taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
3C3587DEFF86457D7DCCF9910B9B93B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5280596/files/figure.png	https://doi.org/10.5281/zenodo.5280596	FIGURE 5. Principal components analysis (PCA) based on nine craniometric variables indicating scatter of individuals on PC1 and PC2 (a) and PC2 and PC3 (b) from six pre-defined taxon-groups of Ethiopian Otomys in relation to existing type specimens of fortior and degeni and new holotypes designated for new species (see Taxonomic Summary). Open squares = O. typus s.s., open triangles = O. fortior; open diamonds = O. yaldeni sp. nov. (= Sp. A); closed squares = O. helleri (= Sp. B); open circles = O. cheesmani sp. nov. (= Sp.C); closed diamonds = O. simiensis sp. nov. (= Sp. D); cross = degeni. Type specimens indicated; normal font indicates existing holotypes; bold font represents newly designated holotypes (see Taxonomic Summary).	FIGURE 5. Principal components analysis (PCA) based on nine craniometric variables indicating scatter of individuals on PC1 and PC2 (a) and PC2 and PC3 (b) from six pre-defined taxon-groups of Ethiopian Otomys in relation to existing type specimens of fortior and degeni and new holotypes designated for new species (see Taxonomic Summary). Open squares = O. typus s.s., open triangles = O. fortior; open diamonds = O. yaldeni sp. nov. (= Sp. A); closed squares = O. helleri (= Sp. B); open circles = O. cheesmani sp. nov. (= Sp.C); closed diamonds = O. simiensis sp. nov. (= Sp. D); cross = degeni. Type specimens indicated; normal font indicates existing holotypes; bold font represents newly designated holotypes (see Taxonomic Summary).	2011-09-13	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita		Zenodo	biologists	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita			
3C3587DEFF86457D7DCCF9910B9B93B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5280598/files/figure.png	https://doi.org/10.5281/zenodo.5280598	FIGURE 6. Canonical variates analysis (plot of first two canonical variates) of six defined taxon-groups of Ethiopian Otomys based on nine craniometric variables. Position of individuals form “unknown” sample from Oromiya, Jimma (asterisks) predicted a posteriori. Open squares = O. typus s.s. (Mt Guna); Closed squares = O. fortior; open circles = O. helleri (= “Sp. B”); closed circles = O. yaldeni sp. nov. (= “Sp. A”); open triangles = O. cheesmani sp. nov. (= “Sp. C”); closed triangles = O. simiensis sp. nov. (= “Sp. D”).	FIGURE 6. Canonical variates analysis (plot of first two canonical variates) of six defined taxon-groups of Ethiopian Otomys based on nine craniometric variables. Position of individuals form “unknown” sample from Oromiya, Jimma (asterisks) predicted a posteriori. Open squares = O. typus s.s. (Mt Guna); Closed squares = O. fortior; open circles = O. helleri (= “Sp. B”); closed circles = O. yaldeni sp. nov. (= “Sp. A”); open triangles = O. cheesmani sp. nov. (= “Sp. C”); closed triangles = O. simiensis sp. nov. (= “Sp. D”).	2011-09-13	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita		Zenodo	biologists	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita			
3C3587DEFF86457D7DCCF9910B9B93B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5280622/files/figure.png	https://doi.org/10.5281/zenodo.5280622	FIGURE 17(a–c). Photographs comparing dorsal, ventral and lateral views of the skulls of holotype of Otomys yaldeni (ZMMU 162594).	FIGURE 17(a–c). Photographs comparing dorsal, ventral and lateral views of the skulls of holotype of Otomys yaldeni (ZMMU 162594).	2011-09-13	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita		Zenodo	biologists	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita			
3C3587DEFF80457A7DCCFEA90E59950C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5280586/files/figure.png	https://doi.org/10.5281/zenodo.5280586	FIGURE 1. Map of east African montane and alpine zones showing distribution of type localities (large, open labeled squares) of forms previously attributed to (a) Otomys typus s. l. and (b) O. tropicalis s. l. Closed squares represent Museum cranial material examined. Circles enclosed localities assigned to OTU's for this study (in Ethiopia these represent hypothesized taxa based on cytogenetic, molecular and morphological data); numbers indicate OTU numbers in Table 1. Arrows indicate localities from which cytochrome-b sequences were available for this study. The asterisk indicates material from a specimen of uncertain identity assigned to cf. sungae.	FIGURE 1. Map of east African montane and alpine zones showing distribution of type localities (large, open labeled squares) of forms previously attributed to (a) Otomys typus s. l. and (b) O. tropicalis s. l. Closed squares represent Museum cranial material examined. Circles enclosed localities assigned to OTU's for this study (in Ethiopia these represent hypothesized taxa based on cytogenetic, molecular and morphological data); numbers indicate OTU numbers in Table 1. Arrows indicate localities from which cytochrome-b sequences were available for this study. The asterisk indicates material from a specimen of uncertain identity assigned to cf. sungae.	2011-09-13	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita		Zenodo	biologists	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita			
3C3587DEFF80457A7DCCFC340E3A9006.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5280586/files/figure.png	https://doi.org/10.5281/zenodo.5280586	FIGURE 1. Map of east African montane and alpine zones showing distribution of type localities (large, open labeled squares) of forms previously attributed to (a) Otomys typus s. l. and (b) O. tropicalis s. l. Closed squares represent Museum cranial material examined. Circles enclosed localities assigned to OTU's for this study (in Ethiopia these represent hypothesized taxa based on cytogenetic, molecular and morphological data); numbers indicate OTU numbers in Table 1. Arrows indicate localities from which cytochrome-b sequences were available for this study. The asterisk indicates material from a specimen of uncertain identity assigned to cf. sungae.	FIGURE 1. Map of east African montane and alpine zones showing distribution of type localities (large, open labeled squares) of forms previously attributed to (a) Otomys typus s. l. and (b) O. tropicalis s. l. Closed squares represent Museum cranial material examined. Circles enclosed localities assigned to OTU's for this study (in Ethiopia these represent hypothesized taxa based on cytogenetic, molecular and morphological data); numbers indicate OTU numbers in Table 1. Arrows indicate localities from which cytochrome-b sequences were available for this study. The asterisk indicates material from a specimen of uncertain identity assigned to cf. sungae.	2011-09-13	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita		Zenodo	biologists	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita			
3C3587DEFF8245787DCCFEC20F2B9157.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5280588/files/figure.png	https://doi.org/10.5281/zenodo.5280588	FIGURE 2. Giemsa stained karyotypes of (a) Otomys helleri (= “Sp. B”; n=6), O. fortior (n=6) and O.typus s.s. (n=1) (2n=58, NFa=58), (b) Otomys yaldeni sp. nov. (= “Sp. A”; n=1; 2n=56, NFa=54), and (c) Otomys simiensis sp. nov. (= “Sp. D”; n=4; 2n=54, NFa=54).	FIGURE 2. Giemsa stained karyotypes of (a) Otomys helleri (= “Sp. B”; n=6), O. fortior (n=6) and O.typus s.s. (n=1) (2n=58, NFa=58), (b) Otomys yaldeni sp. nov. (= “Sp. A”; n=1; 2n=56, NFa=54), and (c) Otomys simiensis sp. nov. (= “Sp. D”; n=4; 2n=54, NFa=54).	2011-09-13	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita		Zenodo	biologists	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita			
3C3587DEFF8245787DCCFEC20F2B9157.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5280596/files/figure.png	https://doi.org/10.5281/zenodo.5280596	FIGURE 5. Principal components analysis (PCA) based on nine craniometric variables indicating scatter of individuals on PC1 and PC2 (a) and PC2 and PC3 (b) from six pre-defined taxon-groups of Ethiopian Otomys in relation to existing type specimens of fortior and degeni and new holotypes designated for new species (see Taxonomic Summary). Open squares = O. typus s.s., open triangles = O. fortior; open diamonds = O. yaldeni sp. nov. (= Sp. A); closed squares = O. helleri (= Sp. B); open circles = O. cheesmani sp. nov. (= Sp.C); closed diamonds = O. simiensis sp. nov. (= Sp. D); cross = degeni. Type specimens indicated; normal font indicates existing holotypes; bold font represents newly designated holotypes (see Taxonomic Summary).	FIGURE 5. Principal components analysis (PCA) based on nine craniometric variables indicating scatter of individuals on PC1 and PC2 (a) and PC2 and PC3 (b) from six pre-defined taxon-groups of Ethiopian Otomys in relation to existing type specimens of fortior and degeni and new holotypes designated for new species (see Taxonomic Summary). Open squares = O. typus s.s., open triangles = O. fortior; open diamonds = O. yaldeni sp. nov. (= Sp. A); closed squares = O. helleri (= Sp. B); open circles = O. cheesmani sp. nov. (= Sp.C); closed diamonds = O. simiensis sp. nov. (= Sp. D); cross = degeni. Type specimens indicated; normal font indicates existing holotypes; bold font represents newly designated holotypes (see Taxonomic Summary).	2011-09-13	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita		Zenodo	biologists	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita			
3C3587DEFF8245787DCCFEC20F2B9157.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5280598/files/figure.png	https://doi.org/10.5281/zenodo.5280598	FIGURE 6. Canonical variates analysis (plot of first two canonical variates) of six defined taxon-groups of Ethiopian Otomys based on nine craniometric variables. Position of individuals form “unknown” sample from Oromiya, Jimma (asterisks) predicted a posteriori. Open squares = O. typus s.s. (Mt Guna); Closed squares = O. fortior; open circles = O. helleri (= “Sp. B”); closed circles = O. yaldeni sp. nov. (= “Sp. A”); open triangles = O. cheesmani sp. nov. (= “Sp. C”); closed triangles = O. simiensis sp. nov. (= “Sp. D”).	FIGURE 6. Canonical variates analysis (plot of first two canonical variates) of six defined taxon-groups of Ethiopian Otomys based on nine craniometric variables. Position of individuals form “unknown” sample from Oromiya, Jimma (asterisks) predicted a posteriori. Open squares = O. typus s.s. (Mt Guna); Closed squares = O. fortior; open circles = O. helleri (= “Sp. B”); closed circles = O. yaldeni sp. nov. (= “Sp. A”); open triangles = O. cheesmani sp. nov. (= “Sp. C”); closed triangles = O. simiensis sp. nov. (= “Sp. D”).	2011-09-13	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita		Zenodo	biologists	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita			
3C3587DEFF8245787DCCFEC20F2B9157.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5280622/files/figure.png	https://doi.org/10.5281/zenodo.5280622	FIGURE 17(a–c). Photographs comparing dorsal, ventral and lateral views of the skulls of holotype of Otomys yaldeni (ZMMU 162594).	FIGURE 17(a–c). Photographs comparing dorsal, ventral and lateral views of the skulls of holotype of Otomys yaldeni (ZMMU 162594).	2011-09-13	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita		Zenodo	biologists	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita			
3C3587DEFF8245787DCCFEC20F2B9157.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5280590/files/figure.png	https://doi.org/10.5281/zenodo.5280590	FIGURE 3a. Bayesian phylogeny inferred from cyt b sequences partitioned by codon position for 34 East African specimens of O. typus s.l. and O. tropicalis s.l. in relation to representatives of seven Otomys species and Parotomys brantsii from eastern, western and southern Africa. Representatives of nine murine taxa served as outgroups (see Appendix 1). Nodal support indices for the main clades (annotated 1 to 8) and subclades (a, b, c, d and e), as well as supported deeper nodes (A, H, I, M, N, O) are given beside the phylogram in the following order: Bayesian posterior probabilities (BPP)/ bootstrap support from 1000 replicates for maximum likelihood (MLbs), under an unpartitioned GTR+I+G model/ unweighted parsimony (UwPbs), six-parameter-weighted parsimony partitioned by codon positions (6PPbs)/ neighbour-joining using uncorrected p-distances (Njpbs)/ neighbour-joining using GTR+I+G-corrected distances (Njcbs). Estimated divergence dates (Myr) for the major lineages identified in the cyt b phylogeny are indicated to the left of each node. Bayesian estimation of divergence times was conducted under a relaxed clock model using BEAST v.1.4. Four well established murine fossil calibration points were specified (Pocock 1987; Rowe et al. 2008, see text).	FIGURE 3a. Bayesian phylogeny inferred from cyt b sequences partitioned by codon position for 34 East African specimens of O. typus s.l. and O. tropicalis s.l. in relation to representatives of seven Otomys species and Parotomys brantsii from eastern, western and southern Africa. Representatives of nine murine taxa served as outgroups (see Appendix 1). Nodal support indices for the main clades (annotated 1 to 8) and subclades (a, b, c, d and e), as well as supported deeper nodes (A, H, I, M, N, O) are given beside the phylogram in the following order: Bayesian posterior probabilities (BPP)/ bootstrap support from 1000 replicates for maximum likelihood (MLbs), under an unpartitioned GTR+I+G model/ unweighted parsimony (UwPbs), six-parameter-weighted parsimony partitioned by codon positions (6PPbs)/ neighbour-joining using uncorrected p-distances (Njpbs)/ neighbour-joining using GTR+I+G-corrected distances (Njcbs). Estimated divergence dates (Myr) for the major lineages identified in the cyt b phylogeny are indicated to the left of each node. Bayesian estimation of divergence times was conducted under a relaxed clock model using BEAST v.1.4. Four well established murine fossil calibration points were specified (Pocock 1987; Rowe et al. 2008, see text).	2011-09-13	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita		Zenodo	biologists	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita			
3C3587DEFF9F45627DCCFE8A0FDA9466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5280588/files/figure.png	https://doi.org/10.5281/zenodo.5280588	FIGURE 2. Giemsa stained karyotypes of (a) Otomys helleri (= “Sp. B”; n=6), O. fortior (n=6) and O.typus s.s. (n=1) (2n=58, NFa=58), (b) Otomys yaldeni sp. nov. (= “Sp. A”; n=1; 2n=56, NFa=54), and (c) Otomys simiensis sp. nov. (= “Sp. D”; n=4; 2n=54, NFa=54).	FIGURE 2. Giemsa stained karyotypes of (a) Otomys helleri (= “Sp. B”; n=6), O. fortior (n=6) and O.typus s.s. (n=1) (2n=58, NFa=58), (b) Otomys yaldeni sp. nov. (= “Sp. A”; n=1; 2n=56, NFa=54), and (c) Otomys simiensis sp. nov. (= “Sp. D”; n=4; 2n=54, NFa=54).	2011-09-13	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita		Zenodo	biologists	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita			
3C3587DEFF9F45627DCCFE8A0FDA9466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5280596/files/figure.png	https://doi.org/10.5281/zenodo.5280596	FIGURE 5. Principal components analysis (PCA) based on nine craniometric variables indicating scatter of individuals on PC1 and PC2 (a) and PC2 and PC3 (b) from six pre-defined taxon-groups of Ethiopian Otomys in relation to existing type specimens of fortior and degeni and new holotypes designated for new species (see Taxonomic Summary). Open squares = O. typus s.s., open triangles = O. fortior; open diamonds = O. yaldeni sp. nov. (= Sp. A); closed squares = O. helleri (= Sp. B); open circles = O. cheesmani sp. nov. (= Sp.C); closed diamonds = O. simiensis sp. nov. (= Sp. D); cross = degeni. Type specimens indicated; normal font indicates existing holotypes; bold font represents newly designated holotypes (see Taxonomic Summary).	FIGURE 5. Principal components analysis (PCA) based on nine craniometric variables indicating scatter of individuals on PC1 and PC2 (a) and PC2 and PC3 (b) from six pre-defined taxon-groups of Ethiopian Otomys in relation to existing type specimens of fortior and degeni and new holotypes designated for new species (see Taxonomic Summary). Open squares = O. typus s.s., open triangles = O. fortior; open diamonds = O. yaldeni sp. nov. (= Sp. A); closed squares = O. helleri (= Sp. B); open circles = O. cheesmani sp. nov. (= Sp.C); closed diamonds = O. simiensis sp. nov. (= Sp. D); cross = degeni. Type specimens indicated; normal font indicates existing holotypes; bold font represents newly designated holotypes (see Taxonomic Summary).	2011-09-13	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita		Zenodo	biologists	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita			
3C3587DEFF9F45627DCCFE8A0FDA9466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5280598/files/figure.png	https://doi.org/10.5281/zenodo.5280598	FIGURE 6. Canonical variates analysis (plot of first two canonical variates) of six defined taxon-groups of Ethiopian Otomys based on nine craniometric variables. Position of individuals form “unknown” sample from Oromiya, Jimma (asterisks) predicted a posteriori. Open squares = O. typus s.s. (Mt Guna); Closed squares = O. fortior; open circles = O. helleri (= “Sp. B”); closed circles = O. yaldeni sp. nov. (= “Sp. A”); open triangles = O. cheesmani sp. nov. (= “Sp. C”); closed triangles = O. simiensis sp. nov. (= “Sp. D”).	FIGURE 6. Canonical variates analysis (plot of first two canonical variates) of six defined taxon-groups of Ethiopian Otomys based on nine craniometric variables. Position of individuals form “unknown” sample from Oromiya, Jimma (asterisks) predicted a posteriori. Open squares = O. typus s.s. (Mt Guna); Closed squares = O. fortior; open circles = O. helleri (= “Sp. B”); closed circles = O. yaldeni sp. nov. (= “Sp. A”); open triangles = O. cheesmani sp. nov. (= “Sp. C”); closed triangles = O. simiensis sp. nov. (= “Sp. D”).	2011-09-13	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita		Zenodo	biologists	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita			
3C3587DEFF9F45627DCCFE8A0FDA9466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5280622/files/figure.png	https://doi.org/10.5281/zenodo.5280622	FIGURE 17(a–c). Photographs comparing dorsal, ventral and lateral views of the skulls of holotype of Otomys yaldeni (ZMMU 162594).	FIGURE 17(a–c). Photographs comparing dorsal, ventral and lateral views of the skulls of holotype of Otomys yaldeni (ZMMU 162594).	2011-09-13	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita		Zenodo	biologists	Taylor, Peter J.;Lavrenchenko, Leonid A.;Carleton, Michael D.;Verheyen, Erik;Bennett, Nigel C.;Oosthuizen, Carel J.;Maree, Sarita			
