taxonID	type	description	language	source
433341A9E8785D64B97F48A3AFF1B9D5.taxon	description	Figs 4, 5, 17, 18	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
433341A9E8785D64B97F48A3AFF1B9D5.taxon	diagnosis	Diagnosis. The nominate subspecies cannot be distinguished from S. brevisunca yunnanensis Wu & Fang, 2009 (Fig. 6) externally but the morphology of male genitalia is diagnostic. In the male genitalia, the left process of juxta is long and the right process is finger-shaped apically in S. brevisunca brevisunca (Figs 17, 18), whereas the left process of juxta is short and the right process is blunt apically, without the finger-shaped apex in S. brevisunca yunnanensis (Fig. 15).	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
433341A9E8785D64B97F48A3AFF1B9D5.taxon	distribution	Distribution. China (Hainan, Guangxi, Yunnan), Vietnam (Fig. 25).	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
888A6FFC00AB5BE1A614EAFD233764D6.taxon	description	Figs 4, 5, 6, 15, 17, 18	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
704BB84D33C153E8A6F416F904E0E5E2.taxon	description	Figs 9, 12, 21, 23	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
704BB84D33C153E8A6F416F904E0E5E2.taxon	materials_examined	Specimens examined. 1 ♂, China, Prov. Yunnan, Pu'er City, Manxieba Village, 3. VI. 2018, leg. HL. Han, J. Wu and MR. Li, genit. prep. WuJ- 109 - 1 (NEFU); 1 ♂, China, Chongqing Municipality, Mt. Simian, 23. VII- 6. VIII. 2018, leg. GX. Wang and WJ. Li, genit. prep. WuJ- 539 - 1 (NEFU); 2 ♂, China, Chongqing Municipality, Mt. Simian, 24 - 30. VII. 2019, leg. TT. Zhao and SC. Deng, genit. prep. WuJ- 538 - 1 (NEFU); 4 ♀, China, Chongqing Municipality, Mt. Simian, 29. VII. - 2. VIII. 2020, leg. HL. Han and J. Wu, genit. prep. WuJ- 540 - 2 and 541 - 2 (NEFU). Female genitalia (Fig. 23). Papillae anales flattened, foot-shaped, covered with dense hairs on surface. Postvaginal plate flattened, strongly sclerotised. Apophysis anterioris highly modified, short, tongue-shaped; apophysis posterioris long and slender, c. 3 x length of apophysis anterioris. Ductus bursae long, membranous, not spiral-shaped. Corpus bursae pear-shaped, twisted in its apical part, with a strongly sclerotised, nearly elliptical central signum.	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
704BB84D33C153E8A6F416F904E0E5E2.taxon	distribution	Distribution. China (Hubei, Chongqing, Sichuan, Yunnan, Xizang); Thailand, Myanmar (Fig. 25).	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
B2C69A8E880C55DCB577BDEAC1A42EAF.taxon	description	Figs 1, 2, 13, 14	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
B2C69A8E880C55DCB577BDEAC1A42EAF.taxon	materials_examined	Material examined. Holotype. ♂, China, Xizang Autonomous Region, Linzhi (= Nyingchi) City, Motuo (= Medog) County, Gedang Countryside, 25. V. - 5. VI. 2021, leg. J. Wu and JJ. Fan, genit. prep. WuJ- 519 - 1 (NEFU). Paratypes. 4 ♂, same data as for holotype, genit. prep. WuJ- 520 - 1 (NEFU).	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
B2C69A8E880C55DCB577BDEAC1A42EAF.taxon	diagnosis	Diagnosis. The new species can be easily distinguished from the known species by its appearance: the antennae are broadly bipectinated at basal 3 / 4 in male; the thorax is black mixed with a little yellow; the base and costal margin area of forewing are dark brown to black; the rounded patch located on the outside of the cell is blurry; the abdomen bears two distinct black hair tufts dorsally. In the other Asian congeners (Figs 3 - 12) the male antennae are bipectinated only in the basal half; the ground colour of the thorax and forewing is yellow mixed with black; the medial patch of the forewing is more distinct than in S. medogensis sp. nov. (Figs 1, 2); the dorsal black hair tuft on the abdomen is weak. The male genitalia are clearly different from those of the other congeners: in S. medogensis sp. nov. (Figs 13, 14), the valva has a well-developed saccular process and the juxta is symmetrical. However, the same structures in other Asian species (Figs 15 - 22) are very different: the valvae without saccular processes; the juxta are asymmetrical with a lateral process.	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
B2C69A8E880C55DCB577BDEAC1A42EAF.taxon	description	Description. Adult (Figs 1, 2). Wingspan 30 - 32 mm in male. Head yellow; labial palpus short, yellow; male antennae brown, broadly bipectinated in basal 3 / 4 then serrate. Thorax and tegula black mixed with a little yellow. Scales on legs black to yellow. Forewing broad, ground colour brown, wing base and costal margin area dark brown to black, outer margin area pale brown; a large, silky reddish brown, rounded medial patch located at outside of cell; subterminal line narrow, black, smoothly curved, running from costal margin near the apex to the tornus, almost parallel to outer margin, bearing a dent in the region between vines R 4 and R 5 towards base; wing venation visible in outer margin area, black; fringe yellow. Hindwing brown; fringe yellow. Abdomen yellow, with two distinct black dorsal hair tufts and long black hairs at terminal area. Male genitalia (Figs 13, 14). Uncus short, weakly bifid apically. Gnathos rod-shaped, blunt apically. Tegumen broad. Valva broad at base, cucullus rounded; costa slightly concave, bearing a triangular process at c. 1 / 3 distance from base that is covered by dense long setae; sacculus inflated, slightly sclerotised, densely covered with setae in upper half; saccular process strongly sclerotised, curved inwardly in a hook-shaped, bifid process near middle and tapering from base to apex, pointed apically. Juxta sclerotised, symmetrical, horseshoe-shaped. Saccus not obvious. Phallus slender, smoothly curved; vesica without cornuti. Female genitalia. Unknown.	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
B2C69A8E880C55DCB577BDEAC1A42EAF.taxon	distribution	Distribution. China (Xizang: Motuo) (Fig. 25).	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
B2C69A8E880C55DCB577BDEAC1A42EAF.taxon	etymology	Etymology. The species is named Squamosa medogensis for its type-locality in Motuo County, Xizang Autonomous Region, China.	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
764B2FDB71225524B2DD7FF46F93F48F.taxon	description	Figs 7, 8, 19, 20	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
764B2FDB71225524B2DD7FF46F93F48F.taxon	materials_examined	Material examined. Holotype. ♂, China, Xizang Autonomous Region, Linzhi (= Nyingchi) City, Motuo (= Medog) County, Beibeng Countryside, Dergong Village, 850 m a. s. l., 25. V. - 4. VI. 2021, leg. HL. Han, genit. prep. WuJ- 518 - 1 (NEFU). Paratypes. 1 ♂, India, sept. or. W. Meghalaya, Garo Hills, Nokrek Nat. Park, 25 ° 40 ' N, 91 ° 04 ' E, 2 - 13. VII. 1997, 1150 m a. s. l., leg. Afonin and Siniaev, genit. prep. 16149 (MWM / ZSM); 1 ♂, Myanmar, Putao, 550 m a. s. l., 27. IV. 1998, leg. Murzin and Sinjaev, genit. prep. 16169 (MWM / ZSM); 1 ♂, Myanmar, 16 km E Putao, 500 m a. s. l., 28 - 30. IV. 1998, leg. Murzin and Sinjaev, genit. prep. 16230 (MWM / ZSM); 1 ♂, Myanmar, 21 km E Putao, Nan Sa Bon Village, 550 m a. s. l., 1 - 5. V. 1998, leg. Murzin and Sinjaev, genit. prep. 16150 (MWM / ZSM); 1 ♂, Myanmar, 25 km E Putao, env. Nan Sa Bon Village, 800 m a. s. l., 6 - 9. V. 1998, leg. Murzin and Sinjaev, genit. prep. 16231 (MWM / ZSM).	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
764B2FDB71225524B2DD7FF46F93F48F.taxon	diagnosis	Diagnosis. The new species is very similar to S. chalcites (Figs 9, 12) in appearance, but it can be distinguished from the latter by the following characters: middle of costal margin area of forewing is covered by dense black scales and the medial patch of forewing is conspicuous. In S. chalcites, the forewing only bears small scattered black scales and the medial patch of forewing is blurry.	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
764B2FDB71225524B2DD7FF46F93F48F.taxon	description	Description. Adult (Figs 7, 8). Wingspan 35 - 39 mm in male. Head brown; labial palpus short, brown; male antennae brown, bipectinated at basal half then serrate. Thorax dark brown to black dorsally, mesothorax with conspicuous tuft of long black hairs anteriorly; tegula brown. Scales on legs dark brown to pale yellow. Forewing distinct elongate, ground colour dark brown mixed with numerous black scales, especially dense in middle part of costal margin area, outer margin area pale brown; a conspicuous, large, silky, rounded medial patch located at outside of cell, inner half bluish black, outer half reddish brown with an arched bright line embedded in middle; subterminal line narrow, black, with depressions in the region of vein R 4, and slightly concave between veins M 3 and CuP; two distinct black spots at middle of veins CuP and 1 A + 2 A; fringe brown. Hindwing ground colour greyish brown to dark brown, anal margin area is darker; fringe pale brown. Abdomen brown to dark brown, mixed with little black hairs dorsally and long black hairs at terminal area. Male genitalia (Figs 19, 20). Uncus short, with apex deeply bifid, strongly sclerotised. Gnathos finger-shaped or slightly widened near apex, blunt apically. Tegumen broad. Valva short and broad; sacculus swollen at base, without saccular process; costa slightly concave at middle; cucullus narrow and rounded. Juxta asymmetrical, horseshoe-shaped, central depression V-shaped; left process strongly sclerotised, long plate-shaped, bearing a distinct spur near base in some individuals; right process plate-shaped, strongly sclerotised apically. Vinculum narrow. Saccus not obvious. Phallus thick, conspicuously waved, slightly thinner near apex, strongly sclerotised and somewhat bifid terminally. Female genitalia. Unknown.	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
764B2FDB71225524B2DD7FF46F93F48F.taxon	distribution	Distribution. China (Xizang: Motuo), India (Meghalaya), Myanmar (Kachin) (Fig. 25).	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
764B2FDB71225524B2DD7FF46F93F48F.taxon	etymology	Etymology. The species is named Squamosa undulophallus after its distinctly wavy phallus.	en	Wu, Jun, Solovyev, Alexey V., Han, Hui-Lin (2022): Two new species of Squamosa Bethune-Baker, 1908 (Lepidoptera, Limacodidae) and first female record of S. chalcites Orhant, 2000 from southern Asia. ZooKeys 1090: 149-159, DOI: http://dx.doi.org/10.3897/zookeys.1090.75823, URL: http://dx.doi.org/10.3897/zookeys.1090.75823
