taxonID	type	description	language	source
3D1487E00A324121FF7FFD7997EE3F99.taxon	materials_examined	Material examined (23 ex.). - Morocco, Bouârfa, 32.4909 lat. - 2.569 lon., ID 14994, 20 / III / 2013 (L. Bacon), ECWP; - Morocco, Bouârfa, 32.47079 lat. - 2.05724 lon., ID 15533, 18 / III / 2014 (J. Gabbard), ECWP; - Morocco, Bouârfa, Tamlelt, 32.4308 lat. - 2.57385 lon., ID 4677, 29 / V / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.51403 lat. - 2.48135 lon., ID 10527, 26 / VI / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.5661 lat. - 2.27995 lon., ID 10531, 22 / VIII / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.50673 lat. - 2.3373 lon., ID 10532, 08 / V / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.52544 lat. - 2.55285 lon., ID 10534, 05 / VI / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.43242 lat. - 2.708695 lon., ID 10535, 19 / IX / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.56644 lat. - 2.279845 lon., ID 10536, 10 / VII / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.50707 lat. - 2.33679 lon., ID 10537, 05 / IX / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.53565 lat. - 2.55283 lon., ID 10539, 12 / VI / 2008 (K. Bourass, S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.27 lat. - 2.4297 lon., ID 10540, 03 / X / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.53385 lat. - 2.5066 lon., ID 10543, 01 / V / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.53385 lat. - 2.55285 lon., ID 10544, 22 / VIII / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.52603 lat. - 2.5526 lon., ID 10545, 05 / VI / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.506294 lat. - 2.33689 lon., ID 10550, 26 / VI / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.43275 lat. - 2.70845 lon., ID 10551, 22 / VIII / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.41818 lat. - 2.27401 lon., ID 10552, 28 / VIII / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.5337 lat. - 2.50666 lon., ID 10553, 28 / VIII / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.56644 lat. - 2.27984 lon., ID 10555, 19 / VI / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.53385 lat. - 2.5066 lon., ID 10591, 26 / IX / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.44563 lat. - 2.39064 lon., ID 21697, 31 / III / 2015 (J. Gabbard), ECWP; - Morocco, Bouârfa, Tamlelt, 32.47249 lat. - 2.07223 lon., ID 23246, 25 / V / 2015 (A. François, M. García-Paris, J. L. Ruiz), ECWP.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A324121FC67FE19971E3D2A.taxon	materials_examined	Material examined (2 ex.). - Morocco, Tendrara, N 17, 32.89624 lat. - 2.05353 lon., ID 15534, 15 / IX / 2013 (J. Gabbard), ECWP; - Morocco, Tendrara, 32.74671 lat. - 2.36698 lon., ID 15845, 02 / V / 2014 (L. Bacon), ECWP.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A324121FC67FE19971E3D2A.taxon	distribution	Distribution and remarks. – A. australis, a species widely distributed in the neighboring Algeria, was recorded for the first time in Morocco by Geniez (2009) in the south-east of the Oriental region, between Bouarfa and Tendrara (Touloun et al., 2014, Dupré, 2017, Touloun, 2019, Ythier & Lourenço, 2022). The examined material extends its distribution 30 km to the west (Fig. 1).	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A324123FC6BFC6C91953F99.taxon	materials_examined	Material examined (54 ex.). - Morocco, Bouânane, 32.13901 lat. - 2.8941 lon., ID 8337, 14 / V / 2011 (J. Gabbard), ECWP; - Morocco, Bouârfa, 32.40921 lat. - 2.59711 lon., ID 564, 23 / IV / 2008 (D. Lijnen, S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, ID 565, 23 / VI / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.41434 lat. - 2.597 lon., ID 566, 01 / V / 2008 (Y. Hingrat), ECWP; - Morocco, Bouârfa, Tamlelt, ID 567, 10 / V / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.48212 lat. - 2.7282 lon., ID 10590, 22 / VIII / 2008 (S. Touil), ECWP; - Morocco, Maatarka, Kehf Tabakhoucht, 32.84563 lat. - 3.43927 lon., ID 16145, 16 / VII / 2014 (T. Dieuleveut), ECWP; - Morocco, Saidia, Moulouya mouth, 35.11125 lat. - 2.35659 lon., ID 21827, 09 / VIII / 2015 (A. François, B. Michel), ECWP; - Morocco, Tendrara, 32.74744 lat. - 2.51924 lon., ID 15844, 16 / V / 2014 (L. Bacon), ECWP; - Morocco, Enjil, ECWP, 33.14769 lat. - 4.58389 lon., ID 21824, 29 / VIII / 2014 (A. François), ECWP; - Morocco, Enjil, ECWP, 33.14769 lat. - 4.58389 lon., ID 21826, 12 / VI / 2014 (J. M. Touzet), ECWP; - Morocco, Enjil, ECWP, 33.13723 lat. - 4.58888 lon., ID 5706, 13 / VII / 2010 (J. Boisard), ECWP; - Morocco, Enjil, ECWP, 33.13723 lat. - 4.58888 lon., ID 6440, 07 / VIII / 2010 (J. Boisard), ECWP; - Morocco, Enjil, ECWP, 33.13723 lat. - 4.58888 lon., ID 10431, 13 / XII / 2011 (R. Guérin), ECWP; - Morocco, Enjil, ECWP, 33.13723 lat. - 4.58888 lon., ID 12532, 27 / IX / 2012 (I. Smaini), ECWP; - Morocco, Enjil, ECWP, 33.12162 lat. - 4.60675 lon., ID 15033, 17 / IX / 2013 (S. Gomez), ECWP; - Morocco, Enjil, ECWP, 33.14769 lat. - 4.58389 lon., ID 22915, 07 / VI / 2014 (G. Le Loc’h), ECWP; - Morocco, Lamjalil, 32.8938 lat. - 3.95663 lon., ID 10567, 18 / VI / 2010 (H. Hdidou), ECWP; - Morocco, Missour, 33.01061 lat. - 4.11166 lon., ID 10566, 20 / VII / 2009 (H. Hdidou), ECWP; - Morocco, Missour, Al Baten, 33.1997 lat. - 3.92071 lon., ID 560, 03 / IV / 2002 (J. Yvernault), ECWP; - Morocco, Missour, Al Baten, 33.2069 lat. - 3.869 lon., ID 561, 28 / V / 2002 (J. Yvernault), ECWP; - Morocco, Missour, AlBaten, 33.28932 lat. - 3.91184 lon., ID 562, 27 / VI / 2002 (J. Yvernault), ECWP; - Morocco, Missour, Al Baten, 33.31958 lat. - 3.87254563 lon., ID 563, 27 / VI / 2002 (J. Yvernault), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 558, 17 / III / 2008 (S. Boullenger), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 10525, 15 / VII / 2009 (A. François, C. Leret), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 4633, 19 / VIII / 2009 (C. Leret), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 5314, 19 / V / 2010 (S. Boullenger), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 5705, 10 / VII / 2010 (S. Boullenger), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 6438, 19 / VIII / 2010 (A. François), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 7025, 19 / VIII / 2010 (A. François), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 7026, 17 / IX / 2010 (H. Chaker), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 7035, 04 / X / 2010 (J. Talha), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 8823, 01 / VII / 2011 (A. François), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 8824, 02 / VII / 2011 (A. François, M. Sbai), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 9653, 29 / VII / 2011 (( H. Chaker), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 9654, 07 / VIII / 2011 (( H. Chaker), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 21825, 23 / VIII / 2015 (S. Lefèvre), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 21828, 10 / IX / 2015 (S. Lefèvre), ECWP; - Morocco, Missour, ECWP, 33.00739 lat. - 4.08819 lon., ID 10565, 15 / VI / 2009 (H. Hdidou), ECWP; - Morocco, Missour, ECWP, 33.00739 lat. - 4.08819 lon., ID 11499, 04 / V / 2012 (B. El Bachra), ECWP; - Morocco, Missour, ECWP, 33.00739 lat. - 4.08819 lon., ID 11501, 25 / V / 2012 (H. Chaker, H. Hdidou), ECWP; - Morocco, Missour, ECWP, 33.00739 lat. - 4.08819 lon., ID 12237, 03 / X / 2012 (A. François, M. Sbai), ECWP; - Morocco, Missour, ECWP, 33.00739 lat. - 4.08819 lon., ID 22460, 16 / IX / 2012 (M. Chamidou), ECWP; - Morocco, Missour, ECWP, 33.00739 lat. - 4.08819 lon., ID 24233, 29 / VIII / 2021 (G. Monchaux), ECWP; - Morocco, Missour, ECWP, 33.00739 lat. - 4.08819 lon., ID 24281, 12 / X / 2021 (A. Haddad, ECWP; - Morocco, Missour, ECWP, near offices office, 33.00722 lat. - 4.0976 lon., ID 559, 21 / VI / 2008 (A. Haddad), ECWP; - Morocco, Missour, ECWP, in an office, 33.00722 lat. - 4.0976 lon., ID 4630, 12 / VIII / 2009 (A. François), ECWP; - Morocco, Missour, ECWP, villa F 2 - 4, 33.00722 lat. - 4.0976 lon., ID 8076, 26 / IV / 2011 (P. Salanova), ECWP; - Morocco, Missour, ECWP, botanicallaboratory, 33.00722 lat. - 4.0976 lon., ID 10432, 03 / XI / 2011 (M. Bidat), ECWP; - Morocco, Missour, ECWP, villa, 33.00722 lat. - 4.0976 lon., ID 8645, 24 / VII / 2012 (C. Leret), ECWP; - Morocco, Missour, ECWP, restaurant, 33.00722 lat. - 4.0976 lon., ID 14652 / 14653, 12 / VII / 2013 (A. François), ECWP; - Morocco, Missour, ECWP, villa F 2 - 7, 33.00722 lat. - 4.0976 lon., ID 14771, 24 / VII / 2013 (J. F. Léger), ECWP; - Morocco, Missour, ECWP, villa F 2 - 7, 33.00722 lat. - 4.0976 lon., ID 15532, 19 / XI / 2013 (A. François), ECWP.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A30412DFC09FDDB90AF366A.taxon	materials_examined	Material examined (54 ex.). - Morocco, Bouârfa, Aïn Larak, 32.60435 lat. - 2.50391 lon., ID 18770, 11 / III / 2014 (A. François), ECWP; - Morocco, Debdou, 34.06452 lat. - 2.98199 lon., ID 14938, 08 / VI / 2013 (A. François, L. Castro), ECWP; - Morocco, betweenAïn Bni Mattar and Maatarka, 34.020708 lat. - 2.12397 lon., ID 12440, 05 / VII / 2012 (A. François, C. Galkowski, M. Sbai), ECWP; - Morocco, between Outat el Haj and Maatarka, 33.58163 lat. - 3.31162 lon., ID 8338, 24 / V / 2011 (A. François, H. Hdidou), ECWP; - Morocco, between Outat el Haj and Maatarka, 33.57178 lat. - 3.36583 lon., ID 8339, 24 / V / 2011 (A. François, H. Hdidou), ECWP; - Morocco, between Outat el Haj and Maatarka, 33.56384 lat. - 3.23189 lon., ID 8340, 24 / V / 2011 (A. François, H. Hdidou), ECWP; - Morocco, Maatarka, 33.21395 lat. - 2.78115 lon., ID 8822, 26 / IV / 2011 (M. Chambouleyron, H. Hdidou), ECWP; - Morocco, Maatarka, 33.23927 lat. - 2.74412 lon., ID 10556, 15 / VI / 2010 (H. Hdidou), ECWP; - Morocco, Maatarka, 33.23927 lat. - 2.74412 lon., ID 10560, 11 / VI / 2010 (H. Hdidou), ECWP; - Morocco, Oriental, 32.9859 lat. - 3.60719 lon., ID 21741, 14 / IV / 2015 (T. Dieuleveut, C. Lucas), ECWP; - Morocco, Almis des Marmoucha, Jbel Tsiouant, 32.2745 lat. - 4.17783 lon., ID 14992, 15 / IV / 2013 (A. François, J. Gabbard), ECWP; - Morocco, Enjil, 33.12101 lat. - 4.59676 lon., ID 10570, 12 / VI / 2010 (H. Hdidou), ECWP; - Morocco, Enjil, 33.2113 lat. - 4.6638 lon., ID 18623, 15 / IV / 2014 (A. François), ECWP; - Morocco, Enjil, ECWP, 33.14769 lat. - 4.58389 lon., ID 21769 / 21783, 20 / V / 2014 (J. M. Touzet), ECWP; - Morocco, Enjil, ECWP, 33.15998 lat. - 4.58624 lon., ID 10569, 09 / VI / 2010 (H. Hdidou), ECWP; - Morocco, Enjil, ECWP, 33.15049 lat. - 4.606 lon., ID 22916, 16 / V / 2014 (M. Chambouleyron, H. Hdidou), ECWP; - Morocco, Enjil, ECWP, 33.14769 lat. - 4.58389 lon., ID 22917, 20 / V / 2014 (J. M. Touzet), ECWP; - Morocco, Enjil, ECWP, gazelle pen, 33.17226 lat. - 4.41112 lon., ID 10568, 13 / VI / 2010 (H. Hdidou), ECWP; - Morocco, Enjil, Enjil lake, hillbetween lake andvillage, 33.22819 lat. - 4.66391 lon., ID 585, 21 / III / 2008 (A. François), ECWP; - Morocco, Lamjalil, 33.89885 lat. - 3.94569 lon., ID 10571, 20 / VII / 2009 (H. Hdidou), ECWP; - Morocco, Lamjalil, 33.87449 lat. - 3.98817 lon., ID 10573, 18 / VI / 2010 (H. Hdidou), ECWP; - Morocco, Midelt Aouli, abandoned mining village, 32.81851 lat. - 4.58497 lon., ID 14993, 05 / VI / 2013 (L. Bacon), ECWP; - Morocco, Missour, Al Baten, 33.22951 lat. - 4.07098 lon., ID 582, 15 / VI / 2002 (Y. Hingrat, J. Yvernault), ECWP; - Morocco, Missour, Al Baten, 33.20759 lat. - 4.05286 lon., ID 588, 19 / XII / 2001 (Y. Hingrat, J. Yvernault), ECWP; - Morocco, Missour, Al Baten, 33.22951 lat. - 4.07098 lon., ID 589, 15 / II / 2002 (Y. Hingrat, J. Yvernault), ECWP; - Morocco, Missour, Al Baten, 33.22951 lat. - 4.07098 lon., ID 590, 17 / V / 2002 (Y. Hingrat, J. Yvernault), ECWP; - Morocco, Missour, Al Baten, 33.29897 lat. - 4.80173 lon., ID 593, 22 / V / 2002 (Y. Hingrat, J. Yvernault), ECWP; - Morocco, Missour, Al Baten, 33.1997 lat. - 3.92071 lon., ID 2795, 16 / IV / 2002 (J. Yvernault), ECWP; - Morocco, Missour, Al Baten, 33.17877 lat. - 3.94202 lon., ID 2796, 22 / IV / 2002 (J. Yvernault), ECWP; - Morocco, Missour, Al Baten, 33.2294 lat. - 4.07156 lon., ID 2797, 22 / IV / 2002 (J. Yvernault), ECWP; - Morocco, Missour, Al Baten, 33.16887 lat. - 3.94317 lon., ID 2809, 16 / V / 2002 (Y. Hingrat), ECWP; - Morocco, Missour, Douar Igli, 33.00586 lat. - 3.94871 lon., ID 10557, 15 / VI / 2010 (( H. Hdidou), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 23250, 09 / V / 2016 (M. Sbai), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 581, 24 / IV / 2008 (A. François, E. Le Nuz, G. Levêque), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 586, 15 / VI / 2000 (Y. Hingrat), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 7105, 07 / X / 2010 (H. Chaker, H. Hdidou, J. F. Léger), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 10011, 30 / IX / 2011 (A. Chlih), ECWP; - Morocco, Missour, ECWP, 33.00251 lat. - 4.10432 lon., ID 10572, 07 / VI / 2010 (H. Hdidou), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 13382, 19 / X / 2012 (G. Levêque), ECWP; - Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 24202, 23 / VIII / 2021 (M. Sriri), ECWP; - Morocco, Missour, ECWP, insemination laboratory, 33.00722 lat. - 4.0976 lon., ID 8829, 07 / VII / 2011 (N. Abbou), ECWP; - Morocco, Missour, ECWP, villa F 2.10, 33.00722 lat. - 4.0976 lon., ID 15034, 05 / IX / 2013 (A. François), ECWP; - Morocco, Missour, ECWP, villa F 2.3, 33.00722 lat. - 4.0976 lon., ID 14937, 10 / VIII / 2013 (E. Le Nuz), ECWP; - Morocco, Missour, ECWP, villa F 2.6, 33.0119 lat. - 4.09692 lon., ID 6450, 20 / VII / 2010 (A. François), ECWP; - Morocco, Missour, ECWP, villa F 2.6, 33.0119 lat. - 4.09692 lon., ID 10433, 22 / X / 2011 (A. François), ECWP; - Morocco, Missour, Jbel Missour, pic, 33.00133 lat. - 4.16963 lon., ID 5703, 24 / VI / 2010 (A. François, K. Benlafkih, M. Sbai), ECWP; - Morocco, Ouled Ali Atchana, 33.5307 lat. - 3.92263 lon., ID 14902 / 14935, 03 / VIII / 2013 (A. François, A. Achou, C. Heyd, V. Liéron), ECWP; - Morocco, OuledAli BouNaceur, Tizi n’Silla, 33.56052 lat. - 3.90465 lon., ID 14936, 04 / VIII / 2013 (A. François, A. Achou, C. Heyd, V. Liéron), ECWP; - Morocco, Tighnest, 33.4051 lat. - 3.7733 lon., ID 10559, 17 / VI / 2010 (H. Hdidou), ECWP; - Morocco, Tighnest, 33.40538 lat. - 3.77373 lon., ID 10563, 07 / VI / 2010 (H. Hdidou), ECWP; - Morocco, Tighnest, 33.42168 lat. - 3.79447 lon., ID 10564, 10 / VI / 2010 (H. Hdidou), ECWP; - Morocco, Zerzaia, 33.65271 lat. - 3.47184 lon., ID 21784, 20 / IV / 2015 (A. François), ECWP.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A3E412DFC0DFF3C97733C9A.taxon	materials_examined	Material examined (5 ex.). - Morocco, Bou Iblane, 33.64595 lat. - 4.10403 lon., ID 592, 11 / X / 2008 (A. François, H. Hdidou), ECWP; - Morocco, Taffert, refuge, 33.68344 lat. - 4.01114 lon., ID 4969 / 4970, 12 / IV / 2010 (A. François, H. Hdidou), ECWP; - Morocco, Taffert, refuge, 33.64949 lat. - 4.16266 lon., ID 5704, 01 / V / 2010 (A. François, C. Galkowski, M. Sbai), ECWP; - Morocco, Almis des Marmoucha, ID 587, 31 / III / 2002 (J. Yvernault), ECWP.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A3E412DFC0DFF3C97733C9A.taxon	distribution	Distribution and remarks. – B. lienhardi, described from the high Atlas in the south of Marrakech, is widely distributed throughout the highAtlas (Vachon, 1952, Lourenço, 2003, 2017, Touloun et al., 2014, Dupré, 2017, ElHidan etal., 2017, Sousa et al., 2017, Touloun, 2019). Thisisthe first reportof B. lienhardi from the Oriental region, where it was collected in the Djebel Bou Iblane (Fig. 2).	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A3E412DFC0AFD1C96823652.taxon	materials_examined	Material examined (12 ex.). - Morocco, Jerada, pass between Guenfouda and Jerada, 34.37752 lat. - 2.17113 lon., ID 12436, 04 / VII / 2012 (A. François, C. Galkowski, M. Sbai), ECWP; - Morocco, Nador, Gourougou, 35.20045 lat. - 3.00725 lon., ID 13380, 06 / X / 2012 (A. François), ECWP; - Morocco, Oujda, Beni Snassen, forest house El Aïn Almou, 34.84735 lat. - 2.20283 lon., ID 12430, 01 / VII / 2012 (A. François, C. Galkowski, M. Sbai), ECWP; - Morocco, Oujda, Jbel Bou Keltoum, pic, 34.3701 lat. - 2.08187 lon., ID 12435, 04 / VII / 2012 (A. François, C. Galkowski, M. Sbai), ECWP; - Morocco, Oujda, Tafoughalt, Beni Snassen, 34.84771 lat. - 2.33716 lon., ID 12429 / 12464, 01 / VII / 2012 (A. François, C. Galkowski, M. Sbai), ECWP; - Morocco, Taourirt, Beni Snassen West, 34.81288 lat. - 2.70916 lon., ID 12434, 03 / VII / 2012 (A. François, C. Galkowski, M. Sbai), ECWP; - Morocco, Taourirt, Jbel Khouali, antenna, 34.3351 lat. - 2.56463 lon., ID 12433, 02 / VII / 2012 (A. François, C. Galkowski, M. Sbai), ECWP; - Morocco, Taourirt, Tanacherfi, Horsts range, 34.33006 lat. - 2.60019 lon., ID 12431 / 12432, 02 / VII / 2012 (A. François, C. Galkowski, M. Sbai), ECWP; - Morocco, Saka, 34.5543 lat. - 3.48417 lon., 1558 m alt., ID 25071, 27 / XI / 2022 (A. François, G. Liénart, C. Landsmann, O. Chabasseur, J. F. Lima Barbero, R. Triguero Ocana), ECWP; - Morocco, Saka, 34.58319 lat. - 3.51407 lon., 1790 m alt., ID 25071, 27 / XI / 2022 (A. François, G. Liénart, C. Landsmann, O. Chabasseur, J. F. Lima Barbero, R. Triguero Ocana), ECWP; - Morocco, 9 km N Tafoughalt, 240 m alt., EY 0381, X / 2014 (O. Touloun), W. R. Lourenço leg., 26 / III / 2022, EYPC (misidentified as B. confluens (Lourenço, Touloun & Boumezzough, 2011) in Touloun et al., 2014).	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A3C412FFF22FF4190FD3AEA.taxon	materials_examined	Material examined (11 ex.). - Morocco, Bouârfa, Tamlelt, 32.50613 lat. - 2.6698 lon., ID 10526, 19 / VI / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.51384 lat. - 2.48154 lon., ID 10528, 10 / VII / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.50613 lat. - 2.6698 lon., ID 10529, 29 / V / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.31528 lat. - 2.32572 lon., ID 10538, 28 / VIII / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.53419 lat. - 2.50649 lon., ID 10542, 19 / VI / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.48212 lat. - 2.7282 lon., ID 10546, 17 / VII / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.25143 lat. - 2.22581 lon., ID 10547, 16 / VII / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.51384 lat. - 2.48154 lon., ID 10548, 03 / VII / 2008 (S. Touil), ECWP; - Morocco, Bouârfa, Tamlelt, 32.53385 lat. - 2.5066 lon., ID 12541, 03 / VII / 2008 (S. Touil), ECWP; - Morocco, Bouânane, Zelmou dam, 32.13496 lat. - 2.8807 lon., ID 8482, 11 / VI / 2011 (A. François, C. Landsmann), ECWP; - Morocco, between Bouârfa and Figuig, 32.25457 lat. - 1.71231 lon., ID 22918, 24 / V / 2015 (A. François, M. García-Paris, J. L. Ruiz), ECWP.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A3C412FFF26FAF193E13889.taxon	materials_examined	Material examined (3 ex.). - Morocco, Anoual, 32.64221 lat. - 3.12263 lon., ID 580, 24 / V / 2008 (A. François, F. Lagarde), ECWP; - Morocco, Bouârfa, Aïn Larak, 32.60435 lat. - 2.50391 lon., ID 18812, 11 / III / 2014 (A. François), ECWP; - Morocco, Oued Bour, 32.73369 lat. - 3.21239 lon., ID 21823, 07 / VII / 2015 (T. Dieuleveut), ECWP.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A3C4129FEEFF88996CF3AE9.taxon	description	(Fig. 5 - 8, 13 - 14, 17 - 18, 21, 29, 32) ZooBank: http: // zoobank. org / 761 CC 1 D 8 - 7 C 61 - 48 D 0 - A 282 - 9 DE 3791 A 55 CB	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A3C4129FEEFF88996CF3AE9.taxon	materials_examined	Holotype, ♂, Morocco, Bouârfa, Tamlelt, 32.43128 lat. - 2.57354 lon., ID 10549, 11 / III / 2015 (S. Touil), deposited in the MNHN. Paratypes (3 ex.). - 1 ♂, Morocco, Bouârfa, Tamlelt, 32.44389 lat. - 2.29296 lon., ID 10541, 11 / III / 2015 (S. Touil), deposited in the ECWP; - 1 ♂, Morocco, Bouârfa, Tamlelt, 32.44350 lat. - 2.29312 lon., ID 10554, 11 / III / 2015 (S. Touil), deposited in the ECWP; - 1 ♀, Morocco, betweenAïnBniMattar and Maatarka, 34.02078 lat. - 2.12397 lon., ID 12441, 31 / XII / 2012 (A. François, C. Galkowski, M. Sbai), deposited in the MNHN. Other material examined (9 ex.). - 1 ♂, Morocco, Bouârfa, Tamlelt, 32.53565 lat. - 2.55283 lon., ID 10530, 22 / VIII / 2008 (S. Touil), ECWP; - 1 ♂, Morocco, Bouârfa, Tamlelt, 32.43095 lat. - 2.57379 lon., ID 10533, 22 / VIII / 2008 (S. Touil), ECWP; - 1 ♂, Morocco, Bouârfa, Tamlelt, 32.43033 lat. - 2.43403 lon., ID 10592, 26 / VI / 2008 (S. Touil), ECWP; - 1 ♂, Morocco, Bouârfa, Tamlelt, 32.44563 lat. - 2.39064 lon., ID 20682, 31 / III / 2015 (J. Gabbard), ECWP; - 1 ♂, Morocco, betweenAïnBniMattar and Maatarka, 34.02078 lat. - 2.12397 lon., ID 12438, 06 / VII / 2012 (A. François, C. Galkowski, M. Sbai), ECWP; - 1 ♂, Morocco, Tendrara, MHNL 47036246, 1984 (G. Chavanon), MHNL; - 1 pre-adult ♂, Morocco, Figuig, MHNL 47036248, 1979 (G. Chavanon), MHNL; - 1 ♂, Morocco, Anoual, 32.65387 lat. - 3.13153 lon., ID 574, 24 / V / 2008 (A. François), ECWP; - 1 ♂, Morocco, Léthima, 32.72258 lat. - 2.74343 lon., ID 577, 13 / III / 2008 (M. Lelièvre, L. Pomarède), ECWP. Comparative material examined (2 ex.). Scorpio punicus Fet, 2000: - Tunisia, Gafsa, 1 ♂, MHNL 44003128, 1903 (Fracques), MHNL; - Tunisia, El Djennah, 1 ♀, EY 0264, J. - B. Lacroix leg. (No. 148), 1993, EYPC.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A3C4129FEEFF88996CF3AE9.taxon	etymology	Etymology. – The specific name honours Mr. Soufiane Touil, staff member of ECWP, who recently passed away. He used to be involved in ECWP’s arthropods sampling, and was greatly appreciated for his engagement and conscientiousness. He is also collector of the holotype of the new species.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A3C4129FEEFF88996CF3AE9.taxon	diagnosis	Diagnosis. – Scorpion of moderate to large size for the genus, with a total length of 61 - 64 mm for the males holotype and paratypes and 57 mm for the female paratype. Coloration basically yellowish to yellowish brown, without delimited darker pigmentation on carapace and tergites; chela manus of similar colour as body. Genital operculum suboval, formed by two plates having a semi-triangular shape; convex anteriorly and posteriorly in both sexes. Pectinal plate weakly divided in two parts, the posterior part slightly wider than the anterior part; pectines longer than length of third coxa and largely exceeding the distal end of fourth coxa in male, slightly longer than length of third coxa and slightly exceeding the distal end of fourth coxa in female; pectinal count 9 - 11 teeth in male, 9 - 11 in female. Leg IV tarsus with 9 internal and 7 external spines. Hemispermatophore with distal lamina curved, terminus enlarged and almost flat; laminar hook close to lamina, long and almost straight.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A3C4129FEEFF88996CF3AE9.taxon	description	Description (based on male holotype and female paratype; measurements after the description). Coloration. – Prosoma: carapace yellowish brown with diffuse brownish variegated spots on the entire surface; median and lateral ocular tubercles marked with dark pigments. Mesosoma: tergites yellowish brown; sternites yellowish. Coxapophysis and sternum yellow; genital operculum andpectines yellow in male, pale yellow in female. Metasoma: yellowish with some greyishpigmentation on ventral face of segments I-III inmale, on ventralface of all segments and on lateral and dorsal faces of segment V in female. Telson yellow; aculeus yellowish at its base and black at its extremity. Chelicerae yellow without any variegated spots in male, with some greyish pigmentation at the base in female; fingers yellow with reddish teeth. Pedipalps: femur and patella yellowish brown; chela yellowish brown with carinae and fingers darker, reddish black in male, reddish brown in female; dentate margins of fingers black. Legs yellowish. Morphology. – Carapace acarinate, without any granulations; anterior margin with a moderately (male) to strongly (female) pronounced concavity; posterior furrows moderatelypronounced; median ocular tubercle in the centre of the carapace; three pairs of lateral eyes; the first two of equal size, the third slightly reduced. Mesosoma: tergites acarinate and almost smooth, with only some sparse minute granulation on posterior and lateral sides. Sternum pentagonal, slightly wider than high. Venter: genital operculum suboval, formed by two plates having a semi-triangular shape; convex anteriorly and posteriorly in both sexes. Pectinal plate weakly divided in two parts, the posterior part slightly wider than the anterior part. Pectines longer than length of third coxa and largely exceeding the distal end of fourth coxa in male, slightly longer than length of third coxa and slightly exceeding the distal end of fourth coxa in female; pectinal tooth count 9 - 10 in male, 9 - 11 in female; fulcra strongly developed. Sternites smooth and shiny; VII with four moderately marked carinae; spiracleslinear andconspicuous. Metasomawithmoderatelytostrongly markedcarinaeonsegmentsI to IV; granulation becomes spiniform on segment V; ventral and latero-ventral carinae intensely spinoid on V; all intercarinal surfaces weakly granular. Telson globular and strongly granular on ventral side with four ventralcarinae formedbystrong spinoid granules; aculeus shorter than vesicle and moderately curved. Cheliceral dentition characteristic of the Scorpionidae (Vachon, 1963); movable finger with one subdistal tooth and conspicuous basal teeth. Pedipalps: femur with four incomplete carinae, intercarinal surfaces smooth to weakly granulated; patella with dorsal carina almost complete, intercarinal surfaces smooth to weakly granulated; chela with weakly marked ventral carinae; dorsal carinae moderately marked; dorso-external aspect of the manus coarsely granular, especially in male. Dentate margin on fixed and movable fingers with a series of granules divided by 5 strong accessory granules. Trichobothriotaxy of type C; orthobothriotaxic (Vachon, 1974); femur with 3 trichobothria, patella with 19 and chela with 26. Legs: tarsi of legs I to IV with 7 / 5, 8 - 9 / 6, 9 / 7, 9 / 7 internal and external spines arranged in series. Hemispermatophore: distal lamina curved, terminus enlarged and almost flat; laminar hook close to lamina, long and almost straight. Morphometric values (mm) (male holotype and female paratype). – Total length (including telson): 63.63 / 57.15. – Carapace length, 10.25 / 8.13; anterior width, 5.63 / 5.38; posterior width, 8.75 / 8.13. – Mesosoma: length, 22.50 / 22.63. – Metasomal segments I: length, 3.63 / 3.13; width, 4.50 / 3.88; II: length, 4.0 / 3.50; width, 4.0 / 3.63; III: length, 4.50 / 3.75; width, 4.0 / 3.38; IV: length, 5.25 / 4.38; width, 3.88 / 3.25; V: length, 7.25 / 6.0; width, 3.63 / 2.63; depth, 3.25 / 2.75. – Telson length, 6.25 / 5.63; vesicle width, 2.75 / 2.63; depth, 2.50 / 2.25. – Pedipalp femur length, 5.63 / 4.88; width, 2.63 / 2.38; patella length, 5.63 / 6.0; width, 3.13 / 2.75; chela length, 11.75 / 11.38; width, 8.38 / 6.88; depth, 4.13 / 3.88; – Movable finger length, 6.75 / 6.75. Comparisons. – By its light coloration and the general shape of its pectinal plate and male genital operculum plate, Scorpio touili sp. n. seems to be more closely related to Scorpio punicus Fet, 2000 (formerly known as Scorpio maurus tunetanus Birula, 1910), a yellowish brown species largely distributed in the high plateaus, the Saharan Atlas and Aures mountains and the northern edge of the Sahara Desert in Tunisia and Algeria, and the new species might represent the ‘ occidental form’ of S. punicus (to which it has been referred in Touloun et al., 2014). S. touili sp. n. can however be easily distinguished from S. punicus notably by the following main features: (i) pectinal plate with posterior part slightly wider than anterior part (same width in S. punicus; Fig. 21, 25, 29 - 30); (ii) male pectine largely exceeding the distal end of the fourth coxa (reaching or only slightly exceeding the distal end of the fourth coxa in S. punicus); (iii) female pectine length slightly longer than length of third coxa (as long as third coxa in S. punicus); (iv) male genital operculum plate generally slightly less flattened with high / length ratio 0.7 (high / length ratio 0.6 in S. punicus; Fig. 21, 25); (v) female genital operculum plate more convex anteriorly (more flattened anteriorly in S. punicus; Fig. 29 - 30); (vi) hemispermatophore with laminar hook almost straight (more curved in S. punicus, Fig. 32, 34); (vii) leg IV tarsus with 9 internal and 7 external spines (n = 4) (8 internal and 6 - 7 external spines in S. punicus). S. touili sp. n. can also be easily distinguished from another new species described in this work, S. moulouya sp. n., mainly occurring in the middle Moulouya river basin, but with some potential zones of sympatry between both species (Fig. 4), notably by the following main features: (i) an overall lighter coloration, yellowish to yellowish brown (brownish to dark brown in S. moulouya sp. n.); (ii) chela manus of similar colour as body (lighter than body in S. moulouya sp. n.); (iii) hemispermatophore with distal lamina more curved (almost straight in S. moulouya sp. n.), lamina terminus enlarged and almost flat (not enlarged and ending with a 45 ° angle in S. moulouya sp. n.) and laminar hook longer and closer to lamina (smaller and further apart from lamina in S. moulouya sp. n., Fig. 32 - 33). Finally, S. touili sp. n. can be easily distinguished from Scorpio atlasensis Khammassi, Harris & Sadine, 2023, a species recently described from the Tellian Atlas of north-western Algeria (Khammassi et al., 2023) and reported in Morocco from two locations in the high plateaus and Saharan Atlas, by the following main features: (i) carapace and mesosoma without delimited darker pigmentation (dark triangular zone on carapace and median dark line on mesosoma in S. atlasensis sp. n.); (ii) female genital operculum plate more convex anteriorly (more flattened anteriorly in S. atlasensis; Fig. 29, 31); (iii) leg IV tarsus with 9 internal and 7 external spines (n = 4) (8 internal and 7 external spines in S. atlasensis. NB. internal and external spine numbers seem to be inversed in Khammassi et al., 2023).	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A3A412BFBC6FB2996233D52.taxon	description	(Fig. 9 - 10, 15, 19, 22) ZooBank: http: // zoobank. org / 5598657 E- 7 F 40 - 4 CBA- 9 BC 8 - 0 C 68 C 1 ED 1 EC 8	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A3A412BFBC6FB2996233D52.taxon	materials_examined	Holotype, ♂, Morocco, Guerbous pass, MHNL 47036249, 11 / III / 1984 (G. Chavanon), deposited in the MHNL. Paratypes (2 ex.). - 1 ♂, Morocco, Guerbous pass, MHNL 47036247, 26 / II / 1984 (G. Chavanon), deposited in the MHNL; - 1 ♀ (right chela), Morocco, Beni Snassen, Tafoughalt, 34.84771 lat. - 2.33716 lon., ID 16569, 01 / VII / 2015 (A. François, C. Galkowski, M. Sbai), deposited in theECWP. Comparative material examined (3 ex.). Scorpio maurus Linnaeus, 1758: Algeria, Zurich (= Sidi Amar), 2 ♂, 1 ♀, MHNL 44003122, 1889 (Massard), MHNL.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A3A412BFBC6FB2996233D52.taxon	etymology	Etymology. – The specific name is placed in apposition to the generic name and refers to the Beni Snassen (Aït Iznassen in Tamazight) tribe, living in the area where the new species occurs.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A3A412BFBC6FB2996233D52.taxon	diagnosis	Diagnosis. – Scorpion of moderate size for the genus, with a total length of 50 - 58 mm for the males holotype and paratype. Coloration basically blackish brown. Genital operculum suboval, formed by two plates having a semi-triangular shape; convex anteriorly and flattened posteriorly. Pectinal plate strongly constricted medially; pectines slightly longer than length of third coxa and slightly exceeding the distal end of fourth coxa in male; pectinal count 9 - 10 teeth in male. Chela fixed finger internal length shorter than manus ventral length in male. Leg IV tarsus with 9 internal and 7 external spines.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A3A412BFBC6FB2996233D52.taxon	description	Description (based on male holotype; measurements after the description). Coloration. – Prosoma: carapace blackish brown with diffuse black variegated spots on the entire surface; median and lateral ocular tubercles marked with black pigments. Mesosoma: tergites blackish brown; sternites reddish black. Coxapophysis and sternum reddish yellow; genital operculum and pectines yellowish. Metasoma: dark brown with blackish pigmentation on ventral carinae of all segments. Telson yellowish brown with some brownish pigmentation on ventral and lateral surfaces; aculeus reddish at its base and black at its extremity. Chelicerae yellowish with brownish variegated spots close to the fingers; fingers yellowish with reddish teeth. Pedipalps: femur and patella brownish with carinae blackish; chela reddish brown with carinae and fingers blackish; dentate margins of fingers black. Legs brownish. Morphology. – Carapace acarinate, without any granulations; anterior margin with a strongly pronounced concavity; posterior furrows moderately pronounced; median ocular tubercle in the centre of the carapace; three pairs of lateral eyes; the first two of equal size, the third slightly reduced. Mesosoma: tergites acarinate and minutely granular, coarser on VII. Sternum pentagonal, slightly wider than high. Venter: genital operculum suboval, formed by two plates having a semi-triangular shape; convex anteriorly and flattened posteriorly. Pectinal plate strongly constricted medially. Pectines slightly longer than length of third coxa and slightly exceeding the distal end of fourth coxa; pectinal tooth count 9 - 10 in male; fulcra strongly developed. Sternites smooth and shiny; VII with four moderately marked carinae; spiracles linear and conspicuous. Metasoma with moderately to strongly marked carinae on segments I to IV; granulation becomes spiniform on segment V; ventral and latero-ventral carinae intensely spinoid on V; all intercarinal surfaces weakly granular. Telson globular and strongly granular on ventral side with four ventral carinae formed by strong spinoid granules; aculeus shorter than vesicle and moderately curved. Cheliceral dentition characteristic of the Scorpionidae (Vachon, 1963); movable finger with one subdistal tooth and conspicuous basal teeth. Pedipalps: femur with four incomplete carinae, intercarinal surfaces smooth to weakly granulated; patella with dorsal carina almost complete, intercarinal surfaces smooth to weakly granulated; chela with weakly marked ventral carinae; dorsal carinae moderately marked; dorso-external aspect of the manus coarsely granular. Dentate margin on fixed and movable fingers with a series of granules divided by 5 strong accessory granules; fixed finger internal length shorter than manus ventral length. Trichobothriotaxy of type C; orthobothriotaxic (Vachon, 1974); femur with 3 trichobothria, patella with 19 and chela with 26. Legs: tarsi of legs I to IV with 7 / 5, 8 / 6, 8 / 7, 9 / 7 internal and external spines arranged in series. Morphometric values (mm) (male holotype). – Total length (including telson): 57.78. – Carapace length, 9.88; anterior width, 5.38; posterior width, 11.13. – Mesosoma: length, 16.0. – Metasomal segments I: length, 3.63; width, 5.13; II: length, 4.38; width, 4.88; III: length, 4.75; width, 4.75; IV: length, 5.63; width, 4.38; V: length, 6.88; width, 3.50; depth, 3.50. – Telson length, 6.63; vesicle width, 3.25; depth, 2.75. – Pedipalp femur length, 6.50, width, 3.0; patella length, 7.75, width, 3.75; chela length, 14.38, width, 9.50, depth, 6.0; movable finger length, 8.88. Comparisons. – By its general coloration and its pectinal plate strongly constricted medially, Scorpio iznassen sp. n. seems to be more closely related to Scorpio maurus Linnaeus, 1758, a dark species largely distributed on the northern flanks of Tellian Atlas in Tunisia and eastern Algeria, and the new species might represent the ‘ occidental form’ of S. maurus. S. iznassen sp. n. can however be easily distinguished from S. maurus notably by the following main features: (i) an overall darker coloration, blackish brown (reddish brown to dark reddish brown in S. maurus); (ii) a smaller overall size in male with 50 - 58 mm (60 - 70 mm in S. maurus); (iii) male pectine length slightly longer than length of third coxa (as long as third coxa in S. maurus); (iv) male genital operculum plate more flattened posteriorly (more convex posteriorly in S. maurus; Fig. 22, 24); (v) male chela fixed finger internal length shorter than chela manus ventral length (as long as manus ventral length in S. maurus). S. iznassen sp. n. can also be easily distinguished from the two other dark species present in northern Morocco, namely Scorpio birulai Fet, 1997 (formerly known as Scorpio maurus subtypicus Birula, 1910) described from the Fahs-Anjra Province and Scorpio hesperus Birula, 1910 described from the surroundings of Tanger, notably by the following main features: (i) general coloration blackish brown (reddish brown to dark reddish brown in S. birulai); (ii) pectinal plate strongly constricted medially (only slightly constricted in S. birulai and S. hesperus; Fig. 22, 26, 28); (iii) male pectine length slightly longer than length of third coxa (as long as third coxa in S. hesperus); (iv) male genital operculum plate suboval, flattened posteriorly (strongly subtriangular posteriorly in S. birulai and S. hesperus; Fig. 22, 26, 28); (v) male chela fixed finger internal length shorter than chela manus ventral length (longer than manus ventral length in S. hesperus); (vi) moreover, the new species present a totally allopatric distribution from S. birulai and S. hesperus, separated notably by the Moulouya river basin.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A384135FC3DFCD3908C3D32.taxon	description	(Fig. 11 - 12, 16, 20, 23, 33) ZooBank: http: // zoobank. org / 1 EE 98653 - BDC 8 - 469 D-BC 78 - B 8 A 488 E 19720	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A384135FC3DFCD3908C3D32.taxon	materials_examined	Holotype, ♂, Morocco, Missour, ECWP, in front of the Ecology laboratory, 33.00722 lat. - 4.09760 lon., ID 5313, 13 / III / 2015 (A. François, H. Hdidou), deposited in the MNHN. Paratypes (3 ex.). - 1 ♂, Morocco, Missour, 33.00768 lat. - 4.08838 lon., ID 10558, 11 / III / 2015 (H. Hdidou), deposited in the ECWP; - 1 ♂, Morocco, Lamjalil, 33.89382 lat. - 3.95681 lon., ID 10561, 11 / III / 2015 (H. Hdidou), deposited in the MNHN; - 1 ♂, Morocco, Tenzil, 33.59808 lat. - 3.16118 lon., ID 15016, 31 / XII / 2013 (T. Dieuleveut), deposited in the ECWP. Other material examined (9 ex.). - 1 ♂, Morocco, Missour, ECWP, 33.01021 lat. - 4.09602 lon., ID 12080, 15 / VIII / 2012 (A. François, B. Michel), ECWP; - 1 ♂, Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 12198, 15 / X / 2012 (C. Robin), ECWP; - 1 ♂, Morocco, Missour, ECWP, 33.00722 lat. - 4.0976 lon., ID 21822, 05 / VI / 2014 (S. Boullenger), ECWP; - 1 ♂, Morocco, Missour, ECWP, villa F 2 - 6, 33.01191 lat. - 4.09693 lon., ID 24231, 28 / VIII / 2021 (A. François), ECWP; - 1 ♂, Morocco, Missour, Al Baten, 33.16383 lat. - 4.01174 lon., ID 573, 10 / V / 2002 (J. Yvernault), ECWP; - 1 ♂, Morocco, Missour, Al Baten, 33.22951 lat. - 4.07098 lon., ID 2870, 17 / V / 2002 (Y. Hingrat, J. Yvernault), ECWP; - 1 ♂, Morocco, Lamjalil, Ouizghet, ID 576, 17 / VI / 2000 (A. François, G. Chavanon), ECWP; - 1 ♂, Morocco, Lamjalil, Ouizghet, 33.8872 lat. - 3.9641 lon., ID 4634, 24 / VIII / 2009 (H. Hdidou), ECWP; - 1 ♂, Morocco, Tenzil, ID 575, 17 / VI / 2000 (Y. Hingrat), ECWP. Comparative material examined (3 ex.). Scorpio fuliginosus (Pallary, 1928): Morocco, Djebel Toubkal, 1 ♂, 2 ♀, EY 0052, 2010 (E. Ythier), EYPC.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A384135FC3DFCD3908C3D32.taxon	etymology	Etymology. – The specific name is placed in apposition to the generic name and refers to the Moulouya River Basin, where the new species mainly occurs.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A384135FC3DFCD3908C3D32.taxon	diagnosis	Diagnosis. – Scorpion of moderate to large size for the genus, with a total length of 53 - 67 mm for the males holotype and paratypes. Coloration basically brownish to dark brown, without delimited darker pigmentation on carapace and tergites; chela manus lighter than body. Genital operculum suboval, formed by two plates having a semi-triangular shape; convex anteriorly and posteriorly. Pectinal plate weakly divided in two parts, the posterior part slightly wider than the anterior part; pectines longer than length of third coxa and largely exceeding the distal end of fourth coxa in male; pectinal count 9 - 11 teeth in male. Leg IV tarsus with 9 internal and 7 external spines. Hemispermatophore with distal lamina almost straight, terminus not enlarged and ending with a 45 ° angle; laminar hook apart from lamina, small and almost straight.	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
3D1487E00A384135FC3DFCD3908C3D32.taxon	description	Description (based on male holotype; measurements after the description). Coloration. – Prosoma: carapace brownish with diffuse dark brown variegated spots on the entire surface; median and lateral ocular tubercles marked with dark pigments. Mesosoma: tergites brownish to dark brown; sternites reddish yellow. Coxapophysis and sternum reddish yellow; genital operculum and pectines yellowish. Metasoma: yellowish brown with some blackishpigmentation on ventralface of all segments and onlateraland dorsal faces of segment V. Telson yellowish brown with some brownish pigmentation on ventral and lateral surfaces; aculeus yellowish at its base and black at its extremity. Chelicerae yellow with greyish variegated spots close to the fingers; fingers yellowish with reddish teeth. Pedipalps: femur and patella yellowish brown; chela yellowish brown with carinae and fingers reddish black; dentate margins of fingers black. Metasoma: yellowish brown. Morphology. – Carapace acarinate, without any granulations; anterior margin with a moderately pronounced concavity; posterior furrows moderately pronounced; median ocular tubercle in the centre of the carapace; three pairs of lateral eyes; the first two of equal size, the third slightly reduced. Mesosoma: tergites acarinate and almost smooth, with only some sparse minute granulation on posterior and lateral sides. Sternum pentagonal, slightly wider than high. Venter: genital operculum suboval, formed by two plates having a semi-triangular shape; convex anteriorly and posteriorly. Pectinal plate weakly divided in two parts, the posterior part slightly wider than the anterior part. Pectines longer than length of third coxa and largely exceeding the distal end of fourth coxa; pectinal tooth count 10 - 11 in male; fulcra strongly developed. Sternites smooth and shiny; VII with four moderately marked carinae; spiracles linear andconspicuous. Metasoma with moderatelyto strongly marked carinae on segments I to IV; granulation becomes spiniform on segmentV; ventral and latero-ventral carinae intensely spinoid on V; all intercarinal surfaces weakly granular. Telsonglobular and strongly granular on ventral side with four ventral carinae formed by strong spinoid granules; aculeus shorter than vesicle and moderately curved. Cheliceral dentition characteristic of the Scorpionidae (Vachon, 1963); movable finger with one subdistal tooth and conspicuous basal teeth. Pedipalps: femur with four incomplete carinae, intercarinal surfaces smooth to weakly granulated; patella with dorsal carina almost complete, intercarinal surfaces smooth to weakly granulated; chela with weakly marked ventral carinae; dorsal carinae moderately marked; dorso-external aspect of the manus coarsely granular. Dentate margin on fixed and movable fingers with a series of granules divided by 5 strong accessory granules. Trichobothriotaxy of type C; orthobothriotaxic (Vachon, 1974); femur with 3 trichobothria, patella with 19 and chela with 26. Legs: tarsi of legs I to IV with 7 / 5, 8 / 6, 9 / 7, 9 / 7 internal and external spines arranged in series. Hemispermatophore: distal lamina almost straight, terminus not enlarged and ending with a 45 ° angle; laminar hook apart from lamina, small and almost straight. Morphometric values (mm) (male holotype). – Total length (including telson): 66.71. – Carapace length, 9.50; anterior width, 6.50; posterior width, 9.75. – Mesosoma: 25.13. – Metasomal segments I: length, 3.69; width, 5.0; II: length, 4.13; width, 4.63; III: length, 4.38; width, 4.25; IV: length, 5.50; width, 3.88; V: length, 7.50; width, 3.13; depth, 3.13. – Telson length, 6.88; vesicle width, 3.25; depth, 2.88. – Pedipalp femur length, 6.25, width, 3.13; patella length, 6.25, width, 3.13; chela length, 13.75, width, 9.63, depth, 5.0; movable finger length, 7.88. Comparisons. – By the general shape of its pectinal plate and male genital operculum plate, Scorpio moulouya sp. n. seems to be more closely related to another new species described in this work, Scorpio touili sp. n., mainly occurring in the high plateaus of the Oriental region, but with some potential zones of sympatry between both species (Fig. 4). S. moulouya sp. n. can however be easily distinguished from S. touili sp. n. notably by the following main features: (i) an overall darker coloration, brownish to dark brown (yellowish to yellowish brown in S. touili sp. n.); (ii) chela manus lighter than body (similar colour as body in S. touili sp. n.); (iii) hemispermatophore with distal lamina almost straight (more curved in S. touili sp. n.), lamina terminus not enlarged and ending with a 45 ° angle (enlarged and almost flat in S. touili sp. n.) and laminar hook smaller and further apart from lamina (longer and closer to lamina in S. touili sp. n., Fig. 32 - 33); S. moulouya sp. n. can also be easily distinguished from another dark species to which it has previously been referred to (Touloun et al., 2014), Scorpio fuliginosus (Pallary, 1928) described from High Atlas flanks and notably the region of Toubkal, notably by the following main features: (i) an overall lighter coloration, brownish to dark brown (similar coloration but always darker in S. fuliginosus); (ii) chela manus lighter than body (same colour or only slightly lighter than body in S. fuliginosus) and vesicle lighter than metasoma (same colour as metasoma in S. fuliginosus); (iii) pectinal plate with posterior part slightly wider than anterior part (same width in S. fuliginosus; Fig. 23, 27); (iv) male pectine length longer than length of third coxa (as long as third coxa in S. fuliginosus) and largely exceeding the distal end of the fourth coxa (reaching the distal end of the fourth coxa in S. fuliginosus); (v) male genital operculum plate more convex anteriorly (more flattened anteriorly in S. fuliginosus; Fig. 23, 27); (vi) hemispermatophore with distal lamina terminus ending with a 45 ° angle in the opposite way of S. fuliginosus and with laminar hook smaller and further apart from lamina (longer and closer to lamina in S. fuliginosus, Fig. 33, 35); (vii) moreover, both species present a totally allopatric distribution. Finally, S. moulouya sp. n. can be easily distinguished from Scorpio atlasensis Khammassi, Harris & Sadine, 2023, a species recently described from the Tellian Atlas of north-western Algeria (Khammassi et al., 2023) and reported in Morocco from one location in the middle Moulouya river basin, by the following main features: (i) carapace and mesosoma brownish to dark brown without delimited darker pigmentation (yellowish to yellowish brown with a dark triangular zone on carapace and a median dark line on mesosoma in S. atlasensis); (ii) chela manus lighter than body (similar colour as body in S. atlasensis); (iii) leg IV tarsus with 9 internal and 7 external spines (n = 4) (8 internal and 7 external spines in S. atlasensis. NB. internal and external spine numbers seem to be inversed in Khammassi et al., 2023).	en	Ythier, Eric, François, Alexandre (2023): The scorpion fauna of the Oriental region in Morocco (Scorpiones: Buthidae, Scorpionidae) with description of three new species of the genus Scorpio Linnaeus, 1758. Faunitaxys 11 (3): 1-15, DOI: 10.57800/faunitaxys-11(3), URL: http://dx.doi.org/10.5281/zenodo.10622561
