taxonID	type	description	language	source
446F87A7322B8A71FF77825DB96EFA88.taxon	description	(Figs. 1 – 4)	en	Li, Zhu, Cuccodoro, Giulio, Chen, Li (2021): On the identity of Oberea acuta Gressitt, 1951, O. bisbipunctata Pic, 1916, O. notata Pic, 1936 and O. shimomurai Kurihara & N. Ohbayashi, 2007 (Coleoptera, Cerambycidae, Lamiinae) with a new synonym. Zootaxa 5039 (3): 352-362, DOI: 10.11646/zootaxa.5039.3.2
446F87A7322B8A71FF77825DB96EFA88.taxon	description	Redescription. Body (Figs. 1 – 3) 12.0 – 14.0 mm long and 2.0 mm wide. Head black, antennae dark brown to black, scape black; pronotum testaceous or red brown, with four or five black disctal spots: two round black spots beside center, two black markings at base on each side, and one small black spot in the middle at base, the latter occasionally absent, all those markings sometimes fused (Fig. 2); scutellum black or yellow brown; elytra black except for testaceous area near scutellum; abdomen testaceous except occasionally parts of sides of first two or first three segments (Fig. 3 e – h); legs predominantly reddish brown, tibiae and tarsi darker, hind tibiae black. Body clothed with short pale gray pubescence. Vertex distinctly depressed at middle with a groove; eyes very large, lower lobes twice as long as gena in male and 1.5 times as long as gena in female. Frons as long as wide in male (Fig. 3 a) and 2 times as long as wide in female (Fig. 3 b). Antennae almost reaching apical 7 / 8 th of elytra in male (Fig. 1 g), shorter in female, reaching apical 3 / 4 th of elytra (Fig. 1 e & 1 f); third segment longer than scape and fourth. Pronotum slightly wider than long or as long as wide, with lateral outlines weakly convex and subparallel in dorsal view; disc slightly raised in middle, finely and densely punctured. Scutellum almost squared, slightly emarginated. Elytra nearly 3.5 times as long as humeral width, and 3.5 times as long as head and prothorax combined, slightly narrowed in middle and broadened preapically; elytral apex obliquely emarginate with acute external angle; disc with fine punctures arranged in six longitudinal rows, gradually finer and irregular towards apex. Abdominal ventrites with punctures at sides. Hind femora reaching middle of abdominal segment II; hind tibiae almost 1.5 times as long as tarsi. Abdominal ventrite V with shallow triangular depression in male (Fig. 3 c) and medially grooved in female (Fig. 3 d). Male terminalia. Tergite VIII broader than long, apex truncated and moderately emarginated, clothed with short hairs (Fig. 4 a); tegmen curved in lateral view; parameres elongate, mostly covered with long hair on apical half, base of each paramere transversely and obliquely ridged ventrally, the latter ridge covered with dense hairs (Fig. 4 b); aedeagal median lobe slightly curved in lateral view, median struts 3 / 5 times as long as whole median lobe, dorsal plate slightly longer than ventral plate, median foramen rounded (Fig. 4 c); endophallus with 2 pairs of rods on apical portion, longer pair slender baculiform, about 2 times as long as shorter pair (Fig. 4 d – e). Female genitalia. Capsule of spermatheca with sclerotized rounded apical lobe and a stalk; spermathecal duct longer than capsule; spermathecal gland extended from strongly sclerotized ring attached to spermathecal duct (Fig. 4 f).	en	Li, Zhu, Cuccodoro, Giulio, Chen, Li (2021): On the identity of Oberea acuta Gressitt, 1951, O. bisbipunctata Pic, 1916, O. notata Pic, 1936 and O. shimomurai Kurihara & N. Ohbayashi, 2007 (Coleoptera, Cerambycidae, Lamiinae) with a new synonym. Zootaxa 5039 (3): 352-362, DOI: 10.11646/zootaxa.5039.3.2
446F87A7322B8A71FF77825DB96EFA88.taxon	materials_examined	Material examined. CHINA: SICHUAN: Holotype, ♀, Chengdu, China, alt. 500 met., 1940, leg. T. L. Sie (SYSU); 1 ♀, Mt. Omei, 31. VII. 1938, leg. C. S. Tai (SYSU); 1 ♀, Sichuan Pronvince, Chengdu, 1941 (SWU); 1 ♀, China, Sichuan, Emei Mountain, Guangfusi, 19. V. 1962, leg. Lin Duan-fang et al. (SWU); 2 ♂♂, China, Sichuan, Mt. Emei, Baoguosi, 9. V. 1957, leg. Cheng Han-Hua (NKU); 1 ♂ 1 ♀, ditto, but 11. V. 1957 (NKU); 1 ♂ 1 ♀, Sichuan, Emeishan, 6. VI. 1955, leg. Ke-Ren Huang, Gen-Tao Jin (IZCAS); 1 ♀, Sichuan, Emeishan, 580 m, 24. VI. 1955, leg. Tian-Rong Huang (IZCAS); 1 ♂, same data but 25. VI. 1955, leg. Huai-Ju Leng (IZCAS); 1 ♀, Sichuan, Baoguosi, 700 m, 20. IV. 1957, You-Cai Lu (IZCAS); 4 ♂♂ 3 ♀♀, Sichuan, Emeishan, Baoguosi, 550 - 750 m, 5 - 9. V. 1957, leg. Ke-Ren Huang (IZCAS); 1 ♀, same data but 12. V. 1957 (IZCAS); 1 ♀, Sichuan, Emeishan, Baoguosi, 550 - 750 m, 2. VI. 1957, leg. Ke-Ren Huang (IZCAS); 2 ♂♂ 1 ♀, same data but 3. VI. 1957 (IZCAS); 1 ♂ 1 ♀, same data but 7 - 8. VI. 1957 (IZCAS); 1 ♀, Sichuan, Emeishan, Baoguosi, 550 - 750 m, 12. VI. 1957, leg. You-Cai Lu (IZCAS); 7 ♂♂ 1 ♀, same data but 6 - 11. V. 1957 (IZCAS); 1 ♀, same data but 17. V. 1957 (IZCAS); 1 ♀, same data but 22. VI. 1957 (IZ- CAS); 1 ♂, same data but 5. VII. 1957; 2 ♂♂ 1 ♀, Sichuan, Emeishan, Baoguosi, 550 - 750 m, 7 - 8. VI. 1957, leg. Zong- Yuan Wang (IZCAS); 1 ♀, same data but 26. V. 1957; 1 ♀, same data but 9. VI. 1957 (IZCAS); 3 ♂♂ 1 ♀, same data but 25 - 26. V. 1957, leg. Fu-Xing Zhu; 3 ♂♂, same data but 20 - 24. V. 1957; 2 ♂♂, same data but 11 - 21. VI. 1957; 2 ♂♂, Sichuan, Emeishan, Qingyinge, 23. V. 1957, leg. Ke-Ren Huang (IZCAS); YUNNAN: 1 ♂, Yunnan Province, Cangyuan County, VII. 2007, leg. Wu Gui-Yi (SWU); 1 ♂, Yunnan Province, Yanjin, 6. V. 1920, leg. Yang Fu-Xing (SWU).	en	Li, Zhu, Cuccodoro, Giulio, Chen, Li (2021): On the identity of Oberea acuta Gressitt, 1951, O. bisbipunctata Pic, 1916, O. notata Pic, 1936 and O. shimomurai Kurihara & N. Ohbayashi, 2007 (Coleoptera, Cerambycidae, Lamiinae) with a new synonym. Zootaxa 5039 (3): 352-362, DOI: 10.11646/zootaxa.5039.3.2
446F87A7322B8A71FF77825DB96EFA88.taxon	distribution	Distribution. China (Sichuan, Yunnan).	en	Li, Zhu, Cuccodoro, Giulio, Chen, Li (2021): On the identity of Oberea acuta Gressitt, 1951, O. bisbipunctata Pic, 1916, O. notata Pic, 1936 and O. shimomurai Kurihara & N. Ohbayashi, 2007 (Coleoptera, Cerambycidae, Lamiinae) with a new synonym. Zootaxa 5039 (3): 352-362, DOI: 10.11646/zootaxa.5039.3.2
446F87A7322B8A71FF77825DB96EFA88.taxon	discussion	Remarks. In the original description Gressitt (1951) stated that the holotype was male, however it is actually female (Fig. 1 a, b). In the materials from Mt. Emei we found another female with the same color patterns as the holotype (Fig. 1 c), but others were identified as O. notata or O. bisbipunctata. However, after careful examination of all these specimens we came to the conclusion that they are all conspecific, and that the species is just quite variable with respect to the color pattern on pronotum and abdomen (Figs. 2, 3 e – h). This species is very similar to O. notata, from which it differs mainly by the abdominal ventrite V yellowish brown rather than black, by the rods of the intersac and by the shape of the spermatheca.	en	Li, Zhu, Cuccodoro, Giulio, Chen, Li (2021): On the identity of Oberea acuta Gressitt, 1951, O. bisbipunctata Pic, 1916, O. notata Pic, 1936 and O. shimomurai Kurihara & N. Ohbayashi, 2007 (Coleoptera, Cerambycidae, Lamiinae) with a new synonym. Zootaxa 5039 (3): 352-362, DOI: 10.11646/zootaxa.5039.3.2
446F87A7322E8A72FF7785C6BFBFFB79.taxon	description	(Figs. 5 – 6)	en	Li, Zhu, Cuccodoro, Giulio, Chen, Li (2021): On the identity of Oberea acuta Gressitt, 1951, O. bisbipunctata Pic, 1916, O. notata Pic, 1936 and O. shimomurai Kurihara & N. Ohbayashi, 2007 (Coleoptera, Cerambycidae, Lamiinae) with a new synonym. Zootaxa 5039 (3): 352-362, DOI: 10.11646/zootaxa.5039.3.2
446F87A7322E8A72FF7785C6BFBFFB79.taxon	diagnosis	Diagnosis. Head black; antennae dark brown to reddish brown, scape black; pronotum testaceous or red brown with two pairs of black spots: one pair beside center and the other at base, scutellum yellow brown; elytra black except testaceous area near scutellum; abdomen testaceous except abdominal ventrite V largely black, sometimes ventrite II or III with black spots laterally; legs predominantly reddish brown or black, with tibiae and tarsi darker, hind tibiae reddish brown to black. Body clothed with short pale gray pubescence. Antennae almost reaching elytral apex in male, shorter in female, reaching apical 5 / 6 th of elytra; third segment longer than scape and fourth. Prothorax slightly wider than long; pronotum slightly raised in middle, finely and densely punctured. Scutellum almost squared, slightly emarginated. Elytra nearly 3.5 times as long as humeral width, and 3.5 times as long as head and prothorax combined; disc with fine punctures. Hind femora reaching middle of abdominal segment II; hind tibiae almost 1.5 times as long as tarsi. Male terminalia. Tergite VIII broader than long, apex truncated and slightly emarginated, densely clothed with short hairs (Fig. 6 d); tegmen curved in lateral view; parameres elongate, mostly covered with long setae on apical half; each paramere with transverse oblique ridge covered with dense fine hairs on ventral side (Fig. 6 a – c); median lobe 1.3 times as long as tegmen and slightly curved in lateral view; median struts nearly 3 / 5 times as long as whole median lobe; dorsal plate slightly longer than ventral plate; apex of ventral plate slightly pointed; median foramen rounded (Fig. 6 e); endophallus with 2 pairs of rods on apical portion, longer pair very slender and baculiform, about 2 times as long as shorter pair (Fig. 6 f). Female genitalia (Fig. 6 g): Capsule of spermatheca with sclerotized rounded apical lobe and a stalk; spermathecal duct longer than capsule; spermathecal gland extended from strongly sclerotized ring attached to spermathecal duct. Material examined. CHINA: SHANGHAI: Holotype, ♀, Prov. Kiangsu, Shanghai (MNHN); 1 ♂, ZoSe, 12. VI. 1931 (IZCAS); 1 ♂, ZoSe, 1. VI. 1934, leg. A. Savio (IZCAS); 1 ♂, ZoSe, 3. VI. 1934, leg. A. Savio (IZCAS); 1 ♂ 1 ♀, ZoSe, 4. VI. 1934, leg. A. Savio (IZCAS); 1 ♂, ZoSe, 6. VI. 1934, leg. A. Savio (IZCAS); 1 ♀, Chine, Prov. Kiangsu, Shanghai (MHNG); JIANGSU: 1 ♀, Nanking, V. 1933 (NAU); 1 ♂, Yangcheou, 20. IV. 1930 (IZCAS); 1 ♀, Yangcheou, 15. VI. 1930 (IZCAS); ANHUI: 1 ♀, Anhwei, Ningkwo, 18. VII. 1920 (IZCAS); JIANGXI: 2 ♂♂, / Kiujiang, Chekiang / [holotype and a mislabeled paratype of O. notata v. rufoantennata Breuning] (MHNG); GUIZHOU: 2 ♀♀, Kweichow (MHNG); Guangdong: 1 ♂ 1 ♀, Kwangtung, S. China, Yaoshan, Lin-hsien (Dist.), 12. V. 1934, leg. F. K. To (SYSU, Ce- 004390 (♂), Ce- 004393 (♀ )); 1 ♀, Kwangtung, S. China, Lung Ping Hui, Lin-hsien (District), 15. V. 1934, leg. F. K. To (SYSU, Ce- 004392); 1 ♂, Kwangtung, S. China, Lung Ping Hui, Lin-hsien (District), 16 - 17. V. 1934, leg. F. K. To (SYSU, Ce- 004376); 1 ♂, Kwangtung, S. China, Sing Taz Chan, Lin-hsien (District), 22 - 23. V. 1934, leg. F. K. To (SYSU, Ce- 004388); 1 ♂, Kwangtung, S. China, Taam yuen Tung, Lin-hsien (District), 4 - 6. VI. 1934, leg. F. K. To (SYSU, Ce- 004389); GUANGXI: 1 ♂, Yangshuo, 25. V. 1948 (SYSU, Ce- 004075); 1 ♂, Yongle, 16. IV. 1959 (SYSU, Ce- 004394); HUNAN: 1 ♀, Hunan Prov., S. China, Tai Kwong Village, Lam Mo District, 26 - 28. VI. 1934, leg. F. K. To (SYSU, Ce- 004074).	en	Li, Zhu, Cuccodoro, Giulio, Chen, Li (2021): On the identity of Oberea acuta Gressitt, 1951, O. bisbipunctata Pic, 1916, O. notata Pic, 1936 and O. shimomurai Kurihara & N. Ohbayashi, 2007 (Coleoptera, Cerambycidae, Lamiinae) with a new synonym. Zootaxa 5039 (3): 352-362, DOI: 10.11646/zootaxa.5039.3.2
446F87A7322E8A72FF7785C6BFBFFB79.taxon	distribution	Distribution. China: Anhui (new record), Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangsu, Jiangxi, Shanghai, Sichuan, Taiwan, Zhejiang.	en	Li, Zhu, Cuccodoro, Giulio, Chen, Li (2021): On the identity of Oberea acuta Gressitt, 1951, O. bisbipunctata Pic, 1916, O. notata Pic, 1936 and O. shimomurai Kurihara & N. Ohbayashi, 2007 (Coleoptera, Cerambycidae, Lamiinae) with a new synonym. Zootaxa 5039 (3): 352-362, DOI: 10.11646/zootaxa.5039.3.2
446F87A7322E8A72FF7785C6BFBFFB79.taxon	discussion	Remarks. It is newly recorded from Anhui province based on specimens deposited in IZCAS. Breuning (1962) described Oberea notata var. rufoantennata Breuning based on the specimens with reddish brown antennae from Kiukiang, Chekiang. However, Kiukiang is in Jiangxi rather than Chekiang (= Zhejiang, Now). And O. notata var. kwangtungensis, based on specimens from Kwangtung, Lien-ping, with the reddish antenna and black hind tibias. Both are unavailable infrasubspecific names (Li, Cuccodoro & Chen, 2015). Based on the examination on the specimens from Guangdong, Guangxi, the antennae are usually reddish brown and hind tibiae black. Kurihara & Ohbayashi (2007) described O. shimomurai from Taiwan, and subsequently Kurihara (2009) redescribed O. notata mentioning that the two species were very similar, but could be distinguished by their body form and male genitalia. However, the redescription of O. notata by Kurihara (2009) is based on a female from Baoguoshi, Mt. Emei, Sichuan, which is the type locality of O. acuta, which does not have the abdominal ventrite V black. And after comparing his description and drawing of male genitalia with O. acuta, we conclude that the specimen used by Kurihara to described O. notata belongs to O. acuta. And finally, after carefully comparing the female holotype and male genitalia of O. notata with his description and drawings of O. shimomurai, we concluded that the latter is a junior synonym of O. notata.	en	Li, Zhu, Cuccodoro, Giulio, Chen, Li (2021): On the identity of Oberea acuta Gressitt, 1951, O. bisbipunctata Pic, 1916, O. notata Pic, 1936 and O. shimomurai Kurihara & N. Ohbayashi, 2007 (Coleoptera, Cerambycidae, Lamiinae) with a new synonym. Zootaxa 5039 (3): 352-362, DOI: 10.11646/zootaxa.5039.3.2
446F87A7322D8A7CFF7784EAB884FE9A.taxon	description	(Fig. 7)	en	Li, Zhu, Cuccodoro, Giulio, Chen, Li (2021): On the identity of Oberea acuta Gressitt, 1951, O. bisbipunctata Pic, 1916, O. notata Pic, 1936 and O. shimomurai Kurihara & N. Ohbayashi, 2007 (Coleoptera, Cerambycidae, Lamiinae) with a new synonym. Zootaxa 5039 (3): 352-362, DOI: 10.11646/zootaxa.5039.3.2
446F87A7322D8A7CFF7784EAB884FE9A.taxon	diagnosis	Diagnosis. Head black; antennae dark brown; pronotum testaceous or red brown with two pairs of black spots: one pair near center and the other at base; scutellum yellow brown; elytra testaceous except for apical 1 / 5 black; abdomen testaceous except abdominal ventrite V mostly black and abdominal ventrite I partly black; legs predominantly reddish brown, with tibiae and tarsi darker, and hind tibiae dark brown. Antennae distinctly shorter than body, reaching 3 / 4 th of elytra; third segment longer than scape and fourth. Elytra nearly 4 times as long as humeral width, and 3.5 times as long as head and prothorax combined; elytral disc with fine punctures. Hind tibiae almost 2 times as long as tarsi.	en	Li, Zhu, Cuccodoro, Giulio, Chen, Li (2021): On the identity of Oberea acuta Gressitt, 1951, O. bisbipunctata Pic, 1916, O. notata Pic, 1936 and O. shimomurai Kurihara & N. Ohbayashi, 2007 (Coleoptera, Cerambycidae, Lamiinae) with a new synonym. Zootaxa 5039 (3): 352-362, DOI: 10.11646/zootaxa.5039.3.2
446F87A7322D8A7CFF7784EAB884FE9A.taxon	materials_examined	Material examined. VIETNAM: Holotype, ♂, Hanoi (MNHN); 1 ♀, Tonkin, Hanoi, 18. v. 1987. Jeanvoine leg. (MHNG); 1 ♀, Tonkin, Langsong (MHNG); 1 ♀, Vietnam, Annam (MHNG).	en	Li, Zhu, Cuccodoro, Giulio, Chen, Li (2021): On the identity of Oberea acuta Gressitt, 1951, O. bisbipunctata Pic, 1916, O. notata Pic, 1936 and O. shimomurai Kurihara & N. Ohbayashi, 2007 (Coleoptera, Cerambycidae, Lamiinae) with a new synonym. Zootaxa 5039 (3): 352-362, DOI: 10.11646/zootaxa.5039.3.2
446F87A7322D8A7CFF7784EAB884FE9A.taxon	distribution	Distribution. Vietnam: Central and Northern.	en	Li, Zhu, Cuccodoro, Giulio, Chen, Li (2021): On the identity of Oberea acuta Gressitt, 1951, O. bisbipunctata Pic, 1916, O. notata Pic, 1936 and O. shimomurai Kurihara & N. Ohbayashi, 2007 (Coleoptera, Cerambycidae, Lamiinae) with a new synonym. Zootaxa 5039 (3): 352-362, DOI: 10.11646/zootaxa.5039.3.2
446F87A7322D8A7CFF7784EAB884FE9A.taxon	discussion	Remarks. In the titan database, O. bisbipunctata ssp. discoreducta Breuning, 1969 is treated as subspecies. However, according to Lin & Tavakilian (2019), it is the synonym of O. bisbipunctata. According to the original description and comparison with two types, the subspecies is similar to the O. bisbipunctata, but lack of the two black premedian disc spots on the pronotum. The pronotal spots could vary within species. It is proper to be treated as the junior synonym. Gressitt (1939) reported Oberea bisbipunctata as new to China based on specimens from Guangdong. However, according to the description and pictures provided by Gressitt (1939, Plate III, fig. 11 & 12) and the examination of the specimens from SYSU, these specimens are clearly O. notata, whose distribution includes also “ Kwangtung (Yimna Shan, Yao Shan. Lien), Szechuan (Suifu = [Yibin, now], Omei = [Emei, now]), Kwangsi (Yangso = [Yangshuo, now]) ” (Gressitt, 1951). The specimens from Guangdong and Guangxi recorded in Gressitt (1951) are O. notata, and those from Sichuan are most likely O. acuta. According to the description and picture (Pu, 1980, Plate XII, fig 170), the O. bisbipunctata recorded in “ Economic Insect Fauna of China Fasc, 19 ” (Pu, 1980) belongs to O. acuta. The picture of O. bisbipunctata from California Academy of Sciences, determined by Gressitt (Bezark, 2020) belongs to O. notata. Wang (2003) reported the distribution of the species includes Liaoning, NE China, but the figure used was from Pu (1980). The distribution in China of O. bisbipunctata reported by Hua (2002) as from “ NE China, Zhejiang, Guangdong, Guangxi, Guizhou, Sichuan ” is based on the mistaken records, and there are no details of his records from Zhejiang and Guizhou. And the entire specimen we examined identified as O. bisbipunctata are O. notata or O. acuta. Our conclusion is that at present there is no evidence that specimens of O. bisbipunctata were found outside of Vietnam.	en	Li, Zhu, Cuccodoro, Giulio, Chen, Li (2021): On the identity of Oberea acuta Gressitt, 1951, O. bisbipunctata Pic, 1916, O. notata Pic, 1936 and O. shimomurai Kurihara & N. Ohbayashi, 2007 (Coleoptera, Cerambycidae, Lamiinae) with a new synonym. Zootaxa 5039 (3): 352-362, DOI: 10.11646/zootaxa.5039.3.2
