taxonID	type	description	language	source
3F1087F9FF8FFFABFF0F311AD40EFD85.taxon	description	(Figs 1 – 11) Synonyms: Taenia festiva Rudolphi, 1817; Moniezia festiva: Blanchard, 1891; Hepatotaenia festiva: Nybelin, 1917; Cittotaenia festiva: Theiler, 1924.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF8FFFABFF0F311AD40EFD85.taxon	materials_examined	Type host: Macropus giganteus Shaw, 1790 (Marsupialia: Macropodidae). Other hosts: Macropus dorsalis (Gray), M. rufus (Desmarest), M. robustus Gould, M. parryi Bennett, M. antilopinus (Gould), Onychogalea unguifera (Gould). Site in host: bile ducts. Types: MNB 1888. Material examined: From Macropus giganteus (eastern grey kangaroo): types; Queensland: 1 specimen, Townsville (AHC 20875 (S )); 2 specimens, Hervey’s Range, Townsville (AHC 29409 (S), 7665 (W); 3 specimens, Woodstock (AHC 29143 (S), 7361 (W); 4 specimens, Harvest Home Stn via Charters Towers (AHC 22287 - 9 (S )); 1 specimen, Pallamana Stn via Charters Towers (AHC 22291 (S )); 2 specimens, Clermont (AHC 29390 (S )); 2 specimens, Emu Park (AHC 21499 (S )); 1 specimen, Theodore (AHC 29397 (S )); 1 specimen, Miles (AHC 29386 (S )); 2 specimens, Chinchilla (AHC 29410 (S), 9780 (W )); 1 specimen, 40 km N of Charleville (AHC 29399 (S )); 1 specimen, 66 km E of Moonie (AHC 29403 (S )); 1 specimen, 30 km E of Bollon (AHC 29405 (S )); New South Wales: 1 specimen, 50 km N of Moree (AHC 29385 (S )); 2 specimens, Weilmoringle (AHC 29401 (S), 34854 (W )); 1 specimen, 85 km N of Bourke (AHC 29404 (S )); 1 specimen, 50 km N of Bourke (AHC 29395 (S )); 2 specimens, Warraweena Stn via Bourke (AHC 28944, 29373 (S )); 1 specimen, 20 km NW of Nyngen (AHC 29407 (S), 27265 (W )); 1 specimen, Ebor (AHC 29384 (S), 10417 (W )); 1 specimen, Armidale (AHC 29411 (S), 9887 (W )); 1 specimen, Coonabarabran (AHC 29375 (S )); 1 specimen, Nowra (AHC 29379 (S), 34850 (W )); 1 specimen, Bondo State Forest (AHC 22092 (S )); Victoria: 1 specimen, Picola North (AHC 20994 (S )); 1 specimen, Barmah (AHC 29393 (S )); 1 specimen, Nariel (AHC 29387 (S )); 1 specimen, Dartmouth (AHC 20945 (S )); 3 specimens, Mitta Mitta (AHC 29398 (S), 12057 (W )); 2 specimens, Boho South (AHC 29377 (S), 34853 (W )); 1 specimen, Dadswell’s Bridge (AHC 29378) (S )); 1 specimen, Pomonal (AHC 29380 (S )); 4 specimens, Zumsteins (AHC 20906 (S )); 3 specimens. Mirranatwa (AHC 29389 (S), 10888 (W )); 3 specimens, Dunkeld (AHC 29388 (S), 10886 (W )); 1 specimen, Sutton Grange (AHC 29402 (S), 19775 (W )); 4 specimens, Fraser National Park (AHC 20855, 21463 (S )); 8 specimens, Eildon (AHC 29391 - 2 (S), 9839, 19065 (W )); 6 specimens, Yan Yean (AHC 20913 (S )); 3 specimens, Healesville (AHC 29383 (S), 34852 (W )); 1 specimen, Chum Creek (AHC 29406 (S), 30374 (S )); 1 specimen, Bacchus Marsh (AHC 20795 (S )); 1 specimen, Brisbane Ranges (AHC 20856 (S )); 1 specimen, You Yang Range (AHC 29400 (S), 10166 (W )); 1 specimen, Staughton Vale (AHC 29412 (S), 10852 (W )); 1 specimen, Avalon (AHC 29382 (S )); 1 specimen, Lara (AHC 29381 (S), 34851 (W )); 2 specimens, Marlo (AHC 20852, 21446 (S )); 1 specimen, Cape Conran (AHC 29394 (S), 10428 (W )); 1 specimen, Gippsland Lakes (AHC 29396 (S )); 2 specimens, Bellbird (AHC 29408 (S), 9711 (W )); 1 specimen, Mt Hickeyard (AHC 20810 (S )); Tasmania: 1 specimen, Ross (AHC 29376 (S), 11028 (W )). From Macropus dorsalis (black-stripe wallaby): Queensland: 6 specimens, Warrawee Stn via Charters Towers (SAM 17953, 22285 - 6 (S )); 5 specimens, Harvest Home Stn via Charters Towers (SAM 22279 - 82 (S )). From Macropus parryi (whiptail wallaby): Queensland: fragments, Mt Fox (SAM 29414 (S )); 1 specimen, Mt Surprise (SAM 7172 (W )); 2 specimens, Townsville (SAM 29416 (S), 7186 (W )); 1 specimen, Reid River (SAM 29417 (S), 7164 (W )); 2 specimens, Mt Elliott (SAM 29418 (S), 7159 (W )); 4 specimens, Inkerman (SAM 29419 (S), 7178 (W )); 3 specimens, Mingela (SAM 29421 (S), 7215 (W )); 2 specimens, Yeppoon (SAM 28957 (S )); fragments, Keppel Sands (SAM 29415); 4 specimens, Banana (SAM 29420 (S), 19782 (W )); 2 specimens, Thangool (SAM 29422 (S )); fragments, Eidsvold (SAM 20069 (S )); New South Wales: 1 specimen, Rivertree (SAM 20761 (S )). From Macropus rufus (red kangaroo): Queensland: 1 specimen, Devoncourt Stn via Cloncurry (SAM 29433 (S), 34832 (W )); 1 specimen, Kiora Stn via Pentland (SAM 29440 (S), 34831 (W )); 1 specimen, 50 km S of Charleville (SAM 29437 (S )); New South Wales: 1 specimen, 10 km SE of Bourke (SAM 29423 (S )); 1 specimen, 40 km SE of Bourke (SAM 29438 (S )); 1 specimen, 50 km SE of Bourke (SAM 29436 (S )); 1 specimen, Byrock (SAM 29426 (S )); 1 specimen, Sturt National Park (SAM 29439 (S), 34834 (W); 1 specimen, Narriara Stn via Broken Hill (SAM 29434 (S), 34833 (W); 1 specimen, Thackaringa Stn via Broken Hill (SAM 29428 (S), 34836 (W )); 6 specimens, Menindee (SAM 20853, 20868 (S )); 8 specimens, Kinchega National Park (SAM 20854 (S )); 6 specimens, Warrananga Stn via Wentworth SAM 20759, 20763 (S )); Victoria: 1 specimen, Lake Cullulleraine (SAM 29432 (S), 19912 (W )); South Australia: 2 specimens, Wilpoorinna Stn via Marree (SAM 29429 (S), 8138, 34837 (W )); 3 specimens, The Twins Stn via Kingoonya (SAM 29430 (S), 8848 (W )); 1 specimen, Wirrealpa Stn via Blinman (SAM 29431 (S), 8149 (W )); 1 specimen, Leigh Creek (SAM 29441 (S), 8620 (W )); 2 specimens, Myrtle Springs Stn via Leigh Creek (SAM 29435 (S), 8132 (W )); 1 specimen, Coondambo Stn via Kingoonya (SAM 29427 (S), SAM 8852 (W )); 2 specimens, Wallerberdina Stn via Pt Augusta (SAM 29425 (S), SAM 19798 (W )); 1 specimen, Melrose (SAM 21373 (S )); Western Australia: 4 specimens, Yalgoo (SAM 29424 (S), 34838 (W), BMNH 2008.6. 3.3 (S )). From Macropus robustus (wallaroo): Northern Territory: 2 specimens, 40 km N of Katherine (SAM 29444 (S )); Queensland: 1 specimen, 24 km E of Georgetown (SAM 29442 (S )); 1 specimen, Bluewater Springs (SAM 28949 (S )); 1 specimen, 83 km N of Charters Towers (SAM 28960 (S )); 1 specimen, 30 km N of Charters Towers (SAM 28955 (S )); 1 specimen, Fletcher View Stn via Charters Towers (SAM 29452 (S), 7412 (W )); 1 specimen, Mingela (SAM 29449 (S), 34842 (W )); 1 specimen, Maida Vale Stn via Mingela (SAM 29453 (S), 7328 (W )); 1 specimen, Koonkool Stn via Hughenden (SAM 29450 (S), 34839 (W )); 1 specimen, Woodbine Stn via Prairie (SAM 29451 (S), 34843 (W )); 6 specimens, 70 km W of Cloncurry (SAM 29455 (S), 34847 (W )); 3 specimens, 76 km W of Cloncurry (SAM 29456 (S), 34836 (W )); 4 specimens, Devoncourt Stn via Cloncurry (SAM 29458 (S), 34844 - 5 (W )); 9 specimens, Oban Stn via Mt Isa (SAM 29457 (S), 11665 (W )); 9 specimens, Pallamana Stn via Charters Towers (SAM 22307 (S )); 12 specimens, Warrawee Stn via Charters Towers (SAM 19755 - 6, 22292 - 22301 (S )); 4 specimens, Harvest Home Stn via Charters Towers (SAM 22302 - 22305 (S )); 3 specimens, Coolagh Stn via Blackall (SAM 29447 (S), 34840 - 1 (W )); 1 specimen, 7 km E of Barcaldine (SAM 29454 (S), 34849 (W )); 1 specimen, 17 km E of Jericho (SAM 29448 (S), 34848 (W )); 1 specimen, 20 km N of Augathella (SAM 29443 (S )); New South Wales: 2 specimens, 30 km E of Armidale (SAM 20793 (S )); 1 specimen, Kingstown (SAM 20791 (S )); South Australia: 2 specimens, Mt Lyndhurst (SAM 29445 (S )), 8617 (W )); 2 specimens, Hawker (SAM 21392 (S )); 4 specimens, Nonning Stn via Pt Augusta (SAM 22306 (S )); Western Australia: 1 specimen, Wialki (SAM 20068 (S )); 1 specimen, Barrow Island (SAM 29446 (S), 22958 (W )); 2 specimens, Beverley Springs Stn via Derby (SAM 21491 - 2 (S )); 3 specimens, Yalgoo (SAM 28959 (S )); 4 specimens, Northampton (SAM 28956 (S )). From Macropus antilopinus (antilopine wallaroo): Northern Territory: 2 specimens, 50 km SW of Katherine (SAM 28958 (S )); 2 specimens, 87 km SW of Katherine (SAM 28954 (S )); 4 specimens, 8 km N of Mataranka (SAM 29460 (S )); Queensland: 1 specimen, Mt Surprise (SAM 29461 (S), 7172 (W )); 8 specimens, Burlington Stn via Mt Surprise (SAM 29462 (S), 6644 (W )); Western Australia: 4 specimens, Kimberley region (SAM 21488, 21490, 21454 (S), 7663 (W )). From Onychogalea unguifera (northern nail-tail wallaby): Western Australia: 2 specimens, Kimberley region (SAM 21440 (S )).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF8FFFABFF0F311AD40EFD85.taxon	description	Description: Based on specimens from M. giganteus. Slender, elongate cestodes, 154 – 251 (214, n = 5) mm long, maximum width up to 8 mm with 437 – 578 (476, n = 5) segments in gravid strobilae. Scolex 0.683 – 1.11 (0.913) in diameter, prominently lobed; suckers circular, 0.244 – 0.325 (0.294) in diameter. Neck prominent. First mature segment 170 – 205 th (190, n = 5). Mature segments craspedote, 1.71 – 2.80 (2.23) wide, 0.163 – 0.569 (0.330) long. Genital atrium small, posterior to middle of lateral segment margin; cirrus sac small, elongate, extending to or barely beyond osmoregulatory canals, 0.163 – 0.325 (0.266) long, 0.052 – 0.065 (0.060) wide; cirrus unarmed; internal seminal vesicle occupies approx. 2 / 3 – 3 / 4 of volume of cirrus sac, 0.078 – 0.202 (0.134) long, 0.026 – 0.052 (0.034) wide; elongate, spiral, external seminal vesicle, covered with layer of glandular cells, 0.065 – 0.195 (0.131) long, 0.033 – 0.117 (0.070) wide; vas deferens coils medially, dorsal to uterus; testes arranged in anterior half of segment between osmoregulatory canals; no prominent space between canals and testis fields; testes arranged in 2 dorso-ventral layers, almost invariably in 2 separate groups. Testes 0.026 – 0.072 (0.049) in diameter; testis number 71 – 93 (82) in each segment. Vagina tubiform, lacking glandular investment, opens to genital atrium posterior to cirrus sac, leads to ovoid to subspherical seminal receptacle 0.091 – 0.247 (0.166) x 0.085 – 0.182 (0.127). Ovary small, flabelliform, medial to seminal receptacle 0.052 – 0.098 (0.072) x 0.033 – 0.078 (0.055); vitellarium reniform, posterior to ovary, 0.072 – 0.143 (0.102) x 0.052 – 0.117 (0.067); Mehlis’ gland anterior and medial to vitellarium. Uteri paired in each segment, transverse, tubular; early uterus extends from beyond level of ovary to near proximal pole of cirrus sac; developing uterus extends medially and laterally, lateral extensions crossing osmoregulatory canals dorsally; medially uteri of each segment do not meet in mid-line. Gravid segments 0.244 – 0.683 (0.406) long, 2.93 – 5.82 (3.88) wide. Egg spherical, shell smooth, 0.045 – 0.050 (0.048) in diameter; pyriform apparatus terminating in numerous reflexed filaments; oncosphere 0.013 – 0.018 (0.015) in diameter. Osmoregulatory canals paired; ventral canal generally wider than dorsal, medial to it; ventral canal 0.020 – 0.085 (0.044) in diameter, dorsal canal 0.010 – 0.033 (0.018) in diameter; transverse canal connects ventral canal at posterior margin of each segment; accessory canals present in some specimens.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF8FFFABFF0F311AD40EFD85.taxon	discussion	Remarks. The redescription of this species is based exclusively upon specimens from Macropus giganteus, identified by Rudolphi (1819) as the host of the type specimens. The type specimens were collected from a grey kangaroo, M. giganteus, which died in the Schönbrunn Zoological Gardens, Vienna, and were collected by Bremser and sent to Rudolphi in Berlin. Putting aside the reliability of the identification of species of Macropus in European zoos during the early nineteenth century, the genetic study of Beveridge et al. (2007), based on the cox 1 gene, indicated that specimens of bile duct cestodes collected from this host species over a broad geographical range in eastern Australia belonged to the same genetic clade and therefore potentially to the same species. A wide range of specimens was also available from this host species covering most of its geographical range (Fig. 12) and all were relatively uniform morphologically. The cestodes were long (> 150 mm), composed of> 400 segments, with the first mature segment being about the 170 – 205 th segment from the scolex. The scolex was usually prominently four-lobed (Fig. 3), though in incompletely relaxed specimens, the lobes of the scolex were shortened (Fig. 2). In the mature segments, the testes were almost invariably separated into two equal groups, with 71 – 93 testes per segment, and the testis fields abutted the longitudinal osmoregulatory canals. The cirrus sac was small and reached the osmoregulatory canals but rarely extended significantly into the medulla (Fig. 8). The external seminal vesicle was spirally coiled and was surrounded by gland cells; when filled, several coils were expanded, giving the appearance of a multichambered external seminal vesicle (Fig. 5). Specimens of P. festiva from M. rufus formed a distinctive clade in the genetic analysis of Beveridge et al. (2007). Morphologically, these specimens (with a single exception dealt with below) were also uniform throughout their geographic range (Fig. 14), and were similar to the specimens from M. giganteus, differing only in the more pronounced extension of the cirrus sac into the medulla in some specimens (Fig. 9). However, there was substantial variation in this feature within specimens from M. rufus and there was no difference in the mean size of the cirrus sac between specimens from M. giganteus and M. rufus (Table 1). Consequently, no significant and consistent morphological differences were observable between specimens from these two host species. Collections from M. rufus were relatively extensive, but were concentrated in the eastern part of the host range (Fig. 14). Additional collecting is required in central and western Australia to ensure that the genetic clade is uniform across its entire range. In the genetic analysis, all specimens from M. parryi also formed a distinctive clade. Morphologically, these specimens, which were obtained from across a wide geographical range (Fig. 13), were relatively uniform morphologically, but differed in one feature. In almost all specimens, the external seminal vesicle was fusiform (Fig. 6) rather than spirally coiled. The testes were in separate groups, a feature of all specimens in this group, except for two specimens, one of which was immature (SAM 29414) and the other was severely contracted (SAM 20069). In all well-preserved specimens examined, there was a distinctive space between the osmoregulatory canals and the lateral extent of the testis fields (Fig. 10), thereby distinguishing these specimens from those found in M. giganteus and M. rufus. Specimens from M. robustus and M. antilopinus occurred in four distinct clades in the genetic analysis of Beveridge et al. (2007), and these differences were reflected in part in their morphological features. Specimens collected from M. robustus in South Australia and Western Australia (type 1 in Fig. 15) were morphologically identical with those from M. giganteus and M. rufus in lacking a space between the testis field and the osmoregulatory canals. Specimens from M. robustus from north western Queensland (Mt Isa, Cloncurry) (type 2 in Fig. 15) were distinguishable in having a distinct space between the osmoregulatory canals and the testis fields as in the specimens from M. parryi (Fig. 10), but were distinguishable from specimens in M. parryi in having a spiral external seminal vesicle. This clade was also found in M. dorsalis in north-eastern Queensland and in M. rufus from Cloncurry and Pentland, Queensland, but in each case, this genetic form was identifiable by the space between the testes and the osmoregulatory canals and by the spiral external seminal vesicle. Specimens with similar morphological features were seen in M. robustus from Armidale, NSW, Barrow Island and the Kimberley, WA, but since no material was available from these locations for genetic analysis, the relationships of these cestodes remain to be delineated. The genetic clade (3 in Fig. 15) identified from coastal north-eastern Queensland was characterised morphologically by the lack of a space between the testis fields and the osmoregulatory canals and was therefore not distinguishable from the clade found in south and western Australia (1 in Fig. 15). All material examined from M. antilopinus in the Northern Territory and all but one specimen examined from Queensland was morphologically distinctive in having a distinct space between the osmoregulatory canals and the testis fields and a greater number of testes per segment than in other specimens (Fig. 11) (Table 1). However, the space between testis fields and osmoregulatory canals was somewhat variable and was small in one specimen. This morphological form was also seen in a single M. robustus from the Northern Territory, in an area in which M. antilopinus and M. robustus are sympatric (4 in Fig. 15). No entire specimens of this morphotype were collected and therefore although it appears to represent a distinctive species, it is not described as a new species herein. In the Mt Surprise region of northern Queensland, M. antilopinus and M. robustus are also sympatric (Fig. 15). The fixed material from M. antilopinus from this area is similar to that from the same host species in the Northern Territory. However, the single specimen of P. festiva from M. antilopinus from north Queensland used in the genetic analysis appears to be identical with the morphological form predominant in M. robustus in that region suggesting that both morphotypes may occur in the two host species in this region. More extensive collections are needed to resolve the morphotypes present in wallaroos in this region. Metric data of P. festiva from various host species are presented in Table 1. Whilst there are some differences in measurements, particularly for specimens designated as clade 4, initial inspection of the data suggested that detailed statistical analysis was not warranted as it was unlikely to yield useful taxonomic information. Fragmentary material was available from Onychogalea unguifera from the Kimberley region of Western Australia. While generally referable to P. festiva, additional specimens are required to determine if it represents an additional species.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF84FFADFF0F34BDD344FC9E.taxon	description	(Figs 16 – 23) Synonyms: Hepatotaenia fellicola Nybelin, 1917.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF84FFADFF0F34BDD344FC9E.taxon	biology_ecology	Host: Macropus agilis (Gould) (Marsupialia: Macropodidae). Types: SMNH 93642 - 45, Noonkanbah, Western Australia (18 ° 31 ’ S 124 ° 48 ’ E), coll. E. Mjöberg, 31. xi. 1911. Site in host: bile ducts.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF84FFADFF0F34BDD344FC9E.taxon	materials_examined	Material examined: From M. agilis (agile wallaby): Western Australia: types, 2 specimens, Beverley Springs Stn via Derby (SAM 21497, 21489 (S), 7510, 7661 (W )); Northern Territory: 2 specimens, Darwin (SAM 20794 (S )), 16 specimens, Katherine (SAM 29851 - 3, 29339 - 40, 29354 (S), BMNH 2008.6.3.4 - 5 (S )), 6 specimens, Tipperary Stn (SAM 9952, 29344 (S), 9952 (W )), 8 specimens, Elizabeth Downs Stn (SAM 20841 (S), 10181 (W )); 2 specimens, Mataranka (SAM 29353 (S )); Queensland: 10 specimens, Townsville (SAM 29342 - 3, 29348, 29350 (S), 7563, 7575 (W )); 3 specimens, Kelso (SAM 29352 (S), 34855 (W )); 4 specimens, Oonoonba (SAM 29345 (S), 7601 (W )), 2 specimens, Yabulu (SAM 29349 (S), 7494 (W )), 3 specimens, Inkerman (SAM 29347 (S), 7556 (W )), 1 specimen, Woodstock (SAM 29341), 2 specimens, Wenlock River, Cape York Peninsula (SAM 29346 (S), 7564 (W )); Papua New Guinea: 1 specimen, Bula Plain, Bensbach Province (SAM 29351 (S )).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF84FFADFF0F34BDD344FC9E.taxon	description	Description: Short, robust worms, 20 – 75 (47) long, maximum width 2 – 6 (3.6) with 96 – 337 (167) segments in gravid specimens. Scolex 0.65 – 0.89 (0.77) in diameter, squat, mildly 4 – lobed; suckers circular, 0.23 – 0.48 (0.30) in diameter. Neck extremely short or absent. First mature segment 25 – 45 th (34). Mature segments craspedote, 1.54 – 3.25 (2.31) wide, 0.15 – 0.33 (0.22) long. Genital atrium small, in middle or slightly posterior to middle of lateral segment margin; cirrus sac elongate, extending well beyond osmoregulatory canals, 0.156 – 0.468 (0.200) long, 0.033 – 0.111 (0.060) wide; cirrus unarmed; internal seminal vesicle occupies approx. 2 / 3 – 3 / 4 of volume of cirrus sac, 0.091 – 0.293 (0.200) long, 0.033 – 0.111 (0.060) wide; elongate external seminal vesicle, covered with glandular cells, 0.098 – 0.208 (0.150) long, 0.033 – 0.072 (0.060) wide; vas deferens coils medially, dorsal to uterus; testes arranged in anterior half of segment between osmoregulatory canals; prominent space between canals and testis fields; testes arranged in 2 dorso-ventral layers, in single band or 2 separate groups; segments with single band or 2 groups occur in same specimen. Testes 0.033 – 0.065 (0.040) in diameter; testis number 73 – 95 (88). Vagina tubiform, lacking glandular investment, opens to genital atrium posterior to cirrus sac, leads to ovoid seminal receptacle 0.078 – 0.293 (0.200) x 0.078 – 0.260 (0.150). Ovary very small, flabelliform, medial to seminal receptacle 0.052 – 0.228 (0.100) x 0.033 – 0.104 (0.060), fully developed in only 4 – 8 segments, involutes within 2 segments of achieving full size; vitellarium reniform, posterior to ovary, 0.098 – 0.228 (0.140) x 0.065 – 0.130 (0.090); Mehlis’ gland anterior and medial to vitellarium, c. 0.08 in diameter. Uteri paired in each segment, transverse, tubular; early uterus extends from level of ovary to near proximal pole of cirrus sac; developing uterus extends medially and laterally, lateral extensions crossing osmoregulatory canals dorsally; uteri of each segment do not meet in mid-line. Gravid segments 0.228 – 0.520 (0.340) long, 1.54 – 3.54 (2.69) wide. Egg spherical, shell smooth, 0.045 – 0.065 (0.054) in diameter; pyriform apparatus conical, terminating in numerous fine filaments at apex; oncosphere 0.015 – 0.023 (0.020) in diameter. Osmoregulatory canals paired; ventral canal generally wider than dorsal, medial to it; ventral canal 0.013 – 0.033 (0.020) in diameter, dorsal canal 0.010 – 0.026 (0.020) in diameter; transverse canal connects ventral canal at posterior margin of each segment; accessory canals absent.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF84FFADFF0F34BDD344FC9E.taxon	discussion	Remarks. This species was first described by Nybelin (1917) under the name Hepatotaenia fellicola, based on specimens collected from Macropus agilis in the Kimberley region of Western Australia. It has subsequently been treated as a synonym of P. festiva (for summary, see Beveridge 1976) being reported from the same host in the Northern Territory (Beveridge 1976) and Queensland (Speare et al. 1983). Allozyme studies (Baverstock et al. 1985) suggested that cestodes from M. agilis from Queensland were genetically distinct from those found in other macropodid hosts (other than for one specimen collected from M. dorsalis) and these findings were confirmed using DNA- based methods which found no differences between specimens from the Northern Territory and Queensland (Beveridge et al. 2007). In the latter study, all specimens collected from M. dorsalis were genetically distinct from those found in M. agilis, casting some doubt on the earlier allozyme study by Baverstock et al. (1985). Progamotaenia fellicola is distinguishable from P. festiva in having a quadrangular scolex with either no neck or an extremely short neck in fully relaxed specimens, in having the first mature segment occur in segments 25 – 45, in the cirrus sac invariably extending across the osmoregulatory canals, well into the medulla, and in having a distinct space between the osmoregulatory canals and the lateral extremities of the testis fields. In this species, the occurrence of segments with testes arranged in two groups or a single band within the same strobila is commonly observed. However, where two groups of testes are present, the space between them is small. This species is most similar to P. aemulans sp. nov. from Macropus dorsalis, described below The differentiation of these two species is discussed following the description of the new species.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF83FFAFFF0F3683D75EF94B.taxon	description	(Figs 24 – 30) Synonyms: P. festiva (Rudolphi, 1819) sensu Spratt et al. (1991).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF83FFAFFF0F3683D75EF94B.taxon	biology_ecology	Host: Macropus irma (Jourdan) (Marsupialia: Macropodidae). Site in host: bile ducts. Types: Holotype Jandakot, Western Australia (32 ° 07 ’ S 115 ° 50 ’ E), coll. L. Jue Sue, 13. v. 1982, SAM 29358; paratypes, same data, SAM 22308 (S), 11889 (W).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF83FFAFFF0F3683D75EF94B.taxon	etymology	Etymology: from adspersus, meaning scattered.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF83FFAFFF0F3683D75EF94B.taxon	materials_examined	Material examined: From Macropus irma (black gloved wallaby): Western Australia: types; fragments of several specimens, Collie (SAM 28948, 29359, AM 495419); 1 specimen and fragments of others, Mt Trio (SAM 28947 (S), BMNH 2008. 6.3.8 (S), AM 495420).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF83FFAFFF0F3683D75EF94B.taxon	description	Description: Short, slender worms, 46 – 72 (59, n = 4) long, maximum width 3 – 4 (3.6, n = 4) with 165 – 214 (188, n = 4) segments in gravid specimens. Scolex 0.618 – 0.975 (0.760) in diameter, squat, mildly 4 - lobed; suckers circular, 0.179 – 0.260 (0.230) in diameter. Neck short. First mature segment 62 – 94 th (76). Mature segments craspedote, 1.22 – 2.31 (1.75) wide, 0.21 – 0.31 (0.25) long. Genital atrium small, slightly posterior to middle of lateral segment margin; cirrus sac small, elongate, extending just beyond osmoregulatory canals, 0.078 – 0.195 (0.110) long, 0.039 – 0.059 (0.050) wide; cirrus unarmed; internal seminal vesicle occupies approx. 2 / 3 – 3 / 4 of volume of cirrus sac, 0.085 – 0.163 (0.120) long, 0.026 – 0.039 (0.030) wide; elongate external seminal vesicle, variable in shape, covered with glandular cells, 0.065 – 0.150 (0.100) long, 0.033 – 0.078 (0.060) wide, extending medially from cirrus sac, sometimes overlying cirrus sac; vas deferens coils medially, dorsal to uterus; testes arranged in anterior half of segment between osmoregulatory canals; no space between canals and testis fields; testes arranged in 2 – 3 dorso-ventral layers, in single band or 2 slightly separate groups; segments with single band or 2 groups occur in same specimen. Testes 0.033 – 0.065 (0.050) in diameter; testis number 63 – 96 (80). Vagina tubiform, lacking glandular investment, opens to genital atrium posterior to cirrus sac, leads to large ovoid to sub-spherical seminal receptacle 0.117 – 0.241 (0.170) x 0.078 – 0.143 (0.110). Ovary flabelliform, medial to seminal receptacle 0.065 – 0.124 (0.090) x 0.046 – 0.117 (0.070), fully developed in few segments only; vitellarium reniform, posterior to ovary, 0.052 – 0.163 (0.120) x 0.052 – 0.130 (0.090); Mehlis’ gland anterior and medial to vitellarium. Uteri paired in each segment, transverse, tubular; early uterus extends from level of ovary to or beyond proximal pole of cirrus sac; developing uterus extends medially and laterally, lateral extensions crossing osmoregulatory canals dorsally; uteri of each segment do not meet in mid-line. Gravid segments 0.244 – 0.406 (0.310) long, 1.54 – 2.76 (2.26) wide. Egg spherical, shell smooth, 0.058 – 0.070 (0.064) in diameter; pyriform apparatus conical, terminating in numerous fine filaments at apex; oncosphere 0.015 – 0.018 (0.016) in diameter. Osmoregulatory canals paired; ventral canal generally wider than dorsal, medial to or overlapping dorsal canal; ventral canal 0.020 – 0.065 (0.030) in diameter, dorsal canal 0.007 – 0.020 (0.012) in diameter; transverse canal connects ventral canal at posterior margin of each segment; tiny accessory canals present overlying ventral canal.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF83FFAFFF0F3683D75EF94B.taxon	discussion	Remarks. This species differs from P. festiva in its small size, from P. aemulans sp. nov., P. dilatata sp. nov., P. fellicola, P. onychogale sp. nov. and P. vombati sp. nov. in having a relatively short cirrus sac which does not extend significantly into the medulla and from P. fellicola and P. aemulans in lacking a distinct space between the testis fields and the osmoregulatory canals. The scolex is smaller than that of P. onychogale and it differs from both P. fellicola and P. onychogale in possessing a short neck. It is similar to P. festiva in being a slender cestode, but differs in the short neck and in not having the suckers on “ stalks ” as well as in having testes almost meeting in the midline in most segments. It differs from P. corniculata sp. nov. in having a very short neck and from P. pulchella sp. nov. in having groups of testes which may meet in the midline. This species occurs only in M. irma, a species restricted to the southwest of Western Australia. In this region, the host is in broad sympatry with M. fuliginosus, M. eugenii and Setonix brachyurus. No bile duct cestodes are known from M. eugenii while M. fuliginosus is parasitised by P. effigia and P. pulchella occurs in S. brachyurus.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF80FFB1FF0F30FBD548FE91.taxon	description	(Figs 31 – 37) Synonyms: P. festiva (Rudolphi, 1819) sensu Spratt et al. (1991), Beveridge et al. 1998, Turni and Smales 2001.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF80FFB1FF0F30FBD548FE91.taxon	biology_ecology	Host: Macropus dorsalis (Gray) (Marsupialia: Macropodidae). Site in host: bile ducts. Types: Holotype, Taunton Stn via Dingo, Queensland (23 º 39 ’ 149 º 20 ’), coll. C. Turni, 1996, SAM 29360 (S); paratypes, same data, SAM 29361 (S), BMNH 2008.6.3.9 - 10 (S).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF80FFB1FF0F30FBD548FE91.taxon	etymology	Etymology: rivalling or equalling, indicating close similarity to P. fellicola.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF80FFB1FF0F30FBD548FE91.taxon	materials_examined	Material examined: From Macropus dorsalis: Queensland: types; 1 specimen, 15 km N of Marlborough (SAM 28945, AM 495438); 1 specimen, 48 km E of Moonie (SAM 29362); 1 specimen, Milman (SAM 21508); 2 specimens, Charters Towers (SAM 29363); 1 specimen, Pallamana Stn via Charters Towers (SAM 22284); 1 specimen, Harvest Home Stn via Charters Towers (SAM 22283).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF80FFB1FF0F30FBD548FE91.taxon	description	Description: Small, elongate worms, 20 – 80 (41) long, maximum width 2 – 5 (4) with 84 – 192 (136) segments in gravid specimens. Scolex 0.88 – 1.14 (1.01) in diameter, squat, mildly 4 – lobed; suckers circular, 0.293 – 0.374 (0.333) in diameter. Neck short. First mature segment 40 – 70 th (51). Mature segments craspedote, 1.01 – 3.04 (2.02) wide, 0.163 – 0.374 (0.245) long. Genital atrium small, in posterior part of lateral segment margin; cirrus sac elongate, extending well beyond osmoregulatory canals, 0.228 – 0.423 (0.341) long, 0.052 – 0.085 (0.062) wide; cirrus unarmed; internal seminal vesicle occupies approx. 2 / 3 – 3 / 4 of volume of cirrus sac, 0.078 – 0.247 (0.147) long, 0.013 – 0.059 (0.036) wide; elongate external seminal vesicle, covered with glandular cells, 0.078 – 0.293 (0.153) long, 0.026 – 0.098 (0.064) wide; vas deferens coils medially, dorsal to uterus; testes arranged in anterior half of segment between osmoregulatory canals; space between canals and testis fields; testes arranged in 2 dorso-ventral layers, almost invariably in single band or more rarely in 2 separate groups separated by small space; segments with single band or 2 groups occur in same specimen; in 10 specimens, 20 of 353 segments had testes in 2 groups. Testes 0.046 – 0.078 (0.060) in diameter; testis number 78 – 109 (89). Vagina tubiform, lacking glandular investment, opens to genital atrium posterior to cirrus sac, leads to ovoid to sub-spherical seminal receptacle 0.150 – 0.293 (0.194) x 0.098 – 0.247 (0.137). Ovary flabelliform, medial to seminal receptacle 0.052 – 0.111 (0.80) x 0.046 – 0.111 (0.066), fully developed in c. 12 segments; vitellarium reniform, posterior to ovary, 0.078 – 0.195 (0.140) x 0.046 – 0.130 (0.082); Mehlis’ gland anterior and medial to vitellarium, c. 0.06 in diameter. Uteri paired in each segment, transverse, tubular; early uterus extends from level of ovary to or slightly beyond proximal pole of cirrus sac; developing uterus extends medially and laterally, lateral extensions crossing osmoregulatory canals dorsally; uteri of each segment do not meet in mid-line. Gravid segments 0.228 – 0.569 (0.389) long, 1.84 – 3.82 (3.08) wide. Egg spherical, shell smooth, 0.024 – 0.029 (0.026) in diameter; pyriform apparatus conical, terminating in numerous fine filaments at apex; oncosphere 0.010 – 0.015 (0.014) in diameter. Osmoregulatory canals paired; ventral canal generally wider than dorsal, medial to it; ventral canal 0.013 – 0.033 (0.022) in diameter, dorsal canal 0.010 – 0.033 (0.016) in diameter; transverse canal connects ventral canal at posterior margin of each segment; accessory canals absent.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF80FFB1FF0F30FBD548FE91.taxon	discussion	Remarks. This species most closely resembles P. fellicola in overall size and number of segments, in the cirrus sac projecting well into the medulla and in having a distinctive space between the osmoregulatory canals and the testis fields. It is distinguished from all congeners by these characters. The two species, P. aemulans and P. fellicola, scarcely differ in morphological characters, but are quite distinct genetically and are apparently host specific (Beveridge et al. 2007). The minor differences observable are in scolex diameter, which is 0.65 – 0.89 mm in P. fellicola, and 0.89 – 1.14 mm in P. aemulans and in differences in the rate of maturation of segments, the first mature segment appearing between segments 25 – 45 (34) in P. fellicola and between 40 – 70 (51) in P. aemulans. These are quite minor morphological differences and alone would scarcely warrant the erection of a new species. However, the genetic differentiation between the two is based on both MEE and DNA sequence data (Baverstock et al. 1985; Beveridge et al. 2007), leaving little doubt that the cestodes from these two hosts represent distinct species. The material upon which the new species is based came from the same localities as “ M. dorsalis parasite type 1 ” of Baverstock et al. (1985) and sequences numbers 27 and 28 in Beveridge et al. (2007). The data of Baverstock et al. (1985) suggested that P. fellicola could also occur in M. dorsalis, although the subsequent study of Beveridge et al. (2007) did not confirm this finding. If the data of Baverstock et al. (1985) are correct then it provides compelling evidence that two species are involved as genetic distinction can be maintained within the same host species. A new species is therefore proposed for the material from M. dorsalis and is differentiated from P. fellicola primarily on the basis of scolex size and rate of maturation of the genitalia. At several of the localities from which this species was collected, M. dorsalis was also parasitised by P. festiva. However, it proved possible to separate the two species quite readily based on testis distribution and the extent to which the cirrus sac extended into the medulla.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9FFFB3FF0F3683D5A6FA73.taxon	description	(Figs 38 – 43) Synonyms: Progamotaenia festiva (Rudolphi, 1819) sensu Beveridge et al. (1992), Spratt et al. (1991). Host: Lagorchestes conspicillatus Gould (Marsupialia: Macropodidae). Site in host: bile ducts. Types: Holotype, Inkerman, Queensland (19 ° 44 ’ S 147 ° 28 ’ E), coll. P. M. Johnson, 5. x. 1971, SAM 29355; 20 paratypes, same data, SAM 20760 (S), 10376 (W), BMNH 2008.3. 6.11 (S). Etymology: corniculata means horned and refers to the structure of the pyriform apparatus.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9FFFB3FF0F3683D5A6FA73.taxon	materials_examined	Material examined: From Lagorchestes conspicillatus (spectacled hare wallaby): Queensland: types; 5 specimens, Inkerman (SAM 29356 (S), 7162 (W )), fragments of 15 specimens, Rubyvale (SAM 29357 (S), 10879 (W )); 3 specimens, Blackbraes National Park (SAM 28946 (S), AM 495425); fragments, 80 km N of Moorinya (SAM 28961 (S), AM 495424).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9FFFB3FF0F3683D5A6FA73.taxon	description	Description: Short, slender worms, 50 – 106 (73) long, maximum width 2.5 – 4.0 (3.3) with 158 – 315 (215) segments in gravid specimens. Scolex small 0.49 – 0.68 (0.54) in diameter, weakly 4 - lobed; suckers circular, 0.146 – 0.228 (0.195) in diameter. Neck slender. First mature segment 72 – 90 th (81). Mature segments craspedote, 0.81 – 1.69 (1.34) wide, 0.14 – 0.31 (0.19) long. Genital atrium small, in middle of lateral segment margin; cirrus sac elongate, extending well beyond osmoregulatory canals, 0.228 – 0.390 (0.302) long, 0.026 – 0.065 (0.047) wide; cirrus unarmed; internal seminal vesicle occupies approx. 2 / 3 – 3 / 4 of volume of cirrus sac, 0.085 – 0.195 (0.140) long, 0.033 – 0.098 (0.060) wide; elongate external seminal vesicle, covered with glandular cells, 0.052 – 0.150 (0.081) long, 0.033 – 0.059 (0.044) wide, sometimes overlying cirrus sac; vas deferens coils medially, dorsal to uterus; testes arranged in anterior half of segment between osmoregulatory canals; no or very slight space between canals and testis fields; testes arranged in 2 – 3 dorsoventral layers, usually in single band, less frequently in 2 separate groups; segments with single band or 2 groups occur in same specimen. Testes 0.039 – 0.059 (0.048) in diameter; testis number 50 – 63 (57). Vagina tubiform, lacking glandular investment, opens to genital atrium posterior to cirrus sac, leads to ovoid seminal receptacle 0.078 – 0.163 (0.109) x 0.010 – 0.098 (0.059). Ovary flabelliform, medial to seminal receptacle 0.046 – 0.098 (0.064) x 0.026 – 0.065 (0.043); vitellarium reniform, posterior to ovary, 0.065 – 0.163 (0.094) x 0.033 – 0.072 (0.054); Mehlis’ gland anterior and medial to vitellarium, c. 0.08 in diameter. Uteri paired in each segment, transverse, tubular; early uterus extends from level of ovary to near proximal pole of cirrus sac; developing uterus extends medially and laterally, lateral extensions crossing osmoregulatory canals dorsally; uteri of each segment do not meet in mid-line. Gravid segments 0.195 – 0.488 (0.369) long, 1.41 – 3.41 (2.08) wide. Egg spherical, shell smooth, 0.039 – 0.043 (0.041) in diameter; pyriform apparatus conical, with 2 tapering arms, without filaments at apices; oncosphere 0.012 – 0.016 (0.014) in diameter. Osmoregulatory canals paired; ventral canal generally wider than dorsal, medial to it; ventral canal 0.020 – 0.052 (0.029) in diameter, dorsal canal 0.013 – 0.026 (0.016) in diameter; transverse canal connects ventral canal at posterior margin of each segment; accessory canals absent.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9FFFB3FF0F3683D5A6FA73.taxon	discussion	Remarks. This species differs from P. festiva in being a small species, 50 – 106 mm long, with the cirrus sac extending across the osmoregulatory canals, into the medulla. It differs from P. fellicola and P. aemulans in that there is no space between the testis fields and osmoregulatory canals and it differs from these species as well as from P. dilatata sp. nov., P. onychogale sp. nov. and P. pulchella sp. nov. in having an elongate neck. It also differs from all species of the complex other than P. pulchella in having a pyriform apparatus terminating in two horns rather than in numerous reflexed filaments.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9CFFB5FF0F3013D44BFABD.taxon	description	(Figs 44 – 50) Synonyms: Progamotaenia festiva (Rudolphi, 1819) in part, Beveridge (1976), Beveridge et al. (1985, 1998), Spratt et al. (1991).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9CFFB5FF0F3013D44BFABD.taxon	biology_ecology	Host: Wallabia bicolor (Desmarest) (Marsupialia: Macropodidae). Site in host: bile ducts. Types: Holotype, Nariel, Victoria (36 º 20 ' S 147 º 48 ' E), coll. D. M. Spratt, 29. v. 1974, SAM 29364; 2 paratypes, same data SAM 29365.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9CFFB5FF0F3013D44BFABD.taxon	etymology	Etymology: from dilatatus (= widened) referring to the broad anterior end of this species.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9CFFB5FF0F3013D44BFABD.taxon	materials_examined	Material examined: From Wallabia bicolor (swamp wallaby): Queensland: 1 specimen, Dingo Beach (SAM 29370 (S )), 1 specimen, Airlie Beach (SAM 29376 (S )); 2 specimens, Rockhampton (SAM 19757 (S )); New South Wales: 6 specimens, Dubbo (SAM 29368 (S )), 1 specimen, Ourimbah (SAM 20067 (S), 1116 (W )), 1 specimen, Nowra (SAM 21489 (S )); Australian Capital Territory: 1 specimen, Tidbinbilla (WLHC S 310 (S), C 235 (W), BMNH 2008.3. 6.12 (S )); Victoria: fragments, Nariel (SAM 29366 (S), 25408 (W )), 3 specimens, Cudgewa (SAM 20825, WLHC C 6, C 12 (S )), 2 specimens, Dartmouth (SAM 20928); 1 specimen, Dixon’s Creek (SAM 29369 (S )).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9CFFB5FF0F3013D44BFABD.taxon	description	Description: Broad worms, 12 – 58 (35) long, maximum width 3 – 8 (5.7) with 92 – 228 (167) segments in gravid specimens. Scolex 0.85 – 1.30 (1.06) in diameter, squat, weakly 4 - lobed; suckers circular, 0.293 – 0.569 (0.436) in diameter. Neck very short. First mature segment 22 – 41 th (32, n = 7). Mature segments craspedote, 1.46 – 2.68 (2.11) wide, 0.065 – 0.179 (0.125) long. Genital atrium small, in middle or slightly posterior to middle of lateral segment margin; cirrus sac elongate, extending well beyond osmoregulatory canals, 0.163 – 0.598 (0.423) long, 0.046 – 0.117 (0.079) wide; cirrus unarmed; internal seminal vesicle occupies approx. 2 / 3 – 3 / 4 of volume of cirrus sac, 0.065 – 0.377 (0.243) long, 0.020 – 0.052 (0.040) wide; elongate external seminal vesicle, covered with layer of glandular cells, 0.072 – 0.208 (0.134) long, 0.033 – 0.143 (0.070) wide; vas deferens coils medially, dorsal to uterus; testes arranged in anterior half of segment between osmoregulatory canals; no prominent space between canals and testis fields; testes arranged in 2 dorso-ventral layers, in single band or in 2 separate groups separated by small space. Testes 0.033 – 0.065 (0.050) in diameter; testis number 68 – 78 (75, n = 5). Vagina tubiform, lacking glandular investment, opens to genital atrium posterior to cirrus sac, leads to ovoid to subspherical seminal receptacle 0.130 – 0.293 (0.212) x 0.052 – 0.195 (0.104). Ovary flabelliform, medial to seminal receptacle 0.065 – 0.124 (0.085) x 0.033 – 0.078 (0.051), fully developed in few segments; vitellarium reniform, posterior to ovary, 0.078 – 0.195 (0.124) x 0.046 – 0.098 (0.066); Mehlis’ gland anterior and medial to vitellarium. Uteri paired in each segment, transverse, tubular; early uterus extends from level of ovary to or beyond proximal pole of cirrus sac; developing uterus extends medially and laterally, lateral extensions crossing osmoregulatory canals dorsally; uteri of each segment do not meet in mid-line. Gravid segments 0.147 – 0.406 (0.241) long, 2.28 – 4.23 (3.27) wide. Egg spherical, shell smooth, 0.055 – 0.062 (0.058) in diameter; pyriform apparatus conical, terminating in numerous fine filaments at apex; oncosphere 0.012 – 0.016 (0.013) in diameter. Osmoregulatory canals paired; ventral canal generally wider than dorsal, medial to it; ventral canal 0.013 – 0.046 (0.023) in diameter, dorsal canal 0.007 – 0.026 (0.013) in diameter; transverse canal connects ventral canal at posterior margin of each segment; accessory canals absent.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9CFFB5FF0F3013D44BFABD.taxon	discussion	Remarks. P. dilatata differs from P. festiva in having a very short neck and a cirrus sac which extends well into the medulla. It differs from P. fellicola and P. aemulans in lacking a distinctive space between the testis fields and the osmoregulatory canals, and from P. pulchella in that the testes are usually distributed in a single band rather than being invariably distributed in two separate groups. It can be distinguished from P. corniculata by the long neck of the latter species and by the fact that the pyriform apparatus of P. corniculata terminates in two separate horns without reflexed filaments. P. dilatata differs from P. onychogale in being an elongate cestode rather than being short and foliiform; P onychogale has up to 84 segments while P. dilatata has 92 – 228 segments. P. dilatata differs from P. adspersa in M. irma in that the cirrus sac of the latter species extends just across the osmoregulatory canals in to the medulla while approximately 50 % of the cirrus sac of P. dilatata lies within the medulla. In addition, the size of the suckers of P. dilatata (0.29 – 57 (0.44) mm) exceeds those of P. adspersa (0.18 – 0.26 (0.23) mm). P. dilatata is readily distinguishable from P. vombati in the shape of the scolex (Figs. 45, 46) and in the number of testes per segment which are 83 – 122 (98) in P. vombati and 68 – 78 (75) in P. dilatata.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9AFFB7FF0F33AED78BFA09.taxon	description	(Figs 51 – 58) Synonyms: Progamotaenia festiva (Rudolphi, 1817) Nybelin, 1917 of Beveridge et al. (1992).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9AFFB7FF0F33AED78BFA09.taxon	biology_ecology	Host: Onychogalea unguifera (Gould) (Marsupialia: Macropodidae). Site in host: bile ducts. Types: Holotype, Springfield Stn via Mt Surprise, Queensland (17 ° 58 ’ S 144 ° 26 ’ E), coll. I. Beveridge, 8. xi. 1994 (SAM 28950, AM 495418); paratype, same data (SAM 29334).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9AFFB7FF0F33AED78BFA09.taxon	etymology	Etymology: the name of the new species is derived from the generic name of the host.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9AFFB7FF0F33AED78BFA09.taxon	materials_examined	Material examined: From O. unguifera: Queensland: types, 45 specimens, Chadshunt Stn via Burketown (SAM 21375, 21388, 29335 - 7 (S), 6633, 6638 - 40, 6655 - 6 (W), BMNH 2008.6.3.6 - 7 (S )), 10 specimens, Wernadinga Stn via Burketown (SAM 21393, 29338 (S), 6660 - 3 (W )); fragments, Bylong Stn via Richmond (SAM 7920 (W )).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9AFFB7FF0F33AED78BFA09.taxon	description	Description: Short, robust worms of variable shape, quadrangular to foliiform, 12 – 24 (15) long, maximum width 4 – 10 (8.2) with 54 – 84 (69) segments in gravid specimens. Scolex large 1.14 – 1.74 (1.40) in diameter, squat, weakly 4 - lobed; suckers circular, 0.406 – 0.640 (0.519) in diameter. Neck absent. First mature segment 18 – 22 th (19). Mature segments craspedote, 3.58 – 8.83 (6.26) wide, 0.08 – 0.21 (0.16) long. Genital atrium small, slightly posterior to middle of lateral segment margin; cirrus sac elongate, extending well beyond osmoregulatory canals, 0.65 – 0.1.36 (0.87) long, 0.065 – 0.111 (0.093) wide; cirrus unarmed; internal seminal vesicle occupies approx. 2 / 3 – 3 / 4 of volume of cirrus sac, 0.228 – 0.867 (0.456) long, 0.065 – 0.111 (0.093) wide; elongate external seminal vesicle, covered with glandular cells, 0.130 – 0.459 (0.289) long, 0.052 – 0.194 (0.089) wide, sometimes overlying cirrus sac; vas deferens coils medially, dorsal to uterus; testes arranged in anterior half of segment between osmoregulatory canals; no space between canals and testis fields; testes arranged in 2 – 3 dorso-ventral layers, in single band or 2 separate groups; segments with single band or 2 groups occur in same specimen; testes clustered lateral to seminal receptacle, arranged in single file, often sparsely anterior to receptacle, single row or 2 rows medial to receptacle. Testes 0.046 – 0.092 (0.063) in diameter; testis number 93 – 128 (110). Vagina tubiform, lacking glandular investment, opens to genital atrium posterior to cirrus sac, leads to enormous ovoid seminal receptacle 0.224 – 1.22 (0.696) x 0.078 – 0.235 (0.183); seminal receptacle ovoid, greatly elongated, often bilobed, extending from transverse osmoregulatory canal to anterior margin of segment; testes frequently compressed between adjacent receptacles. Ovary small, flabelliform, medial to seminal receptacle 0.078 – 0.495 (0.197) x 0.052 – 0.204 (0.108), fully developed in only 5 – 15 segments; vitellarium reniform, posterior to ovary, 0.059 – 0.235 (0.140) x 0.046 – 0.153 (0.081); Mehlis’ gland anterior and medial to vitellarium, c. 0.08 in diameter. Uteri paired in each segment, transverse, tubular; early uterus extends from level of ovary to or beyond proximal pole of cirrus sac; developing uterus extends medially and laterally, lateral extensions crossing osmoregulatory canals dorsally; uteri of each segment do not meet in mid-line. Gravid segments 0.130 – 0.461 (0.214) long, 4.48 – 9.98 (7.70) wide. Egg spherical, shell smooth, 0.050 – 0.060 (0.056) in diameter; pyriform apparatus conical, with 2 tapering arms, terminating in numerous fine filaments at apex; oncosphere 0.015 – 0.025 (0.019) in diameter. Osmoregulatory canals paired; ventral canal generally wider than dorsal, medial to it; ventral canal 0.010 – 0.051 (0.034) in diameter, dorsal canal 0.006 – 0.020 (0.014) in diameter; transverse canal connects ventral canal at posterior margin of each segment; accessory canals absent.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF9AFFB7FF0F33AED78BFA09.taxon	discussion	Remarks. This distinctive species was found only in O. unguifera in Queensland. It is much shorter and broader than any other species though the variation in shape may be due in part to differing methods of fixation used. The scolex and suckers of this species are much larger than any other members of the complex and a neck is lacking. This species is distinguished by its long cirrus sac extending well beyond the osmoregulatory canals, by the very large, elongate seminal receptacle and by the large number of testes (93 – 128) per segment. The testes were extremely difficult to count in this species as they occur in several dorso-ventral layers; counting was achieved by drawing the testes and then counting them from the drawing. The egg of this species is also distinctive with the pyriform apparatus terminating in two arms. Among congeners, a similar pyriform apparatus is known only from P. diaphana, a parasite of the bile ducts of the southern hairy-nosed wombat, Lasiorhinus latifrons, P. pulchella sp. nov. from the quokka, Setonix brachyurus and P. corniculata sp. nov. from the bile ducts of the spectacled hare wallaby, Lagorchestes conspicillatus. The type specimens have been selected from the material used in the molecular study of Beveridge et al. (2007) which showed that the single specimen examined from O. unguifera was genetically distinct from all other bile duct cestodes. Beveridge et al. (1992) reported this cestode in 100 % of the 13 O. unguifera examined for parasites in northern Queensland.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF98FFB9FF0F303AD7B1FB8F.taxon	description	(Figs 59 – 64) Synonyms: Progamotaenia festiva (Rudolphi, 1819) sensu Beveridge (1976), Baverstock et al. (1985), Spratt et al. (1991).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF98FFB9FF0F303AD7B1FB8F.taxon	biology_ecology	Host: Setonix brachyurus (Quoy & Gaimard) (Marsupialia: Macropodidae). Site in host: bile ducts. Types: Holotype, Rottnest Island, Western Australia (31 º 59 ' S 115 º 31 ' E), coll. S. Barker, 19. x. 1959 (SAM 20052, 29372); 9 paratypes, same locality, coll. 19. x. 1959, 30. xii. 1959 (SAM 20052 - 3).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF98FFB9FF0F303AD7B1FB8F.taxon	etymology	Etymology: diminutive of pulcher which means beautiful.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF98FFB9FF0F303AD7B1FB8F.taxon	materials_examined	Material examined: From Setonix brachyurus (quokka): Western Australia: types; 5 specimens, Rottnest Island (AHC 20790, 21376).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF98FFB9FF0F303AD7B1FB8F.taxon	description	Description: Slender, elongate cestodes, 27 – 70 (39.9) long, maximum width 1.5 – 4.0 (2.6) with 115 – 243 (173) segments in gravid strobilae. Scolex 0.358 – 0.618 (0.518) in diameter, barely lobed at apex; suckers circular, 0.065 – 0.211 (0.176) in diameter. Neck short. First mature segment 53 – 63 th (58, n = 6). Mature segments craspedote, 0.650 – 1.54 (1.09) wide, 0.146 – 0.276 (0.213) long. Genital atrium small, in middle of or slightly posterior to middle of lateral segment margin; cirrus sac small, elongate, extending beyond osmoregulatory canals, 0.176 – 0.260 (0.207) long, 0.020 – 0.065 (0.030) wide; cirrus unarmed; internal seminal vesicle occupies approx. 2 / 3 – 3 / 4 of volume of cirrus sac, 0.078 – 0.124 (0.103) long, 0.033 – 0.072 (0.052) wide; elongate external seminal vesicle, covered with layer of glandular cells, 0.052 – 0.091 (0.068) long, 0.026 – 0.078 (0.041) wide; vas deferens coils medially, dorsal to uterus; testes arranged in anterior half of segment between osmoregulatory canals; no prominent space between canals and testis fields; testes arranged in 2 dorso-ventral layers, in 2 separate groups. Testes 0.026 – 0.046 (0.040) in diameter; testes number 34 – 47 (39, n = 5). Vagina tubiform, lacking glandular investment, opens to genital atrium posterior to cirrus sac, leads to ovoid to sub-spherical seminal receptacle 0.085 – 0.228 (0.144) x 0.065 – 0.163 (0.114). Ovary flabelliform, medial to seminal receptacle 0.039 – 0.098 (0.070) x 0.033 – 0.085 (0.055); vitellarium reniform, posterior to ovary, 0.052 – 0.182 (0.105) x 0.039 – 0.098 (0.071); Mehlis’ gland anterior and medial to vitellarium. Uteri paired in each segment, transverse, tubular; early uterus extends from level of antiporal margin of ovary to near proximal pole of cirrus sac; developing uterus extends medially and laterally, lateral extensions crossing osmoregulatory canals dorsally; uteri of each segment do not meet in mid-line. Gravid segments 0.195 – 0.406 (0.301) long, 0.98 – 1.95 (1.49) wide. Egg spherical, shell smooth, 0.041 – 0.047 (0.043) in diameter; pyriform apparatus terminating in 2 horn-like projections; oncosphere 0.012 – 0.014 (0.013) in diameter. Osmoregulatory canals paired; ventral canal generally wider than dorsal, medial to it; ventral canal 0.013 – 0.026 (0.018) in diameter, dorsal canal 0.007 – 0.020 (0.012) in diameter; transverse canal connects ventral canal at posterior margin of each segment; accessory canals absent.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF98FFB9FF0F303AD7B1FB8F.taxon	discussion	Remarks. This species is differentiated from P. festiva by its small size and from P. festiva and P. corniculata in having a short neck. It is distinguished from all of the new species described here in having the testes invariably distributed in two distinct groups within each segment, a character which it shares with P. festiva. It is further distinguished from P. festiva by having a pyriform apparatus which terminates in two horns rather than being conical and terminating with numerous reflexed filaments as occurs in P. festiva. It differs from all bile duct-inhabiting cestodes in having a smaller average number of testes per segment (39). The only species with which the number of testes overlaps is P. diaphana, found in the southern hairy-nosed wombat, Lasiorhinus latifrons, in which the scolex contains black, pigmented areas. Material of this species was included in the MEE study by Baverstock et al. (1985), but was not available for the molecular study of Beveridge et al. (2007).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF96FFBBFF0E32B8D365FC4D.taxon	description	(Figs 65 – 70) Synonyms: Progamotaenia festiva (Rudolphi, 1819) sensu Beveridge (1976), Baverstock et al. (1985), Spratt et al. (1991).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF96FFBBFF0E32B8D365FC4D.taxon	biology_ecology	Host: Vombatus ursinus (Shaw) (Marsupialia: Vombatidae). Site in host: bile ducts. Types: Holotype, Little Bombay, New South Wales (35 ° 24 ' S, 149 ° 40 ' E), coll. D. M. Spratt, 3. ii. 1976, SAM 29371; paratypes (fragments), same data, SAM V 758.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF96FFBBFF0E32B8D365FC4D.taxon	etymology	Etymology: the species name is derived from the generic name of the host.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF96FFBBFF0E32B8D365FC4D.taxon	materials_examined	Material examined: From V. ursinus (common wombat): New South Wales: types; 2 scoleces and fragments, Bondo State Forest (SAM 22093 (S), WLHC C 181, C 225 (S )); Australian Capital Territory: 1 specimen, Condor Creek (WLHC C 121 (S )), 1 scolex, fragments, Brindabella Range (SAM V 759 (S )); Victoria: 1 scolex, fragments, Bairnsdale (SAM 21334 (S )), fragments, Sarsfield (SAM 21504 (S )).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF96FFBBFF0E32B8D365FC4D.taxon	description	Description: Worms of variable width, no intact specimens. Scolex 0.76 – 1.38 (1.13, n = 3) in diameter, weakly 4 - lobed at apex; suckers circular, 0.406 – 0.455 (0.423, n = 3) in diameter. Neck prominent. First mature segment 85 th (n = 1). Mature segments craspedote, 1.02 – 3.49 (2.59) wide, 0.163 – 0.569 (0.356) long. Genital atrium small, in posterior third of lateral segment margin; cirrus sac elongate, extending well beyond osmoregulatory canals, 0.228 – 0.650 (0.435) long, 0.052 – 0.130 (0.085) wide; cirrus unarmed; internal seminal vesicle occupies approx. 2 / 3 – 3 / 4 of volume of cirrus sac, 0.098 – 0.410 (0.271) long, 0.020 – 0.046 (0.034) wide; elongate external seminal vesicle, covered with layer of glandular cells, 0.065 – 0.390 (0.222) long, 0.033 – 0.195 (0.117) wide; vas deferens coils medially, dorsal to uterus; testes arranged in anterior half of segment between osmoregulatory canals; no prominent space between canals and testis fields; testes arranged in 2 dorso-ventral layers, in single band or in 2 separate groups separated by small space. Testes 0.026 – 0.078 (0.063) in diameter; testis number 83 – 112 (98, n = 10). Vagina tubiform, lacking glandular investment, opens to genital atrium posterior to cirrus sac, leads to ovoid seminal receptacle 0.065 – 0.423 (0.278) x 0.039 – 0.312 (0.190). Ovary flabelliform, medial to seminal receptacle 0.046 – 0.195 (0.112) x 0.033 – 0.150 (0.087); vitellarium reniform, posterior to ovary, 0.078 – 0.293 (0.182) x 0.052 – 0.228 (0.115); Mehlis’ gland anterior and medial to vitellarium. Uteri paired in each segment, transverse, tubular; early uterus extends from level of antiporal margin of ovary to near proximal pole of cirrus sac; developing uterus extends medially and laterally, lateral extensions crossing osmoregulatory canals dorsally; uteri of each segment do not meet in mid-line. Gravid segments 0.455 – 0.861 (0.661, n = 9) long, 2.19 – 3.41 (2.72, n = 9) wide. Egg spherical, shell smooth, 0.045 – 0.049 (0.048) in diameter; pyriform apparatus conical, terminating in numerous fine filaments at apex; oncosphere 0.014 – 0.020 (0.017) in diameter. Osmoregulatory canals paired; ventral canal generally wider than dorsal, medial to it; ventral canal 0.013 – 0.052 (0.034) in diameter, dorsal canal 0.013 – 0.033 (0.018) in diameter; transverse canal connects ventral canal at posterior margin of each segment; accessory canals absent.	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
3F1087F9FF96FFBBFF0E32B8D365FC4D.taxon	discussion	Remarks. This species is described from a limited range of material, including only a single complete specimen. Consequently, the metric data available for this new species are limited. P. vombati differs from P. festiva in being a small species with the first mature segment at about segment number 85 compared with 170 – 205 in P. festiva. It differs from P. fellicola and P. aemulans in lacking a distinct space between the testis fields and the osmoregulatory canals and from P. pulchella in having the testes usually arranged in a single band. It is further differentiated from P. pulchella, as well as from P. corniculata and P. onychogale in having a conical pyriform apparatus rather than one which terminates in paired horns. P. vombati differs from P. adspersa in which the cirrus sac barely penetrates into the medulla and from P. dilatata in the number of testes per segment (68 – 78 in P. dilatata, 83 – 112 in P. vombati). Material of this species was included in the MEE study by Baverstock et al. (1985), but was not available for the molecular study of Beveridge et al. (2007).	en	Beveridge, I., Shamsi, S. (2009): Revision of the Progamotaenia festiva species complex (Cestoda: Anoplocephalidae) from Australasian marsupials, with the resurrection of P. fellicola (Nybelin, 1917) comb. nov. Zootaxa 1990: 1-29
