taxonID	type	description	language	source
3852D52EFFC43B20FF6EFA22B539A172.taxon	discussion	Remarks. Since its description, the genus consists of two species, A. chong Ingrisch, 2006 from Thailand and Singapore (Ingrisch, 2006; Tan, 2012, 2017) and A. kinabalu Ingrisch, 2006 from Borneo.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC43B23FF6EF94AB212A63E.taxon	description	(Fig. 1)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC43B23FF6EF94AB212A63E.taxon	materials_examined	Material examined. 1 male (SDK. 19.103) and 1 female (SDK. 19.104), EAST MALAYSIA, Sandakan, Kabili Sepilok Forest Reserve, N 5.87084, E 117.93764, 98.7 ± 6.1 m. a. s. l., mating on foliage, 3 October 2019, 2058 h, coll. M. K. Tan & J. L. Yukang (FRC).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC43B23FF6EF94AB212A63E.taxon	distribution	Distribution. Borneo (Sabah: Sorinsim on Kinabalu, Sepilok in Sandakan)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC43B23FF6EF94AB212A63E.taxon	discussion	Remarks. The specimens reported agree with the original description by Ingrisch (2006) in general habitus, maxillary palps, male abdominal apex, male phallic complex and female ovipositor (Fig. 1). The locality Sepilok is a new record for this species that was described and previously known only from Kinabalu. While Kinabalu is a mountainous terrain, Sandakan has a lower and flatter terrain.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC63B22FF6EFF7FB250A13D.taxon	description	(Figs. 2 – 4)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC63B22FF6EFF7FB250A13D.taxon	materials_examined	Material examined. Holotype, male (SDK. 19.20), EAST MALAYSIA, Sabah, Sandakan, Sepilok, Rainforest Discovery Centre, N 5.87580, E 117.94299, 39.1 ± 5.4 m. a. s. l., leaf litter, 9 January 2019, 1911 h, coll. M. K. Tan, R. Japir, M. Binti, J. L. Yukang (FRC). Paratypes, 2 females: 1 female (SDK. 19.19), same locality, N 5.87356, E 117.94172, 49.1 ± 5.2 m. a. s. l., leaf litter, 9 January 2019, 1105 h, coll. M. K. Tan, R. Japir, M. Binti, J. L. Yukang; 1 female (SDK. 19.35), same locality, N 5.87553, E 117.94116, 33.4 ± 4.9 m. a. s. l., 9 January 2019, 2040 h, coll. M. K. Tan, R. Japir, M. Binti, J. L. Yukang (all FRC).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC63B22FF6EFF7FB250A13D.taxon	discussion	Remarks. Ingrisch (2006) described the female of this species as Cycloptiloides sp. 1 without providing a species name. Here, we provide a formal name, assign a male holotype and describe the male.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC63B22FF6EFF7FB250A13D.taxon	diagnosis	Diagnosis. The male differs from congeners by the supra-anal plate trapezoidal in shape with apical margin broadly rounded, with dark scales and yellow brown setae laterally and shiny pale in the middle (forming two spots). The female differs from congeners by shorter maxillary palps and ovipositor and more transverse supra-anal plate (Ingrisch, 2006).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC63B22FF6EFF7FB250A13D.taxon	etymology	Etymology. The species name refers to the two spots in the middle of the supra-anal plate; bi = two in Latin, maculata = spotted in Latin.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC63B22FF6EFF7FB250A13D.taxon	description	Description. Habitus of male as shown in Figs. 2, 3. Dorsum of head flattened (Fig. 3 A). Frontal rostrum 1.2 times wider than scapus. Maxillary palps with apical (= fifth) a little longer than subapical (= fourth) and third segments, subapical and third segments subequal lengths; apical segment little widened (Fig. 4 A). Pronotum about 1.5 times longer than wide with anterior dorsal margin distinctly concave; lateral margin subparallel in anterior half and faintly widening posteriorly; covering tegmen entirely; posterior margin convex. Fore tibia with internal tympanum large and elongate oval (Fig. 4 B); without external tympanum. Hind femur 1.4 times longer than hind tibia; hind tibia 2.1 times longer than hind metatarsus. Male. Supra-anal plate trapezoidal with apical margin broadly rounded, with setae at lateral parts (Figs. 4 C, 4 D). Subgenital plate trapezoidal, posterior margin slightly convex (Fig. 4 F). Epiphallus and phallic complex hyaline (Fig. 4 E). Female. Already sufficiently described in Ingrisch (2006). Our female specimens from Sandakan agree with the description. Colouration. General colouration similar to female described in Ingrisch (2006). Male supra-anal plate with dark scales laterally, middle shining pale coloured (forming two spots) (Figs. 4 C, 4 D). Cercus (both male and females) with brown scales. Measurements (in mm). Holotype BL = 3.9; FRW = 0.27; SW = 0.14; EW = 0.14; PL = 1.9; PW = 1.3; HFL = 2.5; HFW = 0.9; HTL = 1.8; HML = 0.8.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC63B22FF6EFF7FB250A13D.taxon	distribution	Distribution. Borneo (Sabah: Kinabalu, Sandakan)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC63B22FF6EF987B25DA0EE.taxon	synonymic_list	Ectatoderus angusticollis Chopard, 1969 from Singapore Ectatoderus apterus (Chopard, 1925) from Java Ectatoderus argentatus Ingrisch, 2006 from Thailand Ectatoderus marginatus Bey-Bienko, 1966 from Lesser Sunda Islands Ectatoderus pallidegeniculatus Brunner von Wattenwyl, 1893 from Myanmar [probably Micrornebius] Ectatoderus samui Ingrisch, 2006 from Thailand Ectatoderus sordidus (Walker, 1869) from New Guinea	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC03B28FF6EF932B3F2A6F1.taxon	description	(Figs. 5 – 7)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC03B28FF6EF932B3F2A6F1.taxon	materials_examined	Material examined. Holotype, male (BRU. 19.70), BRUNEI DARUSSALAM, Belait District, Jalan Labi near Andulau Forest Reserve, N 4.63354, E 114.51047, 76.6 ± 5.0 m. a. s. l., on rattan foliage, 7 July 2019, 2042 h, coll. M. K. Tan (UBDM).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC03B28FF6EF932B3F2A6F1.taxon	diagnosis	Diagnosis. The new species is characterised by the scapus being dorsally black but ventrally and laterally brown, and the antennae being wholly black; the head has a distinct black band behind eyes; and the male phallic complex has the medial valve elongated, not curved to a spiral, sinuous both dorso-ventrally and laterally and with the ventro-external sclerite of the lateral valve being elongated, in basal third broader, and tapering into a narrow lamella with subacute apex. The new species is closest in distribution to E. angusticollis Chopard, 1969 from Singapore. In both species the pronotum is more strongly widening apicad, and fore and middle tibiae are provided with black dorsal stripes. The new species differs by the scapus and antennae being black instead of testaceous, by the head being provided with a distinct black band behind eyes and by the male phallic complex having the medial valve not curved to a spiral. The new species is most similar to E. argentatus Ingrisch, 2006 from Thailand having the pronotum strongly widening apicad and the frons dark; it differs by the darked scapus and basal antennal segments instead of being yellow brown, by the male phallic complex which is more elongated and having the medial valve sinuous rather than curving upward (basally) and by the shape of the apical sclerotization of the medial valve. This new species also differs from E. samui Ingrisch, 2006 from Thailand by the pronotum being more strongly widening apicad and by the male phallic complex having the medial valve not curved to a spiral; from E. marginatus Bey-Bienko, 1966 from Lesser Sunda Islands it differs by the pronotum being more strongly widening apicad.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC03B28FF6EF932B3F2A6F1.taxon	etymology	Etymology. The species name refers to the black band posterior of eyes and the black scapus (dorsally only) and antennae; nigro = black in Latin, fascia = band in Latin.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC03B28FF6EF932B3F2A6F1.taxon	description	Description. Habitus of male as shown in Fig. 5. Dorsum of head flattened (Figs. 6 A, 6 B). Frontal rostrum 1.3 times wider than scapus (Fig. 6 A). Maxillary palps with apical (= fifth) and subapical (= fourth) segments of subequal length, third segment longer than apical and subapical segments; apical segment triangular and widened apically (Fig. 6 B). Pronotum about 1.7 times longer than wide with anterior dorsal margin narrow and straight; lateral margin widening posteriorly; strongly produced posteriorly and covering near-entirely tegmen; posterior margin convex. Fore tibia with internal tympanum small and oval; without external tympanum. Hind femur 1.1 times longer than hind tibia; hind tibia 4.4 times longer than hind metatarsus. Male. Supra-anal plate with last abdominal tergite and epiproct distinctly separated by a transverse suture; last abdominal tergite transverse, with hind margin emarginate; supra-anal plate less transverse than last abdominal tergite, flattened, with fewer setae, apex rounded (Fig. 6 E). Genitalia as in Fig. 7. Epiphallus elongated and membranous. Medial valve elongated, not curved to a spiral, sinuous both dorso-ventrally and laterally. Lateral valves weakly sclerotised, forming stout and truncated lobes at apex. Ventro-external sclerite of lateral valve elongate, basal third broader, tapering into a narrow lamella with subacute apex. Female. Unknown. Colouration. Head dorsum light brown, with white stripe running along lateral margin from posterior end, along inner margin of eyes to frontal rostrum (Figs. 6 B, 6 D). Scapus dorsally black, otherwise brown (Figs. 6 A, 6 D); basal antennal segments black, thereafter brown with some segments black (Figs. 6 A, 6 D). Frons and mouthparts dark coloured (Fig. 6 C). Maxillary and labial palps yellow brown (Figs. 6 B, 6 C). Lateral parts of head, including genae, light brown; with a black horizontal band behind eyes. Pronotal dorsal disc brown, apical margin with white scales. Pronotal lateral lobes also with white scales and with brown margins (Fig. 6 D). Tegmen mostly infumated white, but infumated grey at apical area of lateral field (Fig. 6 D). Femur generally pale brown to brown. Fore and middle tibiae dorsally black, otherwise brown. Tarsi with basal segment brown, with middle and apical segments black. Hind femur generally brown with knee dark. Hind tibia brown, dorsally pale yellow. Abdomen with tergites black covered with greyish brown scales; sternites black with whitish scales especially on those nearer to thorax. Abdominal apex black; last abdominal tergite back with brown setae along posterior margin (Fig. 6 E). Cercus brown. Measurements (in mm). Holotype BL = 7.6; FRW = 0.28; SW = 0.22; EW = 0.26; PL = 3.9; PW = 2.2; TL = 2.3; HFL = 4.3; HFW = 1.3; HTL = 4.0; HML = 0.9.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFC03B28FF6EF932B3F2A6F1.taxon	distribution	Distribution. Borneo (Brunei Darussalam)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFCC3B28FF6EFDCAB594A470.taxon	materials_examined	Material examined (image). Holotype, male, SINGAPORE, coll. C. F. Baker (MNHN-EO-ENSIF 4487). Material examined. 2 males (F 4. F. 12, H 12. F. 05), SINGAPORE, Mandai, secondary forest between Mandai Road and Upper Seletar Reservoir, 16 – 26 November 2015, night, coll. M. K. Tan (ZRC); 2 males (ZRC. ORT. 467), SINGAPORE, Lower Pierce Reservoir Park, secondary forest, 10 August 2012, coll. M. K. Tan.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFCC3B28FF6EFC04B452A3FD.taxon	discussion	Remarks. This species group currently consists of Ornebius fastus Yang & Yen, 2001 from Taiwan, Ornebius flori Ingrisch, 1998 and Ornebius marginatus Ingrisch, 1998 from Sabah, Ornebius pullus Ingrisch, 2006 from Brunei, and Ornebius xinyao Tan, 2015 from Singapore. The species are characterised by the male phallic complex having the lateral valves membranous, dorso-ventrally compressed and acute apically; internal sclerites of medial valve almost forming a tube and with apex of ejaculatory duct truncated and little notched (He et al., 2021).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFCC3B2DFF6EFAC7B1ABA181.taxon	description	(Figs. 8 – 10)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFCC3B2DFF6EFAC7B1ABA181.taxon	materials_examined	Material examined. 1 male (BRU. 19.13), BRUNEI DARUSSALAM, Belait District, Jalan Labi near Andulau Forest Reserve, N 4.62238, E 114.51047, 101.3 ± 6.4 m. a. s. l., on foliage, 24 February 2019, 2052 h, coll. M. K. Tan & H. Yeo (ZRC); 1 male (BRU. 19.74), BRUNEI DARUSSALAM, Belait District, Jalan Labi near Andulau Forest Reserve, N 4.63352, E 114.51061, 89.0 ± 7.1, on foliage, 8 July 2019, 2119 h, coll. M. K. Tan (UBDM). Other material examined. 1 male (D 18. F. 03), SINGAPORE, Mandai, secondary forest between Mandai Road and Upper Seletar Reservoir, 13 November 2015, night, coll. M. K. Tan (ZRC).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFCC3B2DFF6EFAC7B1ABA181.taxon	discussion	Remarks. The male specimens from Belait District agree with the original description of O. pullus (see Ingrisch, 2006) from Brunei-Muara District of Brunei Darussalam by habitus, size (Figs. 8, 9), shapes and colours of the epiproct and the paraproct process, as well as by the shape of phallic complex. However, the colour of the tegmen can be variable. While in one male (BRU. 19.13) the colouration of the tegmen agrees with that of the holotype from Brunei-Muara, in the other male (BRU. 19.74) the colour of the tegmen is somewhat intermediate between those of O. pullus and O. flori from Kinabalu in Sabah. Tan & Ingrisch (2013) previously noted that this species may have also occurred in Singapore, although there are some variations from the Bruneian specimens. Specifically, the apices of the membranous lateral valves of the male phallic complex can vary in their length and prominence and the tegmen colouration of the Singapore specimens is darker than those from Brunei (Fig. 10). We postulate that O. pullus from Brunei and Singapore may either represent a species complex or a polymorphic species. Molecular phylogeny may be useful to resolve this problem.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
3852D52EFFCC3B2DFF6EFAC7B1ABA181.taxon	distribution	Distribution. Within Brunei Darussalam, this species can be found in the western areas (i. e., Belait and Brunei-Muara Districts, it is probably also found in Tutong District between Belait and Brunei-Muara), but not in the eastern part of Brunei (i. e., Temburong District) (see Tan & Wahab, 2018). One potential explanation is that O. pullus is found in the flatter terrains in Belait and Brunei-Maura Districts rather than the hilly terrains of Temburong District.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Wahab, Rodzay Bin Haji Abdul (2021): New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo. Zootaxa 5048 (3): 407-421, DOI: 10.11646/zootaxa.5048.3.6
