taxonID	type	description	language	source
3E641D3CFFC2FFFCFEADFDABFE92F926.taxon	description	(Figures 1 – 3)	en	Akira Shimizu, Raymond Wahis, Anthony C. Harris, James P. Pitts (2016): An extraordinary new genus and species of spider wasps (Insecta: Hymenoptera: Pompilidae) from Southeast Asia. Journal of Natural History 50: 1549-1559, DOI: 10.1080/00222933.2016.1155781
3E641D3CFFC2FFFCFEADFDABFE92F926.taxon	diagnosis	Diagnosis The female has unique characteristics among Pompilidae in having a remarkably shaped head, its outline rhomboid in frontal view (Figure 1 A) and triangular in profile (Figure 1 B); upper frons strongly produced anteriorly, hence supra-antennal area overhanging radicle of scape; lower frons markedly receding posteriorly, greatly narrowed below, flattened. The following characteristics are also very specialised and, together with the above traits, distinguish this genus from others: (1) body very slender (Figure 2 A); (2) mesosoma and metasoma compressed laterally; (3) upper frons with several long, stout, suberect setae (Figures 1 A, B); (4) inner orbits strongly arcuate and remarkably convergent below; (5) eye with numerous erect setulae, those being much longer than facet width; (6) clypeus small, entirely flattened; (7) antenna not convolute (Figures 2 A, 3); (8) juncture of anterior and posterior faces of vertex sharply edged; (9) posterior faces of vertex and gena concave, smooth and highly polished, without occipital carina (Figure 1 C); (10) pronotum much longer than mesoscutum at midline (Figure 1 D); (11) pronotal collar slightly depressed below level of its dorsum (Figure 2 B); (12) oblique sulcus on mesepisternum subobsolete (Figure 2 C); (13) mesepimeron entirely obsolete, save for its upper margin (Figure 2 C, arrow); (14) metapostnotum completely invaginated dorsally; (15) lower metapleuron and lateral side of propodeum fairly flattened, smooth and highly polished (Figure 2 C); (16) pterostigma very small and triangular, its base much shorter than cross vein 2 r-rs (Figure 2 D); (17) marginal cell very long and narrow, strongly tapering apically (Figure 2 E); (18) vein M reaching outer wing margin; (19) hind wing with only one basal hamulus; (20) hind wing cross vein cu-a short and almost straight, originating very basally to fork of vein M + CuA, forming an obtuse angle with vein 1 A (Figure 2 F); (21) jugal lobe very small; (22) outer face of fore coxa flattened, with a fine, semi-circularly curved groove posteriorly from base to apex (Figure 2 G, arrow); (23) fore femur enormously incrassate (Figure 2 G); (24) hind coxa much larger than mid coxa, strongly compressed laterally, greatly raised and lamellate dorso-innerly, smooth and polished, forming the same plane with both lower metapleuron and lateral face of propodeum (Figure 2 B); (25) outer apicoventral corner of hind femur rounded; (26) orbicula minute, with its pecten consisting of a few radiating, almost straight weak setulae; (27) all tarsal claws edentate (Figure 1 E); (28) tergum I narrowed and parallel sided basally (Figure 2 A); and (29) sternum VI strongly compressed laterally, with a median carina (Figure 2 H).	en	Akira Shimizu, Raymond Wahis, Anthony C. Harris, James P. Pitts (2016): An extraordinary new genus and species of spider wasps (Insecta: Hymenoptera: Pompilidae) from Southeast Asia. Journal of Natural History 50: 1549-1559, DOI: 10.1080/00222933.2016.1155781
3E641D3CFFC2FFFCFEADFDABFE92F926.taxon	description	Description Body smooth and impunctate. Ocelli forming low obtuse-angled triangle (Figure 1 F). Malar space much longer than one-half width of mandibular base (Figure 1 B). Labrum small, fairly exposed beneath clypeus (Figure 1 A); apical margin feebly convex without median emargination. Mandible small, thin antero-posteriorly with small tooth subapically on inner margin, without lamina or fimbriate groove along outer margin. Maxilla with palpus elongate (Figures 1 G, H); last three palpomeres thinner than third, fourth longer than third. Labium with prementum without median cleft apically or circular or heart- or spade-shaped membranous area subapically (Figure 1 I). Antennal socket removed from upper margin of clypeus by much more than its long axis. Scape long, flattened and curved outward on outer face (Figure 1 F). Pronotum with anterior declivity much shorter than dorsum, gradually and uniformly merging into dorsum, thus never differentiated from dorsum (Figure 2 B); lateral face almost vertical and concave (Figure 2 A), juncture of lateral and dorsal faces narrowly rounded. Dorsal surface of pronotum, mesoscutum and scutellar disc flattened, almost conforming to the same plane (Figure 2 B). Posterolateral margin of mesoscutum not raised. Axilla (cuneoli) practically absent. Propodeal dorsum much longer than declivity, flattened, and gradually merging into latter (Figure 2 C); declivity somewhat concave posteromedially. Third abscissa of fore wing vein M (basal vein) curved anteriorly (Figure 2 E). Cross veins 2 and 3 rs-m almost straightly slanting and subparallel. Cross vein 2 m-cu meeting vein CuA 1 much less than half of distance from base of CuA 1 to wing margin. Vein CuA 1 slightly deflected downward basally. Cross vein cu-a originating basally to fork of vein M + CuA. Mid coxa sharply raised and lamellate on dorso-outer surface. Fore tarsomere I shorter than tarsomeres II – IV combined. Mid and hind femora with basal ring, and small spines posterodorsally. Hind femur compressed laterally, its inner face flattened. Hind tibia with many short but strong spines dorsally, without scale-like teeth; apical spines equal in thickness but unequal in length, weakly splayed out (Figure 2 I). Apicoventral spine on hind tarsomere V and apicodorsal spine on hind tarsomere IV somewhat broadened basally and narrowly triangular. Metasoma slender, fusiform (Figure 2 A). Tergum 1 with lateral crease. Sternum II without transverse groove.	en	Akira Shimizu, Raymond Wahis, Anthony C. Harris, James P. Pitts (2016): An extraordinary new genus and species of spider wasps (Insecta: Hymenoptera: Pompilidae) from Southeast Asia. Journal of Natural History 50: 1549-1559, DOI: 10.1080/00222933.2016.1155781
3E641D3CFFC2FFFCFEADFDABFE92F926.taxon	distribution	Distribution Southeast Asia (Malaysia, Thailand).	en	Akira Shimizu, Raymond Wahis, Anthony C. Harris, James P. Pitts (2016): An extraordinary new genus and species of spider wasps (Insecta: Hymenoptera: Pompilidae) from Southeast Asia. Journal of Natural History 50: 1549-1559, DOI: 10.1080/00222933.2016.1155781
3E641D3CFFC2FFFCFEADFDABFE92F926.taxon	biology_ecology	Biology Unknown.	en	Akira Shimizu, Raymond Wahis, Anthony C. Harris, James P. Pitts (2016): An extraordinary new genus and species of spider wasps (Insecta: Hymenoptera: Pompilidae) from Southeast Asia. Journal of Natural History 50: 1549-1559, DOI: 10.1080/00222933.2016.1155781
3E641D3CFFC6FFFFFEBBFF04FE45FD8F.taxon	description	(Figures 1 – 3) Description of the holotype female Length: body 8.8 mm; fore wing 6.1 mm. Body devoid of long bristles, except for several stout bristles on upper frons and a few weak setae on sternum VI. Pubescence on body long and rather sparse, hence integument is visible. Body black, yellow and orange. Head and mesosoma predominantly black with the following light yellow: V-shaped broad marking along inner orbits on upper frons (Figure 2 A), lower frons, except for ill-defined brown markings below antennal sockets, clypeus, labrum, mandible, gena (Figure 2 B), pair of large spots anterolaterally and pair of transverse spots posteriorly on pronotum, spots on upper and lower mesepisterna dorsally (Figure 2 C), tegula and two pairs of large anterolateral spots on metasomal terga II and III; metasoma with the following orange-red: metasomal tergum I apically, terga II and basal two-thirds of tergum III, except for two pairs of yellow spots, sternum I apically and sterna II – III, remainder black. Antenna mostly reddish brown, darker toward apex dorsally; scape yellow laterally; scape, pedicel and flagellomere I yellowish brown ventrally. Maxillary galea, lacinia, palpus and labial palpus light yellow; other parts of labio-maxillary complex dark brown. All coxae except fore and hind coxal bases, fore and mid femora dorsally, except for bases, fore tibia dorsally, dorsobasal halves of mid and hind tibiae light yellow; fore and mid tarsi, mid tibia except dorsobasal half, hind femur apically and hind tibia internally and ventrally orangered; others blackish brown; fore tibial spur yellow; mid and hind tibial spurs ivorywhite. Fore- and hind wing bases light yellow; fore wing (Figure 2 E) hyaline with yellowish tint, apically with a broad fuscous band covering apical three-quarters of marginal cell, SMCs 3 and 4, discal cell 3, apical margins of SMC 2 and discal cell 2, and apical two-thirds of subdiscal cell 2 (Figure 2 E). Head in frontal view 1.4 times as broad as long. Vertex moderately convex between eye tops (Figure 1 A). Frontal line very short and faint near antennal sockets. Ratio of UID: MID: LID = 8.3: 10: 4.4. Eye 0.59 times half of MID. Ocelli large (Figure 1 F); ocellar area not raised. Ratio of POD: OOD = 1: 0.24; POD 3.1 times as long as width of anterior ocellus. OVD / POD = 0.36. Gena in dorsal view very short, almost linear (Figure 1 F), in lateral view 0.3 times eye width, strongly narrowed above, gradually broadened below (Figure 1 B). Malar space as long as thickness of flagellum. Clypeus 1.5 times as broad as long; anterolateral corner subangulate; apical rim very short, set off from main part of clypeus by fine groove; apical margin arcuately concave. Length ratio of palpomeres I – VI = 7: 14: 31: 35: 25: 31. Length ratio of scape, pedicel, and flagellomeres I and II = 12.9: 5: 10: 10; flagellomere I 3.4 times as long as broad, 0.47 times as long as UID. Posterior margin of vertex weakly concave in dorsal view (Figure 1 F). Pronotum with shoulder slightly produced laterally (Figure 2 A); lateral face smooth and strongly polished; posterior margin obtusely subangulate medially. Mesoscutum with parapsidal sulcus finely impressed, divergent anteriorly. Scutellum with lateral declivity very steep, smooth and strongly polished. Metanotum smooth and strongly polished laterally with disc declined posteriorly. Posterior margin of mesosternum remarkably produced posteriorly into pair of median lobes in front of mid coxae. Metapleuron with ventral and posterior rims distinctly set off from its main part by groove and reflexed (Figure 2 C). Propodeum 1.1 times as long as broad in dorsal view, gradually narrowing caudad with small triangular, strongly polished area and short, sharply impressed median groove basimedially. Fore wing with fifth abscissa of vein M, cross veins 2 and 3 rs-m and 2 m-cu mostly unpigmented for most part (Figure 2 E). Marginal cell removed from wing tip by 0.17 times its own length. Last three abscissae of vein Rs almost straight. Length ratio of cross vein 2 r-rs and last three abscissae of vein Rs = 10: 6.7: 14.2: 25.8. SMC 2 receiving cross vein 1 m-cu at basal 0.29. SMC 3 0.87 times as long as SMC 2 on vein M, 2.1 times as long as SMC 2 on vein Rs, narrowed on vein Rs by 0.90 times its own length on vein M, receiving cross vein 2 m-cu almost at its base. Vein M with fifth abscissa deflected downward; last three abscissae almost straight. Last abscissa of hind wing vein M obsolete (Figure 2 F). Fore tarsomere I almost as long as tarsomere V. Mid tibia with few minute spines dorsally. Hind tibia with dorsal spines much longer and more abundant than those on mid tibia. Longer spur of hind tibia 0.68 times as long as basitarsus. Hind tarsomere V with few small, irregularly arranged spines ventrally. Metasoma broadest at apex of tergum II (Figure 2 A). Sternum I strongly compressed laterally, smooth and strongly polished. Tergum VI and sternum VI compressed laterally (Figure 2 H). Variation based on the paratype female Length: body 5.7 mm; fore wing 4.3 mm. Body more extensively black than in holotype (Figure 3); the following light yellow: V-shaped narrow marking along inner orbits on upper frons, lateral narrow marking along inner orbit on lower frons, pair of apicolateral spots on clypeus, labrum, mandible, gena, pair of large spots anterolaterally and pair of transverse spots posteriorly on pronotum, tegula and two pairs of anterolateral spots on metasomal terga II and III; the following orange: metasomal tergum I apically, terga II and basal two-thirds of tergum III, except for two pairs of yellow spots, sternum I apically and sterna II – III. Antenna dark brown dorsally, darker toward apex; scape laterally and ventrally, and pedicel and flagellomere I ventrally yellowish brown; others light brown. Fore coxa anteriorly, all trochanters apically, fore femur and tibia anteriorly, fore tarsus, mid coxa, dorsoapical half of mid femur, mid and hind tibiae basally light yellow; mid tibia except basally, mid tarsus, hind femur apically, hind tibia ventromedially and hind tarsus orange-red; remainder dark brown. Head in frontal view 1.5 times as broad as long. Ratio of UID: MID: LID = 8.4: 10: 4.9. Eye 0.53 times half of MID. Ratio of POD: OOD = 1: 0.26; POD 3.8 times as long as width of anterior ocellus. OVD / POD = 0.33. Gena in lateral view 0.3 times eye width. Clypeus 1.8 times as broad as long. Length ratio of scape, pedicel, and flagellomeres I and II = 14.3: 6.7: 10: 10.5; flagellomere I 2.8 times as long as thick, 0.38 times as long as UID. Propodeum 0.99 times as long as broad. Length ratio of fore wing cross vein 2 r-rs and last three abscissae of vein Rs = 10: 5.7: 16.5: 25.2. SMC 3 1.4 times as long as SMC 2 on vein M, 3.0 times as long as SMC 2 on vein Rs, narrowed on vein Rs by 0.88 times its own length on vein M. Longer spur of hind tibia 0.67 times as long as basitarsus.	en	Akira Shimizu, Raymond Wahis, Anthony C. Harris, James P. Pitts (2016): An extraordinary new genus and species of spider wasps (Insecta: Hymenoptera: Pompilidae) from Southeast Asia. Journal of Natural History 50: 1549-1559, DOI: 10.1080/00222933.2016.1155781
3E641D3CFFC6FFFFFEBBFF04FE45FD8F.taxon	materials_examined	Type material Holotype, ♀, ‘ Penong MALAYA Batu Feringgi Catchment area 29. iii. 1959 H. T. Pagden ’ ‘ on tree trunk close to nests of Trigona atripes & Trigona collina’ ‘ Pres by Com Inst Ent B M 1960 - 3 ' ‘ C. I. E. COLL. NO. 16593 ' ‘ H. T. Pagden Coll. B. M. 1971 - 461 ' (NHMUK). Paratype, 1 ♀, Thailand, Phetchabun, Nam Nao NP, Checkpoint 16 º 43.687 ' N, 101 º 33.754 ' E, 924 m, Malaise trap, 1 – 8 March 2007, Leng Janteab leg., T 2284 (USU).	en	Akira Shimizu, Raymond Wahis, Anthony C. Harris, James P. Pitts (2016): An extraordinary new genus and species of spider wasps (Insecta: Hymenoptera: Pompilidae) from Southeast Asia. Journal of Natural History 50: 1549-1559, DOI: 10.1080/00222933.2016.1155781
3E641D3CFFC6FFFFFEBBFF04FE45FD8F.taxon	etymology	Etymology The species is named after Mr. M. C. Day, who showed AS the holotype specimen deposited at the NHMUK.	en	Akira Shimizu, Raymond Wahis, Anthony C. Harris, James P. Pitts (2016): An extraordinary new genus and species of spider wasps (Insecta: Hymenoptera: Pompilidae) from Southeast Asia. Journal of Natural History 50: 1549-1559, DOI: 10.1080/00222933.2016.1155781
