identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
3D096322D43E5646FF5BFD767BA7FD9B.text	3D096322D43E5646FF5BFD767BA7FD9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Meyliidae De Coninck 1965	<div><p>Family Meyliidae De Coninck, 1965</p><p>Diagnosis: Adapted from Decraemer and Rho (2014). Body slender or fusiform. Body with fine striation, appearing smooth; with or without ornamentation. Ornamentation either with transverse rows of fine spines, or longitudinal differentiations of ridges or spines; transverse striae may be interrupted by a narrow lateral field; rarely smooth cuticle, i.e., without striae. Few somatic setae, not differentiated. Amphidial fovea large vesicular with or without inner spiral structure, mostly located on the cephalic region, rarely postcephalic. Cephalic cuticle usually smooth, either thickened and cuticularised or thin. Cephalic setae on short peduncle or directly inserted into cephalic cuticle. Pharynx anteriorly muscular, posteriorly tapering, usually with large overlapping dorsal gland. Pigment spots rarely present. Pseudocoelomocytes usually present. Male reproductive system with two testes, the posterior one reflexed; spicules arched; gubernaculum with apophysis, rarely absent. Female reproductive system didelphic-amphidelphic with ovaries reflexed. Phasmata-like sructures present in Erebussau .</p><p>Type genus: Meylia Gerlach, 1956 . Other genera: Boucherius, Gerlachius, Noffsingeria, and Erebussau nom. nov. in present article.</p></div>	https://treatment.plazi.org/id/3D096322D43E5646FF5BFD767BA7FD9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bezerra, Tania Nara;Pape, Ellen;Hauquier, Freija;Vanreusel, Ann	Bezerra, Tania Nara, Pape, Ellen, Hauquier, Freija, Vanreusel, Ann (2021): Description and distribution of Erebussau nom. nov. pro Erebus Bussau, 1993 nec Erebus Latreille, 1810 with description of a new specie, and of Odetenema gesarae gen. nov., sp. nov. (Nematoda: Desmoscolecida) from nodule-bearing abyssal sediments in the Pacific. Zootaxa 4903 (4): 542-562, DOI: 10.11646/zootaxa.4903.4.4
3D096322D43F5646FF5BF8667B35FA43.text	3D096322D43F5646FF5BF8667B35FA43.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Erebussau Bezerra & Pape & Hauquier & Vanreusel 2021	<div><p>Genus Erebussau nom. nov. pro Erebus Bussau, 1993 nec Erebus Latreille, 1810</p><p>Diagnosis: Adapted from Decraemer and Rho (2014). Body cuticle annulated, without ornamentation. Distinct body annulation at the anterior and posterior end. Smooth annules near the anterior end with triangular outline. No lateral differentiation. Cephalic region swollen and offset. Cephalic region and lips thin-walled. Smooth labial surface, labial sensilla not visible. Four cephalic setae with pointed tip. Amphidial fovea vesicular, just posterior to the cephalic region. Stoma small, without teeth; pharynx contour not clear, obscured by the presence of glandular cells. Pigment spots present along the intestine with irregular number, location and shape. Pseudocoelomocytes present. Female reproductive system didelphic, amphidelphic, with reflexed ovaries. Male with two reflexed testes or just the posterior one reflexed. Two round phasmata-like structures, laterally, opposed to each other, on the last posterior third of the tail. Tail conical.</p><p>Type species. Erebussau tenebricosus nom. nov., comb. nov.</p><p>One other species in genus: Erebussau profundus sp. nov.</p><p>Differential diagnosis: Erebussau nom. nov. has a unique appearance when compared with the other genera of the family Meyliidae . It has a distinct body annulation at the anterior and posterior end, without lateral field differentiation as found in Noffsingeria . The peculiar offset cephalic region of Erebussau is unique among the Meyliidae . Pigment spots were not described so far for Meyliidae but they are clearly seen in Erebussau . The presence of phasmata-like structures are also described for the first time within the family Meyliidae .</p><p>Etymology: The name is a combination of Erebus (lat.) underworld, referring to the deep-sea habitat of the specimens; and an honor to Dr. Christian Bussau, who first described the genus giving the previous name, Erebus .</p><p>Type habitat and distribution: Deep-sea with sediment grain size dominated by silt, containing nodules.</p><p>Material examined: 2 ³, 3 ♀ and 4 juveniles from the GSRNOD15–A cruise; 2 ³, 6 ♀ and 2 juveniles from JPI Oceans “Ecological aspects of deep-sea mining cruise” SO239 cruise ; 3 ♀ from Peru Basin, cruise SO242-2 (Table 1) .</p></div>	https://treatment.plazi.org/id/3D096322D43F5646FF5BF8667B35FA43	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bezerra, Tania Nara;Pape, Ellen;Hauquier, Freija;Vanreusel, Ann	Bezerra, Tania Nara, Pape, Ellen, Hauquier, Freija, Vanreusel, Ann (2021): Description and distribution of Erebussau nom. nov. pro Erebus Bussau, 1993 nec Erebus Latreille, 1810 with description of a new specie, and of Odetenema gesarae gen. nov., sp. nov. (Nematoda: Desmoscolecida) from nodule-bearing abyssal sediments in the Pacific. Zootaxa 4903 (4): 542-562, DOI: 10.11646/zootaxa.4903.4.4
3D096322D43D564BFF5BFFEA7FA6F844.text	3D096322D43D564BFF5BFFEA7FA6F844.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Erebussau tenebricosus (Bezerra & Pape & Hauquier & Vanreusel 2021) Bezerra & Pape & Hauquier & Vanreusel 2021	<div><p>Erebussau tenebricosus nom. nov., comb. nov.</p><p>(Figs 2, 3 and 4; Tables 1 and 2)</p><p>Material examined: Three males, nine females and five juveniles. Some of these adult specimens used for the proposal of the new replacement nomen are deposited in the Ghent University Zoological Museum—Museum voor Dierkunde (UGMD) under the codes: Male 1 (Inventory No. UGMD 104398), Male 2 (Inventory No.104399), Female 1 (Inventory No. 104400) and Female 2 (Inventory No. 104401).</p><p>Habitat and locality: Deep-sea with sediment grain size dominated by silt, containing polymetallic nodules. Specimen Male 1, was found in a sample from the GSR license area in the CCFZ during expedition SO239, Station 146, Core 11-A, 0–5 cm at an average of 4544 m depth . The species was previously found by Bussau (1993) in the Peru Basin, in February 1989 at an average of 4137 m depth .</p><p>Description: Males: Body elongated, strongly annulated, especially at anterior and posterior ends. Annulations of the anterior part of the body with triangular smooth outline.A pair of somatic setae is present on the tail. Cephalic region marked by a narrower anterior region of the body. Four stout cephalic setae are present. Amphids vesicular, with blistered fovea. Buccal cavity small and unarmed. Pharynx surrounded by many glands, impairing the description of the muscular disposition. Two or three well defined pigment spots, ventrally and subventrally disposed at the anterior portion of the intestine. Pseudocoelomocytes present, surrounding the anterior portion of the intestine. Anterior testis directed towards the anterior end, well developed. Posterior testis much smaller and reflexed. Spicules curved. Gubernaculum with caudal apophysis. No precloacal supplements. Three caudal glands contained in the tail. Phasmata-like structures located laterally at the last posterior third of the tail region, on each side of the body. These phasmata-like structures can be easily overlooked. Tail conical, long with spinneret.</p><p>Females: Similar to males in most aspects, although generally longer. Cephalic shape more round and pronounced off-set than in males. Cuticle annulation more evident than in males. No pigment spots visible. Reproductive system didelphic with reflexed ovaries. Reproductive system with a clear posterior ovary and a much smaller anterior ovary, not well developed, apparently not functional, refleting a possible pseudomonodelphic condition. Vulva with non-protruding lips, positioned close to the mid body, slightly posteriorly in most cases. No perivulvar glands observed. Uterus not well distinguished. Oviduct indistinct; ovaries antidromous with reflexed tips. Phasmata-like structures larger than in males. Female tail conical, longer than in males. Terminal spinneret with same morphology as in males but slightly elongated.</p><p>Juveniles: Similar to adults in most morphological aspects.</p><p>...Continued on the next page</p></div>	https://treatment.plazi.org/id/3D096322D43D564BFF5BFFEA7FA6F844	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bezerra, Tania Nara;Pape, Ellen;Hauquier, Freija;Vanreusel, Ann	Bezerra, Tania Nara, Pape, Ellen, Hauquier, Freija, Vanreusel, Ann (2021): Description and distribution of Erebussau nom. nov. pro Erebus Bussau, 1993 nec Erebus Latreille, 1810 with description of a new specie, and of Odetenema gesarae gen. nov., sp. nov. (Nematoda: Desmoscolecida) from nodule-bearing abyssal sediments in the Pacific. Zootaxa 4903 (4): 542-562, DOI: 10.11646/zootaxa.4903.4.4
3D096322D433564EFF5BFCBD7815FD9A.text	3D096322D433564EFF5BFCBD7815FD9A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Erebussau profundus Bezerra & Pape & Hauquier & Vanreusel 2021	<div><p>Erebussau profundus sp. nov.</p><p>(Figs 5, 6 and 7; Tables 1 and 2)</p><p>Material examined for description: One male, three females and one juvenile. One holotype male (Inventory No. UGMD 104402) and one paratype female (Inventory No. UGMD 104403) are deposited in the Zoology Museum of Ghent University—Museum voor Dierkunde (UGMD) .</p><p>Type habitat and locality: Deep-sea with sediment grain size dominated by silt, containing nodules. Male holotype was found in a sample from the GSR license area in the CCFZ during expedition SO239, Station 141, Core 1, 0–1 cm, at an average of 4509 m depth .</p><p>Description: Males: Body elongated, strongly annulated until mid-body. Posterior end also strongly annulated. Annulations of the anterior part of the body with triangular smooth outline. No somatic setae are visible. Cephalic region clearly offset followed by a narrow constriction about 50% narrower than the section behind this constriction. Four cephalic setae are present. Amphids vesicular, with a blistered amphidial fovea. Buccal cavity small and unarmed. Pharynx surrounded by many glands. Many pigment spots present, located at the end of the pharynx and beginning of intestine. Pseudocoelomocytes present, at the end of the pharynx and beginning of the intestine. Testes reflexed. Anterior testis well developed, posterior one much smaller. Spicules just slightly curved. Gubernaculum with caudal apophysis. Three setae inserted on elevations of the body cuticle, present as pre-cloacal supplements, plus one additional post-cloacal supplement. Small phasmata-like structures located laterally at the last posterior third of the tail region, on each side of the body. These structures can easily be overlooked. Three caudal glands contained in the tail. Tail conical, long with spinneret.</p><p>Females: Similar to males in most aspects, although generally longer. No sexual dimorphism detected. Reproductive system didelphic, amphidelphic, with reflexed ovaries. Reproductive system with a clear posterior ovary and a smaller anterior ovary. Vulva with non-protruding lips, positioned close to the mid body, slightly posteriorly in most cases. No perivulvar glands observed. Vagina transverse. Uterus not well-distinguished, appearing to be uniform. Oviduct indistinct; ovaries antidromously with reflexed tips. Phasmata-like structures, laterally at the last posterior third of the tail region, on each side of the body, larger than in males. Female tail conical, longer than in males. Terminal spinneret with same morphology as in males but slightly elongated.</p><p>Juveniles: Similar to adults in most morphological aspects.</p><p>Etymology: Profundus (Latin, adjective class I) = deep, profound.</p><p>Differential diagnosis: Erebussau profundus sp. nov. closely resembles Erebussau tenebricosus nom. nov., comb. nov., but males of Erebussau profundus sp. nov. are on average 27% longer in total length. This difference is reduced to 16% when females of both species are compared. Pigment spots are more numerous in Erebussau profundus sp. nov. Five to seven pigment spots were observed, in contrast with two in Erebussau tenebricosus nom. nov., comb. nov. . Other distinct characteristics are the presence of pre- and post-cloacal supplements and the shape of the spicules. The spicules are just slightly curved and 51 % longer when compared to the clearly curved and smaller ones of Erebussau tenebricosus nom. nov., comb. nov. (Table 2). The triangular outline shape of cuticle annulation at the anterior part of the bod of Erebussau profundus sp. nov. is less pronounced than that in Erebussau tenebricosus nom. nov., comb. nov. and extends further until mid-body, although this could be the result of movement, such as contraction of the longitudinal body muscles and friction with the sediment. These movements can cause rings to telescope more or less, proving a more triangular outline.</p><p>General remarks: Analyses of different specimens allowed us to observe intra- and interspecific differences during the new denomination of Erebussau nom. nov., resulting in an update of the description of the family Meyliidae .</p><p>Blister-like amphids are a synapomorphy for Desmoscolecida (Lorenzen, 1981, 1994) and the previous descriptions mentioned that members of Meyliidae differ from the other desmoscolecids by the vesicular fovea showing an inner spiral structure which is not the case for Erebussau nom. nov. Bussau (1993) described a blistered heartshaped amphid vesicular structure, this was not visible in the analyzed Erebussau nom. nov. specimens.</p><p>Another finding was the presence of pigment spots. Pigment spots with different sizes and shapes were found in the specimens of both species of Erebussau . The absence of lens-like structures refrains us from the use of the term ocelli. Bussau (1993) observed in Tricoma and Desmoscolex that the pigment spots were located at the level of the ampulla-like widening of the fusus amphidialis and hypothesized that the pigment spots are in fact concentrated secretions from the amphidial sheath cell, explaining their differences in size and shape among specimens. Lorenzen (1994) described pigment spots as resembling droplets of fluid and noted that their presence appears independent from the biotope in which the animals occur, stating that they were not present in deep-sea species. However, Decraemer (1989) found pigment spots in deep-sea Tricoma (Quadricoma) loricata specimens from the flat of Cap Vert (tropical Atlantic Ocean) revoking Lorenzen’s (1994) statement.</p><p>Phasmata were considered absent in Meyliidae, but phasmata-like structures are present in Erebussau nom. nov.</p><p>Pseudocoelomocytes are observed at the end of the pharynx, beginning of intestine, both in E. tenebricosus nom. nov., comb. nov. and E. profundus sp. nov., although they are more conspicuous in E. profundus sp. nov. To date, Erebussau comb. nov., together with Noffsingeria and Meylia vangoethemi, are the sole representatives of Meyliidae which possess pseudocoelomocytes.</p><p>The females of the described sibling species of Erebussau comb. nov. did not show striking features which can be used to separate the two species, but they could be distinguished by the sharp offset cephalic region, together with the greater length of E. profundus sp. nov. being 16 % longer on average, the triangular outline shape of cuticle annulation which extends further until mid-body and the well-developed anterior ovary observed in the two described specimens of E. profundus sp. nov.</p></div>	https://treatment.plazi.org/id/3D096322D433564EFF5BFCBD7815FD9A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bezerra, Tania Nara;Pape, Ellen;Hauquier, Freija;Vanreusel, Ann	Bezerra, Tania Nara, Pape, Ellen, Hauquier, Freija, Vanreusel, Ann (2021): Description and distribution of Erebussau nom. nov. pro Erebus Bussau, 1993 nec Erebus Latreille, 1810 with description of a new specie, and of Odetenema gesarae gen. nov., sp. nov. (Nematoda: Desmoscolecida) from nodule-bearing abyssal sediments in the Pacific. Zootaxa 4903 (4): 542-562, DOI: 10.11646/zootaxa.4903.4.4
3D096322D437564EFF5BFE167F20F91A.text	3D096322D437564EFF5BFE167F20F91A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Odetenema Bezerra & Pape & Hauquier & Vanreusel 2021	<div><p>Genus Odetenema gen. nov.</p><p>Diagnosis: Body slender, tapering towards both extremities. Cuticle strongly annulated anteriorly and posteriorly, without desmens. Mid body cuticle almost smooth. Row of thick somatic setae on peduncles, ventrally, on the last body third, from the spicules directed to the anterior end. Cephalic region longer than wide, with cuticularised walls on lateral view. Cephalic setae on small peduncles. Smooth labial surface. Labial ring just anterior the cephalic setae peduncles. Buccal cavity small and unarmed. Amphidial fovea large and vesicular. Pharynx cylindrical. Anterior pharyngeal part surrounded by elongated glands. Pigment spots in different sizes at the anterior part of the intestine. Male reproductive system diorchic, with posterior testis reflexed. Spicules slightly arcuate, distally tapered, without capitulum. Gubernaculum without apophysis. Female reproductive system didelphic, amphidelphic with outstretched ovaries. Phasmata laterally, on the last third of the tail. Tail conical. Caudal glands opening in two separate tubular outlets.</p><p>Type species: Odetenema gesarae sp. nov.</p><p>Differential diagnosis: Comparing with the other genera of the subfamily Tricominae, the very large number of body rings with great difference between dorsal and ventral body side and absence of desmen point the new genus towards Paratricoma but without ornamentation of the body rings. The shape of the end ring with two tubular outlets is unique to the Tricominae .</p><p>Etymology: The genus is named in honour of Odete Campinas Bezerra, mother of the first author.</p></div>	https://treatment.plazi.org/id/3D096322D437564EFF5BFE167F20F91A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bezerra, Tania Nara;Pape, Ellen;Hauquier, Freija;Vanreusel, Ann	Bezerra, Tania Nara, Pape, Ellen, Hauquier, Freija, Vanreusel, Ann (2021): Description and distribution of Erebussau nom. nov. pro Erebus Bussau, 1993 nec Erebus Latreille, 1810 with description of a new specie, and of Odetenema gesarae gen. nov., sp. nov. (Nematoda: Desmoscolecida) from nodule-bearing abyssal sediments in the Pacific. Zootaxa 4903 (4): 542-562, DOI: 10.11646/zootaxa.4903.4.4
3D096322D437564DFF5BFCE67BFBF829.text	3D096322D437564DFF5BFCE67BFBF829.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Odetenema gesarae Bezerra & Pape & Hauquier & Vanreusel 2021	<div><p>Odetenema gesarae gen. nov., sp. nov.</p><p>(Figs 8, 9 and 10; Tables 1 and 3)</p><p>Material examined for description: One male and two females. The holotype male (Inventory No. UGMD 104404) and one paratype female (Inventory No. UGMD 104405) are deposited in the Zoology Museum of Ghent Univer-sity—Museum voor Dierkunde (UGMD) .</p><p>Type habitat and locality: Deep-sea with sediment grain size dominated by silt, containing nodules. The male holotype was found in a sample from the GSR license area in the CCFZ at about 4500 m depth, during expedition GSRNOD15–A, MUC006, 0-5 cm deep in the sediment .</p><p>Description: Male: Body slender, tapering anteriorly and posteriorly. Anterior width 50 % of the maximum body diameter. Cuticle with 184 dorsal and 247 ventral rings. The first 23 (dorsal) and 52 (ventral) rings, as well as the last 16 (dorsal) and 25 (ventral) have a well defined outline. The first 6 (dorsal) and 4 (ventral) rings, together with the last 5 ones, show the most explicit defined outline on the body. Desmen not present. Inversion ring not visible. Cuticle at mid-body almost smooth impairing the ring counting. Few somatic setae, relatively thick on short peduncles. Cephalic region longer than wide, with cuticularised walls on lateral view. Cuticle without concretions or ornaments. External labial sensilla indistinct. Cephalic setae located on small peduncles. Amphidial fovea large and vesicular, reaching posteriorly the first ring. Round fovea clearly visible. Buccal cavity small, unarmed. Pharynx anteriorly surrounded by elongated glands and slightly enlarged posteriorly. Nerve ring at 39 µm from the anterior end, 74 % of pharyngeal length. Pseudocoelomocytes not recognizable. Pigment spots of different sizes at the anterior part of the intestine on both sides of the body. Phasmata small, can be easily overlooked. Reproductive system diorchic, testes opposed. Posterior testis reflexed. Vas deferens not clearly distinguished. Spicules slightly curved. Gubernaculum without apophysis. Two small rounded gland cells with distinct nucleus posterior to the gubernaculum. A row of four thick setae on small peduncles are ventrally positioned from the spicules towards the anterior end. Three caudal glands can be observed but two separate tubular outlets, close to one another, are present. Tail conical, end ring with two tubular outlets.</p><p>Females: Longer, but otherwise similar to males in most aspects. Nerve ring not visible on females due to the amount of glands covering the pharynx. Pseudocoelomocytes visible ventrally of the intestine. No sexual dimorphism detected. Reproductive system didelphic, amphidelphic, with outstretched ovaries. Vulva positioned at 56 % of the body from the anterior end. No perivulvar glands observed. Vagina short, transverse and thick-walled. Uterus not well distinguished, appearing to be uniform. Oviduct indistinct. Ovaries outstretched, anterior one more developed than the posterior one with a single fully grown oocyte. Phasmata larger than in males. Terminal caudal glands opening with same morphology as in males, showing two separate tubular outlets. Tail conical.</p><p>Juvenile: Similar to adults in most morphological aspects.</p><p>Etymology: The species is named after the Global Sea Mineral Resources (GSR), the deep-sea exploratory division of the DEME Group.</p><p>Differential diagnosis: Odetenema gesarae gen. nov., sp. nov. differs from all other Tricominae genera described to date, by the special appearance of the cuticle, almost smooth at mid body. The long cephalic region, not offset, tapering to a narrow, and elongated anterior end, forming a kind of rostrum, combined with the short cephalic setae and the long amphid result in an exclusive anterior end. The two separate caudal gland outlets are unique to the order Desmoscolecida .</p><p>Although other specimens were observed, they were not well preserved and perhaps the measurements would not be accurate. For this reason we decided to measure the three specimens which were better preserved and could be measured with accuracy.</p></div>	https://treatment.plazi.org/id/3D096322D437564DFF5BFCE67BFBF829	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bezerra, Tania Nara;Pape, Ellen;Hauquier, Freija;Vanreusel, Ann	Bezerra, Tania Nara, Pape, Ellen, Hauquier, Freija, Vanreusel, Ann (2021): Description and distribution of Erebussau nom. nov. pro Erebus Bussau, 1993 nec Erebus Latreille, 1810 with description of a new specie, and of Odetenema gesarae gen. nov., sp. nov. (Nematoda: Desmoscolecida) from nodule-bearing abyssal sediments in the Pacific. Zootaxa 4903 (4): 542-562, DOI: 10.11646/zootaxa.4903.4.4
