taxonID	type	description	language	source
391787E3FFA4B635FE17D6A3C980F9E6.taxon	description	Description. Male. Integument Color. Antenna, head, thorax, abdomen, and tarsi black; femora and tibiae brown with apices black. Vestiture. Antenna black with gray pubescence, median carina glossy, nearly covered by gray and pale yellow hairs on frons; pronotum with mixed gray, brown and black, with four spots, two anterior, two posterior; antescutellar area whitish-gray; scutellum covered with white pubescence; elytral pubescence composed of gray, pale yellow, and black hairs in contrasting pattern (Fig. 3); sides and venter with gray and pale yellow hairs in mottled patterns; short tuft of gray hairs in middle of first abdominal sternum. Legs gray with brown spot or band near apex of each femur and in middle of lateral face of each tibia. Pygidial vestiture gray-pale yellow with short, transverse dark bar in middle and paired longitudinal marks near margin of apex. Head. Antenna with first four segments and the last filiform, segment 5 – 10 subserrate; frons with very fine median carina extending from interocular fovea nearly to the epistomal suture; vertex finely punctate; labrum slightly sulcate; eyes rounded, dark brown to black; ocular sinus 0.50 to 0.53 width of eye. Prothorax. Subconical, pleura slightly concave; disk with a gray cruciform indentation separating four dark brown or black gibbosities Figs. 3, 4); gibbosity with vaguely defined paired brown blotches; base lobed at middle; apex rounded. Mesothorax and Metathorax. Scutellum small, triangular. Elytron evenly convex, slightly depressed in scutellar region; humerus prominent, with a fine, serrate, transverse carina connecting bases of striae 6 and 7, striae deep, narrow, strial punctures distinct, unisetose. Metepisternum microfoveolate with some scattered foveolae, unisetose. Hind coxa microfoveolate and setose, short area near trochanteral insertion with setose foveolae, small elongate lustrous area on anterior portion. Hind femur constricted basally and apically, expanded medially to about width of coxa, with a very shallowly sulcate ventral margin, with one sharp, small, spine on inner margin of sulcus one-fourth from apex of femur (Fig. 2); tibia with lateral and ventral carinae, inner face convex; apex with five spines, the outer lateral and inner ventral subequal, the others smaller; claws lobed at base. Abdomen. First abdominal sternum three times as long as second; second, third and fourth subequal; fifth slightly longer, deeply and broadly emarginate mesally; all segments, except the first, with a line of strong, black setae. Pygidium ovate, convex. Size. Length (pronotum-elytra) 2.04 to 3.45 mm. Width 1.11 to 2.16 mm. Maximum thoracic depth 1.14 to 2.25 mm. Genitalia. Median lobe with ventral valve U-shaped, with setae flanking anterior portion, dorsal valve less sclerotized and U-shaped; armature of internal sac with two anterior masses of spines, a median mass of fine spines and two basal subtriangular sclerites (Fig. 5). Lateral lobes cleft to 0.33 their length (Fig. 6). Female. Similar to male except the inner face of hind tibia flat, with lateral, dorsal and ventral carinae well developed, first sternum without tuft of hairs; fifth sternum not emarginate, pygidium somewhat gibbous at apex, gray-pale yellow with dark T-shaped mark in apical half connected with vague spots near apex, narrow median line gray-pale yellow. Size. Length (pronotum-elytra) 3.15 to 3.36 mm. Width 2.04 to 2.13 mm. Maximum thoracic depth 2.07 to 2.10 mm.	en	Romero, Jesus, Johnson, Clarence Dan (2003): Revision of the Genus NELTUMIUS Bridwell (Coleoptera: Bruchidae). The Coleopterists Bulletin 57 (2): 219-236, DOI: 10.1649/550, URL: http://dx.doi.org/10.1649/550
391787E3FFA4B635FE17D6A3C980F9E6.taxon	biology_ecology	Host Plants. Old records. Prosopis chilensis (Molina) Stuntz (Kingsolver 1964: 106; Ward et al. 1977: 5; Udayagiri and Wadhi 1989: 95). P. glandulosa Torr. (Kunhikannan 1923: 20; Zacher 1952: 471). P. juliflora (Sw.) DC. (Kingsolver 1964: 106;	en	Romero, Jesus, Johnson, Clarence Dan (2003): Revision of the Genus NELTUMIUS Bridwell (Coleoptera: Bruchidae). The Coleopterists Bulletin 57 (2): 219-236, DOI: 10.1649/550, URL: http://dx.doi.org/10.1649/550
391787E3FFA1B639FF37D15CC870FC10.taxon	description	1976: 196; Kingsolver et al. 1977: 115; Johnson 1978: 438, 1980: 28; Ward et al. 1977: 6; Johnson and Kingsolver 1981: 416; Johnson 1983 a: 28; Johnson 1983 b: 35; Johnson 1983 c: 32; Udayagiri and Wadhi 1989: 95. N. gibbothorax (sic): Bradley 1947: 36; Kingsolver 1964: 105. N. gibbothorae (sic): Blackwelder and Blackwelder 1948: 45. Description. Male. Integument Color. Head, thorax, abdomen, and tarsi black; femora and tibiae brown; antenna dark brown. Vestiture. Frons densely clothed with gray and pale yellow hairs; antenna black with gray pubescence; disk densely covered with pale yellow and gray hairs; antescutellar area lighter gray; scutellum covered with white pubescence; elytra clothed with brown, pale yellow, and gray hairs in distinctive pattern (Fig. 7); sides and venter with intermixed gray and pale yellow hairs, short tuft of gray hairs in middle of first abdominal sternum; legs evenly clothed with gray hairs; pygidium uniformly clothed with gray and pale yellow intermixed hairs, sometimes with indistinct darker spot in middle. Head. Antenna with first four segments and the last filiform, segments 5 – 10 subserrate; frons with very fine median carina extending from interocular fovea nearly to the epistomal suture; vertex finely punctate; labrum finely granulate; eyes rounded, dark brown; ocular sinus 0.55 width of eye. Prothorax. Subconical, pleura slightly concave; gibbosity with vaguely defined paired brown blotches; base lobed at middle; apex rounded. Mesothorax and Metathorax. Scutellum small, triangular. Elytron evenly convex, slightly depressed in scutellar region; humerus prominent, with a fine, serrate, transverse carina connecting bases of striae 6 and 7, striae deep, narrow, strial punctures unisetose. Metepisternum finely microfoveolate. Hind coxa microfoveolate, setose, short area near trochanteral insertion with setose foveolae, small elongate lustrous area on anterior portion. Hind femur constricted basally and apically, expanded medially to about width of coxa; with very shallowly sulcate ventral margin and one small, sharp spine on medial margin of sulcus 0.25 from apex of femur (Fig. 2); tibia sometimes with but usually without lateral, medial and ventral carinae; apex with five spinules, outer lateral spinule longer than others; claws lobed at base. Abdomen. First abdominal sternum three times as long as second, with a carina 0.5 of the length of the sternum width near anterior margin; second, third and fourth sterna subequal; fifth slightly longer, deeply, broadly emarginate mesally; sterna 2 – 5 with a line of strong, black setae. Pygidium ovate and convex. Size. Length (pronotum-elytra) 2.67 mm. Width 1.14 mm. Maximum thoracic depth 1.5 mm. Genitalia. Median lobe with ventral valve U-shaped, with a group of setae flanking anterior portion, dorsal valve less sclerotized and U-shaped; armature of internal sac with two anterior masses of spines, a median mass of fine spines mixed with fine scale-shaped spines and two basal subtriangular sclerites (Fig. 9). Lateral lobes cleft to 0.33 their length (Fig. 10). Female. Similar to male except first sternum without tuft of hairs; fifth sternum not emarginate, pygidium gibbous at apex. Size. Length (pronotum-elytra) 2.37 mm. Width 1.47 mm. Maximum thoracic depth 1.5 mm.	en	Romero, Jesus, Johnson, Clarence Dan (2003): Revision of the Genus NELTUMIUS Bridwell (Coleoptera: Bruchidae). The Coleopterists Bulletin 57 (2): 219-236, DOI: 10.1649/550, URL: http://dx.doi.org/10.1649/550
391787E3FFA1B639FF37D15CC870FC10.taxon	biology_ecology	Host Plants. Old records: Prosopis odorata Torr. and Frém. (Kingsolver 1964: 106; Kingsolver et al. 1977: 115; Ward et al. 1977: 6; Udayagiri and Wadhi 1989: 95). P. pubescens Bentham (Kingsolver 1964: 106; Center and Johnson 1976: 200; Ward et al. 1977: 5; Johnson 1978: 438; Johnson 1980: 28; Udayagiri and Wadhi 1989: 95)	en	Romero, Jesus, Johnson, Clarence Dan (2003): Revision of the Genus NELTUMIUS Bridwell (Coleoptera: Bruchidae). The Coleopterists Bulletin 57 (2): 219-236, DOI: 10.1649/550, URL: http://dx.doi.org/10.1649/550
391787E3FFA1B639FF37D15CC870FC10.taxon	distribution	Distribution. Old Records: USA (Arizona, California, Nevada, Utah) and Mexico (Kingsolver 1964).	en	Romero, Jesus, Johnson, Clarence Dan (2003): Revision of the Genus NELTUMIUS Bridwell (Coleoptera: Bruchidae). The Coleopterists Bulletin 57 (2): 219-236, DOI: 10.1649/550, URL: http://dx.doi.org/10.1649/550
391787E3FFA1B639FF37D15CC870FC10.taxon	discussion	Discussion. This species is closely related to N. arizonensis, from which it can easily be distinguished by the uniformly gray tibiae, anteriorly placed prothoracic gibbosity (Fig. 8), predominantly gray vestiture, the gibbosities are more markedly defined (Fig. 8), and vaguely marked pygidium. The male genitalia of both are almost indistinguishable, except that N. gibbithorax has a group of setae flanking the anterior portion that are more numerous, the median mass of fine spinules is mixed with fine scale-shaped spines, and the two basal subtriangular sclerites are slightly more elongated. There is considerable flux at present in the taxonomy of the genus Prosopis, but Prosopis pubescens is considered to be the valid name of this host, the screwbean mesquite. Some specialists in the genus advocate that there are only two species of mesquite (screwbeans and the others) and the rest of the 68 names are subspecies (Hultine 2001). According to the available evidence, there is no interbreeding between the screwbean group and the others. Thus, N. gibbithorax and N. arizonensis follow taxonomic lines in their host preferences.	en	Romero, Jesus, Johnson, Clarence Dan (2003): Revision of the Genus NELTUMIUS Bridwell (Coleoptera: Bruchidae). The Coleopterists Bulletin 57 (2): 219-236, DOI: 10.1649/550, URL: http://dx.doi.org/10.1649/550
391787E3FFADB63CFE25D5BCC9B1FB5B.taxon	description	Description. Male. Integument Color. Antennae, head, thorax, abdomen, and tarsi black; femora and tibiae brown with apices black. Vestiture. Antennae black with white pubescence; frons sparsely covered with white and pale yellow hairs; prothorax white with vaguely defined paired brown spots at apex; scutellum covered with white pubescence; elytral pubescence white and black hairs in contrasting pattern; sides and venter with white hairs in mottled patterns (Fig. 11); legs clothed with white and pale yellow hairs with median brown band on posterolateral face of each tibia; without tuft of hair on first abdominal sternum. Pygidium evenly convex, whitish-gray in basal third, mixed white and pale yellow with narrow median gray line in apical third. Head. Antenna with first four segments and the last filiform, segments 5 – 10 subserrate, frons with median carina glossy black, head finely punctate; labrum bare, somewhat sulcate, yellowish brown at apex; eyes rounded, dark brown; ocular sinus 0.6 to 0.62 width of eye. Prothorax. Subconical, pleura slightly concave; disk convex with paired brown, obsolete tuberosities near base flanking triangular white antescutellar patch; base strongly lobed at middle, sinuate laterally, posterior corners acute; apex rounded. Mesothorax and Metathorax. Scutellum small, subquadrate; elytron evenly convex; humerus prominent, granulate, lacking serrate carina connecting bases of striae 6 and 7; striae narrow, deep with setiferous punctures hardly discernible. Metepisternum microfoveolate with some scattered foveolae unisetose. Hind coxa microfoveolate, setose, short area near trochanteral insertion with setose foveolae, small elongate lustrous area on anterior portion. Hind femur constricted basally and apically, expanded medially to about width of coxa, with very shallowly sulcate ventral margin, with one small, sharp spine on inner margin of sulcus one-fourth from apex of femur (Fig. 2); tibia with lateral and ventral carinae, inner face convex; apex with five spinules, outer lateral and inner ventral spinules subequal, others smaller; claws lobed at base. Abdomen. First abdominal sternum three times as long as second; second, third and fourth subequal; fifth slightly longer and slightly emarginate mesally; all segments, except the first, with a line of strong, black setae. Pygidium ovate and convex. Size. Length (pronotum-elytra) 1.5 to 2.25 mm. Width 0.87 to 1.5 mm. Maximum thoracic depth 1.11 to 1.29 mm. Genitalia. Median lobe with ventral valve deltoid, dorsal valve membranous; internal sac clothed anteriorly with minute spines, armature formed by large cluster of strong spines interrupted mesally by group of finely dentate scales (Fig. 13). Lateral lobes armed at apices with stout spines, lobes cleft to 0.45 their length (Fig. 14). Female. Similar to male except inner face of hind tibia flat, with lateral, dorsal and ventral carinae well developed, fifth sternum not emarginate; pygidium with triangular gray spot in middle of base flanked by pale yellow spot, faint brown, transverse median band interrupted by median line of gray; paired vague brown spots near apical margin. Size. Length (pronotum-elytra) 2.04 to 2.10 mm. Width 1.2 mm. Maximum thoracic depth 1.05 to 1.2 mm.	en	Romero, Jesus, Johnson, Clarence Dan (2003): Revision of the Genus NELTUMIUS Bridwell (Coleoptera: Bruchidae). The Coleopterists Bulletin 57 (2): 219-236, DOI: 10.1649/550, URL: http://dx.doi.org/10.1649/550
391787E3FFADB63CFE25D5BCC9B1FB5B.taxon	biology_ecology	Host Plants. Old records: Condalia correllii M. C. Johnston (Johnson 1978: 436; Udayagiri and Wadhi 1989: 96). C. globosa I. M. Johnston (Hetz and Johnson 1988: 137). C. globosa var. pubescens I. M. Johnston (Center and Johnson 1976: 200; Johnson 1978: 436; Udayagiri and Wadhi 1989: 96). C. hookeri M. C. Johnston (Johnson 1995: 42). C. hookeri var. hookeri M. C. Johnston (Johnson 1978: 436; Udayagiri and Wadhi 1989: 96). C. spathulata A. Gray (Johnson 1978: 435; Hetz and Johnson 1988: 135; Udayagiri and Wadhi 1989: 96). C. warnockii var. kearneyana M. C. Johnston (Johnson 1978: 434; Udayagiri and Wadhi 1989: 96).	en	Romero, Jesus, Johnson, Clarence Dan (2003): Revision of the Genus NELTUMIUS Bridwell (Coleoptera: Bruchidae). The Coleopterists Bulletin 57 (2): 219-236, DOI: 10.1649/550, URL: http://dx.doi.org/10.1649/550
391787E3FFADB63CFE25D5BCC9B1FB5B.taxon	distribution	Distribution. Old Records: USA (Arizona, California, and Texas) (Johnson 1978; Kingsolver 1964). New Records: Mexico (Nuevo Leon, Queretaro, San Luis Potosi, and Veracruz).	en	Romero, Jesus, Johnson, Clarence Dan (2003): Revision of the Genus NELTUMIUS Bridwell (Coleoptera: Bruchidae). The Coleopterists Bulletin 57 (2): 219-236, DOI: 10.1649/550, URL: http://dx.doi.org/10.1649/550
391787E3FFADB63CFE25D5BCC9B1FB5B.taxon	discussion	Discussion. Neltumius texanus is more similar to N. dospatrias than to N. arizonensis and N. gibbithorax. See discussion of N. dospatrias for differences between the two species. The great difference in the Rhamnaceous host plants preferred by this species and N. arizonensis and N. gibbithorax that prefer seeds of Prosopis is remarkable. Johnson (1978, 1989, 1995) discussed the probable origins of these closely related species feeding in plants in different families, and the ecology of N. texanus in more detail.	en	Romero, Jesus, Johnson, Clarence Dan (2003): Revision of the Genus NELTUMIUS Bridwell (Coleoptera: Bruchidae). The Coleopterists Bulletin 57 (2): 219-236, DOI: 10.1649/550, URL: http://dx.doi.org/10.1649/550
391787E3FFABB63EFEA2D1B9C968FD22.taxon	description	Description. Male. Integument Color. Antennae, head, thorax, abdomen, and tarsi black; femora and tibiae brown with apices black. Vestiture. Antennae black with white pubescence; frons covered with sparse to moderately dense intermixed white and brown hairs, vertex with sparse reddish brown hairs; prothorax with recumbent reddish-brown hairs interrupted by 4 – 6 patches of white hairs, antescutellar area covered with dense white hairs forming a small triangle; scutellum covered with sparse white pubescence; elytral pubescence white, reddish brown, and black hairs in contrasting pattern (Fig. 15); sides and venter with dense white and reddish brown hairs in mottled pattern; legs clothed with sparse white hairs with median brown band on posterolateral face of each tibia; without tuft of hair on first abdominal sternum; pygidium evenly convex, small, medial, triangular patch of white hairs at base, sometimes apex with thin line of white hairs extending to 0.5 length of pygidium, remainder of pygidium with reddish-brown hairs, sometimes with small, intermixed patches of white hairs. Head. Antenna with segments 1 – 4 and 11 filiform, segments 5 – 10 subserrate, 11 acute at apex; frons with strong median glabrous carina extending from frontoclypeal suture to vertex, head with many deep punctations; labrum finely punctate, not sulcate, yellowish brown at apex; eyes rounded, dark brown; ocular sinus 0.6 to 0.62 width of eye. Prothorax. Subconical, pleura slightly concave; disk convex with paired brown, obsolete tuberosities near base flanking triangular white antescutellar patch; base strongly lobed at middle, sinuate laterally, posterior corners acute; apex rounded. Mesothorax and Metathorax. Scutellum very small, subquadrate; elytron evenly convex; humerus prominent, granulate, dark brown to black, lacking serrate carina connecting bases of striae 6 and 7; striae narrow, deep with setiferous punctures hardly discernible. Metepisternum microfoveolate with some scattered foveolae unisetose. Hind coxa microfoveolate, setose, short area near trochanteral insertion with setose foveolae, small elongate lustrous area on anterior portion. Hind femur constricted basally and apically, expanded medially to about width of coxa, with very shallowly sulcate ventral margin, with one small, sharp spine on inner margin of sulcus one-fourth from apex of femur (Fig. 2); tibia with dorsomesal, lateral and ventral carinae, inner face convex; apex with about five spinules, outer lateral and inner ventral spinules usually subequal, others shorter; claws lobed at base. Abdomen. First abdominal sternum three times as long as second; 2 – 4 subequal; fifth slightly longer, slightly emarginate apically; sterna without lines of strong, black setae. Pygidium ovate and convex. Size. Length (pronotum-elytra) 1.5 to 2.3 mm. Width 0.8 to 1.3 mm. Maximum thoracic depth 0.7 to 1.2 mm. Genitalia. Median lobe elongate; in ventral view, ventral valve broader than apex of median lobe, sides slightly concave, apex broadly blunt; dorsal valve membranous, gently rounded at apex; armature of internal sac with narrow, medial chain of spines near apex, chain attached to a broad, elongate cluster of small spines occupying medial portion of internal sac, interrupted near base by group of finely dentate scales attached to large cluster of minute spines at base (Fig. 17). Lateral lobes expanded at apex with many short setae, lateroventral surface of apices strongly sclerotized on periphery, without stout spines, lobes cleft to 0.8 their length (Fig. 18). Female. Similar to male except fifth sternum not emarginate. Size. Length (pronotum-elytra) 1.9 to 2.2 mm. Width 1.1 to 1.2 mm. Maximum thoracic depth 1.0 to 1.2 mm.	en	Romero, Jesus, Johnson, Clarence Dan (2003): Revision of the Genus NELTUMIUS Bridwell (Coleoptera: Bruchidae). The Coleopterists Bulletin 57 (2): 219-236, DOI: 10.1649/550, URL: http://dx.doi.org/10.1649/550
391787E3FFABB63EFEA2D1B9C968FD22.taxon	biology_ecology	Host Plants. Unknown.	en	Romero, Jesus, Johnson, Clarence Dan (2003): Revision of the Genus NELTUMIUS Bridwell (Coleoptera: Bruchidae). The Coleopterists Bulletin 57 (2): 219-236, DOI: 10.1649/550, URL: http://dx.doi.org/10.1649/550
391787E3FFABB63EFEA2D1B9C968FD22.taxon	materials_examined	Type Series. Male holotype, female allotype, and eight paratypes: Mexico. Nuevo Leon: 79 mi. S. Linares, Hwy. 57, VI- 23 1971, 6,7009, C. O’Brien and Marshall. Fourteen paratypes: Mexico. Queretaro; 8 km de la carr. a San Joaquín, San Joaquín, 18 / V / 2001; 208549220 N, 998409100 W; Col. J. L. Cozar; sobre flores de Condalia mexicana Schltdl. One paratype: USA. Arizona: Pima Co., Tucson, 28 mi. N. VI- 17 - 63, J. Doyen, Collector. Holotype, allotype, and six paratypes deposited in the USNMNH. One paratype deposited in the C. D. Johnson Collection. Other paratypes deposited in the following collections in Mexico: J. Romero Collection; Coleccion Entomologica de la Universidad Autonoma de Queretaro, Queretaro; Coleccion de la Universidad Nacional Autonoma de Mexico; Coleccion Entomologica del Instituto de Ecología A. C., Xalapa, Veracruz; Coleccion Nacional de Insectos, Instituto Nacional de Investigaciones Forestales y Agropecuarias, Campo Experimental El Bajio, Celaya, Guanajuato.	en	Romero, Jesus, Johnson, Clarence Dan (2003): Revision of the Genus NELTUMIUS Bridwell (Coleoptera: Bruchidae). The Coleopterists Bulletin 57 (2): 219-236, DOI: 10.1649/550, URL: http://dx.doi.org/10.1649/550
