taxonID	type	description	language	source
3B2987ECF26AFFF2FE6D7A3FDF9AFDB9.taxon	description	Male. Length. Body: 3.19 – 3.57 mm, wing: 3.52 – 3.70 mm. Head (Figure 2 (a, b )). Frons, fronto-orbital plate, parafacial and gena black with brownish-grey pollinosity. Frons longer than wide, with frontal triangle short, not reaching lunula, visible with shifting reflections according to the direction of light (birefringence). Gena with strong black hairs, and with height less than the width of the postpedicel. Arista with its longest microtrichia hardly longer than its basal diameter. Thorax (Figure 2 (c )). Black with grey pollinosity, with two fine dark-brown vitta along dorsocentral row of setae. Chaetotaxy: acrostichal setae short and strong, the anterior presutural pair as long as the eaDC; dorsocentrals 1 + 3, with an eaDC seta, less than one third of the aDC; intra-alars 1 + 2, postsutural setae similar in length; supra-alars 1 + 2, the posterior seta very short. Katepisternum with 2 – 3 setula. Wing. Tegula yellow. Legs. Coxae black with grey pollinosity and black setae. Trochanters reddish brown with black setae. Femora black with grey pollinosity, apex of all femora and all tibia yellow, tarsus black. Mid femur with a row of fine ventral setae, 2 strong setae on anterior surface, and 2 preapical setae on posterodorsal to posterior surface; mid tibia with one anterodorsal and one posterodorsal median setae, anterodorsal seta longer and positioned below of the posterodorsal seta. Hind femur with a complete row of anterodorsal and a row of anteroventral setae on apical half. Abdomen (Figure 2 (d )). Grey with square lateral dark-brown marks on tergites 3 – 5. Sternite 5 with similar width and lenght, basal margin convex, apical margin strongly concave (‘ V’ shape), without a membrane; setae concentrated on the lobes not extended to basal margin (Figure 2 (e )). Terminalia. Cercus longer than broad with a slight constriction on apical third, sclerotised, setulose (hair-like seta), with apical margin concave with 2 strongly sclerotised teeth (Figure 2 (f-h )). Surstyli longer than broad, slightly curved at the distal tip towards cercus (Figure 2 (g, i )). Hypandrium tubular, longer than wide, with lateral prolongations, distal extreme not exposing the phallapodeme. Aedeagus with phallapodeme straight, strongly sclerotised, and a little longer than hypandrium in lateral view; pregonite developed, kidney-shaped, ventrally fused with the hypandrium; postgonite developed; epiphalus slightly sclerotised, and distiphallus tubular, and slightly sclerotised at base (Figure 2 (j-k )). Female unknown.	en	Patitucci, Luciano Damián, Couri, Marcia Souto, Mulieri, Pablo Ricardo (2021): The killer flies Coenosia Meigen (Diptera: Muscidae) of southern South America: the Coenosia chaetosa species group with two new species, and notes on male terminalia patterns. Journal of Natural History 55 (23 - 24): 1499-1519, DOI: 10.1080/00222933.2021.1954711, URL: http://dx.doi.org/10.1080/00222933.2021.1954711
3B2987ECF26AFFF2FE6D7A3FDF9AFDB9.taxon	materials_examined	Type material. Holotype male, pinned, damaged. http: // data. nhm. ac. uk / object / 8 e 084 b 8 e- 63 cc- 49 af- 8937 - d 282366 f 88 b 1 Other specimens examined. ARGENTINA: Chubut: PNLP: 20 males, Intendencia, − 42.085077, − 71.614662, I- 2011, Mulieri & Patitucci leg. (MACN). Neuquén: 1 male, Lago Moquehue, 10 - II- 2018, − 38.962947, − 71.347913, Mulieri leg. (MACN).	en	Patitucci, Luciano Damián, Couri, Marcia Souto, Mulieri, Pablo Ricardo (2021): The killer flies Coenosia Meigen (Diptera: Muscidae) of southern South America: the Coenosia chaetosa species group with two new species, and notes on male terminalia patterns. Journal of Natural History 55 (23 - 24): 1499-1519, DOI: 10.1080/00222933.2021.1954711, URL: http://dx.doi.org/10.1080/00222933.2021.1954711
3B2987ECF26AFFF2FE6D7A3FDF9AFDB9.taxon	distribution	Distribution (Figure 3 (a )). ARGENTINA: Chubut (new record), Neuquén (new record). CHILE: Región de Valparaíso.	en	Patitucci, Luciano Damián, Couri, Marcia Souto, Mulieri, Pablo Ricardo (2021): The killer flies Coenosia Meigen (Diptera: Muscidae) of southern South America: the Coenosia chaetosa species group with two new species, and notes on male terminalia patterns. Journal of Natural History 55 (23 - 24): 1499-1519, DOI: 10.1080/00222933.2021.1954711, URL: http://dx.doi.org/10.1080/00222933.2021.1954711
3B2987ECF26AFFF2FE6D7A3FDF9AFDB9.taxon	discussion	Remarks. Malloch (1934) described C. chaetosa with a single male specimen from Los Andes, Chile. For this species, this author highlights the presence and development of the presutural acrostichal and dorsocentral setae. The number of the presutural dorsocentral setae in Coenosiinae (one or two setae) and the interpretation of the second one (as seta or setula) have been analysed in different ways. All Coenosiinae have a developed aDC (generally with the same length as the postsutural dorsocentral setae), which is placed in the anterior half of the prescutum. The ground-plan state in Muscidae includes two presutural dorsocentral setae, being the absence or reduction of the second presutural dorsocentral seta (pDC), placed in the posterior half of the prescutum, a character observed in Coenosiinae (Couri and Pont 2000). Some Coenosiinae species present a second or extra anterior presutural dorsocentral seta (eaDC), located anterior to the developed aDC. Both setae are in the anterior middle half of the prescutum. In previous works focused on Coenosia, the presence of eaDC was studied in different ways. Malloch (1934) compared for C. chaetosa the size of the eaDC in relation to the prestural acrostical setae, and observed that eaDC setae are very close to the aDC. Other authors mentioned the eaDC as an ‘ accessory presut dc setulae’ (Snyder 1957), or just compared with the size of the aDC (Gregor et al. 2002). Sorokina (2009) presented an interesting discussion on the presence and size of this character for the genera Macrochis Rondani and Dexiopsis Pokorny, currently synonymous with Coenosia. Important to mention that other Coenosiinae genera, as the endemic Apsil Malloch and Reynoldsia Malloch of the Subantarctic sub-region, present this eaDC (Malloch 1934). A detailed study with a phylogenetic analysis could give more information on this character.	en	Patitucci, Luciano Damián, Couri, Marcia Souto, Mulieri, Pablo Ricardo (2021): The killer flies Coenosia Meigen (Diptera: Muscidae) of southern South America: the Coenosia chaetosa species group with two new species, and notes on male terminalia patterns. Journal of Natural History 55 (23 - 24): 1499-1519, DOI: 10.1080/00222933.2021.1954711, URL: http://dx.doi.org/10.1080/00222933.2021.1954711
3B2987ECF265FFF0FE427A69DDADFADE.taxon	description	Male. Length. Body: 3.18 – 3.37 mm, wing: 3.62 – 3.80 mm. Head (Figure 4 (a, b )). Frons black with golden pollinosity, fronto-orbital plate, parafacial and gena black with grey pollinosity. Frons longer than wide, with frontal triangle long, reaching lunula. Gena with thin black hairs, and with height greater than the width of the postpedicel. Arista with its longest hairs hardly longer than its basal diameter. Thorax (Figure 4 (c )). Black with grey pollinosity, with two fine dark-brown vittae along dorsocentral row of setae. Chaetotaxy: acrostichal setae short and strong, the anterior presutural pair as long as the eaDC; dorsocentrals 1 + 3, with a short eaDC pair which is less than one fifth of the aDC; intra-alars 1 + 2, postsutural setae similar in length; supraalars 1 + 2, the posterior seta short. Katepisternum with 1 – 3 setula. Wing. Tegula yellow. Legs. Coxae black with grey pollinosity and black hairs. Trochanters yellow-reddish with black hairs. Femora black with grey pollinosity, tibia yellow, and tarsus dark brown. Mid femur with a row of fine ventral setae, a row of fine setae on anterior surface, and 2 preapical setae on posterodorsal to posterior surface; mid tibia with one anterodorsal and one posterodorsal median setae, anterodorsal seta shorter and positioned below of the posterodorsal seta. Hind femur with a complete row of anterodorsal and a row of anteroventral black hairs. Abdomen (Figure 4 (d )). Grey with sub-triangular lateral dark-brown marks on tergites 3 – 5. Sternite 5 with similar width and lenght, basal margin convex, apical margin strongly concave (‘ V’ shape), without a membrane; setae concentrated on the lobes and a little spread over the basal margin (Figure 4 (e )). Terminalia. Cercus longer than broad with a slightly constriction on apical third, sclerotised, setulose (hair-like seta), with apical margin concave with 2 teeth strongly sclerotised (Figure 4 (f-h )). Surstyli longer than broad, slightly curved towards at the distal tip towards cercus (Figure 4 (g )). Hypandrium tubular, longer than wide, without lateral prolongation, distal extreme not exposing the phallapodeme. Aedeagus with phallapodeme straight, strongly sclerotised, and a little longer than hypandrium in lateral view; pregonite developed, kidney-shaped, ventrally fused with the hypandrium; postgonite developed; epiphalus sclerotised, and distiphallus tubular, and sclerotised at base (Figure 4 (i-j )). Female unknown.	en	Patitucci, Luciano Damián, Couri, Marcia Souto, Mulieri, Pablo Ricardo (2021): The killer flies Coenosia Meigen (Diptera: Muscidae) of southern South America: the Coenosia chaetosa species group with two new species, and notes on male terminalia patterns. Journal of Natural History 55 (23 - 24): 1499-1519, DOI: 10.1080/00222933.2021.1954711, URL: http://dx.doi.org/10.1080/00222933.2021.1954711
3B2987ECF265FFF0FE427A69DDADFADE.taxon	materials_examined	Type material examined. Holotype male, pinned, labels: ‘ Arg. Chubut, P. N. Los Alerces, / Cabaña La Cascasa 532 m (Malaise) / 42 ° 53 ʹ 19,17 ’ S, 71 ° 35 ʹ 34,37 ” W / 5 - II- 2013 Olea, Mulieri & Patitucci leg. ” print. on white paper; ‘ MB 2 ’ print on white paper, black frame, ‘ MACN-En / 34714 ’ print. on white paper. Paratypes: male, pinned, labels: Arg. Chubut, P. N. Los Alerces, / Cabaña La Cascasa 532 m (Malaise) / 42 ° 53 ʹ 19,17 ” S, 71 ° 35 ʹ 34,37 ” W / 5 - II- 2013 Olea, Mulieri & Patitucci leg. ” print. on white paper; ‘ MB 3 ’ print on white paper, black frame, ‘ MACN-En / 34715 ’ print. on white paper; male, pinned, labels: Arg. Chubut, P. N. Los Alerces, / Cabaña La Cascasa 532 m (Malaise) / 42 ° 53 ʹ 19,17 ” S, 71 ° 35 ʹ 34,37 ” W / 5 - II- 2013 Olea, Mulieri & Patitucci leg. ” print. on white paper; ‘ MA 4 ’ print on white paper, black frame, ‘ MACN-En / 347146 ’ print. on white paper. Other specimens examined. ARGENTINA: Chubut: PNLP: 2 males, Río Azul, I- 2012, − 42.0916, − 71.6155, Mulieri & Patitucci leg. (MACN). Neuquén: PNL: 2 males, Quillen, 11 - I- 2013, − 39.3613, − 71.2188, Patitucci leg. (MACN).	en	Patitucci, Luciano Damián, Couri, Marcia Souto, Mulieri, Pablo Ricardo (2021): The killer flies Coenosia Meigen (Diptera: Muscidae) of southern South America: the Coenosia chaetosa species group with two new species, and notes on male terminalia patterns. Journal of Natural History 55 (23 - 24): 1499-1519, DOI: 10.1080/00222933.2021.1954711, URL: http://dx.doi.org/10.1080/00222933.2021.1954711
3B2987ECF265FFF0FE427A69DDADFADE.taxon	distribution	Distribution (Figure 3 (b )). ARGENTINA: Chubut, Neuquén.	en	Patitucci, Luciano Damián, Couri, Marcia Souto, Mulieri, Pablo Ricardo (2021): The killer flies Coenosia Meigen (Diptera: Muscidae) of southern South America: the Coenosia chaetosa species group with two new species, and notes on male terminalia patterns. Journal of Natural History 55 (23 - 24): 1499-1519, DOI: 10.1080/00222933.2021.1954711, URL: http://dx.doi.org/10.1080/00222933.2021.1954711
3B2987ECF265FFF0FE427A69DDADFADE.taxon	etymology	Etymology. The name of this species is dedicated to John R. Malloch for his great contribution to the knowledge of Patagonian Muscidae.	en	Patitucci, Luciano Damián, Couri, Marcia Souto, Mulieri, Pablo Ricardo (2021): The killer flies Coenosia Meigen (Diptera: Muscidae) of southern South America: the Coenosia chaetosa species group with two new species, and notes on male terminalia patterns. Journal of Natural History 55 (23 - 24): 1499-1519, DOI: 10.1080/00222933.2021.1954711, URL: http://dx.doi.org/10.1080/00222933.2021.1954711
3B2987ECF265FFF0FE427A69DDADFADE.taxon	discussion	Remarks. Coenosia mallochi sp. nov. is distinguished from its congeners by the following combination of characters: frontal triangle long, reaching lunula, visible; a short eaDC which is less than one fifth of the aDC; trochanters yellow-reddish with black hairs; mid tibia with one anterodorsal seta and one posterodorsal median seta, anterodorsal seta shorter and positioned below the posterodorsal seta; hind tibia with one long anterodorsal seta and one thin anteroventral seta; cercus longer than broad, setulose (hair-like seta); distiphallus tubular, and sclerotised at base. In Stein’s key (1911), C. mallochi runs to C. iniqua, but the presence of vittae on the scutum and spots on the abdomen can separate it from C. iniqua. In Malloch´s key (1934), C. mallochi runs to C. inaequalis, but the presence of black strong setae on coxae can separate it from the male of C. inaequalis.	en	Patitucci, Luciano Damián, Couri, Marcia Souto, Mulieri, Pablo Ricardo (2021): The killer flies Coenosia Meigen (Diptera: Muscidae) of southern South America: the Coenosia chaetosa species group with two new species, and notes on male terminalia patterns. Journal of Natural History 55 (23 - 24): 1499-1519, DOI: 10.1080/00222933.2021.1954711, URL: http://dx.doi.org/10.1080/00222933.2021.1954711
3B2987ECF267FFF4FE457CDBDDDFFE06.taxon	description	Male. Length. Body: 2.71 – 3.22 mm, wing: 3.03 – 3.19 mm. Head (Figure 5 (a-b )). Frons black, fronto-orbital plate, parafacial and gena black with silver-grey pollinosity. Frons longer than wide, without frontal triangle. Gena with strong black hairs, and with height greater than the width of the postpedicel. Arista with its longest microtrichia shorter than its basal diameter. Thorax (Figure 5 (c )). Black without vittae; postpronotal lobe and lateral plates black with white-grey pollinosity. Chaetotaxy: acrostichal setae long and strong, the anterior presutural pair as long as the eaDC; dorsocentrals 1 + 3, with an aDC seta that is less than one third of the eaDC; intra-alars 1 + 2, posterior postsutural seta shorter; supra-alars 1 + 1. Katepisternum without setula. Wing. Tegula black. Legs. Coxae black with grey pollinosity and black setae. Trochanters yellow with black setae. Femora and tibiae metallic black, apex of all tibiae yellow, tarsi dark brown. Mid femur with 3 – 4 ventral setae, and 2 preapical setae on posterodorsal to posterior surface; mid tibia with one anterodorsal and one posterodorsal median setae, anterodorsal seta shorter and positioned below of the posterodorsal seta. Hind femur with a complete row of anterodorsal, a row of strong anteroventral setae on basal half, and a row of strong posteroventral setae on basal half, with two preapical setae (anterodorsal and posterodorsal); some specimens a short dorsal preapical seta. Abdomen (Figure 5 (d )). Black with brown pollinosity, without spots or central line, tergite 6 and epandrium metallic black. Epandrium globular. Sternite 5 similar in width and lenght, basal margin convex, apical margin strongly concave (‘ U’ shape), with a depressed area at the base; lobes with a membrane on apical margin; setae concentrated on the lobes and extended to basal margin (Figure 5 (e )). Terminalia. Cercus metallic black, longer than broad without a constriction on apical third, sclerotised, setulose only on basal third (hair-like seta), with apical margin concave with two strong and wide sclerotised teeth (Figure 5 (f-h )). Surstyli longer than broad, slightly curved towards at the distal tip towards cercus (Figure 5 (g, i )). Hypandrium tubular, longer than wide. Aedeagus with phallapodeme straight, strongly sclerotised, and longer than hypandrium in lateral view; pregonite developed, kidney-shaped, ventrally fused with the hypandrium; postgonite developed; epiphalus slightly sclerotised, and distiphallus tubular with hairs on ventral surface, and slightly sclerotised at base (Figure 5 (j-k )). Female (Figure 6 (a )). Length. Body: 3.75 – 4.0 mm, wing: 4.03 – 4.20 mm. Differs from male as follows: Legs. Femora and tibiae metallic black, the metallic colour in some specimens is not so evident. Mid femur with an anteroventral row of setae, 3 – 4 strong setae on anterior surface, and with 3 – 4 ventral setae on basal third. mid tibia with one anterodorsal and one posterodorsal median setae, anterodorsal seta longer and positioned below of the posterodorsal seta. Ovipositor: Segments longer than wide. Tergites 6, 7, and 8 with 2 long and parallel scletotized plates, plates on tergite 6 wider than in the others (Figure 6 (b )); epiproct triangular, with hair-like setae, cercus longer than epiproct (Figure 6 (d )). Sternites 6 and 7 as a long and rectangular plate; sternite 8 divided into 2 small and sclerotised plates each with 4 – 5 setae on distal margin (Figure 6 (c )); hypoproct triangular, setulose, with several strong setae on distal margin (Figure 6 (e )). Three spermathecae, ovoid shape.	en	Patitucci, Luciano Damián, Couri, Marcia Souto, Mulieri, Pablo Ricardo (2021): The killer flies Coenosia Meigen (Diptera: Muscidae) of southern South America: the Coenosia chaetosa species group with two new species, and notes on male terminalia patterns. Journal of Natural History 55 (23 - 24): 1499-1519, DOI: 10.1080/00222933.2021.1954711, URL: http://dx.doi.org/10.1080/00222933.2021.1954711
3B2987ECF267FFF4FE457CDBDDDFFE06.taxon	materials_examined	Type material examined. Holotype male, pinned, labels: ‘ Arg. Neuquén, P. N. Lanín / (Malaise open Bandurrias) 800 m / 40 ° 08,728 ¨ ’ S, 71 ° 30,826 ʹ W / 21 - II- 2011 Mulieri & Patitucci leg. ’ print. on white paper; ‘ MACN-En / 34717 ’ print. on white paper. Paratypes: male, pinned, labels: ‘ Arg. Neuquén, P. N. Lanín / Quillen 993 m (Malaise 2) / 39 ° 31 ʹ 40 ’ S, 71 ° 13 ʹ 13 ” W / I- 2013 Olea, Mulieri & Patitucci leg. ” print. on white paper; ‘ MACN-En / 34718 ’ print. on white paper; 2 females, pinned, labels: ‘ Arg. Neuquén, P. N. Lanín / Quillen 993 m (Malaise 2) / 39 ° 31 ʹ 40 ’ S, 71 ° 13 ʹ 13 ” W / I- 2013 Olea, Mulieri & Patitucci leg. ” print. on white paper; ‘ MACN-En / 34719 - 20 ’ print. on white paper. Two males, pinned, labels: ‘ Arg. Neuquén, Lagunas de / Epulaufquen / 1502 m. a. s. l. transecta / - 36.815642, − 71,064096 / 12 - II - 2018 Patitucci leg’. (IFML). Two females and one male, pinned, labels: ‘ Arg. Neuquén, P. N. Lanín / Seccional Bandurrias II- 2011 / - 40.144240, − 71.342986 / Mulieri & Patitucci leg’. print. on white paper (MLP). Other specimens examined. ARGENTINA: Chubut: PNLA: 2 females, 3 males, Cab. La Cascada, II- 2013, − 42.888499, − 71.592376, Mulieri, Patitucci & Olea leg. (MACN); 1 female, Lag. Futalaufquen, II- 2013, − 42.840982, − 71.632934, Patitucci & Olea leg. (MACN). Neuquén: ABM: 1 male, I- 2013, − 38.845457, − 71.093002, Patitucci leg. (MACN), Cerrro Chapelco, 103 females, 15 males, 1983, − 40.197050, − 71.298453, Gentili leg. (IFML); ALE: 26 females, 3 males, 12 - II- 2018, − 36.815.642, − 71.084.096, Mulieri & Patitucci leg. (MACN); PNL: 3 females, 2 males, Quillen, 11 - I- 2013, − 39.3613, − 71.2188, Mulieri, Patitucci & Olea leg. (MACN); Villa Pehuenia: 5 females, 10 / 11 - II- 2018, − 38.884503, − 71.166194, Mulieri & Patitucci leg. (MACN). Río Negro: PNNH: 3 females, Mallín, Playa Negra, II- 2009, - 41357809, − 71571683, Garré & Montes de Ocaleg. (MLP).	en	Patitucci, Luciano Damián, Couri, Marcia Souto, Mulieri, Pablo Ricardo (2021): The killer flies Coenosia Meigen (Diptera: Muscidae) of southern South America: the Coenosia chaetosa species group with two new species, and notes on male terminalia patterns. Journal of Natural History 55 (23 - 24): 1499-1519, DOI: 10.1080/00222933.2021.1954711, URL: http://dx.doi.org/10.1080/00222933.2021.1954711
3B2987ECF267FFF4FE457CDBDDDFFE06.taxon	distribution	Distribution (Figure 3 (c )). ARGENTINA: Chubut, Neuquén, Río Negro.	en	Patitucci, Luciano Damián, Couri, Marcia Souto, Mulieri, Pablo Ricardo (2021): The killer flies Coenosia Meigen (Diptera: Muscidae) of southern South America: the Coenosia chaetosa species group with two new species, and notes on male terminalia patterns. Journal of Natural History 55 (23 - 24): 1499-1519, DOI: 10.1080/00222933.2021.1954711, URL: http://dx.doi.org/10.1080/00222933.2021.1954711
3B2987ECF267FFF4FE457CDBDDDFFE06.taxon	etymology	Etymology. The name refers to the striking black metallic colour of the legs.	en	Patitucci, Luciano Damián, Couri, Marcia Souto, Mulieri, Pablo Ricardo (2021): The killer flies Coenosia Meigen (Diptera: Muscidae) of southern South America: the Coenosia chaetosa species group with two new species, and notes on male terminalia patterns. Journal of Natural History 55 (23 - 24): 1499-1519, DOI: 10.1080/00222933.2021.1954711, URL: http://dx.doi.org/10.1080/00222933.2021.1954711
3B2987ECF267FFF4FE457CDBDDDFFE06.taxon	discussion	Remarks. Coenosia metalleg sp. nov. is distinguished from its congeners by the following combination of characters: thorax and abdomen black without vittae or spots, coxae black, trochanters yellow, and femora and tibiae metallic black, mid tibia with one anterodorsal and one posterodorsal median setae, anterodorsal seta shorter and positioned below the posterodorsal seta, cercus metallic black, longer than broad with hair-like seta, and distiphallus with hairs on ventral surface In Stein’s key (Stein 1911), C. metalleg runs to C. setiventris. These two species are distinguished by differences in the male terminalia of C. setiventris (Couri and Pont 2020), and their general colouration, particularly the colour of the legs. In Malloch´key (Malloch 1934), C. metalleg runs to C. inaequalis, but the metallic black colour of the femora can separate it from C. inaequalis.	en	Patitucci, Luciano Damián, Couri, Marcia Souto, Mulieri, Pablo Ricardo (2021): The killer flies Coenosia Meigen (Diptera: Muscidae) of southern South America: the Coenosia chaetosa species group with two new species, and notes on male terminalia patterns. Journal of Natural History 55 (23 - 24): 1499-1519, DOI: 10.1080/00222933.2021.1954711, URL: http://dx.doi.org/10.1080/00222933.2021.1954711
