taxonID	type	description	language	source
450587FEFFEEF44BFEC14A8AFD68F870.taxon	type_taxon	Type species: Dolichoiulus polyzonus Attems, 1911, by subsequent designation by Jeekel (1970). Tentative diagnosis: A genus of the tribe Pachyiulini characterized by a (more or less distinctly) spoonshaped promere with two apical denticles, and with an opisthomere deeply divided into a mesomeral and a solenomeral branch, the latter lacking fovea or pseudofovea and pseudoflagellum. Different from the similar genera Afropachyiulus, Amblyiulus, Dolichoiulus, and Japanoiulus mostly by a typically (but not always) weakly pronounced anterior lamella of the solenomere, apically never drawn into a distinct process, and by the solenomere apex being blunt or concave, sometimes bearing minute apical processes or / and ciliae, but not tapering into a single, pointed tip.	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEEF44FFC144E0AFBC7FBEF.taxon	description	Figs 1 - 16	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEEF44FFC144E0AFBC7FBEF.taxon	materials_examined	Material: MNHG; male holotype (fragmented into head, rings 1 - 6, left part of ring 7 and rest of body; gonopods, left antenna, leg-pair 1, leg-pair 2, penis, right leg 3, right edge of ring 7, and left mid-body leg dissected; left opisthomere prepared for SEM), sample field number 7314; label “ Iran (Gilan), Siahkal, tamis., mousse, creux gros arbres ” [Iran, Gilan Province, Siahkal, sifting, moss, old, hollow trees]; 06. JUL. 1973; leg. A. Senglet. – MNHG; 1 adult female paratype (fragmented into head to ring 3 and rest of body; right vulva dissected); 1 juvenile female paratype (unbroken); same collecting data as holotype.	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEEF44FFC144E0AFBC7FBEF.taxon	type_taxon	Type locality: Iran, Gilan Province, Siahkal.	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEEF44FFC144E0AFBC7FBEF.taxon	etymology	Etymology: Named in memory of Hans Lohmander (1896 - 1961), a prominent millipede taxonomist who described six species currently assigned to Syrioiulus.	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEEF44FFC144E0AFBC7FBEF.taxon	diagnosis	Diagnosis: A blind species of Syrioiulus, particularly similar to S. astrabadensis (Lohmander, 1932 a) and S. persicus (Golovatch, 1983) in gonopod and vulva structure. Different from S. astrabadensis mostly by its larger size and darker colouration, and by mesomeral process being subequal to rather than significantly shorter than solenomere. Different from S. persicus by absence of metazonal setae, and by presence of typical, pointed mesal apical denticle of promere (the same being a rounded lobe in the latter species).	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEEF44FFC144E0AFBC7FBEF.taxon	description	Description Measurements: Holotype with 60 + 1 + T body rings, length 29 mm, mid-body vertical diameter 1.65 mm. Adult female paratype with 59 + 1 + T body rings, length 28 mm, mid-body vertical diameter 1.9 mm. Juvenile female paratype with 49 + 4 + T body rings. Colouration (after more than 40 years of ethanol preservation) (Figs 1 - 4): Mostly ochre brown, prozonae greyish with purple tinges; head with a dark, blurred, oval spot mesally to each antenna, and with a dark, narrow, axial line on vertex; body without axial line; legs and antennae light yellow to completely pallid; pre-anal ring and paraprocts darker ventrally. General morphology (Figs 1 - 4, 5): Ommatidia absent; vertigial setae absent; 5 supralabral setae in the adult specimens, and 4 in the juvenile; 20 - 26 labral setae. Antennae (Fig. 5) about 1.2 times as long as head in both sexes; antennomeres 3, 4, and 5 subequal in length, about 0.8 times as long as 2, and 1.5 times as long as 6, 5 considerably thicker than 6; a whorl of sensilla basiconica at distal margins of antennomeres 5 and 6, those on 5 nearly as long as the four apical cones, somewhat longer than those on 6. Gnathochilarium of normal julid appearance; promentum relatively short: considerably less than half the length of lingual lamellae; each lingual lamella bearing 4 - 5 setae. Collum smooth, some specimens with one to several very shallow grooves near posterolateral corners. Prozonae with very short and shallow, scattered, longitudinal striae. Metazonae rather shallowly and somewhat irregularly striated, with 5 - 8 striae in a square with sides equal to metazonal length just below ozopore level; hind margins slightly turned outwards, bearing no setae. Ozopores set in metazonae, at 1 / 4 - 1 / 3 of metazonal length behind pro-metazonal suture; the latter indistinct, especially on dorsal side, recognizable only as a barely visible concentric groove running through middle of ring. Mid-body legs 1.55 and 0.95 times as long as midbody diameter in the adult male and in the adult female, respectively. Tarsus of mid-body legs 1.2 - 1.3 times as long as tibia, and about 3 times as long as apical claw. Telson (Fig. 4): Pre-anal ring dorsally with a few short setae; epiproct absent; hypoproct short and broadly rounded, tightly fitting under paraprocts in both sexes, with a pair of disto-median setae and a transverse row of around 10 setae proximally to the former. Paraprocts densely covered with long setae, without rows of short and stiff setae along caudal margins. Male sexual characters: Mandibular stipites (see Fig. 1) forming rather small, rounded, frontoventral lobes. Legpair 1 (Fig. 6) developed as compact hooks pointing mostly mesad, with a tapering tibial outgrowth and a minute, knob-like, tarsal remnant. Leg-pair 2 (Fig. 7) with two crested adhesive pads, one each on postfemur and tibia. Leg-pair 3 (Fig. 8) and following pairs additionally with a vestigial pad on femur; all pads gradually diminishing in size towards telson, completely disappearing in posterior third of body. Pleurotergum 7 (Fig. 9) ventrally forming large, rounded lobes originating from the border between pro- and metazona, protruding mostly ventrad. Penis soft, hyaline, with deeply divided, conoidal lobes ending up in fingertip-like terminal lamellae turned slightly caudad. Gonopods (Figs 11 - 16): Promere (Fig. 11, P in Fig. 12) relatively slender and spoon-like, tapering, as high as opisthomere; width of shaft about 0.7 times that of bowl; ridge (r) massive and rounded, without freely protruding apical part; apical denticles well developed, clearly separated, mesal one running vertical, lateral one somewhat oblique, both with blunt tips pointing caudobasad. Opisthomere (Figs 12 - 16) approximately halfway divided into mesomeral process (m) and solenomere (s), these being subequal in height, forming a broad sinus between them; mesomeral process running mostly straight, at considerable distance from solenomere, apically broadened and spade-like, with a strongly elongated mesal corner turned caudad, partly concealing solenomere, and with a nearly right-angled lateral corner bearing several minute spines / denticles; caudomesal lamella (cl) well-developed, forming a narrow ridge protruding completely mesad, thus being clearly visible in frontal and caudal views; solenomere slender, gradually bent anteriad, ending in two small processes [an anterior, fan-like (fp) and a posterior, shield-like one (sp)] surrounding a deep central concavity; a minute central filiform process (ff) visible in certain aspects; anterior lamella vestigial, recognizable only by several apical fringes. Female sexual characters: Leg-pair 1 considerably, legpair 2 slightly flattened antero-caudally. Vulva (Fig. 10) strongly compressed in the sagittal plane; bursa mostly symmetric: lateral valve (lv) slightly higher and more narrow than mesal one (mv); median cleft relatively broad; operculum (op) by far exceeding bursa, apical margin with a distinct mid-lateral incision and with several bumps along mesal section; setation moderately dense, randomly spread across whole surface of vulva. Receptaculum seminis consisting of a narrow, nearly straight central tube (ct) supporting a very short mesal tube (mt) ending in a small, ovoid, mesal ampulla (ma).	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEEF44FFC144E0AFBC7FBEF.taxon	discussion	Remarks: The type locality of the new species lies in the foothills of the Alborz Mountains, the mountain range where most Syrioiulus species from Iran were exclusively recorded (Fig. 18). The only exception is S. zarudnyi (Lohmander, 1932 a) which is known also from two localities in the central part of the country. Considering the absence of ommatidia and the method (sifting dead organic matter) by which the studied specimens were collected, S. lohmanderi sp. nov. is likely an euedaphic species.	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEAF44EFCA94B82FEF4FD76.taxon	description	Fig. 17	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEAF44EFCA94B82FEF4FD76.taxon	materials_examined	Material: GNM; 2 (supposed) female syntypes of “ Amblyiulus astrabadensis n. sp. ”; Persien: Astrabad (nr. 28); 02. May. 1905, E. Filippowitsch. – GNM; 2 slide preparations of (supposed) female syntypes: Amblyiul. nr. 28, I: vulvae, legs, and mouthparts, and Amblyiul. nr. 28, II: mouthparts and vulvae. – GNM; 2 slide preparations of (supposed) male syntypes: Amblyiul. nr. 28: gonopods, mouthparts, legs, antennae, edges of pleurotergum 7, and Amblyiul. nr. 28, II: labrum, mandible, legs. Type locality: Iran, Golestan province, Gorgan (formerly Esterabad).	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEAF44EFCA94B82FEF4FD76.taxon	discussion	Taxonomic notes: Promere virtually indistinguishable from that of S. lohmanderi sp. nov. Caudomesal lamella (cl) directed meso-anteriad rather than completely mesad. Mesomeral process (m) considerably shorter than solenomere (s), without strongly protruding apical corners; solenomere apically forming more shallow concavity in comparison with S. lohmanderi sp. nov., and with an anterior process directed mostly distad rather than bent caudad. A very fine central filiform process (ff), similar to that of S. lohmanderi sp. nov., visible. Remarks: Despite the conspicuous similarities between the gonopods of S. astrabadensis and S. lohmanderi sp. nov., the two species can be rather easily distinguished by overall appearance: S. astrabadensis is less than 25 mm in length and entirely pallid, whereas S. lohmanderi sp. nov. is 28 - 29 mm long and has a brownish pigmentation. Likewise, the very similar gonopod conformation in S. persicus is in a way countervailed by the setose metazonal hind margins in this species; both S. astrabadensis and S. lohmanderi sp. nov. lack metazonal setae. Examples of species with (almost) completely identical gonopods but otherwise obvious differences in size and colouration are not uncommon in Pachyiulini (e. g. Frederiksen et al., 2012; Golovatch, 2018).	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEBF44EFF4F4C5AFDF4FB72.taxon	discussion	Known records: Ajara [geographic region], region of Batumi, “ Bortschcha ” [probably Borçka, Turkey, about 15 km south of the border with Georgia] (type locality). Note: The original gonopod drawings clearly show an apically concave solenomere which lacks both a rodlike mesal process and an anterior lamella, or the latter is very weakly pronounced. Thus, the species is here transferred from Amblyiulus to Syrioiulus.	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEBF44EFF2E4A2EFDADF970.taxon	distribution	Known records: All from Israel: Rehovot (type locality); by the Sea of Galilee (Verhoeff, 1923); Allone [Alonei] Abba, Beqaot S of Bet Shean [Beit She’an], Allonim, Nilit (Golovatch, 2018).	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEBF44EFED7482CFF5AF87B.taxon	distribution	Known records: Greece, Paros Island, Marathi (type locality).	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEBF44EFCA94E0AFA87FE64.taxon	distribution	Known records: Iran, Gorgan (type locality).	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEBF44EFC9E4D19FC3BFB40.taxon	distribution	Known records: Turkey, Keğubat near Kayseri (type locality).	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEBF44EFC9E4D19FC3BFB40.taxon	discussion	Note: The solenomere in this species ends up with a broad and blunt tip, while it lacks a rod-like mesal process, and the anterior lamella does not form an apical protrusion. This combination of characters does not correspond to the latest concept of Amblyiulus (see Golovatch, 2018) but it fits the current diagnosis of Syrioiulus. Thus the species is here transferred to Syrioiulus. However, the pronouncedly spoon-shaped promere, with a lateral prominence and the mesomeral process tightly contiguous with the solenomere, partly concealing the latter in lateral view, are untypical of Syrioiulus, but are rather common within the genera Amblyiulus and Dolichoiulus. Hence the question mark applied.	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEBF44EFC444A3CFC3CF9D7.taxon	distribution	Known records: Azerbaijan, Lenkoran [Lankaran] (type locality); Iran, the vicinity of Gorgan (Lohmander, 1932 a).	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFEBF44EFC4649CAFC0DF823.taxon	distribution	Known records: Iran, Garmab Plain near Gorgan (type locality).	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFE5F440FF0E4BA4FE65F9EE.taxon	distribution	Known records: Iran, eastern slopes of Bogrov Dāgh [Talysh Mts], “ Altschaly ” (type locality).	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFE5F440FF0E4BA4FE65F9EE.taxon	discussion	Note: Despite the unknown male characters, the vulval structure in this species is so similar with that observed in the other three blind representatives of Syrioiulus known from Iran that there can be no doubt that incarnatus is their congener.	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFE5F440FECD498DFF5AF943.taxon	distribution	Known records: Iran, Gilan Province, Siahkal (type locality).	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFE5F440FF244839FF5AF84F.taxon	distribution	Known records: Iran, 10 km N of Keredj [Karaj] (type locality).	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFE5F440FC774BA4FBF7FA62.taxon	distribution	Known records: Syria, Berzé [Barzeh, today part of Damascus] (type locality).	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFE5F443FC57491EFD22FE4F.taxon	description	Note: Athough the solenomere in this species seems to correspond with the current diagnosis of Syrioiulus, the intermediate lobe connecting the proximal sections of mesomeral process and solenomere, and especially the complete absence of apical denticles on the promere are characters that set S. posthirsutus apart from all remaining congeners, hence the question mark.	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFE6F443FED74D27FE62FCAE.taxon	description	Known records: Azerbaijan, Lenkoran [Lankaran], area of Talysh Mts (type locality).	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
450587FEFFE6F442FF314C47FE22FEF7.taxon	discussion	Note: This is another species of Syrioiulus deviating from the majority of congeners by the conformations of promere and mesomeral processes which are alike those of S. cappadocius. Furthermore, S. zarudnyi has a prominent anterior lamella of the solenomere (albeit apically not drawn into a process) which is also untypical of the genus. However, before a comprehensive revision and phylogeny of Syrioiulus and related genera is made, it seems better that this and the other two aforementioned doubtful species are placed in Syrioiulus. According to Lohmander (1932 a) the type locality of S. zarudnyi is the “ Tschemle-Rogan ” Spring in Arabistan (today Khuzestan Province). The toponym obviously refers to “ Tscheschme-Rogan ” [“ Cheshme ” (English transliteration) meaning “ spring ” in Persian], “ in the land of the Bakhtiary people ”. This is one of the sites visited by the original collector of this species, the Russian zoologist Nikolai Zarudny, during his forth expedition to Iran (Birula, 1905). The exact location of that spring remains Key to the species of Syrioiulus known from Iran based on characters of gonopods, vulvae and external somatic morphology	en	Vagalinski, Boyan (2020): A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae). Revue suisse de Zoologie 127 (1): 83-94, DOI: 10.35929/RSZ.0008, URL: http://dx.doi.org/10.35929/rsz.0008
