taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
4511E41DD826FFC2FDCAFA05F388FB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806774/files/figure.png	https://doi.org/10.5281/zenodo.3806774	Figs 1-5. Lectotype of Ennea radius Preston, 1910, type species of Dadagulella gen. nov. 1. Apical view. 2. Apertural view. 3. Last two whorls, from left side. 4. Last two whorls, from right side. 5. Umbilical view.	Figs 1-5. Lectotype of Ennea radius Preston, 1910, type species of Dadagulella gen. nov. 1. Apical view. 2. Apertural view. 3. Last two whorls, from left side. 4. Last two whorls, from right side. 5. Umbilical view.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD826FFC2FDCAFA05F388FB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806778/files/figure.png	https://doi.org/10.5281/zenodo.3806778	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD826FFC2FDCAFA05F388FB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806780/files/figure.png	https://doi.org/10.5281/zenodo.3806780	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD826FFC2FDCAFA05F388FB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806784/files/figure.png	https://doi.org/10.5281/zenodo.3806784	Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD826FFC2FDCAFA05F388FB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806788/files/figure.png	https://doi.org/10.5281/zenodo.3806788	Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996).	Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996).	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD826FFC2FDCAFA05F388FB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806792/files/figure.png	https://doi.org/10.5281/zenodo.3806792	Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza.	Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD826FFC2FDCAFA05F388FB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806796/files/figure.png	https://doi.org/10.5281/zenodo.3806796	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD826FFC2FDCAFA05F388FB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806800/files/figure.png	https://doi.org/10.5281/zenodo.3806800	Figs 65-68. Radulae of Dadagulella gen. nov. species. 65. D. radius radius, Amboni. 66. D. pembensis sp. nov., Ngezi, radula in situ on odontophore, ventral view. 67. D. pembensis sp. nov., Ngezi. 68. D. pembensis sp. nov., Ngezi, teeth from ventral end of radular ribbon. Abbreviations: c = central tooth; ii, iii, and iv = examples of bi-, tri- and quadricuspid teeth respectively. All scalebars = 10 μm.	Figs 65-68. Radulae of Dadagulella gen. nov. species. 65. D. radius radius, Amboni. 66. D. pembensis sp. nov., Ngezi, radula in situ on odontophore, ventral view. 67. D. pembensis sp. nov., Ngezi. 68. D. pembensis sp. nov., Ngezi, teeth from ventral end of radular ribbon. Abbreviations: c = central tooth; ii, iii, and iv = examples of bi-, tri- and quadricuspid teeth respectively. All scalebars = 10 μm.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD826FFC2FDCAFA05F388FB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806804/files/figure.png	https://doi.org/10.5281/zenodo.3806804	Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens.	Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD826FFC2FDCAFA05F388FB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806808/files/figure.png	https://doi.org/10.5281/zenodo.3806808	Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens.	Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD826FFC2FDCAFA05F388FB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD821FFCFFE7DFA86F34AF9FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806774/files/figure.png	https://doi.org/10.5281/zenodo.3806774	Figs 1-5. Lectotype of Ennea radius Preston, 1910, type species of Dadagulella gen. nov. 1. Apical view. 2. Apertural view. 3. Last two whorls, from left side. 4. Last two whorls, from right side. 5. Umbilical view.	Figs 1-5. Lectotype of Ennea radius Preston, 1910, type species of Dadagulella gen. nov. 1. Apical view. 2. Apertural view. 3. Last two whorls, from left side. 4. Last two whorls, from right side. 5. Umbilical view.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD821FFCFFE7DFA86F34AF9FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806778/files/figure.png	https://doi.org/10.5281/zenodo.3806778	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD821FFCFFE7DFA86F34AF9FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806792/files/figure.png	https://doi.org/10.5281/zenodo.3806792	Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza.	Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD821FFCFFE7DFA86F34AF9FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806796/files/figure.png	https://doi.org/10.5281/zenodo.3806796	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD821FFCFFE7DFA86F34AF9FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806800/files/figure.png	https://doi.org/10.5281/zenodo.3806800	Figs 65-68. Radulae of Dadagulella gen. nov. species. 65. D. radius radius, Amboni. 66. D. pembensis sp. nov., Ngezi, radula in situ on odontophore, ventral view. 67. D. pembensis sp. nov., Ngezi. 68. D. pembensis sp. nov., Ngezi, teeth from ventral end of radular ribbon. Abbreviations: c = central tooth; ii, iii, and iv = examples of bi-, tri- and quadricuspid teeth respectively. All scalebars = 10 μm.	Figs 65-68. Radulae of Dadagulella gen. nov. species. 65. D. radius radius, Amboni. 66. D. pembensis sp. nov., Ngezi, radula in situ on odontophore, ventral view. 67. D. pembensis sp. nov., Ngezi. 68. D. pembensis sp. nov., Ngezi, teeth from ventral end of radular ribbon. Abbreviations: c = central tooth; ii, iii, and iv = examples of bi-, tri- and quadricuspid teeth respectively. All scalebars = 10 μm.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD821FFCFFE7DFA86F34AF9FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806804/files/figure.png	https://doi.org/10.5281/zenodo.3806804	Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens.	Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD821FFCFFE7DFA86F34AF9FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806808/files/figure.png	https://doi.org/10.5281/zenodo.3806808	Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens.	Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD821FFCFFE7DFA86F34AF9FE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82CFFCDFE08F9C2F151F95E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806778/files/figure.png	https://doi.org/10.5281/zenodo.3806778	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82CFFCDFE08F9C2F151F95E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806796/files/figure.png	https://doi.org/10.5281/zenodo.3806796	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82CFFCDFE08F9C2F151F95E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82EFFCBFE16F960F0C8FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806778/files/figure.png	https://doi.org/10.5281/zenodo.3806778	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82EFFCBFE16F960F0C8FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806792/files/figure.png	https://doi.org/10.5281/zenodo.3806792	Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza.	Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82EFFCBFE16F960F0C8FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806804/files/figure.png	https://doi.org/10.5281/zenodo.3806804	Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens.	Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82EFFCBFE16F960F0C8FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806808/files/figure.png	https://doi.org/10.5281/zenodo.3806808	Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens.	Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82EFFCBFE16F960F0C8FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82EFFCBFE16F960F0C8FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806774/files/figure.png	https://doi.org/10.5281/zenodo.3806774	Figs 1-5. Lectotype of Ennea radius Preston, 1910, type species of Dadagulella gen. nov. 1. Apical view. 2. Apertural view. 3. Last two whorls, from left side. 4. Last two whorls, from right side. 5. Umbilical view.	Figs 1-5. Lectotype of Ennea radius Preston, 1910, type species of Dadagulella gen. nov. 1. Apical view. 2. Apertural view. 3. Last two whorls, from left side. 4. Last two whorls, from right side. 5. Umbilical view.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82EFFCBFE16F960F0C8FB62.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806796/files/figure.png	https://doi.org/10.5281/zenodo.3806796	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD828FFCAFD84FB2CF1CEFD01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806778/files/figure.png	https://doi.org/10.5281/zenodo.3806778	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD828FFCAFD84FB2CF1CEFD01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806796/files/figure.png	https://doi.org/10.5281/zenodo.3806796	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD828FFCAFD84FB2CF1CEFD01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD829FFC9FDAEFC8DF3A5FD8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806778/files/figure.png	https://doi.org/10.5281/zenodo.3806778	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD829FFC9FDAEFC8DF3A5FD8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806796/files/figure.png	https://doi.org/10.5281/zenodo.3806796	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD829FFC9FDAEFC8DF3A5FD8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82AFFC9FE51FD17F3E1F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806780/files/figure.png	https://doi.org/10.5281/zenodo.3806780	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82AFFC9FE51FD17F3E1F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82BFFC8FDC4FE8BF085F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806780/files/figure.png	https://doi.org/10.5281/zenodo.3806780	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82BFFC8FDC4FE8BF085F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806792/files/figure.png	https://doi.org/10.5281/zenodo.3806792	Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza.	Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82BFFC8FDC4FE8BF085F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806796/files/figure.png	https://doi.org/10.5281/zenodo.3806796	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82BFFC8FDC4FE8BF085F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD82BFFC8FDC4FE8BF085F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806778/files/figure.png	https://doi.org/10.5281/zenodo.3806778	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD835FFD6FDC4FE8BF650FA2E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806780/files/figure.png	https://doi.org/10.5281/zenodo.3806780	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD835FFD6FDC4FE8BF650FA2E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806796/files/figure.png	https://doi.org/10.5281/zenodo.3806796	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD835FFD6FDC4FE8BF650FA2E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD835FFD5FDC1FA70F745F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806780/files/figure.png	https://doi.org/10.5281/zenodo.3806780	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD835FFD5FDC1FA70F745F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806792/files/figure.png	https://doi.org/10.5281/zenodo.3806792	Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza.	Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD835FFD5FDC1FA70F745F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806796/files/figure.png	https://doi.org/10.5281/zenodo.3806796	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD835FFD5FDC1FA70F745F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD837FFD4FE46FE8BF0D2FA35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806780/files/figure.png	https://doi.org/10.5281/zenodo.3806780	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD837FFD4FE46FE8BF0D2FA35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806796/files/figure.png	https://doi.org/10.5281/zenodo.3806796	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD837FFD4FE46FE8BF0D2FA35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD837FFD2FDF8FA7AF196FD75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806780/files/figure.png	https://doi.org/10.5281/zenodo.3806780	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD837FFD2FDF8FA7AF196FD75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806792/files/figure.png	https://doi.org/10.5281/zenodo.3806792	Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza.	Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD837FFD2FDF8FA7AF196FD75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806796/files/figure.png	https://doi.org/10.5281/zenodo.3806796	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD837FFD2FDF8FA7AF196FD75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806800/files/figure.png	https://doi.org/10.5281/zenodo.3806800	Figs 65-68. Radulae of Dadagulella gen. nov. species. 65. D. radius radius, Amboni. 66. D. pembensis sp. nov., Ngezi, radula in situ on odontophore, ventral view. 67. D. pembensis sp. nov., Ngezi. 68. D. pembensis sp. nov., Ngezi, teeth from ventral end of radular ribbon. Abbreviations: c = central tooth; ii, iii, and iv = examples of bi-, tri- and quadricuspid teeth respectively. All scalebars = 10 μm.	Figs 65-68. Radulae of Dadagulella gen. nov. species. 65. D. radius radius, Amboni. 66. D. pembensis sp. nov., Ngezi, radula in situ on odontophore, ventral view. 67. D. pembensis sp. nov., Ngezi. 68. D. pembensis sp. nov., Ngezi, teeth from ventral end of radular ribbon. Abbreviations: c = central tooth; ii, iii, and iv = examples of bi-, tri- and quadricuspid teeth respectively. All scalebars = 10 μm.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD837FFD2FDF8FA7AF196FD75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806804/files/figure.png	https://doi.org/10.5281/zenodo.3806804	Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens.	Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD837FFD2FDF8FA7AF196FD75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806808/files/figure.png	https://doi.org/10.5281/zenodo.3806808	Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens.	Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD837FFD2FDF8FA7AF196FD75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD837FFD2FDF8FA7AF196FD75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806778/files/figure.png	https://doi.org/10.5281/zenodo.3806778	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD831FFD1FE1EFD39F6B1FD9E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806780/files/figure.png	https://doi.org/10.5281/zenodo.3806780	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD831FFD1FE1EFD39F6B1FD9E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD832FFD1FDE8FD21F0B7F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806780/files/figure.png	https://doi.org/10.5281/zenodo.3806780	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD832FFD1FDE8FD21F0B7F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806796/files/figure.png	https://doi.org/10.5281/zenodo.3806796	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD832FFD1FDE8FD21F0B7F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD833FFDFFDDAFE8BF691FEAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806780/files/figure.png	https://doi.org/10.5281/zenodo.3806780	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD833FFDFFDDAFE8BF691FEAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806804/files/figure.png	https://doi.org/10.5281/zenodo.3806804	Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens.	Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD833FFDFFDDAFE8BF691FEAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806808/files/figure.png	https://doi.org/10.5281/zenodo.3806808	Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens.	Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD833FFDFFDDAFE8BF691FEAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD83CFFDEFE26FEF0F07BFBF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806784/files/figure.png	https://doi.org/10.5281/zenodo.3806784	Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD83CFFDEFE26FEF0F07BFBF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD83DFFDDFE63FBB7F6E2FD5C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806784/files/figure.png	https://doi.org/10.5281/zenodo.3806784	Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD83DFFDDFE63FBB7F6E2FD5C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD83EFFDCFE00FD63F6D1F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806784/files/figure.png	https://doi.org/10.5281/zenodo.3806784	Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD83EFFDCFE00FD63F6D1F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806796/files/figure.png	https://doi.org/10.5281/zenodo.3806796	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD83EFFDCFE00FD63F6D1F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD838FFDAFDA1FE8BF394FBA8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806784/files/figure.png	https://doi.org/10.5281/zenodo.3806784	Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD838FFDAFDA1FE8BF394FBA8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806796/files/figure.png	https://doi.org/10.5281/zenodo.3806796	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD838FFDAFDA1FE8BF394FBA8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD839FFD8FE7EFBF6F6ECFCF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806788/files/figure.png	https://doi.org/10.5281/zenodo.3806788	Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996).	Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996).	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD839FFD8FE7EFBF6F6ECFCF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD83BFFE6FE7BFCBAF724F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806788/files/figure.png	https://doi.org/10.5281/zenodo.3806788	Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996).	Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996).	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD83BFFE6FE7BFCBAF724F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806812/files/figure.png	https://doi.org/10.5281/zenodo.3806812	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD83BFFE6FE7BFCBAF724F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806780/files/figure.png	https://doi.org/10.5281/zenodo.3806780	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD83BFFE6FE7BFCBAF724F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806784/files/figure.png	https://doi.org/10.5281/zenodo.3806784	Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD83BFFE6FE7BFCBAF724F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806796/files/figure.png	https://doi.org/10.5281/zenodo.3806796	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD83BFFE6FE7BFCBAF724F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806778/files/figure.png	https://doi.org/10.5281/zenodo.3806778	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
4511E41DD83BFFE6FE7BFCBAF724F80C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3806774/files/figure.png	https://doi.org/10.5281/zenodo.3806774	Figs 1-5. Lectotype of Ennea radius Preston, 1910, type species of Dadagulella gen. nov. 1. Apical view. 2. Apertural view. 3. Last two whorls, from left side. 4. Last two whorls, from right side. 5. Umbilical view.	Figs 1-5. Lectotype of Ennea radius Preston, 1910, type species of Dadagulella gen. nov. 1. Apical view. 2. Apertural view. 3. Last two whorls, from left side. 4. Last two whorls, from right side. 5. Umbilical view.	2013-02-04	Rowson, Ben;Tattersfield, Peter		Zenodo	biologists	Rowson, Ben;Tattersfield, Peter			
