taxonID	type	description	language	source
421787CFF667C879D9F8FB0AFB34F8E2.taxon	description	Cybocephalus nipponicus Endrödy-Younga, 1971: 244, f. 1. Type locality: Kawasawa, Japan. Cybocephalus nipponicus: Hayashi, 1978: 32 [description of larva]; — Hisamatsu, 1985: 172, pl. 27, fig. 27 [note, photo]; — Kirejtshuk, 1992: 209 [in key, figure]; — Tian & Yu, 1994 a: 120 [redescription]; — Yu & Tian, 1995: 35 [note]; — Tian & Peng, 1997: 35 [note]; — Lupi, 2002: 463 [note]; — Tian, 2006: 63 [list]; — Smith & Cave, 2006 a: 786 [redescription]; — Smith & Cave, 2007 a: 170 [note]; — Smith & Bailey, 2007: 506 [note]. Cybocephalus (Cybocephalus) nipponicus: Jelínek & Audisio, 2007: 490 [catalogue]. Specimens examined. [ISHIKAWA] 4 exs., Campus of Kanazawa Univ., Kanazawa City, 8. X. 1982, S. Hisamatsu leg. [TOCHIGI] 2 exs., Shiriuchi, Tochigi City, 3. I. 1993, H. Yoshitomi leg. (under the bark of Zelkova serrata (Thunb.) Makino.) [KANAGAWA] 2 exs., Odawara, 9. V. 1957, Y. Hirano leg.; 1 ex., Odawara, 22. I. 1973, Y. Furuki leg.; 1 ex., Oishi, 8. IX. 1979, Y. Hirano leg.; 1 ex., Kanogawa, Tanzawa, 18. V. 1986, Y. Hirano leg.; 1 ex., Myojin tôge, 12. VIII. 1993, Y. Hirano leg.; 2 exs., Sarugashima, Atsugi City, 25. IV. 1996, Y. Hirano leg. [TOKYO] 6 exs., Kinshi Park, 1. VIII. 1967, K. Morimoto leg.; 1 ex., Ohshima Is., 17. IV. 1974, Y. Furuki leg. [WAKAYAMA] 6 exs., Arita, 11. X. 1963, S. Nakao leg. [HYÔGO] 1 ex., Mt. Joryuji, nr. Toshima, Awajishima Is., 16. XI. 1972, S. Hisamatsu leg. [TOTTORI] 2 exs., Kurayoshi, 14. VIII. 1974, Y. Furuki leg. [YAMAGUCHI] 9 exs., Hagi, 25. X. 1967, S. Nakao et al. leg.; 1 ex., Mt. Shizukisan, Hagi, 9. IX. 1970, S. Hisamatsu leg. [HIROSHIMA] 10 exs., Ikuchijima Is., 18 ~ 21, IX. 1972, Y. Furuki leg.; 2 exs., Innoshima Is., 5 ~ 6. V. 1973, M. Sakai leg. [EHIME] 6 exs., Matsuyama City, 8. I. 1951, M. Miyatake leg.; 6 exs., Tarumi, Matsuyama City, 19. VI. 1953, T. Ishihara leg.; 1 ex., Matsuyama City, 4. VI. 1955, T. Tachikawa leg. (host: Pseudaulacaspis pentagona Targ.); 2 Ƥ, Matsuyama City, 16. IX. 1955, S. Hisamatsu leg.; 6 exs., Matsuyama, 15. VI. 1956, S. Hisamatsu leg.; 2 exs., Matsuyama City, 27. VI. 1956, T. Tachikawa leg. (collected from Pseudaulacaspis pentagona Targ.); 232 Ƥ, Matsuyama City, 20. VII. 1956, T. Tachikawa leg. (host: Pseudaulacaspis pentagona Targ.); 3 exs., Higashino, Matsuyama City, 9. IV. 1957, M. Miyatake leg.; 1 ex., Matsuyama City, 28. V. 1959, M. Miyatake leg.; 1 ex., Ohmishima Is., 16. VII. 1959, M. Miyatake leg; 2 exs., Ohmishima Is., 16. VII. 1959, M. Sato leg.; 2 exs., Uwajima, 27. IV. 1961, T. Tachikawa leg.; 1 ex., Matsuyama City, 15. VII. 1967, M. Miyatake leg.; 2 exs., Matsuyama City, 28. II. 1968, K. Hatta leg.; 2 exs., Matsuyama City, 11. IX. 1972, Y. Furuki leg. (collected from Pseudaulacaspis pentagona Targ.); 2 exs., Inokuchi, Ohmishima Is., 2. XII. 1973, K. Ito & TK. Ishihara leg.; 3 exs., Jo-hen, 25. VII. 1974, Y. Furuki leg.; 1 ex., Shiroyama, Matsuyama City, 21. II. 1974, Y. Furuki leg.; 34 exs., Tokunomori, Ohzu City, 13. X. 1994, Y. Utsunomiya leg.; 8 exs., Matsubara-izumi, Morimatsu-machi, Matsuyama City, 22. VIII. 2007, T. Kurihara leg.; 23 exs., Morimatsu-machi, Matsuyama City, 24. VIII. 2007, S-T. Hisamatsu leg.; 21 exs., Morimatsu-machi, Matsuyama City, 22. X. 2007, T. Kurihara leg. [KOCHI] 2 exs., Muroto, 7. VI. 1959, M. Miyatake leg.; 1 ex., Ashizurimisaki, 24. VII. 1960, M. Miyatake leg.; 1 ex., Okinoshima Is., 15. VI. 1961, S. Hisamatsu leg. [OITA] 1 ex., Tsukumi, 12. V. 1957, T. Tachikawa leg. [KUMAMOTO] 2 exs., Konose, Kuma Vil., 6. VI. 1967, S. Hisamatsu leg. [FUKUOKA] 6 exs., Tachibana, 13. IV. 1956, S. Nakao leg.; 1 ex., Itoshima, 14. V. 1968, M. Iga leg. (preying Pseudococcus citriculus Green) [NAGASAKI] 8 exs., Omura, 31. VIII. 1963, S. Nakao leg.; 60 exs., Kuchinotsu, 15. XI. 1977, M. Tanaka leg.; 3 exs., Mogi, 20. V. 1971, S. Hisamatsu leg.; 3 exs., Mt. Konpira ~ Suwa Shrine, 12. VII. 1973, S. Kinoshita leg.; 1 ex., Hirado, 5. VII. 1979, A. Oda leg.; 1 ex., Izuhara, Tsushima Is., 25. VIII. 1966, S. Nomoto leg.; 1 ex., Mt. Nanatsudake, Fukue Is., 2. V. 1972, S. Kinoshita leg. [KAGOSHIMA] 9 exs., Nagashima, 14. X. 1971, S. Nakao leg.; 1 ex., Miyanoura, 25. IV. 1975, S. Nagai leg.; 32 exs., Uragami, Amami-Ohshima Is., 20. IX. 1975, S. Nakao leg.; 1 ex., Nakazato, Kikaijima Is., 11. VII. 1996, K. Aita leg.; 1 ex., Kumano, Nakatane-chô, Tanegashima Is., 8. VII. 1974, S. Hisamatsu leg.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF667C879D9F8FB0AFB34F8E2.taxon	diagnosis	Diagnosis. This species differs from other Japanese congeners by the following characteristics: male with yellow head and pronotum; elytra with large punctures (Fig. 3 B), larger than an eye-facet at middle; median lobe of male genitalia (Fig. 4 C) in lateral view thin, strongly sinuate. Redescription. Length 1.38 – 1.64 (1.50) mm (n = 10), width 0.74 – 0.88 (0.82) mm (n = 10), height 0.46 – 0.44 (0.45) mm (n = 5).	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF667C879D9F8FB0AFB34F8E2.taxon	description	Male. Body convex, strongly shiny; dorsal surface almost glabrous. Coloration black; dorsal- and ventral side of head (excluding blackish mandibles and genae), antennae, pronotum, mesoventrite, pro- and mesothoracic legs yellow; margins of pronotum translucent; anterior margins of elytra also translucent, yellowish brown. Hind legs dark yellowish brown. Head extremely wide; vertex with few, minute punctures, sparsely covered with short, recumbent, yellowish setae. Anterior margin of clypeus (Fig. 4 H) with shallow, arcuate emargination at middle. Eyes large, prominent, distinctly visible in ventral aspect. Antennae (Fig. 4 I) with club asymmetrical, 3 - segmented; ninth and tenth antennal segments serrated along inner margin; apical margin of eleventh segment truncate; approximate ratio of each segment (n = 1) is 2.14: 1.68: 2.04: 1.16: 1.43: 1.09: 1.13: 1.00: 1.36: 1.55: 1.50. Pronotum convex, strongly transverse, 1.70 – 1.78 (1.74) times as wide as long (n = 5); anterior angles rounded; posterior angles widely rounded; dorsal disc with minute punctures; interspaces smooth. Scutellum (Fig. 2 D) subtriangular, apex broadly rounded. Elytra conjointly 0.84 – 0.93 (0.88) times as long as wide (n = 5), 1.50 – 1.60 (1.56) times as long as pronotum (n = 5); disc (Fig. 3 B) with two kinds of punctures, of which one is as large as an eye-facet at middle and another one is minute; the larger one possessing two lateral extensions and an assoicated recumbent seta; interspaces smooth. Prosternum short, about as wide as width of prosternal process, strongly carinate at middle; prosternal process about as long as procoxal length. Metaventrite (Fig. 2 B) convex, densely covered with long, recumbent yellowish setae, devoid of setae at basal areas. Inter-metacoxal distance wide, separated by 1.75 – 2.67 (2.21) times as wide as inter- mesocoxal distance (n = 4). Abdominal sternites (Fig. 2 B) densely covered with long, recumbent yellowish setae; inter-metacoxal area of abdominal sternite III wide, apical margin widely rounded; femoral lines (Fig. 2 B) joined with anterior margin of abdominal sternite III. Legs (Fig. 4 E – G) rather thin and flat; protibiae (Fig. 4 E) with slightly sinuate inner margins, outer apical corner rounded. Claws simple. Male genitalia with tegmen (Fig. 4 A) possessing widely spaced setae at apical margin, attenuate in ventral view. Median lobe (Fig. 4 C) in lateral view thin and strongly sinuate, in ventral view strongly converging at apical third, apical margin narrowly truncate. Internal sac sclerites of male genitalia as figured (Fig. 4 D). Female. Head and pronotum black, without yellowish lateral margins of pronotum. Apical margin of abdominal tergite and sternite VII rounded. Ovipositor (Fig. 4 K) with short styli near the apex; gonocoxites feebly sclerotized, deeply cleft and appearing separate and unfused; apex of stylus with 2 to 3 setae; 9 to 10 setae present along apico-lateral margin; spermatheca (Fig. 4 J) oval, sclerotized; spermathecal gland long, curved, weakly sclerotized; sperm duct coiled. Bionomics. Cybocephalus (C.) nipponicus usually feed on armored scale insects (Diaspididae). At least fourteen species of armored scales have been recorded as hosts (Smith & Cave 2007 a), and other insect prey were listed by Smith & Cave (2006 a). Tanaka & Inoue (1980) reported C. (C.) nipponicus as a predator of citrus red mite, Panonychus citri (McGregor) (Acari, Tetranychidae). According to breeding experiments, when C. (C.) nipponicus was given larvae of Unaspis yanonensis (Kuwana) (Hemiptera, Diaspididae), they laid three eggs per female. However, if they were fed eggs of Panonychus citri, their ovaries were observed to be undeveloped and they did not lay eggs. Thus, scale insects are likely the primary food of C. (C.) nipponicus, and citrus red mites are secondarily consumed.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF667C879D9F8FB0AFB34F8E2.taxon	distribution	Distribution. This species is present in Asia, southern Europe, Micronesia, eastern North America, South Africa, and the West Indies (Smith & Cave 2006 a). Cybocephalus (C.) nipponicus is apparently native to Southeast Asia and the South Pacific (Endrödy-Younga 1971; Tian 2006), but has been introduced all over the world (Smith & Bailey 2007). Note. The male pronotum, which is usually yellow, may rarely become dark brown to black.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF660C874D9F8FF2FFE2EFDF9.taxon	description	Cybocephalus politissimus: Endrödy-Younga, 1968: 77 [redescription, figure]; — Kirejtshuk, 1992: 209 [in key, figure]. Cybocephalus (Cybocephalus) politissimus: Jelínek & Audisio, 2007: 490 [catalogue]. Cybocephalus flavicapitus Tian & Yu, 1994 b: 209. Type locality: Shixing, Guangdong Province, China. Synonymized by Kirejtshuk & Fallahzadeh, 2008: 85. Cybocephalus flavocapitis Smith & Bailey, 2007: 504. Type locality: Kenting, Taiwan, syn. nov. Specimens examined. TAIWAN [TAINAN] 1 ex., Kuantzuling, 28. V. 1969, S. Hisamatsu leg. [NANTOU] 1 ex., Wushe, 24. V. 1972, M. Sakai leg. JAPAN [MIYAGI] 1 ex., Takiyama Park, Yamoto-chô, 2. VIII. 2009, T. Ozaki leg. [TOKYO] 3 exs., Izu-oshima Is., 19. V. 1974, Y. Hirano leg.; 1 ex., Niisima, 21. VI. 1992, Y. Hirano leg. [KANAGAWA] 2 exs., Odawara, 9. V. 1957, Y. Hirano leg.; 1 ex., Oishi, 8. IX. 1979, Y. Hirano leg.; 2 exs., Sarugashima, Atsugi City, 25. IV. 1996, Y. Hirano leg.; 1 ex., Tanzawa, 18. V. 1986, Y. Hirano leg.; 1 ex., Myoshintoge, 12. VIII. 1993, Y. Hirano leg.; 1 ex., Sengoku, Hakone, 4. VII. 1992, Y. Hirano leg.; 1 ex., Odawara, Kuno, 9. V. 1957, Y. Hirano leg.; 1 ex., Kamitsuruma 1 chôme, Sagamihara City, 20. IX. 1997, R. Iwata leg. [SHIMANE] 1 ex., Yoshiji-chô, Hamada City, 14. V. 1996, Y. Hirano leg. [EHIME] 12 exs., Kurotaki, nr. Matsuyama City, 18. V. 1958, M. Miyatake leg.; 1 ex., Mt. Sara, 16. V. 1968, M. Sakai leg.; 1 ex., Hantaji, Matsuyama City, 4. V. 1974, K. Ito leg.; 1 ex., Yama-jinjya, Matsuyama City, 23. V. 1973, K. Ito leg.; 1 ex., Tarumi, Matsuyama City, 22. VII. 1995, N. Ohbayashi leg. [KUMAMOTO] 1 ex., Mt. Ichibusa, 9. VI. 1967, S. Hisamatsu leg. [NAGASAKI] 1 ex., Mogi, 20. V. 1971, S. Hisamatsu leg.; 1 ex., Chiziwa, 10. XI. 1965, S. Nakao leg. [KAGOSHIMA] 2 exs., Amagi, Tokunoshima Is., 18. X. 1965, S. Nakao leg.; 2 exs., Mugio, Yaku-chô, 18. III. 1974, TK. Ishihara leg. [OKINAWA] 1 ex., Sueyoshi Park, Naha City, 8. III. 2003, Y. Kikuhara leg.; 2 exs., Syuri, 17. V. 1989, Y. Hirano leg.; 2 exs., Omoto, Ishigaki Is., 27. II. 1993, Y. Hirano leg.; 2 exs., Uegusuku, Kumejima Is., 21. IX. 2001, Y. Hirano leg.; 1 ex., Darumayama, Kumejima Is., 21. IX. 2001, Y. Hirano leg.; 1 ex., Oku, Kunigami-son, 4. IV. 2002, Y. Hirano leg.; 1 ex., Yamada, Onnason, 19. IV. 2009, Y. Hirano leg.; 4 exs., Miyako Is., 5 ~ 7. V. 1998, M. Satô leg.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF660C874D9F8FF2FFE2EFDF9.taxon	diagnosis	Diagnosis. The species is distinguished from other Japanese congeners by the following characteristics: disc of elytra smooth or shagreened; males with yellow head and black pronotum; punctures on elytra (Figs. 3 C – D) smaller than an eye-facet at middle; apical margin of median lobe (Fig. 5 K – O) in ventral view with strongly acute projection at middle. Redescription. Length 1.24 – 1.62 (1.43) mm (n = 7), width 0.70 – 0.82 (0.76) mm (n = 5), height 0.40 – 0.56 (0.49) mm (n = 7).	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF660C874D9F8FF2FFE2EFDF9.taxon	description	Male. Body convex; dorsal disc almost glabrous, shiny. Coloration black; dorsal- and ventral side of head (excluding blackish mandibles and apical half of genae), antennae, lateral margins of pronotum, mesoventrite, pro- and mesothoracic legs (excluding blackish mesotibiae) yellow; lateral margins and posterior corners of pronotum, anterior margins of elytra translucent, yellowish brown; hind legs dark yellowish brown. Head extremely wide; vertex minutely punctured; interspaces smooth or shagreened; labrum transverse, with apical margin shallowly emarginate at middle. Eyes large, visible from ventral aspect. Antennae (Fig. 6 F) similar to those of C. (C.) nipponicus; club 3 - segmented, apical margin of eleventh antennal segment truncate; approximate ratio of each segment (n = 1) is 2.13: 1.65: 1.87: 1.13: 1.25: 1.06: 1.06: 1.00: 1.37: 1.52: 1.40. Pronotum convex, strongly transverse, 1.67 – 1.73 (1.71) times as wide as long (n = 5); anterior angles rounded; posterior angles widely rounded; dorsal disc minutely punctated; interspaces smooth or shagreened. Scutellum with apex broadly rounded. Elytra conjointly 0.86 – 0.97 (0.92) times as long as wide (n = 5), 1.50 – 1.73 (1.60) times as long as pronotum (n = 5); disc (Figs. 3 C – D) with two kinds of punctures, one large, but smaller than an eye-facet, the larger one possessing two lateral extensions and an associated recumbent seta; the other puncture minute; interspaces smooth or shagreened. Metaventrite convex, densely covered with long, recumbent yellowish setae, but without setae at basal areas. Inter-metacoxal distance wide, separated by 1.50 (1.50) times as wide as inter- mesocoxal distance (n = 2). Abdominal sternites densely covered with long, recumbent yellowish setae; inter-metacoxal area of abdominal sternite III with widely rounded apical margin; femoral lines joined with anterior margin of abdominal sternite III. Legs (Fig. 6 C – E) rather slim and flat, similar to those of C. (C.) nipponicus; protibiae (Fig. 6 C) with slightly sinuate inner margins, outer apical corner rounded. Claws simple. Male genitalia (Fig. 5) with some local variations in shape (see ‘ Note’ below); tegmen strongly or uniformly attenuate, scattered with sparse setae at apical margin; median lobe in lateral view strongly thickened, more or less square in shape with strongly acute projection at middle of apical margin in ventral aspect. Internal sac sclerites of male genitalia as figured (Fig. 5 A – E). Female. Coloration of head black. Apical margin of abdominal tergite and sternite VII rounded. Female genitalia (Fig. 6 G) similar to those of C. (C.) nipponicus; coxites feebly sclerotized, appearing separate and unfused, with short styli near apex; apex of stylus with 2 ~ 3 setae; 9 ~ 10 setae present along apico-lateral margin; spermatheca (Fig. 6 A) oval, sclerotized; spermathecal gland long, curved, weakly sclerotized; sperm duct coiled. Bionomics. According to Smith & Bailey (2007), this species feeds upon Aulacaspis yasumatsui Takagi. Distribution. This species occurs in Europe (France, Germany, Italy), Russia (East and West Siberia, Far East), China (Guangdong) (Jelínek & Audisio 2007), Taiwan (new record), and Japan (new record). Taxonomy. Cybocephalus (C.) flavocapitis Smith, 2007 is treated herein as a junior synonym of C. (C.) politissimus. Japanese and Taiwanese specimens of C. (C.) politissimus were compared with type specimens of C. (C.) flavocapitis by Dr. T. R. Smith, and he agrees with this emendation (Smith pers. comm.). Cybocephalus (C.) flavocapitis and C. (C.) politissimus share the following characteristics: head yellowish; apical margin of eleventhantennal segment truncated; tegmen in ventral view more or less attenuate; median lobe in lateral view strongly truncate, in ventral view apical margin with strongly acute projection at middle; internal sac sclerites of same shape. Based on these characteristics, C. (C.) flavocapitis should be treated as a junior synonym of C. (C.) politissimus. Note. This species has some variation in the shape of the male genitalia (Fig. 5) and punctation of the dorsal surface (Fig. 3 C – D). Nevertheless, the basic structure of the male genitalia is identical, especially the thickened median lobe (in lateral aspect) and shape of the internal sac sclerites. Therefore, these variable anatomical features should be considered as intraspecific variation. Coloration of the male pronotum is usually black, but rarely becoming yellowish brown.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF66FC877D9F8FD3DFB3FFE69.taxon	materials_examined	Type series. Holotype: 3, NAZE / Amami Is. / 5. VI. 1960 / Taichi Shibata // HOLOTYPE / Cybocephalus (Cybocephalus) / chlorocapitis S-T. Hisamatsu, 2011 /. Paratype: [OKINAWA] 1 Ƥ, Miyako Is., 5 ~ 7. V. 1998, M. Satô leg. Type depository. The holotype and paratype are deposited in the EUM.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF66FC877D9F8FD3DFB3FFE69.taxon	etymology	Etymology. The specific epithet is derived from the Latin wards “ chloros ” meaning green + “ caput ” meaning head.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF66FC877D9F8FD3DFB3FFE69.taxon	diagnosis	Diagnosis. Surface of head and pronotum (Fig. 1 C) with metallic-green sheen in both sexes, female sheen much reduced. Protibiae (Fig. 7 D) wide (L / W = 3.90 (n = 1 )), outer angle broadly rounded. Apical area of female gonocoxites (Fig. 7 G) sclerotized.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF66FC877D9F8FD3DFB3FFE69.taxon	description	Description. Length 1.70 mm (n = 1), width 0.93 mm (n = 1), height 0.50 mm (n = 1). Male. Body convex; dorsal disc almost glabrous, strongly shining. Coloration black; base of mandibles, labrum, anterior half of head, marginal area of pronotum, and lateral and apical margins of elytra with metallicgreen sheen; labial and maxillary palpi, antennae (except brownish first and eleventh segments), lateral margins of pronotum, and tarsi yellow. Head extremely wide; vertex rather sparsely punctate; interspaces smooth. Mentum trapezoidal, anterior margin deeply emarginate. Eyes large, visible from ventral aspect. Antennae (Fig. 7 I) with club asymmetrical, 3 - segmented; apical margin of eleventh segment widely rounded; approximate ratio of each segment (n = 1) is 2.90: 1.88: 2.02: 1.33: 1.44: 1.10: 1.17: 1.00: 1.40: 1.25: 1.71. Pronotum convex, strongly transverse, 1.70 times as wide as long (n = 1); anterior angles rounded; posterior angles widely rounded; dorsal disc minutely punctuate; interspaces smooth. Scutellum with apex broadly rounded. Elytra conjointly 0.89 times as long as wide (n = 1), 1.65 times as long as pronotum (n = 1); disc with two kinds of punctures; the larger one about as large as an eye-facet, possessing two lateral extensions and an associated recumbent seta; interspaces smooth. L / W = 3.90 (n = 1). Metaventrite convex at anterior half, posterior half depressed for reception of metafemora; disc covered with long, recumbent yellowish setae. Inter-metacoxal distance wide, separated by 1.33 (1.33) times as wide as inter- mesocoxal distance (n = 1). Abdominal sternites densely covered with long, recumbent yellowish setae; inter-metacoxal area of abdominal sternite III with widely rounded apical margin; femoral lines joined with anterior margin of abdominal sternite III. Legs (Fig. 7 D – F) with protibiae wide (L / W = 3.90 (n = 1 )); protibiae with broadly rounded outer apical angle. Claws simple. Male genitalia with tegmen (Fig. 7 A) gently narrowing to apex, setose around margins; lateral margins somewhat constricted at middle; apical margin widely rounded. Median lobe (Fig. 7 C) in lateral view gently curved, strongly narrowing in apical third in ventral view; apical margin with protruded apex. Female. Metallic sheen less developed than in male: base of mandibles, labrum, clypeus, anterior and posterior margins of pronotum with feebly metallic-green sheen. Apical margin of abdominal tergite and sternite VII rounded. Ovipositor (Fig. 7 G) elongate and slender; coxite strongly sclerotized, deeply cleft and appearing separated, long styli present near apex; apex of stylus with 3 ~ 4 setae; 5 ~ 6 setae present along apico-lateral margin; spermatheca (Fig. 7 H) oval, sclerotized; spermathecal gland straight, weakly sclerotized. Bionomics. Biology unknown.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF66FC877D9F8FD3DFB3FFE69.taxon	distribution	Distribution. This species is known only from the type locality on Amami and Miyako Islands, Japan. Remark. This new species is closely allied to C. (C.) liui Tian, known from Yunnan, China, in having metallic-green sheen on surface of head and pronotum and similar shape of median lobe, but can be distinguished by short, widened protibiae and the shape of tegmen. The present new species is also similar to C. (C.) chinensis Yu, known from Guangdong, China, C. (C.) festivus Erichson, known from Europe, and C. (C.) minimus Grouvelle, known from Madagascar and Seychelles, in having metallic-green sheen on surface of head and pronotum, but differs from them by the broadly rounded outer angle of protibiae and the shape of male genitalia.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF66DC870D9F8FEFDFE4EFDD2.taxon	materials_examined	Type series. Holotype: 3, (RYUKYU: JAPAN) / Shirahama / Iriomote / 10. X. 1988 / M. Sakai leg. / HOLOTYPE / Pastillus eminentithorax / S-T. Hisamatsu, 2011 /. Paratypes: 20 exs.: [MIYAGI] 1 ex., Takiyama Park, Yamotochô, 2. VIII. 2009, T. Ozaki leg. [SAITAMA] 1 Ƥ, Urawa City, 18. VI. 1957, H. Kajimura leg. [KANAGAWA] 1 Ƥ, Mt. Kôbô, 22. IV. 1989, Y. Hirano leg. [TOKYO] 1 Ƥ, Sanbancho, 4. X. 2008, K. Masumoto leg.; 1 Ƥ, Shimizudani, Megasaki City, 16. III. 2001, Y. Hirano leg. [AICHI] 1 Ƥ, Araike, Tempaku Ward, Nagoya City, 6. V. 1995, H. Yoshitomi leg.; 1 Ƥ, Hadenoshi-Hassawa, 27. IV. 1996, Y. Hirano leg. [HIROSHIMA] 1 Ƥ, Mt. Haigamine, Kure City, 8. IX. 2002, I. Okamoto leg. [SHIMANE] 1 Ƥ, Kushiro-chô, Masuda City, 15. V. 1996, Y. Hirano leg. [EHIME] 1 Ƥ, Ohmishima Is., 16. VII. 1959, M. Miyatake leg.; 6 Ƥ, Shiroyama, Matsuyama City, 5. V. 1973, M. Sakai leg. [KOCHI] 1 Ƥ, Cape Ashizuri, 13. VI. 1961, S. Hisamatsu leg. [NAGASAKI] 1 Ƥ, Unzen Spa, 5. VI. 1967, S. Hisamatsu leg.; 1 Ƥ, Mt. Fugen ~ Mt. Myoken, 7. X. 1970, S. Hisamatsu leg. [KAGOSHIMA] 1 Ƥ, Sendai City, 28 ~ 24. III. 2001, Y. Hirano leg. Type depository. The holotype and 15 paratypes are deposited in the EUM. Five paratypes are deposited in PCYH.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF66DC870D9F8FEFDFE4EFDD2.taxon	etymology	Etymology. The specific name means “ projecting pronotum (eminens + thorax) ”, which is derived from the strongly projecting lateral margins of the pronotum in lateral aspect.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF66DC870D9F8FEFDFE4EFDD2.taxon	diagnosis	Diagnosis. Body strongly convex, almost hemispherical. Antennae (Fig. 8 K) 10 - segmented, with 2 - segmented club; scape strongly enlarged, widest at middle. Lateral margins of pronotum in lateral view strongly projecting. Inter-metacoxal lobe of abdominal sternite III (Fig. 2 A) strongly narrowed.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF66DC870D9F8FEFDFE4EFDD2.taxon	description	Description. Length 1.66 – 1.83 (1.78) mm (n = 5), width 0.70 – 0.80 (0.61) mm (n = 5), height 0.63 – 0.70 (0.67) mm (n = 5). Male. Body highly convex and shining, contractile; dorsal disc almost glabrous. Coloration black to dark brown; antennal segments 2 – 8, maxillary and labial palpi, prosternum, mesoventrite, procoxae, protrochanters, profemora, mesocoxae and mesotrochanters yellowish. Head wide; vertex distinctly punctuate; interspaces smooth; apical margin distinctly bordered, with deep emaginations for reception of antennal scape. Labrum (Fig. 8 G) about as wide as long, with truncate apical margin. Eyes large, but scarcely visible in ventral aspect. Mentum (Fig. 8 J) trapezoidal. Antennal grooves long and deep. Antennae (Fig. 8 K) 10 - segmented, with 2 - segmented club; scape strongly swollen, widest at middle; approximate ratio of each segment (n = 1) is 2.55: 1.96: 1.25: 1.11: 1.24: 1.02: 1.05: 1.00: 1.51: 4.33. Pronotum strongly convex, 1.70 – 1.78 (1.74) times as wide as long (n = 5); lateral margins strongly projecting in lateral aspect; dorsal disc with minute punctures and short recumbent sparse setae; interspaces smooth. Scutellum (Fig. 2 C) triangular, with acuminate apex. Elytra strongly convex in lateral aspect, conjointly 1.17 – 1.24 (1.19) times as long as wide (n = 5), 1.53 – 1.73 (1.61) times as long as pronotum (n = 5); disc (Fig. 3 A) with minute, uniform punctures; interspaces smooth. Prosternum strongly carinate at middle. Metaventrite (Fig. 2 A) depressed, short, about as wide as metafemora; disc asetose. Inter-metacoxal distance strongly narrow, separated by 1.28 (1.28) times as wide as inter-mesocoxal distance (n = 1). Inter-mesocoxal distance strongly narrowed. Abdominal sternites (Fig. 2 A) covered with long, recumbent yellowish setae; inter-metacoxal area of abdominal sternite III strongly narrowed, triangular; femoral lines (Fig. 2 A) strongly expressed beyond anterior margin of abdominal sternite III. Legs (Fig. 8 D – F) with protibiae slender, inner and outer margin slightly sinuate; metatibiae flat and wide, strongly widened from basal third to apex. Claws simple. Male genitalia with tegmen (Fig. 8 A) in ventral view constricted at middle; anterior margin of tegmen with two projections. Median lobe (Fig. 8 B) in ventral view subparallel-sided from base to middle, abruptly attenuate anteriorly; anterior margin more or less truncate in ventral aspect; apico-lateral corners with tuft of long setae in ventral aspect. Internal sac sclerites of male genitalia (Fig. 8 C) as figured. Female. Apical margin of abdominal tergite and sternite VII rounded. Ovipositor (Fig. 8 I) elongate and slender; coxites feebly sclerotized, deeply cleft and appearing separated, apex of stylus with 3 ~ 4 setae; 5 ~ 6 setae present along apico-lateral margin. Spermatheca (Fig. 8 H) C-shaped, spermathecal gland distinct, sperm duct coiled. Bionomics. Biology unknown.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF66DC870D9F8FEFDFE4EFDD2.taxon	distribution	Distribution. This species is known from Honshû, Shikoku, Kyûshû, and Iriomote Island, Japan.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
421787CFF66DC870D9F8FEFDFE4EFDD2.taxon	discussion	Remark. This new species is similar to Pastillus conflexus Endrödy-Younga, known from Sierra Leone, but can be distinguished from the latter by the ten-segmented antennae and peculiar shape of male genitalia. Pastillus eminentithorax sp. nov. is widely distributed in Japan. Twenty-one specimens were examined, with only one male found, which was collected on Iriomote Is., Okinawa Pref. This preliminary data on sex ratios may indicate a parthenogenetic lifestyle of the species. However, further examination and field data are needed to corroborate this hypothesis. Pastillus is a small genus, consisting of three species, with previously known distributions limited to localities in the Afrotropical Region. This new species represents the fourth species of the genus, and is the first recorded from Palaearctic Region.	en	Hisamatsu, Sadatomo (2013): A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616 (3): 253-267, DOI: 10.11646/zootaxa.3616.3.3
