taxonID	type	description	language	source
BED1863A60AD56B79439BE3B05604323.taxon	description	Figs 1, 2, 3	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
BED1863A60AD56B79439BE3B05604323.taxon	diagnosis	Diagnosis. Body with 21 LBS, and LBS 7 th with spiracles (Fig. 2 E). Antennae (Fig. 2 A, B) composed of 20 – 21 articles, of which three basal ones glabrous dorsally. Cephalic plate smooth, lateral and posterior marginations absent; the posterior edge straight, and covered by tergite 1 (Fig. 2 A). Forcipular trochanteroprefermur with median tooth; tooth-plates with 4 + 4 teeth clearly, sharply (Fig. 2 B). Tergite 1 wider than the cephalic plate and twice as long as tergite 2 (Fig. 2 A); tergites 3 – 19 (20) with complete paramedian sutures, and tergites 12 (13) – 20 with lateral marginations incomplete or nearly complete (Figs 2 E, 3 A, B). Sternites 3 – 20 with complete paramedian sutures (Fig. 2 C, D). Coxopleuron with 1 apical, 2 subapical, and 1 lateral spine, without dorsal spine (Fig. 3 D). Leg 1 with prefemoral and femoral spurs (Fig. 2 B). Ultimate legs (Fig. 3 C, D) long and slender, with ratio between the length and width of prefermur 4.5: 1; prefemoral spines on ultimate leg: 4 VL, 2 – 3 VM, 3 – 4 M, and 1 – 2 DM spines.	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
BED1863A60AD56B79439BE3B05604323.taxon	distribution	Distribution. This species has only been recorded in Southeast Asia, including Thailand, Laos, Cambodia, and Vietnam (Tran et al. 2013; Siriwut et al. 2018). In Vietnam it was recorded from Ha Nam Province (Phu Ly) and Ca Mau Province (Attems 1930; Tran et al. 2013), and Ba Ria – Vung Tau Province and Dong Thap Province in this study. The distribution of A. calcaratus was further updated (Fig. 1).	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
1406D32EE8DC5912AC88C72B65C99D02.taxon	type_taxon	Type species. Alluropus demangei Silvestri, 1911, by monotypy.	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
1406D32EE8DC5912AC88C72B65C99D02.taxon	diagnosis	Diagnosis. According to Schileyko et al. (2020: p. 50), the genus can be recognized by the following characters: median tooth of labrum well developed; forcipular tooth-plates present; trochantero-prefemur with well-developed process; tergites never with longitudinal keels; sternites with incomplete paramedian sutures (in some species much shortened, sometimes also with median and lateral depressions of various sizes / shapes; LBS 7 with spiracles; legs with tarsal spur (s), legs 1 with prefemoral spur; coxopleural process ranging from short to very long and much enlarged with spines (at least with apical ones); prefemur of the ultimate leg with spines; ultimate pretarsus well-developed, with accessory spines.	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
1406D32EE8DC5912AC88C72B65C99D02.taxon	distribution	Distribution. Vietnam, Cambodia, Laos, and Thailand.	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
C89ADC33546457969E237C952A505C35.taxon	description	Figs 1, 6, 7	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
C89ADC33546457969E237C952A505C35.taxon	diagnosis	Diagnosis. Body with 21 leg-bearing segments. LBS 7 th with spiracles. Antennae (Fig. 6 A) composed of 18 – 20 articles, of which three basal ones glabrous dorsally. Cephalic plate without lateral and posterior marginations; posterior edge straight, and covered by tergite 1 (Fig. 6 A). Tooth-plates with 4 (5) + 4 (5) teeth prominently; trochanteroprefemora of forcipular strong, with 3 + 4 prominent teeth (one apical and two-three lateral teeth )); coxosternite with short median suture starting from toothplate (Fig. 6 C). Tergites (Figs 6 B, 7 C) without punctate and short median sulcus; lateral margination absent (except for tergite 21, complete); tergites 6 – 20 with complete paramedian sutures and incomplete on tergites 3 – 5. Sternites 2 – 19 with incomplete (only 10 – 15 % length of sternites) paramedian sutures on anterior and posterior parts (Fig. 6 D). Coxopleural process with 1 (2) apical and 1 (2) subapical spine, lateral and dorsal spines absent (Fig. 7 B). Leg 1 with 1 fermoral, 1 tibial, and 2 tarsal spur (s) (Fig. 7 A). Ultimate leg (Fig. 7 D) prefemora with 1 – 2 VL, 0 – 1 VM, 0 – 1 M, and 0 – 2 DM spines.	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
C89ADC33546457969E237C952A505C35.taxon	distribution	Distribution. This species occurs in Africa (Congo, Sudan), South America (Bolivia, Cuba, El Salvador, Guatemala, Peru, Venezuela, and Uruguay), and South and Southeast Asia (Taiwan, Cambodia, Indonesia, Kalimantan, Maluku, Myanmar, the Philippines, Singapore, Sri Lanka, Sumatera, Papua New Guinea, Malaysia, India, Nepal, Thailand, and Laos) (Bonato et al. 2016; Schileyko and Stoev 2016; Siriwut et al. 2018). In Vietnam it was recorded in Quang Ninh Province (Ha Long Archipelago) and Nghe An Province (Vinh city) by Attems (1938), and Ha Giang, Dien Bien, Son La, Hoa Binh, Phu Tho, Vinh Phuc, Hai Phong, Thai Binh, Ha Noi, and Dong Thap in this study. The distribution of R. immarginata was further updated (Fig. 1).	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
4CFCAE0B8E105EDF82BC2CF559D24BA1.taxon	description	Figs 1, 4, 5	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
4CFCAE0B8E105EDF82BC2CF559D24BA1.taxon	diagnosis	Diagnosis. Body with 21 LBS. LBS 7 th with spiracles. Antennae (Fig. 4 A, B) composed of 18 articles, of which three basal ones glabrous dorsally. Cephalic plate (Fig. 4 A) without lateral and posterior marginations; posterior edge straight and covered by tergite 1. Forcipular segment (Fig. 4 B) smooth; tooth plates with 4 + 4 teeth obviously; coxosternite with short median suture starting from toothplate. Tergites without short median sulcus; paramedian sutures complete on tergites 4 (5) – 19 (Figs 4 C, 5 A); lateral marginations incomplete on tergites 6 (10) – 20 and complete on tergite 21 (Figs 4 C, 5 A). Sternites 3 – 20 with incomplete (10 – 15 % length of sternites) paramedian sutures (Figs 4 B – D, 5 B). Coxopleural process long with 2 apical, 1 subapical, and 1 lateral spine, without dorsal spine (Fig. 5 B). Leg 1 present or absent femoral spur (Fig. 4 A, B). Prefemoral spines on ultimate legs with 2 – 3 VL, 2 – 5 VM, 0 – 1 M, and 1 – 3 DM spines; prefemoral process of ultimate legs very short with 1 spine (Fig. 5 C).	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
4CFCAE0B8E105EDF82BC2CF559D24BA1.taxon	distribution	Distribution. This species is widespread in tropical mainland and island regions. It occurs in Africa (Madagascar, Seychelles, and Mauritius), Central and South America (Mexico, Peru, Cuba, and Haiti), and Central, East, and Southeast Asia (China, Taiwan, Nepal, India, Sri Lanka, Chagos Islands, Yemen, Pakistan, Iran, Myanmar, Thailand, Cambodia, Malay Peninsula, Philippines, and Papua New Guinea) (Siriwut et al. 2018; Joshi et al. 2019). In Vietnam, it has been recorded from Thua Thien Hue (Hai Van Pass); Dong Nai (Ma Da); Khanh Hoa (Spratly Islands); and Kien Giang (Phu Quoc Island) (Attems, 1938; Schileyko, 2007; Son et al. 2022); and in Dien Bien, Thai Binh, Hanoi, Ha Nam, and Quang Binh (in this study). The distribution of R. longipes was further updated (Fig. 1).	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
855610FE43C859B592E133D707581960.taxon	description	Figs 1, 8, 9	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
855610FE43C859B592E133D707581960.taxon	diagnosis	Diagnosis. Body with 21 leg-bearing segments. LBS 7 th with spiracles. Antennae composed of 18 articles, of which three basal ones glabrous dorsally. Tooth-plate with five short teeth. Tergites with lateral marginations and 5 – 7 keels and many spinulations scattered on tergites, more obvious in the posterior tergites, without complete paramedian sutures. Sternites with paramedian suture incomplete on anterior part (on some sternites). Coxopleural process with 2 apical, 1 lateral spines and without dorsal spine.	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
855610FE43C859B592E133D707581960.taxon	description	Description. Body length 58 mm. Antennae composed of 21 articles, of which three basal ones glabrous dorsally, following segments densely covered by minute setae (Fig. 8 A, B). Antenna length reaching to 6 th body segment. Cephalic smooth, without puncta and setae; lateral and posterior margination absent, the posterior border adjacent to anterior border of tergite 1. Four big ocelli present, nearly equal in size. The median suture on the top of cephalic plate long, shallow (Fig. 8 A). Forcipular segment (Fig. 8 B), coxosternite, trochanteroprefemora, and the basal part of tarsungulae smooth, without punctate and setae; coxosternite without median suture. Tooth-plates slightly wider than high and nearly semicircular, with 5 short teeth and well chitinized; the distance between the tooth – plates wide; each tooth-plate with straight, transverse basal suture. Forcipular trochanteroprefemoral process bearing 3 short denticles, the latter being indistinct, position of denticles higher than tooth – plates. Maxillae with sparse short setae, and segment 2 of second maxillary telopodite with spur; chitin-line very short. Tergites with very small punctae; lateral margination incomplete on tergites 9 – 20 and complete on tergite 21; paramedian sutures absent on all tergites; tergites 1 – 7 smooth, and tergites 8 – 20 with 5 – 7 keels and many spinulations which scattered on tergites, more obvious in the posterior tergites; tergite of ultimate leg-bearing segment sub-quadrangular and convex posteriorly (Figs 8 D, 9 A). Sternite surface smooth (without punctae), lacking any median sulci. Sternites shield-shaped, last sternite trapezoid (or nearly triangular). Incomplete paramedian suture on anterior part of some sternites (<10 % of their width) or absent, lacking depression (Figs 8 B, C, 9 B). Coxopleuron densely covered by pores (except the apical part); coxopleural process with 2 apical and 1 lateral spine, without dorsal spine (Fig. 9 B). Locomotory legs usually bearing small, short setae; legs 1 – 4 with 1 tibia spur, legs 1 – 18 with 2 tarsal spurs, 19 – 20 with 1 tarsal spur. Ultimate legs very thin and long; the ratio of lengths of prefemur, femur, tibia, tarsus 1, and tarsus 2 as follows: 2.8: 2.7: 2.5: 2.0: 1.0; ratio of length and width of prefemur 6: 1. Prefemoral spines on ultimate leg: 4 VL, 3 VM, 3 M, and 1 DM spines; prefemoral process of ultimate legs very short with 1 spine; all of segment smooth (Fig. 9 C).	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
855610FE43C859B592E133D707581960.taxon	etymology	Etymology. The species name is named after the Joint Vietnam-Russia Tropical Science and Technology Research Center (VRTC), an institution that has significantly contributed to biodiversity research in Vietnam.	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
855610FE43C859B592E133D707581960.taxon	distribution	Distribution. Only known in Lao Cai province and Son La province in Vietnam (Fig. 2).	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
A9F5EF03B7805D64815DE99341A1804D.taxon	type_taxon	Type species. Rhysida lithobioides (Newport, 1845).	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
A9F5EF03B7805D64815DE99341A1804D.taxon	diagnosis	Diagnosis. Body with 21 LBS. LBS 7 th with spiracles. Antennae composed of 16 – 21 articles, of which 2.8 – 3 basal ones glabrous dorsally. Cephalic plate without lateral and posterior marginations. Forcipular tooth-plates present; tooth-plates with 4 – 7 teeth; trochantero-prefemur with well-developed process; coxosternite with short median suture starting from tooth-plate. Sternites with incomplete paramedian sutures. Coxopleural process long with 1 – 2 apical, 0 – 2 subapical, and 0 – 1 lateral spines, without dorsal spines. Leg 1 with 1 femoral, 1 tibial, and 2 tarsal spurs.	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
A9F5EF03B7805D64815DE99341A1804D.taxon	distribution	Distribution. Neotropics, Africa and Indomalaya, and Australia.	en	Le, Son Xuan, Do, Thinh Tat, Tran, Binh T. T., Nguyen, Anh D. (2025): The centipede genera Alluropus Silvestri, 1911 and Rhysida Wood, 1862 (Scolopendromorpha, Scolopendridae) in Vietnam, with a description of a new species. Zoosystematics and Evolution 101 (4): 1873-1885, DOI: 10.3897/zse.101.156915
