taxonID	type	description	language	source
3F45F66CFFC0EF10FF77FE67FE555C4D.taxon	materials_examined	Material examined. PAKISTAN, Khyber Pakhtunkhwa: 3 specimens, Abbottabad, 11. VII. 2007, leg. & det. M. Abbas, 7 specimens, Kohat, 11. VIII. 2010, leg. & det. M. Abbas, 4 specimens, Kurram Agency, 11. VII. 2012, leg. & det. M. Abbas, 5 specimens, Besham, 11. VIII. 2012, leg. & det. M. Abbas, 3 specimens, Peshawar, 11. VIII. 2013, leg. & det. M. Abbas, 1 specimen, Buner, 11. VIII. 2013, leg. & det. M. Abbas, 1 specimen, Malakand, 1. VII-VIII. 2000, leg. M. Bashir, det. S. Khaliq; Sindh: 1 specimen, Karachi, 1. V. 1980, leg. Anonymous, det. S. Khaliq.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC0EF10FF77FE67FE555C4D.taxon	distribution	Distribution. Khyber Pakhtunkhwa: Abbottabad, Besham, Kohat, Kurram Agency (Abbas et al., 2015; Moretto & Minetti, 2022).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC0EF10FF76FAC1FD615A91.taxon	materials_examined	Material examined. INDIA, Assam: 1 specimens, Garo Hill, Tura, 6. VII. 1917, leg. Anonymous, det. S. Khaliq.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC0EF10FF76FAC1FD615A91.taxon	distribution	Distribution. India (not recorded from Pakistan).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC0EF10FF76FC86FEEF5B8E.taxon	materials_examined	Material examined. PAKISTAN, Islamabad Capital: 1 specimen, 14. VII. 1991, leg. Ahmad, det. Khaliq, 1 specimen, 14. XII. 1980, leg. Anonymous, det. S. Khaliq; Punjab: 1 specimen, Khabikki Lake, Khushab, 3 - 7. VIII. 2007, leg. M. Abbas, det. S. Khaliq; Khyber Pakhtunkhwa: 9 specimens, Buner, 11. VIII. 2007, leg. & det. M. Abbas, 1 specimen, Hamidan, Karak, 24. VI. 2020, leg. S. A. Khan, det. S. Khaliq; 3 specimens, Mansehra, 3. VIII. 2006, leg. & det. M. Abbas; Sindh: 1 specimen, Sadiqabad, Hyderabad, 2. XII. 1954, leg. Qutbuddin, det. S. Khaliq.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC0EF10FF76FC86FEEF5B8E.taxon	distribution	Distribution. Balochistan: Quetta; Khyber Pakhtunkhwa: Buner, Dir, Mansehra, Peshawar; Punjab: Chakwal (Kallar Kahar); Sindh: Tharparkar (Chilhar) (Hashmi & Tashfeen, 1992; Siddiqui et al., 2014; Abbas et al., 2015).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC0EF11FF77F9DFFE945D61.taxon	materials_examined	Material examined. PAKISTAN, Islamabad Capital: 1 specimen, 18. VII. 2007, leg. M. Fida, det. S. Khaliq, 1 specimen, Islamabad, 29. VI. 17, leg. M. Shan, det. S. Khaliq, 1 specimen, 30. IX. 1981, leg. M. Fida, det. S. Khaliq, 1 specimen, 29. III. 2005, leg. K. Hussain, det. M. Abbas; Khyber Pakhtunkhwa: 5 specimens, Swat, 29. VII. 2005, leg. & det. M. Abbas, 1 specimen, Dir, 4. VIII. 2006, leg. & det. M. Abbas, 7 specimens, Swabi, 9. IX. 2007, leg. & det. M. Abbas, 7 specimens, Mardan, 12. VI. 2010, leg. & det. M. Abbas, 9 specimens, Chitral, 12. VIII. 2012, leg. & det. M. Abbas, 9 specimens, Abbottabad, 13. VII. 2013, leg. & det. M. Abbas, 20 specimens, Buner, 1. VII-VIII. 2000, leg. M. Bashir, det. S. Khaliq, 1 specimen, Kopar, Malakand, 16. VII. 2020, leg. S. Khan, det. S. Khaliq, 1 specimen, Kopar, Malakand, 18. VII. 2020, leg. S. Khan, det. S. Khaliq, 1 specimen, Saifi Hilly, Mohmand, 29. VI. 2020, leg. M. Nawaz, det. S. Khaliq, 1 specimen, Batal, 2. VII-VIII. 2000, leg. M. Bashir, det. S. Khaliq; Sindh: 1 specimen, Lawrence Road, Karachi, 19. III. 64, leg. M. Ayyub, det. S. Khaliq, 1 specimen, Shah Dadu, 19. II. 1953, leg. M. Khan, det. S. Khaliq; Punjab: 1 specimen, Birgran, Murree, 26. IX. 1938, leg. I. S. Parathi, det. S. Khaliq 1 specimen, Rawat, Rawalpindi, 26. VII. 2005, leg. & det. M. Abbas; INDIA, Assam. 1 specimen, Dalat, South Assam, 1. III-V. 1918, leg. C. Boden Kloss, det. Anonymous.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC0EF11FF77F9DFFE945D61.taxon	distribution	Distribution. Azad Kashmir: Chikar (Muzaffarabad), Palandri, Rawalakot; Khyber Pakhtunkhwa: Dir, Mardan, Swabi, Swat (Kanju), Waziristan; Punjab: Attock (Fateh Jang), Chakwal (Kallar Kahar), Layyah (Fateh Pur); Sindh: Hyderabad (Tandojam), Tharparkar (Chelhar) (Hashmi & Tashfeen, 1992; Siddiqui et al., 2014).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC1EF11FF86FD99FC8A5B6A.taxon	materials_examined	Material examined. PAKISTAN, Islamabad Capital: 2 specimens, 22. V. 2018, leg. M. Khurram, det. S. Khaliq, 1 specimen, 12. VII. 2005, leg. & det. M. Abbas; Khyber Pakhtunkhwa: 3 specimens, Buner, 1. VII-VIII. 2000, leg. M. Bashir, det. S. Khaliq, 17 specimens, Buner, 2. VII-VIII. 2000, leg. M. Bashir, det. S. Khaliq, 2 specimens, Badraga, Malakand, 23. VI. 2020, leg. & det. S. Khan, 1 specimen, Mansehra, 1. VII-VIII. 2000, leg. M. Bashir, det. S. Khaliq, 1 specimen, Kopar, Malakand, 18. VII. 2020, leg. S. Khan, det. M. Abbas; Punjab: 1 specimen, Bari, Chakwal, 14. IV. 2003, leg. K. Mahmood, det. S. Khaliq, 1 specimen, Bari, Chakwal, 19. IX. 2003, leg. K. Mahmood, det. S. Khaliq, 1 specimen, Khabeki Lake, Khushab, 3 - 7. VIII. 2007, leg. & det. M. Abbas, 1 specimen, Rawalpindi, 29. VIII. 82, leg. Anonymous, det. S. Khaliq, 1 specimen, Ratta road, Rawalpindi, 19. X. 51, leg. M. Ali, det. S. Khaliq; BANGLADESH, 2 specimens, Chittagong, 11. I. 1959, leg. M. Ayyub, det. S. Khaliq.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC1EF11FF86FD99FC8A5B6A.taxon	distribution	Distribution. Locality unknown (Ghosh et al., 2021).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC1EF11FF86FBADFCC8591D.taxon	materials_examined	Material examined. PAKISTAN, Islamabad Capital: 5 specimens, 2. VII. 2005, leg. & det. M. Abbas, 1 specimen, 20. X. 1980, leg. Anonymous, det. S. Khaliq; Khyber Pakhtunkhwa: 7 specimens, Peshawar, 13. VII. 2007, leg. & det. M. Abbas, 5 specimens, Abbottabad, 18. VII. 2008, leg. & det. M. Abbas, 7 specimens, Kohat, 28. VII. 2012, leg. & det. M. Abbas, 1 specimen, Kopar, Malakand, 18. VII. 2020, leg. S. Khan, det. S. Khaliq; Punjab: 1 specimen, Khabeki lake, Khushab, 3 - 7. VIII. 2007, leg. & det. M. Abbas; INDIA, 1 specimen, Madhya Pradesh: Harrah land, Rewa, 9. III. 1927, leg. H. S. Pruthi, det. Anonymous, 1 specimen, South India, Floor of Paini Hills, 20 - 23. IX. 1922, leg. S. Ketap, det. Anonymous.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC1EF11FF86FBADFCC8591D.taxon	distribution	Distribution. Azad Kashmir: Palandri; Khyber Pakhtunkhwa: Dera Ismail Khan; Punjab: Chakwal (Kallar Kahar), Sialkot; Sindh: Karachi, Hyderabad (Tandojam), Tharparkar (Chelhar) (Hashmi & Tashfeen, 1992; Hussain et al., 2020; Siddiqui et al., 2014).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC1EF11FF86F964FF205740.taxon	materials_examined	Material examined. PAKISTAN, Islamabad Capital. 3 specimens, Quaid-e-Azam University, 29. VII. 2005, leg. & det. M. Abbas; Khyber Pakhtunkhwa: Abbottabad, 23. VII. 2010, 4 specimens, Chitral, 12. VII. 2007, 5 specimens, Nowshera, 14. VIII. 2013, 4 specimens, Mansehra, 13. VIII. 2006, 6 specimens, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC1EF11FF86F964FF205740.taxon	distribution	Distribution. Azad Kashmir: Pallandri; Khyber Pakhtunkhwa: Swat (Kanju); Punjab: Bhakkar, Chakwal (Kallar Kahar), Dera Ghazi Khan, Faisalabad, Rahim Yar Khan, Toba Tek Singh (Kamalia); Sindh: Malir, Mirpur Khas, Naukot, Hyderabad (Tandojam), Tharparkar (Chelhar) (Arrow, 1931; Siddiqui et al., 2014).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC7EF17FF86FED1FDE85DD7.taxon	materials_examined	Material examined. PAKISTAN, Khyber Pakhtunkhwa: 1 specimen, Chitral, 10. VII. 2011, 1 specimen, Kurram Agency, 09. VIII. 2013, leg. & det. M. Abbas, 4 specimen, Buner, 1. VII-VIII. 2000, leg. M. Bashir, det. S. Khaliq; Punjab: 7 specimens, Murree, 10. V. 2009, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC7EF17FF86FED1FDE85DD7.taxon	discussion	Remarks. This species is found in the Himalayan dry temperate forests, inner valleys in the mountainous environments in sandy to heavy loamy soils and mostly in grasslands and pastures in cooler regime and is particularly good tolerant to frost.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC7EF17FF86FED1FDE85DD7.taxon	distribution	Distribution. Locality unknown (Schoolmeesters, 2024), Khyber Pakhtunkhwa: Buner, Chitral, Kurram Agency; Punjab: Murree (current study).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC7EF17FF86FD63FC005CB8.taxon	materials_examined	Material examined. PAKISTAN, Khyber Pakhtunkhwa: 4 specimens, Chitral, 13. VIII. 2013, 7 specimens, Swat, 12. VII. 2007, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC7EF17FF86FD63FC005CB8.taxon	discussion	Remarks. This species represents the new country record for Pakistan and extends the distribution across India to Pakistan. It is distributed in the Himalayan moist temperate forests in the mountainous environments and is mostly found in the vicinities of human habitations in the rocky habitats.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC7EF17FF86FD63FC005CB8.taxon	distribution	Distribution. Azad Kashmir: Rawalakot (Siddiqui et al., 2014).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC7EF17FF86FBA2FB305804.taxon	materials_examined	Material examined. PAKISTAN, Islamabad Capital: 2 specimens, Fatima Jinnah Park, 24. V. 1981, leg. A. Ghaffor, det. M. Abbas, 2 specimens, Margalla Hills, 29. VII. 2005, leg. & det. M. Abbas; Khyber Pakhtunkhwa: 1 specimen, Bunair, 1. VII-VIII. 2000, leg. M. Bashir, det. M. Abbas, 3 specimens, Swat, 10. VII. 2008, leg. & det. M. Abbas, 4 specimens, Swabi, 26. VII. 2013, leg. & det. M. Abbas, 2 specimens, Saifi hilly Mohmand, 29. VI. 2020, leg. M. Nawaz, det. S. Khaliq, 29 specimens, Mansehra, 1. VII-VIII. 2000, leg. M. Bashir, det. M. Abbas, 7 specimens, Haripur, 1. VII-VIII. 2000, leg. M. Bashir, det. M. Abbas, 1 specimen, Malakand Kot, 27. IX. 2020, leg. S. Khan, det. S. Khaliq; Punjab: 5 specimens, Attock Akhori, 28. V. 2014, leg. Mishkat, det. S. Khaliq, 1 specimen, Ghazi Brotha, Attock, 29. V. 2014, leg. Mishkat, det. M. Abbas, 1 specimen, Bahawalpur, 29. VIII. 2009, leg. & det. Mishkat, 13 specimens, Lal Suhanra National Park, Bahawalpur, 10. IV. 2015, leg. F. Muhammad & K. Hussain, det. M. Abbas, 4 specimens, Lal Suhanra National Park, Bahawalpur, 11. IV. 2015, leg. F. Muhammad & K. Hussain, det. M. Abbas, 4 specimens, Lal Suhanra National Park, Bahawalpur, 12. IV. 2015, leg. F. Muhammad & K. Hussain, det. M. Abbas, 2 specimens, Khabikki Lake, Khushab, 3 - 7. VIII. 2007, leg. & det. M. Abbas, 8 specimens; Sindh: Taj Muhammad Rukh, Ghotki, 16. V. 2009, leg. Mishkat, det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC7EF17FF86FBA2FB305804.taxon	distribution	Distribution. Azad Kashmir: Pallandri, Muzaffarabad (Pir Chanasi); Khyber Pakhtunkhwa: Abbottabad, Buner, Mansehra, Peshawar, Swabi, Swat (Kanju); Punjab: Chakwal (Kallar Kahar), Islamabad Capital, Jhelum (Pind Dadan Khan), Lahore (Changa Manga Forest), Toba Tek Singh (Kamalia); Sindh: Karachi (Gulzar-e-Hijri), Malir, Mithi (Naukot), Thar, Tharparkar (Chelhar, Islamkot) (Siddiqui & Kamaluddin, 2011; Siddiqui et al., 2014; Abbas et al., 2015; Ali et al., 2015; Génier & Moretto, 2017).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC7EF14FF86F85BFA305E66.taxon	materials_examined	Material examined. PAKISTAN, Khyber Pakhtunkhwa: 3 specimens, Kurram Agency, 8. IX. 2007, 3 specimens, Malakand, 20. VIII. 2012, 1 specimen, Buner, 2. IX. 2013, leg. & det. M. Abbas; Punjab: 1 specimen, Rawat, 26. VII. 2005, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC7EF14FF86F85BFA305E66.taxon	distribution	Distribution. Khyber Pakhtunkhwa: Buner, Kurram Agency, Malakand; Punjab: Rawat (New country record).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC4EF14FF77F9CDFB0D587F.taxon	materials_examined	Material examined. PAKISTAN, Khyber Pakhtunkhwa: 5 specimens, Buner, 9. IX. 2007, 5 specimens, Mansehra, 12. VII. 2008, 5 specimens, Besham, 25. VIII. 2012, 6 specimens, Malakand, 6. VIII. 2013, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC4EF14FF77F9CDFB0D587F.taxon	distribution	Distribution. Azad Kashmir: Rawalakot; Khyber Pakhtunkhwa: Dera Ismail Khan; Sindh: Tharparkar (Arrow, 1931; Hashmi & Tashfeen, 1992; Siddiqui et al., 2014; Abbas et al., 2023).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC4EF14FF77F889FD70577E.taxon	materials_examined	Material examined. PAKISTAN, Islamabad Capital: 1 specimen 18. V. 2005, 7 specimens, 12. VI. 2009; Khyber Pakhtunkhwa: 5 specimens, Buner, 18. VII. 2006, 4 specimens, Abbottabad, 2. VII. 2013, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC4EF14FF77F889FD70577E.taxon	distribution	Distribution. Khyber Pakhtunkhwa: Abbottabad, Buner, Peshawar, Swat (Kanju); Punjab: Islamabad Capital (Siddiqui et al., 2014; Abbas et al., 2023).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC4EF14FF77FB40FE495AC5.taxon	materials_examined	Material examined. PAKISTAN, Khyber Pakhtunkhwa: 3 specimens, Chitral, 14. IX. 2007, 5 specimens, Kurram Agency, 2. VIII. 2010, 9 specimens, Mansehra, 13. IX. 2012, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC4EF14FF77FB40FE495AC5.taxon	discussion	Remarks. This species is usually found at higher altitudes in alpine forests as well as in the alpine pastures.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC4EF14FF77FB40FE495AC5.taxon	distribution	Distribution. Locality unknown (Schoolmeesters, 2024), Khyber Pakhtunkhwa: Chitral, Kurram Agency, Mansehra (current study).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC4EF14FF77FD90FBD75C35.taxon	materials_examined	Material examined. PAKISTAN, Islamabad Capital: 7 specimens, 3. IX. 2012; Khyber Pakhtunkhwa: 5 specimens, Swat, 14. IX. 2007; Punjab: 8 specimens, Rawat, Rawalpindi, 26. VII. 2005, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC4EF14FF77FD90FBD75C35.taxon	discussion	Remarks. This species is common in the foothill habitats.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC4EF14FF77FD90FBD75C35.taxon	distribution	Distribution. Azad Kashmir: Pallandri, Muzaffarabad (Pir Chanasi); Khyber Pakhtunkhwa: Dera Ismail Khan; Punjab: Bhakkar, Chakwal; Sindh: Tharparkar (Siddiqui et al., 2014).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC4EF14FF77FC47FF645B34.taxon	materials_examined	Material examined. PAKISTAN, Khyber Pakhtunkhwa: 5 specimens, Besham, 14. IX. 2007, 7 specimens, Upper Swat, 13. IX. 2012; Punjab: 1 specimen, Rawat, 26. VII. 2005, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC4EF14FF77FC47FF645B34.taxon	distribution	Distribution. Sindh: Karachi (Gulzar-e-Hijri), Malir, Tharparkar (Chilhar), Mithi (Naukot) (Siddiqui et al., 2014).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC4EF14FF76FE92FC825D64.taxon	materials_examined	Material examined. PAKISTAN, Khyber Pakhtunkhwa: 2 specimens, Kurram Agency, 12. VIII. 2006, 3 specimens, Chitral, 8. IX. 2008, 5 specimens, Shandur, 20. VIII. 2012, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC4EF14FF76FE92FC825D64.taxon	distribution	Distribution. Locality unknown (Gorodinski, 2023).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC5EF15FF86FA6AFBFC59E2.taxon	materials_examined	Materials examined. PAKISTAN, Islamabad Capital: 1 specimen, 20. V. 2005; Khyber Pakhtunkhwa: 2 specimens, Malakand, 26. II. 2005, 5 specimens, Swat, 20. VII. 2010, 7 specimens, Mardan, 27. I. 2012; Punjab: 9 specimens, Murree, 13. VIII. 2013, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC5EF15FF86FA6AFBFC59E2.taxon	distribution	Distribution. Khyber Pakhtunkhwa: Mansehra, Mardan, Malakand, Swat; Punjab: Islamabad Capital, Rawalpindi (Murree); Sindh: Karachi (Siddiqui et al., 2014; Abbas et al., 2023).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC5EF15FF86FF2FFF205D6F.taxon	materials_examined	Material examined. PAKISTAN, Khyber Pakhtunkhwa: 7 specimens, Mansehra, 1. VII. 2006, 5 specimens, Peshawar, 11. VI. 2010, 9 specimens, Chitral, 20. VII. 2012, 9 specimens, Nowshera, 18. VIII. 2013; Punjab: 1 specimen, Rawa, 26. VII. 2005, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC5EF15FF86FF2FFF205D6F.taxon	distribution	Distribution. Azad Kashmir: Poonch (Rawalakot); Khyber Pakhtunkhwa: Abbottabad, Chitral, Kohat, Nowshera, Mansehra, Peshawar; Punjab: Toba Tek Singh (Kamalia) (Siddiqui et al., 2014; Abbas et al., 2023).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC5EF15FF86FDA1FDE45C8D.taxon	materials_examined	Material examined. PAKISTAN, Islamabad Capital: 3 specimens, 12. VII. 2005; Khyber Pakhtunkhwa: 7 specimens, Abbottabad, 11. V. 2005, 5 specimens, Chitral, 21. VII. 2007, 9 specimens, Peshawar, 23. VIII. 2012, 4 specimens, Kohat, 3. VII. 2013, 5 specimens, Dir, 13. IV. 2014, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC5EF15FF86FDA1FDE45C8D.taxon	distribution	Distribution. Azad Kashmir: Poonch (Rawalakot), Sudhanoti (Palandri); Gilgit-Baltistan: Gilgit (Naltar); Khyber Pakhtunkhwa: Abbottabad, Besham, Chitral, Dir, Kohat, Nowshera, Peshawar, Swat (Marghazar); Punjab: Islamabad Capital, Rawalpindi (Murree), Toba Tek Singh (Kamalia); Sindh: Malir (Siddiqui et al., 2014; Abbas et al., 2023).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC5EF15FF86FBC7FA175A00.taxon	materials_examined	Material examined. PAKISTAN, Islamabad Capital: 1 specimen, Quaid-e-Azam University, 29. VII. 2005, 6 specimens, 2. VII. 2012, leg. & det. M. Abbas; Khyber Pakhtunkhwa: 12 specimena, Mansehra, 1. VII-VIII. 2000, leg. M. Bashir, det. S. Khaliq, 1 specimen, Batal, 1. VII-VIII. 2000, leg. M. Bashir, det. M. Abbas, 5 specimens, Nowshera, 12. V. 2010, 9 specimens, Peshawar, 13. VIII. 2013, leg. & det. M. Abbas; Punjab: 1 specimen, Rawat, Rawalpindi, 26. VII. 2005, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC5EF15FF86FBC7FA175A00.taxon	distribution	Distribution. Khyber Pakhtunkhwa: Abbottabad, Kohat, Mansehra, Mardan, Nowshera, Peshawar, Swat (Kanju); Punjab: Islamabad Capital; Sindh: Tharparkar (Siddiqui et al., 2014; Abbas et al., 2023).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC5EF1AFF86F880FCC15A89.taxon	materials_examined	Material examined. PAKISTAN, Islamabad Capital: 1 specimen, Quaid-e-Azam University, 29. VII. 2005, leg. M. Abbas, det. S. Khaliq; Khyber Pakhtunkhwa: 5 specimens, Heroshah, Malakand, 13. VII. 2020, leg. S. Khan, det. S. Khaliq, 4 specimens, Heroshah, Malakand, 28. VII. 2020, leg. S. Khan, det. S. Khaliq, 1 specimen, Dargai, Malakand, 19. VII. 2020, leg. S. Khan, det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFC5EF1AFF86F880FCC15A89.taxon	distribution	Distribution. Locality unknown (Löbl & Smetana, 2006).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFCAEF1AFF77F9A7FDCB5830.taxon	materials_examined	Material examined. PAKISTAN, Khyber Pakhtunkhwa: 3 specimens, Kurram Agency, 14. VI. 2007, 3 specimens, Dir, Khyber, 19. VI. 2008, 3 specimens, Khyber Agency, 12. VII. 2012, 6 specimens, Mohmand Agency, 23. IV. 2013, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFCAEF1AFF77F9A7FDCB5830.taxon	distribution	Distribution. Azad Kashmir: Pallandri; Khyber Pakhtunkhwa: Bajor Agency (Arrow, 1931; Lobel & Smetana, 2006; Siddiqui et al., 2014).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFCAEF1BFF77F84AFB1A5E7E.taxon	materials_examined	Material examined. PAKISTAN, Khyber Pakhtunkhwa: 4 specimens, Malakand, 23. VI. 2020, leg. S. Khan, det. S. Khaliq.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFCAEF1BFF77F84AFB1A5E7E.taxon	distribution	Distribution. Azad Kashmir: Muzaffarabad; Khyber Pakhtunkhwa: Dera Ismail Khan, Mansehra; Sindh: Tharparkar (Arrow, 1931; Hashmi & Tashfeen, 1992; Siddiqui et al., 2014).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFCBEF1BFF86FE77FC6B5D21.taxon	materials_examined	Material examined. PAKISTAN, Khyber Pakhtunkhwa: 1 specimen, Kohat, 12. VII. 2007, 3 specimens, Kurram Agency, 16. V. 2010, leg. & det. M. Abbas.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFCBEF1BFF86FE77FC6B5D21.taxon	distribution	Distribution. Khyber Pakhtunkhwa: Bannu (Zídek & Pokorný, 2008).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFCBEF1BFF86FD39FE145B50.taxon	materials_examined	Material examined. PAKISTAN, Khyber Pakhtunkhwa: 2 specimens, Malakand, 28. VII. 2020, leg. S. Khan, det. S. Khaliq.	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
3F45F66CFFCBEF1BFF86FD39FE145B50.taxon	distribution	Distribution. Azad Kashmir: Muzaffarabad (Punjkot); Khyber Pakhtunkhwa: Dera Ismail Khan (Siddiqui et al., 2014).	en	Abbas, Muhammad, Khaliq, Sobia, Masroor, Rafaqat, Hassan, Muhammad Asghar (2024): A checklist of the dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) deposited at Pakistan Museum of Natural History. Journal of Insect Biodiversity and Systematics 10 (2): 415-430, DOI: 10.61186/jibs.10.2.415, URL: https://doi.org/10.61186/jibs.10.2.415
