taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
3E6A87DAFFCEFF984993FDDBFE94FB4E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392019/files/figure.png	https://doi.org/10.5281/zenodo.8392019	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCEFF984993FDDBFE94FB4E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392021/files/figure.png	https://doi.org/10.5281/zenodo.8392021	FIGURES 5–8. Lethrus (Mesolethrus) species, habitus. 5, L. microbuccis Ballion, 1871 (9 km S Ai-Kol); 6, L. elisae Nikolajev, 2001 (Shurob); 7–8, L. dyonini A. Shapovalov & Bagaturov, new species (environs of Tashsay), 7—holotype, 8—allotype. 5–7—males, 8—female.	FIGURES 5–8. Lethrus (Mesolethrus) species, habitus. 5, L. microbuccis Ballion, 1871 (9 km S Ai-Kol); 6, L. elisae Nikolajev, 2001 (Shurob); 7–8, L. dyonini A. Shapovalov & Bagaturov, new species (environs of Tashsay), 7—holotype, 8—allotype. 5–7—males, 8—female.	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCEFF984993FDDBFE94FB4E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392033/files/figure.png	https://doi.org/10.5281/zenodo.8392033	FIGURE 45. Lethrus (Mesolethrus) microbuccis Ballion, 1871, habitus (Kyrgyzstan, between Isfana and Golbo; photograph in nature by D. Milko).	FIGURE 45. Lethrus (Mesolethrus) microbuccis Ballion, 1871, habitus (Kyrgyzstan, between Isfana and Golbo; photograph in nature by D. Milko).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCEFF984993FDDBFE94FB4E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392027/files/figure.png	https://doi.org/10.5281/zenodo.8392027	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCEFF984993FA85FC1BF93E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392019/files/figure.png	https://doi.org/10.5281/zenodo.8392019	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCEFF984993FA85FC1BF93E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392027/files/figure.png	https://doi.org/10.5281/zenodo.8392027	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCEFF984993FA85FC1BF93E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392025/files/figure.png	https://doi.org/10.5281/zenodo.8392025	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCEFF9B4993F955FDBFF878.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392019/files/figure.png	https://doi.org/10.5281/zenodo.8392019	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCEFF9B4993F955FDBFF878.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392025/files/figure.png	https://doi.org/10.5281/zenodo.8392025	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCEFF9B4993F955FDBFF878.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392027/files/figure.png	https://doi.org/10.5281/zenodo.8392027	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCEFF9B4993F955FDBFF878.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392021/files/figure.png	https://doi.org/10.5281/zenodo.8392021	FIGURES 5–8. Lethrus (Mesolethrus) species, habitus. 5, L. microbuccis Ballion, 1871 (9 km S Ai-Kol); 6, L. elisae Nikolajev, 2001 (Shurob); 7–8, L. dyonini A. Shapovalov & Bagaturov, new species (environs of Tashsay), 7—holotype, 8—allotype. 5–7—males, 8—female.	FIGURES 5–8. Lethrus (Mesolethrus) species, habitus. 5, L. microbuccis Ballion, 1871 (9 km S Ai-Kol); 6, L. elisae Nikolajev, 2001 (Shurob); 7–8, L. dyonini A. Shapovalov & Bagaturov, new species (environs of Tashsay), 7—holotype, 8—allotype. 5–7—males, 8—female.	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCEFF9B4993F955FDBFF878.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392031/files/figure.png	https://doi.org/10.5281/zenodo.8392031	FIGURE 44. Map of distribution Lethrus (Mesolethrus) species: 1, Tashsay env.; 2, south from Belovutsoy River; 3, Khujand; 4, Golbo; 5, 9 km S Ai-Kol; 6, Shahriston env.; 7, Zaamin env.; 8, 28 km SW Zaamin; 9, Sudzhina; 10, Mullabulak; 11, Asmansay; 12, Jangiqishlak env.; 13, 11 km southeast of Djush; 14, Samarkand; 15, 65 km on Samarkand to Qarshi road; 16, Qaynarbuloq; 17, Ruskishlak; 18, 30 km East to Kitab, Qashqadaryo Riv.; 19, Qamar; 20, Guldara; 21, Qarshi; 22, Gumbulok; 23, Okravot Pass; 24, Shurob; 25, Boysun; 26, 20 km W Oqmachit; 27, Chagam; 28, Oqmachit; 29, Shahrinav; 30, Dushanbe env.; 31, Dahanakiik; 32, Yavon; 33, Chormagzak Pass.; 34, Fayzobod; 35, Nurek; 36, Pashtamazar; 37, Kulob. 1–2, 7–8, 10–28 Uzbekistan; 4–5 Kyrgyzstan, 3, 6, 9, 29–37 Tajikistan. Red circles—localities of L. dyonini A. Shapovalov & Bagaturov, new species (1—type locality: Tashsay); circles with green colour—localities of L. microbuccis Ballion, 1871 (3—type locality: Khujand); circles with white colour—localities of L. shakhristanicus Nikolajev, 2003 (6—type locality: Shahriston env.); circles with blue colour—localities of L. nuratavicus Nikolajev, 1987 (11—type locality: Asmansay); circles with yellow colour— localities of L. elisae Nikolajev, 2001 (24—type locality: Shurob); circles with orange colour—localities of L. inermis Reitter, 1897 (37—type locality: Kulab?); orange asterisk—Kyrgyzstan, Taldyk (erroneous record of L. inermis from Protzenko 1968).	FIGURE 44. Map of distribution Lethrus (Mesolethrus) species: 1, Tashsay env.; 2, south from Belovutsoy River; 3, Khujand; 4, Golbo; 5, 9 km S Ai-Kol; 6, Shahriston env.; 7, Zaamin env.; 8, 28 km SW Zaamin; 9, Sudzhina; 10, Mullabulak; 11, Asmansay; 12, Jangiqishlak env.; 13, 11 km southeast of Djush; 14, Samarkand; 15, 65 km on Samarkand to Qarshi road; 16, Qaynarbuloq; 17, Ruskishlak; 18, 30 km East to Kitab, Qashqadaryo Riv.; 19, Qamar; 20, Guldara; 21, Qarshi; 22, Gumbulok; 23, Okravot Pass; 24, Shurob; 25, Boysun; 26, 20 km W Oqmachit; 27, Chagam; 28, Oqmachit; 29, Shahrinav; 30, Dushanbe env.; 31, Dahanakiik; 32, Yavon; 33, Chormagzak Pass.; 34, Fayzobod; 35, Nurek; 36, Pashtamazar; 37, Kulob. 1–2, 7–8, 10–28 Uzbekistan; 4–5 Kyrgyzstan, 3, 6, 9, 29–37 Tajikistan. Red circles—localities of L. dyonini A. Shapovalov & Bagaturov, new species (1—type locality: Tashsay); circles with green colour—localities of L. microbuccis Ballion, 1871 (3—type locality: Khujand); circles with white colour—localities of L. shakhristanicus Nikolajev, 2003 (6—type locality: Shahriston env.); circles with blue colour—localities of L. nuratavicus Nikolajev, 1987 (11—type locality: Asmansay); circles with yellow colour— localities of L. elisae Nikolajev, 2001 (24—type locality: Shurob); circles with orange colour—localities of L. inermis Reitter, 1897 (37—type locality: Kulab?); orange asterisk—Kyrgyzstan, Taldyk (erroneous record of L. inermis from Protzenko 1968).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCBFF9D4993FD3FFD29F8D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392019/files/figure.png	https://doi.org/10.5281/zenodo.8392019	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCBFF9D4993FD3FFD29F8D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392025/files/figure.png	https://doi.org/10.5281/zenodo.8392025	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCBFF9D4993FD3FFD29F8D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392027/files/figure.png	https://doi.org/10.5281/zenodo.8392027	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCBFF9D4993FD3FFD29F8D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392031/files/figure.png	https://doi.org/10.5281/zenodo.8392031	FIGURE 44. Map of distribution Lethrus (Mesolethrus) species: 1, Tashsay env.; 2, south from Belovutsoy River; 3, Khujand; 4, Golbo; 5, 9 km S Ai-Kol; 6, Shahriston env.; 7, Zaamin env.; 8, 28 km SW Zaamin; 9, Sudzhina; 10, Mullabulak; 11, Asmansay; 12, Jangiqishlak env.; 13, 11 km southeast of Djush; 14, Samarkand; 15, 65 km on Samarkand to Qarshi road; 16, Qaynarbuloq; 17, Ruskishlak; 18, 30 km East to Kitab, Qashqadaryo Riv.; 19, Qamar; 20, Guldara; 21, Qarshi; 22, Gumbulok; 23, Okravot Pass; 24, Shurob; 25, Boysun; 26, 20 km W Oqmachit; 27, Chagam; 28, Oqmachit; 29, Shahrinav; 30, Dushanbe env.; 31, Dahanakiik; 32, Yavon; 33, Chormagzak Pass.; 34, Fayzobod; 35, Nurek; 36, Pashtamazar; 37, Kulob. 1–2, 7–8, 10–28 Uzbekistan; 4–5 Kyrgyzstan, 3, 6, 9, 29–37 Tajikistan. Red circles—localities of L. dyonini A. Shapovalov & Bagaturov, new species (1—type locality: Tashsay); circles with green colour—localities of L. microbuccis Ballion, 1871 (3—type locality: Khujand); circles with white colour—localities of L. shakhristanicus Nikolajev, 2003 (6—type locality: Shahriston env.); circles with blue colour—localities of L. nuratavicus Nikolajev, 1987 (11—type locality: Asmansay); circles with yellow colour— localities of L. elisae Nikolajev, 2001 (24—type locality: Shurob); circles with orange colour—localities of L. inermis Reitter, 1897 (37—type locality: Kulab?); orange asterisk—Kyrgyzstan, Taldyk (erroneous record of L. inermis from Protzenko 1968).	FIGURE 44. Map of distribution Lethrus (Mesolethrus) species: 1, Tashsay env.; 2, south from Belovutsoy River; 3, Khujand; 4, Golbo; 5, 9 km S Ai-Kol; 6, Shahriston env.; 7, Zaamin env.; 8, 28 km SW Zaamin; 9, Sudzhina; 10, Mullabulak; 11, Asmansay; 12, Jangiqishlak env.; 13, 11 km southeast of Djush; 14, Samarkand; 15, 65 km on Samarkand to Qarshi road; 16, Qaynarbuloq; 17, Ruskishlak; 18, 30 km East to Kitab, Qashqadaryo Riv.; 19, Qamar; 20, Guldara; 21, Qarshi; 22, Gumbulok; 23, Okravot Pass; 24, Shurob; 25, Boysun; 26, 20 km W Oqmachit; 27, Chagam; 28, Oqmachit; 29, Shahrinav; 30, Dushanbe env.; 31, Dahanakiik; 32, Yavon; 33, Chormagzak Pass.; 34, Fayzobod; 35, Nurek; 36, Pashtamazar; 37, Kulob. 1–2, 7–8, 10–28 Uzbekistan; 4–5 Kyrgyzstan, 3, 6, 9, 29–37 Tajikistan. Red circles—localities of L. dyonini A. Shapovalov & Bagaturov, new species (1—type locality: Tashsay); circles with green colour—localities of L. microbuccis Ballion, 1871 (3—type locality: Khujand); circles with white colour—localities of L. shakhristanicus Nikolajev, 2003 (6—type locality: Shahriston env.); circles with blue colour—localities of L. nuratavicus Nikolajev, 1987 (11—type locality: Asmansay); circles with yellow colour— localities of L. elisae Nikolajev, 2001 (24—type locality: Shurob); circles with orange colour—localities of L. inermis Reitter, 1897 (37—type locality: Kulab?); orange asterisk—Kyrgyzstan, Taldyk (erroneous record of L. inermis from Protzenko 1968).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCBFF9D4993FD3FFD29F8D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392037/files/figure.png	https://doi.org/10.5281/zenodo.8392037	FIGURE 46. Locality of L. microbuccis Ballion, 1871, L. shakhristanicus Nikolajev, 2003 and L. nuratavicus Nikolajev, 1987, 28 km southwest of Zaamin, Malguzar Mountain Range, Jizzax Region, Uzbekistan, photograph by S. Dyonin.	FIGURE 46. Locality of L. microbuccis Ballion, 1871, L. shakhristanicus Nikolajev, 2003 and L. nuratavicus Nikolajev, 1987, 28 km southwest of Zaamin, Malguzar Mountain Range, Jizzax Region, Uzbekistan, photograph by S. Dyonin.	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCBFF9D4993FD3FFD29F8D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392041/files/figure.png	https://doi.org/10.5281/zenodo.8392041	FIGURE 47. Locality of L. shakhristanicus and L.microbuccis (green lowland infront of the more distant mountains) southwards Ai-Kol, Turkestan Mountain Range, Batken Reg., Kyrgyzstan, photograph by A. Zhdanko.	FIGURE 47. Locality of L. shakhristanicus and L.microbuccis (green lowland infront of the more distant mountains) southwards Ai-Kol, Turkestan Mountain Range, Batken Reg., Kyrgyzstan, photograph by A. Zhdanko.	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCBFF9D4993FF27FBAAFD97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392019/files/figure.png	https://doi.org/10.5281/zenodo.8392019	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCBFF9D4993FF27FBAAFD97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392021/files/figure.png	https://doi.org/10.5281/zenodo.8392021	FIGURES 5–8. Lethrus (Mesolethrus) species, habitus. 5, L. microbuccis Ballion, 1871 (9 km S Ai-Kol); 6, L. elisae Nikolajev, 2001 (Shurob); 7–8, L. dyonini A. Shapovalov & Bagaturov, new species (environs of Tashsay), 7—holotype, 8—allotype. 5–7—males, 8—female.	FIGURES 5–8. Lethrus (Mesolethrus) species, habitus. 5, L. microbuccis Ballion, 1871 (9 km S Ai-Kol); 6, L. elisae Nikolajev, 2001 (Shurob); 7–8, L. dyonini A. Shapovalov & Bagaturov, new species (environs of Tashsay), 7—holotype, 8—allotype. 5–7—males, 8—female.	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCBFF9D4993FF27FBAAFD97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392027/files/figure.png	https://doi.org/10.5281/zenodo.8392027	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCBFF9D4993FF27FBAAFD97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392025/files/figure.png	https://doi.org/10.5281/zenodo.8392025	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCAFF9C4993FF27FC63FBB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392019/files/figure.png	https://doi.org/10.5281/zenodo.8392019	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCAFF9C4993FF27FC63FBB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392025/files/figure.png	https://doi.org/10.5281/zenodo.8392025	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCAFF9C4993FF27FC63FBB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392027/files/figure.png	https://doi.org/10.5281/zenodo.8392027	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCAFF9C4993FF27FC63FBB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392043/files/figure.png	https://doi.org/10.5281/zenodo.8392043	FIGURE 48. Locality of L. dyonini A. Shapovalov & Bagaturov, new species, environs of Tashsay, western foothills of Chatkal Mountain Range, Uzbekistan, photograph by S. Dyonin.	FIGURE 48. Locality of L. dyonini A. Shapovalov & Bagaturov, new species, environs of Tashsay, western foothills of Chatkal Mountain Range, Uzbekistan, photograph by S. Dyonin.	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCAFF9C4993FF27FC63FBB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392031/files/figure.png	https://doi.org/10.5281/zenodo.8392031	FIGURE 44. Map of distribution Lethrus (Mesolethrus) species: 1, Tashsay env.; 2, south from Belovutsoy River; 3, Khujand; 4, Golbo; 5, 9 km S Ai-Kol; 6, Shahriston env.; 7, Zaamin env.; 8, 28 km SW Zaamin; 9, Sudzhina; 10, Mullabulak; 11, Asmansay; 12, Jangiqishlak env.; 13, 11 km southeast of Djush; 14, Samarkand; 15, 65 km on Samarkand to Qarshi road; 16, Qaynarbuloq; 17, Ruskishlak; 18, 30 km East to Kitab, Qashqadaryo Riv.; 19, Qamar; 20, Guldara; 21, Qarshi; 22, Gumbulok; 23, Okravot Pass; 24, Shurob; 25, Boysun; 26, 20 km W Oqmachit; 27, Chagam; 28, Oqmachit; 29, Shahrinav; 30, Dushanbe env.; 31, Dahanakiik; 32, Yavon; 33, Chormagzak Pass.; 34, Fayzobod; 35, Nurek; 36, Pashtamazar; 37, Kulob. 1–2, 7–8, 10–28 Uzbekistan; 4–5 Kyrgyzstan, 3, 6, 9, 29–37 Tajikistan. Red circles—localities of L. dyonini A. Shapovalov & Bagaturov, new species (1—type locality: Tashsay); circles with green colour—localities of L. microbuccis Ballion, 1871 (3—type locality: Khujand); circles with white colour—localities of L. shakhristanicus Nikolajev, 2003 (6—type locality: Shahriston env.); circles with blue colour—localities of L. nuratavicus Nikolajev, 1987 (11—type locality: Asmansay); circles with yellow colour— localities of L. elisae Nikolajev, 2001 (24—type locality: Shurob); circles with orange colour—localities of L. inermis Reitter, 1897 (37—type locality: Kulab?); orange asterisk—Kyrgyzstan, Taldyk (erroneous record of L. inermis from Protzenko 1968).	FIGURE 44. Map of distribution Lethrus (Mesolethrus) species: 1, Tashsay env.; 2, south from Belovutsoy River; 3, Khujand; 4, Golbo; 5, 9 km S Ai-Kol; 6, Shahriston env.; 7, Zaamin env.; 8, 28 km SW Zaamin; 9, Sudzhina; 10, Mullabulak; 11, Asmansay; 12, Jangiqishlak env.; 13, 11 km southeast of Djush; 14, Samarkand; 15, 65 km on Samarkand to Qarshi road; 16, Qaynarbuloq; 17, Ruskishlak; 18, 30 km East to Kitab, Qashqadaryo Riv.; 19, Qamar; 20, Guldara; 21, Qarshi; 22, Gumbulok; 23, Okravot Pass; 24, Shurob; 25, Boysun; 26, 20 km W Oqmachit; 27, Chagam; 28, Oqmachit; 29, Shahrinav; 30, Dushanbe env.; 31, Dahanakiik; 32, Yavon; 33, Chormagzak Pass.; 34, Fayzobod; 35, Nurek; 36, Pashtamazar; 37, Kulob. 1–2, 7–8, 10–28 Uzbekistan; 4–5 Kyrgyzstan, 3, 6, 9, 29–37 Tajikistan. Red circles—localities of L. dyonini A. Shapovalov & Bagaturov, new species (1—type locality: Tashsay); circles with green colour—localities of L. microbuccis Ballion, 1871 (3—type locality: Khujand); circles with white colour—localities of L. shakhristanicus Nikolajev, 2003 (6—type locality: Shahriston env.); circles with blue colour—localities of L. nuratavicus Nikolajev, 1987 (11—type locality: Asmansay); circles with yellow colour— localities of L. elisae Nikolajev, 2001 (24—type locality: Shurob); circles with orange colour—localities of L. inermis Reitter, 1897 (37—type locality: Kulab?); orange asterisk—Kyrgyzstan, Taldyk (erroneous record of L. inermis from Protzenko 1968).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCAFF9C4993FF27FC63FBB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392037/files/figure.png	https://doi.org/10.5281/zenodo.8392037	FIGURE 46. Locality of L. microbuccis Ballion, 1871, L. shakhristanicus Nikolajev, 2003 and L. nuratavicus Nikolajev, 1987, 28 km southwest of Zaamin, Malguzar Mountain Range, Jizzax Region, Uzbekistan, photograph by S. Dyonin.	FIGURE 46. Locality of L. microbuccis Ballion, 1871, L. shakhristanicus Nikolajev, 2003 and L. nuratavicus Nikolajev, 1987, 28 km southwest of Zaamin, Malguzar Mountain Range, Jizzax Region, Uzbekistan, photograph by S. Dyonin.	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCAFF9F4993FBDCFEBFF9DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392021/files/figure.png	https://doi.org/10.5281/zenodo.8392021	FIGURES 5–8. Lethrus (Mesolethrus) species, habitus. 5, L. microbuccis Ballion, 1871 (9 km S Ai-Kol); 6, L. elisae Nikolajev, 2001 (Shurob); 7–8, L. dyonini A. Shapovalov & Bagaturov, new species (environs of Tashsay), 7—holotype, 8—allotype. 5–7—males, 8—female.	FIGURES 5–8. Lethrus (Mesolethrus) species, habitus. 5, L. microbuccis Ballion, 1871 (9 km S Ai-Kol); 6, L. elisae Nikolajev, 2001 (Shurob); 7–8, L. dyonini A. Shapovalov & Bagaturov, new species (environs of Tashsay), 7—holotype, 8—allotype. 5–7—males, 8—female.	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCAFF9F4993FBDCFEBFF9DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392025/files/figure.png	https://doi.org/10.5281/zenodo.8392025	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCAFF9F4993FBDCFEBFF9DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392027/files/figure.png	https://doi.org/10.5281/zenodo.8392027	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCAFF9F4993FBDCFEBFF9DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392033/files/figure.png	https://doi.org/10.5281/zenodo.8392033	FIGURE 45. Lethrus (Mesolethrus) microbuccis Ballion, 1871, habitus (Kyrgyzstan, between Isfana and Golbo; photograph in nature by D. Milko).	FIGURE 45. Lethrus (Mesolethrus) microbuccis Ballion, 1871, habitus (Kyrgyzstan, between Isfana and Golbo; photograph in nature by D. Milko).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCAFF9F4993FBDCFEBFF9DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392019/files/figure.png	https://doi.org/10.5281/zenodo.8392019	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCAFF9F4993FBDCFEBFF9DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392031/files/figure.png	https://doi.org/10.5281/zenodo.8392031	FIGURE 44. Map of distribution Lethrus (Mesolethrus) species: 1, Tashsay env.; 2, south from Belovutsoy River; 3, Khujand; 4, Golbo; 5, 9 km S Ai-Kol; 6, Shahriston env.; 7, Zaamin env.; 8, 28 km SW Zaamin; 9, Sudzhina; 10, Mullabulak; 11, Asmansay; 12, Jangiqishlak env.; 13, 11 km southeast of Djush; 14, Samarkand; 15, 65 km on Samarkand to Qarshi road; 16, Qaynarbuloq; 17, Ruskishlak; 18, 30 km East to Kitab, Qashqadaryo Riv.; 19, Qamar; 20, Guldara; 21, Qarshi; 22, Gumbulok; 23, Okravot Pass; 24, Shurob; 25, Boysun; 26, 20 km W Oqmachit; 27, Chagam; 28, Oqmachit; 29, Shahrinav; 30, Dushanbe env.; 31, Dahanakiik; 32, Yavon; 33, Chormagzak Pass.; 34, Fayzobod; 35, Nurek; 36, Pashtamazar; 37, Kulob. 1–2, 7–8, 10–28 Uzbekistan; 4–5 Kyrgyzstan, 3, 6, 9, 29–37 Tajikistan. Red circles—localities of L. dyonini A. Shapovalov & Bagaturov, new species (1—type locality: Tashsay); circles with green colour—localities of L. microbuccis Ballion, 1871 (3—type locality: Khujand); circles with white colour—localities of L. shakhristanicus Nikolajev, 2003 (6—type locality: Shahriston env.); circles with blue colour—localities of L. nuratavicus Nikolajev, 1987 (11—type locality: Asmansay); circles with yellow colour— localities of L. elisae Nikolajev, 2001 (24—type locality: Shurob); circles with orange colour—localities of L. inermis Reitter, 1897 (37—type locality: Kulab?); orange asterisk—Kyrgyzstan, Taldyk (erroneous record of L. inermis from Protzenko 1968).	FIGURE 44. Map of distribution Lethrus (Mesolethrus) species: 1, Tashsay env.; 2, south from Belovutsoy River; 3, Khujand; 4, Golbo; 5, 9 km S Ai-Kol; 6, Shahriston env.; 7, Zaamin env.; 8, 28 km SW Zaamin; 9, Sudzhina; 10, Mullabulak; 11, Asmansay; 12, Jangiqishlak env.; 13, 11 km southeast of Djush; 14, Samarkand; 15, 65 km on Samarkand to Qarshi road; 16, Qaynarbuloq; 17, Ruskishlak; 18, 30 km East to Kitab, Qashqadaryo Riv.; 19, Qamar; 20, Guldara; 21, Qarshi; 22, Gumbulok; 23, Okravot Pass; 24, Shurob; 25, Boysun; 26, 20 km W Oqmachit; 27, Chagam; 28, Oqmachit; 29, Shahrinav; 30, Dushanbe env.; 31, Dahanakiik; 32, Yavon; 33, Chormagzak Pass.; 34, Fayzobod; 35, Nurek; 36, Pashtamazar; 37, Kulob. 1–2, 7–8, 10–28 Uzbekistan; 4–5 Kyrgyzstan, 3, 6, 9, 29–37 Tajikistan. Red circles—localities of L. dyonini A. Shapovalov & Bagaturov, new species (1—type locality: Tashsay); circles with green colour—localities of L. microbuccis Ballion, 1871 (3—type locality: Khujand); circles with white colour—localities of L. shakhristanicus Nikolajev, 2003 (6—type locality: Shahriston env.); circles with blue colour—localities of L. nuratavicus Nikolajev, 1987 (11—type locality: Asmansay); circles with yellow colour— localities of L. elisae Nikolajev, 2001 (24—type locality: Shurob); circles with orange colour—localities of L. inermis Reitter, 1897 (37—type locality: Kulab?); orange asterisk—Kyrgyzstan, Taldyk (erroneous record of L. inermis from Protzenko 1968).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCAFF9F4993FBDCFEBFF9DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392037/files/figure.png	https://doi.org/10.5281/zenodo.8392037	FIGURE 46. Locality of L. microbuccis Ballion, 1871, L. shakhristanicus Nikolajev, 2003 and L. nuratavicus Nikolajev, 1987, 28 km southwest of Zaamin, Malguzar Mountain Range, Jizzax Region, Uzbekistan, photograph by S. Dyonin.	FIGURE 46. Locality of L. microbuccis Ballion, 1871, L. shakhristanicus Nikolajev, 2003 and L. nuratavicus Nikolajev, 1987, 28 km southwest of Zaamin, Malguzar Mountain Range, Jizzax Region, Uzbekistan, photograph by S. Dyonin.	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFCAFF9F4993FBDCFEBFF9DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392041/files/figure.png	https://doi.org/10.5281/zenodo.8392041	FIGURE 47. Locality of L. shakhristanicus and L.microbuccis (green lowland infront of the more distant mountains) southwards Ai-Kol, Turkestan Mountain Range, Batken Reg., Kyrgyzstan, photograph by A. Zhdanko.	FIGURE 47. Locality of L. shakhristanicus and L.microbuccis (green lowland infront of the more distant mountains) southwards Ai-Kol, Turkestan Mountain Range, Batken Reg., Kyrgyzstan, photograph by A. Zhdanko.	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFC9FF934993F976FCFDFDCB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392021/files/figure.png	https://doi.org/10.5281/zenodo.8392021	FIGURES 5–8. Lethrus (Mesolethrus) species, habitus. 5, L. microbuccis Ballion, 1871 (9 km S Ai-Kol); 6, L. elisae Nikolajev, 2001 (Shurob); 7–8, L. dyonini A. Shapovalov & Bagaturov, new species (environs of Tashsay), 7—holotype, 8—allotype. 5–7—males, 8—female.	FIGURES 5–8. Lethrus (Mesolethrus) species, habitus. 5, L. microbuccis Ballion, 1871 (9 km S Ai-Kol); 6, L. elisae Nikolajev, 2001 (Shurob); 7–8, L. dyonini A. Shapovalov & Bagaturov, new species (environs of Tashsay), 7—holotype, 8—allotype. 5–7—males, 8—female.	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFC9FF934993F976FCFDFDCB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392025/files/figure.png	https://doi.org/10.5281/zenodo.8392025	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFC9FF934993F976FCFDFDCB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392027/files/figure.png	https://doi.org/10.5281/zenodo.8392027	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFC9FF934993F976FCFDFDCB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392019/files/figure.png	https://doi.org/10.5281/zenodo.8392019	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFC9FF934993F976FCFDFDCB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392031/files/figure.png	https://doi.org/10.5281/zenodo.8392031	FIGURE 44. Map of distribution Lethrus (Mesolethrus) species: 1, Tashsay env.; 2, south from Belovutsoy River; 3, Khujand; 4, Golbo; 5, 9 km S Ai-Kol; 6, Shahriston env.; 7, Zaamin env.; 8, 28 km SW Zaamin; 9, Sudzhina; 10, Mullabulak; 11, Asmansay; 12, Jangiqishlak env.; 13, 11 km southeast of Djush; 14, Samarkand; 15, 65 km on Samarkand to Qarshi road; 16, Qaynarbuloq; 17, Ruskishlak; 18, 30 km East to Kitab, Qashqadaryo Riv.; 19, Qamar; 20, Guldara; 21, Qarshi; 22, Gumbulok; 23, Okravot Pass; 24, Shurob; 25, Boysun; 26, 20 km W Oqmachit; 27, Chagam; 28, Oqmachit; 29, Shahrinav; 30, Dushanbe env.; 31, Dahanakiik; 32, Yavon; 33, Chormagzak Pass.; 34, Fayzobod; 35, Nurek; 36, Pashtamazar; 37, Kulob. 1–2, 7–8, 10–28 Uzbekistan; 4–5 Kyrgyzstan, 3, 6, 9, 29–37 Tajikistan. Red circles—localities of L. dyonini A. Shapovalov & Bagaturov, new species (1—type locality: Tashsay); circles with green colour—localities of L. microbuccis Ballion, 1871 (3—type locality: Khujand); circles with white colour—localities of L. shakhristanicus Nikolajev, 2003 (6—type locality: Shahriston env.); circles with blue colour—localities of L. nuratavicus Nikolajev, 1987 (11—type locality: Asmansay); circles with yellow colour— localities of L. elisae Nikolajev, 2001 (24—type locality: Shurob); circles with orange colour—localities of L. inermis Reitter, 1897 (37—type locality: Kulab?); orange asterisk—Kyrgyzstan, Taldyk (erroneous record of L. inermis from Protzenko 1968).	FIGURE 44. Map of distribution Lethrus (Mesolethrus) species: 1, Tashsay env.; 2, south from Belovutsoy River; 3, Khujand; 4, Golbo; 5, 9 km S Ai-Kol; 6, Shahriston env.; 7, Zaamin env.; 8, 28 km SW Zaamin; 9, Sudzhina; 10, Mullabulak; 11, Asmansay; 12, Jangiqishlak env.; 13, 11 km southeast of Djush; 14, Samarkand; 15, 65 km on Samarkand to Qarshi road; 16, Qaynarbuloq; 17, Ruskishlak; 18, 30 km East to Kitab, Qashqadaryo Riv.; 19, Qamar; 20, Guldara; 21, Qarshi; 22, Gumbulok; 23, Okravot Pass; 24, Shurob; 25, Boysun; 26, 20 km W Oqmachit; 27, Chagam; 28, Oqmachit; 29, Shahrinav; 30, Dushanbe env.; 31, Dahanakiik; 32, Yavon; 33, Chormagzak Pass.; 34, Fayzobod; 35, Nurek; 36, Pashtamazar; 37, Kulob. 1–2, 7–8, 10–28 Uzbekistan; 4–5 Kyrgyzstan, 3, 6, 9, 29–37 Tajikistan. Red circles—localities of L. dyonini A. Shapovalov & Bagaturov, new species (1—type locality: Tashsay); circles with green colour—localities of L. microbuccis Ballion, 1871 (3—type locality: Khujand); circles with white colour—localities of L. shakhristanicus Nikolajev, 2003 (6—type locality: Shahriston env.); circles with blue colour—localities of L. nuratavicus Nikolajev, 1987 (11—type locality: Asmansay); circles with yellow colour— localities of L. elisae Nikolajev, 2001 (24—type locality: Shurob); circles with orange colour—localities of L. inermis Reitter, 1897 (37—type locality: Kulab?); orange asterisk—Kyrgyzstan, Taldyk (erroneous record of L. inermis from Protzenko 1968).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFC4FF944993FB1DFAC1FB9F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392021/files/figure.png	https://doi.org/10.5281/zenodo.8392021	FIGURES 5–8. Lethrus (Mesolethrus) species, habitus. 5, L. microbuccis Ballion, 1871 (9 km S Ai-Kol); 6, L. elisae Nikolajev, 2001 (Shurob); 7–8, L. dyonini A. Shapovalov & Bagaturov, new species (environs of Tashsay), 7—holotype, 8—allotype. 5–7—males, 8—female.	FIGURES 5–8. Lethrus (Mesolethrus) species, habitus. 5, L. microbuccis Ballion, 1871 (9 km S Ai-Kol); 6, L. elisae Nikolajev, 2001 (Shurob); 7–8, L. dyonini A. Shapovalov & Bagaturov, new species (environs of Tashsay), 7—holotype, 8—allotype. 5–7—males, 8—female.	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFC4FF944993FB1DFAC1FB9F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392025/files/figure.png	https://doi.org/10.5281/zenodo.8392025	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFC4FF944993FB1DFAC1FB9F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392027/files/figure.png	https://doi.org/10.5281/zenodo.8392027	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFC4FF944993FB1DFAC1FB9F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392031/files/figure.png	https://doi.org/10.5281/zenodo.8392031	FIGURE 44. Map of distribution Lethrus (Mesolethrus) species: 1, Tashsay env.; 2, south from Belovutsoy River; 3, Khujand; 4, Golbo; 5, 9 km S Ai-Kol; 6, Shahriston env.; 7, Zaamin env.; 8, 28 km SW Zaamin; 9, Sudzhina; 10, Mullabulak; 11, Asmansay; 12, Jangiqishlak env.; 13, 11 km southeast of Djush; 14, Samarkand; 15, 65 km on Samarkand to Qarshi road; 16, Qaynarbuloq; 17, Ruskishlak; 18, 30 km East to Kitab, Qashqadaryo Riv.; 19, Qamar; 20, Guldara; 21, Qarshi; 22, Gumbulok; 23, Okravot Pass; 24, Shurob; 25, Boysun; 26, 20 km W Oqmachit; 27, Chagam; 28, Oqmachit; 29, Shahrinav; 30, Dushanbe env.; 31, Dahanakiik; 32, Yavon; 33, Chormagzak Pass.; 34, Fayzobod; 35, Nurek; 36, Pashtamazar; 37, Kulob. 1–2, 7–8, 10–28 Uzbekistan; 4–5 Kyrgyzstan, 3, 6, 9, 29–37 Tajikistan. Red circles—localities of L. dyonini A. Shapovalov & Bagaturov, new species (1—type locality: Tashsay); circles with green colour—localities of L. microbuccis Ballion, 1871 (3—type locality: Khujand); circles with white colour—localities of L. shakhristanicus Nikolajev, 2003 (6—type locality: Shahriston env.); circles with blue colour—localities of L. nuratavicus Nikolajev, 1987 (11—type locality: Asmansay); circles with yellow colour— localities of L. elisae Nikolajev, 2001 (24—type locality: Shurob); circles with orange colour—localities of L. inermis Reitter, 1897 (37—type locality: Kulab?); orange asterisk—Kyrgyzstan, Taldyk (erroneous record of L. inermis from Protzenko 1968).	FIGURE 44. Map of distribution Lethrus (Mesolethrus) species: 1, Tashsay env.; 2, south from Belovutsoy River; 3, Khujand; 4, Golbo; 5, 9 km S Ai-Kol; 6, Shahriston env.; 7, Zaamin env.; 8, 28 km SW Zaamin; 9, Sudzhina; 10, Mullabulak; 11, Asmansay; 12, Jangiqishlak env.; 13, 11 km southeast of Djush; 14, Samarkand; 15, 65 km on Samarkand to Qarshi road; 16, Qaynarbuloq; 17, Ruskishlak; 18, 30 km East to Kitab, Qashqadaryo Riv.; 19, Qamar; 20, Guldara; 21, Qarshi; 22, Gumbulok; 23, Okravot Pass; 24, Shurob; 25, Boysun; 26, 20 km W Oqmachit; 27, Chagam; 28, Oqmachit; 29, Shahrinav; 30, Dushanbe env.; 31, Dahanakiik; 32, Yavon; 33, Chormagzak Pass.; 34, Fayzobod; 35, Nurek; 36, Pashtamazar; 37, Kulob. 1–2, 7–8, 10–28 Uzbekistan; 4–5 Kyrgyzstan, 3, 6, 9, 29–37 Tajikistan. Red circles—localities of L. dyonini A. Shapovalov & Bagaturov, new species (1—type locality: Tashsay); circles with green colour—localities of L. microbuccis Ballion, 1871 (3—type locality: Khujand); circles with white colour—localities of L. shakhristanicus Nikolajev, 2003 (6—type locality: Shahriston env.); circles with blue colour—localities of L. nuratavicus Nikolajev, 1987 (11—type locality: Asmansay); circles with yellow colour— localities of L. elisae Nikolajev, 2001 (24—type locality: Shurob); circles with orange colour—localities of L. inermis Reitter, 1897 (37—type locality: Kulab?); orange asterisk—Kyrgyzstan, Taldyk (erroneous record of L. inermis from Protzenko 1968).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFC4FF944993FB1DFAC1FB9F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392043/files/figure.png	https://doi.org/10.5281/zenodo.8392043	FIGURE 48. Locality of L. dyonini A. Shapovalov & Bagaturov, new species, environs of Tashsay, western foothills of Chatkal Mountain Range, Uzbekistan, photograph by S. Dyonin.	FIGURE 48. Locality of L. dyonini A. Shapovalov & Bagaturov, new species, environs of Tashsay, western foothills of Chatkal Mountain Range, Uzbekistan, photograph by S. Dyonin.	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFC1FF964993FAA3FA05FD35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392019/files/figure.png	https://doi.org/10.5281/zenodo.8392019	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFC1FF964993FAA3FA05FD35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392027/files/figure.png	https://doi.org/10.5281/zenodo.8392027	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFC1FF964993FAA3FA05FD35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392025/files/figure.png	https://doi.org/10.5281/zenodo.8392025	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFC1FF964993FAA3FA05FD35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392021/files/figure.png	https://doi.org/10.5281/zenodo.8392021	FIGURES 5–8. Lethrus (Mesolethrus) species, habitus. 5, L. microbuccis Ballion, 1871 (9 km S Ai-Kol); 6, L. elisae Nikolajev, 2001 (Shurob); 7–8, L. dyonini A. Shapovalov & Bagaturov, new species (environs of Tashsay), 7—holotype, 8—allotype. 5–7—males, 8—female.	FIGURES 5–8. Lethrus (Mesolethrus) species, habitus. 5, L. microbuccis Ballion, 1871 (9 km S Ai-Kol); 6, L. elisae Nikolajev, 2001 (Shurob); 7–8, L. dyonini A. Shapovalov & Bagaturov, new species (environs of Tashsay), 7—holotype, 8—allotype. 5–7—males, 8—female.	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
3E6A87DAFFC1FF964993FAA3FA05FD35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8392033/files/figure.png	https://doi.org/10.5281/zenodo.8392033	FIGURE 45. Lethrus (Mesolethrus) microbuccis Ballion, 1871, habitus (Kyrgyzstan, between Isfana and Golbo; photograph in nature by D. Milko).	FIGURE 45. Lethrus (Mesolethrus) microbuccis Ballion, 1871, habitus (Kyrgyzstan, between Isfana and Golbo; photograph in nature by D. Milko).	2023-09-28	Shapovalov, Andrey M.;Bagaturov, Mikhail F.		Zenodo	biologists	Shapovalov, Andrey M.;Bagaturov, Mikhail F.			
