identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
3D7587E2D17DFFFEFF5EF98819E50685.text	3D7587E2D17DFFFEFF5EF98819E50685.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tytthoscincus Linkem & Diesmos & Brown 2011	<div><p>Tytthoscincus gen. nov.</p> <p>Type species: Tytthoscincus hallieri (Lidth de Jeude, 1905).</p> <p>Definition: The clade comprising Tytthoscincus hallieri (Lidth de Juede, 1905) and all species that share a more recent common ancestor with Tytthoscincus hallieri than with Anomalopus verreauxii, Calyptotis scutirostrum, Coeranoscincus frontalis, Coggeria naufragus, Ctenotus taeniolatus, Eremiascincus richardsonii, Eulamprus quoyiii, Glaphyromorphus isolepis, Gnypetoscincus queenslandiae, Hemiergis decresiensis, Insulasaurus wrighti, Lerista lineata, Lipinia pulchella, Nangura spinosa, Notoscincus ornatus, Ophioscincus australis, Otosaurus cumingi, Papuascincus stanleyanus, Parvoscincus sisoni, Pinoyscincus jagori, Prasinohaema flavipes, Saiphos equalis, Scincella lateralis, and Sphenomorphus melanopogon.</p> <p>Etymology: From the Greek tytthos, meaning ‘small’ and the Latin scincus for lizard; the combination refers to the small sizes of the species in this genus. Suggested common name: diminutive Asian skink.</p> <p>Description: Tytthoscincus can be identified by the following characters: (1) body size diminutive, usually less than 45 mm SVL; (2) temporal scales small, same size and shape as lateral body scales (Fig. 5); and (3) digits small, toe IV slightly longer than, or equal to, toe III.</p> <p>Included species: Tytthoscincus aesculeticolus (Inger et al., 2001), Tytthoscincus atrigularis (Stejneger, 1905), Tytthoscincus biparietalis (Taylor, 1918), Tytthoscincus hallieri (Lidth de Juede, 1905), and Tytthoscincus parvus (Boulenger, 1897).</p> <p>Comment: This clade of diminutive species has unique features that diagnoses it from all other skinks of the Sphenomorphus group. Although we lack genetic data for Tytthoscincus biparentialis, we nonetheless include it in this genus because it shares the unique presence of divided parietal scales with Tytthoscincus hallieri. The diminutive skinks of Malaysia (Grismer, Ahmad &amp; Onn, 2009) should probably also be placed in this new genus, although we prefer to leave that decision in abeyance until a morphological and genetic examination of those taxa are complete. Tytthoscincus parvus (Boulenger, 1897) is one of three species of diminutive skinks described from Sulawesi Island. It is likely that the other diminutive species on Sulawesi, Sphenomorphus temmincki and Sphenomorphus textus are also part of Tytthoscincus. Future examination of temporal scales on small skinks in South-East Asia should reveal the species composition of Tytthoscincus.</p> </div>	https://treatment.plazi.org/id/3D7587E2D17DFFFEFF5EF98819E50685	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Linkem, Charles W.;Diesmos, Arvin C.;Brown, Rafe M.	Linkem, Charles W., Diesmos, Arvin C., Brown, Rafe M. (2011): Molecular systematics of the Philippine forest skinks (Squamata: Scincidae: Sphenomorphus): testing morphological hypotheses of interspecific relationships. Zoological Journal of the Linnean Society 163 (4): 1217-1243, DOI: 10.1111/j.1096-3642.2011.00747.x, URL: http://dx.doi.org/10.1111/j.1096-3642.2011.00747.x
3D7587E2D17DFFE0FCC0F93419CB0697.text	3D7587E2D17DFFE0FCC0F93419CB0697.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pinoyscincus Linkem & Diesmos & Brown 2011	<div><p>Pinoyscincus gen. nov.</p> <p>Type species: Pinoyscincus jagori (Peters, 1864).</p> <p>Definition: The clade comprising Pinoyscincus jagori (Peters, 1864) and all species that share a more recent common ancestor with Pinoyscincus jagori than with Anomalopus verreauxii, Calyptotis scutirostrum, Coeranoscincus frontalis, Coggeria naufragus, Ctenotus taeniolatus, Eremiascincus richardsonii, Eulamprus quoyii, Glaphyromorphus isolepis, Gnypetoscincus queenslandiae, Hemiergis decresiencsis, Insulasaurus wrighti, Lerista lineata, Lipinia pulchella, Lobulia elegans, Nangura spinosa, Notoscincus ornatus, Ophioscincus australis, Otosaurus cumingi, Papuascincus stanleyanus, Parvoscincus sisoni, Prasinohaema flavipes, Saiphos equalis, Scincella lateralis, Sphenomorphus melanopogon, and Tytthoscincus hallieri.</p> <p>Etymology: The word pinoy is a commonly used Tagalog term of endearment amongst Filipinos, referring to an individual Filipino or the nation as a whole. We use it here in conjunction with the Latin scincus, meaning lizard, to name a clade of skinks found on the Philippine Archipelago. Suggested common name: Filipino skinks.</p> <p>Description: Pinoyscincus can be identified by the following combination of characters: (1) body size medium to large (&gt; 42 mm SVL); (2) paravertebral scale rows 56–80; (3) midbody scale rows 30–44; and (4) subdigital lamellae 17–26. In addition to these scale characters, species in this genus share a unique morphology of the hemipenis. The main shaft of the hemipenis, before the bifurcation, is wide with a large bulbous lobe on each lateral side of the shaft (Fig. 6).</p> <p>Included species: Pinoyscincus abdictus (Brown &amp; Alcala, 1980), Pinoyscincus coxi (Taylor, 1915), Pinoyscincus jagori (Peters, 1864), Pinoyscincus llanosi (Taylor, 1919), and Pinoyscincus mindanensis (Taylor, 1922).</p> <p>Comment: This morphologically cohesive genus includes Brown &amp; Alcala’s (1980) Group 5 and Pinoyscincus mindanensis. All of these species are easily diagnosable among the Philippine skink fauna. The morphology of the hemipenis in this genus has been observed in Pinoyscincus mindanensis, Pinoyscincus abdictus, Pinoyscincus jagori, and Pinoyscincus llanosi and has not been observed in any other Philippine skink examined (Otosaurus cumingi, Insulasaurus arborens, Insulasaurus traanorum, Parvoscincus beyeri, Parvoscincus decipiens, Sphenomorphus fasciatus, Sphenomorphus variegatus). We have not examined the hemipenis of Sphenomorphus acutus or Sphenomorphus diwata yet to see if they share the Pinoyscincus character so we prefer to leave them incertae sedis until a more thorough examination can be performed.</p> </div>	https://treatment.plazi.org/id/3D7587E2D17DFFE0FCC0F93419CB0697	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Linkem, Charles W.;Diesmos, Arvin C.;Brown, Rafe M.	Linkem, Charles W., Diesmos, Arvin C., Brown, Rafe M. (2011): Molecular systematics of the Philippine forest skinks (Squamata: Scincidae: Sphenomorphus): testing morphological hypotheses of interspecific relationships. Zoological Journal of the Linnean Society 163 (4): 1217-1243, DOI: 10.1111/j.1096-3642.2011.00747.x, URL: http://dx.doi.org/10.1111/j.1096-3642.2011.00747.x
3D7587E2D163FFE3FCC0F91519640773.text	3D7587E2D163FFE3FCC0F91519640773.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Insulasaurus Taylor 1922	<div><p>Insulasaurus Taylor, 1922</p> <p>Type species: Insulasaurus wrighti Taylor, 1922.</p> <p>Definition: The clade comprising Insulasaurus wrighti Taylor, 1922 and all species that share a more recent common ancestor with Insulasaurus wrighti than with Anomalopus verreauxii, Calyptotis scutirostrum, Coeranoscincus frontalis, Coggeria naufragus, Ctenotus taeniolatus, Eremiascincus richardsonii, Eulamprus quoyii, Glaphyromorphus isolepis, Gnypetoscincus queenslandiae, Hemiergis decresiencsis, Lerista lineata, Lipinia pulchella, Lobulia elegans, Nangura spinosa, Notoscincus ornatus, Ophioscincus australis, Otosaurus cumingi, Papuascincus stanleyanus, Parvoscincus sisoni, Pinoyscincus jagori, Prasinohaema flavipes, Saiphos equalis, Scincella lateralis, Sphenomorphus melanopogon, and Tytthoscincus hallieri.</p> <p>Description: Insulasaurus is diagnosed by the following combination of characters: (1) medium body size, 45–64 mm SVL; (2) paravertebral scale rows 62–78; (3) midbody scale rows 29–41; and (4) subdigital lamellae 15–25.</p> <p>Included species: Insulasaurus arborens (Taylor, 1917), Insulasaurus traanorum (Linkem, Diesmos &amp; Brown, 2010a), Insulasaurus wrighti Taylor, 1925, and Insulasaurus victoria (Brown &amp; Alcala, 1980).</p> <p>Comment: The monotypic genus Insulasaurus was described by Taylor (1925) based on the presence of a divided frontonasal scale. Greer &amp; Parker (1967) found this character to be variable within Insulasaurus wrighti, and subsequently placed Insulasaurus wrighti in the Sphenomorphus variegatus group and synonymized Insulasaurus with Sphenomorphus. We found that Insulasaurus wrighti, Insulasaurus victoria, Insulasaurus traanorum (all from Palawan Island), and Insulasaurus arborens (Panay Island) are monophyletic, and distinct from other Philippine skinks. Our phylogeny suggests that this small, unique, and biogeographically circumscribed clade is more closely related to the genera Lipinia and Papuascincus, but separate from both, and therefore worthy of designation as a unique genus.</p> <p>At this time, we have no data suggesting that other Sphenomorphus species would be properly placed in the genus Insulasaurus, although species in Borneo (e.g. Sphenomorphus kinabaluensis and Sphenomorphus murudensis) are potential candidates should future phylogenetic studies determine that they are more closely related to Insulasaurus than they are to Sphenomorphus s.s.</p></div> 	https://treatment.plazi.org/id/3D7587E2D163FFE3FCC0F91519640773	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Linkem, Charles W.;Diesmos, Arvin C.;Brown, Rafe M.	Linkem, Charles W., Diesmos, Arvin C., Brown, Rafe M. (2011): Molecular systematics of the Philippine forest skinks (Squamata: Scincidae: Sphenomorphus): testing morphological hypotheses of interspecific relationships. Zoological Journal of the Linnean Society 163 (4): 1217-1243, DOI: 10.1111/j.1096-3642.2011.00747.x, URL: http://dx.doi.org/10.1111/j.1096-3642.2011.00747.x
3D7587E2D161FFE2FF5EFF321FC806AF.text	3D7587E2D161FFE2FF5EFF321FC806AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Otosaurus Gray 1845	<div><p>Otosaurus Gray, 1845</p> <p>Type species: Otosaurus cumingi Gray, 1845.</p> <p>Definition: The clade comprising Otosaurus cumingi (Gray, 1845) and all species that share a more recent common ancestor with Otosaurus cumingi than with Anomalopus verreauxii, Calyptotis scutirostrum, Coeranoscincus frontalis, Coggeria naufragus, Ctenotus taeniolatus, Eremiascincus richardsonii, Eulamprus quoyii, Glaphyromorphus isolepis, Gnypetoscincus queenslandiae, Hemiergis decresiencsis, Insulasaurus wrighti, Lerista lineata, Lipinia pulchella, Lobulia elegans, Nangura spinosa, Notoscincus ornatus, Ophioscincus australis, Papuascincus stanleyanus, Parvoscincus sisoni, Pinoyscincus jagori, Prasinohaema flavipes, Saiphos equalis, Scincella lateralis, Sphenomorphus melanopogon, and Tytthoscincus hallieri.</p> <p>Description: Otosaurus is diagnosed by the following combination of characters: (1) body large and robust, with adults being longer than 115 mm SVL; (2) large supranasal scales in contact medially, occluding frontonasal contact with the rostral; and (3) supraoculars seven or eight.</p> <p>Included species: Otosaurus cumingi Gray, 1845.</p> <p>Comments: The species Otosaurus cumingi Gray, 1845 has always been a morphological outlier to the other Philippine skinks. Being the only Sphenomorphus group skink in the region to have large supranasal scales and having an average body size double that of other species (Gray, 1845; Taylor, 1922a, Brown &amp; Alcala, 1980), it has been recognized as phenotypically distinct and unique amongst Philippine skinks. Our genetic and morphological results confirm its uniqueness amongst other lineages. Historically, this species was placed in the genus Otosaurus Gray, 1845 because of its distinctive morphology. As Otosaurus cumingi is the type species for the genus Otosaurus and is found to be both morphologically and genetically distinct, and our phylogenetic analyses place it in a polytomy with the Australian genera of the Sphenomorphus group and with the clade of Parvoscincus and Pinoyscincus, we re-establish Otosaurus as a monotypic genus, moving cumingi from Sphenomorphus to Otosaurus.</p> </div>	https://treatment.plazi.org/id/3D7587E2D161FFE2FF5EFF321FC806AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Linkem, Charles W.;Diesmos, Arvin C.;Brown, Rafe M.	Linkem, Charles W., Diesmos, Arvin C., Brown, Rafe M. (2011): Molecular systematics of the Philippine forest skinks (Squamata: Scincidae: Sphenomorphus): testing morphological hypotheses of interspecific relationships. Zoological Journal of the Linnean Society 163 (4): 1217-1243, DOI: 10.1111/j.1096-3642.2011.00747.x, URL: http://dx.doi.org/10.1111/j.1096-3642.2011.00747.x
3D7587E2D161FFE5FF5EF8EF1F040256.text	3D7587E2D161FFE5FF5EF8EF1F040256.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parvoscincus Ferner, Brown & Greer 1997	<div><p>Parvoscincus Ferner, Brown &amp; Greer, 1997</p> <p>Type species: Parvoscincus sisoni Ferner, Brown &amp; Greer, 1997.</p> <p>Definition: The clade comprising Parvoscincus sisoni (Ferner, Brown &amp; Greer, 1997) and all species that share a more recent common ancestor with Parvoscincus sisoni than with Anomalopus verreauxii, Calyptotis scutirostrum, Coeranoscincus frontalis, Coggeria naufragus, Ctenotus taeniolatus, Eremiascincus richardsonii, Eulamprus quoyii, Glaphyromorphus isolepis, Gnypetoscincus queenslandiae, Hemiergis decresiencsis, Insulasaurus wrigthi, Lerista lineata, Lipinia pulchella, Lobulia elegans, Nangura spinosa, Notoscincus ornatus, Ophioscincus australis, Otosaurus cumingii, Papuascincus stanleyanus, Pinoyscincus jagori, Prasinohaema flavipes, Saiphos equalis, Scincella lateralis, Sphenomorphus melanopogon, and Tytthoscincus hallieri.</p> <p>Description: Parvoscincus is diagnosed by the following combination of characters: (1) body size usually small (&lt;55 mm SVL) but larger in high-elevation species (46 mm &lt;SVL &lt;86 mm); (2) four enlarged supraoculars; (3) paravertebral scales 51–110; (4) midbody scale rows 23–46; and (5) subdigital lamellae 10–20.</p> <p>Included species: Parvoscincus beyeri (Taylor, 1922), Parvoscincus boyingi (Brown et al., 2010), Parvoscincus decipiens (Boulenger, 1894), Parvoscincus hadros (Brown et al., 2010), Parvoscincus igorotorum (Brown et al., 2010), Parvoscincus laterimaculatus (Brown &amp; Alcala, 1980), Parvoscincus leucospilos (Peters, 1872), Parvoscincus lawtoni (Brown &amp; Alcala, 1980), Parvoscincus luzonensis (Boulenger, 1894), Parvoscincus kitangladensis (Brown, 1995), Parvoscincus palawanensis (Brown &amp; Alcala, 1961), Parvoscincus sisoni (Ferner, Brown &amp; Greer, 1997), Parvoscincus steerei (Stejneger, 1908), and Parvoscincus tagapayo (Brown et al., 1999).</p> <p>Comments: The recently described genus Parvoscincus (Ferner, Brown &amp; Greer, 1997) is nested within a large clade of Philippine Sphenomorphus (Clade K). Represented in our phylogeny by the type species, Parvoscincus sisoni, it is clear that this genus is not phylogenetically distinct from other Philippine Sphenomorphus as originally proposed (Ferner, Brown &amp; Greer, 1997). The other species in this genus, Parvoscincus palawanensis, was not sampled; therefore, it is uncertain if it would be related to Parvoscincus sisoni, but we assume that it is until contrary evidence is presented. Clade K is clearly a unique and supported group of mostly small species of Philippine Sphenomorphus. As Parvoscincus is placed within this clade, we recommend that the name Parvoscincus be expanded to include the other small-bodied species in this Philippine clade (Parvoscincus leucospilos, Parvoscincus tagapayao, Parvoscincus luzonensis, Parvoscincus lawtoni, Parvoscincus kitangladensis, Parvoscincus laterimaculatus, Parvoscincus steerei, Parvoscincus decipiens) in addition to the secondarily enlarged, montane forest species (Parvoscincus beyeri, Parvoscincus boyingi, Parvoscincus igorotorum, and Parvoscincus hadros). Two species (Sphenomorphus acutus and Sphenomorphus diwata) in the Philippines are not diagnosable to either Parvoscincus or Pinoyscincus. These morphologically distinct species are genetically most similar to Parvoscincus, but this relationship has low phylogenetic support. We prefer to leave these species incertae sedis until a more thorough examination can be performed.</p> </div>	https://treatment.plazi.org/id/3D7587E2D161FFE5FF5EF8EF1F040256	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Linkem, Charles W.;Diesmos, Arvin C.;Brown, Rafe M.	Linkem, Charles W., Diesmos, Arvin C., Brown, Rafe M. (2011): Molecular systematics of the Philippine forest skinks (Squamata: Scincidae: Sphenomorphus): testing morphological hypotheses of interspecific relationships. Zoological Journal of the Linnean Society 163 (4): 1217-1243, DOI: 10.1111/j.1096-3642.2011.00747.x, URL: http://dx.doi.org/10.1111/j.1096-3642.2011.00747.x
3D7587E2D166FFE5FF6AFD86199305BE.text	3D7587E2D166FFE5FF6AFD86199305BE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scincella Mittleman 1950	<div><p>Scincella Mittleman, 1950</p> <p>Type species: Scincella lateralis (Say, 1823).</p> <p>Definition: The clade comprising Scincella lateralis (Say, 1823) and all species that share a more recent common ancestor with Scincella lateralis than with Anomalopus verreauxii, Calyptotis scutirostrum, Coeranoscincus frontalis, Coggeria naufragus, Ctenotus taeniolatus, Eremiascincus richardsonii, Eulamprus quoyii, Glaphyromorphus isolepis, Gnypetoscincus queenslandiae, Hemiergis decresiencsis, Insulasaurus wrighti, Lerista lineata, Lipinia pulchella, Lissonota maculata, Lobulia elegans, Nangura spinosa, Notoscincus ornatus, Ophioscincus australis, Otosaurus cumingii, Papuascincus stanleyanus, Parvoscincus sisoni, Pinoyscincus jagori, Prasinohaema flavipes, Saiphos equalis, Sphenomorphus melanopogon, Tytthoscincus hallieri.</p> <p>Description: Scincella can be diagnosed by the following combination of characters: (1) body size medium (SVL usually &lt;65 mm); (2) alpha palate (Greer, 1974) with nine premaxillary teeth; (3) long, thin postorbital bone usually present; and (4) with a transparent window in a movable lower eyelid. Transparent window may be lacking in southern populations of Sp. cheerei.</p> <p>Included species: Scincella apraefrontalis Nguyen, Nguyen, Bohme &amp; Ziegler, 2010, Scincella assata (Cope, 1864), Scincella barbouri (Stejneger, 1925), Scincella boettgeri (Van Denburgh, 1912), Scincella capitanea Oubeter, 1986, Scincella caudaequinae (Smith, 1951), Scincella cherriei (Cope, 1893), Scincella doriae (Boulenger, 1887), Scincella forbesora (Taylor, 1937), Scincella formosensis (Van Denburgh, 1912), Scincella gemmingeri (Cope, 1864), Scincella inconspicua (Müller, 1894), Scincella incerta (Stuart, 1940), Scincella kikaapoa Garcia-Vazquez, Canseco- Marquez &amp; Nieto-Montes de Oca, 2010, Scincella lateralis (Say, 1823), Scincella macrotis (Steindachner, 1867), Scincella melanosticta (Boulenger, 1887), Scincella modesta (Günther, 1864), Scincella monticola (Schmidt, 1927), Scincella ochracea (Bourret, 1937), Scincella potanini (Günther, 1896), Scincella przewalskii (Bedriaga, 1912), Scincella punctatolineata (Boulenger, 1893), Scincella rarus (Myers &amp; Donnelly), 1991, Scincella reevesi (Gray, 1838), Scincella rufocaudatus Darevsky &amp; Nguyen, 1983, Scincella rupicola (Smith, 1927), Scincella schmidti (Barbour, 1927), Scincella silvicola (Taylor, 1937), Scincella tsinglingensis (Hu &amp; Djao, 1966), Scincella vandenburghi (Schmidt, 1927), and Scincella victoriana (Shreve, 1940).</p> <p>Comment: The New World species Scincella cherriei and Scincella assata are nested within the genus Scincella, sister to the North American species Scincella lateralis. We predict that Scincella rarus, and Scincella incertus also will be members of this clade. When Greer (1974: 33) revised the genus Leiolepisma, he provided detailed comments about the potential relationships of these Central American skinks. Morphologically, these species are a mix of Sphenomorphus and Scincella, with Scincella assatus and Scincella incertus lacking a postorbital bone but possessing a window in the lower eye (characters of Scincella) and Scincella cherriei possessing a postorbital bone but having population variation in the presence of the lower eyelid window. Greer (1974) inferred that Scincella cherriei was the primitive form of the Central American radiation owing to the possession of the postorbital bone and placed these species in Sphenomorphus. He noted that this did not make sense biogeographically because it inferred a separate migration across the Bering Bridge, but he argued it was more plausible than the re-evolution of the postorbital bone in Scincella cherriei. Our molecular evidence shows that the Central American species are part of the same radiation as North American Scincella, following the biogeographical expectation. It is therefore reasonable to move these Central American skinks to the genus Scincella.</p> </div>	https://treatment.plazi.org/id/3D7587E2D166FFE5FF6AFD86199305BE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Linkem, Charles W.;Diesmos, Arvin C.;Brown, Rafe M.	Linkem, Charles W., Diesmos, Arvin C., Brown, Rafe M. (2011): Molecular systematics of the Philippine forest skinks (Squamata: Scincidae: Sphenomorphus): testing morphological hypotheses of interspecific relationships. Zoological Journal of the Linnean Society 163 (4): 1217-1243, DOI: 10.1111/j.1096-3642.2011.00747.x, URL: http://dx.doi.org/10.1111/j.1096-3642.2011.00747.x
