identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
3B3687C07C769B22FF67F7DE9E5DFEE0.text	3B3687C07C769B22FF67F7DE9E5DFEE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus Ashmead 1893	<div><p>Genus Dissomphalus Ashmead, 1893</p><p>Dissomphalus Ashmead, 1893: 41 –42. Type-species: Dissomphalus xanthopus Ashmead, by original designation.</p><p>Ecitopria Wasmann, 1899: 55–56, 127. Type-species: Ecitopria crassicornis Wasmann, by monotypy. Synonymized by Evans, 1955: 290.</p><p>Psilobethylus Kieffer in Kieffer &amp; Marshall, 1906: 461 –462. Type-species: Psilobethylus luteus Kieffer, by monotypy. Synonymized by Terayama, 1995: 888.</p><p>Thaumatepyris Kieffer, 1910a: 47. Type-species: Thaumatepyris punctatus Kieffer, by monotypy. Synonymized by Evans, 1964: 41.</p><p>Glenobethylus Kieffer, 1910a: 50. Type-species: Glenobethylus montanus Kieffer, by monotypy. Synonymized by Evans, 1964: 41.</p><p>Parisobrachium Kieffer, 1914b: 237, 424. Type-species: Rhabdepyris ? albipes Kieffer, by monotypy. Synonymized by Azevedo, 2008: 784.</p><p>Parecitopria Oglobin, 1930: 15 . Type-species: Parecitopria azarai Oglobin, by monotypy. Synonimized by Evans, 1964: 41.</p><p>Diagnosis. Male. Body robust. Clypeus prominent, median lobe sometimes ill-defined. Metasoma relatively wide and short; second tergo sometimes with tergal process (depressions, pits, tufts of setae, tubercles). Hypopygium with three stalks. Genitalia wide; paramere simple or bilobate; aedeagus complex, composed by ventral ramus and dorsal body. Female. Head longer than wide. Eye small, with from one to about 25 facets. Mesonotum short, transverse, posterior margin gently convex at broad junction with propodeum; anterior margin of propodeum not embracing sides of mesonotum to any appreciable extent. Genitalia with aedeagus divided into dorsal body and ventral ramus.</p></div>	https://treatment.plazi.org/id/3B3687C07C769B22FF67F7DE9E5DFEE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C759B2DFF67F3AB9AF9FAA3.text	3B3687C07C759B2DFF67F3AB9AF9FAA3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus Ashmead 1893	<div><p>Key to Dissomphalus from Central America</p><p>1. Tergal process absent................................................................................... 2</p><p>- Tergal process present.................................................................................. 5</p><p>2. Mandible with four distal teeth; median clypeal lobe ill-defined................................................ 4</p><p>- Mandible with two or three distal teeth; median clypeal lobe trapezoidal.......................................... 3</p><p>3. Mandible with three distal teeth; anterior margin of pronotal disc ecarinate; posterior hypopygeal margin sinuous (Fig. 66); aedeagal ventral ramus shorter than aedeagal dorsal body, inner margin straight, apex crenulate slightly curved outward (Fig. 104)................................................................................. D. asfartus sp. nov.</p><p>- Mandible with two distal teeth; anterior margin of pronotal disc coarse; posterior hypopygeal margin weakly concave (Fig. 80); aedeagal ventral ramus higher than aedeagal dorsal body, inner margin excavated, apex simple and parallel (Fig. 170)........................................................................................ D. raziogus sp. nov.</p><p>4. Posterior hypopygeal margin sinuous (Fig. 65); paramere with dorsal margin straight; aedeagal dorsal body with apical margin acute (Fig. 101); aedeagal ventral ramus abruptly narrowing apicad (Fig.102)....................... D. azagus sp. nov.</p><p>- Posterior hypopygeal margin straight or nearly so (Fig. 67); paramere with dorsal margin sinuous; aedeagal dorsal body with apical margin rounded (Fig. 105); aedeagal ventral ramus progressively narrowing apicad (Fig. 106)...... D. iozus sp. nov.</p><p>5. Tergal process median................................................................................. 6</p><p>- Tergal process lateral or sublateral....................................................................... 22</p><p>6. Tergal process with circular depression, with densely tuft of setae, always without tubercle; aedeagal ventral ramus wide... 7</p><p>- Tergal process and aedeagal ventral ramus not as above...................................................... 10</p><p>7. Tergal process with depression very small and two small tufts of setae............................ D. altivolans Evans</p><p>- Tergal process with very large depression and large tufts of setae............................................... 8</p><p>8. Tergal process with pair of tuft of long setae posterad, fused; paramere as long as basiparamere......... D. unitus Azevedo</p><p>- Tergal process with pair of tuft of short setae, variable direction and distinctly separated; paramere much long than basi- paramere............................................................................................ 9</p><p>9. Median clypeal lobe ill-defined (Fig. 1) with high keel, median carina very high in profile; gena with spine (Fig. 2); dorsal margin of paramere sinuous (Fig. 99); aedeagal ventral ramus bifid (Fig. 100)........................ D. isteus sp. nov.</p><p>- Median clypeal lobe tridentate, without such keel, median carina not as above; gena without spine; dorsal margin of paramere straight; aedeagal ventral ramus simple (Figs 97, 98)...................................... D. subdeformis Azevedo</p><p>10. Tergal process with median depression, transverse, elliptical with pair of tubercles directed each other................. 11</p><p>- Tergal process not as above............................................................................. 12</p><p>11. Aedeagal dorsal ramus shorter than aedeagal dorsal body with subapical dilation (Fig. 196)........... D. dilatatus Azevedo</p><p>- Aedeagal dorsal ramus nearly as long as aedeagal dorsal body, margin smooth (Fig. 199)............... D. vallensis Evans</p><p>12. Depression of tergal process extremely shallow, much wider than long, occupying nearly two median thirds of tergite width..................................................................................................... 13</p><p>- Depression of tergal process not as above.................................................................. 14</p><p>13. Depression of tergal process very shallow, nearly imperceptible; aedeagal dorsal body bidentate, base of ventral margin with small tooth; aedeagal ventral ramus long with projection narrow and flat apical upward............ D. rasissimus Azevedo</p><p>- Depression of tergal process not so shallow; aedeagal dorsal body simple and apex of ventral margin serrated (Figs 171, 173); aedeagal ventral ramus short, wide with two conspicuous and laminar rami, ventral and apical margins serrated (Fig. 172)...................................................................................... D. bilobatus Azevedo</p><p>14. Tergal process with depression deep, wide and subtriangular, with base directed forward, lateral pit very large much arched inward medially, with many long setae mesad........................................... D. curvifoveatus Azevedo</p><p>- Tergal process with depression not as above............................................................... 15</p><p>15. Aedeagal ventral ramus with apical portion as wide as median portion, basal portion of outer margin straight.................................................................................................... D. apertus Kieffer</p><p>- Aedeagal ventral ramus with apex much narrower than median, basal portion of outer margin excavated or projected..... 16</p><p>16. Aedeagal ventral ramus with basal portion of outer margin projected, apex parallel or nearly so (Fig. 220).. D. trisus sp. nov.</p><p>- Aedeagal ventral ramus with basal portion of outer margin excavated, apex much arched outward..................... 17</p><p>17. Pronotal disc with anterior margin coarse.................................................................. 18</p><p>- Pronotal disc with anterior margin carinate................................................................ 18</p><p>18. Tergal process with shallow depression and small tubercle (Fig. 37); apex of apical lobes of aedeagal dorsal body acute (125); apex of aedeagal ventral ramus toothed (Fig. 126).............................................. D. asitius sp. nov.</p><p>- Tergal process with deep depression, without tubercle (Fig. 39); apex of apical lobes of dorsal body rounded (Fig. 129); apex of aedeagal ventral ramus smooth (Fig. 130).................................................. D. tisinus sp. nov.</p><p>19. Tergal process with shallow and subcircular depression, with very small tubercle or without tubercle; apical portion of aedea- gal ventral ramus divergent and long..................................................................... 20</p><p>- Tergal process with deep and large depression; apical portion divergent of aedeagal ventral ramus very short............ 21</p><p>20. Tergal process with tubercle (Fig. 38); posterior hypopygeal margin weakly concave (Fig. 69); outer lobe of aedeagal dorsal body long (Fig. 127); basal inner margin of aedeagal ventral ramus straight (Fig. 128)................. D. ibocius sp. nov.</p><p>- Tergal process without tubercle (Fig. 55); posterior hypopygeal margin straight (Fig. 86); outer lobe of aedeagal dorsal body very long (Fig. 188); basal inner margin of aedeagal ventral ramus sinuous (Fig. 189)................. D. terbius sp. nov.</p><p>21. Mandible with three distal teeth; posterior hypopygeal margin straight; apex of paramere with four projections............................................................................................... D. angulatus Azevedo</p><p>- Mandible with two distal teeth; posterior hypopygeal margin convex; apex of paramere with one projection;.................................................................................................. D. bifoveatus Kieffer</p><p>22. Tergal process sublateral, usually with dense tuft of setae posterad, tubercle absent................................. 23</p><p>- Tergal process lateral, with tuft of setae not as above, tubercle usually present..................................... 31</p><p>23. Mandible with three distal teeth; apical lobes of aedeagal dorsal body with crown-like structure, aedeagal ventral ramus thick................................................................................................... 24</p><p>- Mandible with two teeth; apical lobes of aedeagal dorsal body not as above, aedeagal ventral ramus slender.................................................................................................. D. punctatus (Kieffer)</p><p>24. Pronotal disc with anterior margin carinate............................................................... 25</p><p>- Pronotal disc ecarinate................................................................................ 30</p><p>25. Posterior hypopygeal margin straight..................................................................... 26</p><p>- Posterior hypopygeal margin convex..................................................................... 29</p><p>26. Aedeagal dorsal body with crown-process long or very long................................................... 27</p><p>- Aedeagal dorsal body with crown-process small............................................................ 28</p><p>27. Median clypeal lobe trapezoidal; crown-process with apex dorsad; aedeagal ventral ramus entirely wide with acute apex................................................................................. D. sinatus Alencar &amp; Azevedo</p><p>- Median clypeal lobe subtrapezoidal (Fig. 16); crown-process with apex laterad (Fig. 157); aedeagal ventral ramus with median half wide narrowing apicad (Fig. 158)....................................................... D. afestus sp.nov.</p><p>28. Median clypeal carina strongly developed in profile; aedeagal ventral ramus with apical portion with strongly concavity, apex rounded and wide............................................................ D. galeatus Alencar &amp; Azevedo</p><p>- Median clypeal carina not developed in profile; aedeagal ventral ramus with apical portion without concavity, apex acute and narrow................................................................... D. declinatus Alencar &amp; Azevedo</p><p>29. Apical lobes of aedeagal dorsal body long; base of paramere developed; cuspis with apex acute; aedeagal ventral ramus short than aedeagal dorsal body...................................................... D. ramosus Alencar &amp; Azevedo</p><p>- Apical lobes of aedeagal dorsal body short; base of paramere not as above; cuspis with rounded apex; aedeagal ventral ramus much shorter than dorsal aedeagal dorsal body.................................... D. corontaus Alencar &amp; Azevedo</p><p>30. Paramere longer than basiparamere; apical lobes of aedeagal dorsal body very long; inner margin of aedeagal ventral ramus serrated with apex straight dorsad............................................... D. culteratus Alencar &amp; Azevedo</p><p>- Paramere as long as basiparamere; apical lobes of aedeagal dorsal body regular sized; aedeagal ventral ramus smooth with apex folded laterad............................................................ D. pronus Alencar &amp; Azevedo</p><p>31. Tergal processes with pair conspicuous tubercles with high rim or nearly so and tuft of dense setae.................... 32</p><p>- Tergal processes with pair of small tubercles or even absent, never as main modification of processes.................. 39</p><p>32. Mandible with three distal teeth; median clypeal lobe ill-defined, with one angulate very large tooth (Fig. 12); basiparamere narrower than paramere (Fig. 140)......................................................... D. ibrastus sp. nov.</p><p>- Mandible with two distal teeth; median clypeal lobe trapezoidal, with tooth no so large; basiparamere as narrow as paramere or much wide.......................................................................................... 33</p><p>33. Basiparamere very wide basally (Fig. 145); aedeagal ventral ramus much higher than aedeagal dorsal body (Fig. 146)............................................................................................. D. teraus sp. nov.</p><p>- Basiparamere entirely narrow; aedeagal ventral ramus as long as or shorter than aedeagal dorsal body.................. 33</p><p>34. Paramere much long than basiparamere; apex of aedeagal dorsal body translucent................................. 35</p><p>- Paramere as long as basiparamere; apex of aedeagal dorsal body not translucent................................... 36</p><p>35. Tergal process with very shallow depression; aedeagal ventral ramus with median portion as wide as basal region, apical por- tion with inner margin serrated (Fig. 132)................................................. D. cervoides Azevedo</p><p>- Tergal process without depression (Fig. 42); aedeagal ventral ramus with median portion wider than basal region, apical por- tion with inner margin smooth (Fig. 139)....................................................... D. ettus sp. nov.</p><p>36. Aedeagal dorsal body narrowing abruptly apicad............................................................ 37</p><p>- Aedeagal dorsal body narrowing slightly apicad............................................................ 38</p><p>37. Tergal process with small depression (Fig. 46); inner margin of aedeagal ventral ramus smooth, apex rounded (Fig. 155)........................................................................................... D. urazius sp. nov.</p><p>- Tergal process with large depression (Fig. 44); inner margin of aedeagal ventral ramus serrated, apex truncate with rounded projection (Fig. 143).................................................................... D. racarus sp. nov.</p><p>38. Tergal process with shallow and small depression (Fig. 40); apex of aedeagal ventral ramus truncate and wide, inner margin strongly sinuous and smooth (Fig. 134).................................................... D. ardonius sp. nov.</p><p>- Tergal process with deep and large depression (Fig. 41); apex aedeagal ventral ramus rounded and narrow, inner margin slightly sinuous and crenulate (Fig. 137)..................................................... D. arteus sp. nov.</p><p>39. Tergal process composed of tubercle with wide fovea and thick setae on border of each depression.................... 40</p><p>- Tergal process not as above............................................................................. 45</p><p>40. Mandible with four distal teeth.......................................................................... 41</p><p>- Mandible with two distal teeth.......................................................................... 42</p><p>41. Median clypeal lobe ill-defined (Fig. 20); apical lobes of aedeagal dorsal body very short (180); aedeagal ventral ramus simple (Fig.181)............................................................................. D. aretisus sp. nov.</p><p>- Median clypeal lobe tridentate; apical lobes of aedeagal dorsal body regular sized; aedeagal ventral ramus with three long and conspicuous apical horns................................................................... D. politus Kieffer</p><p>42. Median clypeal carina very high in profile (Fig. 21); anterior margin of pronotal disc carinate; aedeagal ventral ramus with slender and long ventral projection (Fig. 183)................................................... D. citus sp. nov.</p><p>- Median clypeal carina low in profile; anterior margin of pronotal disc ecarinate or coarse; aedeagal ventral ramus without slen- der and long ventral projection.......................................................................... 43</p><p>43. Tergal process with dense tuft of setae on tubercle; posterior hypopygeal margin straight; dorsal body entirely wide or nearly so; aedeagal ventral ramus with not translucent apex......................................................... 44</p><p>- Tergal process with short and sparse tuft of setae on tubercle (Fig. 50); posterior hypopygeal margin concave (Fig. 81); aedea- gal dorsal body entirely narrow (Fig. 178); aedeagal ventral ramus with translucent apex (Fig. 179)........ D. areius sp. nov.</p><p>44. Tergal process with high tubercle and small pit (Fig. 53); aedeagal dorsal body with slightly constriction near apical lobe (Fig. 184); apex of aedeagal ventral ramus toothed (Fig.185).......................................... D. sartus sp. nov.</p><p>- Tergal process with low tubercle and large pit (Fig. 54); aedeagal dorsal body with abruptly constriction near apical lobe (Fig. 186); apex of aedeagal ventral ramus smooth (Fig. 187)......................................... D. teobus sp. nov.</p><p>45. Metasoma with pair of lateral and anterior depressions large, usually more than half of median length of tergite II........ 46</p><p>- Metasoma with pair of lateral depressions not bigger than half lenght of tergite II.................................. 55</p><p>46. Inner margin of basivolsella toothed...................................................................... 47</p><p>- Inner margin of basivolsella not toothed................................................................... 48</p><p>47. Basivolsella with tooth in apical portion; base of digitus produced; apex of aedeagal ventral ramus bidentate (Fig. 108); apex of outer pair of apical lobes of aedeagal dorsal body acute (Figs 107, 109)......................... D. geniculatus Azevedo</p><p>- Basivolsella with tooth in medial portion; base of digitus not produced; apex of aedeagal ventral ramus smooth; apex of outer pair of apical lobes of aedeagal dorsal body truncate........................................... D. octavus Azevedo</p><p>48. Apex of aedeagal ventral ramus with two teeth, resembling fish tale in lateral view (Fig. 117)....... D. piscicercus Azevedo</p><p>- Apex of aedeagal ventral ramus with rounded or acute apex not as above......................................... 49</p><p>49. Apex of aedeagal ventral ramus rounded.................................................................. 50</p><p>- Apex of aedeagal ventral ramus angulate or subangulate...................................................... 51</p><p>50. Aedeagal ventral ramus longer than aedeagal dorsal body, outer margin expanded on median portion; aedeagal dorsal body with four pairs of apical lobes........................................................ D. pedipalpoides Azevedo</p><p>- Aedeagal ventral ramus much longer than aedeagal dorsal body, outer margin excavated on median portion; aedeagal dorsal body with three pairs of apical lobes......................................................... D. trogon Azevedo</p><p>51. Aedeagal ventral ramus with inner margin of apex bifurcated.................................................. 52</p><p>- Aedeagal ventral ramus with inner margin of apex not bifurcated............................................... 53</p><p>52. Posterior hypopygeal margin concave, corners angulate; aedeagal ventral ramus longer than aedeagal dorsal body (Fig. 121); aedeagal dorsal body with three pairs of apical lobes, apex as wide as median and basal portion (Fig. 119).................................................................................................... D. strepsus Azevedo</p><p>- Posterior hypopygeal margin straight, corners rounded; aedeagal ventral ramus shorter than aedeagal dorsal body (Fig. 114); aedeagal dorsal body with two pairs apical lobes, apex narrower than median and basal portion (Fig. 113).. D. latus Azevedo</p><p>53. Posterior hypopygeal margin straight, with corners subangulate; aedeagal ventral ramus as long as aedeagal dorsal body or nearly so, apex straight................................................................................ 54</p><p>- Posterior hypopygeal margin concave with corners rounded; aedeagal ventral ramus shorter than aedeagal dorsal body, apex divergent (Fig. 112)................................................................ D. intradentatus Azevedo</p><p>54. Inner margin of paramere without pegs; apical lobes of aedeagal dorsal body elliptical; apex of aedeagal ventral ramus suban- gulate................................................................................ D. gladius Azevedo</p><p>- Inner margin of paramere with four pegs; apical lobes of aedeagal dorsal body diamond-shaped; apex of aedeagal ventral ramus abruptly acute................................................................ D. alticarinatus Azevedo</p><p>55. Aedeagal ventral ramus tubular; aedeagal dorsal body with pair of basal process tridentate........... D. brasiliensis Kieffer</p><p>- Aedeagal ventral ramus laminar; aedeagal dorsal body not as above............................................. 56</p><p>56. Tergal process with tuft of setae with distinct long lateral setae................................................. 57</p><p>- Tergal process not as above............................................................................. 58</p><p>57. Median clypeal lobe with three small teeth; apical lobes of aedeagal dorsal body long and narrow, narrowing slightly apicad......................................................................................... D. gilvipes Evans</p><p>- Median clypeal lobe with one angulate tooth (Fig. 17); apical lobes of aedeagal dorsal body short and wide but narrowing abruptly apicad (Fig. 165)................................................................ D. arbeius sp. nov.</p><p>58. Tergal process with pair of lateral drop-shaped pits, with elevated rim, sharp anteriorly and rounded posteriorly.................................................................................................. D. guttus Azevedo</p><p>- Metasomal tergite not as above.......................................................................... 59</p><p>59. Tergal processes with pair inclined and linear depression..................................................... 60</p><p>- Tergal processes not as above........................................................................... 61</p><p>60. Aedeagal ventral ramus much shorter than aedeagal dorsal body, apical half evenly arched, apex sharp..... D. infissus Evans</p><p>- Aedeagal ventral ramus as long as aedeagal dorsal body, apical half sinuous, apex narrow but rounded (Fig. 175)............................................................................................... D. rufipalpis Kieffer</p><p>61. Tergal process with tubercle............................................................................ 62</p><p>- Tergal process without tubercle.......................................................................... 69</p><p>62 Apical lobes of aedeagal dorsal body mask-shaped (Figs 202, 204–206)............. D. personatus Redighieri &amp; Azevedo</p><p>- Apical lobes of aedeagal dorsal body without mask-shaped.................................................... 63</p><p>63. Mandible with two distal teeth.......................................................................... 64</p><p>- Mandible with four distal teeth.......................................................................... 66</p><p>64. Hypopygeal margin straight or nearly so; paramere wide; aedeagal dorsal body triangular-shaped..................... 65</p><p>- Hypopygeal margin concave; paramere very wide; aedeagal dorsal body bottle-shaped......... D. pilus Alencar &amp; Azevedo</p><p>65. Tergal process with deep depression (Fig. 61); apical margin of paramere truncate (Fig. 217); aedeagal ventral ramus as long as dorsal body, long, simple, curved outward (Fig. 218)............................................ D. isortus sp. nov.</p><p>- Tergal process with shallow depression; apical margin of paramere convex; aedeagal ventral ramus shorter than aedeagal dor- sal body, wide, with apical margin inclined and straight, inner corner sharpened..................... D. strabus Azevedo</p><p>66. Median clypeal tooth angulate and very short (Fig. 29); tubercle of tergal process without pit on top (Fig. 60); apical lobes of aedeagal dorsal body connected to median portion of aedeagal body by four narrow projections (Fig. 215)...................................................................................................... D. ilarius sp. nov.</p><p>- Median clypeal tooth regular sized; tubercle of tergal process with pit on top; apical lobes of aedeagal dorsal body not con- nected to median portion of aedeagal dorsal body as above.................................................... 67</p><p>67. Median clypeal carina angled in profile; basal margin of dorsal margin of paramere very projected (Fig. 213); aedeagal ventral ramus with apical portion divergent (Fig. 214)................................................ D. geortus sp. nov.</p><p>- Median clypeal carina straight in profile; basal margin of dorsal margin of paramere not as above; aedeagal ventral ramus with apical portion parallel................................................................................. 68</p><p>68. Posterior hypopygeal margin slightly convex; apical portion of dorsal margin of paramere not excavated; apical lobes of aedea- gal dorsal body long................................................................... D. plaummani Evans</p><p>- Posterior hypopygeal margin sinuous (Fig. 89); apical portion of dorsal margin of paramere very excavated (Fig. 211); apical lobes of aedeagal dorsal body very short (Fig. 212).............................................. D. borus sp. nov.</p><p>69. Tergal process with deeply excavated depression covered by membrane (Fig. 63); posterior hypopygeal margin strongly con- cave (Fig. 94); dorsal margin of paramere very excavated (Fig. 221).............................. D. ubracus sp. nov.</p><p>- Tergal process not as above; posterior hypopygeal margin straight or slightly concave; dorsal margin of paramere not very excavated........................................................................................... 70</p><p>70. Depression of tergal processes large and somewhat triangular; aedeagal ventral ramus laminar with apical wing-like expansion (Fig. 201)........................................................................... D. rettenmeyeri Evans</p><p>- Tergal process without depression or with circular or subcircular depression, aedeagal ventral ramus not as above........ 71</p><p>71. Basipamere about 2.0 times as long as size of paramere (Fig. 209); aedeagal dorsal body entirely narrow; aedeagal ventral ramus entirely very narrow (Fig. 210)..................................................... D. arteobius sp. nov.</p><p>- Basipamere as long as paramere or nearly so, aedeagal dorsal body entirely wide; aedeagal ventral ramus wide, at least on basal or/and medial portion............................................................................. 72</p><p>72. Apical lobes of aedeagal dorsal body very short; aedeagal ventral ramus with inner margin concave at apical half with long median filament..................................................................... D. xanthopus Ashmead</p><p>- Apical lobes of aedeagal dorsal body long (Fig. 207); aedeagal ventral ramus with inner margin straight without apical fila- ment (Fig. 208)......................................................................... D. abarus sp. nov.</p></div>	https://treatment.plazi.org/id/3B3687C07C759B2DFF67F3AB9AF9FAA3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7A9B2DFF67F7F29A24F962.text	3B3687C07C7A9B2DFF67F7F29A24F962.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus altivolans Evans 1954	<div><p>altivolans species-group</p><p>Diagnosis. Male. Metasoma with median depression, dense tufts of setae. Digitus serrated with rounded teeth inner upper margin. Aedeagal ventral ramus wide.</p><p>Remarks. It has six species: D. altivolans Evans, D. deformis Evans, D. megomphalus Evans, D. subdeformis Azevedo, D. lanceolatus Azevedo and D. isteus sp. nov. .</p><p>Distribution. Ranging from northern U.S.A. to north of Argentina, including some Caribbean islands.</p></div>	https://treatment.plazi.org/id/3B3687C07C7A9B2DFF67F7F29A24F962	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7A9B2FFF67F5309EFFFF28.text	3B3687C07C7A9B2FFF67F5309EFFFF28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus altivolans Evans 1954	<div><p>Dissomphalus altivolans Evans, 1954</p><p>(Figs 95–96)</p><p>Dissomphalus altivolans Evans, 1954: 294, 297, 301, 307–308 (♂, holotype from Tallulah, USNM, figs 7, 21) 1978: 40, 41–42; Krombein 1979: 1217; Gordh &amp; Móczár 1990: 218, Azevedo 1999a: 356, 357–358, 390.</p><p>Dissomphalus nigrescens Evans, 1954: 294, 297, 299, 308–309; 1962: 68, 75–76; 1964: 45; 1978:40, 41, 47–48; Krombein, 1979: 1217; Gordh &amp; Móczár, 1990: 224, Azevedo 1999a: 357 (designation of synonymy).</p><p>Diagnosis. Mandible tridentate. Clypeus tridentate. Tergal process with median concavity not so large and deep, with pair of lateral subcircular tufts densely covered by short setae. Hypopygium with posterior margin concave, corners rounded. Genitalia: paramere with apex rounded; aedeagal ventral ramus shorter than aedeagal dorsal body, inner margin smooth, parallel and straight; outer margin smooth, slightly convergent; surfaces horizontal; apex rounded, basal margin with median invagination; aedeagal dorsal body with two pairs of apical lobes; outer lobe wide, with rounded apex, inner margin concave, convergent dorsally; inner lobe membranous and setose. Apodeme extending beyond genital ring.</p><p>Remarks. No additional material of this species was not found in this study.</p><p>Material examined. Holotype, ♂, Tallula, La., “Airpl. 794”, take in airplane trap between 20 and 500 feet altitude, P.A. Glick (USNM nº. 62277).</p><p>Distribution (Fig. 225). United States of America (Illinois, Indiana, California, Missouri, Arizona, Arkansas, Alabama, Louisiana and Texas), Mexico (Sonora, Sinaloa, Jalisco, Quintana Roo and Chiapas), Guatemala, Honduras and Panama.</p></div>	https://treatment.plazi.org/id/3B3687C07C7A9B2FFF67F5309EFFFF28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C789B28FF67F5329DE9FE2A.text	3B3687C07C789B28FF67F5329DE9FE2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus subdeformis Azevedo. A 1999	<div><p>Dissomphalus subdeformis Azevedo, 1999</p><p>(Figs 97–98)</p><p>Dissomphalus subdeformis Azevedo, 1999a: 356, 360–361, 382, 391 (♂ holotype from Venezuela, USNM, figs 122–123, 203).</p><p>Diagnosis. Mandible tridentate. Clypeus tridentate. Tergal process with large and deep depression with pair of elliptical lateral tuft of setae directed slightly each other. Hypopygium with posterior margin slightly concave. Genitalia: paramere slightly arched inward; aedeagal ventral ramus shorter than aedeagal dorsal body, wide, narrowing slightly apicad; inner margin diverging and outer margin convergent; aedeagal dorsal body with two pairs of apical lobes; outer lobe wide, with apex rounded, dorsal margin much produced inward; inner lobe setose and serrated. Apodeme extending beyond genital ring.</p><p>Remarks. No additional material of this species was not found in this study.</p><p>Material examined. Holotype, ♂, VENEZUELA, Barinas: Rio Caparo, Research Sta., 32 Km E, El Cornern, seasonal forest, light trap black, 3–5.ii.1978, J.B. Heppener col. (USNM).</p><p>Distribution (Fig. 242). Guatemala, El Salvador, Nicaragua, Costa Rica, Panama, Colombia, Curação, Venezuela, Trinidad &amp; Tobago and Surinam.</p></div>	https://treatment.plazi.org/id/3B3687C07C789B28FF67F5329DE9FE2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7F9B2AFF67F4969C4DF84C.text	3B3687C07C7F9B2AFF67F4969C4DF84C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus isteus Azevedo 2017	<div><p>Dissomphalus isteus Brito &amp; Azevedo, sp. nov.</p><p>(Figs 1–2, 33, 64, 99–100)</p><p>Description. Male. Body length 2.24 mm. Color: body castaneous. Head (Figs 1–2). Mandible with three distal teeth. Median clypeal lobe ill-defined, with one rounded and very large tooth, with high keel; median clypeal carina very high in profile, incomplete apically, straight in profile or nearly so. Gena with spine. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 33). Tergal process median, with very shallow, subcircular and large depression, without tubercle, with short and dense tuft of setae. Posterior hypopygeal margin straight with median concavity (Fig. 64). Genitalia (Figs 99–100): paramere with dorsal margin entirely wide and sinuous; apical margin truncate. Aedeagal ventral ramus as long as dorsal body, wide, progressively narrowing apicad; cross section laminar; ramous bifid; surface horizontal; inner margin straight; outer margin slightly sinuous; apex long, simple, parallel; additional inner ramus short; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, subvertical and narrow, with apical margin angled and posterad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Remarks. This species is easily recognized by having a very large and high median clypeal tooth which shows high keel in profile view, gena with spine, the dorsal margin of paramere very sinuous with long and thick setae and the aedeagal ventral ramus bifid. This species is similar to D. cornutus Evans by having a spine in ventral region of head. However, this species has large spine in gena, whereas D. cornutus has shorter spine in hypostomal carina.</p><p>Material examined. Holotype, ♂, PANAMA, Panama Pr [ovincia]: P[arque] Nac[ional] Altos de Campana, [08º43'15"N, 80º01'43"W], 10–14 ago 1999, Tr[ampa] Malaise, A. Santos, P. González [col.] (MIUP). Paratypes: Coclé Pr [ovincia]: 1♂, El Copé, Par[que] Nac[ional General de División] Omar Torrijos H[errera], 570– 670 m.s.n.m., [08º41'56"N, 80º39'17"W], 26–30 sep. 1999, Tr[ampa] Malaise, A. Santos, P. González [col.] (MIUP); 1♂, same province of holotype, P[arque] Nac[ional] Sob[eranía], Camino, Plantación, [09º09'N, 79º43'W], 24 ago.–2 sep.1999, [Trampa] Malaise, A. Santos, P. González [col.] (MIUP).</p><p>Distribution (Fig. 238). Panama (Coclé and Panama).</p></div>	https://treatment.plazi.org/id/3B3687C07C7F9B2AFF67F4969C4DF84C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7C9B2BFF67F7699D9EF9B6.text	3B3687C07C7C9B2BFF67F7699D9EF9B6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus amplus Redighieri & Azevedo 2006	<div><p>amplus species-group</p><p>Diagnosis. Male. Head regularly wide. Mandible regularly long. Metasomal tergite II without tergal process. Aedeagal ventral ramus without apical filament. Aedeagal dorsal body without apical flagellae.</p><p>Remarks. It has eight species: D. amplus Redighieri &amp; Azevedo, D. clovisi Colombo &amp; Azevedo, D. kuara Colombo &amp; Azevedo, D. pyata Colombo &amp; Azevedo, D. miriamae Colombo &amp; Azevedo, D. azagus sp. nov., D. asfartus sp. nov. and D. iozus sp. nov. .</p><p>Distribution. Panama and Brazil (Espírito Santo).</p></div>	https://treatment.plazi.org/id/3B3687C07C7C9B2BFF67F7699D9EF9B6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7C9B35FF67F4839C64F880.text	3B3687C07C7C9B35FF67F4839C64F880.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus azagus Azevedo 2017	<div><p>Dissomphalus azagus Brito &amp; Azevedo sp. nov.</p><p>(Figs 3, 34, 65, 101–102)</p><p>Description. Male. Body length 2.27 mm. Color: head and mesosoma castaneous; metasoma light castaneous. Head (Fig. 3). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina high in profile, complete or nearly so, angled in profile. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 34). Tergal process absent. Posterior hypopygeal margin sinuous (Fig. 65). Genitalia (Figs 101–102): paramere with dorsal margin wide basally; apical margin rounded. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin slightly sinuous; apex short, simple, parallel; additional inner ramus absent; basal stub short and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and narrow, with apical margin acute and dorsad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Remarks. This species is easily recognized by having the paramere entirely wide with the apex very excavated, the outer lobes of the aedeagal dorsal body with the distal region mesad and L-shaped, and the apex of the aedeagal ventral ramus acute. This species is similar to D. borus sp. nov. by having the apex of paramere very excavated, the dorsal margin almost straight and the basiparamere shorter than the paramere. However, this species has the apical lobes of the aedeagal dorsal body long and the apex of the aedeagal ventral ramus abruptly acute, whereas D. borus sp. nov. has the apical lobes of the aedeagal dorsal body very short and the aedeagal ventral ramus progressively narrowing apicad.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Material</a> examined. Holotype, ♂, PANAMA, Bocas del Toro: Wekso, Teribe, P [arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ampa] Malaise, A. Santos[col.] (MIUP) . Paratypes: 4♂, same data as holotype (MIUP); 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.454994&amp;materialsCitation.latitude=9.47425" title="Search Plazi for locations around (long -82.454994/lat 9.47425)">Par</a> [que] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.454994&amp;materialsCitation.latitude=9.47425" title="Search Plazi for locations around (long -82.454994/lat 9.47425)">Nac</a> [ional] Humedal de San [-]San Pond Sac. {= Sak}, [09°28'27.3"N, 82°27'18"W], 26–28 Oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP) .</p><p>Distribution (Fig. 231). Panama (Bocas del Toro).</p></div>	https://treatment.plazi.org/id/3B3687C07C7C9B35FF67F4839C64F880	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C619B37FF67F6D99AF9FAD2.text	3B3687C07C619B37FF67F6D99AF9FAD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus asfartus Azevedo 2017	<div><p>Dissomphalus asfartus Brito &amp; Azevedo, sp. nov.</p><p>(Figs 4, 35, 66, 103–104)</p><p>Description. Male. Body length 2.03 mm. Color: head and mesosoma castaneous; metasoma light castaneous. Head (Fig. 4). Mandible with three distal teeth. Median clypeal lobe subtrapezoidal, with one angulate tooth; median clypeal carina low in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 35). Tergal process absent. Posterior hypopygeal margin sinuous (Fig. 66). Genitalia (Figs 103–104): paramere with dorsal margin wide basally; apical margin rounded. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin sinuous; apex long, crenulate, slightly curved outward; additional inner ramus absent; basal stub short and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, horizontal and wide, with apical margin straight and dorsad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Description. Female. (Fig. 223). Body length 1.30 mm. Body and legs light castaneous, eye black. Head 1.46 × as long as wide, side parallel slightly convex. Frons densely coriaceous, punctures large, dense and shallow. Mandible with four apical teeth, lower one larger. Clypeus with subtrapezoidal median lobe and median carina high. Eye with two facets, subcircular, distant from mandible 1.66 × its length. Vertex concave; occipital carina visible in dorsal view. Gena 0.42 × length of head. Mesosoma coriaceous. Pronotal collar 0.32 × length of pronotal disc, 1.19 × as long as wide. Mesoscutum 0.48 × length of pronotal disc, posterior margin convex. Propodeal disc long, 1.62 × as long as wide; spiracle placed at dorsolateral of propodeum, attached to lateral carina. Legs not spinose. Metasoma polished, 0.86 × as long as mesosoma. Petiole short with anterior margin in v-shaped and rugose.</p><p>Remarks. This species is easily recognized by having the eyes bulging, the dorsal margin of the paramere very expanded distally downward and the apical margin of the aedeagal dorsal body crenulate. This species is similar to D. krombeini Azevedo by having the apical portion of the dorsal margin of the paramere excavated and the aedeagal dorsal body abruptly narrowing apicad. However, this species has the dorsal margin of the paramere with the distal margin expanded downward, the apical margin of the aedeagal dorsal body crenulate and the tergal process absent, whereas D. krombeini has the dorsal margin of the paramere with the distal margin not expanded downward, the apical margin of the aedeagal dorsal body smooth and has the tergal process with the tuft of setae with one distinctly long seta.</p><p>Material examined. Holotype, ♂, Darién: P[arque] N[acional] Darién, Pirre, 900m, Est [ación] Rancho Frio, 21 mar–4 ab 2000, Tr [ampas] ama[rillas], Cambra, Santos, Bermudez (MIUP) . Allotype in copula, ♀, same data as holotype . Paratypes: Bocas del Toro: 5 ♂, Wekso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Teribe</a>, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP) ; Chiriqui Pr [ovincia]: 28♂, P[arque] I[nternacional] L[a] A[mistad], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.73333&amp;materialsCitation.latitude=8.9" title="Search Plazi for locations around (long -82.73333/lat 8.9)">Jurutungo</a>, 1800 m, [08°54'N, 82°44'W], 30 oct.–5 nov. 1994, Tr [ampa] Malaise (25♂) and Tr [ampas] Amarillas (3♂), A. Rodriguez [col.] (MIUP) ; 10 ♂, 2 km W <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.583336&amp;materialsCitation.latitude=8.85" title="Search Plazi for locations around (long -82.583336/lat 8.85)">Cerro Punta</a>, 1700 m, [08°51'N, 82°35'W], 19. V–8.VI.1977, Peck &amp; Howden [col.] (CNCI) ; 8 ♂, Finca La Suiza near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 25–6.VI.2000, F[light] I[nterception] T[rap], H. &amp; A. Howden [col.] (7♂) and 2.VI.2000, Howden [col.] (1♂) (CNCI) ; 3♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.24944&amp;materialsCitation.latitude=8.744167" title="Search Plazi for locations around (long -82.24944/lat 8.744167)">La</a> Fortuna Dam, 1200 m, wet forest, [08°44'39"N, 82°14'58"W], 14–VI–16.VII– 1982, F[light] I[nterception] T[rap], B. Gill [col.] (1♂, PMAE), (2♂, CNCI) ; 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.20833&amp;materialsCitation.latitude=8.662777" title="Search Plazi for locations around (long -82.20833/lat 8.662777)">Quebrada de la Mina</a>, 1249 m, 8°39'46"N, 82°12'30"W, 27–28. V.2000, Y[ellow] P[an] T[rap], B. &amp; J. Gill [col.] (CNCI) ; Veraguas Pr [ovincia]: 8♂, P[arque] N[acional], Isla Coiba, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Cerro la Equis</a>, 200 m, [07º28'09"N, 81º45'24"W], 30–nov–4–dic.1998, [Trampa] Malaise, R. Cambra, S. Bermudez [col.] (1♂) , 0m (1♂), 29 jul – 4 ag 1998, 120m (2♂), 100m (1♂), 8–12 mar 1998, 90m (3♂); [Los Santos]: 2♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.416664&amp;materialsCitation.latitude=7.5166664" title="Search Plazi for locations around (long -80.416664/lat 7.5166664)">Ojo de Agua</a>, [07°31'N, 80°25'W], 30.VI–14.VII.1982, B. Gill [col.] (PMAE) ; Coclé Pr [ovincia]: 3♂ El Copé, Par [que] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">Nac</a> [ional <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">General de División</a>] Omar Torrijos H [errera], 570– 670 m.s.n.m., [08º41'56"N, 80º39'17"W], 26–30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; Panama Pr [ovincia]: 2♂, P[arque] N[atural] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-89.54472&amp;materialsCitation.latitude=8.9925" title="Search Plazi for locations around (long -89.54472/lat 8.9925)">Metropolitano</a>, [08º59'33"N, 89º32'41"W], 24 ago.–3 sep.1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; 1♂, same data as holotype, except Malaise (MIUP); 1♂, same data as holotype, except 1000 m (MIUP); 1♂, same data as holotype, except 580m (MIUP); 1♂, same province of holotype, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-87.61667&amp;materialsCitation.latitude=7.85" title="Search Plazi for locations around (long -87.61667/lat 7.85)">Cana</a>, 1380 m, [07°51'N, 87°37'W], 4–7.VI.1986, F[light] I[nterception] T[rap], [J.] Ashe &amp; [R.] Brooks [col.] (CNCI) ; 1♂, same province of holotype, Estac [ión] Cruce de Mono, P[arque] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-87.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -87.47019/lat 7.8573055)">Nac</a> [ional] Darién, [07°51'26.3"N, 87°28'12.7"W], 6 feb–4 mar 1993, tr[ampa] Malaise, R. Cambra, J. Coronado [col.] (MIUP) .</p><p>Distribution (Fig. 230). Panama (Bocas del Toro, Chiriqui, Veraguas, Los Santos, Coclé, Panama and Darién).</p></div>	https://treatment.plazi.org/id/3B3687C07C619B37FF67F6D99AF9FAD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C609B30FF67F7609CCDFE0F.text	3B3687C07C609B30FF67F7609CCDFE0F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus iozus Azevedo 2017	<div><p>Dissomphalus iozus Brito &amp; Azevedo, sp. nov.</p><p>(Figs 5, 36, 67, 105–106)</p><p>Description. Male. Body length 3.13 mm. Color: body castaneous. Head (Fig. 5). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina low in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 36). Tergal process absent. Posterior hypopygeal margin straight or nearly so (Fig. 67). Genitalia (Figs 105–106): paramere with dorsal margin entirely wide; apical margin rounded. Aedeagal ventral ramus shorter than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin sinuous; outer margin slightly sinuous; apex short, simple, parallel; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and wide, with apical margin rounded and dorsad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Remarks. This species is easily recognized by having the genitalia robust, the lobes of the aedeagal dorsal body with the apical region more expanded, rounded, laminar and the aedeagal ventral ramus very wide. This species is similar to D. abarus sp. nov. by having the dorsal margin of the paramere rounded, the outer lobes of the aedeagal dorsal body long and the aedeagal ventral ramus wide. However, this species has the apex of the apical lobes of the aedeagal dorsal body rounded and laminar, the apex of the aedeagal ventral ramus without projections, whereas D. abarus sp. nov. has the apex of the aedeagal ventral ramus not rounded nor laminar and the apex of the aedeagal ventral ramus with two small rounded projections.</p><p>Material examined. Holotype, ♂, PANAMA, Veraguas Pr [ovincia]: P[arque] N[acional] Coiba, Isla Coiba, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Cerro la Equis</a>, 120 m, [07º28'09"N, 81º45'24"W], 29 jul–4 ag 1998, Malaise, R. Cambra, A. Santos [col.] (MIUP). Paratypes: 1♂, same data as holotype ; Panama Pr [ovincia]: 1♂, [Panama], P[arque] N[atural] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-89.54472&amp;materialsCitation.latitude=8.9925" title="Search Plazi for locations around (long -89.54472/lat 8.9925)">Metropolitano</a>, [08º59'33"N, 89º32'41"W], 24 ago.–3 sep.1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; Darién Pr [ovincia]: 3♂, P[arque] N[acional] Darién, Pirre, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-87.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -87.75/lat 8.0)">Est</a> [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-87.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -87.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 87°45'W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP).</p><p>Distribution (Fig. 237). Panama (Veraguas, Panama and Darién).</p></div>	https://treatment.plazi.org/id/3B3687C07C609B30FF67F7609CCDFE0F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C669B31FF67F2A89CE6FEBC.text	3B3687C07C669B31FF67F2A89CE6FEBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus apertus Kieffer, Alencar & Azevedo 1914	<div><p>apertus species-group</p><p>Diagnosis. Male. Metasomal tergite II with pair of tuft of setae very large and rounded placed in very shallow depression.</p><p>Remarks. It has three species: D. apertus Kieffer, D. subpilosus Azevedo and D. parvus Azevedo. Distribution. Ranging from northeast of the U.S. A to Bolivia.</p></div>	https://treatment.plazi.org/id/3B3687C07C669B31FF67F2A89CE6FEBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C669B31FF67F3899F92FAD2.text	3B3687C07C669B31FF67F3899F92FAD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus apertus Kieffer, Alencar & Azevedo 1914	<div><p>Dissomphalus apertus Kieffer, 1914</p><p>Dissomphalus apertus Kieffer 1914a: 60 –61 (♂ holotype from United States) 1914b: 496, 499, Evans 1954: 295, 297, 306– 307, 1962: 68, 75; Evans 1964: 45, 1978: 40, 41, 42–43; Krombein 1963: 263, 1979: 1217; Gordh &amp; Móczár 1990: 218; Azevedo 1999a: 352 –353, 390.</p><p>Psilobethylus foveolatus Brown &amp; Cheng 1951: 143 –146, Krombein 1979: 1217.</p><p>Dissomphalus foveolatus (Brown &amp; Cheng) Evans 1954: 306 –307 (genus combination); Krombein 1963: 263; Evans 1978: 42 (designation of synonymy).</p><p>Diagnosis. Male. Mandible tridentate. Clypeus tridentate. Tergal process with very shallow depression, pair of very large and rounded depression with dense tuft of setae. Hypopygium with posterior margin straight. Genitalia: paramere with apex rounded; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar, margins smooth, inner margin straight and outer margin slightly convergent; apex wide with outer corner angulate; aedeagal dorsal body with two pairs of apical lobes; outer pair wide, with apex rounded, inner surface concave, convergent dorsally, dorsal margin with rounded expansion inward; inner pair membranous and setose. Apodeme not extending beyond genital ring.</p><p>Remarks. No additional material of this species was not found in this study. The holotype is lost, but the identity of this species is well-established. The diagnosis presented above is based on a male from Mexico as listed below.</p><p>Material examined. MEXICO, Quintana Roo: 3Km, S Puente Morelos, Jd Botânico, 14. XII.1993, L. Masner col. (CNCI).</p><p>Distribution (Fig. 227). United States of America (New York, Pennsylvania, Maryland, Nebrasca, Illinois, Ohio, New Jersey, D.C, Delaware, Kansas, Missouri, Virginia, North and South Carolina, New Mexico, Texas and Florida), Mexico (Jalisco, Vera Cruz, Quintana Roo and Chiapas), Belize, Honduras, Guatemala, Costa Rica and Panama.</p></div>	https://treatment.plazi.org/id/3B3687C07C669B31FF67F3899F92FAD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C669B31FF67F75E9B3DF8AF.text	3B3687C07C669B31FF67F75E9B3DF8AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus bicavatus	<div><p>bicavatus species-group</p><p>Diagnosis. Male. Mandible bidentate. Pronotal disc usually with texture and punctures as frons. Mesoscutum and scutellum with texture weaker, punctures smaller than those of frons. Metasoma with pair of lateral and anterior depression, large usually more than 0.5 × median length of tergite II. Hypopygeal median stalk slender. Female. Mandible tetradentate. Clypeus tridentate. Metapectal propodeal disc with sides nearly parallel.</p><p>Remarks. It has 34 species: D. abruptus Azevedo, D. alticarinatus Azevedo, D. archeatus Azevedo, D. basivolsellus Azevedo, D. bivolsellus Azevedo, D. curvilongus Azevedo, D. elongatus Azevedo, D. falciformes Azevedo, D. flagellatus Azevedo, D. geniculatus, D. gladius Azevedo, D. intradentatus, D. largimanus Azevedo, D. latus Azevedo, D. lobisserratus Azevedo, D. longimerus Azevedo, D. longipilosus Azevedo, D. mantoides Azevedo, D. megadentatus Azevedo, D. microdentatus Azevedo, D. octavus Azevedo, D. pedipalpoides Azevedo, D. piscicercus, D. rectangularis Azevedo, D. retorcerens Azevedo, D. sinuatus Azevedo, D. spiculus Azevedo, D. strepsus, D. subtriangularis Azevedo, D. tetracerutus Azevedo, D. tetralobatus Azevedo, D. trogon Azevedo, D. tropoides Azevedo, D. falcatus Evans and D. bicavatus Evans.</p><p>Distribution. Ranging from South of Mexico to Southeastern Brazil (São Paulo).</p></div>	https://treatment.plazi.org/id/3B3687C07C669B31FF67F75E9B3DF8AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C659B32FF67F2A89D8AFCA6.text	3B3687C07C659B32FF67F2A89D8AFCA6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus geniculatus Azevedo 2001	<div><p>Dissomphalus geniculatus Azevedo, 2001</p><p>(Figs 107–109)</p><p>Dissomphalus geniculatus Azevedo, 2001: 174, 185–186 (♂, holotype from Costa Rica, EMUS, figs 67–68).</p><p>Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with pair of antero-lateral depressions, ovoid, deep, large with small pit covered by tuft of long seate. Hypopygium with posterior margin concave. Genitalia: paramere with oblique and concave apex; aedeagal ventral ramus as long as aedeagal dorsal body, laminar, surface subvertical, apical half convergent, apex bidentate; aedeagal dorsal body with three pairs of apical lobes; outer pair large, laminar, surface horizontal, outer margin with six teeth sharpened progressively, inner margin with projection; dorsal inner pair stout, membranose and setose; ventral inner pair small, laminar, narrowing to apex; basal process of aedeagal dorsal body with pair of lateral teeth. Apodeme extending beyond genital ring.</p><p>Remarks. This species is known to Costa Rica and Panama (Azevedo 2001) and now more specimens are added. In this series the specimens are quite similar to those previously studied, without any variation that worths to mention.</p><p>Material examined. Holotype ♂, COSTA RICA, Guanacaste: 3km SE Rio Naranjo, 1.XI.1991, F. Parker col. (EMUS) ; PANAMA, Bocas del Toro Pr [ovincia]: 1♂, Par [que] Nac [ional] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.454994&amp;materialsCitation.latitude=9.47425" title="Search Plazi for locations around (long -82.454994/lat 9.47425)">Humedal de San</a> [-] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.454994&amp;materialsCitation.latitude=9.47425" title="Search Plazi for locations around (long -82.454994/lat 9.47425)">San Pond Sac</a> {= <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.454994&amp;materialsCitation.latitude=9.47425" title="Search Plazi for locations around (long -82.454994/lat 9.47425)">Sak</a>}, [09°28' 27.3"N, 82°27'18"W], 26–28 Oct. 1999, Tr [ampa] Malaise, A. Santos[col.] (MIUP) ; Darién Pr [ovincia]: 1♂, P[arque] N[acional] Darién, Pirre, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Est</a> [ción] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 77°45'W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP).</p><p>Distribution (Fig. 234). Costa Rica and Panama.</p></div>	https://treatment.plazi.org/id/3B3687C07C659B32FF67F2A89D8AFCA6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C659B32FF67F1F29C56F928.text	3B3687C07C659B32FF67F1F29C56F928.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus intradentatus Azevedo 2001	<div><p>Dissomphalus intradentatus Azevedo, 2001</p><p>(Figs 110–112)</p><p>Dissomphalus intradentatus Azevedo, 2001: 174, 186–187 (♂, holotype from Panama, PMAE, figs 82–88).</p><p>Diagnosis. Mandible bidentate. Clypeus tridentate. Tergal process with pair of antero-lateral depressions, ovoid, deep, large with small pit covered by tuft of long seate. Hypopygium with posterior margin slightly concave. Genitalia: paramere with apex concave; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar, surface sub-horizontal; outer margin concave apically; apex acute, slightly diverging; aedeagal dorsal body with four pairs of apical lobes, outer pair laminar, surface vertical, narrowing at apex and directed downward, apical; dorsal margin with short expansion and small teeth; ventral margin with large tooth, three inner pairs stout, membranose and setose; basal process with pair of lateral teeth directed outward, margin between them straight. Apodeme not extending beyond genital ring.</p><p>Remarks. This species is known to Panama (Azevedo 2001) and here more specimens are added. Some variations are worth to mention: mesosoma and metasoma castaneous; median clypeal tooth angulate; median clypeal carina complete apically; vertex crest straight; anterior margin of pronotal disc ecarinate; basal process of aedeagal dorsal body without teeth.</p><p>Material examined. Holotype ♂, PANAMA, Canal Zone: Barro Colorado Island, 3-13. VI.1983. B . Gill col. (PMAE); PANAMA, Darién Pr[ovincia]: 14♂, P[arque] N[acional] Darién, Pirre, Est [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-87.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -87.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 87°45'W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP); 2♂, Cruce de Mono: Est [ación] I[nstituto] N[acional de] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-87.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -87.47019/lat 7.8573055)">Re</a> [cusos] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-87.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -87.47019/lat 7.8573055)">Na</a> [turales] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-87.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -87.47019/lat 7.8573055)">Re</a> [novables], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-87.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -87.47019/lat 7.8573055)">Parque</a>] nacional Darién, [07°51'26.3"N, 87°28'12.7"W], 5–28 feb 1993, Tr[ampas] Amarillas, R. Cambra, J. Coronado [col.] (MIUP).</p><p>Distribution (Fig. 237). Panama (Panama and Darién).</p></div>	https://treatment.plazi.org/id/3B3687C07C659B32FF67F1F29C56F928	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C659B3CFF67F5759C29FEC2.text	3B3687C07C659B3CFF67F5759C29FEC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus latus Azevedo 2001	<div><p>Dissomphalus latus Azevedo, 2001</p><p>(Figs 113–114)</p><p>Dissomphalus latus Azevedo, 2001: 175, 188–189 (♂ holotype from Trinidad &amp; Tobago, EMUS figs 98–101).</p><p>Diagnosis. Mandible bidentate. Clypeus subtrapezoidal. Tergal process with pair of antero-lateral depressions, ovoid, deep, margin with setae, pit with high border covered by tuft of long setae in middle of depression. Hypopygium with posterior margin straight. Genitalia: paramere with apex oblique and truncate; aedeagal dorsal ramus shorter than aedeagal dorsal body, laminar; surface horizontal, evenly wide; apex bidentate; aedeagal dorsal body with two pairs of apical lobes; outer pair laminar, surface vertical, wide; apex rounded and directed downward; inner pair, membranous, stout and setose. Apodeme extending beyond genital ring.</p><p>Remarks. No additional material of this species was not found in this study. Material examined. Holotype ♂, TRINIDAD &amp; TOBAGO, Trinidad Island: Simila Reserve Station, 2– 15.VI.1981, Hanson &amp; Clemons col. (EMUS).</p><p>Distribution (Fig. 238). Panama and Trinidad &amp; Tobago.</p></div>	https://treatment.plazi.org/id/3B3687C07C659B3CFF67F5759C29FEC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6B9B3CFF67F3509CCDFB45.text	3B3687C07C6B9B3CFF67F3509CCDFB45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus piscicercus Azevedo 2001	<div><p>Dissomphalus piscicercus Azevedo, 2001</p><p>(Figs 115–118)</p><p>Dissomphalus piscicercus Azevedo, 2001: 174, 196–197 (♂, holotype from Panama, PMAE, figs 148–154).</p><p>Diagnosis. Mandible bidentate. Clypeus tridentate. Tergal process with pair of antero-lateral depressions, ovoid, deep, large with small pit covered by tuft of long seate. Hypopygium with posterior margin concave. Genitalia: paramere with apex oblique and slightly concave; aedeagal ventral ramus as long as aedeagal dorsal body, laminar, surface vertical, parallel; apex with two large apical teeth; aedeagal dorsal body with three pairs of apical lobes; outer pair with apex converging, inner margin serrated, ventral margin with three long and narrow teeth; inner pairs stout, membranous and setose; dorsal pair short and the ventral as long as aedeagal dorsal body; basal process of aedeagal dorsal body with pair of short teeth outward and median long tooth. Apodeme extending beyond genital ring.</p><p>Remarks. This species is known only to Panama (Azevedo 2001) and now more specimens are added. In this series the specimens are quite similar to those previously studied, without any variation that worths to mention.</p><p>Material examined. Holotype, ♂, PANAMA, Canal Zone: Barro Colorado Island, 3–13.VI.1983, B. Gill col. (PMAE) ; PANAMA, Veraguas Pr[ovincia]: 1♂, P[arque] N[acional] Coiba, Isla Coiba, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Cerro la Equis</a>, 120 m [07º28'09"N, 81º45'24"W], 29 jul–4 ag 1998, Malaise, R. Cambra, A. Santos [col.] (MIUP) ; 1♂, Isla Coiba, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Estación</a> biológica, [07º28'09"N, 81º45'24"W], 4–8 Agosto 1998, [trampas] Amarillas, Col. L. D., R. C., A. A., A. S. (MIUP) ; Darién Pr[ovincia]: 60♂, P[arque] N[acional] Darién, Pirre, Est [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-87.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -87.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 87°45'W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (59♂) and 24–26 marzo, 450m, col. A. Santos (1♂) (MIUP) .</p><p>Distribution (Fig. 241). Panama (Veraguas, Panama and Darién).</p></div>	https://treatment.plazi.org/id/3B3687C07C6B9B3CFF67F3509CCDFB45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6B9B3CFF67F6D29D8AF8BD.text	3B3687C07C6B9B3CFF67F6D29D8AF8BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus strepsus Azevedo 2001	<div><p>Dissomphalus strepsus Azevedo, 2001</p><p>(Figs 119–121)</p><p>Dissomphalus strepsus Azevedo, 2001: 175, 200–201 (♂ holotype from Costa Rica EMUS, figs 175–179).</p><p>Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with pair of antero-lateral depressions, ovoid, deep, large with small pit covered by tuft of long setae. Hypopygium with posterior margin somewhat concave. Genitalia: paramere with apex oblique and concave; aedeagal dorsal ramus longer than aedeagal dorsal body, parallel, apical third directed downward and apex forward, sharpened and abruptly angled; inner margin with series of small teeth; aedeagal dorsal body with three pairs of apical lobes; outer pair with two rami, dorsal ramus parallel with apex narrow and angled downward, ventral ramus arched inward and dorsal ramus with apices nearly touching each other; inner pairs stout, membranose with rounded apex. Apodeme not extending beyond genital ring.</p><p>Remarks. This species is known to Costa Rica and Panama (Azevedo 2001). Now one specimen is added. In this series the specimens are quite similar to those previously studied, without any variation that worths to mention.</p><p>Material examined. Holotype, ♂, COSTA RICA, Guanacaste: 3km, SE, Rio Naranjo, 13–31.VII.1993, F.D. Parker (EMUS) ; PANAMA, Darién: 1♂ Cana Serrania de Pirre, 1250m, 4–7.vi.1996, J. Ashe, R. Brooks (UFES) .</p><p>Distribution (Fig. 242). Costa Rica and Panama.</p></div>	https://treatment.plazi.org/id/3B3687C07C6B9B3CFF67F6D29D8AF8BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6A9B3DFF67F3E59D8AFAF8.text	3B3687C07C6A9B3DFF67F3E59D8AFAF8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus angulatus Azevedo 1999	<div><p>Dissomphalus angulatus Azevedo, 1999</p><p>(Fig 122–124)</p><p>Dissomphalus angulatus Azevedo, 1999: 331 –332 (♂, holotype from Costa Rica, CNCI, figs 49, 173–174).</p><p>Diagnosis. Mandible tridentate. Clypeus with median lobe ill-defined. Tergal process median with deep and large elliptical depression, occupying slightly more than median third of the width of the tergite with two rounded median tuft of setae. Hypopygium with median concavity. Genitalia: paramere with apex truncare and oblique with four projections low and rounded; aedeagal ventral ramus shorter than aedeagal dorsal body, flattened, narrow, wholly arched; median part outward and apex abruptly angulate outward; aedeagal dorsal body not so wide laterally, narrowing to apex, with sufaces vertical and apex slightly divergent, ventral margin serrated. Apodeme not extending beyond genital ring.</p><p>Remarks. This species was known from Costa Rica (Azevedo 1999a) and is recorded for the first time from Panama. Some variations are worth to mention: metasoma light castaneous and dark castaneous; median clypeal tooth short; anterior margin of pronotal disc coarse; depression of tergal process shallow, more distant each other with few and shorter tuft of setae, some specimens have dense and longer tuft of setae.</p><p>Material examined. Holotype, ♂, COSTA RICA, San Vito de Coto Brus: Las Cruces, 1200m, 17.vii– 12.ix.1982, Malaise trap, B. Gill col. (CNCI) . PANAMA, Chiriqui: 4♂, Finca La Suiza near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 25–6.VI.2000, (3♂) and 2–6.VI.2000 (1♂), F[light] I[nterception] T[rap], H. &amp; A. Howden [col.] (CNCI) ; 2♂, P[arque] I[nternacional] L[a] A[mistad], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.73333&amp;materialsCitation.latitude=8.9" title="Search Plazi for locations around (long -82.73333/lat 8.9)">Jurutungo</a>, 1800 m, [08°54'N, 82°44'W], 30 oct. 5 nov 1994, Tr [ampa] Malaise, (1♂) and 1600 m, 30 oct 1994, T[rampa] Amarilla (1♂), A. Rodriguez [col.] (MIUP) ; 5♂, 2 km W <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.583336&amp;materialsCitation.latitude=8.85" title="Search Plazi for locations around (long -82.583336/lat 8.85)">Cerro Punta</a>, 1700 m, [08°51'N, 82°35'W], 19.V–8.VI.1977, Peck &amp; Howden [col.] (CNCI) ; 1♂, 15 km NW Hato del <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.63333&amp;materialsCitation.latitude=8.766666" title="Search Plazi for locations around (long -82.63333/lat 8.766666)">Volcano</a>, 1200 m, [08°46'N, 82°38'W], 24–31.V.1997, Peck &amp; Howden [col.] (CNCI) .</p><p>Distribution (Fig. 226). Costa Rica and Panama.</p></div>	https://treatment.plazi.org/id/3B3687C07C6A9B3DFF67F3E59D8AFAF8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6A9B3DFF67F2A89BC9FE59.text	3B3687C07C6A9B3DFF67F2A89BC9FE59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus bifoveatus	<div><p>bifoveatus species-group</p><p>Diagnosis. Male. Metasoma with median large depression; pair of tubercles. Aedeagal ventral ramus narrow, much arched outward medially, apex abruptly angulate, straight and diverging.</p><p>Remarks. It has six species: D. bifoveatus Kieffer, D. microculus Azevedo, D. angulatus, D. asitius sp. nov., D. ibocius sp. nov. and D. tisinus sp. nov. .</p><p>Distribution. Mexico (Durango, Morelos, Chiapas), Nicaragua, Costa Rica and Panama.</p></div>	https://treatment.plazi.org/id/3B3687C07C6A9B3DFF67F2A89BC9FE59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6A9B3EFF67F7439C64FEBB.text	3B3687C07C6A9B3EFF67F7439C64FEBB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus asitius Azevedo 2017	<div><p>Dissomphalus asitius Brito &amp; Azevedo, sp. nov.</p><p>(Figs 6, 37, 68, 125–126)</p><p>Description. Male. Body length 2.03 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 6). Mandible with two distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina low in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 37). Tergal process with median, shallow, subcircular and large depression, with small, low and dorsad tubercle placed on center of depression, pit very small, with short and dense tuft of setae. Posterior hypopygeal margin strongly concave (Fig. 68). Genitalia (Figs 125–126): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface subvertical; inner margin sinuous; outer margin sinuous; apex long, toothed, angled outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body two pairs of apical lobes; outer lobe long, horizontal and wide, with apical margin acute and dorsad, inner pair stout, membranous and setose. Apodeme not extending beyond genital ring.</p><p>Remarks. This species is easily recognized by having the dorsal margin of the paramere almost straight, the apex of the outer lobe of the aedeagal dorsal body abruptly acute, the aedeagal ventral ramus with a large fold and the genital ring bifurcated in the distal region in ventral view. This combination of characters is unusual for bifoveatus species-group.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Material</a> examined. Holotype, ♂, PANAMA, Bocas del Toro: Wekso, Teribe, P [arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ampa] Malaise, A. Santos[col.] (MIUP) . Paratypes: 5♂, same data as holotype .</p><p>Distribution (Fig. 228). Panama (Bocas del Toro).</p></div>	https://treatment.plazi.org/id/3B3687C07C6A9B3EFF67F7439C64FEBB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C689B3FFF67F2A89C4DFBD0.text	3B3687C07C689B3FFF67F2A89C4DFBD0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus ibocius Azevedo 2017	<div><p>Dissomphalus ibocius Brito &amp; Azevedo sp. nov.</p><p>(Figs 7, 38, 69, 127–128)</p><p>Description. Male. Body length 2.24 mm. Color: body dark castaneous. Head (Fig. 7). Mandible with two distal teeth. Median clypeal lobe ill-defined, with one angulate very short tooth; median clypeal carina low in profile, incomplete apically, angled in profile. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc coriaceous; anterior margin carinate. Metasoma (Fig. 38). Tergal process with shallow, subcircular and large depression, with very small and very low dorsad tubercle placed on inner area of depression, pit very small with short tuft of setae. Posterior hypopygeal margin weakly concave (Fig. 69). Genitalia (Figs 127–128): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface subvertical; inner margin sinuous; outer margin sinuous; apex long, crenulate, curved outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and wide, with apical margin crenulate and laterad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring.</p><p>Variations. Body length between 1.79 (min.) mm and 2.60 (max.) mm.</p><p>Remarks. This species is similar to D. asitius sp. nov. by having the aedeagal ventral ramus with toothed apex and the paramere with crenulate apex. However, it species has the apical lobes of the aedeagal dorsal body long, with laterad apex and the paramere with the dorsal margin sinuous, whereas D. asitius sp. nov. has the apical lobes of the aedeagal dorsal body shorter with dorsad apex and the paramere with the dorsal margin almost straight.</p><p>Material examined. Holotype, ♂, Coclé Pr [ovincia]: El Copé, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">Par</a> [que] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">Nac</a> [ional <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">General de División</a>] Omar Torrijos H [errera], 570–670 m.s.n.m., [08º41'56"N, 80º39'17"W], 26–30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) . Paratypes: 2♂, same data as holotype; Panama Pr [ovincia]: 2♂, P[arque] Nac [ional] Altos de Campana, [08º43'15"W, 80º01'43"W], 10–14 ago 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) .</p><p>Distribution (Fig. 235). Panama (Coclé and Panama).</p></div>	https://treatment.plazi.org/id/3B3687C07C689B3FFF67F2A89C4DFBD0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C689B38FF67F65D9DD0FF6F.text	3B3687C07C689B38FF67F65D9DD0FF6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus tisinus Azevedo 2017	<div><p>Dissomphalus tisinus Brito &amp; Azevedo, sp. nov.</p><p>(Figs 8, 39, 70, 129–130)</p><p>Description. Male. Body length 3.52 mm. Color: body dark castaneous. Head (Fig. 8). Mandible with three distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina high in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 39). Tergal process with median deep, subcircular and large depression, without tubercle, pit absent, with short and dense tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 70). Genitalia (Figs 129–130): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface subvertical; inner margin slightly sinuous; outer margin sinuous; apex long, simple, angled outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, subhorizontal and narrow, with apical margin abruptly acute and ventrad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring.</p><p>Variations. Metasoma castaneous; median clypeal tooth longer.</p><p>Remarks. This species is easily recognized by having the apical lobes of the aedeagal dorsal body short and triangular, the median portion of the aedeagal ventral ramus sinuous and narrow. This species is similiar to D. asitius sp. nov. by having the paramere with the dorsal margin almost straight and the apex of the aedeagal dorsal body curved outward. However, this species has the apical lobes of the aedeagal dorsal body short, narrowing apicad and the apical portion of the aedeagal ventral ramus slender and smooth, whereas D. asitius sp. nov. has the apical lobes of the aedeagal dorsal body longer, abruptly acute and the apical portion of the aedeagal ventral ramus toothed.</p><p>Material examined. Holotype, ♂, PANAMA, Chiriqui: Finca La Suiza near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 2– 6.VI.2000, F[light] I[nterception] T[rap], H. &amp; A. Howden [col.] (CNCI) . Paratypes: 1♂, same data as holotype; 2♂, same data as holotype except 25–6.VI.2000 (CNCI); 1♂, 15 km NW Hato del <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.63333&amp;materialsCitation.latitude=8.766666" title="Search Plazi for locations around (long -82.63333/lat 8.766666)">Volcano</a>, 1200 m, [08°46'N, 82°38'W], 24–31.V.1997, Peck &amp; Howden [col.] (CNCI).</p><p>Distribution (Fig. 243). Panama (Chiriqui).</p></div>	https://treatment.plazi.org/id/3B3687C07C689B38FF67F65D9DD0FF6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6F9B38FF67F0799DF1FAE2.text	3B3687C07C6F9B38FF67F0799DF1FAE2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus brasiliensis Kieffer 1910	<div><p>Dissomphalus brasiliensis Kieffer, 1910</p><p>Dissomphalus brasiliensis Kieffer, 1910b: 295 (♂, holotype from Pará, CASC); Redighieri &amp; Azevedo, 2006: 319 –320. Dissomphalus bispinulatus Evans, 1969: 13, 19–20. Synonimized by Redighieri &amp; Azevedo, 2006.</p><p>Dissomphalus hastatus Evans, 1979: 276 –277, 281–283. Synonymized by Azevedo, 1999b.</p><p>Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with pair of latero-anterior ovoid and shallow depressions with tubercle and tuft of setae. Hypopygium with posterior margin slightly concave. Genitalia: paramere arched inward, apex produced and narrow; aedeagal ventral longer than aedeagal dorsal body, wide, tubular, parallel, apex rounded; aedeagal dorsal body with three pairs of apical lobes, inner pair elongate with rounded apex; median pair membranous, laminar, entirely, wide; outer pair with apical projection inclined downward with strongly serrated margin. Apodeme not extending beyond genital ring.</p><p>Remarks. This species is known to Panama, Venezuela, British Guiana, Suriname, Ecuador Peru, Brazil (Rondônia, São Paulo) and Argentina (Salta). Here two specimens are added. Some variations are worth to mention: median clypeal lobe ill-defined with one rounded tooth; median clypeal carina high in profile; vertex crest straight; pronotal disc with margin ecarinate; paramere longer than aedeagal dorsal body; basiparamere shorter than paramere; aedeagal dorsal ramus with inner margin concave in median portion.</p><p>Material examined. Holotype ♂, BRAZIL, Pará: Baker col. (CASC, #17054) . PANAMA, Veraguas Pr [ovincia]: 2#, P[arque] N[acional] Coiba, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Isla Coiba</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Cerro La Equis</a>, 90 m, [07º28'09"N, 81º45'24"W], 30– nov.–4dic.1998, Malaise, R. Cambra, S. Bermudez [col.] (MIUP) .</p><p>Distribution (Fig. 228). Panama, Venezuela, Guiana, Suriname, Ecuador, Peru, Brazil (Pará, Rondônia, Acre, Espírito Santo and São Paulo) and Argentina.</p></div>	https://treatment.plazi.org/id/3B3687C07C6F9B38FF67F0799DF1FAE2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6F9B38FF67F33B9E8BFE2C.text	3B3687C07C6F9B38FF67F33B9E8BFE2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus brasiliensis Kieffer 1910	<div><p>brasiliensis species-group</p><p>Diagnosis. Male. Mandible tridentate. Clypeus broad. Aedeagal ventral ramus tubular. Aedeagal dorsal body small.</p><p>Remarks. It has two species: D. brasiliensis and D. tubulatus Redighieri &amp; Azevedo. Distribution. Venezuela, Guiana, Suriname, Ecuador, Peru, Brazil (Rondônia, Acre, Paraíba, Espírito Santo and São Paulo) and Argentina.</p></div>	https://treatment.plazi.org/id/3B3687C07C6F9B38FF67F33B9E8BFE2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6F9B38FF67F7B09D4AF8EC.text	3B3687C07C6F9B38FF67F7B09D4AF8EC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus conicus Azevedo 2003	<div><p>conicus species-group</p><p>Diagnosis. Male. Median clypeal lobe trapezoidal, except D. filus Azevedo and D. ibrastus sp. nov. . Metasoma with pair lateral tergal process, tubercle, pit on top and tuft of setae.</p><p>Remarks. It has 19 species: D. cervoides, D. conicus Azevedo, D. curviventris Azevedo, D. filus, D. manus Azevedo, D. truncatus Azevedo, D. umbilicus Azevedo, D. h-ramus Redighieri &amp; Azevedo, D. laminaris Redighieri &amp; Azevedo, D. verrucosus Redighieri &amp; Azevedo, D. botocudus Colombo &amp; Azevedo, D. ardonius sp. nov., D. arteus sp. nov., D. ettus sp. nov., D. ibrastus sp. nov., D. racarus sp. nov., D. teraus sp. nov. and D. urazius sp. nov. .</p><p>Distribution. Panama, Trinidad &amp; Tobago, Ecuador, Peru, Brazil (Amazonas, Pará, Rondônia, Ceará, Pernambuco, Mato Grosso, Goiás, Distrito Federal, Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo, Paraná and Santa Catarina), Bolivia.</p></div>	https://treatment.plazi.org/id/3B3687C07C6F9B38FF67F7B09D4AF8EC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6E9B39FF67F2A89EC2FBF1.text	3B3687C07C6E9B39FF67F2A89EC2FBF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus cervoides Azevedo 2003	<div><p>Dissomphalus cervoides Azevedo, 2003</p><p>(Figs 131–132)</p><p>Dissomphalus cervoides Azevedo, 2003: 31 –32 (♂, holotype from Pará, MPEG, figs 33–36).</p><p>Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with lateral pair of small and very shallow depressions and pit with dense tuft of setae. Hypopygium with posterior margin straight. Genitalia: paramere longer than basiparamere, dorsal margin straight; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; apical half translucent and thin with top-rounded expansions, apex constricted below; aedeagal dorsal body with three pairs of apical lobes; dorsal pair thick, surface vertical and parallel; dorsal margin angled; ventral pair membranous, stout, rounded, shorter than dorsal one; median pair less stout, less membranous and shorter than median one; basal process with pair of lateral rounded expansions upward. Apodeme extending beyond genital ring.</p><p>Remarks. This species is known to Panama, Trinidad &amp; Tobago, Ecuador, Peru, Brazil (Amazonas, Pará, Rondônia, Mato Grosso, Goiás) and Bolivia (Azevedo 2003). Here more specimens are added. Some variations in this series worth to mention: mesosoma castaneous, metasoma light castaneous; median clypeal tooth angulate; median clypeal carina straight; pronotal disc ecarinate; tergal process without depression.</p><p>Material examined. Holotype ♂, BRAZIL, Pará: Tucuruí, Rio Tocantins, V.1986, Interception Trap, N. Degallier col. (MPEG) ; PANAMA, Colón: 8♂, 14 km N J[un]ct[ion] Escobal r[oa]d and Piña r[oa]d on Piña r[oa]d, ± 20 m, [09°20'N', 79°54'W], 2–3.VI.1995, F[light] I[nterception] T[rap], J. Ashe &amp; R. Brooks [col.] (CNCI); 4♂, S[an]ta Rita [Arriba], Macizo, Parador Sierra Llorona, [09°19'57.1"N, 79°45'1.6"W], 22 dic. 1997, T[rampa] Malaise, D. Quintero [col.] (MIUP); Darién Pr [ovincia]: 1♂, P[arque] N[acional] Darién, Pirre, Est [ación] Rancho Frio, 230 m, [08°00'N, 77°45'W], 21 mar–4 ab 2000, Malaise, Cambra, Santos, Bermudez [col.] (MIUP).</p><p>Distribution (Fig. 229). Panama, Trinidad &amp; Tobago, Ecuador, Peru, Brazil (Amazonas, Pará, Rondônia, Mato Grosso, Goiás), Bolivia.</p></div>	https://treatment.plazi.org/id/3B3687C07C6E9B39FF67F2A89EC2FBF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6E9B3AFF67F6BE9C64FF27.text	3B3687C07C6E9B3AFF67F6BE9C64FF27.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus ardonius Azevedo 2017	<div><p>Dissomphalus ardonius Brito &amp; Azevedo, sp. nov.</p><p>(Figs 9, 40, 71, 133–135)</p><p>Description. Male. Body length 3.22 mm. Color: body castaneous. Head (Fig. 9). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 40). Tergal process with lateral, shallow, circular and small depression, with large, very low and dorsad tubercle placed on center of depression, pit large, with short and sparse tuft of setae. Posterior hypopygeal margin median straight with angled corners (Fig. 71). Genitalia (Figs 133–135): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, entirely wide, cross section laminar; surface vertical; inner margin strongly sinuous; outer margin slightly sinuous; apex long, slightly crenulate, parallel; additional inner ramus long; basal stub long and narrow. Aedeagal dorsal body with two pairs of apical lobes, outer lobe long, vertical and very narrow, with apical margin acute and ventrad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring.</p><p>Variation. Metasoma light castaneous.</p><p>Remarks. This species is easily recognized by having two spines in the ventral margin of the apical lobes of the aedeagal dorsal body with apical spine shorter than distal one, the aedeagal ventral ramus vertical and wide in the median portion with wavy apex. This species is similar to D. urazius sp. nov. by having spine in the ventral margin of apical lobes of the aedeagal dorsal body and the aedeagal ventral ramus entirely wide with the inner margin sinuous. However, this species has the dorsal margin of the paramere sinuous, the apex of the basal bar not projected forward, and the apical margin of the aedeagal dorsal body wavy, whereas D. urazius sp. nov. has the dorsal margin of the paramere not sinuous, the apex of the basal bar projected forward and the apical margin of the aedeagal ventral ramus not wavy.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Material</a> examined. Holotype, ♂, PANAMA, Bocas del Toro: Wekso, Teribe, P [arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ampa] Malaise, A. Santos[col.] (MIUP) . Paratypes: 9♂, same data as holotype .</p><p>Distribution (Fig. 226). Panama (Bocas del Toro).</p></div>	https://treatment.plazi.org/id/3B3687C07C6E9B3AFF67F6BE9C64FF27	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6D9B3AFF67F3739C1CFAF0.text	3B3687C07C6D9B3AFF67F3739C1CFAF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus arteus Azevedo 2017	<div><p>Dissomphalus arteus Brito &amp; Azevedo, sp. nov.</p><p>(Figs 10, 41, 72, 136–137)</p><p>Description. Male. Body length 4.96 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 10). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, complete or nearly so, convex in profile. Frons strongly coriaceous and punctures large. Vertex crest weakly concave. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 41). Tergal process with lateral, deep, subcircular and large depression, with small, low and dorsad tubercle placed on anterior area of depression, pit small, with long and sparse tuft of setae. Posterior hypopygeal margin sinuous (Fig. 72). Genitalia (Figs 136–137): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface vertical; outer margin and inner margin slightly sinuous; apex long, simple, parallel; additional inner ramus long; basal stub short and narrow. Aedeagal dorsal body with pair of apical lobe; outer lobe long, vertical and very narrow, with apical margin crenulate and ventrad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring.</p><p>Variation. Tergal process with very shallow depression and without tubercle.</p><p>Remarks. This species is similar to the D. racarus sp. nov. by having the dorsal margin of the paramere sinuous, the apex of the basal bar projected forward and the inner margin of the aedeagal ventral ramus crenulate. However, this species has the apex of the basal bar smooth, the ventral margin of the apical lobes of the aedeagal dorsal body not excavated and the apical margin of the aedeagal ventral ramus smooth, whereas D. racarus sp. nov. has the apex of basal bar crenulate, the ventral margin of the apical lobes of the aedeagal dorsal body deeply excavated and the apical margin of the aedeagal ventral ramus with a small projection dorsad.</p><p>Material examined. Holotype ♂, PANAMA, Bocas del Toro: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Wekso</a>, Teribe, P [arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ampa] Malaise, A. Santos[col.] (MIUP) . Paratypes: 7♂, same data as holotype; Coclé Pr [ovincia]: 1♂, El Copé, Par [que] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">Nac</a> [ional <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">General de División</a>] Omar Torrijos H [errera], 570–670 m.s.n.m., [08º41'56"N, 80º39'17"W], 26–30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) .</p><p>Distribution (Fig. 229). Panama (Bocas del Toro and Coclé).</p></div>	https://treatment.plazi.org/id/3B3687C07C6D9B3AFF67F3739C1CFAF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6D9B04FF67F7BD9BAFFDE2.text	3B3687C07C6D9B04FF67F7BD9BAFFDE2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus ettus Azevedo 2017	<div><p>Dissomphalus ettus Brito &amp; Azevedo, sp. nov.</p><p>(Figs 11, 42, 73, 138–139)</p><p>Description. Male. Body length 4.0 mm. Color: body dark castaneous. Head (Fig. 11). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina low in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 42). Tergal process lateral, without depression, low and dorsad tubercle, pit small, with short and dense tuft of setae. Posterior hypopygeal margin weakly concave (Fig. 73). Genitalia (Figs 138–139): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus higher than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin excavated; outer margin sinuous; apex short, translucent, wavy, parallel, additional inner ramus short; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, subhorizontal and wide, with apical margin rounded and ventrad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Variations. Tergal process closer to each other; tubercle higher or with few setae.</p><p>Remarks. This species is easily recognized by having the paramere narrow with the dorsal margin straight, very acute apex, the basivolsella wavy and the basal bar in the median portion of the aedeagal dorsal body projected forward. This species is similar to the D. cervoides by having the apex of the aedeagal ventral ramus laminar, translucent with two expansions. However, this species has the paramere narrow, the aedeagal ventral ramus wide with the inner margin of the apex smooth and the basivolsella wavy, whereas D. cervoides has the paramere not so straight, the aedeagal ventral ramus narrower with the inner margin of the apex serrated and the basivolsella not wavy.</p><p>Material examined. Holotype, ♂, PANAMA, Darién Pr [ovincia]: P[arque] N[acional] Darién, Pirre, Est[ación] Rancho Frio, 80 m, [08°00'N, 77°45'W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.]</p><p>(MIUP). Paratypes: Bocas del Toro: 1♂, Wekso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Teribe</a>, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP) ; Chiriqui: 3♂, Finca La Suiza near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 25–6.VI.2000, F[light] I[nterception] T[rap], H. &amp; A. Howden [col.] (CNCI) ; Veraguas Pr [ovincia]: 2♂, P[arque] N[acional] Coiba, Isla Coiba, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Cerro la Equis</a>, 0 m, [07º28'09"N, 81º45'24"W], 30.nov–4.dic.1998, Malaise, R. Cambra, S. Bermudez [col.] (1♂) and 6–11 mar 1998, Tr [ampa] R. Cambra, A. Santos (1♂) (MIUP) ; Panama Pr [ovincia]: 1♂, Cerro Azul, Las Nubes, [09º10'N, 79º25'W], 9–15– oct.1999, Malaise, R. Cambra, A. Santos [col.] (MIUP); 1♂, C[anal] Z[one], B[arro] Colorado Is[land], [09°09'N, 79°51'W], 15–31.VII.1994, F[light] I[nterception] T[rap], D. Banks [col.] (CNCI); 5♂ same data as holotype; 1♂, same province of holotype, Cruce de Mono: Est [ación] I[nstituto] N[acional de] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Re</a> [cusos] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Na</a> [turales] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Re</a> [novables], [07°51'26.3"N, 77°28'12.7"W], 6.II–4.III.1993, t[rap] Malaise, R. Cambra, J. Coronado [col.] (MIUP).</p><p>Distribution (Fig. 233). Panama (Bocas del Toro, Chiriqui, Veraguas, Panama and Darién).</p></div>	https://treatment.plazi.org/id/3B3687C07C6D9B04FF67F7BD9BAFFDE2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C539B04FF67F0B09DD0F990.text	3B3687C07C539B04FF67F0B09DD0F990.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus ibrastus Azevedo 2017	<div><p>Dissomphalus ibrastus Brito &amp; Azevedo, sp. nov.</p><p>(Figs 12, 43, 74, 140–141)</p><p>Description. Male. Body length 3.10 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 12). Mandible with three distal teeth. Median clypeal lobe ill- defined, with one angulate very large tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly convex. Pronotal disc weakly coriaceous; anterior margin carinate. Metasoma (Fig. 43). Tergal process with lateral, deep, subcircular and large depression, with small, low and dorsad tubercle placed on center of depression, pit small, with short and sparse tuft of setae. Posterior hypopygeal margin weakly concave (Fig. 74). Genitalia (Figs 140–141): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin slightly sinuous; apex long, simple, parallel; additional inner ramus absent; basal stub short and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and wide, with apical margin rounded and ventrad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Variation. Tergal process with very shallow depression.</p><p>Remarks. This species is easily recognized by having the median clypeal tooth very long, the base of the outer margin of the apical lobes of the aedeagal dorsal body with two small rounded and dorsad projections. This species is similar to D. longiclypeus Azevedo by having the median clypeal tooth long. However, this species has the tergal process lateral, the apical lobes of the aedeagal dorsal body long and the aedeagal ventral ramus with the inner margin of the apex excavated, whereas D. longiclypeus has the tergal process median, the aedeagal dorsal body short and the aedeagal ventral ramus not excavated.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.73333&amp;materialsCitation.latitude=8.9" title="Search Plazi for locations around (long -82.73333/lat 8.9)">Material</a> examined. Holotype, ♂, PANAMA, Chiriqui Pr [ovincia]: P[arque] I[nternacional] L[a] A[mistad], Jurutungo, 1800 m, [08°54'N, 82°44'W], 30 oct–5 nov.1994, Tr [ampas] Amarillas, A. Rodriguez [col.] (MIUP) . Paratypes: 2♂, same data as holotype; 1♂, 2 km W <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.583336&amp;materialsCitation.latitude=8.85" title="Search Plazi for locations around (long -82.583336/lat 8.85)">Cerro Punta</a>, 1700 m, [08°51'N, 82°35'W], 19.V–8.VI.1977, Peck &amp; Howden [col.] (CNCI) .</p><p>Distribution (Fig. 235). Panama (Chiriquí).</p></div>	https://treatment.plazi.org/id/3B3687C07C539B04FF67F0B09DD0F990	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C539B05FF67F4A29CE1FC82.text	3B3687C07C539B05FF67F4A29CE1FC82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus racarus Azevedo 2017	<div><p>Dissomphalus racarus Brito &amp; Azevedo, sp. nov.</p><p>(Figs 13, 44, 75, 142–144)</p><p>Description. Male. Body length 4.78 mm. Color: body castaneous. Head (Fig. 13). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin coarse. Metasoma (Fig. 44). Tergal process lateral with shallow, subcircular and large depression, with small, low and dorsad tubercle placed on anterior area of depression, pit large, with short and sparse tuft of setae. Posterior hypopygeal margin sinuous (Fig. 75). Genitalia (Figs 142–144): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, entirely wide with small projection apicad; cross section laminar; surface vertical; inner margin sinuous, outer margin sinuous; apex short, simple, parallel; additional inner ramus long; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe, long, vertical and very narrow, with apical margin crenulate and posterad; inner pair, stout, membranous and setose. Apodeme not extending beyond genital ring.</p><p>Variations. Body length 3.31 (min.) mm and 6.06 mm (max.). Depression of tergal process deep or very shallow.</p><p>Remarks. This species is easily recognized by having the ventral margin of the apical lobes of the aedeagal dorsal body deeply excavated, the apex of the basal with apex projected forward, the aedeagal ventral ramus vertical and the apex with a small projection dorsad. This species is similar to D. arteus sp. nov. by having the dorsal margin of the paramere sinuous, the apex of the basal bar projected forward and the inner margin of the aedeagal ventral ramus crenulate. However, this species has the apex of the basal bar crenulate, the ventral margin of the aedeagal dorsal body deeply excavated and the apical margin of the aedeagal dorsal body with small dorsad projection, whereas D. arteus sp. nov. has the apex of the basal bar smooth, the ventral margin of the apical lobes of the aedeagal dorsal body not excavated and the aedeagal ventral ramus without projection.</p><p>Material examined. Holotype, ♂, PANAMA, Darién Pr [ovincia]: P[arque] N[acional] Darién, Pirre, Est[ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 77°45'W], 16 nov [20] 00–17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP) . Paratypes: Bocas del Toro: 1♂, Wekso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Teribe</a>, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ap] Malaise, A. Santos [col.] (MIUP) ; 4♂, same data as holotype; 1♂, same data as holotype except 450m, 24–26 marzo, col. A. Santos (MIUP).</p><p>Distribution (Fig. 243). Panama (Bocas del Toro and Darién).</p></div>	https://treatment.plazi.org/id/3B3687C07C539B05FF67F4A29CE1FC82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C529B06FF67F1989B7BFE73.text	3B3687C07C529B06FF67F1989B7BFE73.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus teraus Azevedo 2017	<div><p>Dissomphalus teraus Brito &amp; Azevedo, sp. nov.</p><p>(Figs 14, 45, 76, 145–153)</p><p>Description. Male. Body length 4.33 mm. Color: head and mesosoma dark castaneous; metasoma light castaneous. Head (Fig. 14). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly convex. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 45). Tergal process with lateral, shallow, circular and large depression, with small, low and laterad tubercle placed on inner area of depression, pit small, with short tuft of setae. Posterior hypopygeal margin sinuous (Fig. 76). Genitalia (Figs 145–153): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus higher than dorsal body, wide, progressively narrowing apicad; cross section laminar; ramous bifid; surface horizontal; inner margin sinuous; outer margin sinuous; apex long, simple, curved outward; additional inner ramus long; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and narrow, with apical margin rounded and mesad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Variations. Head dark castaneous; metassoma castaneous; frons coriaceous; median clypeal lobe subtrapezoidal; depression of tergal process very shallow or without depression, with very low, very small and without tubercle; apical lobe narrow in sickle-shaped with teeth in inner margin and apical lobes with acute apex and inner margin smooth.</p><p>This species has two different aedeagal ventral ramus and apical lobes of the aedeagal dorsal body. In one kind, the aedeagal ventral ramus can be lower than the aedeagal dorsal body; the apical margin entirely wide with rounded and dorsad apex; in the other kind the aedeagal ventral ramus can have the apex narrow and two small and laterad projections.</p><p>Remarks. This species is similar to D. piscicercus by having the paramere entirely wide and the basiparamere very wide. However, this species has the tergal process with tubercle, small and shallow depression, the aedeagal ventral ramus with simple apex and the apex of the basal bar straight with two laterad teeth, whereas D. piscicercus has the tergal process with ovoid, deep, large depressions nearly touching the posterior margin of the tergite I, with pit, tuft of long setae, the aedeagal ventral ramus with two large apical teeth and the basal bar without two laterad teeth.</p><p>Material examined. Holotype, ♂, PANAMA, Chi [riqui]: Hartmans Finca, 28.VI–3.VII.1981 (PMAE) . Paratypes: Bocas del Toro: 7 ♂, Wekso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Teribe</a>, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP) ; 5♂, same data as holotype; 6♂ same data as holotype except 3–VII1981, B. Gill [col.] (PMAE); 1♂, same province of holotype, Finca La Suiza near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 2–6.VI.2000, F[light] I[nterception] T[rap], H. &amp; A. Howden [col.] (CNCI) ; 1♂, same province of holotype, 5,6 km N <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.42166&amp;materialsCitation.latitude=8.823055" title="Search Plazi for locations around (long -82.42166/lat 8.823055)">Boquete</a>, La Culebra trail, [08°49'23"N, 82°25'18"W], F[light] I[nterception] T[rap], J. Ashe &amp; R. Brooks [col.] (CNCI) ; 2♂, same province of holotype, Finca La Suiza near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 25–6.VI.2000, F[light] I[nterception] T[rap], H. &amp; A. Howden [col.] (CNCI) ; [Los Santos]: 2♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.416664&amp;materialsCitation.latitude=7.5166664" title="Search Plazi for locations around (long -80.416664/lat 7.5166664)">Ojo de Agua</a>, [07°31'N, 80°25'W], 30.VI–14.VII.1982, B. Gill [col.] (PMAE) .</p><p>Distribution (Fig. 244). Panama (Bocas del Toro, Chiriqui and Los Santos).</p></div>	https://treatment.plazi.org/id/3B3687C07C529B06FF67F1989B7BFE73	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C519B06FF67F4329AF8F82A.text	3B3687C07C519B06FF67F4329AF8F82A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus coronatus , Alencar & Azevedo 2006	<div><p>coronatus species-group</p><p>Diagnosis. Male. Mandible tridentate, uppermost tooth usually inconspicuous. Median clypeal lobe trapezoidal except D. afestus sp. nov. . Metasoma with sublateral tergal process, wich have a pair of circular depressions with dense tuft of convergent setae directed backward, antero-lateral margin of depression with setae, tubercle absent. Genitalia with inner apical surface of paramere generally with one setae distinctly longer than others. Aedeagal ventral ramus shorter than dorsal body with at least half part inclined and tending to be convergente mesad of genitalia. Aedeagal dorsal body wide with crown-like structure.</p><p>Remarks. It has fourteen species: D. aculetaus Alencar &amp; Azevedo, D. auritus Alencar &amp; Azevedo, D. coronatus, Alencar &amp; Azevedo, D. culteratus Alencar &amp; Azevedo, D. declinatus Alencar &amp; Azevedo, D. decussatus Alencar &amp; Azevedo, D. flexuosus Alencar &amp; Azevedo, D. galeatus, Alencar &amp; Azevedo, D. hirtus, Alencar &amp; Azevedo, D. pronus, Alencar &amp; Azevedo, D. ramosus Alencar &amp; Azevedo, D. sinatus Alencar &amp; Azevedo, D. uber Alencar &amp; Azevedo and D. afestus sp. nov. .</p><p>Distribution. Dominica, Costa Rica, Panama, Venezuela, Trinidad &amp; Tobago, Brazil (Paraná), Ecuador and Peru.</p></div>	https://treatment.plazi.org/id/3B3687C07C519B06FF67F4329AF8F82A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C519B06FF67F3C09C56FA60.text	3B3687C07C519B06FF67F3C09C56FA60.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus urazius Azevedo 2017	<div><p>Dissomphalus urazius Brito &amp; Azevedo, sp. nov.</p><p>(Figs 15, 46, 77, 154–156)</p><p>Description. Male. Body length 3.70 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 15). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc coriaceous; anterior margin ecarinate. Metasoma (Fig. 46). Tergal process with lateral, shallow, circular and small depression, with small, very low and dorsad tubercle placed on inner area of depression, pit small, with short and sparse tuft of setae. Posterior hypopygeal margin weakly concave (Fig. 77). Genitalia (Figs 154–156). Paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, entirely wide; cross section laminar; surface vertical; inner margin slightly sinuous with small projection in apical portion; outer margin slightly sinuous; apex long, simple, parallel; additional inner ramus long; basal stub short and narrow. Aedeagal dorsal body with pair of apical lobe; outer lobe long, subvertical and narrow, with apical margin rounded and ventrad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring.</p><p>Variations. Body length 3.07 mm (min.) 4.45 mm (max). Tooth of median clypeal lobe rounded; anterior margin of pronotal disc coarse; depression of tergal process very shallow and pit of tubercle large.</p><p>Remarks. This species is similar to D. ardonius sp. nov. by having spine in the ventral margin of the apical lobes of the aedeagal dorsal body and the ventral ramus entirely wide with the inner margin sinuous. However, this species has the dorsal margin of the paramere straight, the apex of the basal bar projected forward and the apical margin of the aedeagal ventral ramus smooth, whereas D. ardonius sp. nov. has the dorsal margin of the paramere sinuous, the apex of the basal bar not projected forward and the apical margin of the aedeagal ventral ramus wavy.</p><p>Material examined. Holotype, ♂, PANAMA, Panama Pr [ovincia]: P[arque] Nac[ional] Altos de Campana, [08º43'15"N, 80º01'43"W], 10–14 agos 1999, Malaise, P. González, A. Santos [col.] (MIUP). Paratypes: 1♂ same data as holotype; Darién: 4♂, Serrania de Pierre, Cana, 1380 m, [07°51'N, 77°37'W], 4–7.VI.1986, F[light] I[nterception] T[rap], [J.] Ashe &amp; [R.] Brooks [col.] (2♂) (CNCI) and 1250m, 4–7.vi.1996, (2♂) (UFES).</p><p>Distribution (Fig. 246). Panama (Panama and Darién).</p></div>	https://treatment.plazi.org/id/3B3687C07C519B06FF67F3C09C56FA60	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C509B00FF67F5659DC1FC5F.text	3B3687C07C509B00FF67F5659DC1FC5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus afestus Azevedo 2017	<div><p>Dissomphalus afestus Brito &amp; Azevedo sp. nov.</p><p>(Figs 16, 47, 78, 157–158)</p><p>Description. Male. Body length 2.70 mm. Color: head and mesosoma dark castaneous; metasoma castaneous.</p><p>Head (Fig. 16). Mandible with three distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina low in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc coriaceous; anterior margin carinate. Metasoma (Fig. 47). Tergal process lateral, without depression, without tubercle, with short tuft of setae. Posterior hypopygeal margin straight (Fig. 78). Genitalia (Figs 157–158): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface subhorizontal; inner margin concave; outer margin sinuous; apex long, simple, slightly curved inward; additional inner ramus absent; basal stub long and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe horizontal and wide, with apical margin angled and laterad; inner pair stout, membranous and setose; crown-process very long. Apodeme not extending beyond genital ring.</p><p>Variations. Pronotal disc coarse; tergal process with very short tuft of setae, without depression.</p><p>Remarks. This species is easily recognized by having the genitalia wide, the apical lobes of the aedeagal dorsal body very long and the inner lobe of the aedeagal dorsal body very big, extending through the outer lobes of the aedeagal dorsal body. This species is similar to D. culteratus by having apical lobes of aedeagal dorsal body long with apex laterad and paramere longer than basiparamere. However, this species has the aedeagal ventral ramus with inner margin smooth, basal half wide and apical half narrow with apex mesad, whereas D. culteratus has the aedeagal ventral ramus narrowing at apex, inner margin microserrated and apex parallel.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Material</a> examined. Holotype ♂, PANAMA, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Darién Pr</a> [ovincia]: P[arque] N[acional] Darién, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Pirre</a>, Est [ación] Rancho Frio, 1000 m, [08°00'N, 77°45'W], 21 mar–4 ab 2000, Malaise, Cambra, Santos, Bermudez [col.] (MIUP) . Paratypes: 4 ♂, same data as holotype .</p><p>Distribution (Fig. 225). Panama (Darién).</p></div>	https://treatment.plazi.org/id/3B3687C07C509B00FF67F5659DC1FC5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C579B00FF67F1EB9A5EF969.text	3B3687C07C579B00FF67F1EB9A5EF969.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus coronatus , Alencar & Azevedo 2006	<div><p>Dissomphalus coronatus Alencar &amp; Azevedo, 2006</p><p>Dissomphalus coronatus Alencar &amp; Azevedo, 2006: 7 –9 (♂, holotype from Panama, MIUP, figs 7–9).</p><p>Diagnosis. Mandible tridentate. Clypeus trapezoidal. Tergal process with pair of circular, large and deep lateral depressions with dense tuft of convergent setae directed backward. Hypopygium with posterior margin with median convexity. Genitalia: paramere wide, dorsal margin straight and ventral margin concave; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; surface horizontal, angulate medially laterad; basal half narrow and convergent mesad; aedeagal dorsal body laminar, surface vertical, base wide narrowing to apex; apex rounded with sinuous and serrate crest-like projection, directed ventrally; crown-process small. Apodeme not extending beyond genital ring.</p><p>Remarks. This species is known to Costa Rica, Panama, Venezuela, Dominica, Trinidad &amp; Tobago, Ecuador (Alencar &amp; Azevedo 2006). In this series the specimens are quite similar to those previously studied, without any variation that worths to mention.</p><p>Material examined. Holotype, ♂, PANAMA, Veraguas: Coiba N [acional] P[arque], Isla Coibita [<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.70416&amp;materialsCitation.latitude=7.643611" title="Search Plazi for locations around (long -81.70416/lat 7.643611)">Rancheria</a>], [7º38’37”N 81º42’15”W], 29.VII. –4.VIII.1998, Malaise trap, R. Cambra &amp; A. Santos col. (MIUP) . PANAMA, Veraguas Pr [ovincia]: 13♂, P[arque] N[acional] Coiba, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Isla Coibita</a>, [07º28'09"N, 81º45'24"W], 27–nov–4- dic.1998, (5♂), 0m, 30–nov. –4dic.1998 (5♂) and 120m (3♂) Malaise, R. Cambra, S. Bermudez [col.] (MIUP) ; Darién Pr [ovincia]: 2♂, P[arque] N[acional] Darién, Pirre, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Est</a> [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 77°45'W], 16 nov–17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP) .</p><p>Distribution (Fig. 231). Costa Rica, Panama, Venezuela, Dominica, Trinidad &amp; Tobago and Ecuador.</p></div>	https://treatment.plazi.org/id/3B3687C07C579B00FF67F1EB9A5EF969	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C579B01FF67F5369DD0FD80.text	3B3687C07C579B01FF67F5369DD0FD80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus culteratus Alencar & Azevedo 2006	<div><p>Dissomphalus culteratus Alencar &amp; Azevedo, 2006</p><p>Dissomphalus culteratus Alencar &amp; Azevedo, 2006: 09–11 (♂, holotype from Panama, PMAE, figs 10–11).</p><p>Diagnosis. Mandible tridentate. Clypeus trapezoidal. Tergal process with pair of circular, large and deep lateral depressions with dense tuft of convergent setae directed backward. Hypopygium with posterior margin with median convexity. Genitalia: paramere with dorsal margin straight and ventral margin concave; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar, surface horizontal and arched ventrally; basal half wide; inner margin sinuous; apical margin with outer margin straight and inner margin microserrate; aedeagal dorsal body wide, narrowing to apex; apical part with concavity mesad, apex with two arms; crown-process with long lateral prolongation in middle of aedeagal dorsal body. Apodeme extending beyond genital ring.</p><p>Remarks. This species is known to Panama (Alencar &amp; Azevedo). In this series, the specimens are quite similar to those previously studied, without any variation that worths to mention.</p><p>Material examined. Holotype, ♂, PANAMA, Chiriquí: Hatman’s Farm, [8º51’N 82º46’W], 3.VII.1981, B. Gill col (PMAE); PANAMA, Chiriqui Pr [ovincia]: 2♂, P[arque] I[nternacional] L[a] A[mistad], Jurutungo, 1800 m, [08°54'N, 82°44'W], 30oci. 5nov.1994, Tr[ampa] Malaise, A. Rodriguez [col.] (MIUP); 9♂, 5.6 km N Boquete, La Culebra trail, [08°49'23"N, 82°25'18"W], (8♂) and [08° 54,7'N, 82° 47,3 W], 17–18.vi. 1996, 1875m (1♂), F[light] I[nterception] T[rap], J. Ashe &amp; R. Brooks [col.] (CNCI); [Los Santos]: 1♂, Ojo de Agua, [07°31'N, 80°25'W], 30.VI–14.VII.1982, B. Gill [col.] (PMAE).</p><p>Distribution (Fig. 229). Costa and Panama.</p></div>	https://treatment.plazi.org/id/3B3687C07C579B01FF67F5369DD0FD80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C569B01FF67F08B9DB4FB45.text	3B3687C07C569B01FF67F08B9DB4FB45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus declinatus Alencar & Azevedo 2006	<div><p>Dissomphalus declinatus Alencar &amp; Azevedo, 2006</p><p>Dissomphalus declinatus Alencar &amp; Azevedo, 2006: 11 –13, 18 (♂, holotype from Ecuador, UCDC, figs 12–14).</p><p>Diagnosis. Mandible tridentate. Clypeus trapezoidal. Tergal process with pair of circular, large and deep lateral depressions, with tuft of setae convergent directed backward. Hypopygium with posterior margin straight. Genitalia: paramere with apex straight; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; surface horizontal; basal half long; inner margin sinuous; outer margin straight; apical half narrow dorsally and converging mesad; aedeagal dorsal body laminar; surface vertical; apex wide and inclined ventrally; crown-process small; apodeme extending beyond genital ring.</p><p>Remarks. No additional material of this species was not found in this study.</p><p>Material examined. Holotype, ♂, ECUADOR, Napo: Huahua Sumaco, 44km on Hollin Loreto R[oa]d, [0º37’S 77º35’W], 15–22.XII.1989, M[alaise] T[rap], H. Real, M. and J. Wasbauer col. (UCDC);</p><p>Distribution (Fig. 233). Panama and Ecuador.</p></div>	https://treatment.plazi.org/id/3B3687C07C569B01FF67F08B9DB4FB45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C569B01FF67F6D29CADF810.text	3B3687C07C569B01FF67F6D29CADF810.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus galeatus , Alencar & Azevedo 2006	<div><p>Dissomphalus galeatus Alencar &amp; Azevedo, 2006</p><p>Dissomphalus galeatus Alencar &amp; Azevedo, 2006: 15 –16 (♂, holotype from Costa Rica, CNCI, figs 20–21).</p><p>Diagnosis. Mandible tridentate. Clypeus trapezoidal, median carina complete and strongly developed in profile. Tergal process with pair of circular, large and deep lateral depressions with dense tuft of convergent setae directed backward. Hypopygium with posterior margin straight. Genitalia: paramere wide, dorsal margin straight and ventral margin concave; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; surface horizontal, and angulate ventrally; basal half wide; inner margin with strong concavity before apex; outer margin convex above; aedeagal dorsal body laminar, surface vertical; base wide, narrowing to rounded apex; ventral margin with long projection directed to apical part of aedeagal ventral ramus; dorsal margin sinuous and serrate; crown-process small. Apodeme not extending beyond genital ring.</p><p>Remarks. This species is known to Costa Rica and Brazil (Rondônia), and now it is recorded for the first time from Panama. Some variations are worth to mention: median clypeal tooth shorter and angulate; median carina of clypeus complete apically; depression of tergal process very shallow.</p><p>Material examined. Holotype, ♂, COSTA RICA, [Puntarenas]: San Vito de Coto Brus, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.57&amp;materialsCitation.latitude=8.469999" title="Search Plazi for locations around (long -82.57/lat 8.469999)">Las Cruces</a>, 1200m, [8º28’12”N 82º34’12”W], 17.VIII–12.IX.1982, Malaise Trap, B. Gill col. (CNCI) ; PANAMA, Chiriqui: 3♂, Finca La Suiza near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 25–6.VI.2000, F[light] I[nterception] T[rap], H. &amp; A. Howden [col.] (CNCI) ; 4♂, 15 km NW Hato del <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.63333&amp;materialsCitation.latitude=8.766666" title="Search Plazi for locations around (long -82.63333/lat 8.766666)">Volcano</a>, 1200 m, [08°46'N, 82°38'W], 24–31.V.1997, Peck &amp; Howden [col.] (CNCI).</p><p>Distribution (Fig. 234). Costa Rica, Panama and Brazil (Rondônia).</p></div>	https://treatment.plazi.org/id/3B3687C07C569B01FF67F6D29CADF810	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C559B02FF67F4859A1AF859.text	3B3687C07C559B02FF67F4859A1AF859.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus curvifoveatus Azevedo 1999	<div><p>curvifoveatus species-group</p><p>Diagnosis. Male. Metasoma with tergal process median, large, deep, triangular depression with base toward; pair of very large lateral pits, arched inward medially, covered by a lot of long setae inward.</p><p>Remarks. It has a single species, D. curvifoveatus .</p><p>Distribution. Belize, Costa Rica, Panama, Venezuela, Ecuador and Brazil (Amazonas, Pará and Rondônia).</p></div>	https://treatment.plazi.org/id/3B3687C07C559B02FF67F4859A1AF859	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C559B02FF67F2A89D8AFBD6.text	3B3687C07C559B02FF67F2A89D8AFBD6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus pronus , Alencar & Azevedo 2006	<div><p>Dissomphalus pronus Alencar &amp; Azevedo, 2006</p><p>Dissomphalus pronus Alencar &amp; Azevedo, 2006: 19 –20 (♂, holotype from Costa Rica, PMAE, figs 24–27).</p><p>Diagnosis. Mandible tridentate. Clypeus trapezoidal. Tergal process with pair of circular, large and deep lateral depressions with dense tuft of convergent setae directed backward. Hypopygium with posterior margin convex. Genitalia: paramere wide; dorsal margin straight and ventral margin concave; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; surface horizontal and angulate ventrally; basal half long and wide; inner margin sinuous; apical half narrowing to apex and converging mesad; apex folded laterad; aedeagal dorsal body wide; surface vertical; apical part with concavity mesad, apex with two arms; crown-process small. Apodeme not extending beyond genital ring.</p><p>Remarks. This species is known to Costa Rica (Alencar &amp; Azevedo 2006) and now it is recorded for the first time from Panama. Some variations are worth to mention: head, mesosoma and metasoma castaneous; clypeal carina not developed in profile; anterior margin of pronotal disc carinate and followed by foveae; apex of aedeagal ventral ramus folded.</p><p>Material examined. Holotype, ♂, COSTA RICA, Monteverde: 1520m, 25.VI–2.VII.1983, D.H Lindeman col. (PMAE) ; PANAMA, Colón: 1♂, 14 km N J[un]ct[ion] Escobal r[oa]d and Piña r[oa]d on <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.9&amp;materialsCitation.latitude=9.333333" title="Search Plazi for locations around (long -79.9/lat 9.333333)">Piña</a> r[oa]d, ± 20 m, [09°20'N', 79°54'W], 2–3.VI.1995, F[light] I[nterception] T[rap], J. Ashe &amp; R. Brooks [col.] (CNCI) ; Coche {= Coclé}: 1♂, El Cope {= <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.58417&amp;materialsCitation.latitude=8.619722" title="Search Plazi for locations around (long -80.58417/lat 8.619722)">El Copé</a>}, [08°37'11"N, 80°35'03"W], 19.XI.1994, Windson &amp; Edwards [col.] (CNCI) ; Panama: 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.833336&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -79.833336/lat 8.666667)">Cerro Campana</a>, 08°40'N, 79°50'W, 7–14.V.1973 (CNCI) ; 1♂, C[anal] Z[one], B[arro] Colorado Is[land], [09°09'N, 79°51'W], 1–14.VII.1994, F[light] I[nterception] T[rap], D. Banks [col.] (CNCI); Darién Pr [ovincia]: 58♂, P[arque] N[acional] Darién, Pirre, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Est</a> [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 77°45'W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP); 2♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.683334&amp;materialsCitation.latitude=7.75" title="Search Plazi for locations around (long -77.683334/lat 7.75)">Cana</a>, 7°45'N, 77°41'W, 3–7. VI. 96, 550m, F[light] I[nterception] T[rap] J. Ashe, R. Brooks (UFES) .</p><p>Distribution (Fig. 239). Costa Rica and Panama.</p></div>	https://treatment.plazi.org/id/3B3687C07C559B02FF67F2A89D8AFBD6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C559B02FF67F6629CBFF9B8.text	3B3687C07C559B02FF67F6629CBFF9B8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus ramosus Alencar & Azevedo 2006	<div><p>Dissomphalus ramosus Alencar &amp; Azevedo, 2006</p><p>Dissomphalus ramosus Alencar &amp; Azevedo, 2006: 20 –21, 28 (♂ holotype from Ecuador, PMAE, figs 28–30).</p><p>Diagnosis. Mandible tridentate. Clypeus trapezoidal. Tergal process with pair of circular, inclined and deep lateral depressions with tuft of setae directed backward. Hypopygium with posterior margin convex. Genitalia: paramere dorsal margin straight; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; surface horizontal; basal half wide; inner margin irregular; apical half narrowing to apex; aedeagal dorsal body wide; surface vertical; apical part narrow and mesad; apex with three arms; crown-process with long lateral prolongation in middle of aedeagal dorsal body. Apodeme not extending beyond genital ring.</p><p>Remarks. No additional material of this species was not found in this study.</p><p>Material examined. Holotype, ♂, ECUADOR, Pichincha: 47km S. S[an]to Domingo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.15&amp;materialsCitation.latitude=-0.5" title="Search Plazi for locations around (long -79.15/lat -0.5)">Rio Palenque Res</a> [erve] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.15&amp;materialsCitation.latitude=-0.5" title="Search Plazi for locations around (long -79.15/lat -0.5)">Sta</a> [tion], 160–180m, rainforest, [0º30’S 79º9’W], 29.IV–5.V.1987, 2MT, B.V, Brown and L. D Coote col. (PMAE).</p><p>Distribution (Fig. 241). Costa Rica, Panama, Venezuela and Ecuador.</p></div>	https://treatment.plazi.org/id/3B3687C07C559B02FF67F6629CBFF9B8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C549B03FF67F2A89E70FC05.text	3B3687C07C549B03FF67F2A89E70FC05.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus curvifoveatus Azevedo 1999	<div><p>Dissomphalus curvifoveatus Azevedo, 1999</p><p>(Fig 159–164)</p><p>Dissomphalus curvifoveatus Azevedo, 1999a: 328 –329, 368, 386–387 (♂, holotype from Brazil, Amazonas, INPA, figs: 40–42, 165); Azevedo, 2003: 60.</p><p>Diagnosis. Mandible bidentate. Clypeus subtrapezoidal. Tergal process median with wide, deep and triangular depression, with very large pit densely covered with setae. Hypopygium with posterior margin with very short invagination. Genitalia: paramere bidigitiform with median longitudinal invagination very deep; volsella with digitus very long 0.5X the length of paramere; aedeagal dorsal ramus longer than aedeagal dorsal body, surfaces vertical; apex narrow and smooth; inner margin excavated medially and straight basally; outer margin convex; aedeagal dorsal body with two pairs of apical lobes; ventral pair narrow, lanceolate with apex sharpened; dorsal pair narrowing to apex divergent, with two teeth. Apodeme not extending beyond genital ring.</p><p>Remarks. This species is known to Belize, Costa Rica, Panama, Venezuela, Ecuador and Brazil (Acre, Amazônia, Pará, Rondônia and Paraná) (Azevedo 1999a) and here more specimens are added. Some variations are worth to mention: median clypeal tooth angulate; anterior margin of pronotal disc coarse; one specimen with depressions of tergal process with few setae and tergal process more distant from each other.</p><p>Material examined. Holotype, ♂, BRAZIL, Amazonas: Reserve 1112, 8.xi. 1985, Malaise trap, B. Klein col. (INPA) ; PANAMA, [Panama Province]: 1♂, Nusagandi Sta [tion] n[ea]r Columbia INA <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.53333&amp;materialsCitation.latitude=8.966666" title="Search Plazi for locations around (long -79.53333/lat 8.966666)">Igar Trail</a>, [08°58'N, 79°32'W], 12–13. V.1994, A. Windson [col.] (CNCI) ; Darién Pr [ovincia]: 3♂, P[arque] N[acional] Darién, Pirre, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Est</a> [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 77°45'W], 16 nov 00 – 17 ene 2001, [Trampa] Malaise, R. Cambra, A. Santos [col.] (MIUP).</p><p>Distribution (Fig. 232). Belize, Costa Rica, Panama, Venezuela, Ecuador and Brazil (Amazonas, Pará, Rondônia).</p></div>	https://treatment.plazi.org/id/3B3687C07C549B03FF67F2A89E70FC05	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C549B03FF67F6199CADFAC9.text	3B3687C07C549B03FF67F6199CADFAC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus gilvipes Evans 1979	<div><p>gilvipes species-group</p><p>Diagnosis. Male. Clypeus broadly projected forward. Metasoma with lateral tergal process, tufts of setae with most lateral setae longer than others.</p><p>Remarks. It has five species: D. gilvipes Evans, D. krombeini Azevedo, D. altyclypeatus Azevedo, D. bicerutus Azevedo, and D. arbeius sp. nov. .</p><p>Distribution. Ranging from Southern Florida to northern Argentina.</p></div>	https://treatment.plazi.org/id/3B3687C07C549B03FF67F6199CADFAC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C549B0CFF67F7639CF1FD77.text	3B3687C07C549B0CFF67F7639CF1FD77.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus arbeius Azevedo 2017	<div><p>Dissomphalus arbeius Brito &amp; Azevedo, sp. nov.</p><p>(Figs 17, 48, 79, 165–166)</p><p>Description. Male. Body length 2.00 mm. Color: body castaneous. Head (Fig. 17). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina low in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 48). Tergal process, with lateral, shallow, linear and small depression, without tubercle, pit absent, with very short tuft of setae with one distinctly long seta. Posterior hypopygeal margin weakly concave (Fig. 79). Genitalia (Figs 165–166): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin slightly sinuous; apex short, simple, angled outward; additional inner ramus absent; basal stub short and very narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, subvertical and wide with apical margin abruptly acute and dorsad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring.</p><p>Variation. Head, mesosoma and metasoma light castaneous.</p><p>Remarks. This species is similar to D. gilvipes by having the dorsal margin of the paramere expanded distally and the aedeagal ventral ramus abruptly narrowing apicad. However, this species has the apical lobes of the aedeagal dorsal body short and wider and the apical margin of the paramere crenulate, whereas D. gilvipes has the apical lobes of the aedeagal dorsal body long and narrow and the apical margin of the paramere smooth.</p><p>Material examined. Holotype, ♂, PANAMA, Chiriqui: 2 km W <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.583336&amp;materialsCitation.latitude=8.85" title="Search Plazi for locations around (long -82.583336/lat 8.85)">Cerro Punta</a>, 1700 m, [08°51'N, 82°35'W], 19. V–8.VI.1977, Peck &amp; Howden [col.] (CNCI) . Paratypes: 12♂, same data of holotype; 1♂, STRI n[ea]r <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.24027&amp;materialsCitation.latitude=8.72611" title="Search Plazi for locations around (long -82.24027/lat 8.72611)">Futurna</a> {= Fortuna}, 1260 m, 8°43'34"N, 82°14'25"W, 30.V.2000, Y[ellow] P[an] T[rap], B. &amp; J. Gill [col.] (CNCI) ; 15♂, P[arque] I[nternacional] L[a] A[mistad], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.73333&amp;materialsCitation.latitude=8.9" title="Search Plazi for locations around (long -82.73333/lat 8.9)">Jurutungo</a>, 1800 m, [08°54'N, 82°44'W], 30oct.5 nov 1994, Tr [ampa] Malaise (13♂) and Tr [ampas] Amarillas (2♂) A. Rodriguez [col.] (MIUP) ; 2♂, 15 km NW Hato del <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.63333&amp;materialsCitation.latitude=8.766666" title="Search Plazi for locations around (long -82.63333/lat 8.766666)">Volcano</a>, 1200 m, [08°46'N, 82°38'W], 24–31. V.1997, Peck &amp; Howden [col.] (CNCI) ; 3♂, Fortuna, Div. Cont. Chiriqui- B[ocas del] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.19092&amp;materialsCitation.latitude=8.77125" title="Search Plazi for locations around (long -82.19092/lat 8.77125)">Toro</a>, 1070–1090 m, [08°46'16.5"N, 82°11'27.3"W], 8–11.abr.1999, Tr [ampas] Amar [illas], R. Cambra, A. Santos [col.] (MIUP) ; Coclé Pr [ovincia]: 1♂, El Copé, Par [que] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">Nac</a> [ional <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">General de División</a>] Omar Torrijos H [errera], 570–670 m.s.n.m., [08º41'56"N, 80º39'17"W], 26–30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; Panama Pr [ovincia]: 4♂, P[arque] Nac [ional] Altos de Campana, [08º43'15"W, 80º01'43"W], 10–14 ago 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; 2♂, P[arque] Nac [ional] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.71667&amp;materialsCitation.latitude=9.15" title="Search Plazi for locations around (long -79.71667/lat 9.15)">Sob</a> [eranía], Camino, Plantación, [09º09'N, 79º43'W], 24 ago.–2 sep.1999 (MIUP) ; 1♂, Cerro Azul, Las Nubes, [09º10'N, 79º25'W], 9–15–oct.1999, R. Cambra, A. Santos [col.] (MIUP); 2♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.833336&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -79.833336/lat 8.666667)">Cerro Campana</a>, 08°40'N, 79°50'W, 7–14. V.1973 (CNCI).</p><p>Distribution (Fig. 228). Panama (Chiriqui, Coclé and Panama).</p></div>	https://treatment.plazi.org/id/3B3687C07C549B0CFF67F7639CF1FD77	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5B9B0CFF67F0C39C8EFA28.text	3B3687C07C5B9B0CFF67F0C39C8EFA28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus gilvipes Evans 1979	<div><p>Dissomphalus gilvipes Evans, 1979</p><p>Dissomphalus gilvipes Evans, 1979: 278, 279, 283 (♂, holotype from Colombia, FSCA, fig. 7); Gordh &amp; Móczár, 1990: 222, Azevedo, 1999b: 929, 931–932; 2003: 22; Redighieri &amp; Azevedo, 2004: 330; 2006: 309, Colombo &amp; Azevedo, 2016: 16 – 17.</p><p>Diagnosis. Mandible tetradentate. Clypeus broadly projected toward with three small teeth. Tergal process lateral, with pair of subcircular depressions with distinct long lateral setae. Hypopygium with posterior margin straight, with small rounded median projection. Genitalia: paramere with dorsal margin developed inward; aedeagal ventral ramus as long as aedeagal dorsal body, laminar with surfaces vertical; basal half wide and apical narrow; apex blunt and oblique; aedeagal dorsal body with pair of parallel lobe, narrowing apically; inner lobe setose; base of genitalia with pair of laminar, rectangular expansion. Apodeme not extending beyond genital ring.</p><p>Remarks. This species is known to Panama, Colombia, Ecuador, Peru, Brazil (Amazonas, Para, Rondônia, Mato Grosso, Pernambuco, Minas Gerais, Distrito Federal, São Paulo and Espírito Santo) (Colombo &amp; Azevedo 2016) and here two specimens are added. Some variations are worth to mention: outer lobe of apical lobes of aedeagal dorsal body entirely narrow, laminar in lateral view, convergent apically; aedeagal ventral ramus with apex slightly laterad.</p><p>Material examined. Holotype ♂, COLOMBIA, Dept. Valle: Central de Anchicaya, 30km E Buenaventura, 560m, 14–16 July 1975, R. Wilkerson, Malaise trap (FSCA) . PANAMA, Chiriqui: 2#, 15 km NW Hato del <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.63333&amp;materialsCitation.latitude=8.766666" title="Search Plazi for locations around (long -82.63333/lat 8.766666)">Volcano</a>, 1200 m, [08°46'N, 82°38'W], 24–31.V.1997, Peck &amp; Howden [col.] (CNCI) .</p><p>Distribution (Fig. 235). Panama, Colombia, Ecuador, Peru, Brazil (Amazonas, Para, Rondônia, Mato Grosso, Pernambuco, Minas Gerais, Distrito Federal, São Paulo and Espírito Santo).</p></div>	https://treatment.plazi.org/id/3B3687C07C5B9B0CFF67F0C39C8EFA28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5B9B0CFF67F4759DAFF8C9.text	3B3687C07C5B9B0CFF67F4759DAFF8C9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus guttus Azevedo 2003	<div><p>guttus species-group</p><p>Diagnosis. Male. Metasoma with lateral tergal process with pair of lateral drop-shaped pits, high rim, sharpened anteriorly and rounded posteriorly.</p><p>Remarks. It has two species: D. guttus Azevedo and D. latimerus Azevedo. Distribution. Costa Rica, Panama and Brazil.</p></div>	https://treatment.plazi.org/id/3B3687C07C5B9B0CFF67F4759DAFF8C9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5A9B0EFF67F53A9D8AFCE8.text	3B3687C07C5A9B0EFF67F53A9D8AFCE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus guttus Azevedo 2003	<div><p>Dissomphalus guttus Azevedo, 2003</p><p>(Fig 167–168)</p><p>Dissomphalus guttus Azevedo, 2003: 48 –50 (♂, holotype from Costa Rica, CNCI, figs 91–93).</p><p>Diagnosis. Mandible tridentate. Clypeus subtrapezoidal. Tergal process with pair of droped-shaped pits with outer rim, sharpened anteriorly and rounded posteriorly, with tuft of setae backward and very shallow depression.</p><p>Hypopygium with posterior margin straight. Genitalia: paramere wide, apex rounded; aedeagal ventral ramus shorter than aedeagal dorsal body; apical half narrow and strongly curved downward; aedeagal dorsal body with two pairs of apical lobes; outer pair laminar; surface vertical, wide with rounded apex in lateral view; apex tridentate, directed downward; ventral margin with anterior elongate basal tooth; inner pair membranous, stout and setose; base of aedeagus strongly dilated upward. Apodeme extending beyond genital ring.</p><p>Remarks. This species is known to Costa Rica and Panama (Azevedo 2003) and here more specimens are added. Some variations are worth to mention: median clypeal tooth angulate; median clypeal carina incomplete apically; anterior margin of pronotal disc coarse.</p><p>Material examined. Holotype, ♂, COSTA RICA, Sant Vito de Coto Brus: Las Cruces, 1200m, 9. VII.1982, Malaise trap, B. Gill col. (CNCI) . PANAMA, Bocas del Toro: 4♂, Wekso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Teribe</a>, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP) ; Chiriqui: 6♂, Finca La Suiza near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 25–6.VI.2000, F[light] I[nterception] T[rap], H. &amp; A. Howden [col.] (CNCI) ; 1♂, 15 km NW Hato del <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.63333&amp;materialsCitation.latitude=8.766666" title="Search Plazi for locations around (long -82.63333/lat 8.766666)">Volcano</a>, 1200 m, [08°46'N, 82°38'W], 24–31.V.1997, Peck &amp; Howden [col.] (CNCI) ; Veraguas Pr [ovincia]: 1♂, Calovebora, Guazarito, 15–18 feb. 2000, Malaise, Col. S. Bermudez (MIUP); Darién Pr [ovincia]: 7♂, P[arque] N[acional] Darién, Pirre, Est [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 77°45'W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP); 1♂, Est [ación] I[nstituto] N[acional de] Re [cusos] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Na</a> [turales] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Re</a> [novables], [07°51'26.3"N, 77°28'12.7"W], 5–28 feb 1993, R. Cambra [col.] (MIUP).</p><p>Distribution (Fig. 236). Costa Rica and Panama.</p></div>	https://treatment.plazi.org/id/3B3687C07C5A9B0EFF67F53A9D8AFCE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C599B0FFF67F7BD9C29FE2A.text	3B3687C07C599B0FFF67F7BD9C29FE2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus pilus Alencar & Azevedo 2008	<div><p>Dissomphalus pilus Alencar &amp; Azevedo, 2008</p><p>Dissomphalus pilus Alencar &amp; Azevedo, 2008: 0 4, 0 9, 20–22 (♂, holotype from Trinidad &amp; Tobago, CNCI, figs 71–77).</p><p>Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with pair of minute tubercle with small pit on top, and few setae directed backward. Hypopygium with posterior margin concave. Genitalia: paramere very wide and apex rounded; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; surface horizontal; inner margin straight outer margin convex; filament inserted subapically; aedeagal dorsal body with two pairs of apical lobes; outer pair laminar; surface vertical and wide; apex straight; inner pair irregular in lateral view, membranous, setose and granulate in ventral view; basal bar triangular; conector with lateral expansion concave. Apodeme extending beyond genital ring.</p><p>Remarks. This species is known to Trinidad and Tobago (Alencar &amp; Azevedo 2008). Now it is recorded for the first time from Panama. Some variations are worth to mention: apical lobes of aedeagal dorsal body very projected forward and apical margin truncate.</p><p>Material examined. Holotype, ♂, TRINIDAD [and TOBAGO]: [Saint George]: 16 km N Arima, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-61.266666&amp;materialsCitation.latitude=10.5" title="Search Plazi for locations around (long -61.266666/lat 10.5)">Andrews Trace</a>, 620m up. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-61.266666&amp;materialsCitation.latitude=10.5" title="Search Plazi for locations around (long -61.266666/lat 10.5)">Montane</a>, rain for[est], [~ 10º30’N 61º 16’ W], 7–24.vi.1993 . FIT, S. &amp; J. Peck col. (CNCI). PANAMA, Veraguas Pr [ovincia] : 12♂, P[arque] N[acional] Coiba, Isla Coiba, [07º28'09"N, 81º45'24"W], 27– nov–4–dic.1998 (1♂), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Cerro la Equis</a>, 130m , 8–12 mar [zo] 1998 (1♂), 29 jul–4 ag 1998, 100m (1♂), 120m (1♂), Malaise, R. Cambra, A. Santos [col.] ; 8-12 mar [zo] 1998, 0m (1♂), 90m (1♂), 30–nov–4–dic. 1998, 100m (1♂), 120m (1♂), 0m (1♂), R. Cambra, S. Bermudez [col.]; Est [ación] Biol [ogica] Coiba, 26 nov–4 dic 1998 (1♂) 5-6 ag 1998 (1♂), R. Cambra, A. Santos [col.] (MIUP); Panama Pr [ovincia]: 5♂, P[arque] Nac [ional] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.71667&amp;materialsCitation.latitude=9.15" title="Search Plazi for locations around (long -79.71667/lat 9.15)">Sob</a> [eranía], Camino, Plantación, [09º09'N, 79º43'W], 24 ago.–2 sep.1999, Malaise, A. Santos, P. González [col.] (MIUP) ; 3♂, P[arque] N[atural] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.54472&amp;materialsCitation.latitude=8.9925" title="Search Plazi for locations around (long -79.54472/lat 8.9925)">Metropolitano</a>, [08º59'33"N, 79º32'41"W], 24 ago.–3 sep.1999, Malaise, A. Santos, P. González [col.] (MIUP) ; 1♂, Cerro Azul, Las Nubes, [09º10'N, 79º25'W], 9–15.oct.1999, R. Cambra, A. Santos [col.] (MIUP); Coclé: 1♂, Falda Cerro La Vieja, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.201385&amp;materialsCitation.latitude=8.669723" title="Search Plazi for locations around (long -82.201385/lat 8.669723)">Chiguirí Arriba</a>, [08º40'11"N, 82º12'05"W], 24 abril 1994, A . Rodriguez, R. Cambra [col.] (MIUP).</p><p>Distribution (Fig. 239). Panama and Trinidad &amp; Tobago.</p></div>	https://treatment.plazi.org/id/3B3687C07C599B0FFF67F7BD9C29FE2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C589B0FFF67F44B9B83F8BC.text	3B3687C07C589B0FFF67F44B9B83F8BC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus punctatus (Kieffer 1910)	<div><p>punctatus species-group</p><p>Diagnosis. Male. Tergal process with circular depression, dense tufts of setae directed backward, tubercle absent. Remarks. It has four species: D. napo Evans, D. punctatus, D. bahiensis Redighieri &amp; Azevedo and D. tupinikim Colombo &amp; Azevedo.</p><p>Distribution. Costa Rica, Panama, Ecuador, Peru, Brazil (Amazonas, Acre, Sergipe, Alagoas, Pernambuco, Paraíba, Mato Grosso, Bahia, Espírito Santo, Rio de Janeiro, São Paulo and Paraná) and Paraguay.</p></div>	https://treatment.plazi.org/id/3B3687C07C589B0FFF67F44B9B83F8BC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C589B0FFF67F0789CF0F9F4.text	3B3687C07C589B0FFF67F0789CF0F9F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus raziogus Azevedo 2017	<div><p>Dissomphalus raziogus Brito &amp; Azevedo sp. nov.</p><p>(Figs 18, 49, 80, 169–170)</p><p>Description. Male. Body length 2.27 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 18). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina low in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest strongly concave. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 49). Tergal process absent. Posterior hypopygeal margin weakly concave (Fig. 80). Genitalia (Figs 169–170): paramere with dorsal margin wide basally; apical margin rounded. Aedeagal ventral ramus higher than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin excavated; outer margin slightly sinuous; apex long, simple, parallel; additional inner ramus absent; basal stub short and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, subhorizontal and narrow, with apical margin acute and mesad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Variations. Mesosoma dark castaneous; median clypeal lobe short.</p><p>Remarks. This species is similar to D. isortus sp. nov. by having the aedeagal dorsal body narrowing slightly apicad, the apical lobes of the aedeagal dorsal body in conical-shaped. However, this species has the tergal process absent, the paramere narrow, the apex of the apical lobes of the aedeagal dorsal body mesad and the aedeagal ventral ramus with filament inserted in the median part, characteristic diagnosis for microsticus species-group, whereas D. isortus sp. nov. has the tergal process with a tubercle, pit with thick setae the paramere wide, the apex of the apical lobes of the aedeagal dorsal body dorsad and the aedeagal ventral ramus without filament.</p><p>Material examined. Holotype, ♂, PANAMA, Bocas del Toro: Wekso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Teribe</a>, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ap] Malaise, A. Santos[col.] (MIUP) . Paratypes: 8♂, same data as holotype; Chiriquí Pr [ovincia]: 1♂, P[arque] I[nternacional] L[a] A[mistad], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.73333&amp;materialsCitation.latitude=8.9" title="Search Plazi for locations around (long -82.73333/lat 8.9)">Jurutungo</a>, 1800 m, [08°54'N, 82°44'W], 30 oct. 5 nov. 1994, Tr [ampa] Malaise, A. Rodriguez [col.] (MIUP) ; 1♂, 15 km NW Hato del <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.63333&amp;materialsCitation.latitude=8.766666" title="Search Plazi for locations around (long -82.63333/lat 8.766666)">Volcano</a>, 1200 m, [08°46'N, 82°38'W], 24–31.V.1997, Peck &amp; Howden [col.] (CNCI) ; 1♂, Fortuna: Div. Cont. Chiriqui-B[ocas del] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.19092&amp;materialsCitation.latitude=8.77125" title="Search Plazi for locations around (long -82.19092/lat 8.77125)">Toro</a>, 1070–1090 m, [08°46'16.5"N, 82°11'27.3"W], 8–11.abr.1999, Malaise, R. Cambra, A. Santos [col.] (MIUP) .</p><p>Distribution (Fig. 241). Panama (Bocas del Toro and Chiriquí).</p></div>	https://treatment.plazi.org/id/3B3687C07C589B0FFF67F0789CF0F9F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5F9B08FF67F2A89B70FB39.text	3B3687C07C5F9B08FF67F2A89B70FB39.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus punctatus (Kieffer 1910)	<div><p>Dissomphalus punctatus (Kieffer, 1910)</p><p>Thaumatepyris punctatus Kieffer, 1910a: 47 (♂, holotype from Peru, ZMHB).</p><p>Dissomphalus punctatus: Evans, 1964: 47 (♂, new combination, figs 24, 26, 33); Azevedo, 2003: 28 –30; Redighieri &amp; Azevedo, 2006: 310 –311; Azevedo, 1999b: 492.</p><p>Glenobethylus montanus Kieffer, 1910a: 50 (♂, holotype, Pérou, Départ. de Cuzco, Plateau de Cosnipata, altitude de 1000 mètres, en janvier (Garlepp); ZMBH). Synonymized by Azevedo, 1999b: 492.</p><p>Dissomphalus montanus: Evans, 1964: 46, 48 (♂, new combination).</p><p>Dissomphalus napo: Redighieri &amp; Azevedo, 2004: 330, part.</p><p>Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with very shallow, small and rounded depressions covered with dense tuft of setae directed backward. Hypopygium with dorsal margin straight. Genitalia: paramere with rounded apex with dorsal margin straight; aedeagal ventral ramus slightly shorter than aedeagal dorsal body, sinuous and wide; with apical filament directed outward; aedeagal dorsal body with two pairs of apical lobes; dorsal lobe with filament with sharpened apex, inner surface setose, vertical; ventral lobe longer, membranous, wide with rounded apex in lateral view, surfaces vertical; inner lobe densely setose. Apodeme extending beyond genital ring.</p><p>Remarks. This species is known to Costa Rica, Panama, Ecuador, Peru and Brazil (Amazonas, Acre, Pernambuco, Alagoas, Sergipe, Bahia, Espírito Santo, Rio de Janeiro, São Paulo and Paraná). Now some specimens are added. Some variations are worth to mention: median clypeal tooth angulate; median clypeal carina straight and complete apically; anterior margin of pronotal disc coarse; tergal process with very shallow depression; paramere short, wider and apical margin straight; basivolsella with a rounded expansion.</p><p>Material examined. Holotype ♂, PERU, Cuzco: Cajon, Bergland, 1500m, 11.I.1901, Garlepp col. (ZMHB, #195); PANAMA, Chiriqui: 1♂, Santa Clara, Finca Hartmann, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.74555&amp;materialsCitation.latitude=8.860556" title="Search Plazi for locations around (long -82.74555/lat 8.860556)">Ojo de Agua</a>, [08°51'38"N, 82°44'44"W], 8– 9.XI.2001, M[alaise] T[rap], N. Schiff [col.] (UCDC) ; 1♂ Res [erva Florestal] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.237495&amp;materialsCitation.latitude=8.744167" title="Search Plazi for locations around (long -82.237495/lat 8.744167)">La</a> Fortuna, Viveno, [08°44'39"N, 82°14'15"W], 14–18.VI.1994, A. R. Gillogly [col.] (CNCI) ; Darién Pr [ovincia]: 28♂, P[arque] N[acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m , [08°00'N, 77°45'W], 16 nov–17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (25♂) and 1000 m, 21 mar–4 ab 2000, Cambra, Santos, Bermudez [col.] (2♂) (MIUP); 4♂, Serrania de Pierre, Cana, 1380 m , [07°51'N, 77°37'W], 4–7.VI.1986 F[light] I[nterception] T[rap], [J.] Ashe &amp; [R.] Brooks [col.] (2♂) (CNCI) and 1450m, 4–7.vi. 1995 (2♂) (UFES).</p><p>Distribution (Fig. 242). Costa Rica, Panama, Ecuador, Peru, and Brazil (Amazonas, Acre, Pernambuco, Alagoas, Sergipe, Bahia, Espírito Santo, Rio de Janeiro, São Paulo and Paraná).</p></div>	https://treatment.plazi.org/id/3B3687C07C5F9B08FF67F2A89B70FB39	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5F9B08FF67F7069CACF9F7.text	3B3687C07C5F9B08FF67F7069CACF9F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus rasissimus	<div><p>rasissimus species-group</p><p>Diagnosis. Male. Metasoma with median tergal proces, with very shallow, very large, depression occupying two median thirds of tergite, reaching broadly posterior margin of tergite I; half-moon-shaped. Paramere deeply divided medially forming two distinct lobes.</p><p>Remarks. It has two species: D. rasissimus Azevedo and D. bilobatus Azevedo. Distribution. Mexico (Chiapas and Oaxaca), Belize and Costa Rica.</p></div>	https://treatment.plazi.org/id/3B3687C07C5F9B08FF67F7069CACF9F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5F9B09FF67F4439D8AFC33.text	3B3687C07C5F9B09FF67F4439D8AFC33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus bilobatus Azevedo 1999	<div><p>Dissomphalus bilobatus Azevedo, 1999</p><p>(Figs 171–174)</p><p>Dissomphalus bilobatus Azevedo, 1999a: 0 4, 0 9, 20–22 (♂, holotype from Trinidad &amp; Tobago, CNCI, figs 71–77).</p><p>Diagnosis. Mandible oblique without teeth. Tergal process with very large depression hlaf-moon-shaped, with two low median tubercles and tuft of setae. Hypopygium with posterior margin slightly convex. Genitalia: paramere with deep median longitudinal invagination, dorsal lobe larger, apex wide, concave with corners rounded; ventral lobe thin, small, apex with setae; aedeagal ventral ramus shorter than aedeagal dorsal body, wide with apex truncate; surfaces divergent dorsally; ventral part of apex divided longitudinally with two serrated lamina; outer lamina small and narrowed apically and inner lamina larger; aedeagal dorsal body not developed narrowing apically; inner margin straight and parallel and outer margin convergent. Apodeme not extending beyond genital ring.</p><p>Remarks. This species is known to Costa Rica (Azevedo 1999). Now it is recorded for the first time from Panama. Some variations are worth to mention: mesosoma light castaneous; median clypeal tooth angulate; median clypeal carina complete; vertex crest straight; anterior margin of pronotal disc coarse; pit of tergal process with very short tuft of setae.</p><p>Material examined. Holotype, ♂, COSTA RICA, San Vito de Coto Brus: Las cruces, 1200m, 17.viii– 12.ix.1982, Malaise trap, B. Gill col. (CNCI) . PANAMA, Chiriquí: 1♂, 5.6 km N Boquete, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.42166&amp;materialsCitation.latitude=8.823055" title="Search Plazi for locations around (long -82.42166/lat 8.823055)">La Culebra</a> trail, [08°49'23"N, 82°25'18"W], F[light] I[nterception] T[rap], J. Ashe &amp; R. Brooks [col.] (CNCI) ; 1♂, [Santa Clara], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.74555&amp;materialsCitation.latitude=8.860556" title="Search Plazi for locations around (long -82.74555/lat 8.860556)">Harman's Finca</a>, [08°51'38"N, 82°44'44"W], 3.VII.1981, B. Gill [col.] (PMAE) ; Veraguas Pr [ovincia]: 2♂, Santa Fé, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.108444&amp;materialsCitation.latitude=8.518194" title="Search Plazi for locations around (long -81.108444/lat 8.518194)">Altos De Piedra</a>, [08°31'5.5"N, 81°06'30.4"W], 13–17 nov. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; Colón: 3♂, 14 km N J[un]ct[ion] Escobal r[oa]d and Piña r[oa]d on <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.9&amp;materialsCitation.latitude=9.333333" title="Search Plazi for locations around (long -79.9/lat 9.333333)">Piña</a> r[oa]d, ± 20 m, [09°20'N', 79°54'W], 2–3.VI.1995, F[light] I[nterception] T[rap], J. Ashe &amp; R. Brooks [col.] (CNCI) ; Coclé Pr [ovincia]: 2♂, El Copé, Par [que] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">Nac</a> [ional <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">General de División</a>] Omar Torrijos H [errera], 570–670 m.s.n.m., [08º41'56"N, 80º39'17"W], 26–30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; [Kuna Yala]: 1♂, Nusagandi Sta [tion] n[ea]r Columbia INA <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.53333&amp;materialsCitation.latitude=8.966666" title="Search Plazi for locations around (long -79.53333/lat 8.966666)">Igar Trail</a>, [08°58'N, 79°32'W], 12–13.V.1994, A. Windson [col.] (CNCI) ; Panamá Pr [ovincia]: 1♂, Parq [ue] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.71667&amp;materialsCitation.latitude=9.15" title="Search Plazi for locations around (long -79.71667/lat 9.15)">Nac</a> [ional] Soberanía, Camino, Plantación, [09º09'N, 79º43'W], 23–29 feb. 2000, P. González, A. Santos [col.] (MIUP) ; Comarca Kuna Yala: 6♂, Ustupu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.92667&amp;materialsCitation.latitude=9.1325" title="Search Plazi for locations around (long -77.92667/lat 9.1325)">Rio Abudi</a>, [09º07'57"N, 77º55'36"W], 6–12 dic. 1999, P. González [col.] (MIUP) ; Darién Pr [ovincia]: 57♂, P[arque] N[acional] Darién, Pirre, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Est</a> [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 77°45'W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (51♂), 1000m, 21 mar–4 ab 2000 (4♂) and 900 m (2♂) Cambra, Santos, Bermudez [col.] (MIUP); 3♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.61667&amp;materialsCitation.latitude=7.85" title="Search Plazi for locations around (long -77.61667/lat 7.85)">Cana</a>, 1380 m, [07°51'N, 77°37'W], 4–7.VI.1986 (2♂) , (CNCI) and 550 m, 7º 45'N 77° 41'W, 3–7.vi.96, (1♂), (UFES) F[light] I[nterception] T[rap], [J.] Ashe &amp; [R.] Brooks [col.].</p><p>Distribution (Fig. 231). Costa Rica and Panama.</p></div>	https://treatment.plazi.org/id/3B3687C07C5F9B09FF67F4439D8AFC33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5E9B09FF67F6009C88FAF0.text	3B3687C07C5E9B09FF67F6009C88FAF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus rufipalpis Kieffer 1910	<div><p>rufipalpis species-group</p><p>Diagnosis. Male. Metasoma with lateral tergal process, with pair of elliptical, inclined, elongated depression with linear and oblique tufts of setae.</p><p>Remarks. It has five species: D. ellipticus Evans, D. rufipalpis Kieffer, D. inclinatus Redighieri &amp; Azevedo, D. infissus Evans and D. guarani Colombo &amp; Azevedo.</p><p>Distribution. Ranging from the southern Florida to northern Argentina.</p></div>	https://treatment.plazi.org/id/3B3687C07C5E9B09FF67F6009C88FAF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5E9B0AFF67F7BD9EC8FE9F.text	3B3687C07C5E9B0AFF67F7BD9EC8FE9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus infissus Evans 1969	<div><p>Dissomphalus infissus Evans, 1969</p><p>Dissomphalus infissus Evans, 1969: 14, 19, 22 (♂, holotype from Argentina, FIML, figs 14, 20, 28); Gordh &amp; Móczár, 1990: 226; Azevedo, 1999b: 929, 933–934; 2003: 27; Redighieri &amp; Azevedo, 2006: 311, Colombo &amp; Azevedo, 2016: 28 –29.</p><p>Diagnosis. Mandible tridentate. Clypeus trapezoidal. Tergal process lateral with elliptical and inclined depressions and linear and oblique tuft of setae. Hypopygium with posterior margin straight. Genitalia: paramere short and wide; apex concave and very produced dorsally; aedeagal ventral very short; base wide; apical half narrow, slightly convergent with apex very sharpened; inner margin sinuous; outer margin expanded laterally; aedeagal dorsal body very wide, with apex laminar in lateral view; inner lamina sinuous, setose inside; outer lamina with series of grooves; base of aedeagal dorsal body slightly wider. Apodeme not extending beyond genital ring.</p><p>Remarks. This species is known to Brazil (Paraíba, Pernambuco, Espírito Santo, São Paulo, Paraná and Santa Catarina) and Argentina (Colombo &amp; Azevedo 2016). Now it is recorded for the first time from Panama. Some variations are worth to mention: head dark castaneous; frons strongly coriaceous; median clypeal tooth angulate; anterior margin of pronotal disc coarse; depression of tergal process very large; apex of paramere strongly curved inward.</p><p>Material examined. Holotype ♂, ARGENTINA, Oran: Abra Grande, Salta, 16–23 Feb. 1967, R. Golbach (FIML); PANAMA, Colón: 1♂ 14 km N J[un]ct[ion] Escobar r[oad] on Piña r[oa]d, ± 20 m, 09°20'N', 79°54'W], 2–3.VI.1995, F[light] I[nterception] T[rap], J. Ashe &amp; R. Brooks [col.] (CNCI); Darién Pr [ovincia]: 1♂, P[arque] N[acional] Darién, Pirre, Est[ación] Rancho Frio, 80 m, [08°00'N, 77°45'W], 16 nov–17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP).</p><p>Distribution (Fig. 237). Panama, Brazil (Paraíba, Pernambuco, Espírito Santo, São Paulo, Paraná and Santa Catarina) and Argentina.</p></div>	https://treatment.plazi.org/id/3B3687C07C5E9B0AFF67F7BD9EC8FE9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5C9B0BFF67F2A89EFAFB55.text	3B3687C07C5C9B0BFF67F2A89EFAFB55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus rufipalpis Kieffer 1910	<div><p>Dissomphalus rufipalpis Kieffer, 1910</p><p>(Figs 175–177)</p><p>Dissomphalus rufipalpis Kieffer, 1910b: 44 –45, 1914b: 496, 500, Evans, 1962: 69, 74–75, 1964: 46, Gordh &amp; Móczár, 1990: 225, Azevedo, 1999c: 944 –945; Azevedo, 2003: 28, Redighieri &amp; Azevedo, 2006: 311. Dissomphalus obliquos Evans, 1979: 277, 278, 279–280; Gordh &amp; Móczár, 1990: 224, Azevedo, 1999c: 9944 (designation of synonymy).</p><p>Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with elliptical and and inclined depressions and linear and oblique tuft of small setae. Hypopygium with posterior margin nearly straight. Genitalia: paramere with dorsal margin developed with base projected downward and small tooth; aedeagal ventral ramus as long as aedeagal dorsal body, laminar; surfaces vertical; basal half wide, narrowing at sinuous apex; aedeagal dorsal body with two pairs of apical lobes; outer pair laminar; surface vertical and setose; inner pair slightly inclined upward with setose surface. Apodeme extending beyond genital ring.</p><p>Remarks. This species is known to Mexico (Vera Cruz, Chiapas), Belize, Guatemala, Honduras, Costa Rica, Panama, Colombia and Ecuador (Azevedo 1999b) and here more specimens are added. In this series the specimens are quite similar to those previously studied, without any variation that worths to mention.</p><p>Material examined. Holotype ♂, BELIZE, #4 (CASC, #17055) ; PANAMA, Chiriqui: 3♂, Finca La Suiza near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 25–6.VI.2000, F[light] I[nterception] T[rap], H. &amp; A. Howden [col.] (CNCI) ; 1♂, 15 km NW Hato del <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.63333&amp;materialsCitation.latitude=8.766666" title="Search Plazi for locations around (long -82.63333/lat 8.766666)">Volcano</a>, 1200 m, [08°46'N, 82°38'W], 24–31.V.1997, Peck &amp; Howden [col.] (CNCI) ; Veraguas Pr [ovincia]: 1♂, P[arque] N[acional] Coiba, Isla Coita {= <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Coiba</a>}, [07º28'09"N, 81º45'24"W], 27nov– 4dic 1998, Malaise, R. Cambra, S. Ber [mudez col.] (MIUP) ; 1♂, Santa Fé D[is]tr[ict], [08°31'N, 81°06'W], 24–26 marzo 1999, B. Amarillas, col: A. Santos, L. D. (MIUP); Comarca Kuna Yala: 1♂, Ustupu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.92667&amp;materialsCitation.latitude=9.1325" title="Search Plazi for locations around (long -77.92667/lat 9.1325)">Rio Abudi</a>, [09º07'57"N, 77º55'36"W], 6–12 dic. 1999, Malaise, P. González [col.] (MIUP) ; Darién Pr [ovincia]: 12♂, P[arque] N[acional] Darién, Pirre, Est [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 450m, [08°00'N, 77°45'W], 24–26 marzo 2001, Malaise, col. A. Santos (2♂) , 230m, 21 mar–4 ab 2000, Cambra, Santos, Bermudez [col.] (1♂), 80m, 16 nov 00 – 17 ene 2001 R. Cambra, A. Santos [col.] (9♂) (MIUP);</p><p>Distribution (Fig. 244). Mexico (Vera Cruz, Chiapas), Belize, Guatemala, Honduras, Costa Rica, Panama, Colombia and Ecuador.</p></div>	https://treatment.plazi.org/id/3B3687C07C5C9B0BFF67F2A89EFAFB55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5C9B14FF67F4B39B2EFC7B.text	3B3687C07C5C9B14FF67F4B39B2EFC7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus areius Azevedo 2017	<div><p>Dissomphalus areius Brito &amp; Azevedo, sp. nov.</p><p>(Figs 19, 50, 81, 178–179)</p><p>Description. Male. Body length 1.55 mm. Color: body castaneous. Head (Fig. 19). Mandible with two distal teeth. Median clypeal lobe subtrapezoidal, with one angulate tooth; median clypeal carina low in profile; incomplete apically; straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 50). Tergal process with lateral, very shallow, subcircular and small depression, with small, high and laterad tubercle placed on inner area of depression, pit small, with short and sparse tuft of setae. Posterior hypopygeal margin weakly concave (Fig. 81). Genitalia (Figs 178–179): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus higher than dorsal body, evenly narrow; cross section laminar; surface subhorizontal; inner margin sinuous; outer margin slightly sinuous; apex short, simple, translucent, slightly curved inward; additional inner ramus absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, horizontal and wide, with apical margin rounded and laterad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring.</p><p>Variations. Metasoma light castaneous, tubercle low and with dense tuft of setae, ventral projection of apical lobe of aedeagal dorsal body longer, narrower or even absent.</p><p>Remarks. This species is easily recognized by having the apical lobes of the aedeagal dorsal body with a small laterad expansion bear-ear-shaped. This species is similar to D. sartus sp. nov. by having the paramere very excavated apically and the aedeagal dorsal body with three narrow, long filaments which connect the apical lobes of the aedeagal dorsal body to the basal portion of the aedeagal dorsal body. However, this species has the apical lobes of the aedeagal dorsal body with a small laterad projection, the aedeagal ventral ramus with the apex translucent and smooth, whereas D. sartus sp. nov. has the apical lobes of the aedeagal dorsal body with the apex abruptly acute and the aedeagal ventral ramus with the apex not translucent and toothed.</p><p>Material examined. Holotype, ♂, PANAMA, Panama Pr [ovincia]: P[arque] Nac[ional] Sob[eranía], Camino, Plantación, [09º09'N, 79º43'W], 24 ago.–2 sep.1999, Malaise, A. Santos, P. González [col.] (MIUP). Paratypes: Bocas del Toro: 2♂, Wekso, Teribe, P[arque] I[nternacional] L[a] A[mistad], 50 msnm,[09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr[ampa] Malaise, A. Santos[col.] (MIUP); Chiriqui: 1♂, Finca La Suiza near Hornitos, [08°40'N, 82°14'W], 2–6.VI.2000, F[light] I[nterception] T[rap], H. &amp; A. Howden [col.] (CNCI); Coche {= Coclé}: 1♂, El Cope {=El Copé}, [08°37'11"N, 80°35'03"W], 19.XI.1994, Windson &amp; Edwards [col.] (CNCI). 2♂, same data as holotype; 2♂, same province of holotype, P[arque] N[atural] Metropolitano, [08º59'33"N, 79º32'41"W], 24 ago.–3 sep.1999 (MIUP);</p><p>Distribution (Fig. 227). Panama (Bocas del Toro, Chiriqui, Coclé and Panama).</p></div>	https://treatment.plazi.org/id/3B3687C07C5C9B14FF67F4B39B2EFC7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C439B15FF67F1D09C24FF6F.text	3B3687C07C439B15FF67F1D09C24FF6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus aretisus Azevedo 2017	<div><p>Dissomphalus aretisus Brito &amp; Azevedo, sp. nov.</p><p>(Figs 20, 51, 82, 180–181)</p><p>Description. Male. Body length 1.73 mm. Color: head castaneous; mesosoma and metasoma light castaneous. Head (Fig. 20). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one rounded very short tooth; median clypeal carina high in profile, incomplete apically, convex in profile. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 51). Tergal process with lateral, shallow, circular and large depression, with small, low and dorsad tubercle placed on center of depression, pit small, with short tuft of setae. Posterior hypopygeal margin strongly concave (Fig. 82). Genitalia (Figs 180–181): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; ramous bifid, surface horizontal; inner margin sinuous; outer margin sinuous; apex short, simple, parallel; additional inner ramus absent; basal stub short and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe very short, horizontal and narrow, with apical margin abruptly acute and laterad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Remarks. This species is the most different in setosus species-group by having the aedeagal dorsal body elongated and very short apical lobes of the aedeagal dorsal body. These characters are easy to be recognized. This species is similar to D. filus, by having the aedeagal dorsal body elongated with small apical lobes of the aedeagal dorsal body. However, this species has the paramere the same size as the basiparamere, the apical lobes of the aedeagal dorsal body very short and wide and the aedeagal ventral ramus bifid whereas D. filus has the paramere longer than basiparamere, the apical lobes of the aedeagal dorsal body longer and narrow and the aedeagal ventral ramus not bifid. The base of the aedeagal ventral ramus has small lamelar projections that are difficult to see in magnifying glass, but easily seem in microscope. In this species-group just this species and D. w-aedeagus has the ventral ramus bifid.</p><p>Material examined. Holotype, ♂, PANAMA, Veraguas Pr [ovincia]: P[arque] N[acional] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Coiba</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Isla Coiba</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Est</a> [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Biologica Coiba</a>, [07º28'09"N, 81º45'24"W], 5–6 ag 1998, Malaise, R. Cambra, A. Santos [col.] (MIUP) . Paratypes: 3♂, same data as holotype; Darién: 1♂, Est [acion] I[nstituto] N[acional de] Re [cusos] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Na</a> [turales] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Re</a> [novables], P[arque] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Nac</a> [ional] Darién, [07°51'26.3"N, 77°28'12.7"W], 5–28 feb 1993, Malaise, R. Cambra [col.] (MIUP) .</p><p>Distribution (Fig. 226). Panama (Veraguas and Darién).</p></div>	https://treatment.plazi.org/id/3B3687C07C439B15FF67F1D09C24FF6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C429B15FF67F33B9CF0FAF0.text	3B3687C07C429B15FF67F33B9CF0FAF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus citus Azevedo 2017	<div><p>Dissomphalus citus Brito &amp; Azevedo, sp. nov.</p><p>(Figs 21, 52, 83, 182–183)</p><p>Description. Male. Body length 1.55 mm. Color: body castaneous. Head (Fig. 21). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina very high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures large. Vertex crest straight. Pronotal disc weakly coriaceous; anterior margin carinate. Metasoma (Fig. 52). Tergal process with lateral, very shallow, subcircular and small depression, with very small, low and posterad tubercle placed on center of depression, pit very small, with short and dense tuft of setae. Posterior hypopygeal margin strongly concave (Fig. 83). Genitalia (Figs 182–183): paramere with dorsal margin entirely wide; apical margin rounded. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin sinuous; apex long with sinuous, slender ventral projection, curved outward; additional inner ramus absent; basal stub long and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, subhorizontal and narrow, with apical margin rounded and dorsad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Variations. Head, mesosoma and metasoma light castaneous; anterior margin of pronotal disc coarse; tergal process without depression.</p><p>Remarks. This species is easily recognized by having the aedeagal ventral ramus with slender ventral projection and the carina of the median clypeal tooth very high in profile and the ventral margin of the paramere thick with thick setae. This species is similar to D. magnus by having the dorsal margin of the paramere almost straight with distinct thick setae apical and longer than the basiparamere. However, this species has the aedeagal ventral ramus abruptly narrow apicad with one ventral slender projection and the apical lobes of the aedeagal dorsal body subhorizontal whereas D. magnus has the aedeagal ventral ramus divided in two tubular, sinuous, projections and the apical lobes of the aedeagal dorsal body horizontal.</p><p>Material examined. Holotype, ♂, PANAMA, [Chiriquí]: Fortuna, Div. Cont. Chiriqui-B[ocas del] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.190834&amp;materialsCitation.latitude=8.771111" title="Search Plazi for locations around (long -82.190834/lat 8.771111)">Toro</a>, 1070–1090 m, [08°46'16"N, 82°11'27"W], 8–11.abr.1999, Malaise, R. Cambra, A. Santos [col.] (MIUP) . Paratypes: 5♂, same data as holotype; 2♂, same of holotype except 9–11.abr.1999, Tr[ampas] Amar[illas] (MIUP); Bocas del Toro: 1♂, Wekso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Teribe</a>, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17-24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP) .</p><p>Distribution (Fig. 232). Panama (Bocas del Toro and Chiriquí).</p></div>	https://treatment.plazi.org/id/3B3687C07C429B15FF67F33B9CF0FAF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C429B17FF67F7489B5FFE2A.text	3B3687C07C429B17FF67F7489B5FFE2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus sartus Azevedo 2017	<div><p>Dissomphalus sartus Brito &amp; Azevedo, sp. nov.</p><p>(Figs 22, 53, 84, 184–185)</p><p>Description. Male. Body length 1.85 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 22). Mandible with two distal teeth. Median clypeal lobe subtrapezoidal, with one angulate tooth; median clypeal carina low in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc coriaceous; anterior margin ecarinate. Metasoma (Fig. 53). Tergal process with lateral, shallow, subcircular and small depression, with small, high and dorsad tubercle placed on center of depression, pit small, with short and dense tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 84). Genitalia (Figs 184–185): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus as long as dorsal body, wide, progressively narrowing apicad; cross section laminar; surface subhorizontal; inner margin straight; outer margin slightly sinuous; apex long, simple, slightly curved inward; additional inner ramus absent; basal stub short and very narrow. Aedeagal dorsal body with pair of apical lobe; outer lobe short, horizontal and very narrow, with apical margin abruptly acute and ventrad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Variations. Depression of tergal process very shallow, tubercle low; apical lobe of aedeagal dorsal body longer and narrower.</p><p>Remarks. This species is similar to D. areius sp. nov. by having the paramere excavated apically and expanded basally and the aedeagal dorsal body with three narrow and long filaments which connect the apical lobes of the aedeagal dorsal body to the basal portion of the aedeagal dorsal body. However, this species has the apex of apical lobes of the aedeagal dorsal body abruptly acute and the aedeagal ventral ramus with the apex toothed, whereas D. areius sp. nov. has the apex of the apical lobes of the aedeagal dorsal body with a small laterad projection and the aedeagal ventral ramus with the apex translucent and smooth.</p><p>Material examined. Holotype, ♂, PANAMA, Darién Pr [rovincia]: P[arque] N[acional] Darién, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Pirre</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Est</a> [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 77°45'W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP). Paratypes: Bocas del Toro: 3♂, Wekso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Teribe</a>, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP) ; Coclé Pr [ovincia]: 1♂, El Copé, Par [que] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">Nac</a> [ional <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">General de División</a>] Omar Torrijos H [errera], 570–670 m.s.n.m., [08º41'56"N, 80º39'17"W], 26–30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; Panama Pr [ovincia]: 1♂, P[arque] Nac [ional] Altos de Campana, [08º43'15"W, 80º01'43"W], 10–14 ago 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; 3♂ same data as holotype; 1♂, same of holotype except 1000 m, 21 mar–4 ab 2000 (MIUP).</p><p>Distribution (Fig. 245). Panama (Bocas del Toro, Coclé, Panama and Darién).</p></div>	https://treatment.plazi.org/id/3B3687C07C429B17FF67F7489B5FFE2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C409B10FF67F0029E41FEC2.text	3B3687C07C409B10FF67F0029E41FEC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus teobus Azevedo 2017	<div><p>Dissomphalus teobus Brito &amp; Azevedo, sp. nov.</p><p>(Figs 23, 54, 85, 186–187)</p><p>Description. Male. Body length 2.63 mm. Color: body castaneous. Head (Fig. 23). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina low in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc weakly coriaceous; anterior margin coarse. Metasoma (Fig. 54). Tergal process with lateral, deep, circular and large depression, with small, low and dorsad tubercle placed on center of depression, pit large, with short and dense tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 85). Genitalia (Figs 186–187): paramere with dorsal margin entirely wide; apical margin rounded. Aedeagal ventral ramus as long as dorsal body, evenly narrow; cross section laminar; surface horizontal; inner margin sinuous; outer margin slightly sinuous; apex short, simple, curved outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, subhorizontal and narrow, with apical margin abruptly acute and posterad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Variation. Tergal process with pit of tubercle with few setae.</p><p>Remarks. This species is easily recognized by having the paramere wide basally and the triangular shape of the outer lobe of the aedeagal dorsal body which is very projected forward. This species is not similar to anyone of the setosus species-group.</p><p>Material examined. Holotype, ♂, Darién Pr [ovincia]: P[arque] N[acional] Darién, Pirre, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Est</a> [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 77°45'W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP). Paratypes: Bocas del Toro, Wekso: 1♂, Teribe, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Tr</a> [ampas] Amarillas, A. Santos[col.] (MIUP) ; Chiriqui: 1♂, Finca La Suiza near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 25–6.VI.2000, F[light] I[nterception] T[rap], H. &amp; A. Howden [col.] (CNCI) ; 1♂, 15 km NW Hato del <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.63333&amp;materialsCitation.latitude=8.766666" title="Search Plazi for locations around (long -82.63333/lat 8.766666)">Volcano</a>, 1200 m, [08°46'N, 82°38'W], 24–31.V.1997, Peck &amp; Howden [col.] (CNCI) ; Veraguas Pr [ovincia]: 14♂, P[arque] N[acional] Coiba, Isla Coiba, Cerro la Equis 0m, (3♂), 90m (3♂), [07º28'09"N, 81º45'24"W], 8–12 mar [zo] 1998, Malaise, R. Cambra, A. Santos [col.], 120m, 29jul–4 ag 1998 (5♂), 100 m, 30– nov–4–dic.1998, R. Cambra, S. Bermudez [col.] (2♂) and Playa Blanca-Cativales, 31 jul–6 ago 1998, R . Cambra, A. Santos [col.] (1♂) (MIUP) ; Colón Pr [ovincia]: 1♂ P[a]r[que] Nac [ional] Portobelo, San Antonio, 16–29 feb. 2000, Tr [ampa] Malaise, P. González, A. Santos [col.] (MIUP) ; 1♂, 14 km N J[un]ct[ion] Escobal r[oa]d and Piña r[oa]d on <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.9&amp;materialsCitation.latitude=9.333333" title="Search Plazi for locations around (long -79.9/lat 9.333333)">Piña</a> r[oa]d, ± 20 m, [09°20'N', 79°54'W], 2–3.VI.1995, F[light] I[nterception] T[rap], J. Ashe &amp; R. Brooks [col.] (CNCI) ; Comarca Kuna Yala: 6♂, Ustupu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.92667&amp;materialsCitation.latitude=9.1325" title="Search Plazi for locations around (long -77.92667/lat 9.1325)">Rio Abudi</a>, [09º07'57"N, 77º55'36"W], 6–12 dic. 1999, Malaise, P. González [col.] (MIUP) ; Panama Pr [ovincia]: 11♂, P[arque] N[atural] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.54472&amp;materialsCitation.latitude=8.9925" title="Search Plazi for locations around (long -79.54472/lat 8.9925)">Metropolitano</a>, [08º59'33"N, 79º32'41"W], 24 ago.–3 sep.1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; 1♂, P[arque] Nac [ional] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.71667&amp;materialsCitation.latitude=9.15" title="Search Plazi for locations around (long -79.71667/lat 9.15)">Sob</a> [eranía], Camino, Plantación, [09º09'N, 79º43'W], 24 ago.–2 sep.1999 Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; 2♂, Chilibre, Parque Nacional Chagres, Campo Chagres, [09º25'54"N, 79º30'54"W], 24–28.enero.2000 Tr[ampa] Malaise, A. Santos, P. González [col.] (MIUP); 3♂, C[anal] Z[one], B[arro] Colorado Is[land], [09°09'N, 79°51'W], 15–31.VII.1994, (2♂) and 1–14.VII.1994 (1♂), F[light] I[nterception] T[rap], D. Banks [col.] (CNCI); 27♂, same data as holotype; 9♂, same province of holotype Est [ación] I[nstituto] N[acional de] Re [cusos] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Na</a> [turales] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Re</a> [novables] [07°51'26.3"N, 77°28'12.7"W], 5–28 feb 1993, Malaise, R. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Cambra</a> [col.] (MIUP) ; 6♂, Cana, Serrania de Pirre, 1250m, 4–7.vi.1996, J. Ashe, R. Brooks. (UFES) ; 5♂, Estac [ión] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Cruce de Mono</a>, [07°51'26.3"N, 77°28'12.7"W], 6 feb–4 mar 1993, Tr [ampa] malaise, R.</p><p>Cambra, J. Coronado [col.] (MIUP); 2♂, Cruce de Mono: Estac. I[nstituto] N[acional de] Re[cusos] Na[turales] Re[novables], 6.II–4.III. 1993 (MIUP).</p><p>Distribution (Fig. 246). Panama (Bocas del Toro, Chiriqui, Veraguas, Colón, Panama, Comarca Kuna Yala and Darién).</p></div>	https://treatment.plazi.org/id/3B3687C07C409B10FF67F0029E41FEC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C479B10FF67F3559D9AFB71.text	3B3687C07C479B10FF67F3559D9AFB71.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus politus Ashmead 1894	<div><p>Dissomphalus politus Ashmead, 1894</p><p>Dissomphalus politus Ashmead, 1894: 193, 195 (♂ holotype from Saint Vicent). Dalla Torre 1898: 558, Kieffer 1908: 36, 1914b: 496, 501, Evans, 1964: 46, Gordh &amp; Móczár 1990: 225, Azevedo 1999b: 54, 1999b: 927; Azevedo, 2003: 20, 71.</p><p>Diagnosis. Mandible tetradentate. Clypeus tridentate. Tergal process with pair of very shallow depressions, with minute tubercle with setae on top, outer margin of depression with group of thick setae. Hypopygium with posterior margin concave. Genitalia: paramere wide basally, narrowing to the rounded apex in lateral view, with strong setae above; aedeagal ventral ramus higher than aedeagal dorsal body, with three long and conspicuous apical horns; aedeagal dorsal body wide, with two pairs of apical lobes; outer pair stout and membranous; inner pair laminar, vertical with apex bidentate and directed downward.</p><p>Remarks. This species is known to Saint Vicent (Azevedo 2003). Now it is recorded for the first time from Panama. Some variations are worth to mention: head, mesosoma and metasoma light castaneous; median clypeal carina inconspicuous; median clypeal tooth angulate; anterior margin of pronotal disc slightly coarse; tergal process with tubercle, thick setae on pit and in outer margin of depression; aedeagal ventral ramus shorter than aedeagal dorsal body.</p><p>Material examined. Lectotype male: S[AIN]T VICENT, W[est] I[ndies]: 1000 feet, H. H. Smith col (USNM, #2497). PANAMA, Veraguas: 10♂, P[arque] N[acional] Coiba, Isla Coibita, [07º28'09"N, 81º45'24"W], 6–11 mar 1998 (1♂), 29 jul. 4 ag 1998, (3♂), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Cerro la Equis</a>, 120 m (1♂), Playa Blanca-Cativales 31 jul–6 ago 1998 (3♂), Est [ación] Biologica Coiba, 5–6 ag 1998 (2♂), Malaise, R. Cambra, A. Santos [col.] (MIUP); Darién : 1♂, Est [ación] I[nstituto] N[acional de] Re [cusos] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Na</a> [turales] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Re</a> [novables], P[arque] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Nac</a> [ional] Darién, [07°51'26.3"N, 77°28'12.7"W], 5–28 feb 1993, Malaise, R. Cambra [col.] (MIUP) ; 1♂, Cana 7º45' N, 77º 41 W, 3– 7.vi. 96, 550m, J. Ashe, R. Brooks. (UFES).</p><p>Distribution (Fig. 240). Panama and Saint Vicent.</p></div>	https://treatment.plazi.org/id/3B3687C07C479B10FF67F3559D9AFB71	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C479B10FF67F6C39D80FA1C.text	3B3687C07C479B10FF67F6C39D80FA1C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus strabus Azevedo 2003	<div><p>strabus species-group</p><p>Diagnosis. Tergal process lateral with pair of sublateral depressions, tubercle and tuft slightly directed toward each other.</p><p>Remarks. It has two species: D. strabus Azevedo and D. thysanus Azevedo. Distribution. Dominican Republic and Panama.</p></div>	https://treatment.plazi.org/id/3B3687C07C479B10FF67F6C39D80FA1C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C479B10FF67F42D9D25F82A.text	3B3687C07C479B10FF67F42D9D25F82A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus strabus Azevedo 2003	<div><p>Dissomphalus strabus Azevedo, 2003</p><p>Dissomphalus strabus Azevedo, 2003: 12, 52–53 (♂ holotype from Panama, MIUP, figs 100–103).</p><p>Diagnosis. Mandible bidentate. Clypeus tridentate. Tergal process with pair of rounded and shallow depression with pit, high him and tuft of setae slightly directed each other. Hypopygium with posterior margin straight. Genitalia: paramere with dorsal margin slightly convex; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; surface horizontal; apex with two rami; aedeagal dorsal body with two pairs of apical lobes, the outer pair narrow with apex sharpened strongly angled downward; inner margin with large and triangular basal expansion; inner pair membranous, stout and setose. Apodeme extending beyond the genital ring.</p><p>Material examined. Holotype, ♂, PANAMA, Fortuna: Cont. Chiriqui, B. Toro, 1070–1090m, 8–11.IV.1999, Malaise trap, R. Cambra &amp; A. Santos col. (MIUP) .</p><p>Remarks. No additional material of this species was not found in this study.</p><p>Distribution (Fig. 243). Panama.</p></div>	https://treatment.plazi.org/id/3B3687C07C479B10FF67F42D9D25F82A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C469B11FF67F3E59F91F9C0.text	3B3687C07C469B11FF67F3E59F91F9C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus plaumanni Evans 1964	<div><p>Dissomphalus plaumanni Evans, 1964</p><p>Dissomphalus plaumanni Evans, 1964: 47, 52, 58–59 (♂, holotype from Santa Catarina, BMNH, figs 19, 27, 32, 35); Azevedo, 1999b: 939 –941; 2003: 26–27; Redighieri &amp; Azevedo, 2004: 330; 2006: 312; Colombo &amp; Azevedo, 2016: 33 –34.</p><p>Diagnosis. Mandible tetradentate. Clypeus trapezoidal. Tergal process lateral with shallow depressions, pair of turbecles with pit of setae on top. Hypopygium with posterior margin convex with small conical expansion. Genitalia: paramere with acute apex and arched inward, dorsal margin developed; aedeagal ventral ramus short than aedeagal dorsal body, as high as base of cuspis; wide; laminar; inner and outer margins straight and slightly convergent; apex acute with outer margin slightly concave with ventral surface slightly directed outward; aedeagal dorsal body with pair of apical lobes, laminar, wide with rounded apex in lateral view, compressed in dorsal view, below the apex. Apodeme extending beyond genital ring.</p><p>Remarks. This is one of the most widespread species through the Neotropical region (Azevedo 1999b) and here more specimens are added. Some variations are worth to mention: median clypeal tooth angulate; median carina clypeal straight and complete; vertex crest straight; tergal process with very small and very shallow depression; pit of tubercle with few setae.</p><p>Material examined. Holotype ♂, BRAZIL, Santa Catarina: Nova Teutonia, 26 February 1945, Fritz Plaumann (BMNH) ; PANAMA, Chiriqui: 17♂, Finca La Suiza near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 2–6.VI.2000 (6♂) , 25–6.VI.2000 (11♂), F[light] I[nterception] T[rap], H. &amp; A. Howden [col.] (CNCI); 2♂ Res [erva Florestal] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.237495&amp;materialsCitation.latitude=8.744167" title="Search Plazi for locations around (long -82.237495/lat 8.744167)">La</a> Fortuna, Viveno, [08°44’39”N, 82°14’15”W], 14–18.VI.1994, A. R. Gillogly [col.] (CNCI) ; 1♂ 15 km NW Hato del <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.63333&amp;materialsCitation.latitude=8.766666" title="Search Plazi for locations around (long -82.63333/lat 8.766666)">Volcano</a>, 1200 m [08°46'N, 82°38'W], 24–31.V.1997, Peck &amp; Howden [col.] (CNCI) ; Veraguas Pr [ovincia]: 6♂, P[arque] N[acional] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Coiba</a>, Isla Coibita, [07º28'09"N, 81º45'24"W], 27–nov–4–dic.1998, (2♂) , Cerro La Equis, 120 m, 30–nov.–4dic.1998 (2♂) , 100m (1♂), 90m (1♂), Malaise, R. Cambra, S. Bermudez [col.] (MIUP); Colón: 2♂, 14 km N J[un]ct[ion] Escobal r[oa]d and Piña r[oa]d on <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.9&amp;materialsCitation.latitude=9.333333" title="Search Plazi for locations around (long -79.9/lat 9.333333)">Piña</a> r[oa]d, ± 20 m, [09°20'N', 79°54'W], 2–3.VI.1995, F[light] I[nterception] T[rap], J. Ashe &amp; R. Brooks [col.] (CNCI) ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.0&amp;materialsCitation.latitude=9.416667" title="Search Plazi for locations around (long -80.0/lat 9.416667)">Old Gamboa</a>: 1♂, C[anal] Z[one], [09°25'N, 80°00'W], 14.XI.1994, F[light] I[nterception] T[rap], P. Windson [col.] (CNCI) ; Darién Pr [ovincia]: 30♂ P[arque] N[acional] Darién, Pirre, Est [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-87.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -87.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 87°45'W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (29♂), 21 mar–4 ab 2000, Tr[ampa] Amarilla, Cambra, Santos, Bermudez [col.] (1♂) (MIUP).</p><p>Distribution (Fig. 240). Costa Rica, Panama, Ecuador, Peru and Brazil (Pará, Espírito Santo, São Paulo and Paraná).</p></div>	https://treatment.plazi.org/id/3B3687C07C469B11FF67F3E59F91F9C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C469B11FF67F2A89E41FE58.text	3B3687C07C469B11FF67F2A89E41FE58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus tuberculatus	<div><p>tuberculatus species-group</p><p>Diagnosis. Male. Clypeus trapezoidal. Metasoma with lateral tergal process, with pair of tubercles slightly directed each other, placed in pair of shallow depression or in horizontal surface.</p><p>Remarks. It has two species: D. plaumanni Evans and D. tuberculatus Ashmead. Distribution. Saint Vincent, Costa Rica, Panama, Ecuador, Peru and Brazil (Pará, Espírito Santo, São Paulo and Paraná).</p></div>	https://treatment.plazi.org/id/3B3687C07C469B11FF67F2A89E41FE58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C469B11FF67F44D9E6BF875.text	3B3687C07C469B11FF67F44D9E6BF875.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus ulceratus	<div><p>ulceratus species-group</p><p>Diagnosis. Male. Tergal process median with depression, pair of tufts of setae. Basal portion of dorsal margin of paramere much developed. Digitus wide medially. Cuspis elongated. Aedeagal ventral ramus with inner margin sinuous basally, wide, narrowing apically.</p><p>Remarks. It has six species: D. ulceratus Evans, D. dentiformis Azevedo, D. concavatus Azevedo, D. rectilineus Azevedo, D. congo Colombo &amp; Azevedo and D. terbius sp. nov. .</p><p>Distribution. Panama, Brazil (Distrito Federal, Espírito Santo, Rio de Janeiro, São Paulo and Paraná) and Argentina.</p></div>	https://treatment.plazi.org/id/3B3687C07C469B11FF67F44D9E6BF875	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C459B12FF67F2A89C64FC34.text	3B3687C07C459B12FF67F2A89C64FC34.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus terbius Azevedo 2017	<div><p>Dissomphalus terbius Brito &amp; Azevedo, sp. nov.</p><p>(Figs 24, 55, 86, 188–189)</p><p>Description. Male. Body length 2.39 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 24). Mandible with three distal teeth. Median clypeal lobe ill-defined, with one angulate very short tooth; median clypeal carina low in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc coriaceous; anterior margin carinate. Metasoma (Fig. 55). Tergal process with median, shallow, subcircular and large depression, without tubercle, with pit large, short and dense tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 86). Genitalia (Figs 188–189): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface vertical; inner margin straight; outer margin sinuous; apex long, simple, angled outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe very long, horizontal and wide, with apical margin angled and laterad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Variations. Head castaneous and mesosoma light castaneous; median clypeal tooth inconspicuous.</p><p>Remarks. This species is similar to D. afestus sp. nov. by having the apical lobes of the aedeagal dorsal body very long and wide and the apical half of the aedeagal ventral ramus narrow. However, this species has the dorsal margin of the paramere rounded with the apex crenulate and the apex of the aedeagal ventral ramus angled outward, whereas D. afestus sp. nov. has the paramere with the dorsal margin no so rounded with the apex smooth and the apex of the aedeagal ventral ramus not angled outward.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Material</a> examined. Holotype, ♂, PANAMA, Bocas del Toro: Wekso, Teribe, P [arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ampa] Malaise, A. Santos[col.] (MIUP) . Paratypes: 8 ♂ same data as holotype (MIUP) .</p><p>Distribution (Fig. 247). Panama (Bocas del Toro).</p></div>	https://treatment.plazi.org/id/3B3687C07C459B12FF67F2A89C64FC34	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C459B12FF67F6019A49FAD5.text	3B3687C07C459B12FF67F6019A49FAD5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus unitus Azevedo. Seven 1999	<div><p>unitus species-group</p><p>Diagnosis. Male. Tergal process median, rounded depression nearly totally covered by pair of joined tufts of long setae directed backward.</p><p>Remarks. It has two species: D. unitus and D. delgatus Azevedo.</p><p>Distribution. Mexico (Chiapas, Vera Cruz), Belize, Guatemala, Costa Rica, Panama and Venezuela.</p></div>	https://treatment.plazi.org/id/3B3687C07C459B12FF67F6019A49FAD5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C459B13FF67F7629A2EFD5B.text	3B3687C07C459B13FF67F7629A2EFD5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus unitus Azevedo. Seven 1999	<div><p>Dissomphalus unitus Azevedo, 1999</p><p>(Fig 190–194)</p><p>Dissomphalus unitus Azevedo, 1999a: 350 –351, 380–381, 390; (♂, holotype from Costa Rica, CNCI, figs 112–116, 193–194); Azevedo, 2003: 62.</p><p>Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with shallow and large depression with pair of joined tuft of long setae backward. Hypopygium with posterior margin with three small concavities. Genitalia: paramere with apex arched inward and truncate; aedeagal ventral ramus slightly longer than aedeagal dorsal body narrow, diverging dorsally; inner margin sinuous and parallel; outer margin with broad median concavity; aedeagal dorsal body with two pairs of apical lobes; outer lobe laminar with rounded apex; inner lobe stout densely setose. Apodeme extending beyond genital ring.</p><p>Remarks. This species is known to Mexico (Chiapas, Vera Cruz), Belize, Guatemala, Costa Rica and Panama (Azevedo 1999a) and here more specimens are added. Some variations are worth to mention: metasoma light castaneous; median clypeal tooth angulate; aedeagal ventral ramus as longer as aedeagal dorsal body; volsella with rounded expansion medially and some specimens without it.</p><p>Material examined. Holotype, ♂, COSTA RICA, San Coto de Vito Brus: Las Cruces, 1200m, 2–9.ix.1982, Malaise trap, B. Gill col. (CNCI) . PANAMA, Chiriqui: 4♂, 15 km NW Hato del <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.63333&amp;materialsCitation.latitude=8.766666" title="Search Plazi for locations around (long -82.63333/lat 8.766666)">Volcano</a>, 1200 m, [08°46'N, 82°38'W], 24–31.V.1997, Peck &amp; Howden [col.] (CNCI) ; 6♂, Res [erva Florestal] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.236664&amp;materialsCitation.latitude=8.690833" title="Search Plazi for locations around (long -82.236664/lat 8.690833)">La</a> Fortuna, Viveno, [08°41'27"N, 82°14'12"W], 14–18.VI.1994, A. R. Gillogly [col.] (CNCI) ; 1♂, water shed, hydro trail, 1100 m, [08°44'21"N, 82°14'36"W], 24.I.1994 – 27.I.1994, D. M. Windson [col.] (CNCI); 1♂, Finca La Suiza near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 2–6.VI.2000, F[light] I[nterception] T[rap], H. &amp; A. Howden [col.] (CNCI) ; 1♂, Santa Clara, Finca Hartman, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.74555&amp;materialsCitation.latitude=8.860556" title="Search Plazi for locations around (long -82.74555/lat 8.860556)">Ojo de Agua</a>, [08°51'38"N, 82°44'44"W], 8–9.XI.2001, M[alaise] T[rap], N. Schiff [col.] (UCDC) ; Veraguas: 1♂, Pr [ovincia] Santa Fé], Santa Fé, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.10833&amp;materialsCitation.latitude=8.518055" title="Search Plazi for locations around (long -81.10833/lat 8.518055)">Altos De Piedra</a>, [08°31'5"N, 81°06'30"W], 13–17 nov. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; Coclé Pr [ovincia]: 2♂, El Copé, Par[que] Nac [ional General de División] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">Omar Torrijos</a> H[errera], 570 – 670 m.s.n.m., [08º41'56"N, 80º39'17"W], 26–30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; Panamá Pr [ovincia]: 1♂, P[arque] N[atural] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.54472&amp;materialsCitation.latitude=8.9925" title="Search Plazi for locations around (long -79.54472/lat 8.9925)">Metropolitano</a>, [08º59'33"N, 79º32'41"W], 24 ago. –3 sep.1999 (MIUP) ; Darién Pr [ovincia]: 33♂, P[arque] N[acional] Darién, Pirre, Est [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 77°45'W], 16 nov–17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (26♂) , 1000 m, 21 mar–4 ab 2000, Cambra, Santos, Bermudez [col.] (5♂) and 580 m (1♂) (MIUP), 1450m, 4–7. vi. 1995, F[light] I[ntrception] T[rap] J. Ashe, R. Brooks (1♂) (UFES); 1♂, Nusagandi Sta [tion] n[ea]r Columbia INA <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.53333&amp;materialsCitation.latitude=8.966666" title="Search Plazi for locations around (long -79.53333/lat 8.966666)">Igar Trail</a>, [08°58'N, 79°32'W], 12–13.V.1994, A . Windson [col.] (CNCI).</p><p>Distribution (Fig. 247). Mexico (Vera Cruz and Chiapas), Belize, Guatemala, Costa Rica and Panama.</p></div>	https://treatment.plazi.org/id/3B3687C07C459B13FF67F7629A2EFD5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C449B13FF67F60B9CB8F832.text	3B3687C07C449B13FF67F60B9CB8F832.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus dilatatus Azevedo 1999	<div><p>Dissomphalus dilatatus Azevedo, 1999</p><p>(Figs 195–196)</p><p>Dissomphalus dilatatus Azevedo, 1999: 324 –325 (♂ holotype from Costa Rica, LACM, figs 32–35, 159–160).</p><p>Diagnosis. Mandible tridentate. Clypeus with median tooth. Tergal process with elliptical, deep and large depression with pair of tubercles and tuft of setae directed each other. Hypopygium with posterior margin concave. Genitalia: paramere arched inward apically and rounded; aedeagal ventral ramus short than aedeagal dorsal body with dilation below the apex; inner margin straight; outer margin convex; aedeagal dorsal body with two pair of apical lobes; outer lobe large with apex folded inward; inner lobe circular and setose. Apodeme extending beyond genital ring.</p><p>Remarks. This species is known from Mexico (Vera Cruz), Costa Rica and Panama (Azevedo 1999). Here more specimens are added. In this series the specimens are quite similar to those previously studied, without any variation that is worth to mention.</p><p>Material examined. Holotype, ♂ COSTA RICA , Puntarenas: Monteverde, 1400m, 10º43’N 84º02W, 12– 15.vi.1974, Malaise trap, W.A. Harding &amp; J. P Donahue col. (LACM) ; PANAMA, Bocas del Toro: 1♂, Wekso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Teribe</a>, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP) ; 1♂, Fortuna, Div. Cont. Chiriqui-B[ocas del] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.19092&amp;materialsCitation.latitude=8.77125" title="Search Plazi for locations around (long -82.19092/lat 8.77125)">Toro</a>, 1070–1090 m, [08°46'16.5"N, 82°11'27.3"W], 9–11.abr.1999, Tr [ampas] Amar., R. Cambra, A. Santos [col.] (MIUP) ; Chiriqui: 5♂, Finca La Suiza near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 25–6.VI.2000, F[light] I[nterception] T[rap], H. &amp; A. Howden [col.] (CNCI) ; Colón: 1♂, 14 km N J[un]ct[ion] Escobal r[oa]d and Piña r[oa]d on <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.9&amp;materialsCitation.latitude=9.333333" title="Search Plazi for locations around (long -79.9/lat 9.333333)">Piña</a> r[oa]d, ± 20 m, [09°20'N, 79°54'W], 2–3.VI.1995, F[light] I[nterception] T[rap], J. Ashe &amp; R. Brooks [col.] (CNCI); [Panama] : 1♂, C[anal] Z[one], B[arro] Colorado Is[land], [09°09'N, 79°51'W], 15–31.VII.1994, F[light] I[nterception] T[rap], D. Banks [col.] (CNCI); Darién Pr [ovincia]: 20♂, P[arque] N[acional] Darién, Pirre, Est[ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 77°45'W], 16 nov–17 ene 2001, Malaise, R. Cambra, A. Santos [col.] and 450m, 24–26 marzo 2001, col. A. Santos (2♂) (MIUP);</p><p>Distribution (Fig. 233). Mexico (Vera Cruz), Costa Rica and Panama.</p></div>	https://treatment.plazi.org/id/3B3687C07C449B13FF67F60B9CB8F832	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C449B13FF67F0E89BDEFC34.text	3B3687C07C449B13FF67F0E89BDEFC34.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus vallensis Evans. Eighteen 1979	<div><p>vallensis species-group</p><p>Diagnosis. Male. Clypeus with only median tooth. Tergal process median, depression transverse and elliptical. Pair of lateral tubercles, directed each other.</p><p>Remarks. It has 11 species: D. bisserratus Azevedo, D. polidentatus Azevedo, D. planus Azevedo, D. extrarramis Azevedo, D. triangularis Azevedo, D. dilatatus, D. strictus Azevedo, D. bifurcatus Azevedo, D. digitatus Azevedo, D. vallensis and D. rosangelae Colombo &amp; Azevedo.</p><p>Distribution. Ranging from Mexico to south of Brazil, including some Caribbean islands.</p></div>	https://treatment.plazi.org/id/3B3687C07C449B13FF67F0E89BDEFC34	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4B9B1CFF67F2A89D86FC63.text	3B3687C07C4B9B1CFF67F2A89D86FC63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus vallensis Evans. Eighteen 1979	<div><p>Dissomphalus vallensis Evans, 1979</p><p>(Figs 197–199)</p><p>Dissomphlaus vallensis Evans, 1979: 277, 281 (♂ holotype from USNM); Gordh &amp; Móczár, 1990: 226; Azevedo, 1999a: 327 – 328, 162–163.</p><p>Diagnosis. Mandible bidentate. Clypeus wih median lobe ill-defined. Tergal process median with large, elliptical and deep depression with pair of tubercles with tuft of setae directed each other. Hypopygium with posterior margin concave. Genitalia: paramere with apex truncate and oblique; aedeagal ventral ramus as long as aedeagal dorsal body; surface smooth; inner margin straight; outer margin convex; apical half narrow and divergent; aedeagal dorsal body with apical half arched downward; ventral side with median fold; dorsal margin of apical half arched outward and pair of very small lobes upward in base of aedeagal dorsal body. Apodeme not extending beyond genital ring.</p><p>Remarks. This species is known from Colombia (Azevedo 1999a) and is recorded for the first time from Panama. Some variations are worth to mention: median clypeal tooth angulate; median clypeal carina conspicuous and complete; vertex crest straight; anterior margin of pronotal disc coarse; aedeagal ventral ramus shorter than aedeagal dorsal body; median region of aedeagal dorsal body wide, apical portion elongated and narrow, outer margin sinuous; genital ring bifurcated distally in ventral view.</p><p>Material examined. Holotype ♂, COLOMBIA, Peñas Blancas: 10km, W Cali, Dept. Valle, 26–28 1975, Malaise trap, R. Wilkerson (FSCA) ; PANAMA, Coche {= Coclé}: 1♂, El Cope {= <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.58417&amp;materialsCitation.latitude=8.619722" title="Search Plazi for locations around (long -80.58417/lat 8.619722)">El Copé</a>}, 730 m, [08°37'11"N, 80°35'03"W], 19–20.XI.1994, F[light] I[nterception] T[rap], D. Windson Edwards [col.] (CNCI) ; Darién: 5♂, Cana, Serraria {= Serrania} de Pirre 1450m, 4–7.vi. 1995, J. Ashe, R. Brooks F[light] I[nterception] T[rap] (UFES).</p><p>Distribution (Fig. 244). Colombia and Panama.</p></div>	https://treatment.plazi.org/id/3B3687C07C4B9B1CFF67F2A89D86FC63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4B9B1DFF67F6699A09FCCA.text	3B3687C07C4B9B1DFF67F6699A09FCCA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus rettenmeyeri Evans 1964	<div><p>Dissomphalus rettenmeyeri Evans, 1964</p><p>(Figs 200–201)</p><p>Dissomphalus rettenmeyeri Evans, 1964: 45, 52, 53–55; (♂, holotype from Panama, USNM, figs 28–30); Gordh &amp; Móczár, 1990: 25; Azevedo, 2003: 70 –71.</p><p>Diagnosis. Mandible bidentate. Clypeus broadly projected forward, with small median tooth paralleled by small rounded expansion. Tergal process with pair of triangular and shallow lateral depressions, drop-shaped pit with tuft of very short setae. Genitalia: paramere unusually very wide, apex truncate; aedeagal ventral ramus short than aedeagal dorsal body, with arrow apical expansion; aedeagal dorsal body with two pairs of apical lobes; outer pair elongate dome-shaped lobe; inner pair membranous, stout and setose.</p><p>Remarks. This species is known only from the holotype and allotype from Panama. In this study, it was the most abundant species with 273 specimens and it has the largest distribution. All the specimens when compared with description by Evans (1964) and redescription by Azevedo (2003) have no variations that worth to mention.</p><p>Material examined. Holotype, ♂, PANAMA, Barro Colorado Island: Canal Zone, 26.II.1955, Berlese refuse, C.W Rettenmeyer col. (USNM #64996) . PANAMA, Bocas del Toro: 61♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Teribe</a>, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP) ; Chiriqui: 33#, Finca La Suiza near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 25–6.VI.2000, F[light] I[nterception] T[rap], H. &amp; A. Howden [col.] (CNCI) ; 2–6.VI.2000 (CNCI) (9♂) and (UFES) (2♂); 1♂, La Fortuna Dam, 1200 m wet forest 14–VI–16–VII–1982, B . Gill. (CNCI); Veraguas: 2♂, Pr [ovincia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.108444&amp;materialsCitation.latitude=8.518194" title="Search Plazi for locations around (long -81.108444/lat 8.518194)">Santa Fé Distri</a>], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.108444&amp;materialsCitation.latitude=8.518194" title="Search Plazi for locations around (long -81.108444/lat 8.518194)">Santa Fé</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.108444&amp;materialsCitation.latitude=8.518194" title="Search Plazi for locations around (long -81.108444/lat 8.518194)">Altos de Piedra</a>, 770 m, [08°31'5.5"N, 81°06'30.4"W], 22–26 marzo 1999, Malaise, Col. A. Santos, L . D. (1♂) and 13–17 nov. 1999, A. Santos, P. González [col.] (1♂) (MIUP) ; Colón Pr[ovincia]: 3♂, P[a]r[que] Nac [ional] Portobelo, San Antonio, 16–29 feb. 2000, Tr [ampa] Malaise, P. González, A. Santos [col.] (MIUP) ; 3♂, S[an]ta Rita [<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.750275&amp;materialsCitation.latitude=9.3324995" title="Search Plazi for locations around (long -79.750275/lat 9.3324995)">Arriba</a>], Macizo, Parador Sierra Llorona, [09°19'57"N, 79°45'1"W], 22 dic. 1997, D . Quintero [col.] (MIUP); 3♂, 14 km N J[un]ct[ion] Escobal r[oa]d and Piña r[oa]d on Piña r[oa]d, ± 20 m , [09°20'N', 79°54'W], 2–3.VI.1995, F[light] I[nterception] T[rap], J. Ashe &amp; R. Brooks [col.] (CNCI); Coclé Pr[ovincia]: 3♂, El Copé, Par [que] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">Nac</a> [ional <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">General de División</a>] Omar Torrijos H [errera], 570–670 m.s.n.m. [08º41'56"N, 80º39'17"W], 26–30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; Panama Pr[ovincia]: 6♂, P[arque] Nac [ional] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.71667&amp;materialsCitation.latitude=9.15" title="Search Plazi for locations around (long -79.71667/lat 9.15)">Sob</a> [eranía], Camino, Plantación, [09º09'N, 79º43'W], 24 ago.–2 sep.1999, Malaise, P. González, A. Santos [col.] (MIUP) ; 2♂, P[arque] Nac [ional] Altos de Campana, [08º43'15"W, 80º01'43"W], 10–14 ago 1999 Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; 1♂, Chilibre, Parque Nacional Chagres, Campo Chagres, [09º25'54"N, 79º30'54"W], 24–28.enero.2000 Tr[ampa] Malaise, A. Santos, P. González [col.] (MIUP); 1♂, P[arque] N[atural] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.54472&amp;materialsCitation.latitude=8.9925" title="Search Plazi for locations around (long -79.54472/lat 8.9925)">Metropolitano</a>, [08º59'33"N, 79º32'41"W], 24 ago.–3 sep.1999 Tr [ampa] Malaise, A. Santos, P. González [col] (MIUP) ; 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.833336&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -79.833336/lat 8.666667)">Cerro Campana</a>, 08°40'N, 79°50'W, 7–14.V.1973 (CNCI) ; 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.85&amp;materialsCitation.latitude=9.15" title="Search Plazi for locations around (long -79.85/lat 9.15)">Barro Colorado</a> I[sland], [09°09'N, 79°51'W], 23.III.1981, R. B. &amp; L. S. Kimsey [col.] (UCDC) ; 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.0&amp;materialsCitation.latitude=9.416667" title="Search Plazi for locations around (long -80.0/lat 9.416667)">Old Gamboa</a>, C[anal] Z[one], [09°25'N, 80°00'W], 14.XI.1994, F[light] I[nterception] T[rap], P. Windson [col.] (CNCI) ; 5♂, C[anal] Z[one], B[arro] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.85&amp;materialsCitation.latitude=9.15" title="Search Plazi for locations around (long -79.85/lat 9.15)">Colorado Is</a> [land], [09°09'N, 79°51'W], 15–31.VII.1994 (1♂) , 14.VII.1994 (3♂) D. Banks [col.] and 3–13– VI–1983, B Gill (1♂) (CNCI) ; Darién Pr[ovincia]: 134♂, P[arque] N[acional] Darién, Pirre, Est [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 77°45'W], 16 nov 00 – 17 ene 2001, Malaise (131♂), 18–24 ene 2001 Tr[ampa] Amarilla (1♂) R. Cambra, A. Santos [col.], 450m, 24–26 marzo 2001, Malaise, col. A. Santos (1♂) and 560m, 21 mar–4 ab 2000, Cambra, Santos, Bermudez [col.] (1♂) (MIUP); 5♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.61667&amp;materialsCitation.latitude=7.85" title="Search Plazi for locations around (long -77.61667/lat 7.85)">Cana</a>, 1380 m, [07°51'N, 77°37'W], 4–7.VI.1986, F[light] I[nterception] T[rap], [J.] Ashe &amp; [R.] Brooks [col.] (CNCI) ; 7♂, Serraria {Serrania} de Pirre, 1450m, 4– 7.vi.1995, J. Ashe, R. Brooks F[light] I[nterception] T[rap] (6♂) (UFES) and 1456 m, 7–9.VI.1996 (1♂) (CNCI).</p><p>Distribution (Fig. 249). Panama (Bocas del Toro, Chiriquí, Veraguas, Colón, Coclé, Panamá and Darién).</p></div>	https://treatment.plazi.org/id/3B3687C07C4B9B1DFF67F6699A09FCCA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4A9B1DFF67F1589CA3F8FD.text	3B3687C07C4A9B1DFF67F1589CA3F8FD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus personatus Redighieri & Azevedo 2006	<div><p>Dissomphalus personatus Redighieri &amp; Azevedo, 2006</p><p>(Figs 202–206)</p><p>Dissomphalus personatus Redighieri &amp; Azevedo, 2006: 327 (♂ holotype from Paraná MZSP, figs 43–44).</p><p>Diagnosis. Mandible tetradentate. Clypeus subtrapezoidal. Tergal process with lateral, shallow and subcircular depressions, with small tubercles and tuft of setae. Hypopygium with posterior margin concave. Genitalia: paramere acute with dorsal and ventral margins straight; aedeagal ventral ramus longer than aedeagal dorsal body, laminar; surface horizontal; basal half wide and fused; apical half with two laminas narrowing at apex; aedeagal dorsal body with two pairs of apical lobes; outer pair laminar; surface vertical; apex acute with horizontal projection masked-shaped; inner pair membranous, setose and longer than outer lobe.</p><p>Remarks. This species is known to Brazil (São Paulo and Paraná) (Redighieri &amp; Azevedo 2006). Now it is recorded for the first time from Panama. Some variations in this series worth to mention: head, mesosoma and metasoma castaneous; aedeagal ventral ramus shorter than aedeagal dorsal body; apical lobes of aedeagal dorsal body with outer margin angulate and inner margin very excavated in some specimens; outer margin of apical lobes of aedeagal dorsal body subquadrate and inner margin almost straight in some specimens.</p><p>Material examined. Holotye, ♂, BRAZIL, Paraná: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.8925&amp;materialsCitation.latitude=-25.574306" title="Search Plazi for locations around (long -48.8925/lat -25.574306)">Morretes</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.8925&amp;materialsCitation.latitude=-25.574306" title="Search Plazi for locations around (long -48.8925/lat -25.574306)">Parque Estadual do Pau Oco</a>, 25º34’27.5”S 48º53’33”W, 10–13.iv.2002, armadilha Malaise, M.T. Tavares e eq.col. (MZSP) ; PANAMA, Bocas del Toro: 1♂, Wekso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Teribe</a>, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP) ; [Chiriquí]: 1♂, Fortuna, Div. Cont. Chiriqui-B[ocas del] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.19092&amp;materialsCitation.latitude=8.77125" title="Search Plazi for locations around (long -82.19092/lat 8.77125)">Toro</a>, 1070– 1090 m, [08°46'16.5"N, 82°11'27.3"W], 8–11.abr.1999, Malaise, R. Cambra, A. Santos [col.] (MIUP) ; Coclé Pr [ovincia]: 3♂, El Copé, Par [que] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">Nac</a> [ional <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">General de División</a>] Omar Torrijos H [errera], 570–670 m.s.n.m., [08º41'56"N, 80º39'17"W], 26–30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; Darién Pr [ovincia]: 4♂, P[arque] N[acional] Darién, Pirre, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Est</a> [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 900 m, [08°00'N, 77°45'W], 21 mar– 4 ab 2000, Malaise, Cambra, Santos, Bermudez [col.] (1♂) , 80 m, 16 nov–17ene 2001, R. Cambra, A. Santos [col.] (3♂) (MIUP); 1♂, Cana, Serrania de Pirre, 1250m, 4–7.vi.1996, J. Ashe, R. Brooks (UFES) .</p><p>Distribution (Fig. 239). Panama and Brazil (São Paulo and Paraná).</p></div>	https://treatment.plazi.org/id/3B3687C07C4A9B1DFF67F1589CA3F8FD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C499B1FFF67F53E9DD0FC7A.text	3B3687C07C499B1FFF67F53E9DD0FC7A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus abarus Azevedo 2017	<div><p>Dissomphalus abarus Brito &amp; Azevedo sp. nov.</p><p>(Figs 25, 56, 87, 207–208)</p><p>Description. Male. Body length 4.54 mm. Color: body dark castaneous. Head (Fig 25). Mandible with three distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, incomplete apically, straight in profile or nearly so. Frons strongly coriaceous and punctures small. Vertex crest straight. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 56). Tergal process with lateral, shallow, subcircular and large depression, without tubercle, with short and dense tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 87). Genitalia (Figs 207–208): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin sinuous; apex short, translucent, with two small projections, slightly curved outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and wide, with apical margin rounded and slightly mesad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Variations. Head, metasoma and mesosoma castaneous; median clypeal lobe subtrapezoidal; pit of tergal process with few setae.</p><p>Remarks. This species is easily recognized by having the genitalia robust, the aedeagal ventral ramus very wide with the apex translucent with two small projections. This species is similar to D. iozus sp. nov. by having the aedeagal dorsal body robust and the aedeagal ventral ramus very wide. However, this species has the paramere, basiparamere and the apex of apical lobes of the aedeagal dorsal body narrow and the apex of the aedeagal ventral ramus translucent with small projections whereas, D. iozus sp. nov. has the paramere and the basiparamere wide, the apex of the aedeagal dorsal body wide and rounded and the apex of the aedeagal ventral ramus simple and not translucent.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.78833&amp;materialsCitation.latitude=8.911667" title="Search Plazi for locations around (long -82.78833/lat 8.911667)">Material</a> examined. Holotype ♂, PANAMA, Chiriquí: 08° 54,7' N 82° 47,3'W 17–18.vi. 1996, 1875m, J. Ashe, R. Brooks F[light] I[interception] T[trap] (UFES) . Paratypes: 3♂, same data as holotype; 1♂ 2 km W <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.583336&amp;materialsCitation.latitude=8.85" title="Search Plazi for locations around (long -82.583336/lat 8.85)">Cerro Punta</a>, 1700 m, [08°51'N, 82°35'W], 19.V–8.VI.1977, Peck &amp; Howden [col.] (CNCI) ; 1♂, P[arque] I[nternacional] L[a] A[mistad], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.73333&amp;materialsCitation.latitude=8.9" title="Search Plazi for locations around (long -82.73333/lat 8.9)">Jurutungo</a>, 1800 m, [08°54'N, 82°44'W], 30oci. 5nov.1994, Tr [ampa] Malaise, A. Rodriguez [col.] (MIUP) .</p><p>Distribution (Fig. 225). Panama (Chiriquí).</p></div>	https://treatment.plazi.org/id/3B3687C07C499B1FFF67F53E9DD0FC7A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C489B1FFF67F1C89DC1F894.text	3B3687C07C489B1FFF67F1C89DC1F894.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus arteobius Azevedo 2017	<div><p>Dissomphalus arteobius Brito &amp; Azevedo, sp. nov.</p><p>(Figs 26, 57, 88, 209–210)</p><p>Description. Male. Body length 3.13 mm. Color: head dark castaneous; mesosoma and metasoma castaneous. Head (Fig. 26). Mandible with two distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina low in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures large. Vertex crest straight. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 57). Tergal proces with lateral, shallow, subcircular and small depression, without tubercle, with very short tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 88). Genitalia (Figs 209–210): paramere with dorsal margin entirely wide; apical margin rounded. Aedeagal ventral ramus shorter than dorsal body, evenly narrow; cross section laminar; surface subvertical; inner margin concave; outer margin sinuous; apex short, simple, parallel; additional inner ramus absent; basal stub short and very narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and narrow, with apical margin abruptly acute and ventrad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Variations. Mesosoma dark castaneous; median clypeal tooth shorter; depression of tergal process very shallow.</p><p>Remarks. This species is easily recognized by having the very long basiparamere about 2 × the size of the paramere. This characteristic does not have in other species in Dissomphalus . The paramere is wide and short, the apex of the apical lobe of the aedeagal dorsal body is articulated and the eyes are bulging.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Material</a> examined. Holotype, ♂, PANAMA, Darién Pr [ovincia]: P[arque] N[acional] Darién, Pirre, Est[ación] Rancho Frio, 80 m, [08°00'N, 77°45'W], 16 nov–17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP) . Paratypes: 2♂, same data as holotype; 3♂, same of holotype except 1000m, 21 mar–4 ab 2000 (2♂), 580m, 21 mar–4 ab 2000 (1♂) (MIUP).</p><p>Distribution (Fig. 227). Panama (Darién).</p></div>	https://treatment.plazi.org/id/3B3687C07C489B1FFF67F1C89DC1F894	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4F9B18FF67F2A89B6DFAF0.text	3B3687C07C4F9B18FF67F2A89B6DFAF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus borus Azevedo 2017	<div><p>Dissomphalus borus Brito &amp; Azevedo, sp. nov.</p><p>(Figs 27, 58, 89, 211–212)</p><p>Description. Male. Body length 1.73 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 27). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one rounded tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 58). Tergal process lateral, without depression, with very small, low and dorsad tubercle, pit very small, with short tuft of setae. Posterior hypopygeal margin sinuous (Fig. 89). Genitalia (Figs 211–212): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus higher than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin sinuous; apex long, simple, parallel; additional inner ramus absent; basal stub very short and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe very short, horizontal and narrow, with apical margin abruptly acute and slightly mesad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Variation. Pit of tubercle of tergal process without setae.</p><p>Remarks. This species is easily recognized by having the dorsal margin of the paramere straight and the apex very excavated in the apical region and the apical lobes of the aedeagal dorsal body very short. This species is similar to D. azagus sp. nov. by having the dorsal margin of the paramere straight and the apex very excavated. However, this species has the tergal process present, the apical lobes of the aedeagal dorsal body very short and the aedeagal ventral ramus progressively narrowing apicad, whereas D. azagus sp. nov has the tergal process absent, the apical lobes of the aedeagal dorsal body long and the apex of the aedeagal ventral ramus narrowing abruptly apicad.</p><p>Material examined. Holotype, ♂, PANAMA, Veraguas Pr [ovincia]: P[arque] N[acional] Coiba, Isla Coiba, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Cerro la Equis</a>, 120 m, [07º28'09"N, 81º45'24"W], 29 jul–4 ag 1998, Malaise, R. Cambra, S. Bermudez [col.] (MIUP) . Paratypes: 1♂ same as holotype except 30–nov.–4dic.1998 (MIUP); 1♂ same as holotype except 200m (MIUP); 1♂, Santa Fé, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.108444&amp;materialsCitation.latitude=8.518194" title="Search Plazi for locations around (long -81.108444/lat 8.518194)">Altos de Piedra</a>, 770 m [08°31'5.5"N, 81°06'30.4"W], 22–26 marzo 1999, Col. A. Santos, L. D. (MIUP) ; [Chiriqui]: 1♂, Formosa, Finca La Suiza 1 Km se. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 28.V–2.VI.1994, Malaise Trap, F. Andrews &amp; A. Gilbert [col.] (UCDC) ; Coclé Pr [ovincia]: 4♂, El Coclé, Par [que] Nac [ional <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">General de División</a>] Omar Torrijos H [errera], 570–670 m.s.n.m.[08º41'56"N, 80º39'17"W], 26–30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; [Panama Province]: 1♂, C[anal] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.85&amp;materialsCitation.latitude=9.15" title="Search Plazi for locations around (long -79.85/lat 9.15)">Zone Barro Colorado</a> I[sland], [09°09'N, 79°51'W], V.11.1981, R. B. &amp; L. S. Kimsey [col.] (UCDC) .</p><p>Distribution (Fig. 232). Panama (Chiriqui, Veraguas, Coclé and Panama).</p></div>	https://treatment.plazi.org/id/3B3687C07C4F9B18FF67F2A89B6DFAF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4F9B19FF67F7429DDBFEBB.text	3B3687C07C4F9B19FF67F7429DDBFEBB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus geortus Azevedo 2017	<div><p>Dissomphalus geortus Brito &amp; Azevedo sp. nov.</p><p>(Figs 28, 59, 90, 213–214)</p><p>Description. Male. Body length 1.16 mm. Color: head and metasoma castaneous; metasoma light castaneous. Head (Fig. 28). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one rounded tooth; median clypeal carina high in profile, complete or nearly so, angled in profile. Frons weakly coriaceous and punctures small. Vertex crest strongly convex. Pronotal disc polished; anterior margin ecarinate. Metasoma (Fig. 59). Tergal process lateral, without depression, low and dorsad tubercle, pit small, with long and dense tuft of setae. Posterior hypopygeal margin sinuous (Fig. 90). Genitalia (Figs 213–214): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus higher than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin sinuous; apex short, simple, slightly curved outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and narrow, with apical margin rounded and posterad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Remarks. This species is similar to D. krombeini by having the paramere expanded basally and the aedeagal ventral ramus abruptly narrowing apicad. However, this species has the tergal process with tubercle with tuft of setae and the apical lobes of the aedeagal dorsal body longer, whereas the D. krombeini has the apical lobe of the aedeagal dorsal body shorter and the tergal process linear with very short tuft of setae with one distincty long seta.</p><p>Material examined. Holotype, ♂, PANAMA, Veraguas: P[arque] N[acional] Coiba, Isla Coiba, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Cerro la Equis</a>, 90 m, [07º28'09"N, 81º45'24"W], 30–nov–4–dic.1998, Malaise, R. Cambra, S. Bermudez [col.] (MIUP). Paratypes: 3♂, same data as holotype ; 5♂ same as holotype except, 120 m (2♂), 200m (1♂) (MIUP); 3♂ same as holotype except 90 m, 8–12mar [zo] 1998 (1♂), 270 m, 7–13mar 1998, R. Cambra, A. Santos [col.] (1♂), 29 jul–4 ago 1998 (1♂) (MIUP).</p><p>Distribution (Fig. 234). Panama (Veraguas).</p></div>	https://treatment.plazi.org/id/3B3687C07C4F9B19FF67F7429DDBFEBB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4D9B1BFF67F6C99C99FD12.text	3B3687C07C4D9B1BFF67F6C99C99FD12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus ilarius Azevedo 2017	<div><p>Dissomphalus ilarius Brito &amp; Azevedo sp. nov.</p><p>(Figs 29, 60, 91, 215–216)</p><p>Description. Male. Body length 2.18 mm. Color: head and mesosoma castaneous; metasoma light castaneous. Head (Fig. 29). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one angulate very short tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 60). Tergal process lateral, without depression, with small, very low and dorsad tubercle, pit absent, with short and sparse tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 91). Genitalia (Figs 215–216): paramere with dorsal margin wide basally; apical margin rounded. Aedeagal ventral ramus shorter than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin sinuous; apex long, simple, parallel; additional inner ramus absent; basal stub very short and very narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, horizontal and wide, with apical margin abruptly acute and posterad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Variations. Head dark castaneous; median clypeal tooth inconspicuous; pit of tubercle of tergal process with short setae.</p><p>Remarks. This species is easily recognized by having the paramere very expanded basally and four narrow projections that connect the apical lobes of the aedeagal dorsal body with the median portion of the aedeagal dorsal body and the tergal process with one minute tubercle without pit with few setae. This species is similar to the D. microtuberculatus, Azevedo by having the tergal process with two minutes tubercles. However, this species has the basiparamere shorter than the paramere, the apical lobes of the aedeagal dorsal body without flagella and the apex of the aedeagal ventral ramus progressively narrowing apicad, whereas D. microtuberculatus has the basiparamere longer than the paramere, the apical lobes of the aedeagal dorsal body with flagella and the apex of the aedeagal ventral ramus abruptly acute.</p><p>Material examined. Holotype, ♂, PANAMA, Coclé Pr [ovincia]: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">El Copé</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">Par</a> [que] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">Nac</a> [ional <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">General de División</a>] Omar Torrijos H [errera], 570–670 m.s.n.m., [08º41'56"N, 80º39'17"W], 26–30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) . Paratypes: Chiriqui: 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.24944&amp;materialsCitation.latitude=8.744167" title="Search Plazi for locations around (long -82.24944/lat 8.744167)">La</a> Fortuna Dam, 1200 m, wet forest, [08°44'39"N, 82°14'58"W], 14–VI–16.VII–1982, F[light] I[nterception] T[rap], B. Gill [col.] (PMAE) ; 1♂, Fortuna, Div. Cont. Chiriqui-B[ocas del] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.19092&amp;materialsCitation.latitude=8.77125" title="Search Plazi for locations around (long -82.19092/lat 8.77125)">Toro</a>, 1070–1090 m, [08°46'16.5"N, 82°11'27.3"W], 8–11.abr.1999, Malaise, R. Cambra, A. Santos [col.] (MIUP) ; Veraguas Pr [ovincia]: 9♂ P[arque] N[acional] Coiba, Isla Coiba, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.75667&amp;materialsCitation.latitude=7.4691668" title="Search Plazi for locations around (long -81.75667/lat 7.4691668)">Cerro La Equis</a>, 90 m (8♂) and 200m (1♂) [07º28'09"N, 81º45'24"W], 30–nov.–4dic.1998, Malaise, R. Cambra, S. Bermudez [col.] (MIUP) ; 1♂, Pr [ovincia] Santa Fé, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.108444&amp;materialsCitation.latitude=8.518194" title="Search Plazi for locations around (long -81.108444/lat 8.518194)">Altos De Piedra</a>, [08°31'5.5"N, 81°06'30.4"W], 13–17 nov. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; 1♂, [Rio] Calovébora, [<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.21667&amp;materialsCitation.latitude=8.7" title="Search Plazi for locations around (long -81.21667/lat 8.7)">Rio</a>] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.21667&amp;materialsCitation.latitude=8.7" title="Search Plazi for locations around (long -81.21667/lat 8.7)">Guazarito</a>, [08º42'N, 81º13'W], 15–18 feb. 2000, Col. S. Bermudez (MIUP) ; 1♂ same data as holotype; Darién: 2♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-87.61667&amp;materialsCitation.latitude=7.85" title="Search Plazi for locations around (long -87.61667/lat 7.85)">Cana</a>, 600 m, [07°51'N, 87°37'W], 3–7.VI.1996, J. Ashe &amp; R. Brooks [col.] (CNCI);</p><p>Distribution (Fig. 236). Panama (Veraguas, Darién, Coclé and Chiriquí).</p></div>	https://treatment.plazi.org/id/3B3687C07C4D9B1BFF67F6C99C99FD12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4C9B1BFF67F1209C79F8F8.text	3B3687C07C4C9B1BFF67F1209C79F8F8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus isortus Azevedo 2017	<div><p>Dissomphalus isortus Brito &amp; Azevedo sp. nov.</p><p>(Figs 30, 61, 92, 217–218)</p><p>Description. Male. Body length 2.42 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 30). Mandible with two distal teeth. Median clypeal lobe subtrapezoidal, with one angulate tooth; median clypeal carina high in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin coarse. Metasoma (Fig. 61). Tergal process lateral, with deep, circular and large depression, with large, high and dorsad tubercle placed on center of depression, pit large, with short and dense tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 92). Genitalia (Figs 217–218): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus as long as dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin slightly sinuous; outer margin sinuous; apex long, simple, curved outward; additional inner ramus absent; basal stub short and very narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, subvertical and narrow, with apical margin rounded and posterad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Variation. Depression of tergal process shallow.</p><p>Remarks. This species is easily recognized by having the paramere very wide, the apical lobes of the aedeagal dorsal body conical-shaped, the cuspis very small and the aedeagal ventral ramus wide basally. This species is similar to D. raziogus sp. nov. by having the aedeagal dorsal body narrowing slightly apicad and the apical lobes of the aedeagal dorsal body in conical-shaped. However, this species has the tergal process present, the paramere very wide and the apex of apical lobes of the aedeagal dorsal body dorsad, whereas D. raziogus sp. nov. has the tergal process absent, the paramere not wide and the apex of apical lobes of the aedeagal dorsal body mesad.</p><p>Material examined. Holotype, ♂, PANAMA, Coclé Pr [ovincia]: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">El Copé</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">Par</a> [que] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">Nac</a> [ional <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">General de División</a>] Omar Torrijos H [errera], 570–670 m.s.n.m., [08º41'56"N, 80º39'17"W], 26–30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) . Paratypes: 5♂, same data as holotype; Darién Pr [ovincia]: 1♂, P[arque] N[acional] Darién, Pirre, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Est</a> [ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 1000 m, [08°00'N, 77°45'W], 21mar–4 ab 2000, Malaise, Cambra, Santos, Bermudez [col.] (MIUP) ; 2♂, Cana, Serraria {Serrania} de Pirre, 1450 m, 4–7.vi.1995 F[light] I[interception] T[rap], J Ashe, R. Brooks (UFES).</p><p>Distribution (Fig. 238). Panama (Coclé and Darién).</p></div>	https://treatment.plazi.org/id/3B3687C07C4C9B1BFF67F1209C79F8F8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4C9B64FF67F5509C02FC17.text	3B3687C07C4C9B64FF67F5509C02FC17.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus trisus Azevedo 2017	<div><p>Dissomphalus trisus Brito &amp; Azevedo, sp. nov.</p><p>(Figs 31, 62, 93, 219–220)</p><p>Description. Male. Body length 5.16 mm. Color: head and mesosoma dark castaneous; metasoma castaneous.</p><p>Head (Fig. 31). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina very high in profile, complete or nearly so, convex in profile. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc weakly coriaceous; anterior margin coarse. Metasoma (Fig. 62). Tergal process median, without depression, with small, low and dorsad tubercle, pit small, with very short and dense tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 93). Genitalia (Figs 219–220): paramere with dorsal margin entirely wide; apical margin rounded. Aedeagal ventral ramus higher than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface subhorizontal; inner margin straight; outer margin sinuous; apex long, simple, slightly curved outward, additional inner ramus absent, basal stub long and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and narrow, with apical margin abruptly acute and ventrad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.</p><p>Variations. Body length 3.88 mm (min.) 5.16 mm (max.).</p><p>Remarks. This species is easily recognized by having the aedeagal ventral ramus with median half wide and apical half narrow and parallel. The tergal process median, characterized by having two small tubercles without depressions and small tuft of setae. This species is similar to D. scamatus Azevedo by having the median tergal process, without depression, with tubercle, tuft of setae and the basiparamere narrow. However, this species has the apical lobes of the aedeagal dorsal body long with acute apex and aedeagal ventral ramus with inner margin straight, abruptly narrowing apicad, whereas D. scamatus has the apical lobes of the aedeagal dorsal body short, narrowing at apex, aedeagal ventral ramus with inner margin concave, with acute apex.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.78833&amp;materialsCitation.latitude=8.911667" title="Search Plazi for locations around (long -82.78833/lat 8.911667)">Material</a> examined. Holotype, ♂, PANAMA, Chiriquí: 08°54,7'N, 82° 47,3'W 17–18.vi. 1996, 1875 m, F[light] I[interception] T[rap], J. Ashe, R. Brooks (UFES) . Paratypes: 1♂, same data as holotype; 1♂, Finca La Suiza near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.23333&amp;materialsCitation.latitude=8.666667" title="Search Plazi for locations around (long -82.23333/lat 8.666667)">Hornitos</a>, [08°40'N, 82°14'W], 25–6.VI.2000, H. &amp; A. Howden [col.] (CNCI) ; [Los Santos]: 3♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.416664&amp;materialsCitation.latitude=7.5166664" title="Search Plazi for locations around (long -80.416664/lat 7.5166664)">Ojo de Agua</a>, [07°31'N, 80°25'W], 30.VI–14.VII.1982, B. Gill [col.] (PMAE) .</p><p>Distribution (Fig. 245). Panama (Chiriquí and Los Santos).</p></div>	https://treatment.plazi.org/id/3B3687C07C4C9B64FF67F5509C02FC17	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C339B65FF67F6239E38FC5F.text	3B3687C07C339B65FF67F6239E38FC5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus ubracus Azevedo 2017	<div><p>Dissomphalus ubracus Brito &amp; Azevedo sp. nov.</p><p>(Figs 32, 63, 94, 221–222)</p><p>Description. Male. Body length 1.61 mm. Color: body castaneous. Head (Fig. 32). Mandible with three distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina high in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 63). Tergal process lateral, with deeply excaveted, subcircular and large depression covered by a membrane, without tubercle, pit absent. Posterior hypopygeal margin strongly concave (Fig. 94). Genitalia (Figs 221–222): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin slightly sinuous; apex short, simple, slightly curved outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, horizontal and wide, with apical margin abruptly acute and dorsad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring.</p><p>Variations. Mesosoma light castaneous; metasoma castaneous; tergal process more laterad and longer.</p><p>Remarks. This species is easily recognized by having the median clypeal carina with angled elevation, the paramere very excavated and the tergal process with deeply excaveted, subcircular and large depression covered by a membrane. This kind tergal process has been never found in Dissomphalus and it is easily recognized in the genus. This species is similar to the D. gilvipes by having the paramere wide basally and the aedeagal ventral ramus short with acute apex. However, this species has the apical margin of the paramere crenulate, the apical lobes of the aedeagal dorsal body wide and horizontal and the tergal process with very excavated depression covered by a membrane, whereas D. gilvipes has the paramere smooth, the apical lobes of the aedeagal dorsal body vertical and narrow and the tergal process with a tuft of setae with distinct long lateral setae.</p><p>Material examined. Holotype, ♂, PANAMA, Darién Pr [ovincia]: P[arque] N[acional] Darién, Pirre, Est[ación] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=8.0" title="Search Plazi for locations around (long -77.75/lat 8.0)">Rancho Frio</a>, 80 m, [08°00'N, 77°45'W], 16 nov–17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP) . Paratypes: Bocas del Toro: 1♂, Wekso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.93889&amp;materialsCitation.latitude=9.406944" title="Search Plazi for locations around (long -82.93889/lat 9.406944)">Teribe</a>, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [09º24'25"N, 82º56'20"W], 17–24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP) ; [Chiriquí]: 2♂, Fortuna, Div. Cont. Chiriqui-B[ocas del] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-82.19092&amp;materialsCitation.latitude=8.77125" title="Search Plazi for locations around (long -82.19092/lat 8.77125)">Toro</a>, 1070–1090 m, [08°46'16.5"N, 82°11'27.3"W], 8–11.abr.1999, R. Cambra, A. Santos [col.] (MIUP) ; Veraguas Pr [ovincia]: 1♂, Calovebora, Guazarito, 15–18 feb. 2000, malaise, Col. S. Bermudez (MIUP); 1♂, Santa Fé, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.108444&amp;materialsCitation.latitude=8.518194" title="Search Plazi for locations around (long -81.108444/lat 8.518194)">Altos de Piedra</a>, 770 m, [08°31'5.5"N, 81°06'30.4"W], 22–26 marzo 1999, Col. A. Santos, L. D. (MIUP) ; Colón: 5♂, 14 km N J[un]ct[ion] Escobal r[oa]d and Piña r[oa]d on <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.9&amp;materialsCitation.latitude=9.333333" title="Search Plazi for locations around (long -79.9/lat 9.333333)">Piña</a> r[oa]d, ± 20 m, [09°20'N', 79°54'W], 2–3.VI.1995, F[light] I[nterception] T[rap], J. Ashe &amp; R. Brooks [col.] (CNCI) ; Coche {= Coclé}: 1♂, El Cope {= <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.58417&amp;materialsCitation.latitude=8.619722" title="Search Plazi for locations around (long -80.58417/lat 8.619722)">El Copé</a>}, [08°37'11"N, 80°35'03"W], 19.XI.1994, Windson &amp; Edwards [col.] (CNCI) ; 6♂, Par [que] Nac [ional <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">General de División</a>] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.654724&amp;materialsCitation.latitude=8.698889" title="Search Plazi for locations around (long -80.654724/lat 8.698889)">Omar Torrijos</a> H[errera], 570– 670 m.s.n.m., [08º41'56"N, 80º39'17"W], 26–30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP) ; Panama Pr [ovincia]: 14♂, P[arque] Nac [ional] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.71667&amp;materialsCitation.latitude=9.15" title="Search Plazi for locations around (long -79.71667/lat 9.15)">Sob</a> [eranía], Camino, Plantación, [09º09'N, 79º43'W], 24 ago.–2 sep.1999, Malaise, A. Santos, P. González [col.] (MIUP) ; 5♂, P[arque] Nac [ional] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.02862&amp;materialsCitation.latitude=8.720833" title="Search Plazi for locations around (long -80.02862/lat 8.720833)">Altos de Campana</a>, [08º43'15"N, 80º01'43"W], 10–14 ago. 1999 (MIUP) ; 1♂, P[arque] N[atural] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.54472&amp;materialsCitation.latitude=8.9925" title="Search Plazi for locations around (long -79.54472/lat 8.9925)">Metropolitano</a>, [08º59'33"N, 79º32'41"W], 24 ago.–3 sep.1999 (MIUP) ; 1♂, Cerro Azul, Las Nubes, [09º10'N, 79º25'W], 9–15.oct.1999, R. Cambra, A. Santos [col.] (MIUP); 7♂, C[anal] Z[one], B[arro] Colorado Is[land], [09°09'N, 79°51'W], 15–31.VII.1994 (5♂), 1–14.VII.1994 (1♂) and 16– 24.VIII.1994 (1♂) F[light] I[nterception] T[rap], D. Banks [col.] (CNCI); Comarca Kuna Yala: 1♂, Ustupu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.92667&amp;materialsCitation.latitude=9.1325" title="Search Plazi for locations around (long -77.92667/lat 9.1325)">Rio Abudi</a>, [09º07'57"N, 77º55'36"W], 6–12 dic. 1999, Malaise, P. González [col.] (MIUP) ; 12♂, same data as holotype; 1♂, same as holotype except 1000 m, 21 mar–4 ab 2000, Cambra, Santos, Bermudez [col.] (MIUP); 1♂, same as holotype except 450m, 24–26 marzo 2001, col. A. Santos (MIUP); 19♂, Darién, Cruce de Mono, Est [ación] I[nstituto] N[acional de] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Re</a> [cusos] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Na</a> [turales] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Re</a> [novables], P[arque] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.47019&amp;materialsCitation.latitude=7.8573055" title="Search Plazi for locations around (long -77.47019/lat 7.8573055)">Nac</a> [ional] Darién, [07°51'26.3"N, 77°28'12.7"W], 5–28 feb 1993, R. Cambra [col.] (MIUP) ;</p><p>Distribution (Fig. 248). Panama (Bocas del Toro, Chiriquí, Veraguas, Coclé, Colón, Panama, Comarca Kuna Yala and Darién).</p></div>	https://treatment.plazi.org/id/3B3687C07C339B65FF67F6239E38FC5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Azevedo, Celso O.	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
