments and diagnosis. Integument of mesoscutum smooth, with two anterior yellow bands; each side of vertex with one yellow spot (Fig. 12). Female: metasoma with yellow bands on T 1, T 3 - T 5 (Fig. 21). This species is very similar with C. argentinum differing by the metasoma with shallower, smaller and sparser punctures. Ctenanthidium nigripes also differs by the presence of only two yellow bands on metasoma while C. argentinum has the metasoma with four yellow bands.	en	Parizotto, Daniele R., Urban, Danúncia (2013): Contribution to the knowledge of the Anthidulum Michener and Ctenanthidium Urban (Hymenoptera, Apidae) with new species from Argentina and Peru. Zootaxa 3609 (3): 311-318, DOI: 10.11646/zootaxa.3609.3.5
3B0087DD8C7D043AD3BABC56EB85D096.taxon	description	Description male. Approximate body length 6.1 mm; forewing length 5.2 mm; head width 2.0 mm; eye length 1.3 mm. Color: Integument black except yellow as follows: mandible except teeth and margins; clypeus with subapical macula, wider at middle; supraclypeal area with U-shaped macula including juxtantennal carina; lower paraocular area, narrowing near antennal sockets. One spot on each side of vertex, behind eyes. Antenna with scape, pedicel and first two basal segments amber; pedicel blackened dorsally (Fig. 11). Mesoscutum with an anterior band on each side, wider near middle; axilla and scutellum with subapical band and translucent margin. Tegula amber, wing membrane blackened, veins black, darker at base and at costal margin (Fig. 12). Legs with coxae, trochanters and basal half of femora blackened; fore leg with femur amber on external surface and apex; middle and hind legs with apices of femora, tibiae and tarsomeres amber. Terga black; T 1 with large yellow band, slightly narrow at middle; T 2 with yellow lateroventral macula; T 3 - T 5 with discal yellow band, distal half of terga yellowish amber with translucent margin, larger on T 3; T 6 - T 7 with marginal yellow band (Fig. 23). S 1 with lateral spots; S 2 with apical half yellow and stained by two discal black spot; S 3 with two large lateral yellow spots and translucent medial margin. Pilosity. Predominantly white, hairs longer than ocellar diameter, sparser and longer on paraocular area, clypeal apex, scutellar apex, sides of terga, sterna and legs. Second and third sterna with apical fringe of plumose hairs laterally. Punctures. Head and mesosoma densely punctate; punctures on metasoma shallower and sparser than in mesosoma. Punctures of first and second terga sparser and smaller than in the remaining terga. Material examined. One male: “ BOLIVIA, Cochabamba: / Peña Colorada, 1800 m. / February 21,1976 / L. E. Peña ” (AMNH); ARGENTINA: one male, Jujuy, Tumbaya, II / 84, Fritz; one male and one female from Salta, El Maray, I / 94, Fritz (AMNH); one female from Salta, Cuesta, Obispo, II / 85 (AMNH); one male and one female from Salta, Chorrillos, III / 93 (AMNH); one male from El Alisal, III / 93 (DZUP) and one female from El Alisal III / 94, Fritz, (DZUP). Variation. In the other males examined following variations in the supraclypeal area were noted: the specimen from El Alisal has a U-shaped macula, including the juxtantennal carina; the male from El Maray has a yellow spot near the epistomal sulcus; the male from Tumbaya has a small yellow band near the epistomal sulcus. Females differ from holotype by having larger ferruginous areas.	en	Parizotto, Daniele R., Urban, Danúncia (2013): Contribution to the knowledge of the Anthidulum Michener and Ctenanthidium Urban (Hymenoptera, Apidae) with new species from Argentina and Peru. Zootaxa 3609 (3): 311-318, DOI: 10.11646/zootaxa.3609.3.5
