taxonID	type	description	language	source
3B3687C07C769B22FF67F7DE9E5DFEE0.taxon	description	Glenobethylus Kieffer, 1910 a: 50. Type-species: Glenobethylus montanus Kieffer, by monotypy. Synonymized by Evans, 1964: 41.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C769B22FF67F7DE9E5DFEE0.taxon	diagnosis	Diagnosis. Male. Body robust. Clypeus prominent, median lobe sometimes ill-defined. Metasoma relatively wide and short; second tergo sometimes with tergal process (depressions, pits, tufts of setae, tubercles). Hypopygium with three stalks. Genitalia wide; paramere simple or bilobate; aedeagus complex, composed by ventral ramus and dorsal body. Female. Head longer than wide. Eye small, with from one to about 25 facets. Mesonotum short, transverse, posterior margin gently convex at broad junction with propodeum; anterior margin of propodeum not embracing sides of mesonotum to any appreciable extent. Genitalia with aedeagus divided into dorsal body and ventral ramus.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7A9B2DFF67F7F29A24F962.taxon	diagnosis	Diagnosis. Male. Metasoma with median depression, dense tufts of setae. Digitus serrated with rounded teeth inner upper margin. Aedeagal ventral ramus wide.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7A9B2DFF67F7F29A24F962.taxon	discussion	Remarks. It has six species: D. altivolans Evans, D. deformis Evans, D. megomphalus Evans, D. subdeformis Azevedo, D. lanceolatus Azevedo and D. isteus sp. nov..	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7A9B2DFF67F7F29A24F962.taxon	distribution	Distribution. Ranging from northern U. S. A. to north of Argentina, including some Caribbean islands.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7A9B2FFF67F5309EFFFF28.taxon	diagnosis	Diagnosis. Mandible tridentate. Clypeus tridentate. Tergal process with median concavity not so large and deep, with pair of lateral subcircular tufts densely covered by short setae. Hypopygium with posterior margin concave, corners rounded. Genitalia: paramere with apex rounded; aedeagal ventral ramus shorter than aedeagal dorsal body, inner margin smooth, parallel and straight; outer margin smooth, slightly convergent; surfaces horizontal; apex rounded, basal margin with median invagination; aedeagal dorsal body with two pairs of apical lobes; outer lobe wide, with rounded apex, inner margin concave, convergent dorsally; inner lobe membranous and setose. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7A9B2FFF67F5309EFFFF28.taxon	discussion	Remarks. No additional material of this species was not found in this study.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7A9B2FFF67F5309EFFFF28.taxon	materials_examined	Material examined. Holotype, ♂, Tallula, La., “ Airpl. 794 ”, take in airplane trap between 20 and 500 feet altitude, P. A. Glick (USNM nº. 62277).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7A9B2FFF67F5309EFFFF28.taxon	distribution	Distribution (Fig. 225). United States of America (Illinois, Indiana, California, Missouri, Arizona, Arkansas, Alabama, Louisiana and Texas), Mexico (Sonora, Sinaloa, Jalisco, Quintana Roo and Chiapas), Guatemala, Honduras and Panama.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C789B28FF67F5329DE9FE2A.taxon	diagnosis	Diagnosis. Mandible tridentate. Clypeus tridentate. Tergal process with large and deep depression with pair of elliptical lateral tuft of setae directed slightly each other. Hypopygium with posterior margin slightly concave. Genitalia: paramere slightly arched inward; aedeagal ventral ramus shorter than aedeagal dorsal body, wide, narrowing slightly apicad; inner margin diverging and outer margin convergent; aedeagal dorsal body with two pairs of apical lobes; outer lobe wide, with apex rounded, dorsal margin much produced inward; inner lobe setose and serrated. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C789B28FF67F5329DE9FE2A.taxon	discussion	Remarks. No additional material of this species was not found in this study.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C789B28FF67F5329DE9FE2A.taxon	materials_examined	Material examined. Holotype, ♂, VENEZUELA, Barinas: Rio Caparo, Research Sta., 32 Km E, El Cornern, seasonal forest, light trap black, 3 – 5. ii. 1978, J. B. Heppener col. (USNM).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C789B28FF67F5329DE9FE2A.taxon	distribution	Distribution (Fig. 242). Guatemala, El Salvador, Nicaragua, Costa Rica, Panama, Colombia, Curação, Venezuela, Trinidad & Tobago and Surinam.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7F9B2AFF67F4969C4DF84C.taxon	description	Description. Male. Body length 2.24 mm. Color: body castaneous. Head (Figs 1 – 2). Mandible with three distal teeth. Median clypeal lobe ill-defined, with one rounded and very large tooth, with high keel; median clypeal carina very high in profile, incomplete apically, straight in profile or nearly so. Gena with spine. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 33). Tergal process median, with very shallow, subcircular and large depression, without tubercle, with short and dense tuft of setae. Posterior hypopygeal margin straight with median concavity (Fig. 64). Genitalia (Figs 99 – 100): paramere with dorsal margin entirely wide and sinuous; apical margin truncate. Aedeagal ventral ramus as long as dorsal body, wide, progressively narrowing apicad; cross section laminar; ramous bifid; surface horizontal; inner margin straight; outer margin slightly sinuous; apex long, simple, parallel; additional inner ramus short; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, subvertical and narrow, with apical margin angled and posterad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7F9B2AFF67F4969C4DF84C.taxon	discussion	Remarks. This species is easily recognized by having a very large and high median clypeal tooth which shows high keel in profile view, gena with spine, the dorsal margin of paramere very sinuous with long and thick setae and the aedeagal ventral ramus bifid. This species is similar to D. cornutus Evans by having a spine in ventral region of head. However, this species has large spine in gena, whereas D. cornutus has shorter spine in hypostomal carina.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7F9B2AFF67F4969C4DF84C.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Panama Pr [ovincia]: P [arque] Nac [ional] Altos de Campana, [08 º 43 ' 15 " N, 80 º 01 ' 43 " W], 10 – 14 ago 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP). Paratypes: Coclé Pr [ovincia]: 1 ♂, El Copé, Par [que] Nac [ional General de División] Omar Torrijos H [errera], 570 – 670 m. s. n. m., [08 º 41 ' 56 " N, 80 º 39 ' 17 " W], 26 – 30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); 1 ♂, same province of holotype, P [arque] Nac [ional] Sob [eranía], Camino, Plantación, [09 º 09 ' N, 79 º 43 ' W], 24 ago. – 2 sep. 1999, [Trampa] Malaise, A. Santos, P. González [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7F9B2AFF67F4969C4DF84C.taxon	distribution	Distribution (Fig. 238). Panama (Coclé and Panama).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7C9B2BFF67F7699D9EF9B6.taxon	diagnosis	Diagnosis. Male. Head regularly wide. Mandible regularly long. Metasomal tergite II without tergal process. Aedeagal ventral ramus without apical filament. Aedeagal dorsal body without apical flagellae.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7C9B2BFF67F7699D9EF9B6.taxon	discussion	Remarks. It has eight species: D. amplus Redighieri & Azevedo, D. clovisi Colombo & Azevedo, D. kuara Colombo & Azevedo, D. pyata Colombo & Azevedo, D. miriamae Colombo & Azevedo, D. azagus sp. nov., D. asfartus sp. nov. and D. iozus sp. nov..	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7C9B2BFF67F7699D9EF9B6.taxon	distribution	Distribution. Panama and Brazil (Espírito Santo).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7C9B35FF67F4839C64F880.taxon	description	Description. Male. Body length 2.27 mm. Color: head and mesosoma castaneous; metasoma light castaneous. Head (Fig. 3). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina high in profile, complete or nearly so, angled in profile. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 34). Tergal process absent. Posterior hypopygeal margin sinuous (Fig. 65). Genitalia (Figs 101 – 102): paramere with dorsal margin wide basally; apical margin rounded. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin slightly sinuous; apex short, simple, parallel; additional inner ramus absent; basal stub short and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and narrow, with apical margin acute and dorsad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7C9B35FF67F4839C64F880.taxon	discussion	Remarks. This species is easily recognized by having the paramere entirely wide with the apex very excavated, the outer lobes of the aedeagal dorsal body with the distal region mesad and L-shaped, and the apex of the aedeagal ventral ramus acute. This species is similar to D. borus sp. nov. by having the apex of paramere very excavated, the dorsal margin almost straight and the basiparamere shorter than the paramere. However, this species has the apical lobes of the aedeagal dorsal body long and the apex of the aedeagal ventral ramus abruptly acute, whereas D. borus sp. nov. has the apical lobes of the aedeagal dorsal body very short and the aedeagal ventral ramus progressively narrowing apicad.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7C9B35FF67F4839C64F880.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Bocas del Toro: Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP). Paratypes: 4 ♂, same data as holotype (MIUP); 1 ♂, Par [que] Nac [ional] Humedal de San [-] San Pond Sac. {= Sak}, [09 ° 28 ' 27.3 " N, 82 ° 27 ' 18 " W], 26 – 28 Oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C7C9B35FF67F4839C64F880.taxon	distribution	Distribution (Fig. 231). Panama (Bocas del Toro).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C619B37FF67F6D99AF9FAD2.taxon	description	Description. Male. Body length 2.03 mm. Color: head and mesosoma castaneous; metasoma light castaneous. Head (Fig. 4). Mandible with three distal teeth. Median clypeal lobe subtrapezoidal, with one angulate tooth; median clypeal carina low in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 35). Tergal process absent. Posterior hypopygeal margin sinuous (Fig. 66). Genitalia (Figs 103 – 104): paramere with dorsal margin wide basally; apical margin rounded. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin sinuous; apex long, crenulate, slightly curved outward; additional inner ramus absent; basal stub short and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, horizontal and wide, with apical margin straight and dorsad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring. Description. Female. (Fig. 223). Body length 1.30 mm. Body and legs light castaneous, eye black. Head 1.46 × as long as wide, side parallel slightly convex. Frons densely coriaceous, punctures large, dense and shallow. Mandible with four apical teeth, lower one larger. Clypeus with subtrapezoidal median lobe and median carina high. Eye with two facets, subcircular, distant from mandible 1.66 × its length. Vertex concave; occipital carina visible in dorsal view. Gena 0.42 × length of head. Mesosoma coriaceous. Pronotal collar 0.32 × length of pronotal disc, 1.19 × as long as wide. Mesoscutum 0.48 × length of pronotal disc, posterior margin convex. Propodeal disc long, 1.62 × as long as wide; spiracle placed at dorsolateral of propodeum, attached to lateral carina. Legs not spinose. Metasoma polished, 0.86 × as long as mesosoma. Petiole short with anterior margin in v-shaped and rugose.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C619B37FF67F6D99AF9FAD2.taxon	discussion	Remarks. This species is easily recognized by having the eyes bulging, the dorsal margin of the paramere very expanded distally downward and the apical margin of the aedeagal dorsal body crenulate. This species is similar to D. krombeini Azevedo by having the apical portion of the dorsal margin of the paramere excavated and the aedeagal dorsal body abruptly narrowing apicad. However, this species has the dorsal margin of the paramere with the distal margin expanded downward, the apical margin of the aedeagal dorsal body crenulate and the tergal process absent, whereas D. krombeini has the dorsal margin of the paramere with the distal margin not expanded downward, the apical margin of the aedeagal dorsal body smooth and has the tergal process with the tuft of setae with one distinctly long seta.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C619B37FF67F6D99AF9FAD2.taxon	materials_examined	Material examined. Holotype, ♂, Darién: P [arque] N [acional] Darién, Pirre, 900 m, Est [ación] Rancho Frio, 21 mar – 4 ab 2000, Tr [ampas] ama [rillas], Cambra, Santos, Bermudez (MIUP). Allotype in copula, ♀, same data as holotype. Paratypes: Bocas del Toro: 5 ♂, Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP); Chiriqui Pr [ovincia]: 28 ♂, P [arque] I [nternacional] L [a] A [mistad], Jurutungo, 1800 m, [08 ° 54 ' N, 82 ° 44 ' W], 30 oct. – 5 nov. 1994, Tr [ampa] Malaise (25 ♂) and Tr [ampas] Amarillas (3 ♂), A. Rodriguez [col.] (MIUP); 10 ♂, 2 km W Cerro Punta, 1700 m, [08 ° 51 ' N, 82 ° 35 ' W], 19. V – 8. VI. 1977, Peck & Howden [col.] (CNCI); 8 ♂, Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 25 – 6. VI. 2000, F [light] I [nterception] T [rap], H. & A. Howden [col.] (7 ♂) and 2. VI. 2000, Howden [col.] (1 ♂) (CNCI); 3 ♂, La Fortuna Dam, 1200 m, wet forest, [08 ° 44 ' 39 " N, 82 ° 14 ' 58 " W], 14 – VI – 16. VII – 1982, F [light] I [nterception] T [rap], B. Gill [col.] (1 ♂, PMAE), (2 ♂, CNCI); 1 ♂, Quebrada de la Mina, 1249 m, 8 ° 39 ' 46 " N, 82 ° 12 ' 30 " W, 27 – 28. V. 2000, Y [ellow] P [an] T [rap], B. & J. Gill [col.] (CNCI); Veraguas Pr [ovincia]: 8 ♂, P [arque] N [acional], Isla Coiba, Cerro la Equis, 200 m, [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 30 – nov – 4 – dic. 1998, [Trampa] Malaise, R. Cambra, S. Bermudez [col.] (1 ♂), 0 m (1 ♂), 29 jul – 4 ag 1998, 120 m (2 ♂), 100 m (1 ♂), 8 – 12 mar 1998, 90 m (3 ♂); [Los Santos]: 2 ♂, Ojo de Agua, [07 ° 31 ' N, 80 ° 25 ' W], 30. VI – 14. VII. 1982, B. Gill [col.] (PMAE); Coclé Pr [ovincia]: 3 ♂ El Copé, Par [que] Nac [ional General de División] Omar Torrijos H [errera], 570 – 670 m. s. n. m., [08 º 41 ' 56 " N, 80 º 39 ' 17 " W], 26 – 30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); Panama Pr [ovincia]: 2 ♂, P [arque] N [atural] Metropolitano, [08 º 59 ' 33 " N, 89 º 32 ' 41 " W], 24 ago. – 3 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); 1 ♂, same data as holotype, except Malaise (MIUP); 1 ♂, same data as holotype, except 1000 m (MIUP); 1 ♂, same data as holotype, except 580 m (MIUP); 1 ♂, same province of holotype, Cana, 1380 m, [07 ° 51 ' N, 87 ° 37 ' W], 4 – 7. VI. 1986, F [light] I [nterception] T [rap], [J.] Ashe & [R.] Brooks [col.] (CNCI); 1 ♂, same province of holotype, Estac [ión] Cruce de Mono, P [arque] Nac [ional] Darién, [07 ° 51 ' 26.3 " N, 87 ° 28 ' 12.7 " W], 6 feb – 4 mar 1993, tr [ampa] Malaise, R. Cambra, J. Coronado [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C619B37FF67F6D99AF9FAD2.taxon	distribution	Distribution (Fig. 230). Panama (Bocas del Toro, Chiriqui, Veraguas, Los Santos, Coclé, Panama and Darién).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C609B30FF67F7609CCDFE0F.taxon	description	Description. Male. Body length 3.13 mm. Color: body castaneous. Head (Fig. 5). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina low in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 36). Tergal process absent. Posterior hypopygeal margin straight or nearly so (Fig. 67). Genitalia (Figs 105 – 106): paramere with dorsal margin entirely wide; apical margin rounded. Aedeagal ventral ramus shorter than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin sinuous; outer margin slightly sinuous; apex short, simple, parallel; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and wide, with apical margin rounded and dorsad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C609B30FF67F7609CCDFE0F.taxon	discussion	Remarks. This species is easily recognized by having the genitalia robust, the lobes of the aedeagal dorsal body with the apical region more expanded, rounded, laminar and the aedeagal ventral ramus very wide. This species is similar to D. abarus sp. nov. by having the dorsal margin of the paramere rounded, the outer lobes of the aedeagal dorsal body long and the aedeagal ventral ramus wide. However, this species has the apex of the apical lobes of the aedeagal dorsal body rounded and laminar, the apex of the aedeagal ventral ramus without projections, whereas D. abarus sp. nov. has the apex of the aedeagal ventral ramus not rounded nor laminar and the apex of the aedeagal ventral ramus with two small rounded projections.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C609B30FF67F7609CCDFE0F.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Veraguas Pr [ovincia]: P [arque] N [acional] Coiba, Isla Coiba, Cerro la Equis, 120 m, [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 29 jul – 4 ag 1998, Malaise, R. Cambra, A. Santos [col.] (MIUP). Paratypes: 1 ♂, same data as holotype; Panama Pr [ovincia]: 1 ♂, [Panama], P [arque] N [atural] Metropolitano, [08 º 59 ' 33 " N, 89 º 32 ' 41 " W], 24 ago. – 3 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); Darién Pr [ovincia]: 3 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 87 ° 45 ' W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C609B30FF67F7609CCDFE0F.taxon	distribution	Distribution (Fig. 237). Panama (Veraguas, Panama and Darién).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C669B31FF67F2A89CE6FEBC.taxon	diagnosis	Diagnosis. Male. Metasomal tergite II with pair of tuft of setae very large and rounded placed in very shallow depression.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C669B31FF67F2A89CE6FEBC.taxon	discussion	Remarks. It has three species: D. apertus Kieffer, D. subpilosus Azevedo and D. parvus Azevedo. Distribution. Ranging from northeast of the U. S. A to Bolivia.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C669B31FF67F3899F92FAD2.taxon	diagnosis	Diagnosis. Male. Mandible tridentate. Clypeus tridentate. Tergal process with very shallow depression, pair of very large and rounded depression with dense tuft of setae. Hypopygium with posterior margin straight. Genitalia: paramere with apex rounded; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar, margins smooth, inner margin straight and outer margin slightly convergent; apex wide with outer corner angulate; aedeagal dorsal body with two pairs of apical lobes; outer pair wide, with apex rounded, inner surface concave, convergent dorsally, dorsal margin with rounded expansion inward; inner pair membranous and setose. Apodeme not extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C669B31FF67F3899F92FAD2.taxon	discussion	Remarks. No additional material of this species was not found in this study. The holotype is lost, but the identity of this species is well-established. The diagnosis presented above is based on a male from Mexico as listed below.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C669B31FF67F3899F92FAD2.taxon	materials_examined	Material examined. MEXICO, Quintana Roo: 3 Km, S Puente Morelos, Jd Botânico, 14. XII. 1993, L. Masner col. (CNCI).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C669B31FF67F3899F92FAD2.taxon	distribution	Distribution (Fig. 227). United States of America (New York, Pennsylvania, Maryland, Nebrasca, Illinois, Ohio, New Jersey, D. C, Delaware, Kansas, Missouri, Virginia, North and South Carolina, New Mexico, Texas and Florida), Mexico (Jalisco, Vera Cruz, Quintana Roo and Chiapas), Belize, Honduras, Guatemala, Costa Rica and Panama.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C669B31FF67F75E9B3DF8AF.taxon	diagnosis	Diagnosis. Male. Mandible bidentate. Pronotal disc usually with texture and punctures as frons. Mesoscutum and scutellum with texture weaker, punctures smaller than those of frons. Metasoma with pair of lateral and anterior depression, large usually more than 0.5 × median length of tergite II. Hypopygeal median stalk slender. Female. Mandible tetradentate. Clypeus tridentate. Metapectal propodeal disc with sides nearly parallel.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C669B31FF67F75E9B3DF8AF.taxon	discussion	Remarks. It has 34 species: D. abruptus Azevedo, D. alticarinatus Azevedo, D. archeatus Azevedo, D. basivolsellus Azevedo, D. bivolsellus Azevedo, D. curvilongus Azevedo, D. elongatus Azevedo, D. falciformes Azevedo, D. flagellatus Azevedo, D. geniculatus, D. gladius Azevedo, D. intradentatus, D. largimanus Azevedo, D. latus Azevedo, D. lobisserratus Azevedo, D. longimerus Azevedo, D. longipilosus Azevedo, D. mantoides Azevedo, D. megadentatus Azevedo, D. microdentatus Azevedo, D. octavus Azevedo, D. pedipalpoides Azevedo, D. piscicercus, D. rectangularis Azevedo, D. retorcerens Azevedo, D. sinuatus Azevedo, D. spiculus Azevedo, D. strepsus, D. subtriangularis Azevedo, D. tetracerutus Azevedo, D. tetralobatus Azevedo, D. trogon Azevedo, D. tropoides Azevedo, D. falcatus Evans and D. bicavatus Evans.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C669B31FF67F75E9B3DF8AF.taxon	distribution	Distribution. Ranging from South of Mexico to Southeastern Brazil (São Paulo).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C659B32FF67F2A89D8AFCA6.taxon	diagnosis	Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with pair of antero-lateral depressions, ovoid, deep, large with small pit covered by tuft of long seate. Hypopygium with posterior margin concave. Genitalia: paramere with oblique and concave apex; aedeagal ventral ramus as long as aedeagal dorsal body, laminar, surface subvertical, apical half convergent, apex bidentate; aedeagal dorsal body with three pairs of apical lobes; outer pair large, laminar, surface horizontal, outer margin with six teeth sharpened progressively, inner margin with projection; dorsal inner pair stout, membranose and setose; ventral inner pair small, laminar, narrowing to apex; basal process of aedeagal dorsal body with pair of lateral teeth. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C659B32FF67F2A89D8AFCA6.taxon	discussion	Remarks. This species is known to Costa Rica and Panama (Azevedo 2001) and now more specimens are added. In this series the specimens are quite similar to those previously studied, without any variation that worths to mention.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C659B32FF67F2A89D8AFCA6.taxon	materials_examined	Material examined. Holotype ♂, COSTA RICA, Guanacaste: 3 km SE Rio Naranjo, 1. XI. 1991, F. Parker col. (EMUS); PANAMA, Bocas del Toro Pr [ovincia]: 1 ♂, Par [que] Nac [ional] Humedal de San [-] San Pond Sac {= Sak}, [09 ° 28 ' 27.3 " N, 82 ° 27 ' 18 " W], 26 – 28 Oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP); Darién Pr [ovincia]: 1 ♂, P [arque] N [acional] Darién, Pirre, Est [ción] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C659B32FF67F2A89D8AFCA6.taxon	distribution	Distribution (Fig. 234). Costa Rica and Panama.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C659B32FF67F1F29C56F928.taxon	diagnosis	Diagnosis. Mandible bidentate. Clypeus tridentate. Tergal process with pair of antero-lateral depressions, ovoid, deep, large with small pit covered by tuft of long seate. Hypopygium with posterior margin slightly concave. Genitalia: paramere with apex concave; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar, surface sub-horizontal; outer margin concave apically; apex acute, slightly diverging; aedeagal dorsal body with four pairs of apical lobes, outer pair laminar, surface vertical, narrowing at apex and directed downward, apical; dorsal margin with short expansion and small teeth; ventral margin with large tooth, three inner pairs stout, membranose and setose; basal process with pair of lateral teeth directed outward, margin between them straight. Apodeme not extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C659B32FF67F1F29C56F928.taxon	discussion	Remarks. This species is known to Panama (Azevedo 2001) and here more specimens are added. Some variations are worth to mention: mesosoma and metasoma castaneous; median clypeal tooth angulate; median clypeal carina complete apically; vertex crest straight; anterior margin of pronotal disc ecarinate; basal process of aedeagal dorsal body without teeth.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C659B32FF67F1F29C56F928.taxon	materials_examined	Material examined. Holotype ♂, PANAMA, Canal Zone: Barro Colorado Island, 3 - 13. VI. 1983. B. Gill col. (PMAE); PANAMA, Darién Pr [ovincia]: 14 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 87 ° 45 ' W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP); 2 ♂, Cruce de Mono: Est [ación] I [nstituto] N [acional de] Re [cusos] Na [turales] Re [novables], Parque] nacional Darién, [07 ° 51 ' 26.3 " N, 87 ° 28 ' 12.7 " W], 5 – 28 feb 1993, Tr [ampas] Amarillas, R. Cambra, J. Coronado [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C659B32FF67F1F29C56F928.taxon	distribution	Distribution (Fig. 237). Panama (Panama and Darién).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C659B3CFF67F5759C29FEC2.taxon	diagnosis	Diagnosis. Mandible bidentate. Clypeus subtrapezoidal. Tergal process with pair of antero-lateral depressions, ovoid, deep, margin with setae, pit with high border covered by tuft of long setae in middle of depression. Hypopygium with posterior margin straight. Genitalia: paramere with apex oblique and truncate; aedeagal dorsal ramus shorter than aedeagal dorsal body, laminar; surface horizontal, evenly wide; apex bidentate; aedeagal dorsal body with two pairs of apical lobes; outer pair laminar, surface vertical, wide; apex rounded and directed downward; inner pair, membranous, stout and setose. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C659B3CFF67F5759C29FEC2.taxon	discussion	Remarks. No additional material of this species was not found in this study. Material examined. Holotype ♂, TRINIDAD & TOBAGO, Trinidad Island: Simila Reserve Station, 2 – 15. VI. 1981, Hanson & Clemons col. (EMUS).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C659B3CFF67F5759C29FEC2.taxon	distribution	Distribution (Fig. 238). Panama and Trinidad & Tobago.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6B9B3CFF67F3509CCDFB45.taxon	diagnosis	Diagnosis. Mandible bidentate. Clypeus tridentate. Tergal process with pair of antero-lateral depressions, ovoid, deep, large with small pit covered by tuft of long seate. Hypopygium with posterior margin concave. Genitalia: paramere with apex oblique and slightly concave; aedeagal ventral ramus as long as aedeagal dorsal body, laminar, surface vertical, parallel; apex with two large apical teeth; aedeagal dorsal body with three pairs of apical lobes; outer pair with apex converging, inner margin serrated, ventral margin with three long and narrow teeth; inner pairs stout, membranous and setose; dorsal pair short and the ventral as long as aedeagal dorsal body; basal process of aedeagal dorsal body with pair of short teeth outward and median long tooth. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6B9B3CFF67F3509CCDFB45.taxon	discussion	Remarks. This species is known only to Panama (Azevedo 2001) and now more specimens are added. In this series the specimens are quite similar to those previously studied, without any variation that worths to mention.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6B9B3CFF67F3509CCDFB45.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Canal Zone: Barro Colorado Island, 3 – 13. VI. 1983, B. Gill col. (PMAE); PANAMA, Veraguas Pr [ovincia]: 1 ♂, P [arque] N [acional] Coiba, Isla Coiba, Cerro la Equis, 120 m [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 29 jul – 4 ag 1998, Malaise, R. Cambra, A. Santos [col.] (MIUP); 1 ♂, Isla Coiba, Estación biológica, [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 4 – 8 Agosto 1998, [trampas] Amarillas, Col. L. D., R. C., A. A., A. S. (MIUP); Darién Pr [ovincia]: 60 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 87 ° 45 ' W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (59 ♂) and 24 – 26 marzo, 450 m, col. A. Santos (1 ♂) (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6B9B3CFF67F3509CCDFB45.taxon	distribution	Distribution (Fig. 241). Panama (Veraguas, Panama and Darién).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6B9B3CFF67F6D29D8AF8BD.taxon	diagnosis	Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with pair of antero-lateral depressions, ovoid, deep, large with small pit covered by tuft of long setae. Hypopygium with posterior margin somewhat concave. Genitalia: paramere with apex oblique and concave; aedeagal dorsal ramus longer than aedeagal dorsal body, parallel, apical third directed downward and apex forward, sharpened and abruptly angled; inner margin with series of small teeth; aedeagal dorsal body with three pairs of apical lobes; outer pair with two rami, dorsal ramus parallel with apex narrow and angled downward, ventral ramus arched inward and dorsal ramus with apices nearly touching each other; inner pairs stout, membranose with rounded apex. Apodeme not extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6B9B3CFF67F6D29D8AF8BD.taxon	discussion	Remarks. This species is known to Costa Rica and Panama (Azevedo 2001). Now one specimen is added. In this series the specimens are quite similar to those previously studied, without any variation that worths to mention.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6B9B3CFF67F6D29D8AF8BD.taxon	materials_examined	Material examined. Holotype, ♂, COSTA RICA, Guanacaste: 3 km, SE, Rio Naranjo, 13 – 31. VII. 1993, F. D. Parker (EMUS); PANAMA, Darién: 1 ♂ Cana Serrania de Pirre, 1250 m, 4 – 7. vi. 1996, J. Ashe, R. Brooks (UFES).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6B9B3CFF67F6D29D8AF8BD.taxon	distribution	Distribution (Fig. 242). Costa Rica and Panama.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6A9B3DFF67F3E59D8AFAF8.taxon	diagnosis	Diagnosis. Mandible tridentate. Clypeus with median lobe ill-defined. Tergal process median with deep and large elliptical depression, occupying slightly more than median third of the width of the tergite with two rounded median tuft of setae. Hypopygium with median concavity. Genitalia: paramere with apex truncare and oblique with four projections low and rounded; aedeagal ventral ramus shorter than aedeagal dorsal body, flattened, narrow, wholly arched; median part outward and apex abruptly angulate outward; aedeagal dorsal body not so wide laterally, narrowing to apex, with sufaces vertical and apex slightly divergent, ventral margin serrated. Apodeme not extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6A9B3DFF67F3E59D8AFAF8.taxon	discussion	Remarks. This species was known from Costa Rica (Azevedo 1999 a) and is recorded for the first time from Panama. Some variations are worth to mention: metasoma light castaneous and dark castaneous; median clypeal tooth short; anterior margin of pronotal disc coarse; depression of tergal process shallow, more distant each other with few and shorter tuft of setae, some specimens have dense and longer tuft of setae.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6A9B3DFF67F3E59D8AFAF8.taxon	materials_examined	Material examined. Holotype, ♂, COSTA RICA, San Vito de Coto Brus: Las Cruces, 1200 m, 17. vii – 12. ix. 1982, Malaise trap, B. Gill col. (CNCI). PANAMA, Chiriqui: 4 ♂, Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 25 – 6. VI. 2000, (3 ♂) and 2 – 6. VI. 2000 (1 ♂), F [light] I [nterception] T [rap], H. & A. Howden [col.] (CNCI); 2 ♂, P [arque] I [nternacional] L [a] A [mistad], Jurutungo, 1800 m, [08 ° 54 ' N, 82 ° 44 ' W], 30 oct. 5 nov 1994, Tr [ampa] Malaise, (1 ♂) and 1600 m, 30 oct 1994, T [rampa] Amarilla (1 ♂), A. Rodriguez [col.] (MIUP); 5 ♂, 2 km W Cerro Punta, 1700 m, [08 ° 51 ' N, 82 ° 35 ' W], 19. V – 8. VI. 1977, Peck & Howden [col.] (CNCI); 1 ♂, 15 km NW Hato del Volcano, 1200 m, [08 ° 46 ' N, 82 ° 38 ' W], 24 – 31. V. 1997, Peck & Howden [col.] (CNCI).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6A9B3DFF67F3E59D8AFAF8.taxon	distribution	Distribution (Fig. 226). Costa Rica and Panama.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6A9B3DFF67F2A89BC9FE59.taxon	diagnosis	Diagnosis. Male. Metasoma with median large depression; pair of tubercles. Aedeagal ventral ramus narrow, much arched outward medially, apex abruptly angulate, straight and diverging.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6A9B3DFF67F2A89BC9FE59.taxon	discussion	Remarks. It has six species: D. bifoveatus Kieffer, D. microculus Azevedo, D. angulatus, D. asitius sp. nov., D. ibocius sp. nov. and D. tisinus sp. nov..	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6A9B3DFF67F2A89BC9FE59.taxon	distribution	Distribution. Mexico (Durango, Morelos, Chiapas), Nicaragua, Costa Rica and Panama.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6A9B3EFF67F7439C64FEBB.taxon	description	Description. Male. Body length 2.03 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 6). Mandible with two distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina low in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 37). Tergal process with median, shallow, subcircular and large depression, with small, low and dorsad tubercle placed on center of depression, pit very small, with short and dense tuft of setae. Posterior hypopygeal margin strongly concave (Fig. 68). Genitalia (Figs 125 – 126): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface subvertical; inner margin sinuous; outer margin sinuous; apex long, toothed, angled outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body two pairs of apical lobes; outer lobe long, horizontal and wide, with apical margin acute and dorsad, inner pair stout, membranous and setose. Apodeme not extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6A9B3EFF67F7439C64FEBB.taxon	discussion	Remarks. This species is easily recognized by having the dorsal margin of the paramere almost straight, the apex of the outer lobe of the aedeagal dorsal body abruptly acute, the aedeagal ventral ramus with a large fold and the genital ring bifurcated in the distal region in ventral view. This combination of characters is unusual for bifoveatus species-group.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6A9B3EFF67F7439C64FEBB.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Bocas del Toro: Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP). Paratypes: 5 ♂, same data as holotype.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6A9B3EFF67F7439C64FEBB.taxon	distribution	Distribution (Fig. 228). Panama (Bocas del Toro).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C689B3FFF67F2A89C4DFBD0.taxon	description	Description. Male. Body length 2.24 mm. Color: body dark castaneous. Head (Fig. 7). Mandible with two distal teeth. Median clypeal lobe ill-defined, with one angulate very short tooth; median clypeal carina low in profile, incomplete apically, angled in profile. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc coriaceous; anterior margin carinate. Metasoma (Fig. 38). Tergal process with shallow, subcircular and large depression, with very small and very low dorsad tubercle placed on inner area of depression, pit very small with short tuft of setae. Posterior hypopygeal margin weakly concave (Fig. 69). Genitalia (Figs 127 – 128): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface subvertical; inner margin sinuous; outer margin sinuous; apex long, crenulate, curved outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and wide, with apical margin crenulate and laterad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring. Variations. Body length between 1.79 (min.) mm and 2.60 (max.) mm.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C689B3FFF67F2A89C4DFBD0.taxon	discussion	Remarks. This species is similar to D. asitius sp. nov. by having the aedeagal ventral ramus with toothed apex and the paramere with crenulate apex. However, it species has the apical lobes of the aedeagal dorsal body long, with laterad apex and the paramere with the dorsal margin sinuous, whereas D. asitius sp. nov. has the apical lobes of the aedeagal dorsal body shorter with dorsad apex and the paramere with the dorsal margin almost straight.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C689B3FFF67F2A89C4DFBD0.taxon	materials_examined	Material examined. Holotype, ♂, Coclé Pr [ovincia]: El Copé, Par [que] Nac [ional General de División] Omar Torrijos H [errera], 570 – 670 m. s. n. m., [08 º 41 ' 56 " N, 80 º 39 ' 17 " W], 26 – 30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP). Paratypes: 2 ♂, same data as holotype; Panama Pr [ovincia]: 2 ♂, P [arque] Nac [ional] Altos de Campana, [08 º 43 ' 15 " W, 80 º 01 ' 43 " W], 10 – 14 ago 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C689B3FFF67F2A89C4DFBD0.taxon	distribution	Distribution (Fig. 235). Panama (Coclé and Panama).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C689B38FF67F65D9DD0FF6F.taxon	description	Description. Male. Body length 3.52 mm. Color: body dark castaneous. Head (Fig. 8). Mandible with three distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina high in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 39). Tergal process with median deep, subcircular and large depression, without tubercle, pit absent, with short and dense tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 70). Genitalia (Figs 129 – 130): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface subvertical; inner margin slightly sinuous; outer margin sinuous; apex long, simple, angled outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, subhorizontal and narrow, with apical margin abruptly acute and ventrad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring. Variations. Metasoma castaneous; median clypeal tooth longer.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C689B38FF67F65D9DD0FF6F.taxon	discussion	Remarks. This species is easily recognized by having the apical lobes of the aedeagal dorsal body short and triangular, the median portion of the aedeagal ventral ramus sinuous and narrow. This species is similiar to D. asitius sp. nov. by having the paramere with the dorsal margin almost straight and the apex of the aedeagal dorsal body curved outward. However, this species has the apical lobes of the aedeagal dorsal body short, narrowing apicad and the apical portion of the aedeagal ventral ramus slender and smooth, whereas D. asitius sp. nov. has the apical lobes of the aedeagal dorsal body longer, abruptly acute and the apical portion of the aedeagal ventral ramus toothed.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C689B38FF67F65D9DD0FF6F.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Chiriqui: Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 2 – 6. VI. 2000, F [light] I [nterception] T [rap], H. & A. Howden [col.] (CNCI). Paratypes: 1 ♂, same data as holotype; 2 ♂, same data as holotype except 25 – 6. VI. 2000 (CNCI); 1 ♂, 15 km NW Hato del Volcano, 1200 m, [08 ° 46 ' N, 82 ° 38 ' W], 24 – 31. V. 1997, Peck & Howden [col.] (CNCI).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C689B38FF67F65D9DD0FF6F.taxon	distribution	Distribution (Fig. 243). Panama (Chiriqui).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6F9B38FF67F0799DF1FAE2.taxon	diagnosis	Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with pair of latero-anterior ovoid and shallow depressions with tubercle and tuft of setae. Hypopygium with posterior margin slightly concave. Genitalia: paramere arched inward, apex produced and narrow; aedeagal ventral longer than aedeagal dorsal body, wide, tubular, parallel, apex rounded; aedeagal dorsal body with three pairs of apical lobes, inner pair elongate with rounded apex; median pair membranous, laminar, entirely, wide; outer pair with apical projection inclined downward with strongly serrated margin. Apodeme not extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6F9B38FF67F0799DF1FAE2.taxon	discussion	Remarks. This species is known to Panama, Venezuela, British Guiana, Suriname, Ecuador Peru, Brazil (Rondônia, São Paulo) and Argentina (Salta). Here two specimens are added. Some variations are worth to mention: median clypeal lobe ill-defined with one rounded tooth; median clypeal carina high in profile; vertex crest straight; pronotal disc with margin ecarinate; paramere longer than aedeagal dorsal body; basiparamere shorter than paramere; aedeagal dorsal ramus with inner margin concave in median portion.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6F9B38FF67F0799DF1FAE2.taxon	materials_examined	Material examined. Holotype ♂, BRAZIL, Pará: Baker col. (CASC, # 17054). PANAMA, Veraguas Pr [ovincia]: 2 #, P [arque] N [acional] Coiba, Isla Coiba, Cerro La Equis, 90 m, [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 30 – nov. – 4 dic. 1998, Malaise, R. Cambra, S. Bermudez [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6F9B38FF67F0799DF1FAE2.taxon	distribution	Distribution (Fig. 228). Panama, Venezuela, Guiana, Suriname, Ecuador, Peru, Brazil (Pará, Rondônia, Acre, Espírito Santo and São Paulo) and Argentina.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6F9B38FF67F33B9E8BFE2C.taxon	diagnosis	Diagnosis. Male. Mandible tridentate. Clypeus broad. Aedeagal ventral ramus tubular. Aedeagal dorsal body small.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6F9B38FF67F33B9E8BFE2C.taxon	discussion	Remarks. It has two species: D. brasiliensis and D. tubulatus Redighieri & Azevedo. Distribution. Venezuela, Guiana, Suriname, Ecuador, Peru, Brazil (Rondônia, Acre, Paraíba, Espírito Santo and São Paulo) and Argentina.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6F9B38FF67F7B09D4AF8EC.taxon	diagnosis	Diagnosis. Male. Median clypeal lobe trapezoidal, except D. filus Azevedo and D. ibrastus sp. nov.. Metasoma with pair lateral tergal process, tubercle, pit on top and tuft of setae.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6F9B38FF67F7B09D4AF8EC.taxon	discussion	Remarks. It has 19 species: D. cervoides, D. conicus Azevedo, D. curviventris Azevedo, D. filus, D. manus Azevedo, D. truncatus Azevedo, D. umbilicus Azevedo, D. h-ramus Redighieri & Azevedo, D. laminaris Redighieri & Azevedo, D. verrucosus Redighieri & Azevedo, D. botocudus Colombo & Azevedo, D. ardonius sp. nov., D. arteus sp. nov., D. ettus sp. nov., D. ibrastus sp. nov., D. racarus sp. nov., D. teraus sp. nov. and D. urazius sp. nov..	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6F9B38FF67F7B09D4AF8EC.taxon	distribution	Distribution. Panama, Trinidad & Tobago, Ecuador, Peru, Brazil (Amazonas, Pará, Rondônia, Ceará, Pernambuco, Mato Grosso, Goiás, Distrito Federal, Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo, Paraná and Santa Catarina), Bolivia.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6E9B39FF67F2A89EC2FBF1.taxon	diagnosis	Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with lateral pair of small and very shallow depressions and pit with dense tuft of setae. Hypopygium with posterior margin straight. Genitalia: paramere longer than basiparamere, dorsal margin straight; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; apical half translucent and thin with top-rounded expansions, apex constricted below; aedeagal dorsal body with three pairs of apical lobes; dorsal pair thick, surface vertical and parallel; dorsal margin angled; ventral pair membranous, stout, rounded, shorter than dorsal one; median pair less stout, less membranous and shorter than median one; basal process with pair of lateral rounded expansions upward. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6E9B39FF67F2A89EC2FBF1.taxon	discussion	Remarks. This species is known to Panama, Trinidad & Tobago, Ecuador, Peru, Brazil (Amazonas, Pará, Rondônia, Mato Grosso, Goiás) and Bolivia (Azevedo 2003). Here more specimens are added. Some variations in this series worth to mention: mesosoma castaneous, metasoma light castaneous; median clypeal tooth angulate; median clypeal carina straight; pronotal disc ecarinate; tergal process without depression.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6E9B39FF67F2A89EC2FBF1.taxon	materials_examined	Material examined. Holotype ♂, BRAZIL, Pará: Tucuruí, Rio Tocantins, V. 1986, Interception Trap, N. Degallier col. (MPEG); PANAMA, Colón: 8 ♂, 14 km N J [un] ct [ion] Escobal r [oa] d and Piña r [oa] d on Piña r [oa] d, ± 20 m, [09 ° 20 ' N', 79 ° 54 ' W], 2 – 3. VI. 1995, F [light] I [nterception] T [rap], J. Ashe & R. Brooks [col.] (CNCI); 4 ♂, S [an] ta Rita [Arriba], Macizo, Parador Sierra Llorona, [09 ° 19 ' 57.1 " N, 79 ° 45 ' 1.6 " W], 22 dic. 1997, T [rampa] Malaise, D. Quintero [col.] (MIUP); Darién Pr [ovincia]: 1 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 230 m, [08 ° 00 ' N, 77 ° 45 ' W], 21 mar – 4 ab 2000, Malaise, Cambra, Santos, Bermudez [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6E9B39FF67F2A89EC2FBF1.taxon	distribution	Distribution (Fig. 229). Panama, Trinidad & Tobago, Ecuador, Peru, Brazil (Amazonas, Pará, Rondônia, Mato Grosso, Goiás), Bolivia.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6E9B3AFF67F6BE9C64FF27.taxon	description	Description. Male. Body length 3.22 mm. Color: body castaneous. Head (Fig. 9). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 40). Tergal process with lateral, shallow, circular and small depression, with large, very low and dorsad tubercle placed on center of depression, pit large, with short and sparse tuft of setae. Posterior hypopygeal margin median straight with angled corners (Fig. 71). Genitalia (Figs 133 – 135): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, entirely wide, cross section laminar; surface vertical; inner margin strongly sinuous; outer margin slightly sinuous; apex long, slightly crenulate, parallel; additional inner ramus long; basal stub long and narrow. Aedeagal dorsal body with two pairs of apical lobes, outer lobe long, vertical and very narrow, with apical margin acute and ventrad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring. Variation. Metasoma light castaneous.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6E9B3AFF67F6BE9C64FF27.taxon	discussion	Remarks. This species is easily recognized by having two spines in the ventral margin of the apical lobes of the aedeagal dorsal body with apical spine shorter than distal one, the aedeagal ventral ramus vertical and wide in the median portion with wavy apex. This species is similar to D. urazius sp. nov. by having spine in the ventral margin of apical lobes of the aedeagal dorsal body and the aedeagal ventral ramus entirely wide with the inner margin sinuous. However, this species has the dorsal margin of the paramere sinuous, the apex of the basal bar not projected forward, and the apical margin of the aedeagal dorsal body wavy, whereas D. urazius sp. nov. has the dorsal margin of the paramere not sinuous, the apex of the basal bar projected forward and the apical margin of the aedeagal ventral ramus not wavy.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6E9B3AFF67F6BE9C64FF27.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Bocas del Toro: Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP). Paratypes: 9 ♂, same data as holotype.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6E9B3AFF67F6BE9C64FF27.taxon	distribution	Distribution (Fig. 226). Panama (Bocas del Toro).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6D9B3AFF67F3739C1CFAF0.taxon	description	Description. Male. Body length 4.96 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 10). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, complete or nearly so, convex in profile. Frons strongly coriaceous and punctures large. Vertex crest weakly concave. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 41). Tergal process with lateral, deep, subcircular and large depression, with small, low and dorsad tubercle placed on anterior area of depression, pit small, with long and sparse tuft of setae. Posterior hypopygeal margin sinuous (Fig. 72). Genitalia (Figs 136 – 137): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface vertical; outer margin and inner margin slightly sinuous; apex long, simple, parallel; additional inner ramus long; basal stub short and narrow. Aedeagal dorsal body with pair of apical lobe; outer lobe long, vertical and very narrow, with apical margin crenulate and ventrad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring. Variation. Tergal process with very shallow depression and without tubercle.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6D9B3AFF67F3739C1CFAF0.taxon	discussion	Remarks. This species is similar to the D. racarus sp. nov. by having the dorsal margin of the paramere sinuous, the apex of the basal bar projected forward and the inner margin of the aedeagal ventral ramus crenulate. However, this species has the apex of the basal bar smooth, the ventral margin of the apical lobes of the aedeagal dorsal body not excavated and the apical margin of the aedeagal ventral ramus smooth, whereas D. racarus sp. nov. has the apex of basal bar crenulate, the ventral margin of the apical lobes of the aedeagal dorsal body deeply excavated and the apical margin of the aedeagal ventral ramus with a small projection dorsad.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6D9B3AFF67F3739C1CFAF0.taxon	materials_examined	Material examined. Holotype ♂, PANAMA, Bocas del Toro: Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP). Paratypes: 7 ♂, same data as holotype; Coclé Pr [ovincia]: 1 ♂, El Copé, Par [que] Nac [ional General de División] Omar Torrijos H [errera], 570 – 670 m. s. n. m., [08 º 41 ' 56 " N, 80 º 39 ' 17 " W], 26 – 30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6D9B3AFF67F3739C1CFAF0.taxon	distribution	Distribution (Fig. 229). Panama (Bocas del Toro and Coclé).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6D9B04FF67F7BD9BAFFDE2.taxon	description	Description. Male. Body length 4.0 mm. Color: body dark castaneous. Head (Fig. 11). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina low in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 42). Tergal process lateral, without depression, low and dorsad tubercle, pit small, with short and dense tuft of setae. Posterior hypopygeal margin weakly concave (Fig. 73). Genitalia (Figs 138 – 139): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus higher than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin excavated; outer margin sinuous; apex short, translucent, wavy, parallel, additional inner ramus short; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, subhorizontal and wide, with apical margin rounded and ventrad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring. Variations. Tergal process closer to each other; tubercle higher or with few setae.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6D9B04FF67F7BD9BAFFDE2.taxon	discussion	Remarks. This species is easily recognized by having the paramere narrow with the dorsal margin straight, very acute apex, the basivolsella wavy and the basal bar in the median portion of the aedeagal dorsal body projected forward. This species is similar to the D. cervoides by having the apex of the aedeagal ventral ramus laminar, translucent with two expansions. However, this species has the paramere narrow, the aedeagal ventral ramus wide with the inner margin of the apex smooth and the basivolsella wavy, whereas D. cervoides has the paramere not so straight, the aedeagal ventral ramus narrower with the inner margin of the apex serrated and the basivolsella not wavy.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6D9B04FF67F7BD9BAFFDE2.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Darién Pr [ovincia]: P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP). Paratypes: Bocas del Toro: 1 ♂, Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP); Chiriqui: 3 ♂, Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 25 – 6. VI. 2000, F [light] I [nterception] T [rap], H. & A. Howden [col.] (CNCI); Veraguas Pr [ovincia]: 2 ♂, P [arque] N [acional] Coiba, Isla Coiba, Cerro la Equis, 0 m, [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 30. nov – 4. dic. 1998, Malaise, R. Cambra, S. Bermudez [col.] (1 ♂) and 6 – 11 mar 1998, Tr [ampa] R. Cambra, A. Santos (1 ♂) (MIUP); Panama Pr [ovincia]: 1 ♂, Cerro Azul, Las Nubes, [09 º 10 ' N, 79 º 25 ' W], 9 – 15 – oct. 1999, Malaise, R. Cambra, A. Santos [col.] (MIUP); 1 ♂, C [anal] Z [one], B [arro] Colorado Is [land], [09 ° 09 ' N, 79 ° 51 ' W], 15 – 31. VII. 1994, F [light] I [nterception] T [rap], D. Banks [col.] (CNCI); 5 ♂ same data as holotype; 1 ♂, same province of holotype, Cruce de Mono: Est [ación] I [nstituto] N [acional de] Re [cusos] Na [turales] Re [novables], [07 ° 51 ' 26.3 " N, 77 ° 28 ' 12.7 " W], 6. II – 4. III. 1993, t [rap] Malaise, R. Cambra, J. Coronado [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C6D9B04FF67F7BD9BAFFDE2.taxon	distribution	Distribution (Fig. 233). Panama (Bocas del Toro, Chiriqui, Veraguas, Panama and Darién).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C539B04FF67F0B09DD0F990.taxon	description	Description. Male. Body length 3.10 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 12). Mandible with three distal teeth. Median clypeal lobe ill- defined, with one angulate very large tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly convex. Pronotal disc weakly coriaceous; anterior margin carinate. Metasoma (Fig. 43). Tergal process with lateral, deep, subcircular and large depression, with small, low and dorsad tubercle placed on center of depression, pit small, with short and sparse tuft of setae. Posterior hypopygeal margin weakly concave (Fig. 74). Genitalia (Figs 140 – 141): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin slightly sinuous; apex long, simple, parallel; additional inner ramus absent; basal stub short and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and wide, with apical margin rounded and ventrad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring. Variation. Tergal process with very shallow depression.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C539B04FF67F0B09DD0F990.taxon	discussion	Remarks. This species is easily recognized by having the median clypeal tooth very long, the base of the outer margin of the apical lobes of the aedeagal dorsal body with two small rounded and dorsad projections. This species is similar to D. longiclypeus Azevedo by having the median clypeal tooth long. However, this species has the tergal process lateral, the apical lobes of the aedeagal dorsal body long and the aedeagal ventral ramus with the inner margin of the apex excavated, whereas D. longiclypeus has the tergal process median, the aedeagal dorsal body short and the aedeagal ventral ramus not excavated.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C539B04FF67F0B09DD0F990.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Chiriqui Pr [ovincia]: P [arque] I [nternacional] L [a] A [mistad], Jurutungo, 1800 m, [08 ° 54 ' N, 82 ° 44 ' W], 30 oct – 5 nov. 1994, Tr [ampas] Amarillas, A. Rodriguez [col.] (MIUP). Paratypes: 2 ♂, same data as holotype; 1 ♂, 2 km W Cerro Punta, 1700 m, [08 ° 51 ' N, 82 ° 35 ' W], 19. V – 8. VI. 1977, Peck & Howden [col.] (CNCI).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C539B04FF67F0B09DD0F990.taxon	distribution	Distribution (Fig. 235). Panama (Chiriquí).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C539B05FF67F4A29CE1FC82.taxon	description	Description. Male. Body length 4.78 mm. Color: body castaneous. Head (Fig. 13). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin coarse. Metasoma (Fig. 44). Tergal process lateral with shallow, subcircular and large depression, with small, low and dorsad tubercle placed on anterior area of depression, pit large, with short and sparse tuft of setae. Posterior hypopygeal margin sinuous (Fig. 75). Genitalia (Figs 142 – 144): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, entirely wide with small projection apicad; cross section laminar; surface vertical; inner margin sinuous, outer margin sinuous; apex short, simple, parallel; additional inner ramus long; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe, long, vertical and very narrow, with apical margin crenulate and posterad; inner pair, stout, membranous and setose. Apodeme not extending beyond genital ring. Variations. Body length 3.31 (min.) mm and 6.06 mm (max.). Depression of tergal process deep or very shallow.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C539B05FF67F4A29CE1FC82.taxon	discussion	Remarks. This species is easily recognized by having the ventral margin of the apical lobes of the aedeagal dorsal body deeply excavated, the apex of the basal with apex projected forward, the aedeagal ventral ramus vertical and the apex with a small projection dorsad. This species is similar to D. arteus sp. nov. by having the dorsal margin of the paramere sinuous, the apex of the basal bar projected forward and the inner margin of the aedeagal ventral ramus crenulate. However, this species has the apex of the basal bar crenulate, the ventral margin of the aedeagal dorsal body deeply excavated and the apical margin of the aedeagal dorsal body with small dorsad projection, whereas D. arteus sp. nov. has the apex of the basal bar smooth, the ventral margin of the apical lobes of the aedeagal dorsal body not excavated and the aedeagal ventral ramus without projection.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C539B05FF67F4A29CE1FC82.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Darién Pr [ovincia]: P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov [20] 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP). Paratypes: Bocas del Toro: 1 ♂, Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ap] Malaise, A. Santos [col.] (MIUP); 4 ♂, same data as holotype; 1 ♂, same data as holotype except 450 m, 24 – 26 marzo, col. A. Santos (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C539B05FF67F4A29CE1FC82.taxon	distribution	Distribution (Fig. 243). Panama (Bocas del Toro and Darién).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C529B06FF67F1989B7BFE73.taxon	description	Description. Male. Body length 4.33 mm. Color: head and mesosoma dark castaneous; metasoma light castaneous. Head (Fig. 14). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly convex. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 45). Tergal process with lateral, shallow, circular and large depression, with small, low and laterad tubercle placed on inner area of depression, pit small, with short tuft of setae. Posterior hypopygeal margin sinuous (Fig. 76). Genitalia (Figs 145 – 153): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus higher than dorsal body, wide, progressively narrowing apicad; cross section laminar; ramous bifid; surface horizontal; inner margin sinuous; outer margin sinuous; apex long, simple, curved outward; additional inner ramus long; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and narrow, with apical margin rounded and mesad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring. Variations. Head dark castaneous; metassoma castaneous; frons coriaceous; median clypeal lobe subtrapezoidal; depression of tergal process very shallow or without depression, with very low, very small and without tubercle; apical lobe narrow in sickle-shaped with teeth in inner margin and apical lobes with acute apex and inner margin smooth. This species has two different aedeagal ventral ramus and apical lobes of the aedeagal dorsal body. In one kind, the aedeagal ventral ramus can be lower than the aedeagal dorsal body; the apical margin entirely wide with rounded and dorsad apex; in the other kind the aedeagal ventral ramus can have the apex narrow and two small and laterad projections.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C529B06FF67F1989B7BFE73.taxon	discussion	Remarks. This species is similar to D. piscicercus by having the paramere entirely wide and the basiparamere very wide. However, this species has the tergal process with tubercle, small and shallow depression, the aedeagal ventral ramus with simple apex and the apex of the basal bar straight with two laterad teeth, whereas D. piscicercus has the tergal process with ovoid, deep, large depressions nearly touching the posterior margin of the tergite I, with pit, tuft of long setae, the aedeagal ventral ramus with two large apical teeth and the basal bar without two laterad teeth.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C529B06FF67F1989B7BFE73.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Chi [riqui]: Hartmans Finca, 28. VI – 3. VII. 1981 (PMAE). Paratypes: Bocas del Toro: 7 ♂, Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP); 5 ♂, same data as holotype; 6 ♂ same data as holotype except 3 – VII 1981, B. Gill [col.] (PMAE); 1 ♂, same province of holotype, Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 2 – 6. VI. 2000, F [light] I [nterception] T [rap], H. & A. Howden [col.] (CNCI); 1 ♂, same province of holotype, 5,6 km N Boquete, La Culebra trail, [08 ° 49 ' 23 " N, 82 ° 25 ' 18 " W], F [light] I [nterception] T [rap], J. Ashe & R. Brooks [col.] (CNCI); 2 ♂, same province of holotype, Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 25 – 6. VI. 2000, F [light] I [nterception] T [rap], H. & A. Howden [col.] (CNCI); [Los Santos]: 2 ♂, Ojo de Agua, [07 ° 31 ' N, 80 ° 25 ' W], 30. VI – 14. VII. 1982, B. Gill [col.] (PMAE).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C529B06FF67F1989B7BFE73.taxon	distribution	Distribution (Fig. 244). Panama (Bocas del Toro, Chiriqui and Los Santos).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C519B06FF67F4329AF8F82A.taxon	diagnosis	Diagnosis. Male. Mandible tridentate, uppermost tooth usually inconspicuous. Median clypeal lobe trapezoidal except D. afestus sp. nov.. Metasoma with sublateral tergal process, wich have a pair of circular depressions with dense tuft of convergent setae directed backward, antero-lateral margin of depression with setae, tubercle absent. Genitalia with inner apical surface of paramere generally with one setae distinctly longer than others. Aedeagal ventral ramus shorter than dorsal body with at least half part inclined and tending to be convergente mesad of genitalia. Aedeagal dorsal body wide with crown-like structure.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C519B06FF67F4329AF8F82A.taxon	discussion	Remarks. It has fourteen species: D. aculetaus Alencar & Azevedo, D. auritus Alencar & Azevedo, D. coronatus, Alencar & Azevedo, D. culteratus Alencar & Azevedo, D. declinatus Alencar & Azevedo, D. decussatus Alencar & Azevedo, D. flexuosus Alencar & Azevedo, D. galeatus, Alencar & Azevedo, D. hirtus, Alencar & Azevedo, D. pronus, Alencar & Azevedo, D. ramosus Alencar & Azevedo, D. sinatus Alencar & Azevedo, D. uber Alencar & Azevedo and D. afestus sp. nov..	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C519B06FF67F4329AF8F82A.taxon	distribution	Distribution. Dominica, Costa Rica, Panama, Venezuela, Trinidad & Tobago, Brazil (Paraná), Ecuador and Peru.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C519B06FF67F3C09C56FA60.taxon	description	Description. Male. Body length 3.70 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 15). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc coriaceous; anterior margin ecarinate. Metasoma (Fig. 46). Tergal process with lateral, shallow, circular and small depression, with small, very low and dorsad tubercle placed on inner area of depression, pit small, with short and sparse tuft of setae. Posterior hypopygeal margin weakly concave (Fig. 77). Genitalia (Figs 154 – 156). Paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, entirely wide; cross section laminar; surface vertical; inner margin slightly sinuous with small projection in apical portion; outer margin slightly sinuous; apex long, simple, parallel; additional inner ramus long; basal stub short and narrow. Aedeagal dorsal body with pair of apical lobe; outer lobe long, subvertical and narrow, with apical margin rounded and ventrad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring. Variations. Body length 3.07 mm (min.) 4.45 mm (max). Tooth of median clypeal lobe rounded; anterior margin of pronotal disc coarse; depression of tergal process very shallow and pit of tubercle large.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C519B06FF67F3C09C56FA60.taxon	discussion	Remarks. This species is similar to D. ardonius sp. nov. by having spine in the ventral margin of the apical lobes of the aedeagal dorsal body and the ventral ramus entirely wide with the inner margin sinuous. However, this species has the dorsal margin of the paramere straight, the apex of the basal bar projected forward and the apical margin of the aedeagal ventral ramus smooth, whereas D. ardonius sp. nov. has the dorsal margin of the paramere sinuous, the apex of the basal bar not projected forward and the apical margin of the aedeagal ventral ramus wavy.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C519B06FF67F3C09C56FA60.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Panama Pr [ovincia]: P [arque] Nac [ional] Altos de Campana, [08 º 43 ' 15 " N, 80 º 01 ' 43 " W], 10 – 14 agos 1999, Malaise, P. González, A. Santos [col.] (MIUP). Paratypes: 1 ♂ same data as holotype; Darién: 4 ♂, Serrania de Pierre, Cana, 1380 m, [07 ° 51 ' N, 77 ° 37 ' W], 4 – 7. VI. 1986, F [light] I [nterception] T [rap], [J.] Ashe & [R.] Brooks [col.] (2 ♂) (CNCI) and 1250 m, 4 – 7. vi. 1996, (2 ♂) (UFES).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C519B06FF67F3C09C56FA60.taxon	distribution	Distribution (Fig. 246). Panama (Panama and Darién).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C509B00FF67F5659DC1FC5F.taxon	description	Description. Male. Body length 2.70 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 16). Mandible with three distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina low in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc coriaceous; anterior margin carinate. Metasoma (Fig. 47). Tergal process lateral, without depression, without tubercle, with short tuft of setae. Posterior hypopygeal margin straight (Fig. 78). Genitalia (Figs 157 – 158): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface subhorizontal; inner margin concave; outer margin sinuous; apex long, simple, slightly curved inward; additional inner ramus absent; basal stub long and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe horizontal and wide, with apical margin angled and laterad; inner pair stout, membranous and setose; crown-process very long. Apodeme not extending beyond genital ring. Variations. Pronotal disc coarse; tergal process with very short tuft of setae, without depression.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C509B00FF67F5659DC1FC5F.taxon	discussion	Remarks. This species is easily recognized by having the genitalia wide, the apical lobes of the aedeagal dorsal body very long and the inner lobe of the aedeagal dorsal body very big, extending through the outer lobes of the aedeagal dorsal body. This species is similar to D. culteratus by having apical lobes of aedeagal dorsal body long with apex laterad and paramere longer than basiparamere. However, this species has the aedeagal ventral ramus with inner margin smooth, basal half wide and apical half narrow with apex mesad, whereas D. culteratus has the aedeagal ventral ramus narrowing at apex, inner margin microserrated and apex parallel.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C509B00FF67F5659DC1FC5F.taxon	materials_examined	Material examined. Holotype ♂, PANAMA, Darién Pr [ovincia]: P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 1000 m, [08 ° 00 ' N, 77 ° 45 ' W], 21 mar – 4 ab 2000, Malaise, Cambra, Santos, Bermudez [col.] (MIUP). Paratypes: 4 ♂, same data as holotype.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C509B00FF67F5659DC1FC5F.taxon	distribution	Distribution (Fig. 225). Panama (Darién).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C579B00FF67F1EB9A5EF969.taxon	diagnosis	Diagnosis. Mandible tridentate. Clypeus trapezoidal. Tergal process with pair of circular, large and deep lateral depressions with dense tuft of convergent setae directed backward. Hypopygium with posterior margin with median convexity. Genitalia: paramere wide, dorsal margin straight and ventral margin concave; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; surface horizontal, angulate medially laterad; basal half narrow and convergent mesad; aedeagal dorsal body laminar, surface vertical, base wide narrowing to apex; apex rounded with sinuous and serrate crest-like projection, directed ventrally; crown-process small. Apodeme not extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C579B00FF67F1EB9A5EF969.taxon	discussion	Remarks. This species is known to Costa Rica, Panama, Venezuela, Dominica, Trinidad & Tobago, Ecuador (Alencar & Azevedo 2006). In this series the specimens are quite similar to those previously studied, without any variation that worths to mention.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C579B00FF67F1EB9A5EF969.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Veraguas: Coiba N [acional] P [arque], Isla Coibita [Rancheria], [7 º 38 ’ 37 ” N 81 º 42 ’ 15 ” W], 29. VII. – 4. VIII. 1998, Malaise trap, R. Cambra & A. Santos col. (MIUP). PANAMA, Veraguas Pr [ovincia]: 13 ♂, P [arque] N [acional] Coiba, Isla Coibita, [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 27 – nov – 4 - dic. 1998, (5 ♂), 0 m, 30 – nov. – 4 dic. 1998 (5 ♂) and 120 m (3 ♂) Malaise, R. Cambra, S. Bermudez [col.] (MIUP); Darién Pr [ovincia]: 2 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C579B00FF67F1EB9A5EF969.taxon	distribution	Distribution (Fig. 231). Costa Rica, Panama, Venezuela, Dominica, Trinidad & Tobago and Ecuador.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C579B01FF67F5369DD0FD80.taxon	diagnosis	Diagnosis. Mandible tridentate. Clypeus trapezoidal. Tergal process with pair of circular, large and deep lateral depressions with dense tuft of convergent setae directed backward. Hypopygium with posterior margin with median convexity. Genitalia: paramere with dorsal margin straight and ventral margin concave; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar, surface horizontal and arched ventrally; basal half wide; inner margin sinuous; apical margin with outer margin straight and inner margin microserrate; aedeagal dorsal body wide, narrowing to apex; apical part with concavity mesad, apex with two arms; crown-process with long lateral prolongation in middle of aedeagal dorsal body. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C579B01FF67F5369DD0FD80.taxon	discussion	Remarks. This species is known to Panama (Alencar & Azevedo). In this series, the specimens are quite similar to those previously studied, without any variation that worths to mention.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C579B01FF67F5369DD0FD80.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Chiriquí: Hatman’s Farm, [8 º 51 ’ N 82 º 46 ’ W], 3. VII. 1981, B. Gill col (PMAE); PANAMA, Chiriqui Pr [ovincia]: 2 ♂, P [arque] I [nternacional] L [a] A [mistad], Jurutungo, 1800 m, [08 ° 54 ' N, 82 ° 44 ' W], 30 oci. 5 nov. 1994, Tr [ampa] Malaise, A. Rodriguez [col.] (MIUP); 9 ♂, 5.6 km N Boquete, La Culebra trail, [08 ° 49 ' 23 " N, 82 ° 25 ' 18 " W], (8 ♂) and [08 ° 54,7 ' N, 82 ° 47,3 W], 17 – 18. vi. 1996, 1875 m (1 ♂), F [light] I [nterception] T [rap], J. Ashe & R. Brooks [col.] (CNCI); [Los Santos]: 1 ♂, Ojo de Agua, [07 ° 31 ' N, 80 ° 25 ' W], 30. VI – 14. VII. 1982, B. Gill [col.] (PMAE).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C579B01FF67F5369DD0FD80.taxon	distribution	Distribution (Fig. 229). Costa and Panama.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C569B01FF67F08B9DB4FB45.taxon	diagnosis	Diagnosis. Mandible tridentate. Clypeus trapezoidal. Tergal process with pair of circular, large and deep lateral depressions, with tuft of setae convergent directed backward. Hypopygium with posterior margin straight. Genitalia: paramere with apex straight; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; surface horizontal; basal half long; inner margin sinuous; outer margin straight; apical half narrow dorsally and converging mesad; aedeagal dorsal body laminar; surface vertical; apex wide and inclined ventrally; crown-process small; apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C569B01FF67F08B9DB4FB45.taxon	discussion	Remarks. No additional material of this species was not found in this study.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C569B01FF67F08B9DB4FB45.taxon	materials_examined	Material examined. Holotype, ♂, ECUADOR, Napo: Huahua Sumaco, 44 km on Hollin Loreto R [oa] d, [0 º 37 ’ S 77 º 35 ’ W], 15 – 22. XII. 1989, M [alaise] T [rap], H. Real, M. and J. Wasbauer col. (UCDC);	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C569B01FF67F08B9DB4FB45.taxon	distribution	Distribution (Fig. 233). Panama and Ecuador.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C569B01FF67F6D29CADF810.taxon	diagnosis	Diagnosis. Mandible tridentate. Clypeus trapezoidal, median carina complete and strongly developed in profile. Tergal process with pair of circular, large and deep lateral depressions with dense tuft of convergent setae directed backward. Hypopygium with posterior margin straight. Genitalia: paramere wide, dorsal margin straight and ventral margin concave; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; surface horizontal, and angulate ventrally; basal half wide; inner margin with strong concavity before apex; outer margin convex above; aedeagal dorsal body laminar, surface vertical; base wide, narrowing to rounded apex; ventral margin with long projection directed to apical part of aedeagal ventral ramus; dorsal margin sinuous and serrate; crown-process small. Apodeme not extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C569B01FF67F6D29CADF810.taxon	discussion	Remarks. This species is known to Costa Rica and Brazil (Rondônia), and now it is recorded for the first time from Panama. Some variations are worth to mention: median clypeal tooth shorter and angulate; median carina of clypeus complete apically; depression of tergal process very shallow.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C569B01FF67F6D29CADF810.taxon	materials_examined	Material examined. Holotype, ♂, COSTA RICA, [Puntarenas]: San Vito de Coto Brus, Las Cruces, 1200 m, [8 º 28 ’ 12 ” N 82 º 34 ’ 12 ” W], 17. VIII – 12. IX. 1982, Malaise Trap, B. Gill col. (CNCI); PANAMA, Chiriqui: 3 ♂, Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 25 – 6. VI. 2000, F [light] I [nterception] T [rap], H. & A. Howden [col.] (CNCI); 4 ♂, 15 km NW Hato del Volcano, 1200 m, [08 ° 46 ' N, 82 ° 38 ' W], 24 – 31. V. 1997, Peck & Howden [col.] (CNCI).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C569B01FF67F6D29CADF810.taxon	distribution	Distribution (Fig. 234). Costa Rica, Panama and Brazil (Rondônia).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C559B02FF67F4859A1AF859.taxon	diagnosis	Diagnosis. Male. Metasoma with tergal process median, large, deep, triangular depression with base toward; pair of very large lateral pits, arched inward medially, covered by a lot of long setae inward.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C559B02FF67F4859A1AF859.taxon	discussion	Remarks. It has a single species, D. curvifoveatus.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C559B02FF67F4859A1AF859.taxon	distribution	Distribution. Belize, Costa Rica, Panama, Venezuela, Ecuador and Brazil (Amazonas, Pará and Rondônia).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C559B02FF67F2A89D8AFBD6.taxon	diagnosis	Diagnosis. Mandible tridentate. Clypeus trapezoidal. Tergal process with pair of circular, large and deep lateral depressions with dense tuft of convergent setae directed backward. Hypopygium with posterior margin convex. Genitalia: paramere wide; dorsal margin straight and ventral margin concave; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; surface horizontal and angulate ventrally; basal half long and wide; inner margin sinuous; apical half narrowing to apex and converging mesad; apex folded laterad; aedeagal dorsal body wide; surface vertical; apical part with concavity mesad, apex with two arms; crown-process small. Apodeme not extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C559B02FF67F2A89D8AFBD6.taxon	discussion	Remarks. This species is known to Costa Rica (Alencar & Azevedo 2006) and now it is recorded for the first time from Panama. Some variations are worth to mention: head, mesosoma and metasoma castaneous; clypeal carina not developed in profile; anterior margin of pronotal disc carinate and followed by foveae; apex of aedeagal ventral ramus folded.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C559B02FF67F2A89D8AFBD6.taxon	materials_examined	Material examined. Holotype, ♂, COSTA RICA, Monteverde: 1520 m, 25. VI – 2. VII. 1983, D. H Lindeman col. (PMAE); PANAMA, Colón: 1 ♂, 14 km N J [un] ct [ion] Escobal r [oa] d and Piña r [oa] d on Piña r [oa] d, ± 20 m, [09 ° 20 ' N', 79 ° 54 ' W], 2 – 3. VI. 1995, F [light] I [nterception] T [rap], J. Ashe & R. Brooks [col.] (CNCI); Coche {= Coclé}: 1 ♂, El Cope {= El Copé}, [08 ° 37 ' 11 " N, 80 ° 35 ' 03 " W], 19. XI. 1994, Windson & Edwards [col.] (CNCI); Panama: 1 ♂, Cerro Campana, 08 ° 40 ' N, 79 ° 50 ' W, 7 – 14. V. 1973 (CNCI); 1 ♂, C [anal] Z [one], B [arro] Colorado Is [land], [09 ° 09 ' N, 79 ° 51 ' W], 1 – 14. VII. 1994, F [light] I [nterception] T [rap], D. Banks [col.] (CNCI); Darién Pr [ovincia]: 58 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP); 2 ♂, Cana, 7 ° 45 ' N, 77 ° 41 ' W, 3 – 7. VI. 96, 550 m, F [light] I [nterception] T [rap] J. Ashe, R. Brooks (UFES).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C559B02FF67F2A89D8AFBD6.taxon	distribution	Distribution (Fig. 239). Costa Rica and Panama.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C559B02FF67F6629CBFF9B8.taxon	diagnosis	Diagnosis. Mandible tridentate. Clypeus trapezoidal. Tergal process with pair of circular, inclined and deep lateral depressions with tuft of setae directed backward. Hypopygium with posterior margin convex. Genitalia: paramere dorsal margin straight; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; surface horizontal; basal half wide; inner margin irregular; apical half narrowing to apex; aedeagal dorsal body wide; surface vertical; apical part narrow and mesad; apex with three arms; crown-process with long lateral prolongation in middle of aedeagal dorsal body. Apodeme not extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C559B02FF67F6629CBFF9B8.taxon	discussion	Remarks. No additional material of this species was not found in this study.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C559B02FF67F6629CBFF9B8.taxon	materials_examined	Material examined. Holotype, ♂, ECUADOR, Pichincha: 47 km S. S [an] to Domingo, Rio Palenque Res [erve] Sta [tion], 160 – 180 m, rainforest, [0 º 30 ’ S 79 º 9 ’ W], 29. IV – 5. V. 1987, 2 MT, B. V, Brown and L. D Coote col. (PMAE).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C559B02FF67F6629CBFF9B8.taxon	distribution	Distribution (Fig. 241). Costa Rica, Panama, Venezuela and Ecuador.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C549B03FF67F2A89E70FC05.taxon	diagnosis	Diagnosis. Mandible bidentate. Clypeus subtrapezoidal. Tergal process median with wide, deep and triangular depression, with very large pit densely covered with setae. Hypopygium with posterior margin with very short invagination. Genitalia: paramere bidigitiform with median longitudinal invagination very deep; volsella with digitus very long 0.5 X the length of paramere; aedeagal dorsal ramus longer than aedeagal dorsal body, surfaces vertical; apex narrow and smooth; inner margin excavated medially and straight basally; outer margin convex; aedeagal dorsal body with two pairs of apical lobes; ventral pair narrow, lanceolate with apex sharpened; dorsal pair narrowing to apex divergent, with two teeth. Apodeme not extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C549B03FF67F2A89E70FC05.taxon	discussion	Remarks. This species is known to Belize, Costa Rica, Panama, Venezuela, Ecuador and Brazil (Acre, Amazônia, Pará, Rondônia and Paraná) (Azevedo 1999 a) and here more specimens are added. Some variations are worth to mention: median clypeal tooth angulate; anterior margin of pronotal disc coarse; one specimen with depressions of tergal process with few setae and tergal process more distant from each other.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C549B03FF67F2A89E70FC05.taxon	materials_examined	Material examined. Holotype, ♂, BRAZIL, Amazonas: Reserve 1112, 8. xi. 1985, Malaise trap, B. Klein col. (INPA); PANAMA, [Panama Province]: 1 ♂, Nusagandi Sta [tion] n [ea] r Columbia INA Igar Trail, [08 ° 58 ' N, 79 ° 32 ' W], 12 – 13. V. 1994, A. Windson [col.] (CNCI); Darién Pr [ovincia]: 3 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov 00 – 17 ene 2001, [Trampa] Malaise, R. Cambra, A. Santos [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C549B03FF67F2A89E70FC05.taxon	distribution	Distribution (Fig. 232). Belize, Costa Rica, Panama, Venezuela, Ecuador and Brazil (Amazonas, Pará, Rondônia).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C549B03FF67F6199CADFAC9.taxon	diagnosis	Diagnosis. Male. Clypeus broadly projected forward. Metasoma with lateral tergal process, tufts of setae with most lateral setae longer than others.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C549B03FF67F6199CADFAC9.taxon	discussion	Remarks. It has five species: D. gilvipes Evans, D. krombeini Azevedo, D. altyclypeatus Azevedo, D. bicerutus Azevedo, and D. arbeius sp. nov..	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C549B03FF67F6199CADFAC9.taxon	distribution	Distribution. Ranging from Southern Florida to northern Argentina.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C549B0CFF67F7639CF1FD77.taxon	description	Description. Male. Body length 2.00 mm. Color: body castaneous. Head (Fig. 17). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina low in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 48). Tergal process, with lateral, shallow, linear and small depression, without tubercle, pit absent, with very short tuft of setae with one distinctly long seta. Posterior hypopygeal margin weakly concave (Fig. 79). Genitalia (Figs 165 – 166): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin slightly sinuous; apex short, simple, angled outward; additional inner ramus absent; basal stub short and very narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, subvertical and wide with apical margin abruptly acute and dorsad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring. Variation. Head, mesosoma and metasoma light castaneous.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C549B0CFF67F7639CF1FD77.taxon	discussion	Remarks. This species is similar to D. gilvipes by having the dorsal margin of the paramere expanded distally and the aedeagal ventral ramus abruptly narrowing apicad. However, this species has the apical lobes of the aedeagal dorsal body short and wider and the apical margin of the paramere crenulate, whereas D. gilvipes has the apical lobes of the aedeagal dorsal body long and narrow and the apical margin of the paramere smooth.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C549B0CFF67F7639CF1FD77.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Chiriqui: 2 km W Cerro Punta, 1700 m, [08 ° 51 ' N, 82 ° 35 ' W], 19. V – 8. VI. 1977, Peck & Howden [col.] (CNCI). Paratypes: 12 ♂, same data of holotype; 1 ♂, STRI n [ea] r Futurna {= Fortuna}, 1260 m, 8 ° 43 ' 34 " N, 82 ° 14 ' 25 " W, 30. V. 2000, Y [ellow] P [an] T [rap], B. & J. Gill [col.] (CNCI); 15 ♂, P [arque] I [nternacional] L [a] A [mistad], Jurutungo, 1800 m, [08 ° 54 ' N, 82 ° 44 ' W], 30 oct. 5 nov 1994, Tr [ampa] Malaise (13 ♂) and Tr [ampas] Amarillas (2 ♂) A. Rodriguez [col.] (MIUP); 2 ♂, 15 km NW Hato del Volcano, 1200 m, [08 ° 46 ' N, 82 ° 38 ' W], 24 – 31. V. 1997, Peck & Howden [col.] (CNCI); 3 ♂, Fortuna, Div. Cont. Chiriqui- B [ocas del] Toro, 1070 – 1090 m, [08 ° 46 ' 16.5 " N, 82 ° 11 ' 27.3 " W], 8 – 11. abr. 1999, Tr [ampas] Amar [illas], R. Cambra, A. Santos [col.] (MIUP); Coclé Pr [ovincia]: 1 ♂, El Copé, Par [que] Nac [ional General de División] Omar Torrijos H [errera], 570 – 670 m. s. n. m., [08 º 41 ' 56 " N, 80 º 39 ' 17 " W], 26 – 30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); Panama Pr [ovincia]: 4 ♂, P [arque] Nac [ional] Altos de Campana, [08 º 43 ' 15 " W, 80 º 01 ' 43 " W], 10 – 14 ago 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); 2 ♂, P [arque] Nac [ional] Sob [eranía], Camino, Plantación, [09 º 09 ' N, 79 º 43 ' W], 24 ago. – 2 sep. 1999 (MIUP); 1 ♂, Cerro Azul, Las Nubes, [09 º 10 ' N, 79 º 25 ' W], 9 – 15 – oct. 1999, R. Cambra, A. Santos [col.] (MIUP); 2 ♂, Cerro Campana, 08 ° 40 ' N, 79 ° 50 ' W, 7 – 14. V. 1973 (CNCI).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C549B0CFF67F7639CF1FD77.taxon	distribution	Distribution (Fig. 228). Panama (Chiriqui, Coclé and Panama).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5B9B0CFF67F0C39C8EFA28.taxon	diagnosis	Diagnosis. Mandible tetradentate. Clypeus broadly projected toward with three small teeth. Tergal process lateral, with pair of subcircular depressions with distinct long lateral setae. Hypopygium with posterior margin straight, with small rounded median projection. Genitalia: paramere with dorsal margin developed inward; aedeagal ventral ramus as long as aedeagal dorsal body, laminar with surfaces vertical; basal half wide and apical narrow; apex blunt and oblique; aedeagal dorsal body with pair of parallel lobe, narrowing apically; inner lobe setose; base of genitalia with pair of laminar, rectangular expansion. Apodeme not extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5B9B0CFF67F0C39C8EFA28.taxon	discussion	Remarks. This species is known to Panama, Colombia, Ecuador, Peru, Brazil (Amazonas, Para, Rondônia, Mato Grosso, Pernambuco, Minas Gerais, Distrito Federal, São Paulo and Espírito Santo) (Colombo & Azevedo 2016) and here two specimens are added. Some variations are worth to mention: outer lobe of apical lobes of aedeagal dorsal body entirely narrow, laminar in lateral view, convergent apically; aedeagal ventral ramus with apex slightly laterad.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5B9B0CFF67F0C39C8EFA28.taxon	materials_examined	Material examined. Holotype ♂, COLOMBIA, Dept. Valle: Central de Anchicaya, 30 km E Buenaventura, 560 m, 14 – 16 July 1975, R. Wilkerson, Malaise trap (FSCA). PANAMA, Chiriqui: 2 #, 15 km NW Hato del Volcano, 1200 m, [08 ° 46 ' N, 82 ° 38 ' W], 24 – 31. V. 1997, Peck & Howden [col.] (CNCI).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5B9B0CFF67F0C39C8EFA28.taxon	distribution	Distribution (Fig. 235). Panama, Colombia, Ecuador, Peru, Brazil (Amazonas, Para, Rondônia, Mato Grosso, Pernambuco, Minas Gerais, Distrito Federal, São Paulo and Espírito Santo).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5B9B0CFF67F4759DAFF8C9.taxon	diagnosis	Diagnosis. Male. Metasoma with lateral tergal process with pair of lateral drop-shaped pits, high rim, sharpened anteriorly and rounded posteriorly.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5B9B0CFF67F4759DAFF8C9.taxon	discussion	Remarks. It has two species: D. guttus Azevedo and D. latimerus Azevedo. Distribution. Costa Rica, Panama and Brazil.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5A9B0EFF67F53A9D8AFCE8.taxon	diagnosis	Diagnosis. Mandible tridentate. Clypeus subtrapezoidal. Tergal process with pair of droped-shaped pits with outer rim, sharpened anteriorly and rounded posteriorly, with tuft of setae backward and very shallow depression. Hypopygium with posterior margin straight. Genitalia: paramere wide, apex rounded; aedeagal ventral ramus shorter than aedeagal dorsal body; apical half narrow and strongly curved downward; aedeagal dorsal body with two pairs of apical lobes; outer pair laminar; surface vertical, wide with rounded apex in lateral view; apex tridentate, directed downward; ventral margin with anterior elongate basal tooth; inner pair membranous, stout and setose; base of aedeagus strongly dilated upward. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5A9B0EFF67F53A9D8AFCE8.taxon	discussion	Remarks. This species is known to Costa Rica and Panama (Azevedo 2003) and here more specimens are added. Some variations are worth to mention: median clypeal tooth angulate; median clypeal carina incomplete apically; anterior margin of pronotal disc coarse.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5A9B0EFF67F53A9D8AFCE8.taxon	materials_examined	Material examined. Holotype, ♂, COSTA RICA, Sant Vito de Coto Brus: Las Cruces, 1200 m, 9. VII. 1982, Malaise trap, B. Gill col. (CNCI). PANAMA, Bocas del Toro: 4 ♂, Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP); Chiriqui: 6 ♂, Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 25 – 6. VI. 2000, F [light] I [nterception] T [rap], H. & A. Howden [col.] (CNCI); 1 ♂, 15 km NW Hato del Volcano, 1200 m, [08 ° 46 ' N, 82 ° 38 ' W], 24 – 31. V. 1997, Peck & Howden [col.] (CNCI); Veraguas Pr [ovincia]: 1 ♂, Calovebora, Guazarito, 15 – 18 feb. 2000, Malaise, Col. S. Bermudez (MIUP); Darién Pr [ovincia]: 7 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP); 1 ♂, Est [ación] I [nstituto] N [acional de] Re [cusos] Na [turales] Re [novables], [07 ° 51 ' 26.3 " N, 77 ° 28 ' 12.7 " W], 5 – 28 feb 1993, R. Cambra [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5A9B0EFF67F53A9D8AFCE8.taxon	distribution	Distribution (Fig. 236). Costa Rica and Panama.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C599B0FFF67F7BD9C29FE2A.taxon	diagnosis	Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with pair of minute tubercle with small pit on top, and few setae directed backward. Hypopygium with posterior margin concave. Genitalia: paramere very wide and apex rounded; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; surface horizontal; inner margin straight outer margin convex; filament inserted subapically; aedeagal dorsal body with two pairs of apical lobes; outer pair laminar; surface vertical and wide; apex straight; inner pair irregular in lateral view, membranous, setose and granulate in ventral view; basal bar triangular; conector with lateral expansion concave. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C599B0FFF67F7BD9C29FE2A.taxon	discussion	Remarks. This species is known to Trinidad and Tobago (Alencar & Azevedo 2008). Now it is recorded for the first time from Panama. Some variations are worth to mention: apical lobes of aedeagal dorsal body very projected forward and apical margin truncate.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C599B0FFF67F7BD9C29FE2A.taxon	materials_examined	Material examined. Holotype, ♂, TRINIDAD [and TOBAGO]: [Saint George]: 16 km N Arima, Andrews Trace, 620 m up. Montane, rain for [est], [~ 10 º 30 ’ N 61 º 16 ’ W], 7 – 24. vi. 1993. FIT, S. & J. Peck col. (CNCI). PANAMA, Veraguas Pr [ovincia]: 12 ♂, P [arque] N [acional] Coiba, Isla Coiba, [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 27 – nov – 4 – dic. 1998 (1 ♂), Cerro la Equis, 130 m, 8 – 12 mar [zo] 1998 (1 ♂), 29 jul – 4 ag 1998, 100 m (1 ♂), 120 m (1 ♂), Malaise, R. Cambra, A. Santos [col.]; 8 - 12 mar [zo] 1998, 0 m (1 ♂), 90 m (1 ♂), 30 – nov – 4 – dic. 1998, 100 m (1 ♂), 120 m (1 ♂), 0 m (1 ♂), R. Cambra, S. Bermudez [col.]; Est [ación] Biol [ogica] Coiba, 26 nov – 4 dic 1998 (1 ♂) 5 - 6 ag 1998 (1 ♂), R. Cambra, A. Santos [col.] (MIUP); Panama Pr [ovincia]: 5 ♂, P [arque] Nac [ional] Sob [eranía], Camino, Plantación, [09 º 09 ' N, 79 º 43 ' W], 24 ago. – 2 sep. 1999, Malaise, A. Santos, P. González [col.] (MIUP); 3 ♂, P [arque] N [atural] Metropolitano, [08 º 59 ' 33 " N, 79 º 32 ' 41 " W], 24 ago. – 3 sep. 1999, Malaise, A. Santos, P. González [col.] (MIUP); 1 ♂, Cerro Azul, Las Nubes, [09 º 10 ' N, 79 º 25 ' W], 9 – 15. oct. 1999, R. Cambra, A. Santos [col.] (MIUP); Coclé: 1 ♂, Falda Cerro La Vieja, Chiguirí Arriba, [08 º 40 ' 11 " N, 82 º 12 ' 05 " W], 24 abril 1994, A. Rodriguez, R. Cambra [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C599B0FFF67F7BD9C29FE2A.taxon	distribution	Distribution (Fig. 239). Panama and Trinidad & Tobago.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C589B0FFF67F44B9B83F8BC.taxon	diagnosis	Diagnosis. Male. Tergal process with circular depression, dense tufts of setae directed backward, tubercle absent. Remarks. It has four species: D. napo Evans, D. punctatus, D. bahiensis Redighieri & Azevedo and D. tupinikim Colombo & Azevedo.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C589B0FFF67F44B9B83F8BC.taxon	distribution	Distribution. Costa Rica, Panama, Ecuador, Peru, Brazil (Amazonas, Acre, Sergipe, Alagoas, Pernambuco, Paraíba, Mato Grosso, Bahia, Espírito Santo, Rio de Janeiro, São Paulo and Paraná) and Paraguay.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C589B0FFF67F0789CF0F9F4.taxon	description	Description. Male. Body length 2.27 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 18). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina low in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest strongly concave. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 49). Tergal process absent. Posterior hypopygeal margin weakly concave (Fig. 80). Genitalia (Figs 169 – 170): paramere with dorsal margin wide basally; apical margin rounded. Aedeagal ventral ramus higher than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin excavated; outer margin slightly sinuous; apex long, simple, parallel; additional inner ramus absent; basal stub short and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, subhorizontal and narrow, with apical margin acute and mesad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring. Variations. Mesosoma dark castaneous; median clypeal lobe short.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C589B0FFF67F0789CF0F9F4.taxon	discussion	Remarks. This species is similar to D. isortus sp. nov. by having the aedeagal dorsal body narrowing slightly apicad, the apical lobes of the aedeagal dorsal body in conical-shaped. However, this species has the tergal process absent, the paramere narrow, the apex of the apical lobes of the aedeagal dorsal body mesad and the aedeagal ventral ramus with filament inserted in the median part, characteristic diagnosis for microsticus species-group, whereas D. isortus sp. nov. has the tergal process with a tubercle, pit with thick setae the paramere wide, the apex of the apical lobes of the aedeagal dorsal body dorsad and the aedeagal ventral ramus without filament.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C589B0FFF67F0789CF0F9F4.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Bocas del Toro: Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ap] Malaise, A. Santos [col.] (MIUP). Paratypes: 8 ♂, same data as holotype; Chiriquí Pr [ovincia]: 1 ♂, P [arque] I [nternacional] L [a] A [mistad], Jurutungo, 1800 m, [08 ° 54 ' N, 82 ° 44 ' W], 30 oct. 5 nov. 1994, Tr [ampa] Malaise, A. Rodriguez [col.] (MIUP); 1 ♂, 15 km NW Hato del Volcano, 1200 m, [08 ° 46 ' N, 82 ° 38 ' W], 24 – 31. V. 1997, Peck & Howden [col.] (CNCI); 1 ♂, Fortuna: Div. Cont. Chiriqui-B [ocas del] Toro, 1070 – 1090 m, [08 ° 46 ' 16.5 " N, 82 ° 11 ' 27.3 " W], 8 – 11. abr. 1999, Malaise, R. Cambra, A. Santos [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C589B0FFF67F0789CF0F9F4.taxon	distribution	Distribution (Fig. 241). Panama (Bocas del Toro and Chiriquí).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5F9B08FF67F2A89B70FB39.taxon	diagnosis	Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with very shallow, small and rounded depressions covered with dense tuft of setae directed backward. Hypopygium with dorsal margin straight. Genitalia: paramere with rounded apex with dorsal margin straight; aedeagal ventral ramus slightly shorter than aedeagal dorsal body, sinuous and wide; with apical filament directed outward; aedeagal dorsal body with two pairs of apical lobes; dorsal lobe with filament with sharpened apex, inner surface setose, vertical; ventral lobe longer, membranous, wide with rounded apex in lateral view, surfaces vertical; inner lobe densely setose. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5F9B08FF67F2A89B70FB39.taxon	discussion	Remarks. This species is known to Costa Rica, Panama, Ecuador, Peru and Brazil (Amazonas, Acre, Pernambuco, Alagoas, Sergipe, Bahia, Espírito Santo, Rio de Janeiro, São Paulo and Paraná). Now some specimens are added. Some variations are worth to mention: median clypeal tooth angulate; median clypeal carina straight and complete apically; anterior margin of pronotal disc coarse; tergal process with very shallow depression; paramere short, wider and apical margin straight; basivolsella with a rounded expansion.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5F9B08FF67F2A89B70FB39.taxon	materials_examined	Material examined. Holotype ♂, PERU, Cuzco: Cajon, Bergland, 1500 m, 11. I. 1901, Garlepp col. (ZMHB, # 195); PANAMA, Chiriqui: 1 ♂, Santa Clara, Finca Hartmann, Ojo de Agua, [08 ° 51 ' 38 " N, 82 ° 44 ' 44 " W], 8 – 9. XI. 2001, M [alaise] T [rap], N. Schiff [col.] (UCDC); 1 ♂ Res [erva Florestal] La Fortuna, Viveno, [08 ° 44 ' 39 " N, 82 ° 14 ' 15 " W], 14 – 18. VI. 1994, A. R. Gillogly [col.] (CNCI); Darién Pr [ovincia]: 28 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (25 ♂) and 1000 m, 21 mar – 4 ab 2000, Cambra, Santos, Bermudez [col.] (2 ♂) (MIUP); 4 ♂, Serrania de Pierre, Cana, 1380 m, [07 ° 51 ' N, 77 ° 37 ' W], 4 – 7. VI. 1986 F [light] I [nterception] T [rap], [J.] Ashe & [R.] Brooks [col.] (2 ♂) (CNCI) and 1450 m, 4 – 7. vi. 1995 (2 ♂) (UFES).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5F9B08FF67F2A89B70FB39.taxon	distribution	Distribution (Fig. 242). Costa Rica, Panama, Ecuador, Peru, and Brazil (Amazonas, Acre, Pernambuco, Alagoas, Sergipe, Bahia, Espírito Santo, Rio de Janeiro, São Paulo and Paraná).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5F9B08FF67F7069CACF9F7.taxon	diagnosis	Diagnosis. Male. Metasoma with median tergal proces, with very shallow, very large, depression occupying two median thirds of tergite, reaching broadly posterior margin of tergite I; half-moon-shaped. Paramere deeply divided medially forming two distinct lobes.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5F9B08FF67F7069CACF9F7.taxon	discussion	Remarks. It has two species: D. rasissimus Azevedo and D. bilobatus Azevedo. Distribution. Mexico (Chiapas and Oaxaca), Belize and Costa Rica.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5F9B09FF67F4439D8AFC33.taxon	diagnosis	Diagnosis. Mandible oblique without teeth. Tergal process with very large depression hlaf-moon-shaped, with two low median tubercles and tuft of setae. Hypopygium with posterior margin slightly convex. Genitalia: paramere with deep median longitudinal invagination, dorsal lobe larger, apex wide, concave with corners rounded; ventral lobe thin, small, apex with setae; aedeagal ventral ramus shorter than aedeagal dorsal body, wide with apex truncate; surfaces divergent dorsally; ventral part of apex divided longitudinally with two serrated lamina; outer lamina small and narrowed apically and inner lamina larger; aedeagal dorsal body not developed narrowing apically; inner margin straight and parallel and outer margin convergent. Apodeme not extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5F9B09FF67F4439D8AFC33.taxon	discussion	Remarks. This species is known to Costa Rica (Azevedo 1999). Now it is recorded for the first time from Panama. Some variations are worth to mention: mesosoma light castaneous; median clypeal tooth angulate; median clypeal carina complete; vertex crest straight; anterior margin of pronotal disc coarse; pit of tergal process with very short tuft of setae.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5F9B09FF67F4439D8AFC33.taxon	materials_examined	Material examined. Holotype, ♂, COSTA RICA, San Vito de Coto Brus: Las cruces, 1200 m, 17. viii – 12. ix. 1982, Malaise trap, B. Gill col. (CNCI). PANAMA, Chiriquí: 1 ♂, 5.6 km N Boquete, La Culebra trail, [08 ° 49 ' 23 " N, 82 ° 25 ' 18 " W], F [light] I [nterception] T [rap], J. Ashe & R. Brooks [col.] (CNCI); 1 ♂, [Santa Clara], Harman's Finca, [08 ° 51 ' 38 " N, 82 ° 44 ' 44 " W], 3. VII. 1981, B. Gill [col.] (PMAE); Veraguas Pr [ovincia]: 2 ♂, Santa Fé, Altos De Piedra, [08 ° 31 ' 5.5 " N, 81 ° 06 ' 30.4 " W], 13 – 17 nov. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); Colón: 3 ♂, 14 km N J [un] ct [ion] Escobal r [oa] d and Piña r [oa] d on Piña r [oa] d, ± 20 m, [09 ° 20 ' N', 79 ° 54 ' W], 2 – 3. VI. 1995, F [light] I [nterception] T [rap], J. Ashe & R. Brooks [col.] (CNCI); Coclé Pr [ovincia]: 2 ♂, El Copé, Par [que] Nac [ional General de División] Omar Torrijos H [errera], 570 – 670 m. s. n. m., [08 º 41 ' 56 " N, 80 º 39 ' 17 " W], 26 – 30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); [Kuna Yala]: 1 ♂, Nusagandi Sta [tion] n [ea] r Columbia INA Igar Trail, [08 ° 58 ' N, 79 ° 32 ' W], 12 – 13. V. 1994, A. Windson [col.] (CNCI); Panamá Pr [ovincia]: 1 ♂, Parq [ue] Nac [ional] Soberanía, Camino, Plantación, [09 º 09 ' N, 79 º 43 ' W], 23 – 29 feb. 2000, P. González, A. Santos [col.] (MIUP); Comarca Kuna Yala: 6 ♂, Ustupu, Rio Abudi, [09 º 07 ' 57 " N, 77 º 55 ' 36 " W], 6 – 12 dic. 1999, P. González [col.] (MIUP); Darién Pr [ovincia]: 57 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (51 ♂), 1000 m, 21 mar – 4 ab 2000 (4 ♂) and 900 m (2 ♂) Cambra, Santos, Bermudez [col.] (MIUP); 3 ♂, Cana, 1380 m, [07 ° 51 ' N, 77 ° 37 ' W], 4 – 7. VI. 1986 (2 ♂), (CNCI) and 550 m, 7 º 45 ' N 77 ° 41 ' W, 3 – 7. vi. 96, (1 ♂), (UFES) F [light] I [nterception] T [rap], [J.] Ashe & [R.] Brooks [col.].	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5F9B09FF67F4439D8AFC33.taxon	distribution	Distribution (Fig. 231). Costa Rica and Panama.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5E9B09FF67F6009C88FAF0.taxon	diagnosis	Diagnosis. Male. Metasoma with lateral tergal process, with pair of elliptical, inclined, elongated depression with linear and oblique tufts of setae.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5E9B09FF67F6009C88FAF0.taxon	discussion	Remarks. It has five species: D. ellipticus Evans, D. rufipalpis Kieffer, D. inclinatus Redighieri & Azevedo, D. infissus Evans and D. guarani Colombo & Azevedo.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5E9B09FF67F6009C88FAF0.taxon	distribution	Distribution. Ranging from the southern Florida to northern Argentina.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5E9B0AFF67F7BD9EC8FE9F.taxon	diagnosis	Diagnosis. Mandible tridentate. Clypeus trapezoidal. Tergal process lateral with elliptical and inclined depressions and linear and oblique tuft of setae. Hypopygium with posterior margin straight. Genitalia: paramere short and wide; apex concave and very produced dorsally; aedeagal ventral very short; base wide; apical half narrow, slightly convergent with apex very sharpened; inner margin sinuous; outer margin expanded laterally; aedeagal dorsal body very wide, with apex laminar in lateral view; inner lamina sinuous, setose inside; outer lamina with series of grooves; base of aedeagal dorsal body slightly wider. Apodeme not extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5E9B0AFF67F7BD9EC8FE9F.taxon	discussion	Remarks. This species is known to Brazil (Paraíba, Pernambuco, Espírito Santo, São Paulo, Paraná and Santa Catarina) and Argentina (Colombo & Azevedo 2016). Now it is recorded for the first time from Panama. Some variations are worth to mention: head dark castaneous; frons strongly coriaceous; median clypeal tooth angulate; anterior margin of pronotal disc coarse; depression of tergal process very large; apex of paramere strongly curved inward.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5E9B0AFF67F7BD9EC8FE9F.taxon	materials_examined	Material examined. Holotype ♂, ARGENTINA, Oran: Abra Grande, Salta, 16 – 23 Feb. 1967, R. Golbach (FIML); PANAMA, Colón: 1 ♂ 14 km N J [un] ct [ion] Escobar r [oad] on Piña r [oa] d, ± 20 m, 09 ° 20 ' N', 79 ° 54 ' W], 2 – 3. VI. 1995, F [light] I [nterception] T [rap], J. Ashe & R. Brooks [col.] (CNCI); Darién Pr [ovincia]: 1 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5E9B0AFF67F7BD9EC8FE9F.taxon	distribution	Distribution (Fig. 237). Panama, Brazil (Paraíba, Pernambuco, Espírito Santo, São Paulo, Paraná and Santa Catarina) and Argentina.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5C9B0BFF67F2A89EFAFB55.taxon	diagnosis	Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with elliptical and and inclined depressions and linear and oblique tuft of small setae. Hypopygium with posterior margin nearly straight. Genitalia: paramere with dorsal margin developed with base projected downward and small tooth; aedeagal ventral ramus as long as aedeagal dorsal body, laminar; surfaces vertical; basal half wide, narrowing at sinuous apex; aedeagal dorsal body with two pairs of apical lobes; outer pair laminar; surface vertical and setose; inner pair slightly inclined upward with setose surface. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5C9B0BFF67F2A89EFAFB55.taxon	discussion	Remarks. This species is known to Mexico (Vera Cruz, Chiapas), Belize, Guatemala, Honduras, Costa Rica, Panama, Colombia and Ecuador (Azevedo 1999 b) and here more specimens are added. In this series the specimens are quite similar to those previously studied, without any variation that worths to mention.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5C9B0BFF67F2A89EFAFB55.taxon	materials_examined	Material examined. Holotype ♂, BELIZE, # 4 (CASC, # 17055); PANAMA, Chiriqui: 3 ♂, Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 25 – 6. VI. 2000, F [light] I [nterception] T [rap], H. & A. Howden [col.] (CNCI); 1 ♂, 15 km NW Hato del Volcano, 1200 m, [08 ° 46 ' N, 82 ° 38 ' W], 24 – 31. V. 1997, Peck & Howden [col.] (CNCI); Veraguas Pr [ovincia]: 1 ♂, P [arque] N [acional] Coiba, Isla Coita {= Coiba}, [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 27 nov – 4 dic 1998, Malaise, R. Cambra, S. Ber [mudez col.] (MIUP); 1 ♂, Santa Fé D [is] tr [ict], [08 ° 31 ' N, 81 ° 06 ' W], 24 – 26 marzo 1999, B. Amarillas, col: A. Santos, L. D. (MIUP); Comarca Kuna Yala: 1 ♂, Ustupu, Rio Abudi, [09 º 07 ' 57 " N, 77 º 55 ' 36 " W], 6 – 12 dic. 1999, Malaise, P. González [col.] (MIUP); Darién Pr [ovincia]: 12 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 450 m, [08 ° 00 ' N, 77 ° 45 ' W], 24 – 26 marzo 2001, Malaise, col. A. Santos (2 ♂), 230 m, 21 mar – 4 ab 2000, Cambra, Santos, Bermudez [col.] (1 ♂), 80 m, 16 nov 00 – 17 ene 2001 R. Cambra, A. Santos [col.] (9 ♂) (MIUP);	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5C9B0BFF67F2A89EFAFB55.taxon	distribution	Distribution (Fig. 244). Mexico (Vera Cruz, Chiapas), Belize, Guatemala, Honduras, Costa Rica, Panama, Colombia and Ecuador.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5C9B14FF67F4B39B2EFC7B.taxon	description	Description. Male. Body length 1.55 mm. Color: body castaneous. Head (Fig. 19). Mandible with two distal teeth. Median clypeal lobe subtrapezoidal, with one angulate tooth; median clypeal carina low in profile; incomplete apically; straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 50). Tergal process with lateral, very shallow, subcircular and small depression, with small, high and laterad tubercle placed on inner area of depression, pit small, with short and sparse tuft of setae. Posterior hypopygeal margin weakly concave (Fig. 81). Genitalia (Figs 178 – 179): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus higher than dorsal body, evenly narrow; cross section laminar; surface subhorizontal; inner margin sinuous; outer margin slightly sinuous; apex short, simple, translucent, slightly curved inward; additional inner ramus absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, horizontal and wide, with apical margin rounded and laterad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring. Variations. Metasoma light castaneous, tubercle low and with dense tuft of setae, ventral projection of apical lobe of aedeagal dorsal body longer, narrower or even absent.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5C9B14FF67F4B39B2EFC7B.taxon	discussion	Remarks. This species is easily recognized by having the apical lobes of the aedeagal dorsal body with a small laterad expansion bear-ear-shaped. This species is similar to D. sartus sp. nov. by having the paramere very excavated apically and the aedeagal dorsal body with three narrow, long filaments which connect the apical lobes of the aedeagal dorsal body to the basal portion of the aedeagal dorsal body. However, this species has the apical lobes of the aedeagal dorsal body with a small laterad projection, the aedeagal ventral ramus with the apex translucent and smooth, whereas D. sartus sp. nov. has the apical lobes of the aedeagal dorsal body with the apex abruptly acute and the aedeagal ventral ramus with the apex not translucent and toothed.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5C9B14FF67F4B39B2EFC7B.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Panama Pr [ovincia]: P [arque] Nac [ional] Sob [eranía], Camino, Plantación, [09 º 09 ' N, 79 º 43 ' W], 24 ago. – 2 sep. 1999, Malaise, A. Santos, P. González [col.] (MIUP). Paratypes: Bocas del Toro: 2 ♂, Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP); Chiriqui: 1 ♂, Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 2 – 6. VI. 2000, F [light] I [nterception] T [rap], H. & A. Howden [col.] (CNCI); Coche {= Coclé}: 1 ♂, El Cope {= El Copé}, [08 ° 37 ' 11 " N, 80 ° 35 ' 03 " W], 19. XI. 1994, Windson & Edwards [col.] (CNCI). 2 ♂, same data as holotype; 2 ♂, same province of holotype, P [arque] N [atural] Metropolitano, [08 º 59 ' 33 " N, 79 º 32 ' 41 " W], 24 ago. – 3 sep. 1999 (MIUP);	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C5C9B14FF67F4B39B2EFC7B.taxon	distribution	Distribution (Fig. 227). Panama (Bocas del Toro, Chiriqui, Coclé and Panama).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C439B15FF67F1D09C24FF6F.taxon	description	Description. Male. Body length 1.73 mm. Color: head castaneous; mesosoma and metasoma light castaneous. Head (Fig. 20). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one rounded very short tooth; median clypeal carina high in profile, incomplete apically, convex in profile. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 51). Tergal process with lateral, shallow, circular and large depression, with small, low and dorsad tubercle placed on center of depression, pit small, with short tuft of setae. Posterior hypopygeal margin strongly concave (Fig. 82). Genitalia (Figs 180 – 181): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; ramous bifid, surface horizontal; inner margin sinuous; outer margin sinuous; apex short, simple, parallel; additional inner ramus absent; basal stub short and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe very short, horizontal and narrow, with apical margin abruptly acute and laterad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C439B15FF67F1D09C24FF6F.taxon	discussion	Remarks. This species is the most different in setosus species-group by having the aedeagal dorsal body elongated and very short apical lobes of the aedeagal dorsal body. These characters are easy to be recognized. This species is similar to D. filus, by having the aedeagal dorsal body elongated with small apical lobes of the aedeagal dorsal body. However, this species has the paramere the same size as the basiparamere, the apical lobes of the aedeagal dorsal body very short and wide and the aedeagal ventral ramus bifid whereas D. filus has the paramere longer than basiparamere, the apical lobes of the aedeagal dorsal body longer and narrow and the aedeagal ventral ramus not bifid. The base of the aedeagal ventral ramus has small lamelar projections that are difficult to see in magnifying glass, but easily seem in microscope. In this species-group just this species and D. w-aedeagus has the ventral ramus bifid.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C439B15FF67F1D09C24FF6F.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Veraguas Pr [ovincia]: P [arque] N [acional] Coiba, Isla Coiba, Est [ación] Biologica Coiba, [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 5 – 6 ag 1998, Malaise, R. Cambra, A. Santos [col.] (MIUP). Paratypes: 3 ♂, same data as holotype; Darién: 1 ♂, Est [acion] I [nstituto] N [acional de] Re [cusos] Na [turales] Re [novables], P [arque] Nac [ional] Darién, [07 ° 51 ' 26.3 " N, 77 ° 28 ' 12.7 " W], 5 – 28 feb 1993, Malaise, R. Cambra [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C439B15FF67F1D09C24FF6F.taxon	distribution	Distribution (Fig. 226). Panama (Veraguas and Darién).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C429B15FF67F33B9CF0FAF0.taxon	description	Description. Male. Body length 1.55 mm. Color: body castaneous. Head (Fig. 21). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina very high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures large. Vertex crest straight. Pronotal disc weakly coriaceous; anterior margin carinate. Metasoma (Fig. 52). Tergal process with lateral, very shallow, subcircular and small depression, with very small, low and posterad tubercle placed on center of depression, pit very small, with short and dense tuft of setae. Posterior hypopygeal margin strongly concave (Fig. 83). Genitalia (Figs 182 – 183): paramere with dorsal margin entirely wide; apical margin rounded. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin sinuous; apex long with sinuous, slender ventral projection, curved outward; additional inner ramus absent; basal stub long and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, subhorizontal and narrow, with apical margin rounded and dorsad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring. Variations. Head, mesosoma and metasoma light castaneous; anterior margin of pronotal disc coarse; tergal process without depression.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C429B15FF67F33B9CF0FAF0.taxon	discussion	Remarks. This species is easily recognized by having the aedeagal ventral ramus with slender ventral projection and the carina of the median clypeal tooth very high in profile and the ventral margin of the paramere thick with thick setae. This species is similar to D. magnus by having the dorsal margin of the paramere almost straight with distinct thick setae apical and longer than the basiparamere. However, this species has the aedeagal ventral ramus abruptly narrow apicad with one ventral slender projection and the apical lobes of the aedeagal dorsal body subhorizontal whereas D. magnus has the aedeagal ventral ramus divided in two tubular, sinuous, projections and the apical lobes of the aedeagal dorsal body horizontal.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C429B15FF67F33B9CF0FAF0.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, [Chiriquí]: Fortuna, Div. Cont. Chiriqui-B [ocas del] Toro, 1070 – 1090 m, [08 ° 46 ' 16 " N, 82 ° 11 ' 27 " W], 8 – 11. abr. 1999, Malaise, R. Cambra, A. Santos [col.] (MIUP). Paratypes: 5 ♂, same data as holotype; 2 ♂, same of holotype except 9 – 11. abr. 1999, Tr [ampas] Amar [illas] (MIUP); Bocas del Toro: 1 ♂, Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 - 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C429B15FF67F33B9CF0FAF0.taxon	distribution	Distribution (Fig. 232). Panama (Bocas del Toro and Chiriquí).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C429B17FF67F7489B5FFE2A.taxon	description	Description. Male. Body length 1.85 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 22). Mandible with two distal teeth. Median clypeal lobe subtrapezoidal, with one angulate tooth; median clypeal carina low in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc coriaceous; anterior margin ecarinate. Metasoma (Fig. 53). Tergal process with lateral, shallow, subcircular and small depression, with small, high and dorsad tubercle placed on center of depression, pit small, with short and dense tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 84). Genitalia (Figs 184 – 185): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus as long as dorsal body, wide, progressively narrowing apicad; cross section laminar; surface subhorizontal; inner margin straight; outer margin slightly sinuous; apex long, simple, slightly curved inward; additional inner ramus absent; basal stub short and very narrow. Aedeagal dorsal body with pair of apical lobe; outer lobe short, horizontal and very narrow, with apical margin abruptly acute and ventrad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring. Variations. Depression of tergal process very shallow, tubercle low; apical lobe of aedeagal dorsal body longer and narrower.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C429B17FF67F7489B5FFE2A.taxon	discussion	Remarks. This species is similar to D. areius sp. nov. by having the paramere excavated apically and expanded basally and the aedeagal dorsal body with three narrow and long filaments which connect the apical lobes of the aedeagal dorsal body to the basal portion of the aedeagal dorsal body. However, this species has the apex of apical lobes of the aedeagal dorsal body abruptly acute and the aedeagal ventral ramus with the apex toothed, whereas D. areius sp. nov. has the apex of the apical lobes of the aedeagal dorsal body with a small laterad projection and the aedeagal ventral ramus with the apex translucent and smooth.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C429B17FF67F7489B5FFE2A.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Darién Pr [rovincia]: P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP). Paratypes: Bocas del Toro: 3 ♂, Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP); Coclé Pr [ovincia]: 1 ♂, El Copé, Par [que] Nac [ional General de División] Omar Torrijos H [errera], 570 – 670 m. s. n. m., [08 º 41 ' 56 " N, 80 º 39 ' 17 " W], 26 – 30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); Panama Pr [ovincia]: 1 ♂, P [arque] Nac [ional] Altos de Campana, [08 º 43 ' 15 " W, 80 º 01 ' 43 " W], 10 – 14 ago 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); 3 ♂ same data as holotype; 1 ♂, same of holotype except 1000 m, 21 mar – 4 ab 2000 (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C429B17FF67F7489B5FFE2A.taxon	distribution	Distribution (Fig. 245). Panama (Bocas del Toro, Coclé, Panama and Darién).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C409B10FF67F0029E41FEC2.taxon	description	Description. Male. Body length 2.63 mm. Color: body castaneous. Head (Fig. 23). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina low in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc weakly coriaceous; anterior margin coarse. Metasoma (Fig. 54). Tergal process with lateral, deep, circular and large depression, with small, low and dorsad tubercle placed on center of depression, pit large, with short and dense tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 85). Genitalia (Figs 186 – 187): paramere with dorsal margin entirely wide; apical margin rounded. Aedeagal ventral ramus as long as dorsal body, evenly narrow; cross section laminar; surface horizontal; inner margin sinuous; outer margin slightly sinuous; apex short, simple, curved outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, subhorizontal and narrow, with apical margin abruptly acute and posterad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring. Variation. Tergal process with pit of tubercle with few setae.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C409B10FF67F0029E41FEC2.taxon	discussion	Remarks. This species is easily recognized by having the paramere wide basally and the triangular shape of the outer lobe of the aedeagal dorsal body which is very projected forward. This species is not similar to anyone of the setosus species-group.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C409B10FF67F0029E41FEC2.taxon	materials_examined	Material examined. Holotype, ♂, Darién Pr [ovincia]: P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP). Paratypes: Bocas del Toro, Wekso: 1 ♂, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampas] Amarillas, A. Santos [col.] (MIUP); Chiriqui: 1 ♂, Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 25 – 6. VI. 2000, F [light] I [nterception] T [rap], H. & A. Howden [col.] (CNCI); 1 ♂, 15 km NW Hato del Volcano, 1200 m, [08 ° 46 ' N, 82 ° 38 ' W], 24 – 31. V. 1997, Peck & Howden [col.] (CNCI); Veraguas Pr [ovincia]: 14 ♂, P [arque] N [acional] Coiba, Isla Coiba, Cerro la Equis 0 m, (3 ♂), 90 m (3 ♂), [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 8 – 12 mar [zo] 1998, Malaise, R. Cambra, A. Santos [col.], 120 m, 29 jul – 4 ag 1998 (5 ♂), 100 m, 30 – nov – 4 – dic. 1998, R. Cambra, S. Bermudez [col.] (2 ♂) and Playa Blanca-Cativales, 31 jul – 6 ago 1998, R. Cambra, A. Santos [col.] (1 ♂) (MIUP); Colón Pr [ovincia]: 1 ♂ P [a] r [que] Nac [ional] Portobelo, San Antonio, 16 – 29 feb. 2000, Tr [ampa] Malaise, P. González, A. Santos [col.] (MIUP); 1 ♂, 14 km N J [un] ct [ion] Escobal r [oa] d and Piña r [oa] d on Piña r [oa] d, ± 20 m, [09 ° 20 ' N', 79 ° 54 ' W], 2 – 3. VI. 1995, F [light] I [nterception] T [rap], J. Ashe & R. Brooks [col.] (CNCI); Comarca Kuna Yala: 6 ♂, Ustupu, Rio Abudi, [09 º 07 ' 57 " N, 77 º 55 ' 36 " W], 6 – 12 dic. 1999, Malaise, P. González [col.] (MIUP); Panama Pr [ovincia]: 11 ♂, P [arque] N [atural] Metropolitano, [08 º 59 ' 33 " N, 79 º 32 ' 41 " W], 24 ago. – 3 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); 1 ♂, P [arque] Nac [ional] Sob [eranía], Camino, Plantación, [09 º 09 ' N, 79 º 43 ' W], 24 ago. – 2 sep. 1999 Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); 2 ♂, Chilibre, Parque Nacional Chagres, Campo Chagres, [09 º 25 ' 54 " N, 79 º 30 ' 54 " W], 24 – 28. enero. 2000 Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); 3 ♂, C [anal] Z [one], B [arro] Colorado Is [land], [09 ° 09 ' N, 79 ° 51 ' W], 15 – 31. VII. 1994, (2 ♂) and 1 – 14. VII. 1994 (1 ♂), F [light] I [nterception] T [rap], D. Banks [col.] (CNCI); 27 ♂, same data as holotype; 9 ♂, same province of holotype Est [ación] I [nstituto] N [acional de] Re [cusos] Na [turales] Re [novables] [07 ° 51 ' 26.3 " N, 77 ° 28 ' 12.7 " W], 5 – 28 feb 1993, Malaise, R. Cambra [col.] (MIUP); 6 ♂, Cana, Serrania de Pirre, 1250 m, 4 – 7. vi. 1996, J. Ashe, R. Brooks. (UFES); 5 ♂, Estac [ión] Cruce de Mono, [07 ° 51 ' 26.3 " N, 77 ° 28 ' 12.7 " W], 6 feb – 4 mar 1993, Tr [ampa] malaise, R. Cambra, J. Coronado [col.] (MIUP); 2 ♂, Cruce de Mono: Estac. I [nstituto] N [acional de] Re [cusos] Na [turales] Re [novables], 6. II – 4. III. 1993 (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C409B10FF67F0029E41FEC2.taxon	distribution	Distribution (Fig. 246). Panama (Bocas del Toro, Chiriqui, Veraguas, Colón, Panama, Comarca Kuna Yala and Darién).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C479B10FF67F3559D9AFB71.taxon	diagnosis	Diagnosis. Mandible tetradentate. Clypeus tridentate. Tergal process with pair of very shallow depressions, with minute tubercle with setae on top, outer margin of depression with group of thick setae. Hypopygium with posterior margin concave. Genitalia: paramere wide basally, narrowing to the rounded apex in lateral view, with strong setae above; aedeagal ventral ramus higher than aedeagal dorsal body, with three long and conspicuous apical horns; aedeagal dorsal body wide, with two pairs of apical lobes; outer pair stout and membranous; inner pair laminar, vertical with apex bidentate and directed downward.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C479B10FF67F3559D9AFB71.taxon	discussion	Remarks. This species is known to Saint Vicent (Azevedo 2003). Now it is recorded for the first time from Panama. Some variations are worth to mention: head, mesosoma and metasoma light castaneous; median clypeal carina inconspicuous; median clypeal tooth angulate; anterior margin of pronotal disc slightly coarse; tergal process with tubercle, thick setae on pit and in outer margin of depression; aedeagal ventral ramus shorter than aedeagal dorsal body.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C479B10FF67F3559D9AFB71.taxon	materials_examined	Material examined. Lectotype male: S [AIN] T VICENT, W [est] I [ndies]: 1000 feet, H. H. Smith col (USNM, # 2497). PANAMA, Veraguas: 10 ♂, P [arque] N [acional] Coiba, Isla Coibita, [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 6 – 11 mar 1998 (1 ♂), 29 jul. 4 ag 1998, (3 ♂), Cerro la Equis, 120 m (1 ♂), Playa Blanca-Cativales 31 jul – 6 ago 1998 (3 ♂), Est [ación] Biologica Coiba, 5 – 6 ag 1998 (2 ♂), Malaise, R. Cambra, A. Santos [col.] (MIUP); Darién: 1 ♂, Est [ación] I [nstituto] N [acional de] Re [cusos] Na [turales] Re [novables], P [arque] Nac [ional] Darién, [07 ° 51 ' 26.3 " N, 77 ° 28 ' 12.7 " W], 5 – 28 feb 1993, Malaise, R. Cambra [col.] (MIUP); 1 ♂, Cana 7 º 45 ' N, 77 º 41 W, 3 – 7. vi. 96, 550 m, J. Ashe, R. Brooks. (UFES).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C479B10FF67F3559D9AFB71.taxon	distribution	Distribution (Fig. 240). Panama and Saint Vicent.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C479B10FF67F6C39D80FA1C.taxon	diagnosis	Diagnosis. Tergal process lateral with pair of sublateral depressions, tubercle and tuft slightly directed toward each other.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C479B10FF67F6C39D80FA1C.taxon	discussion	Remarks. It has two species: D. strabus Azevedo and D. thysanus Azevedo. Distribution. Dominican Republic and Panama.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C479B10FF67F42D9D25F82A.taxon	diagnosis	Diagnosis. Mandible bidentate. Clypeus tridentate. Tergal process with pair of rounded and shallow depression with pit, high him and tuft of setae slightly directed each other. Hypopygium with posterior margin straight. Genitalia: paramere with dorsal margin slightly convex; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar; surface horizontal; apex with two rami; aedeagal dorsal body with two pairs of apical lobes, the outer pair narrow with apex sharpened strongly angled downward; inner margin with large and triangular basal expansion; inner pair membranous, stout and setose. Apodeme extending beyond the genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C479B10FF67F42D9D25F82A.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Fortuna: Cont. Chiriqui, B. Toro, 1070 – 1090 m, 8 – 11. IV. 1999, Malaise trap, R. Cambra & A. Santos col. (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C479B10FF67F42D9D25F82A.taxon	discussion	Remarks. No additional material of this species was not found in this study.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C479B10FF67F42D9D25F82A.taxon	distribution	Distribution (Fig. 243). Panama.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C469B11FF67F3E59F91F9C0.taxon	diagnosis	Diagnosis. Mandible tetradentate. Clypeus trapezoidal. Tergal process lateral with shallow depressions, pair of turbecles with pit of setae on top. Hypopygium with posterior margin convex with small conical expansion. Genitalia: paramere with acute apex and arched inward, dorsal margin developed; aedeagal ventral ramus short than aedeagal dorsal body, as high as base of cuspis; wide; laminar; inner and outer margins straight and slightly convergent; apex acute with outer margin slightly concave with ventral surface slightly directed outward; aedeagal dorsal body with pair of apical lobes, laminar, wide with rounded apex in lateral view, compressed in dorsal view, below the apex. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C469B11FF67F3E59F91F9C0.taxon	discussion	Remarks. This is one of the most widespread species through the Neotropical region (Azevedo 1999 b) and here more specimens are added. Some variations are worth to mention: median clypeal tooth angulate; median carina clypeal straight and complete; vertex crest straight; tergal process with very small and very shallow depression; pit of tubercle with few setae.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C469B11FF67F3E59F91F9C0.taxon	materials_examined	Material examined. Holotype ♂, BRAZIL, Santa Catarina: Nova Teutonia, 26 February 1945, Fritz Plaumann (BMNH); PANAMA, Chiriqui: 17 ♂, Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 2 – 6. VI. 2000 (6 ♂), 25 – 6. VI. 2000 (11 ♂), F [light] I [nterception] T [rap], H. & A. Howden [col.] (CNCI); 2 ♂ Res [erva Florestal] La Fortuna, Viveno, [08 ° 44 ’ 39 ” N, 82 ° 14 ’ 15 ” W], 14 – 18. VI. 1994, A. R. Gillogly [col.] (CNCI); 1 ♂ 15 km NW Hato del Volcano, 1200 m [08 ° 46 ' N, 82 ° 38 ' W], 24 – 31. V. 1997, Peck & Howden [col.] (CNCI); Veraguas Pr [ovincia]: 6 ♂, P [arque] N [acional] Coiba, Isla Coibita, [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 27 – nov – 4 – dic. 1998, (2 ♂), Cerro La Equis, 120 m, 30 – nov. – 4 dic. 1998 (2 ♂), 100 m (1 ♂), 90 m (1 ♂), Malaise, R. Cambra, S. Bermudez [col.] (MIUP); Colón: 2 ♂, 14 km N J [un] ct [ion] Escobal r [oa] d and Piña r [oa] d on Piña r [oa] d, ± 20 m, [09 ° 20 ' N', 79 ° 54 ' W], 2 – 3. VI. 1995, F [light] I [nterception] T [rap], J. Ashe & R. Brooks [col.] (CNCI); Old Gamboa: 1 ♂, C [anal] Z [one], [09 ° 25 ' N, 80 ° 00 ' W], 14. XI. 1994, F [light] I [nterception] T [rap], P. Windson [col.] (CNCI); Darién Pr [ovincia]: 30 ♂ P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 87 ° 45 ' W], 16 nov 00 – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (29 ♂), 21 mar – 4 ab 2000, Tr [ampa] Amarilla, Cambra, Santos, Bermudez [col.] (1 ♂) (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C469B11FF67F3E59F91F9C0.taxon	distribution	Distribution (Fig. 240). Costa Rica, Panama, Ecuador, Peru and Brazil (Pará, Espírito Santo, São Paulo and Paraná).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C469B11FF67F2A89E41FE58.taxon	diagnosis	Diagnosis. Male. Clypeus trapezoidal. Metasoma with lateral tergal process, with pair of tubercles slightly directed each other, placed in pair of shallow depression or in horizontal surface.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C469B11FF67F2A89E41FE58.taxon	discussion	Remarks. It has two species: D. plaumanni Evans and D. tuberculatus Ashmead. Distribution. Saint Vincent, Costa Rica, Panama, Ecuador, Peru and Brazil (Pará, Espírito Santo, São Paulo and Paraná).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C469B11FF67F44D9E6BF875.taxon	diagnosis	Diagnosis. Male. Tergal process median with depression, pair of tufts of setae. Basal portion of dorsal margin of paramere much developed. Digitus wide medially. Cuspis elongated. Aedeagal ventral ramus with inner margin sinuous basally, wide, narrowing apically.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C469B11FF67F44D9E6BF875.taxon	discussion	Remarks. It has six species: D. ulceratus Evans, D. dentiformis Azevedo, D. concavatus Azevedo, D. rectilineus Azevedo, D. congo Colombo & Azevedo and D. terbius sp. nov..	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C469B11FF67F44D9E6BF875.taxon	distribution	Distribution. Panama, Brazil (Distrito Federal, Espírito Santo, Rio de Janeiro, São Paulo and Paraná) and Argentina.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C459B12FF67F2A89C64FC34.taxon	description	Description. Male. Body length 2.39 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 24). Mandible with three distal teeth. Median clypeal lobe ill-defined, with one angulate very short tooth; median clypeal carina low in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc coriaceous; anterior margin carinate. Metasoma (Fig. 55). Tergal process with median, shallow, subcircular and large depression, without tubercle, with pit large, short and dense tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 86). Genitalia (Figs 188 – 189): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface vertical; inner margin straight; outer margin sinuous; apex long, simple, angled outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe very long, horizontal and wide, with apical margin angled and laterad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring. Variations. Head castaneous and mesosoma light castaneous; median clypeal tooth inconspicuous.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C459B12FF67F2A89C64FC34.taxon	discussion	Remarks. This species is similar to D. afestus sp. nov. by having the apical lobes of the aedeagal dorsal body very long and wide and the apical half of the aedeagal ventral ramus narrow. However, this species has the dorsal margin of the paramere rounded with the apex crenulate and the apex of the aedeagal ventral ramus angled outward, whereas D. afestus sp. nov. has the paramere with the dorsal margin no so rounded with the apex smooth and the apex of the aedeagal ventral ramus not angled outward.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C459B12FF67F2A89C64FC34.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Bocas del Toro: Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP). Paratypes: 8 ♂ same data as holotype (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C459B12FF67F2A89C64FC34.taxon	distribution	Distribution (Fig. 247). Panama (Bocas del Toro).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C459B12FF67F6019A49FAD5.taxon	diagnosis	Diagnosis. Male. Tergal process median, rounded depression nearly totally covered by pair of joined tufts of long setae directed backward.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C459B12FF67F6019A49FAD5.taxon	discussion	Remarks. It has two species: D. unitus and D. delgatus Azevedo.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C459B12FF67F6019A49FAD5.taxon	distribution	Distribution. Mexico (Chiapas, Vera Cruz), Belize, Guatemala, Costa Rica, Panama and Venezuela.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C459B13FF67F7629A2EFD5B.taxon	diagnosis	Diagnosis. Mandible bidentate. Clypeus trapezoidal. Tergal process with shallow and large depression with pair of joined tuft of long setae backward. Hypopygium with posterior margin with three small concavities. Genitalia: paramere with apex arched inward and truncate; aedeagal ventral ramus slightly longer than aedeagal dorsal body narrow, diverging dorsally; inner margin sinuous and parallel; outer margin with broad median concavity; aedeagal dorsal body with two pairs of apical lobes; outer lobe laminar with rounded apex; inner lobe stout densely setose. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C459B13FF67F7629A2EFD5B.taxon	discussion	Remarks. This species is known to Mexico (Chiapas, Vera Cruz), Belize, Guatemala, Costa Rica and Panama (Azevedo 1999 a) and here more specimens are added. Some variations are worth to mention: metasoma light castaneous; median clypeal tooth angulate; aedeagal ventral ramus as longer as aedeagal dorsal body; volsella with rounded expansion medially and some specimens without it.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C459B13FF67F7629A2EFD5B.taxon	materials_examined	Material examined. Holotype, ♂, COSTA RICA, San Coto de Vito Brus: Las Cruces, 1200 m, 2 – 9. ix. 1982, Malaise trap, B. Gill col. (CNCI). PANAMA, Chiriqui: 4 ♂, 15 km NW Hato del Volcano, 1200 m, [08 ° 46 ' N, 82 ° 38 ' W], 24 – 31. V. 1997, Peck & Howden [col.] (CNCI); 6 ♂, Res [erva Florestal] La Fortuna, Viveno, [08 ° 41 ' 27 " N, 82 ° 14 ' 12 " W], 14 – 18. VI. 1994, A. R. Gillogly [col.] (CNCI); 1 ♂, water shed, hydro trail, 1100 m, [08 ° 44 ' 21 " N, 82 ° 14 ' 36 " W], 24. I. 1994 – 27. I. 1994, D. M. Windson [col.] (CNCI); 1 ♂, Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 2 – 6. VI. 2000, F [light] I [nterception] T [rap], H. & A. Howden [col.] (CNCI); 1 ♂, Santa Clara, Finca Hartman, Ojo de Agua, [08 ° 51 ' 38 " N, 82 ° 44 ' 44 " W], 8 – 9. XI. 2001, M [alaise] T [rap], N. Schiff [col.] (UCDC); Veraguas: 1 ♂, Pr [ovincia] Santa Fé], Santa Fé, Altos De Piedra, [08 ° 31 ' 5 " N, 81 ° 06 ' 30 " W], 13 – 17 nov. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); Coclé Pr [ovincia]: 2 ♂, El Copé, Par [que] Nac [ional General de División] Omar Torrijos H [errera], 570 – 670 m. s. n. m., [08 º 41 ' 56 " N, 80 º 39 ' 17 " W], 26 – 30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); Panamá Pr [ovincia]: 1 ♂, P [arque] N [atural] Metropolitano, [08 º 59 ' 33 " N, 79 º 32 ' 41 " W], 24 ago. – 3 sep. 1999 (MIUP); Darién Pr [ovincia]: 33 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (26 ♂), 1000 m, 21 mar – 4 ab 2000, Cambra, Santos, Bermudez [col.] (5 ♂) and 580 m (1 ♂) (MIUP), 1450 m, 4 – 7. vi. 1995, F [light] I [ntrception] T [rap] J. Ashe, R. Brooks (1 ♂) (UFES); 1 ♂, Nusagandi Sta [tion] n [ea] r Columbia INA Igar Trail, [08 ° 58 ' N, 79 ° 32 ' W], 12 – 13. V. 1994, A. Windson [col.] (CNCI).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C459B13FF67F7629A2EFD5B.taxon	distribution	Distribution (Fig. 247). Mexico (Vera Cruz and Chiapas), Belize, Guatemala, Costa Rica and Panama.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C449B13FF67F60B9CB8F832.taxon	diagnosis	Diagnosis. Mandible tridentate. Clypeus with median tooth. Tergal process with elliptical, deep and large depression with pair of tubercles and tuft of setae directed each other. Hypopygium with posterior margin concave. Genitalia: paramere arched inward apically and rounded; aedeagal ventral ramus short than aedeagal dorsal body with dilation below the apex; inner margin straight; outer margin convex; aedeagal dorsal body with two pair of apical lobes; outer lobe large with apex folded inward; inner lobe circular and setose. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C449B13FF67F60B9CB8F832.taxon	discussion	Remarks. This species is known from Mexico (Vera Cruz), Costa Rica and Panama (Azevedo 1999). Here more specimens are added. In this series the specimens are quite similar to those previously studied, without any variation that is worth to mention.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C449B13FF67F60B9CB8F832.taxon	materials_examined	Material examined. Holotype, ♂ COSTA RICA, Puntarenas: Monteverde, 1400 m, 10 º 43 ’ N 84 º 02 W, 12 – 15. vi. 1974, Malaise trap, W. A. Harding & J. P Donahue col. (LACM); PANAMA, Bocas del Toro: 1 ♂, Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP); 1 ♂, Fortuna, Div. Cont. Chiriqui-B [ocas del] Toro, 1070 – 1090 m, [08 ° 46 ' 16.5 " N, 82 ° 11 ' 27.3 " W], 9 – 11. abr. 1999, Tr [ampas] Amar., R. Cambra, A. Santos [col.] (MIUP); Chiriqui: 5 ♂, Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 25 – 6. VI. 2000, F [light] I [nterception] T [rap], H. & A. Howden [col.] (CNCI); Colón: 1 ♂, 14 km N J [un] ct [ion] Escobal r [oa] d and Piña r [oa] d on Piña r [oa] d, ± 20 m, [09 ° 20 ' N, 79 ° 54 ' W], 2 – 3. VI. 1995, F [light] I [nterception] T [rap], J. Ashe & R. Brooks [col.] (CNCI); [Panama]: 1 ♂, C [anal] Z [one], B [arro] Colorado Is [land], [09 ° 09 ' N, 79 ° 51 ' W], 15 – 31. VII. 1994, F [light] I [nterception] T [rap], D. Banks [col.] (CNCI); Darién Pr [ovincia]: 20 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] and 450 m, 24 – 26 marzo 2001, col. A. Santos (2 ♂) (MIUP);	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C449B13FF67F60B9CB8F832.taxon	distribution	Distribution (Fig. 233). Mexico (Vera Cruz), Costa Rica and Panama.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C449B13FF67F0E89BDEFC34.taxon	diagnosis	Diagnosis. Male. Clypeus with only median tooth. Tergal process median, depression transverse and elliptical. Pair of lateral tubercles, directed each other.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C449B13FF67F0E89BDEFC34.taxon	discussion	Remarks. It has 11 species: D. bisserratus Azevedo, D. polidentatus Azevedo, D. planus Azevedo, D. extrarramis Azevedo, D. triangularis Azevedo, D. dilatatus, D. strictus Azevedo, D. bifurcatus Azevedo, D. digitatus Azevedo, D. vallensis and D. rosangelae Colombo & Azevedo.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C449B13FF67F0E89BDEFC34.taxon	distribution	Distribution. Ranging from Mexico to south of Brazil, including some Caribbean islands.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4B9B1CFF67F2A89D86FC63.taxon	diagnosis	Diagnosis. Mandible bidentate. Clypeus wih median lobe ill-defined. Tergal process median with large, elliptical and deep depression with pair of tubercles with tuft of setae directed each other. Hypopygium with posterior margin concave. Genitalia: paramere with apex truncate and oblique; aedeagal ventral ramus as long as aedeagal dorsal body; surface smooth; inner margin straight; outer margin convex; apical half narrow and divergent; aedeagal dorsal body with apical half arched downward; ventral side with median fold; dorsal margin of apical half arched outward and pair of very small lobes upward in base of aedeagal dorsal body. Apodeme not extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4B9B1CFF67F2A89D86FC63.taxon	discussion	Remarks. This species is known from Colombia (Azevedo 1999 a) and is recorded for the first time from Panama. Some variations are worth to mention: median clypeal tooth angulate; median clypeal carina conspicuous and complete; vertex crest straight; anterior margin of pronotal disc coarse; aedeagal ventral ramus shorter than aedeagal dorsal body; median region of aedeagal dorsal body wide, apical portion elongated and narrow, outer margin sinuous; genital ring bifurcated distally in ventral view.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4B9B1CFF67F2A89D86FC63.taxon	materials_examined	Material examined. Holotype ♂, COLOMBIA, Peñas Blancas: 10 km, W Cali, Dept. Valle, 26 – 28 1975, Malaise trap, R. Wilkerson (FSCA); PANAMA, Coche {= Coclé}: 1 ♂, El Cope {= El Copé}, 730 m, [08 ° 37 ' 11 " N, 80 ° 35 ' 03 " W], 19 – 20. XI. 1994, F [light] I [nterception] T [rap], D. Windson Edwards [col.] (CNCI); Darién: 5 ♂, Cana, Serraria {= Serrania} de Pirre 1450 m, 4 – 7. vi. 1995, J. Ashe, R. Brooks F [light] I [nterception] T [rap] (UFES).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4B9B1CFF67F2A89D86FC63.taxon	distribution	Distribution (Fig. 244). Colombia and Panama.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4B9B1DFF67F6699A09FCCA.taxon	diagnosis	Diagnosis. Mandible bidentate. Clypeus broadly projected forward, with small median tooth paralleled by small rounded expansion. Tergal process with pair of triangular and shallow lateral depressions, drop-shaped pit with tuft of very short setae. Genitalia: paramere unusually very wide, apex truncate; aedeagal ventral ramus short than aedeagal dorsal body, with arrow apical expansion; aedeagal dorsal body with two pairs of apical lobes; outer pair elongate dome-shaped lobe; inner pair membranous, stout and setose.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4B9B1DFF67F6699A09FCCA.taxon	discussion	Remarks. This species is known only from the holotype and allotype from Panama. In this study, it was the most abundant species with 273 specimens and it has the largest distribution. All the specimens when compared with description by Evans (1964) and redescription by Azevedo (2003) have no variations that worth to mention.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4B9B1DFF67F6699A09FCCA.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Barro Colorado Island: Canal Zone, 26. II. 1955, Berlese refuse, C. W Rettenmeyer col. (USNM # 64996). PANAMA, Bocas del Toro: 61 ♂, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP); Chiriqui: 33 #, Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 25 – 6. VI. 2000, F [light] I [nterception] T [rap], H. & A. Howden [col.] (CNCI); 2 – 6. VI. 2000 (CNCI) (9 ♂) and (UFES) (2 ♂); 1 ♂, La Fortuna Dam, 1200 m wet forest 14 – VI – 16 – VII – 1982, B. Gill. (CNCI); Veraguas: 2 ♂, Pr [ovincia, Santa Fé Distri], Santa Fé, Altos de Piedra, 770 m, [08 ° 31 ' 5.5 " N, 81 ° 06 ' 30.4 " W], 22 – 26 marzo 1999, Malaise, Col. A. Santos, L. D. (1 ♂) and 13 – 17 nov. 1999, A. Santos, P. González [col.] (1 ♂) (MIUP); Colón Pr [ovincia]: 3 ♂, P [a] r [que] Nac [ional] Portobelo, San Antonio, 16 – 29 feb. 2000, Tr [ampa] Malaise, P. González, A. Santos [col.] (MIUP); 3 ♂, S [an] ta Rita [Arriba], Macizo, Parador Sierra Llorona, [09 ° 19 ' 57 " N, 79 ° 45 ' 1 " W], 22 dic. 1997, D. Quintero [col.] (MIUP); 3 ♂, 14 km N J [un] ct [ion] Escobal r [oa] d and Piña r [oa] d on Piña r [oa] d, ± 20 m, [09 ° 20 ' N', 79 ° 54 ' W], 2 – 3. VI. 1995, F [light] I [nterception] T [rap], J. Ashe & R. Brooks [col.] (CNCI); Coclé Pr [ovincia]: 3 ♂, El Copé, Par [que] Nac [ional General de División] Omar Torrijos H [errera], 570 – 670 m. s. n. m. [08 º 41 ' 56 " N, 80 º 39 ' 17 " W], 26 – 30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); Panama Pr [ovincia]: 6 ♂, P [arque] Nac [ional] Sob [eranía], Camino, Plantación, [09 º 09 ' N, 79 º 43 ' W], 24 ago. – 2 sep. 1999, Malaise, P. González, A. Santos [col.] (MIUP); 2 ♂, P [arque] Nac [ional] Altos de Campana, [08 º 43 ' 15 " W, 80 º 01 ' 43 " W], 10 – 14 ago 1999 Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); 1 ♂, Chilibre, Parque Nacional Chagres, Campo Chagres, [09 º 25 ' 54 " N, 79 º 30 ' 54 " W], 24 – 28. enero. 2000 Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); 1 ♂, P [arque] N [atural] Metropolitano, [08 º 59 ' 33 " N, 79 º 32 ' 41 " W], 24 ago. – 3 sep. 1999 Tr [ampa] Malaise, A. Santos, P. González [col] (MIUP); 1 ♂, Cerro Campana, 08 ° 40 ' N, 79 ° 50 ' W, 7 – 14. V. 1973 (CNCI); 1 ♂, Barro Colorado I [sland], [09 ° 09 ' N, 79 ° 51 ' W], 23. III. 1981, R. B. & L. S. Kimsey [col.] (UCDC); 1 ♂, Old Gamboa, C [anal] Z [one], [09 ° 25 ' N, 80 ° 00 ' W], 14. XI. 1994, F [light] I [nterception] T [rap], P. Windson [col.] (CNCI); 5 ♂, C [anal] Z [one], B [arro] Colorado Is [land], [09 ° 09 ' N, 79 ° 51 ' W], 15 – 31. VII. 1994 (1 ♂), 14. VII. 1994 (3 ♂) D. Banks [col.] and 3 – 13 – VI – 1983, B Gill (1 ♂) (CNCI); Darién Pr [ovincia]: 134 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov 00 – 17 ene 2001, Malaise (131 ♂), 18 – 24 ene 2001 Tr [ampa] Amarilla (1 ♂) R. Cambra, A. Santos [col.], 450 m, 24 – 26 marzo 2001, Malaise, col. A. Santos (1 ♂) and 560 m, 21 mar – 4 ab 2000, Cambra, Santos, Bermudez [col.] (1 ♂) (MIUP); 5 ♂, Cana, 1380 m, [07 ° 51 ' N, 77 ° 37 ' W], 4 – 7. VI. 1986, F [light] I [nterception] T [rap], [J.] Ashe & [R.] Brooks [col.] (CNCI); 7 ♂, Serraria {Serrania} de Pirre, 1450 m, 4 – 7. vi. 1995, J. Ashe, R. Brooks F [light] I [nterception] T [rap] (6 ♂) (UFES) and 1456 m, 7 – 9. VI. 1996 (1 ♂) (CNCI).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4B9B1DFF67F6699A09FCCA.taxon	distribution	Distribution (Fig. 249). Panama (Bocas del Toro, Chiriquí, Veraguas, Colón, Coclé, Panamá and Darién).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4A9B1DFF67F1589CA3F8FD.taxon	diagnosis	Diagnosis. Mandible tetradentate. Clypeus subtrapezoidal. Tergal process with lateral, shallow and subcircular depressions, with small tubercles and tuft of setae. Hypopygium with posterior margin concave. Genitalia: paramere acute with dorsal and ventral margins straight; aedeagal ventral ramus longer than aedeagal dorsal body, laminar; surface horizontal; basal half wide and fused; apical half with two laminas narrowing at apex; aedeagal dorsal body with two pairs of apical lobes; outer pair laminar; surface vertical; apex acute with horizontal projection masked-shaped; inner pair membranous, setose and longer than outer lobe.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4A9B1DFF67F1589CA3F8FD.taxon	discussion	Remarks. This species is known to Brazil (São Paulo and Paraná) (Redighieri & Azevedo 2006). Now it is recorded for the first time from Panama. Some variations in this series worth to mention: head, mesosoma and metasoma castaneous; aedeagal ventral ramus shorter than aedeagal dorsal body; apical lobes of aedeagal dorsal body with outer margin angulate and inner margin very excavated in some specimens; outer margin of apical lobes of aedeagal dorsal body subquadrate and inner margin almost straight in some specimens.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4A9B1DFF67F1589CA3F8FD.taxon	materials_examined	Material examined. Holotye, ♂, BRAZIL, Paraná: Morretes, Parque Estadual do Pau Oco, 25 º 34 ’ 27.5 ” S 48 º 53 ’ 33 ” W, 10 – 13. iv. 2002, armadilha Malaise, M. T. Tavares e eq. col. (MZSP); PANAMA, Bocas del Toro: 1 ♂, Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP); [Chiriquí]: 1 ♂, Fortuna, Div. Cont. Chiriqui-B [ocas del] Toro, 1070 – 1090 m, [08 ° 46 ' 16.5 " N, 82 ° 11 ' 27.3 " W], 8 – 11. abr. 1999, Malaise, R. Cambra, A. Santos [col.] (MIUP); Coclé Pr [ovincia]: 3 ♂, El Copé, Par [que] Nac [ional General de División] Omar Torrijos H [errera], 570 – 670 m. s. n. m., [08 º 41 ' 56 " N, 80 º 39 ' 17 " W], 26 – 30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); Darién Pr [ovincia]: 4 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 900 m, [08 ° 00 ' N, 77 ° 45 ' W], 21 mar – 4 ab 2000, Malaise, Cambra, Santos, Bermudez [col.] (1 ♂), 80 m, 16 nov – 17 ene 2001, R. Cambra, A. Santos [col.] (3 ♂) (MIUP); 1 ♂, Cana, Serrania de Pirre, 1250 m, 4 – 7. vi. 1996, J. Ashe, R. Brooks (UFES).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4A9B1DFF67F1589CA3F8FD.taxon	distribution	Distribution (Fig. 239). Panama and Brazil (São Paulo and Paraná).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C499B1FFF67F53E9DD0FC7A.taxon	description	Description. Male. Body length 4.54 mm. Color: body dark castaneous. Head (Fig 25). Mandible with three distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, incomplete apically, straight in profile or nearly so. Frons strongly coriaceous and punctures small. Vertex crest straight. Pronotal disc coriaceous; anterior margin coarse. Metasoma (Fig. 56). Tergal process with lateral, shallow, subcircular and large depression, without tubercle, with short and dense tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 87). Genitalia (Figs 207 – 208): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin sinuous; apex short, translucent, with two small projections, slightly curved outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and wide, with apical margin rounded and slightly mesad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring. Variations. Head, metasoma and mesosoma castaneous; median clypeal lobe subtrapezoidal; pit of tergal process with few setae.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C499B1FFF67F53E9DD0FC7A.taxon	discussion	Remarks. This species is easily recognized by having the genitalia robust, the aedeagal ventral ramus very wide with the apex translucent with two small projections. This species is similar to D. iozus sp. nov. by having the aedeagal dorsal body robust and the aedeagal ventral ramus very wide. However, this species has the paramere, basiparamere and the apex of apical lobes of the aedeagal dorsal body narrow and the apex of the aedeagal ventral ramus translucent with small projections whereas, D. iozus sp. nov. has the paramere and the basiparamere wide, the apex of the aedeagal dorsal body wide and rounded and the apex of the aedeagal ventral ramus simple and not translucent.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C499B1FFF67F53E9DD0FC7A.taxon	materials_examined	Material examined. Holotype ♂, PANAMA, Chiriquí: 08 ° 54,7 ' N 82 ° 47,3 ' W 17 – 18. vi. 1996, 1875 m, J. Ashe, R. Brooks F [light] I [interception] T [trap] (UFES). Paratypes: 3 ♂, same data as holotype; 1 ♂ 2 km W Cerro Punta, 1700 m, [08 ° 51 ' N, 82 ° 35 ' W], 19. V – 8. VI. 1977, Peck & Howden [col.] (CNCI); 1 ♂, P [arque] I [nternacional] L [a] A [mistad], Jurutungo, 1800 m, [08 ° 54 ' N, 82 ° 44 ' W], 30 oci. 5 nov. 1994, Tr [ampa] Malaise, A. Rodriguez [col.] (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C499B1FFF67F53E9DD0FC7A.taxon	distribution	Distribution (Fig. 225). Panama (Chiriquí).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C489B1FFF67F1C89DC1F894.taxon	description	Description. Male. Body length 3.13 mm. Color: head dark castaneous; mesosoma and metasoma castaneous. Head (Fig. 26). Mandible with two distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina low in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures large. Vertex crest straight. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 57). Tergal proces with lateral, shallow, subcircular and small depression, without tubercle, with very short tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 88). Genitalia (Figs 209 – 210): paramere with dorsal margin entirely wide; apical margin rounded. Aedeagal ventral ramus shorter than dorsal body, evenly narrow; cross section laminar; surface subvertical; inner margin concave; outer margin sinuous; apex short, simple, parallel; additional inner ramus absent; basal stub short and very narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and narrow, with apical margin abruptly acute and ventrad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring. Variations. Mesosoma dark castaneous; median clypeal tooth shorter; depression of tergal process very shallow.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C489B1FFF67F1C89DC1F894.taxon	discussion	Remarks. This species is easily recognized by having the very long basiparamere about 2 × the size of the paramere. This characteristic does not have in other species in Dissomphalus. The paramere is wide and short, the apex of the apical lobe of the aedeagal dorsal body is articulated and the eyes are bulging.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C489B1FFF67F1C89DC1F894.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Darién Pr [ovincia]: P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP). Paratypes: 2 ♂, same data as holotype; 3 ♂, same of holotype except 1000 m, 21 mar – 4 ab 2000 (2 ♂), 580 m, 21 mar – 4 ab 2000 (1 ♂) (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C489B1FFF67F1C89DC1F894.taxon	distribution	Distribution (Fig. 227). Panama (Darién).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4F9B18FF67F2A89B6DFAF0.taxon	description	Description. Male. Body length 1.73 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 27). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one rounded tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 58). Tergal process lateral, without depression, with very small, low and dorsad tubercle, pit very small, with short tuft of setae. Posterior hypopygeal margin sinuous (Fig. 89). Genitalia (Figs 211 – 212): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus higher than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin sinuous; apex long, simple, parallel; additional inner ramus absent; basal stub very short and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe very short, horizontal and narrow, with apical margin abruptly acute and slightly mesad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring. Variation. Pit of tubercle of tergal process without setae.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4F9B18FF67F2A89B6DFAF0.taxon	discussion	Remarks. This species is easily recognized by having the dorsal margin of the paramere straight and the apex very excavated in the apical region and the apical lobes of the aedeagal dorsal body very short. This species is similar to D. azagus sp. nov. by having the dorsal margin of the paramere straight and the apex very excavated. However, this species has the tergal process present, the apical lobes of the aedeagal dorsal body very short and the aedeagal ventral ramus progressively narrowing apicad, whereas D. azagus sp. nov has the tergal process absent, the apical lobes of the aedeagal dorsal body long and the apex of the aedeagal ventral ramus narrowing abruptly apicad.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4F9B18FF67F2A89B6DFAF0.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Veraguas Pr [ovincia]: P [arque] N [acional] Coiba, Isla Coiba, Cerro la Equis, 120 m, [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 29 jul – 4 ag 1998, Malaise, R. Cambra, S. Bermudez [col.] (MIUP). Paratypes: 1 ♂ same as holotype except 30 – nov. – 4 dic. 1998 (MIUP); 1 ♂ same as holotype except 200 m (MIUP); 1 ♂, Santa Fé, Altos de Piedra, 770 m [08 ° 31 ' 5.5 " N, 81 ° 06 ' 30.4 " W], 22 – 26 marzo 1999, Col. A. Santos, L. D. (MIUP); [Chiriqui]: 1 ♂, Formosa, Finca La Suiza 1 Km se. Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 28. V – 2. VI. 1994, Malaise Trap, F. Andrews & A. Gilbert [col.] (UCDC); Coclé Pr [ovincia]: 4 ♂, El Coclé, Par [que] Nac [ional General de División] Omar Torrijos H [errera], 570 – 670 m. s. n. m. [08 º 41 ' 56 " N, 80 º 39 ' 17 " W], 26 – 30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); [Panama Province]: 1 ♂, C [anal] Zone Barro Colorado I [sland], [09 ° 09 ' N, 79 ° 51 ' W], V. 11.1981, R. B. & L. S. Kimsey [col.] (UCDC).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4F9B18FF67F2A89B6DFAF0.taxon	distribution	Distribution (Fig. 232). Panama (Chiriqui, Veraguas, Coclé and Panama).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4F9B19FF67F7429DDBFEBB.taxon	description	Description. Male. Body length 1.16 mm. Color: head and metasoma castaneous; metasoma light castaneous. Head (Fig. 28). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one rounded tooth; median clypeal carina high in profile, complete or nearly so, angled in profile. Frons weakly coriaceous and punctures small. Vertex crest strongly convex. Pronotal disc polished; anterior margin ecarinate. Metasoma (Fig. 59). Tergal process lateral, without depression, low and dorsad tubercle, pit small, with long and dense tuft of setae. Posterior hypopygeal margin sinuous (Fig. 90). Genitalia (Figs 213 – 214): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus higher than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin sinuous; apex short, simple, slightly curved outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and narrow, with apical margin rounded and posterad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4F9B19FF67F7429DDBFEBB.taxon	discussion	Remarks. This species is similar to D. krombeini by having the paramere expanded basally and the aedeagal ventral ramus abruptly narrowing apicad. However, this species has the tergal process with tubercle with tuft of setae and the apical lobes of the aedeagal dorsal body longer, whereas the D. krombeini has the apical lobe of the aedeagal dorsal body shorter and the tergal process linear with very short tuft of setae with one distincty long seta.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4F9B19FF67F7429DDBFEBB.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Veraguas: P [arque] N [acional] Coiba, Isla Coiba, Cerro la Equis, 90 m, [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 30 – nov – 4 – dic. 1998, Malaise, R. Cambra, S. Bermudez [col.] (MIUP). Paratypes: 3 ♂, same data as holotype; 5 ♂ same as holotype except, 120 m (2 ♂), 200 m (1 ♂) (MIUP); 3 ♂ same as holotype except 90 m, 8 – 12 mar [zo] 1998 (1 ♂), 270 m, 7 – 13 mar 1998, R. Cambra, A. Santos [col.] (1 ♂), 29 jul – 4 ago 1998 (1 ♂) (MIUP).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4F9B19FF67F7429DDBFEBB.taxon	distribution	Distribution (Fig. 234). Panama (Veraguas).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4D9B1BFF67F6C99C99FD12.taxon	description	Description. Male. Body length 2.18 mm. Color: head and mesosoma castaneous; metasoma light castaneous. Head (Fig. 29). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one angulate very short tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 60). Tergal process lateral, without depression, with small, very low and dorsad tubercle, pit absent, with short and sparse tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 91). Genitalia (Figs 215 – 216): paramere with dorsal margin wide basally; apical margin rounded. Aedeagal ventral ramus shorter than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin sinuous; apex long, simple, parallel; additional inner ramus absent; basal stub very short and very narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, horizontal and wide, with apical margin abruptly acute and posterad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring. Variations. Head dark castaneous; median clypeal tooth inconspicuous; pit of tubercle of tergal process with short setae.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4D9B1BFF67F6C99C99FD12.taxon	discussion	Remarks. This species is easily recognized by having the paramere very expanded basally and four narrow projections that connect the apical lobes of the aedeagal dorsal body with the median portion of the aedeagal dorsal body and the tergal process with one minute tubercle without pit with few setae. This species is similar to the D. microtuberculatus, Azevedo by having the tergal process with two minutes tubercles. However, this species has the basiparamere shorter than the paramere, the apical lobes of the aedeagal dorsal body without flagella and the apex of the aedeagal ventral ramus progressively narrowing apicad, whereas D. microtuberculatus has the basiparamere longer than the paramere, the apical lobes of the aedeagal dorsal body with flagella and the apex of the aedeagal ventral ramus abruptly acute.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4D9B1BFF67F6C99C99FD12.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Coclé Pr [ovincia]: El Copé, Par [que] Nac [ional General de División] Omar Torrijos H [errera], 570 – 670 m. s. n. m., [08 º 41 ' 56 " N, 80 º 39 ' 17 " W], 26 – 30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP). Paratypes: Chiriqui: 1 ♂, La Fortuna Dam, 1200 m, wet forest, [08 ° 44 ' 39 " N, 82 ° 14 ' 58 " W], 14 – VI – 16. VII – 1982, F [light] I [nterception] T [rap], B. Gill [col.] (PMAE); 1 ♂, Fortuna, Div. Cont. Chiriqui-B [ocas del] Toro, 1070 – 1090 m, [08 ° 46 ' 16.5 " N, 82 ° 11 ' 27.3 " W], 8 – 11. abr. 1999, Malaise, R. Cambra, A. Santos [col.] (MIUP); Veraguas Pr [ovincia]: 9 ♂ P [arque] N [acional] Coiba, Isla Coiba, Cerro La Equis, 90 m (8 ♂) and 200 m (1 ♂) [07 º 28 ' 09 " N, 81 º 45 ' 24 " W], 30 – nov. – 4 dic. 1998, Malaise, R. Cambra, S. Bermudez [col.] (MIUP); 1 ♂, Pr [ovincia] Santa Fé, Altos De Piedra, [08 ° 31 ' 5.5 " N, 81 ° 06 ' 30.4 " W], 13 – 17 nov. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); 1 ♂, [Rio] Calovébora, [Rio] Guazarito, [08 º 42 ' N, 81 º 13 ' W], 15 – 18 feb. 2000, Col. S. Bermudez (MIUP); 1 ♂ same data as holotype; Darién: 2 ♂, Cana, 600 m, [07 ° 51 ' N, 87 ° 37 ' W], 3 – 7. VI. 1996, J. Ashe & R. Brooks [col.] (CNCI);	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4D9B1BFF67F6C99C99FD12.taxon	distribution	Distribution (Fig. 236). Panama (Veraguas, Darién, Coclé and Chiriquí).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4C9B1BFF67F1209C79F8F8.taxon	description	Description. Male. Body length 2.42 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 30). Mandible with two distal teeth. Median clypeal lobe subtrapezoidal, with one angulate tooth; median clypeal carina high in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin coarse. Metasoma (Fig. 61). Tergal process lateral, with deep, circular and large depression, with large, high and dorsad tubercle placed on center of depression, pit large, with short and dense tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 92). Genitalia (Figs 217 – 218): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus as long as dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin slightly sinuous; outer margin sinuous; apex long, simple, curved outward; additional inner ramus absent; basal stub short and very narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, subvertical and narrow, with apical margin rounded and posterad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring. Variation. Depression of tergal process shallow.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4C9B1BFF67F1209C79F8F8.taxon	discussion	Remarks. This species is easily recognized by having the paramere very wide, the apical lobes of the aedeagal dorsal body conical-shaped, the cuspis very small and the aedeagal ventral ramus wide basally. This species is similar to D. raziogus sp. nov. by having the aedeagal dorsal body narrowing slightly apicad and the apical lobes of the aedeagal dorsal body in conical-shaped. However, this species has the tergal process present, the paramere very wide and the apex of apical lobes of the aedeagal dorsal body dorsad, whereas D. raziogus sp. nov. has the tergal process absent, the paramere not wide and the apex of apical lobes of the aedeagal dorsal body mesad.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4C9B1BFF67F1209C79F8F8.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Coclé Pr [ovincia]: El Copé, Par [que] Nac [ional General de División] Omar Torrijos H [errera], 570 – 670 m. s. n. m., [08 º 41 ' 56 " N, 80 º 39 ' 17 " W], 26 – 30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP). Paratypes: 5 ♂, same data as holotype; Darién Pr [ovincia]: 1 ♂, P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 1000 m, [08 ° 00 ' N, 77 ° 45 ' W], 21 mar – 4 ab 2000, Malaise, Cambra, Santos, Bermudez [col.] (MIUP); 2 ♂, Cana, Serraria {Serrania} de Pirre, 1450 m, 4 – 7. vi. 1995 F [light] I [interception] T [rap], J Ashe, R. Brooks (UFES).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4C9B1BFF67F1209C79F8F8.taxon	distribution	Distribution (Fig. 238). Panama (Coclé and Darién).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4C9B64FF67F5509C02FC17.taxon	description	Description. Male. Body length 5.16 mm. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (Fig. 31). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina very high in profile, complete or nearly so, convex in profile. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc weakly coriaceous; anterior margin coarse. Metasoma (Fig. 62). Tergal process median, without depression, with small, low and dorsad tubercle, pit small, with very short and dense tuft of setae. Posterior hypopygeal margin straight or nearly so (Fig. 93). Genitalia (Figs 219 – 220): paramere with dorsal margin entirely wide; apical margin rounded. Aedeagal ventral ramus higher than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface subhorizontal; inner margin straight; outer margin sinuous; apex long, simple, slightly curved outward, additional inner ramus absent, basal stub long and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and narrow, with apical margin abruptly acute and ventrad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring. Variations. Body length 3.88 mm (min.) 5.16 mm (max.).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4C9B64FF67F5509C02FC17.taxon	discussion	Remarks. This species is easily recognized by having the aedeagal ventral ramus with median half wide and apical half narrow and parallel. The tergal process median, characterized by having two small tubercles without depressions and small tuft of setae. This species is similar to D. scamatus Azevedo by having the median tergal process, without depression, with tubercle, tuft of setae and the basiparamere narrow. However, this species has the apical lobes of the aedeagal dorsal body long with acute apex and aedeagal ventral ramus with inner margin straight, abruptly narrowing apicad, whereas D. scamatus has the apical lobes of the aedeagal dorsal body short, narrowing at apex, aedeagal ventral ramus with inner margin concave, with acute apex.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4C9B64FF67F5509C02FC17.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Chiriquí: 08 ° 54,7 ' N, 82 ° 47,3 ' W 17 – 18. vi. 1996, 1875 m, F [light] I [interception] T [rap], J. Ashe, R. Brooks (UFES). Paratypes: 1 ♂, same data as holotype; 1 ♂, Finca La Suiza near Hornitos, [08 ° 40 ' N, 82 ° 14 ' W], 25 – 6. VI. 2000, H. & A. Howden [col.] (CNCI); [Los Santos]: 3 ♂, Ojo de Agua, [07 ° 31 ' N, 80 ° 25 ' W], 30. VI – 14. VII. 1982, B. Gill [col.] (PMAE).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C4C9B64FF67F5509C02FC17.taxon	distribution	Distribution (Fig. 245). Panama (Chiriquí and Los Santos).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C339B65FF67F6239E38FC5F.taxon	description	Description. Male. Body length 1.61 mm. Color: body castaneous. Head (Fig. 32). Mandible with three distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina high in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (Fig. 63). Tergal process lateral, with deeply excaveted, subcircular and large depression covered by a membrane, without tubercle, pit absent. Posterior hypopygeal margin strongly concave (Fig. 94). Genitalia (Figs 221 – 222): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin slightly sinuous; apex short, simple, slightly curved outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, horizontal and wide, with apical margin abruptly acute and dorsad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring. Variations. Mesosoma light castaneous; metasoma castaneous; tergal process more laterad and longer.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C339B65FF67F6239E38FC5F.taxon	discussion	Remarks. This species is easily recognized by having the median clypeal carina with angled elevation, the paramere very excavated and the tergal process with deeply excaveted, subcircular and large depression covered by a membrane. This kind tergal process has been never found in Dissomphalus and it is easily recognized in the genus. This species is similar to the D. gilvipes by having the paramere wide basally and the aedeagal ventral ramus short with acute apex. However, this species has the apical margin of the paramere crenulate, the apical lobes of the aedeagal dorsal body wide and horizontal and the tergal process with very excavated depression covered by a membrane, whereas D. gilvipes has the paramere smooth, the apical lobes of the aedeagal dorsal body vertical and narrow and the tergal process with a tuft of setae with distinct long lateral setae.	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C339B65FF67F6239E38FC5F.taxon	materials_examined	Material examined. Holotype, ♂, PANAMA, Darién Pr [ovincia]: P [arque] N [acional] Darién, Pirre, Est [ación] Rancho Frio, 80 m, [08 ° 00 ' N, 77 ° 45 ' W], 16 nov – 17 ene 2001, Malaise, R. Cambra, A. Santos [col.] (MIUP). Paratypes: Bocas del Toro: 1 ♂, Wekso, Teribe, P [arque] I [nternacional] L [a] A [mistad], 50 msnm, [09 º 24 ' 25 " N, 82 º 56 ' 20 " W], 17 – 24 oct. 1999, Tr [ampa] Malaise, A. Santos [col.] (MIUP); [Chiriquí]: 2 ♂, Fortuna, Div. Cont. Chiriqui-B [ocas del] Toro, 1070 – 1090 m, [08 ° 46 ' 16.5 " N, 82 ° 11 ' 27.3 " W], 8 – 11. abr. 1999, R. Cambra, A. Santos [col.] (MIUP); Veraguas Pr [ovincia]: 1 ♂, Calovebora, Guazarito, 15 – 18 feb. 2000, malaise, Col. S. Bermudez (MIUP); 1 ♂, Santa Fé, Altos de Piedra, 770 m, [08 ° 31 ' 5.5 " N, 81 ° 06 ' 30.4 " W], 22 – 26 marzo 1999, Col. A. Santos, L. D. (MIUP); Colón: 5 ♂, 14 km N J [un] ct [ion] Escobal r [oa] d and Piña r [oa] d on Piña r [oa] d, ± 20 m, [09 ° 20 ' N', 79 ° 54 ' W], 2 – 3. VI. 1995, F [light] I [nterception] T [rap], J. Ashe & R. Brooks [col.] (CNCI); Coche {= Coclé}: 1 ♂, El Cope {= El Copé}, [08 ° 37 ' 11 " N, 80 ° 35 ' 03 " W], 19. XI. 1994, Windson & Edwards [col.] (CNCI); 6 ♂, Par [que] Nac [ional General de División] Omar Torrijos H [errera], 570 – 670 m. s. n. m., [08 º 41 ' 56 " N, 80 º 39 ' 17 " W], 26 – 30 sep. 1999, Tr [ampa] Malaise, A. Santos, P. González [col.] (MIUP); Panama Pr [ovincia]: 14 ♂, P [arque] Nac [ional] Sob [eranía], Camino, Plantación, [09 º 09 ' N, 79 º 43 ' W], 24 ago. – 2 sep. 1999, Malaise, A. Santos, P. González [col.] (MIUP); 5 ♂, P [arque] Nac [ional] Altos de Campana, [08 º 43 ' 15 " N, 80 º 01 ' 43 " W], 10 – 14 ago. 1999 (MIUP); 1 ♂, P [arque] N [atural] Metropolitano, [08 º 59 ' 33 " N, 79 º 32 ' 41 " W], 24 ago. – 3 sep. 1999 (MIUP); 1 ♂, Cerro Azul, Las Nubes, [09 º 10 ' N, 79 º 25 ' W], 9 – 15. oct. 1999, R. Cambra, A. Santos [col.] (MIUP); 7 ♂, C [anal] Z [one], B [arro] Colorado Is [land], [09 ° 09 ' N, 79 ° 51 ' W], 15 – 31. VII. 1994 (5 ♂), 1 – 14. VII. 1994 (1 ♂) and 16 – 24. VIII. 1994 (1 ♂) F [light] I [nterception] T [rap], D. Banks [col.] (CNCI); Comarca Kuna Yala: 1 ♂, Ustupu, Rio Abudi, [09 º 07 ' 57 " N, 77 º 55 ' 36 " W], 6 – 12 dic. 1999, Malaise, P. González [col.] (MIUP); 12 ♂, same data as holotype; 1 ♂, same as holotype except 1000 m, 21 mar – 4 ab 2000, Cambra, Santos, Bermudez [col.] (MIUP); 1 ♂, same as holotype except 450 m, 24 – 26 marzo 2001, col. A. Santos (MIUP); 19 ♂, Darién, Cruce de Mono, Est [ación] I [nstituto] N [acional de] Re [cusos] Na [turales] Re [novables], P [arque] Nac [ional] Darién, [07 ° 51 ' 26.3 " N, 77 ° 28 ' 12.7 " W], 5 – 28 feb 1993, R. Cambra [col.] (MIUP);	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
3B3687C07C339B65FF67F6239E38FC5F.taxon	distribution	Distribution (Fig. 248). Panama (Bocas del Toro, Chiriquí, Veraguas, Coclé, Colón, Panama, Comarca Kuna Yala and Darién).	en	Azevedo, Celso O. (2017): Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species. Zootaxa 4335 (1): 1-72, DOI: 10.11646/zootaxa.4335.1.1
