taxonID	type	description	language	source
4A4D4F09FFBDFFB3F0A5F494FA45F958.taxon	description	Figures 21, 22, 23, 80 E, 80 J, 81 A, 82 A, 84 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFBDFFB3F0A5F494FA45F958.taxon	diagnosis	Diagnosis. Regular and longitudinal striae on head dorsum diverge towards posterior margin; rugose striae on mesosoma and petiole; dorsal margin of petiole discontinuous; ventral surface of node transversely striate; anterior region of node higher than posterior region; profemur posterior surface smooth; striae on protibia extensor surface weakly marked; striation on tergum of first gastral segment restricted to base; branched setae of two types: 1) thin setae, main axis branching off 2 – 4 times, short branches of equal size; 2) thick setae, main axis flattened in its final half, with several branches.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFBDFFB3F0A5F494FA45F958.taxon	description	Redescription. WORKER (n = 16) (Fig. 21 A – C): HL 1 (0.88 – 1.08); HW 0.94 (0.81 – 1.02); ML 0.56 (0.56 – 0.66); SL 0.64 (0.58 – 0.76); MOD 0.26 (0.23 – 0.26); PNW 0.64 (0.55 – 0.72); WL 1.20 (1.10 – 1.39); PSL 0.18 (0.16 – 0.32); PL 0.57 (0.51 – 0.60); PW 0.26 (0.20 – 0.26); PPL 0.33 (0.28 – 0.34); PPW 0.35 (0.30 – 0.37); GL 1 (0.94 – 1.20); TL 4.66 (4.27 – 5.23); CI 94 (90.72 – 95.91); SI 68.08 (68.08 – 74.74); OI 27.66 (25 – 28.40). Small to midsized. Shiny integument. Mesosoma and petiole dark brown; head and postpetiole light brown; yellowish or brownish appendices. Branched setae, midsized to short, suberect to subdecumbent; branched setae of two types: 1) thin setae, main axis branching off 2 – 4 times, short branches of equal size (Fig. 80 E); 2) thick setae, main axis flattened in its final half, with several branches, mainly in the postpetiole and gaster (Fig. 80 J). Head subquadrate; posterior margin straight. Mandible masticatory margin with 5 teeth. Anterior margin of clypeus slightly convex medially, with a pair of small teeth laterally; median area of clypeus with 8 – 9 regular to irregular, longitudinal and thick striae converging to a point on the anterior margin, interspaces smooth or with thinner striation. Frontal triangle with 1 sagittal stria. Short scape, not reaching head posterior margin; apical antennomere as long as previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye reniform, midsized, larger diameter with 12 ommatidia. Regular and longitudinal striae on head dorsum, divergent towards posterior margin; interval between thicker striae filled with thinner longitudinal and few transverse striae. Head lateral region with thin, regular and adjacent striae converging to eye margin; laterodorsal region rugose; gena striate, 2 – 3 regular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma covered with rugose striae of uniform thickness, interspaces smooth. Promesonotal junction and metanotal groove indistinct. Transverse carina indistinct to inconspicuous. Dorsal margin of mesosoma continuous, convex. Propodeal lobe bidentate, dorsal tooth slightly longer and acute than the shorter and blunt ventral tooth; dorsal tooth length is slightly longer than 1 / 2 of propodeal spine length. Propodeal spine midsized, straight (LV), divergent (DV), sculptured on base. Transverse striae of uniform thickness on procoxa; irregular and transverse striae on C 2 and C 3. Profemur smooth. Protibia extensor surface entirely striate. Dorsal margin of petiole discontinuous. Subtriangular projection on mesoventral surface of petiole. Node with transverse striae on ventral and anterior surfaces, interspaces distinguishable; rugose striae on lateral and dorsal surfaces, interspaces smooth. Postpetiole and subpostpetiolar process with regular, longitudinal, weakly marked striae or smooth; subpostpetiolar process weak, convex. First gastral segment striation similar to postpetiole striae; striae on tergum shorter than 1 / 3 of postpetiole length; sternite striation restricted to laterobasal region. QUEEN (n = 5) (Fig. 22 A – C): HL 1 (0.96 – 1.08); HW 0.96 (0.93 – 1.01); ML 0.58 (0.58 – 0.66); SL 0.64 (0.64 – 0.70); MOD 0.26 (0.26 – 0.28); PNW 0.80 (0.76 – 0.88); WL 1.38 (1.32 – 1.48); PSL 0.24 (0.20 – 0.32); PL 0.58 (0.58 – 0.66); PW 0.28 (0.26 – 0.30); PPL 0.32 (0.32 – 0.38); PPW 0.40 (0.36 – 0.41); GL 1.24 (1.24 – 1.38); TL 5.10 (5.10 – 5.56); CI 96 (92.45 – 96.87); SI 66.66 (66.66 – 71.42); OI 27.08 (26 – 28.57). Medium to large-sized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 13 – 14 ommatidia. Scutum longitudinally striate, regular to irregular striae going from an anterior central point towards transcutal suture, interspaces distinguishable. Longitudinal striation mostly regular striae on anepisternum and katepisternum, interspaces distinguishable. Longitudinal striation mostly irregular on axilla, interspaces distinguishable. Scutoscutellar sulcus inconspicuous. Scutellum with same sculpture of scutum. Transverse striae on propodeum (DV). Lateral surface of mesosoma with vermicular striae directed to propodeal spine. Wings as in Fig. 81 A. MALE (first description) (n = 2) (Fig. 23 A – C): HL (0.73 – 0.74); HW (0.64 – 0.65); ML (0.40 – 0.42); SL (0.22 – 0.24); MOD (0.30); PNW (0.74 – 0.78); WL (1.24); PL (0.54 – 0.56); PW (0.22 – 0.23); PPL (0.30 – 0.31); PPW (0.30 – 0.32); GL (1.04 – 1.22); TL (4.26 – 4.48); CI (86.48 – 89.04); SI (34.37 – 36.92); OI (46.15 – 46.87). Brownish body, darker gaster, lighter appendices. Mandible with 4 teeth. Anepisternum and katepisternum mostly with longitudinal striae. Scutellum with irregular to slightly vermicular striae, thicker than scutum striae, interspaces distinguishable. Propodeum with irregular striae assuming multiple directions, interspaces distinguishable. Propodeal lobe rounded. Petiole dorsum partly smooth and partly covered with irregular striae assuming multiple directions, interspaces distinguishable. Postpetiole mostly smooth. Gaster smooth. Wings as in Fig. 82 A.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFBDFFB3F0A5F494FA45F958.taxon	etymology	Etymology. This species was named in honor of Luigi Balzan (1865 – 1893), an Italian naturalist and professor at National Institute, Asuncion, Paraguay, who collected the type specimen.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFBDFFB3F0A5F494FA45F958.taxon	discussion	Comments. Emery (1894) only described the worker of Tetramorium balzani. While visiting MSNG, we found a worker and queen pinned together and labeled “ typus ”. The original description gives a worker size range, but there were no other workers in the collection, and Emery may have mistakenly included the queen in the measurements or the other worker (s) from this series is missing. We designate the worker as lectotype. Borgmeier’s (1937) original description of Lundella speciosa makes reference to the similarity between L. speciosa (RJ, Brazil) and L. balzani (Argentina, Paraguay). Lundella speciosa was separated from L. balzani by the larger body (vs. smaller), bicolored integument — blackish mesosoma, light-brown appendices (vs. brownish), and longer propodeal spine (vs. shorter). Brown (1953) synonymized Lundella under Hylomyrma, and Kempf (1960) considered Hylomyrma balzani as a senior synonym of H. speciosa based on the study of more than 100 specimens (workers and queens) from Argentina, Brazil (PR, RJ, RS, and SC), and Paraguay. According to Kempf (1960), the range of morphological variation observed in H. balzani specimens from the south and southeast regions of Brazil included body measures, propodeal spine length, and the bicolored integument of H. speciosa. The lectotype of H. balzani is concolorous, the propodeal spine is short, there is microsculpture associated with the macrosculpture (rugose striae) laterally on the mesosoma, and the postpetiolar dorsum and base of tergum IV have longitudinal striae. The lectotype of L. speciosa is bicolored, the propodeal spines are longer, there is no microsculpture, and both the postpetiole and tergite IV are smooth. The material gathered for this research, especially specimens from Brazil (BA, CE, RJ, and SP), shows that some characteristics used by Emery and Borgmeier to define these two species can be observed in a single specimen. Thus, we corroborate the synonym proposed by Kempf (1960). The rugose striae characteristic of many H. balzani are weaker in some specimens from different localities in Brazil (Itambé Leoncio, Ibirapitanga, Lauro de Freitas, Lomanto Júnior, Maracás, Mata de São João, Milagres, São João do Paraíso, and Gurupá Mirim, BA; Sooretama, ES; Recife, PE; Itatiaia, RJ; Xaxim, SC; and Areia Branca, and São Cristóvão, SE). Moreover, the sculpture of the propodeal dorsum of these specimens is subopaque and slightly marked, and the sculpture of the petiolar node dorsum can show regular striae forming semi-ellipses. Consequently, these specimens partly resemble H. reitteri, which presents regular to irregular striae that are longitudinal on the mesosomal dorsum and that are transverse on the propodeal dorsum. Hylomyrma balzani and H. reitteri have broad distributions (Figs. 84, 85), occurring sympatrically in Paraguay (Alto Paraná department) and in many localities of Brazil. Both species have been recorded in the same fauna inventory, but in different samples (Santa Teresa, ES; Lavras, and Viçosa, MG; Estrada Velha Guaricana, Irati, Laranjeiras, Morretes, Rondon, and Tuneiras do Oeste, PR; Rio de Janeiro, and Teresópolis, RJ; Nova Petrópolis, and 5 km N of Progresso, RS; Nova Teutônia, and Seara, SC; Botucatu, and Jundiaí, SP, Brazil). Both species were also found in the same sample in Santa Teresa, ES (Winkler 24), Chapecó (Coll. Kempf n ° 2655, Coll. Borgm. Nr. 6501) and Nova Teutônia (13628, 6869), SC, and Frederico Westphalen, RS (5546). Hylomyrma balzani can be distinguished from H. reitteri (characteristic in parentheses) by the rugose striae on the mesosoma (vs. regular to irregular striae), the rugose striae on the petiole (vs. irregular striae), and the ventral surface of petiole with transverse striae (vs. smooth surface). The winged queens of both species are very similar, differing only in the laterodorsal sculpture of the propodeum. In Hylomyrma balzani this region presents rugose striae, whereas H. reitteri has regular to irregular striae in this region. Specimens of H. lopesi have been identified as H. balzani, especially due to the similarity of the sculpture on gena, petiole and postpetiole, and the similar shape of the petiole and postpetiole. But both species can be distinguished by the rugose striae on the mesosoma of H. balzani and the regular striae on the mesosoma of H. lopesi. The distribution of these species cover a broad area in Brazil, both co-occurring in Botucatu (SP) and Milagres (BA). Hylomyrma balzani is more restricted to eastern portions of Brazil, from northeast to southern regions (Fig. 84), whereas H. lopesi is distributed mainly in the center of Brazil from northern (state of Pará) to southern (state of São Paulo) regions (Fig. 89). Hylomyrma balzani is similar to H. primavesi, a species known only from higher elevations in Serra do Cipó, MG, Brazil (Fig. 88). Hylomyrma balzani can be distinguished from H. primavesi by the smaller body, TL 4.27 – 5.23 mm, WL 1.10 – 1.39 mm (vs. larger, TL 5.32 – 5.78 mm, WL 1.34 – 1.48 mm); and the indistinct promesonotal junction and metanotal groove (vs. both distinct). There is considerable variation among H. balzani specimens, mainly because of the presence of intercastes. Out of the 11 Hylomyrma species with intercastes, H. balzani, H. lopesi, and H. reitteri present the larger phenotypic plasticity associated with head and mesosoma modifications. Intercastes with one inconspicuous central ocellus usually have two darker regions on the head similar to the ocelli location of winged queens. Intercastes with a more conspicuous central ocellus may also have these two darker regions on the head; they also have the metanotal groove distinguished by a slight to well-marked depression. In intercastes with three ocelli, gradual changes in the mesosoma are observed. The 3 - ocelli intercaste from Vargito, BA, Brazil, bears also a slight differentiation of the mesonotum in scutellum and prescutellum, and of the metanotum. The specimens from Maranguape, CE, Brazil, in addition to the characteristics described above, present the promesonotal suture (specimens W 49, Mg 50 W), and also the insertion wing region in the mesosoma, as observed in winged queens (specimen P 1).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFBDFFB3F0A5F494FA45F958.taxon	distribution	Distribution. Hylomyrma balzani is known from northern Argentina (Misiones province) to northeastern Brazil, also occurring in Paraguay (Fig. 84). Natural history. There is no information on the collection method or in which phytophysiognomy most known specimens were sampled. Specimens from Brazil inhabit Atlantic forest areas and forested enclaves in the Central-West region (in Serra da Bodoquena, MS). Most specimens were collected with winkler extractors, and a few were sampled with pitfall traps, honey baits, and Berlese-Tüllgren funnels. Apart from the presence of intercastes, and possibly ergatoids (a few intercastes have more queen-like traits), there is no further information available regarding the biology of H. balzani. Additional material examined (278 workers, 39 queens, 2 males, 30 intercastes): ARG [ARGENTINA]: Misiones: Est. Exp. Loreto, 10. v. 1932, Dr. A. A. Oglobin [leg.], # 1957 (1 I 1 Q) [MZSP]; same data, Sammlung [collection] Dr. F. Santschi Kairouan (1 I) [NHMB]; same except 30. vi. 1932, Sammlung [collection] Dr. F. Santschi Kairouan, # 1973 (1 I) [NHMB]. BR [BRAZIL]: AL [Alagoas]: Quebrangulo, 31. viii. 99, 0919 S, 3628 W, Santos J. R. M. dos [leg.] (3 W) [MZSP]; same data, 1283 (3 W 1 Q) (MZSP 67322) [MZSP]; Res. Pedra Talhada, 05. viii. 99, Col. D. S. Santos, 637 (3 W 1 Q) [CEPLAC]; Murici, ESEC Murici, ICMBio, 09 ° 14 ′ 57.04 ″ S, 35 ° 47 ′ 28.88 ″ W, 23. ix. 2014, Lorenzo E. P. col. (1 W) [UFGD]; BA [Bahia]: Maracás, Faz. Mª Inácia, Mata Cipó, 24 – 29. xi. 1990, Brandão, Diniz & Oliveira col., peneira + troncos caídos (13 W 3 I) [MZSP]; same data (2 W) [UTLP]; same except peneira, # 1 (6 W) [MZSP]; isca - mel, dia - solo, 20 (1 W) [MZSP]; berlese (1 W) [MZSP]; emergido de formigueiro artificial (1 M) [MZSP]; Lençóis, 25. iii. 2001, Santos J. R. M., 34 (3 W) [CEPLAC]; Cachoeira, 11.12. [19] 93, J. H. C. Delabie [leg.], 4733 (5 W) [CEPLAC]; Mata São João, Reserva Sapiranga, 12 ° 33 ′ 29.3 ″ S, 33 ° 02 ′ 35.2 ″ W, 21 – 28. vii. 2001, Silva R. R., Brandão C. R. F. cols., Winkler 30 (3 W) [MZSP]; same except Winkler 22 (1 W covered with gold) [MZSP]; Winkler 28 (3 W) [MZSP]; Winkler 4 (3 W) [MZSP]; Winkler 6 (1 W) [MZSP]; Winkler 17 (1 W) [MZSP]; Winkler 5 (1 Q) [MZSP]; Itambé, Área A, 25.07. [20] 03, 145976 S, 0404153 W, Santos J. R. M. [leg.], Mata-W (3 W) [CEPLAC]; same data (1 Q) (MZSP 67324) [MZSP]; same data (1 W 1 Q) [DZUP]; same except Área B, 151734 S, 0402740 W, 19.07. [20] 03 (1 W 1 Q) [CEPLAC]; Maraú-Tremenbé, 07.06.97, Santos J. R. M dos [leg.], Mata-W-A 4 (2 W) [CEPLAC]; Itororó, 11.08. [20] 00, 14 ° 58 ′ 28 ″ S, 40 ° 03 ′ 01 ″ W, Santos J. R. M. dos [leg.], 637 (1 W) [CEPLAC]; Vargito, 22.03. [19] 99, 1524 S, 03933 W, Santos J. R. M. dos [leg.], 1283 (1 W 1 I) [CEPLAC]; Itambé Leoncio, 14.35.67 S, 040.20.23 W, 06.08. [20] 03, Santos J. R. M. dos [leg.], Mta (2 W) [CEPLAC]; Ibirapitanga, A 2, 141139 S, 0392523 W, 22.04. [19] 97, Santos J. R. M. dos [leg.], Mata — W, 1283 (4 W) [CEPLAC]; same data (1 M) (MZSP 67323) [MZSP]; Lomanto Júnior, 14.48.57 S, 039. S 29. S 02 W, 17.11. [19] 97, Santos J. R. M. dos [leg.], Mata-A 6, 1283 (2 W 1 Q) [CEPLAC]; São João do Paraíso, 153854 S, 392630 W, 17.09. [19] 99, Santos J. R. M. dos [leg.], 637 (2 W) [CEPLAC]; Gurupá Mirim, Fazenda Esperança, 05. ii. 2009, Col. Marques T. & estagiários, Floresta Atlântica, Parcela, Arm. 7, Estrato Sol. (1 W) [CEPLAC]; Lauro de Freitas, Busca Vida, 12 ° 51 ′ 51 ″ S, 38 ° 16 ′ 11 ″ W, vi – x. 2010, Travassos, M. L. O. col., # 5641, I 1448 (1 W) [CEPLAC]; Milagres, área 3, S 12 ° 54.411 ′, W 39 ° 50.863 ′, 29. vii. 2010, Ulysséa, M. A., Medina, A. M. e Campos, E. M. leg., Caatinga Arbórea, serapilheira, winkler, W 33 (1 W) [MZSP]; same except 17. i. 2011, W 137 (6 W 1 Q) [MZSP]; 17. i. 2011, W 127 (1 Q 2 W) [MZSP]; área 1, S 12 ° 54.542 ′, W 39 ° 51.279 ′, 12 – 16. vii. 2010, T 4 F, 2 H 1 (1 W) [MZSP]; área 1, S 12 ° 54.542 ′, W 39 ° 51.279 ′, 13. vii. 2010, W 6 (1 W) [MZSP]; área 1, S 12 ° 54.542 ′, W 39 ° 51.279 ′, 10. i. 2011, W 108 (1 W) [MZSP]; área 1, S 12 ° 54.542 ′, W 39 ° 51.279 ′, 23. x. 2010, W 62 (1 Q) [MZSP]; área 1, S 12 ° 54.542 ′, W 39 ° 51.279 ′, W 52 (1 Q) [MZSP]; área 2, S 12 ° 54.294 ′, W 39 ° 52.083 ′, 10. i. 2011, W 121 (1 W covered with gold) [MZSP]; CE [Ceará]: Maranguape, + 875 m, 03 ° 53 ′ S, 38 ° 43 ′ W, viii. 2003, Y. Quinet [leg.], P 1 (1 W 1 I) [MZSP]; same except W 49 (1 W 2 I) [MZSP]; W 50 (2 W) [MZSP]; Trilha Gavião, 3 ° 54 ′ S 38 ° 43 ′ O, + 890 m, 15. vii. 2004, N. Hites col., Winkler, Mata úmida de altitude / Brejo, Mg 50 W (1 I) [MZSP]; Guaramiranga, Pq. Tr., + 900 m, 04 ° 16 ′ S, 38 ° 56 ′ W, 28. ii. 2002, Y. Quinet [leg.], W 44 (1 W) [MZSP]; same except W 46 (1 W) [MZSP]; 25. ii. 2002, W 65 (2 W) [MZSP]; 2. iii. 2002, P 27 (1 W) [MZSP]; Pacoti, 415 S, 3855 W, Mata do Hotel Remanso, alt 700 m, 2001, Nathalie Hites [leg.], Mata secundária, TP- 3 (D) 250 (1 W) [CEPLAC]; ES [Espírito Santo]: Santa Teresa, 900 m, 23. fev. 67, W. L. Brown col., mt. for. (1 W) [MCZC]; Estação Biológica Santa Lucia, 19 ° 58 ′ 09 ″ S, 40 ° 32 ′ 15 ″ W, 20 – 24. i. 2002, Schoereder J. H. & Ribas C. R. cols., Winkler 37 (1 I) [MZSP]; same except Winkler 18 (2 W) [MZSP]; Winkler 30 (1 W 1 Q) [MZSP]; Winkler 24 (1 W) [MZSP]; Winkler 1 (1 W 1 Q) [MZSP]; Winkler 33 (1 W) [MZSP]; Winkler 49 (1 W 1 I) [MZSP]; Sooretama, REBIO Sooretama, 19 ° 04 ′ 21 ″ S, 39 ° 56 ′ 57 ″ W, 14 – 15. v. 2002, Schoereder, J. H. & Ribas, C. R. cols., Winkler 39 (3 I) [MZSP]; same except Winkler 3 (2 W) [MZSP]; Winkler 38 (1 I) [MZSP]; Winkler 37 (1 W) [MZSP]; Winkler 21 (1 Q) [MZSP]; GO [Goiás]: Ouro Verde, Faz. Boa Vista, 16 ° 17 ′ 54.5 ″ S, 49 ° 12 ′ 42.6 ″ W, 20 – 24. i. 2005, Silva R. R. col., Winkler 38 (1 W) [MZSP]; Cavalcante, Serra da Contenda, 13 ° 42 ′ 03.6 ″ S, 47 ° 27 ′ 51.9 ″ W, 16. x. 2004, Silva R. R. & Dietz B. H. cols., Mata Ciliar, Winkler 12 (1 W) [MZSP]; [MG] Minas Gerais: Parque Estadual do Rio Doce, Trilha da Lagoa Preta, Elev. 280 m, 19 ° 47 ′ 49 ″ S, 42 ° 34 ′ 38 ″ W, 28. viii. 2005, TEAM exped. Col., Floresta Atlântica Estacional Semidecidual, ITLP 3.10 Mini-winkler (1 W) [MZSP]; same except TLP 2.1 (1 W) [MZSP]; TLP 2.3 (1 W) [MZSP]; TLP 2.1 (1 W) [MZSP]; TLP 1.2 (1 W) [MZSP]; 23 – 24. viii. 2005, Trilha da Garapa Torta, TGT 1.1 (1 W) [MZSP]; 30. xi. 05, LP 2 - 4 (1 W) [MZSP]; 30. xi. 05, LP 2 - 3 (1 W) [MZSP]; Marlieria, Parque Estadual do Rio Doce, Trilha da Garapa Torta, 280 m, - 19.796944, - 42.577222, 24. viii. 2005, TEAM exped. Col., Floresta Estacional Semidecidual, GT 2 - 7 (1 W) [MZSP]; same except Trilha da Lagoa Preta 30. xi. 2005, LP 3 - 2 (1 W) [MZSP]; Lavras, Fragmento, 06 – 12.2002, Santos M. S. & Dias N. S. [leg.] (2 W 1 Q) [CEPLAC]; Serra do Cipó, i / 2013, PEMII 1, Coletor M. C. Anjos, Winkler (1 W) [UFMG]; Serra do Cipó, próx. Cachoeira da Capivara, entrada p / estátua “ Juquinha ″, 1354 m, - 19.253306, - 43.552531, 13. v. 2016, J. Chaul & S. Epifânio [leg.], epigaeic, Jcwinkler # 019 (1 Q 10 W) [MZSP]; same except 1351 m, 19.252982, - 43.551787, epi + hypogaeic, Jcwinkler # 023 (2 W) [MZSP]; Conceição do Mato Dentro, Serra da Serpentina, 18 – 28. iii. 09, Silva R. R. leg. (1 W) [DZUP]; Viçosa, UFV, Mata da Biologia, 706 m, 20 ° 45 ′ 26.67 ″ S, 42 ° 51 ′ 39.07 ″ W, iii. 2013, J. Chaul col., winkler, UFV LABECOL n ° 000137 (1 Q) [MZSP]; same except 20 ° 45 ′ 26 ″ S, 42 ° 51 ′ 49 ″ W, 17. viii. 2014, winkler of rotten log, UFV LABECOL n ° 000134 (1 W) [MZSP]; Bairro Cristais, 20 ° 46 ′ S, 42 ° 50 ′ W, 20. iv. 2013, Jesus & Chaul [leg.], winkler, JC 0007, UFV LABECOL n ° 000136 (1 Q) [MZSP]; MT [Mato Grosso do Sul]: Ivinhema, Someco, 2005, S. Almeida Lare, Stela, UEMS (1 W) [CEPLAC]; MS [Mato Grosso do Sul]: Bonito, PARNA Serra da Bodoquena, Faz. Harmonia, 21 ° 17 ′ 09.8 ″ S, 56 ° 41 ′ 45.5 ″ W, 08 – 17. x. 2006, Silvestre R. et al. col., 4 ª exp., Winkler A- 05, Hym- 00461 - F (1 I) [UFGD]; [Ponta Porã], Mata Ciliar do Rio Dourados, Fazenda Paineirinha, 22 ° 07 ’ 20.1 ” S, 55 ° 38 ’ 25.5 ” W, 20 / 04 / [20] 07, Silvestre et al. col., A- 11 (1 I) [MZSP]; same except A- 02 (1 W) [UFGD]; A- 18 (1 W) [MZSP]; A- 05 (1 W) [UFGD]; PB [Paraíba]: Conde, APA Tambaba, 07 ° 22 ′ 05.69 ″ S, 34 ° 48 ′ 39.05 ″ W, 24. viii. 2014, Brisa Lunar P. Tavares col. (1 W 1 Q) [UFGD]; PE [Pernambuco]: Recife, Horto Dois Irmãos, 08 ° 00 ′ 32 ″ S, 34 ° 56 ′ 40 ″ W, 15 – 24. vii. 2002, Silva R. R. & Eberhardt F. cols., Winkler 5 (3 W) [MZSP]; same except Winkler 13 (1 W); Winkler 14 (1 I); Winkler 9 (1 W); Winkler 1 (2 W); Winkler 17 (1 Q) [MZSP]; PR [Paraná]: Rondon, iv. 1965, F. Plaumann [leg.], 4772 (1 W) [MZSP]; Laranjeiras, iv. 1965, F. Plaumann [leg.], 4551 (1 W) [MZSP]; Foz do Iguaçu, Pq. Nac. Iguaçu, 6 – 8. i. 1997, Pinto-da-Rocha e Casari [leg.] (1 W) [MZSP]; Pinhão, UHE Segredo, Ribeirão Estreito, 24. i. 1992, R. P. da Rocha [leg]., berlese (2 W) [MZSP]; same data (2 W) [JTLC]; Rib. Estreito, 20. iii. 1992, Rocha & Barreto, Berlese (2 W) [MZSP]; Porto Vitória, 700 m, 26 ° 10 ′ – 51 ° 12 ′, x. 1959, F. Plaumann [leg.], 3199 (2 W) [MZSP]; same data (1 W 1 Q) [MCZC]; Morretes, Parque Estadual do pau-Ôco, 25 ° 34 ′ 33.5 ″ S, 48 ° 53 ′ 19.5 ″ W, 6 – 11. v. 2002, Silva R. R. & Dietz B. H. cols., Winkler 51 (1 W 1 I) [MZSP]; same except Winkler 22 (1 W) [MZSP]; Winkler 29 (2 W) [MZSP]; Estrada Velha Guaricana, ca Cachoeira das Mulas, - 25.7303 ° 49.0911 °, 10. iv. 2017, R. López & J. Lattke [leg.], 3812 (1 W) [DZUP]; same except 3833 (1 W) [DZUP]; 3811 (1 W) [DZUP]; 3818 (1 W) [MZSP]; Tuneiras do Oeste, Reserva Biológica de Perobas, 23 ° 50 ′ 5.64 ″ S, 52 ° 45 ′ 37.52 ″ W, 18. ix. 2015, Busanello, D. & Caron, E. cols. (4 W) [DZUP]; REBIO das Perobas, 540 m, 23 ° 50 ′ S, 52 ° 45 ′ W, 18. ix. 2015, E. Caron col., winkler (2 W) [DZUP]; Palotina, Parque Estadual São Camilo, 24 ° 19.276 ′ S, 53 ° 55.247 ′ W, 23. ix. 2015, Busanello, D. & Caron, E. cols. (3 W) [DZUP]; Irati, Guamirim, Fz. Arroio Grande, Talhão 15 - AII, 25 ° 35 ′ 36.11 ″ S, 50 ° 49 ′ 12.06 ″ W, 09. x. 2014, Marques, C. G. P., Falbot, L. Col., Serapilheira, Funil de Berlese (1 W) [DZUP]; RJ [Rio de Janeiro]: Nova Iguaçu, ReBio Tinguá, BRE, 2. ii. 2002, Ext. Winkler, A 07 (1 W) [MZSP]; same except A 03 (1 W) [MZSP]; A 01 (1 W) [MZSP]; A 08 (1 W) [MZSP]; 4. ii. 2002, A 42 (1 W) [MZSP]; Teresópolis, P. N. Serra dos Órgãos, 23 – 28. xi. 1999, Dietz, Silva & Rocha cols., Winkler (1 W) [MZSP]; Itatiaya [Itatiaia], Borgmeier [leg.], gesiebt [shifted], 10. i. 56 (3 W) [MZSP]; P. N. Itatiaia, 846 m, 22 ° 27 ′ 08.56 ″ S, 44 ° 36. ′ 55.74 ′ W, 20. i. 2015, Lasmar et al. col. (5 W) [DZUP]; same except 1136 m, 22 ° 25 ′ 51.62 ″ S, 44 ° 36. ′ 57.36 ′ W (2 W) [DZUP]; Rio de Janeiro, Floresta da Tijuca, D. Federal, 25. i. 1960, C. A. C Seabra [leg.], Coleção Campos Seabra (1 W) [MZSP]; RS [Rio Grande do Sul]: 5 km north of Progresso, Linha Araçá, near Rio Fão, 300 – 400 m, 52 ° 20 ′ W, 29 ° 10 ′ S, 04. v. 1999, Leg. J. Bihn, Secondary Forest, soil sifting, Winkler 1 qm (2 W, one covered with gold) (MZSP 67434) [MZSP]; same except 29. iv. 1999 (1 W 1 Q) [MZSP]; Frederico Westphalen, UF Santa Maria, 04. xi. 2008, Granzotto F. [leg.], # 5546 (3 W 2 Q) [CEPLAC]; N. Petrópolis, Lat 29 ° 20, Long 51 ° 12, xi. 1959, F. Plaumann [leg.], 3222 (2 W 1 Q) [MZSP]; SC [Santa Catarina]: Chapecó, v. 1957, Plaumann col., Col. Kempf n ° 2655 (5 W 3 I) [MZSP]; same except viii. 1960 (3 W) [MZSP]; xii. 1957, Coll. Borgm. Nr. 6501 (3 W 1 Q) [MZSP]; Concórdia, ix. 1959, F. Plaumann [leg.], 8297 (2 W) [MZSP]; same except vii. 1958, 2732 (6 W) [MZSP]; Nueva [Nova] Teutonia, Fritz Plaumann [leg.] (3 W 1 Q) [IFML]; same except vi. 1960, 8173 (1 W) [MZSP]; vi. 1961, 6868 (5 W 1 Q) [MZSP]; vi. 1961, 6869 (1 Q) [MZSP]; 300 – 500 m, 27 ° 11 ′ B, 52 ° 23 ′ L, iv. 1976, 13627 (7 W 1 Q) [MZSP]; iv. 1976, 13628 (1 W) [MZSP]; 1961 (1 W) [MHNG]; vi. 1959 (2 W) [MZSP]; ix. 1959, 3491 (6 W) [MZSP]; vi. 1957 (3 W 2 Q) [MZSP]; viii. 1957 (1 W) [MZSP]; vii. 1957 (1 Q) [MCZC]; x. 1954 (1 W) [MZSP]; xi. 1954 (1 Q) [MZSP]; Seara, 24 ° 07 ′ S, 52 ° 18 ′ W, vi – vii. 1999, Rogério R. Silva, col., solo, Winkler (1 W) [MZSP]; same except viii. 1958 (2 W) [MZSP]; viii. 1958, 2740 (2 W) [MZSP]; vii. 1958, 2709 (2 W) [MZSP]; v – xii. 1998, Transecto I, Winkler (1 Q) [MZSP]; Linha Facão [near S. Carlos], v. 1957, F. Plaumann (2 W 1 Q) [USNM]; same data (1 Q) [MZSP]; Xaxim, cat., Plaumann [leg.] (1 W) [MZSP]; Xanxerê, xii. 1957, Plaumann [leg.] (1 W) [MZSP]; SE [Sergipe]: Areia Branca, E. E. da Serra de Itabaiana, 10 ° 45 ′ 54 ″ S, 37 ° 19 ′ 57.4 ″ W, 19 – 25. v. 2003, Silva R. R., Dietz B. H. & Ferreira L. S. cols., Winkler 25 (2 W) [MZSP]; S. [São] Cristóvão, 10 ° 54.3 ′ S, 37 ° 11.7 ′ W, 31. i. 2014, Almeida, R. P. S. Leg., IFS, Área APP, Parcela 9, 36 (2 W) [DZUP]; SP [São Paulo]: Cajuru, Faz. Santa Carlota, 21 ° 17 ′ S, 47 ° 18 ′ W, 24 / 04 / 1993, R. Silvestre [leg.], isca mel, dia solo, VI- 13 (1 W) [MZSP]; same except 11 / 6 / 1993, isca sardinha, dia solo (4 W 1 I) [MZSP]; Botucatu, 11 - 03 - 1988, Forti L. C. & Rinaldi I. M. P. col., Armadilha de solo, Forti # 08 (1 W) [MZSP]; same except 13 - 10 - 1987, Forti # 18 (1 W) [MZSP]; 13 - 10 - 1987, Forti # 19 (1 W) [MZSP]; 07 - 07 - 1987, Forti # 33 (1 W) [MZSP]; 07 - 10 - 1986, Forti # 45 (1 W) [MZSP]; 03 - 11 - 1986, Forti # 49 (1 W) [MZSP]; 08 - 02 - 1988, Forti # 09 (1 W) [MZSP]; 10 - 09 - 1987, Forti # 20 (1 W) [MZSP]; 04 - 04 - 1988, Forti # 29 (1 Q) [MZSP]; Jundiaí, Serra do Japi, 23 ° 13 ’ S, 46 ° 58 ’ W, iii. 2011, T. Postali col., pitfall, # 1579 (1 W) [MZSP]; TO [Tocantins]: Palmeirante, 07 ° 52 ′ 25.3 ″ S, 47 ° 57 ′ 07.4 ″ W, 10 - 15. xii. 2001, Albuquerque & Silva cols., Mata Ciliar / Cerradão, Transecto I, Winkler 06 (1 W) [MZSP]. PARAGUAY: Balzan [leg.] (1 Q) [MSNG]. Alto Parana: [San Alberto], Puerto Sta. Teresa, 3. xi. 79, C. Dlouhy [leg.], 47 (1 I) [MHNG]; Arroyo Itaba Guazu, 4. xi. 79, J-L. Perret [leg.] (1 W) [MHNG]; Canendeyú: Ype Jhu, 28. x. 82, V. Mahnert [leg.], 39 (2 W) [MHNG]; R. Carapa-R. Alto Parana, 2. xi. 79, C. Vaucher [leg.], 46 (1 I) [MHNG]; same except F. Baud et al. of MHN-Geneva, 46 (1 W 1 I) [MHNG]; Concepción: [Azote’y], Arroyo Azotey, 10. x. 79, J-L. Perret (1 W) [MHNG]; [Itapúa]: San Benito, 29. x. 82, V. Mahnert [leg.], 82 / 17 (3 W) [MHNG]; nr. Caicisa [Compañía Aceitera de Itapúa Comercial y Industrial S. A.], Arroyo Tembey, 6. xi. 82, F. Baud et al. of MHN-Geneva, forest, leaf litt. & wood, wickler apparat (1 Q 4 W 2 I) [MHNG].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFB3FF89F0A5F30DFC31FB6E.taxon	description	Figures 24, 25, 26, 80 A, 80 F, 82 F, 84 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFB3FF89F0A5F30DFC31FB6E.taxon	diagnosis	Diagnosis. Head dorsum with regular and longitudinal striae, mesial striation in part directed to posterior margin and in part anteriorly divergent and posteriorly convergent, interspaces distinguishable; mesosoma covered with regular to irregular striae of variable thickness, interspaces indistinguishable; dorsal margin of petiole discontinuous, dorsum with same striation of mesosoma lateral; subtriangular projection on mesoventral surface of petiole; subpostpetiolar process weak, subtriangular; transverse striae on profemur posterior surface; protibia extensor surface striate; long striae on tergum of first gastral segment.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFB3FF89F0A5F30DFC31FB6E.taxon	description	Redescription. WORKER (n = 4) (Fig. 24 A – C): HL 0.92 (0.90 – 0.96); HW 0.86 (0.82 – 0.89); ML 0.54 (0.54 – 0.56); SL 0.62 (0.58 – 0.64); MOD 0.24 (0.20 – 0.24); PNW 0.63 (0.62 – 0.64); WL 1.22 (1.20 – 1.23); PSL 0.20 (0.20 – 0.27); PL 0.54 (0.54 – 0.55); PW 0.24 (0.23 – 0.25); PPL 0.32 (0.32 – 0.34); PPW 0.32 (0.32 – 0.33); GL 0.96 (0.96 – 1.06); TL 4.50 (4.50 – 4.69); CI 93.47 (89.36 – 92.70); SI 72.09 (65.16 – 76.19); OI 27.90 (22.47 – 27.90). Midsized. Integument subopaque, except for the shiny gaster. Brownish body, lighter appendices (yellowish). Thin and mostly unbranched setae (Fig. 80 A), long to midsized, suberect to subdecumbent; few branched setae with 2 – 3 or multiple short branches of equal size, especially on petiole (Fig. 80 F). Head subquadrate; posterior margin straight. Mandible masticatory margin with 5 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus striate, with regular and longitudinal striae, being 4 thicker striae, interspaces indistinguishable. Frontal triangle with 1 sagittal stria. Scape short, not reaching head posterior margin; apical antennomere shorter than the 3 previous antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye drop-shaped, small-sized, larger diameter with 10 ommatidia. Regular and longitudinal striae on head dorsum, mesial striation in part directed to posterior margin and in part anteriorly divergent and posteriorly convergent, interspaces between thicker striae in part smooth and int part filled with thinner striae. Head lateral and laterodorsal regions with regular and longitudinal striae converging to eye margin, interspaces indistinguishable; gena striate, irregular and semicircular striae circumscribe the torulus, not reaching eye margin, being 2 – 3 thicker striae, interspaces distinguishable. Interspaces between striae on head ventral surface indistinguishable. Mesosoma covered with regular to irregular striae, thinner striae (microsculpture) of variable thickness superimposed on thicker striae (macrosculpture), interspaces indistinguishable. Transverse striae on pronotum anterior region continuing on lateral surface; longitudinal striae on pronotum posterior region and mesonotum; thinner striae (microsculpture) superimposed on thicker striae (macrosculpture) on mesosoma lateral, striation in part continuing transversely on propodeum dorsum and in part directed to propodeal spine. Promesonotal junction and metanotal groove indistinct. Transverse carina well-marked. Dorsal margin of mesosoma continuous, convex. Propodeal lobe bidentate, acute dorsal tooth slightly longer than the blunt ventral tooth; dorsal tooth length slightly longer than 1 / 2 of propodeal spine length. Propodeal spine midsized, straight (LV), divergent (DV), sculptured. Procoxa striae of uniform thickness; regular and transverse striae on C 2 and C 3. Profemur covered with regular and transverse striae. Regular and longitudinal striae on protibia extensor surface. Dorsal margin of petiole discontinuous. Subtriangular projection on mesoventral surface of petiole. Convex node; regular and transverse striae on ventral and anterior surfaces, interspaces distinguishable; striae on anterior surface and dorsoanterior region continuing on lateral surface; irregular striae of variable thickness on dorsal surface, drop-shaped or semicircular, interspaces indistinguishable. Postpetiole and subpostpetiolar process with regular and longitudinal striae; subpostpetiolar process weak, subtriangular. First gastral segment striation similar to postpetiole striae; long striae on tergum, longer than postpetiole length; sternite striation covering the laterobasal region. QUEEN (n = 4) (Fig. 25 A – C): HL 0.88 (0.88 – 1); HW 0.84 (0.84 – 0.94); ML 0.58 (0.56 – 64); SL 0.63 (0.62 – 0.68); MOD 0.26 (0.24 – 0.27); PNW 0.70 (0.70 – 0.84); WL 1.30 (1.30 – 1.50); PSL 0.22 (0.22 – 0.31); PL 0.58 (0.56 – 0.62); PW 0.25 (0.25 – 0.28); PPL 0.36 (0.36 – 0.41); PPW 0.37 (0.37 – 0.41); GL 1.16 (1.16 – 1.30); TL 4.86 (4.86 – 5.43); CI 95.45 (92.39 – 95.45); SI 75 (72.94 – 75); OI 30.95 (27.95 – 30.95). Medium to large-sized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 12 ommatidia. Striae on scutum longitudinal, regular to irregular, going from an anterior central point towards transcutal suture, interspaces indistinguishable; thinner striae (microsculpture) between thicker striae (macrosculpture). Longitudinal striae of variable thickness on anepisternum and katepisternum, mostly regular on anepisternum and irregular on katepisternum, interspaces indistinguishable. Axilla and scutellum with same sculpture of scutum. Scutoscutellar sulcus inconspicuous. Transverse striae on propodeum (DV). Lateral surface of mesosoma with irregular striae directed to propodeal spine. Wings unknown. MALE (first description) (n = 1) (Fig. 26 A – C): HL 0.72; HW 0.64; ML 0.43; SL 0.22; MOD 0.28; PNW 0.66; WL 1.20; PL 0.54; PW 0.22; PPL 0.32; PPW 0.30; GL 1.12; TL 4.33; CI 88.88; SI 34.37; OI 43.75. Brownish body, lighter appendices (yellowish), darker gaster. Mandible with 5 – 6 teeth. Interspaces between head dorsum striation smooth. Scutum mostly smooth anteriorly and covered with regular to irregular and thinner striae posteriorly. Anepisternum and katepisternum mostly covered with longitudinal, regular and thinner striae, interspaces distinguishable. Scutellum with regular to slightly irregular striae, thicker than scutum striae, interspaces distinguishable. Propodeum with irregular striae assuming multiple directions, interspaces distinguishable. Propodeal lobe subquadrate. Petiole and postpetiole mostly covered with longitudinal, regular and thinner striae, interspaces distinguishable. Tergum of first gastral segment with same striation of postpetiole, striae restricted to basal region. Wings as in Fig. 82 F.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFB3FF89F0A5F30DFC31FB6E.taxon	etymology	Etymology. Blandus (Latin) = smooth tongue, flattering, friendly.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFB3FF89F0A5F30DFC31FB6E.taxon	discussion	Comments. Hylomyrma blandiens, H. dentiloba, and H. jeronimae have similar morphology. Hylomyrma blandiens was described based on two specimens (worker and queen), and H. dentiloba based on one worker. Hylomyrma blandiens has a broad distribution in northwestern South America, from Bolivia to French Guiana (Fig. 84), whereas H. dentiloba and H. jeronimae are restricted to Central America (Figs. 84, 86). Hylomyrma blandiens can be distinguished from H. dentiloba (characteristic in parentheses) in the distinguishable interspaces on head dorsum striation (vs. indistinguishable interspaces), the convex dorsal margin of mesonotum (vs. straight), the propodeal spine slightly longer than the dorsal tooth of propodeal lobe (vs. shorter), and the striae length on the first gastral segment similar to the postpetiole length (vs. shorter than the postpetiole length). Hylomyrma blandiens can be distinguished from H. jeronimae in the regular striae on the mesial region of head dorsum in part directed to posterior margin and in part anteriorly divergent and posteriorly convergent (vs. regular to irregular striae and divergent), the drop-shaped eye (vs. reniform), the longitudinal striation on the mesosomal dorsum (vs. striae assuming multiple directions), the discontinuous dorsal margin of petiole (vs. continuous), and the striae on the tergum of the first gastral segment extending up to its first third (vs. restricted to its basal region). Also, H. blandiens resembles H. macielae in the regular divergent-convergent striae on the mesial region of head dorsum, and the indistinguishable interspaces on the mesosomal striation. Hylomyrma blandiens can be distinguished from H. macielae in the longitudinal striae on the mesosomal dorsum (striae assuming multiple directions in H. macielae). The western limit of the distribution of H. blandiens, the east side of The Andes in Ecuador and Colombia (Fig. 84), overlaps in part with the distribution of H. macielae (Fig. 86). There is morphological variation across the range of H. blandiens. Specimens from Ecuador have longer propodeal spines than elsewhere in the range. Specimens from Ecuador and Tachira, Venezuela, have a more pronounced mesoventral projection on the petiole. Striae on the mesial area of the head dorsum vary from being longitudinal to somewhat angled, divergent anteriorly and convergent posteriorly.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFB3FF89F0A5F30DFC31FB6E.taxon	distribution	Distribution. This species is widespread in northwest South America (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname, and Venezuela) and Trinidad (Trinidad and Tobago) (Fig. 84). Natural history. Hylomyrma blandiens specimens are usually collected in samples of sifted leaf-litter in tropical rainforest. One Hylomyrma blandiens nest with 24 specimens (3 Q 21 W) was collected by M. A. Ulysséa in a fallen twig on the ground in the Peruvian Amazon (Madre de Dios, Estación Biológica Villa Carmen). Apart from the specimens, plant remains and parts of a Curculionidae imago were also found inside the nest, but there were no larvae or pupae. Additional material examined (202 workers, 32 queens, 1 male): BOLIVIA: Cochabamba: 109 k E Cochabamba at Lagunitas, 17 ° 06 ′ 22 ″ S, 65 ° 40 ′ 57 ″ W, 01. ii. 99, R. Anderson, Mont. evergreen forest, litter, # 18644 (2 W) [MZSP]; # 18636 (2 W) [MZSP]; La Paz: Caranari, Sara Ana, 410 msnm, 12. xii. 2011, A. Aduviri col., Bosque, Berlese (1 W) (CBF – Hym (Form) - 004761) [CBF]; [department unknown]: Rio Negro, Jan 1921 – 22, W. M. Mann collector, Mulford BioExpt [Biological Exploration] (2 W) [USNM]. BRAZIL: AC [Acre]: Porto Walter, 08 ° 15 ′ 31.2 ″ S, 72 ° 46 ′ 37.1 ″ W, 05. ii – 17. iv. 1997, J. Caldwell, Bufo typhonius, # 12688 (2 W) [CEPLAC]; same except # 13529 (1 Q) [CEPLAC]; # 13529 (1 W) [DZUP]; AM [Amazonas]: Manaus, Dimona Station, INPA, 100 ha Plot, 130 m, 60.09414 ° ′ W, 2.34121 ° ′ S, 16. i. 2009, J. Sosa-Calvo, rainforest, leaf litter, Winkler sample, JSC 090116 - LS 05 (1 W) [USNM]; same except JSC 090116 - LS 06 (1 W) [USNM]; 104 m, 60.09687 ° ′ W, 2.35365 ° ′ S, 13. i. 2009, JSC 090113 - LS 07 (2 W) [USNM]; PNJ Rio Carabinani, mar. Esq., Cachoeira Paredão, 2 ° 10 ′ 20 ″ S, 61 ° 32 ′ 22 ″ W, 29. vii – 1. viii. 1995, A. Santos col., F. Primária 2, Arm. Pitfall (3) (1 W) [INPA]; Presidente Figueiredo, I. Pe Inchado, L. Balbina, 1 ° 54.45 ′ S, 59 ° 29.75 ′ W, 25 / 02 / [19] 94, [M.] Queiroz col., mata secundária, serapilheira (1 W) [MPEG]; same except 1992 / [19] 94 [CEPLAC]; MT [Mato Grosso]: Alta Floresta, Frag. 155 - C 2, 56 ° 25 ′ 49 ″ E, 88 ° 95 ′ 55 ″ N, 11. vi. 2008, Castuera L. O. (1 W) [MZSP]; same except Frag. 151 - B 3, R. E. Vicente col., winkler (1 Q) [MZSP]; RO [Rondônia]: Porto Velho, Área Abunã, A 11 P 1, 09 ° 38 ′ 05.6 ″ S, 65 ° 27 ′ 11.2 ″ W, 17 – 27. vii. 2013, Mazão G. R. & Probst R. S. cols. (1 W) [DZUP]; same except A 10 P 2, 09 ° 35 ′ 53.1 ″ S, 65 ° 22 ′ 00.1 ″ W, 17 – 27. vii. 2013, Mazão G. R. & Probst R. S. cols. (1 W) [DZUP]; A 11 P 4, 09 ° 38 ′ 05.6 ″ S, 65 ° 27 ′ 11.2 ″ W, 4 – 17. i. 2012, Dattilo, W. F. & Mazão, G. R. cols. (1 W) [MZSP]; A 12 P 1, 09 ° 36 ′ 29.7 ″ S, 65 ° 22 ′ 40.7 ″ W, 17 – 30. vi. 2012, Sanhudo, C. E. D. & Andriolli, F. S. cols. (2 W, one worker just petiole, postpetiole and gaster) [MZSP]; A 12 P 1, 04 – 18. ix. 2012, Ulysséa, M. A. & Prado, L. P. cols. (5 W) [MZSP]; A 12 P 3, 09 ° 36 ′ 29.7 ″ S, 65 ° 22 ′ 40.7 ″ W, 04 – 18. ix. 2012, Ulysséa, M. A. & Prado, L. P. cols. (3 W) [MZSP]; A 12 P 4, 09 ° 36 ′ 29.7 ″ S, 65 ° 22 ′ 40.7 ″ W, 04 – 18. ix. 2012, Ulysséa, M. A. & Prado, L. P. cols. (3 W) [MZSP]; Porto Velho, Área Caiçara, C 1 P 3, 09 ° 26 ′ 14.6 ″ S, 64 ° 49 ′ 58.2 ″ W 17 – 27. vii. 2013, Mazão G. R. & Probst R. S. cols. (1 W) [DZUP]; same except C 1 P 4, 09 ° 26 ′ 31 ″ S, 64 ° 51 ′ 40 ″ W, 02 – 16. x. 2013, Mazão G. R. & Mendonça R. T. T. cols. (1 W) [DZUP]; C 3 P 4, 09 ° 26 ′ 46.8 ″ S, 64 ° 49 ′ 31.1 ″ W, 4 – 18. ix. 2012, Vicente R. E. & Oliveira J. cols. (1 W) [DZUP]; C 3 P 1, 17 – 30. vi. 2012, Sanhudo, C. E. D. & Andriolli, F. S. (1 W) [MZSP]; C 1 P 4, 09 ° 26 ′ 14.6 ″ S, 64 ° 49 ′ 58.2 ″ W, 04 – 18. ix. 2012, Ulysséa, M. A. & Prado, L. P. cols. (1 W) [MZSP]; C 3 P 1, 09 ° 26 ′ 41 ″ S, 64 ° 49 ′ 39 ″ W, 02 – 16. x. 2013, Mazão G. R. & Mendonça R. T. T. cols. (1 W) [DZUP]; Porto Velho, Área Mutum, M 5 P 1, 09 ° 35 ′ 29.5 ″ S, 65 ° 02 ′ 57.6 ″ W, 4 – 18. ix. 2012, Ulysséa, M. A. & Prado, L. P. cols. (1 W) [MZSP]; same except M 6 P 1, 09 ° 35 ′ 56 ″ S, 65 ° 02 ′ 59 ″ W, 17 – 27. vii. 2013, Mazão G. R. & Probst R. S. cols. (1 W) [DZUP]; M 6 P 2, 09 ° 36 ′ 27 ″ S, 65 ° 02 ′ 55 ″ W, 2 – 16. x. 2013, Mazão G. R. & Mendonça R. T. T. cols. (1 W) [DZUP]; M 8 P 1, 09 ° 37 ′ 42.9 ″ S, 65 ° 03 ′ 27.8 ″ W, 4 – 18. ix. 2012, Vicente R. E. & Oliveira J. cols. (4 W) [DZUP]; M 8 P 2, 09 ° 37 ′ 42.9 ″ S, 65 ° 03 ′ 27.8 ″ W, 4 – 18. ix. 2012, Ulysséa, M. A. & Prado, L. P. cols. (1 W) [MZSP]; Ouro Preto do Oeste, 28. iii. 1985, J. Dias, Res. INPA n ° 0170 (1 W) [INPA]. COLOMBIA: Caquetá: [Puerto] Solano, PNN [Serranía de] Chiribiquete, R. Sararamano, B. Verde military, 300 m, 0 ° 10 ′ 48 ″ N, 72 ° 37 ′ 24 ″ W, 9. iv. 2000, E. González leg., Winkler 14 (1 W) [IHVL]; same except 17. iv. 2000, Winkler 2 (1 W) [IHVL]. ECUADOR: Orellana: Tiputini Biodiversity Station, Jan. 2004 – Jan. 2006, A. L. Mertl col., dissertation 2009, litter nest, tropical rainforest, terra firme (1 W) [MCZC]; Napo: 20 km S of Tena, 600 m, 11 Jul 1976, S. & J. Peck, B 360 (2 W) [MCZC]. FRENCH GUIANA: [Cayenne]: Kaw. Mt., Amazon Nature Lodge, 950 ′, 52 ° 12.349 ′ W, 4 ° 33.426 ′ N, T. Schultz, U. Mueller & J. LaPolla, 2 ° forest, leaf litter, Winkler sample, TRS 050719 - 01 - LS 14 (1 W) [USNM]; same except TRS 050719 - 01 - LS 10 (3 W 1 Q) [USNM]; TRS 050719 - 01 - LS 15 (1 W) [USNM]; TRS 050719 - 01 - LS 17 (1 W) [USNM]; Nouragues Field Station, XII Trail, 39 m, 52 ° 40.57 ′ W, 4 ° 5.325 ′ N, 27. vii. 2005, T. R. Schultz, J. S. LaPolla & D. Price, 1 ° forest, litter sample, Winkler sample, TRS 050727 - 02 - LS 12 (2 W) [USNM]; same data (1 M) (USNMENT 00537690) [USNM]; Nouragues Natural Reserve Station, 102 km SW of Cayenne, lat 4.08799, lon - 52.67978, 145 m, 21. viii- 1. ix. 2018, Rainforest, Ant Course 2018 (2 W) [MZSP]; Nouragues, 4 ° 05 ′ 20 ″ N, 52 ° 40 ′ 28 ″ O, 100 – 140 m, 25. iii. 2006, Jérôme Orivel (+ autres), Fourmis de litière, Winkler, GPWTr 10 (1 Q) [MZSP]; [Saint Laurent du Maroni]: Maripassoula, 21 / 23. vi. 2000, Delabie, Durou, Dejean, Corbara & Giberneau [leg.], Winkler (1 W) [CEPLAC]. GUYANA: [Upper Demerara-Berbice]: Mabura Hill, camp at end of rd. from Georgetown to Letham Rd., 64 m, 58 ° 41.982 ′ W, 5 ° 9.313 ′ N, 29. x. 2002, J. S. LaPolla et al., 1 ° forest, litter sample, JSL 021029 - 01 - LS 20 (1 W) [USNM]; JSL 021029 - 01 - LS 15 (1 W) [USNM]; same except JSL 021029 - 01 - LS 08 (1 Q) [USNM]; Upper Takutu-Upper Essequibo: Acarai Mts., nr. Romeo′s Camp, 273 m, 58 ° 56.761 ′ W, 1 ° 23.122 ′ N, 10. x. 2006, T. R. Schultz & J. Sosa-Calvo, 1 ° forest swamp, leaf litter, Winkler sample, JSC 061010 - LS 08 (3 W) [USNM]; same except 263 m, 58 ° 56.772 ′ W, 1 ° 23.136 ′ N, JSC 061010 - LS 07 (1 W) [USNM]; 268 m, 58 ° 56.799 ′ W, 1 ° 23.216 ′ N, JSC 061010 - LS 01 (2 W 1 Q) [USNM]; 314 m, 58 ° 56.787 ′ W, 1 ° 23.137 ′ N, JSC 061010 - LS 06 (1 W) [USNM]; 293 m, 58 ° 56.804 ′ W, 1 ° 23.171 ′ N, JSC 061010 - LS 04 (2 W 1 Q) [USNM]; 303 m, 58 ° 56.808 ′ W, 1 ° 23.191 ′ N, JSC 061010 - LS 03 (2 W 1 Q) [USNM]; 280 m, 58 ° 56.806 ′ W, 1 ° 23.193 ′ N, JSC 061010 - LS 02 (1 W) [USNM]. PERU: Cusco: Estac. Biol. Villa Carmen, trail 12, 800 m, 12.869.043, - 71.407.11 + 300 m, 8. viii. 2013, M. A. Ulysséa col., rainforest, [nest in] rotten log (3 Q 7 W) [MZSP]; same data (14 W) (MZHY 101) [MZSP]; same data (2 W) (MZHY 180, MZHY 2015) [MZSP]; same except 5 – 15. viii. 2013, Ant Course 2013 (1 W) [MZSP]; same data (3 W) [DZUP]; Loreto: Ramón Castillo, 5 km NW Letícia, 23 Feb 1972, S. & J. Peck, forest, litter, berlesete, 231 (7 Q 23 W, one worker covered with gold) [MZSP]; same data (18 W 2 Q) [MCZC]; Madre de Dios: 202 m, 32.34 k SW Puerto Maldonado, 12 ° 49 ′ 36.9 ″ S, 69 ° 22 ′ 16.9 ″ W, 28. vii. 2012, W. & E. Mackay, mini-Winkler litter extraction, Tropical rain forest, tierra firme, reddishbrn clay, # 25167 (1 W) [UTEP]; same data (1 W covered with gold) (MZSP 67435) [MZSP]; same except # 25168 (1 W 1 Q) [UTLP]; # 25168 (1 W 1 Q) [MZSP]; # 25170 (1 W) [MZSP]; 230 m, 26. vii. 2012, # 25137 (1 W) [MZSP]; Los Amigos Field Station, Trail 6, 277 m, 70 ° 6 ′ 3.1 W, 12 ° 34 ′ 8.4 S, 6. x. 2004, T. R. Schultz, C. Marshall & J. Sosa-Calvo, 1 ° forest, litter sample, = JSC 041006 - 16, TRS 041006 - 01 - LS- 16 (2 W) [USNM]; Inkaterra, Reserva Amazónica, 11. vii. 2012, 19 L 0494117 - 19 L 0494099, 8614232 - 8614050, B. M. Jimenez col., B / GW 8 S (4 W) [UNMSM]; same except 8. vii. 2012, 19 L 0494901 - 19 L 0494572, 8614342 - 8613832, AW 2 seca (3 W) [MZSP]; AW 14 S (3 W) [UNMSM]; 5. vii. 2015, 19 L 0494646 - 19 L 0494557, 8614458 - 8614648, DW 14 S (4 W) [MZSP]; Reserva Nacional Tambopata, Sachavacayoc, 210 m, 12 ° 51 ’ 21 ″ S, 69 ° 21 ’ 43 ″ W, 19 – 31. vii. 2010, J. Chaul col., Neotropical Ant Course (1 W) [MZSP]; Tambopata, 15 km NE Puerto Maldonado, Cuzco Amazonica, 200 m, 20. vi. 89, S. P. Cover & J. E. Tobin cols, CA- 483, camp. sifted litter from compost pile by trail to zone I (3 W) [MCZC]. SURINAME: Sipaliwini: Nassau Mts., 514 m, 54 ° 36.24.3 W, 4 ° 49 ′ 1.9 N, 3. xi. 2005, J. Sosa-Calvo & R. Badal, 1 ° forest, litter sample, tall forest, high leaf litter decomposition, JSC 051103 - 01 - LS 11 (1 W) [USNM]; same except JSC 051103 - 01 - LS 19 (1 W) [USNM]; JSC 051103 - 01 - LS 10 (1 W) [USNM]; JSC 051103 - 01 - LS 18 (1 W) [USNM]; JSC 051103 - 01 - LS 13 (1 W) [USNM]; JSC 051103 - 01 - LS 09 (1 Q) [USNM]. [TRINIDAD AND TOBAGO]: TRINIDAD: [Arima]: Arima Valley, Windblow Ridge, 1800 ′, 5.15.88, Cover, Moffett, T- 44, secondary forest, rotten stick (4 W) [MCZC]; [Couva-Tabaquite-Talaro]: Arena Forest, mid Barker Trace, 5 Nov 2003, Wetterer, # 239 (1 W) [MCZC]; N Balata Branch, 31 Oct 2003, Wetterer, # 218 (1 W) [MCZC]; Arena & Brazil Rds, 5 Nov 2003, Wetterer, # 241 (1 W) [MCZC]; W Racoon Ride, 31 Oct 2003, Wetterer, # 216 (1 W) [MCZC]; 0.6 km ESE HQ, 9 July 2004, J. K. Wetterer, # 478 (1 W) [MCZC]; [Princes Town / Rio Calro-Mayaro]: Victoria-Mayaro Res., 2 km N gate, 3 Nov 2003, Wetterer, # 235 (1 Q) [MCZC]; [Rio Claro-Mayaro]: 17 km SE Rio Claro, 28 July 2004, J. K. Wetterer, # 533 (1 Q) [MCZC]; [Rio Claro-Mayaro / Sangre Gandre]: Nariva Swamp, 23. iv. 35, N. A. Weber, 140 (6 W) [MCZC]; [San Juan-Laventille]: North Cost Rd., 4 km from Saddle, 22 Nov 2003, Wetterer, # 249 (1 W 1 Q) [MCZC]; 3 km N Saddle, 2 Nov 2003, Wetterer, # 226 (1 Q) [MCZC]; [Sangre Grande]: Tamana nr cave, 12 July 2004, J. K. Wetterer, # 505 (1 Q) [MCZC]; [Tunapuna-Piarco]: Blanchisseuse Rd., 2 km N Asa Wright, 30 Oct 2003, Wetterer, # 212 (1 W) [MCZC]; La Veronica, 5.7 km up road, 19 June 2004, J. K. Wetterer, # 452 (1 W) [MCZC]. VENEZUELA: Amazonas: Rio Mavaca Cam, 65 ° 06 ′ W, 02 ° 02 ′ N, 16 – 27. iii. 89, Phipps-FUDECI Exped. by Amer. Mus. Nat. Hist., D. A. Grimaldi col. (1 W) [MCZC]; Tachira: Las Cuevas, 44 km WNW Sn. Cristobal, 7 ° 48 ′ N, 71 ° 46 ′ W, 500 m, 29. viii. 1988, col. J. Lattke, hojarasca tamizada (1 W) [MIZA]; Uribante-Caparo, 1217 (4 W) [MIZA]; Media Libra, Rio Negro Rd past Puente Santa Elena, 600 – 630 m, 10 Dec. 1985, J. Latke & W. L. Brown (1 W) [MIZA]; Aragua: P. N. H. Pittier, Valle Sta. María, 3.8 km SSW Cumboto, 615 m, 10 ° 22 ′ N, 67 ° 49 ′ W, 31. viii. 2003, E. Rodríguez, A. Grotto & J. Lattke, Fila El Viento, Bosque Seco, Ex hojarasca, 2803 (1 W) (MIZA 0021688 MZHY 220) [MZSP]; same except 2797 (1 W) (MIZA 0021690) [MIZA]; 2799 (1 W) (MIZA 0021692 MZHY 198) [MZSP]; 2802 (2 Q 1 W, worker covered with gold) (MIZA 0021687, MIZA 0021691, MIZA 0021689) [MIZA]; BA [Barinas]: 17 km SW Cdad Bolivia, 240 m, 8 ° 04 ′ N, 70 ° 48 ′ W, 27. viii. 87, J. Lattke, 1162 (2 W) [MIZA]; Sucre: 7 km S. El Pilar, <10 m, 29. vii. 1987, S. & J. Peck, hojarasca tamizada (2 W 1 Q) (MIZA 0021679, MIZA 0021683) [MIZA]; Zulia: El Tucuco, 51 km SO de Machiques, 420 m, 8. ix. 1984, J. Lattke leg., em hojarasca (1 Q) (MIZA 0021693) [MIZA].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF89FF8CF0A5F0DEFF3DF91D.taxon	description	Figures 27, 28, 85 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF89FF8CF0A5F0DEFF3DF91D.taxon	diagnosis	Diagnosis. Regular and longitudinal striae on head dorsum, mesial striation in part directed to posterior margin and in part anteriorly divergent and posteriorly convergent; irregular to regular striae of variable thickness on mesosoma dorsum, interspaces indistinguishable; mesonotum and metanotal groove region with semicircular striae; dorsal margin of petiole discontinuous, transverse striae on anterior region of dorsal surface continuing on lateral surface; subtriangular projection on mesoventral surface of petiole; subpostpetiolar process weak, convex; transverse striae on profemur posterior surface; protibia extensor surface striate; long striae on tergum of first gastral segment.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF89FF8CF0A5F0DEFF3DF91D.taxon	description	Redescription. WORKER (n = 4) (Fig. 27 A – C): HL 0.92 (0.92 – 1); HW 0.89 (0.89 – 0.94); ML 0.56 (0.56 – 0.62); SL 0.59 (0.59 – 0.70); MOD 0.21 (0.21 – 0.24); PNW 0.62 (0.62 – 0.66); WL 1.19 (1.19 – 1.26); PSL 0.22 (0.22 – 0.24); PL 0.57 (0.54 – 0.60); PW 0.26 (0.24 – 0.26); PPL 0.32 (0.32 – 0.36); PPW 0.35 (0.35 – 0.36); GL 0.91 (0.91 – 1.08); TL 4.50 (4.50 – 4.86); CI 96.47 (94 – 97.91); SI 65.85 (65.85 – 74.46); OI 24.39 (24.39 – 26.66). Small-sized. Shiny integument. Brownish body, yellowish appendices. Thin and unbranched setae, long to short, suberect to subdecumbent. Head subquadrate; posterior margin straight. Mandible masticatory margin with 6 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with 8 – 10 regular, longitudinal and thick striae, interspaces between thicker striae filled with thinner striae. Fontal triangle with 1 sagittal stria. Scape short, not reaching head posterior margin; apical antennomere with equal length or slightly shorter than 3 previous antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye drop-shaped, midsized, larger diameter with 12 ommatidia. Regular and longitudinal striae on head dorsum, mesial striation in part directed to posterior margin and in part anteriorly divergent and posteriorly convergent / parallel, interspaces between thicker striae filled with thinner striae. Striae of variable thickness on head lateral and laterodorsal regions converging to eye margin, interspaces indistinguishable; gena striate, 6 – 7 regular and semicircular striae circumscribe the torulus, almost reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Irregular to regular striae of variable thickness on mesosoma dorsum, interspaces indistinguishable; striae assume multiple directions; transverse striae on pronotum anterior region (DV) continuing on lateral surface; pronotum posterior region, mesonotum, and metanotal groove region (immediate anterior to transverse carina) with semicircular striae; part of the irregular striae on mesepisternum continuing on propodeum, and part is directed to propodeal spine. Promesonotal junction and metanotal groove indistinct. Transverse carina well-marked. Dorsal margin of mesosoma continuous, convex. Propodeal lobe bidentate, dorsal tooth longer and acute than the shorter and blunt ventral tooth; dorsal tooth length 1 / 2 of propodeal spine length. Propodeal spine midsized, straight (LV), divergent (DV), sculptured on base. Thin striae of uniform thickness on procoxa; irregular and transverse striae on C 2 and C 3. Profemur covered with irregular to regular transverse striae. Protibia extensor surface entirely covered with regular and longitudinal striae. Dorsal margin of petiole discontinuous. Subtriangular projection on mesoventral surface of petiole. Convex node; striae of variable thickness, interspaces indistinguishable; regular and transverse striae on anterior surface and on anterior region of dorsal continuing on lateral surface, striae weakly marked; middle and posterior regions of dorsal surface with longitudinal striae; transverse striae on ventral surface weakly marked. Postpetiole and subpostpetiolar process with regular and longitudinal striae; subpostpetiolar process weak, convex. First gastral segment striation similar to postpetiole striae; long striae on tergum, longer than postpetiole length; sternite striation covering the laterobasal region. QUEEN (first description) (n = 1) (Fig. 28 A – C): HL 0.96; HW 0.92; ML 0.60; SL 0.66; MOD 0.27; PNW 0.78; WL 1.38; PSL 0.28; PL 0.62; PW 0.28; PPL 0.38; PPW 0.42; GL 1.18; TL 5.12; CI 95.83; SI 71.74; OI 29.34. Midsized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 15 ommatidia. Longitudinal and regular striae of variable thickness on scutum going from an anterior central point towards transcutal suture, interspaces between thicker striae filled with thinner striae. Longitudinal striae of uniform thickness on anepisternum and katepisternum, interspaces indistinguishable. Longitudinal and mostly regular striae on axilla, interspaces indistinguishable. Scutoscutellar sulcus inconspicuous. Scutellum with the same sculpture as scutum. Transverse striae on propodeum (DV), interspaces indistinguishable. Lateral surface of mesosoma with mostly regular striae directed in part to propodeal dorsum and in part to propodeal spine, interspaces indistinguishable. Wings unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF89FF8CF0A5F0DEFF3DF91D.taxon	etymology	Etymology. The name is in reference to the species type locality, Colombia.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF89FF8CF0A5F0DEFF3DF91D.taxon	discussion	Comments. The sculpture on the mesosomal dorsum of H. columbica is similar to H. dolichops and H. montana. Hylomyrma columbica differs from H. dolichops (characteristic in parentheses) in the regular striae on the mesial region of head dorsum in part directed to posterior margin and in part anteriorly divergent and posteriorly convergent / parallel (vs. vermicular and divergent striae on anterior two-third of mesial area, irregular and parallel on the upper third), and the drop-shaped eye (vs. reniform). Hylomyrma columbica can be distinguished from H. montana in the combination of the regular striae on the mesial region of head dorsum in part directed to the posterior margin and in part anteriorly divergent and posteriorly convergent / parallel (vs. regular to irregular striae and divergent), the discontinuous dorsal margin of petiole (vs. continuous), and the long striae on tergum of the first gastral segment (vs. short striae). All three are allopatric in northwestern South America, but occur in nearby areas near the border between Colombia and Ecuador. Hylomyrma dolichops has a broader distribution (Brazil, Colombia, Ecuador, and Venezuela) (Fig. 87), whereas H. columbica occurs in Colombia and in western Venezuela (Fig. 85), and H. montana has been recorded in Costa Rica, Panama and Ecuador (Fig. 83).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF89FF8CF0A5F0DEFF3DF91D.taxon	distribution	Distribution. This species is only known from Colombia and western Venezuela (Fig. 85). Natural history. Specimens are frequently found in leaf-litter samples with winkler extractors or Berlese-Tüllgren funnels, which suggests that this species nests in the leaf-litter, fallen logs, rotten wood, or inside natural cavities of the superficial soil layers. Also, two specimens were found among plant fibers, which were sourced from Colombia, in a quarantine station in the USA, suggesting that workers may forage on vegetation.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF89FF8CF0A5F0DEFF3DF91D.taxon	materials_examined	Additional material examined (12 workers, 1 queen): COLOMBIA: on fiber plant from Colombia at Quarantine, Washington DC, 27 July 1916, E. R. Sasscer collector (2 W, one covered with gold) [USNM]; Huila: 4 k NE Rivera, 30 Dec 1986, W. Mackay, pitfall trap, forest area, # 9039 (1 W) (MZSP 67315) [MZSP]; [La] Guarija: R. Don Diego, 25 – 50 m, 18. june. 1976, W. L. Brown & R. C. Kugler [leg.], forest leaf litter (3 W 1 Q) [MCZC]; Mag. [Magdalena]: Pueblito - limite sur, Parque Tayrona, 210 – 360 m, 26. i. 78, C. Kugler col., berlese (1 W) [MCZC]; Pueblito, Tayrona PK., 360 m, 11. xi. 76, R. C. Kugler & W. L. Brown cols., leaf litter (1 W) [MCZC]; Meta: 1 km E. La Macarena, Caño Morrocoy, 260 m, 12. jan. 77, C. Kugler col., litter berlesate (1 W) [MCZC]; same except 25. iv. 1976 (1 W) [MCZC]. VENEZUELA: Zulia: El Tucuco, Dto. Perijá, 6. ix. 1984, cols. J. L attke, E. Rubio (1 W) (MZHY 192) [MZSP]; 54 km ESE Yaracal, Falcon, 935 m, Cerro Los Caracoles, 10.8718 – 69.0274 °, 24. iii. 2002, J. Lattke, 2482 (1 W) [DZUP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF8CFF8EF0A5F348FD4EF816.taxon	description	Figures 29, 83 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF8CFF8EF0A5F348FD4EF816.taxon	materials_examined	Holotype: COLOMBIA: Caquetá: San José de Fragua, R. Turayaco, 1250 m, 10 – 15. ix. 2000, E. L. G. [leg.], W 5 (1 W) [IHVL]. Paratypes: same data as holotype (1 W covered with gold) [IHVL]; (1 W) (MZHY 210) [MZSP]; same except 1500 m, 07 – 10. ix. 2000 (1 W) [MCZC]; Putumayo: Mocoa, 01 ° 08 ′ N, 76 ° 38 ′ W, Los Mayos, 1800 m, i. 1999, E. González leg., winkler 11 (1 W covered with gold) [IHVL]; same except winkler 10 (1 W covered with gold) [MZSP]; winkler 1 (1 W) [IHVL]; winkler 14 (2 W, one covered with gold) (MZSP 67317, MZSP 67318) [MZSP]; caida, T 1 T 10 (1 W) [MCZC].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF8CFF8EF0A5F348FD4EF816.taxon	diagnosis	Diagnosis. Irregular striae on head dorsum diverge to posterior margin; rugose striae on mesosoma and petiole; promesonotal junction and metanotal groove distinguishable by a slight depression and altered sculpture; dorsal margin of petiole discontinuous; transverse striae on node ventral surface weakly marked; profemur posterior surface smooth; protibia extensor surface striation weakly marked; striation on tergum of first gastral segment restricted to base.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF8CFF8EF0A5F348FD4EF816.taxon	description	Description. WORKER (n = 3) (Fig. 29 A – C): HL (0.98 – 1.04); HW (0.92 – 1); ML (0.61 – 0.64); SL (0.70 – 0.74); MOD (0.20 – 0.22); PNW (0.65 – 0.68); WL (1.30 – 1.40); PSL (0.27 – 0.30); PL (0.60 – 0.62); PW (0.25 – 0.26); PPL (0.34 – 0.40); PPW (0.36 – 0.38); GL (1.10 – 1.20); TL (4.96 – 5.30); CI (93.87 – 96.15); SI (73.95 – 76.09); OI (21.73 – 22.91). Midsized. Shiny integument. Bicolored body, brownish with lighter leg. Thin and unbranched setae, long to midsized, suberect to subdecumbent. Head subquadrate; posterior margin slightly concave at middle. Mandible masticatory margin with 5 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with 8 striae, regular and longitudinal striae converging to a point on the anterior margin, interspaces distinguishable. Frontal triangle with 1 sagittal stria or 1 stria subdivided towards the posterior margin. Short scape, not reaching head posterior margin; apical antennomere slightly shorter than 3 previous antennomeres together. Frontal carina straight. Eye oval, small-sized, larger diameter with 9 ommatidia. Irregular and longitudinal striae on head dorsum, divergent towards posterior margin, interspaces between thicker striae filled with thinner striae. Head lateral and laterodorsal regions with thin and regular striae converging to eye margin, interspaces indistinguishable; anterior part of laterodorsal region with few irregular and longitudinal striae connecting the region posterior to frontal carina with eye margin; gena striate, 4 – 6 regular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma covered with rugose striae of uniform thickness, transverse on pronotum anterior region (DV), longitudinal in other regions (DV), interspaces between rugose striae smooth. Promesonotal junction and metanotal groove distinguishable by a slight depression (LV) and altered sculpture (DV). Transverse carina inconspicuous. Dorsal margin of mesosoma slightly discontinuous, convex. Propodeal lobe bidentate, dorsal tooth slightly longer and sharper than the shorter and blunt ventral tooth; dorsal tooth length around 1 / 3 shorter than propodeal spine length. Propodeal spine midsized, straight or apex curved downwards (LV), divergent (DV), sculptured on base. Procoxa striae of uniform thickness; irregular and transverse striae on C 2 and C 3. Profemur mostly smooth. Protibia extensor surface entirely striate. Dorsal margin of petiole discontinuous. Convex projection on mesoventral surface of petiole. Convex node; transverse striae on ventral and anterior surfaces weakly marked; rugose striae on lateral and dorsal surfaces. Postpetiole and subpostpetiolar process partly covered with regular and longitudinal striae, weakly marked or smooth; subpostpetiolar process weak, convex. First gastral segment striation similar to postpetiole striae; longitudinal striae on tergum shorter than postpetiole length; sternite smooth. QUEEN Unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF8CFF8EF0A5F348FD4EF816.taxon	etymology	Etymology. The epithet dandarae is a Latin noun in the genitive case created by adding the singular Latin genitive case suffix - e to the first name of a female person. This species is named in honor of Dandara (? – 1694), a leader and warrior in the resistance against slavery during the Brazilian colonial period. She was part of the “ Quilombo dos Palmares ”, the largest settlement of African and Afro-Brazilian peoples who escaped enslavement.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF8CFF8EF0A5F348FD4EF816.taxon	discussion	Comments. Hylomyrma dandarae is similar to H. adelae, H. mariae, and H. wachiperi with respect to the sculpture of the head dorsum and mesosoma, and the profile of the mesosoma. Hylomyrma dandarae can be distinguished from H. adelae (characteristics in parentheses) based on the following characters: oval eye (vs. dropshaped), irregular divergent-longitudinal striae on the head dorsum (vs. rugose striae), distinct promesonotal junction and metanotal groove (vs. both indistinct), and tergum of the first gastral segment with short striae (vs. long striae). The two species occur allopatrically in Bolivia (Fig. 83), H. adelae in Cochabamba and H. dandarae in La Paz. Hylomyrma dandarae differs from H. mariae in the oval eye (vs. drop-shaped), the longer petiolar node (vs. shorter), and the short striae on tergum of the first gastral segment restrict to its first third (vs. long striae). Also, both species occur in nearby geographic areas, H. dandarae in the south (Fig. 83) and H. mariae in the center and north of Colombia (Fig. 88). Hylomyrma dandarae can be distinguished from H. wachiperi in the irregular striae on the head dorsum (vs. regular striae), the rugose striae on the mesosomal dorsum (vs. irregular striae), the distinct metanotal groove (vs. indistinct), the short propodeal spine (vs. long), and the tergum of the first gastral segment with long striae (vs. short striae). Hylomyrma wachiperi occurs in Peru whereas H. dandarae occurs in Bolivia and Colombia. There is morphological variation across the range of H. dandarae. The shape of the propodeal spines vary from being entirely straight to having a downwardly curved apex. The specimens from La Paz, Bolivia, have irregular and transverse striae on the petiolar dorsum.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF8CFF8EF0A5F348FD4EF816.taxon	distribution	Distribution. Hylomyrma dandarae is known from only three localities, two in southwest Colombia and one in La Paz, Bolivia (Fig. 83). Natural history. This species inhabits sites at elevations from 1250 to 2048 m. Specimens were collected with pitfall traps and winkler extractors, which suggests that workers forage on the forest floor, and that nests are located in fallen logs, rotten wood, between leaves, or inside natural cavities of the superficial soil layers.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF8CFF8EF0A5F348FD4EF816.taxon	materials_examined	Additional material examined (2 workers): BOLIVIA: La Paz: Nor Yungas, Coripata, Altuspata, 2048 m, 8198991 S, 639994 W, 5. iii. 2009, borde del bosque, trampa pitfall, Beatriz Mamani col. (1 W) (CBF – Hym (Form) - 004762) [CBF]. COLOMBIA: Putumayo: Mocoa, 01 ° 08 ′ N, 76 ° 38 ′ W, Los Mayos, 1800 m, i. 1999, E. González leg., winkler 10 (1 W) (MZHY 222) [MZSP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF8FFF82F0A5F494FD74FBF4.taxon	description	Figures 30, 31, 80 D, 81 G, 86 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF8FFF82F0A5F494FD74FBF4.taxon	diagnosis	Diagnosis. Head dorsum with regular and longitudinal striae, mesial striation in part directed to posterior margin and in part anteriorly divergent and posteriorly convergent, interspaces distinguishable; mesosoma covered with regular to irregular and adjacent striae of variable thickness; dorsal margin of petiole discontinuous, dorsum with same striation of mesosoma lateral; subtriangular projection on mesoventral surface of petiole; subpostpetiolar process weak, convex; transverse striae on profemur posterior surface; protibia extensor surface striate; striation on tergum of first gastral segment restricted to base.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF8FFF82F0A5F494FD74FBF4.taxon	description	Redescription. WORKER (n = 4) (Fig. 30 A – C): HL 0.85 (0.84 – 0.92); HW 0.76 (0.76 – 0.84); ML 0.52 (0.52 – 0.60); SL 0.55 (0.54 – 0.60); MOD 0.21 (0.21 – 0.22); PNW 0.55 (0.55 – 0.62); WL 1.12 (1.10 – 1.20); PSL 0.15 (0.15 – 0.16); PL 0.52 (0.52 – 0.58); PW 0.22 (0.22 – 0.26); PPL 0.27 (0.27 – 0.32); PPW 0.32 (0.31 – 0.34); GL 0.85 (0.85 – 1.08); TL 4.15 (4.15 – 4.65); CI 89.70 (89.70 – 91.30); SI 72.13 (70.23 – 75); OI 27.87 (25 – 26.31). Small to midsized. Subopaque integument, except for the shiny gaster. Head, antennae and mesosoma light brown; yellowish leg; brownish petiole, postpetiole and gaster. Thin and mostly unbranched setae, long to midsized, suberect to subdecumbent; few branched setae with 2 – 3 or multiple short branches of equal size (Fig. 80 D), specially on gaster. Head subquadrate; posterior margin straight. Mandible masticatory margin with 5 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with regular and longitudinal striae, being 6 – 7 thicker striae, interspaces indistinguishable. Frontal triangle with 2 – 3 striae. Scape short, not reaching head posterior margin; apical antennomere shorter than 3 previous antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye reniform, midsized, larger diameter with 11 ommatidia. Striae of variable thickness on head dorsum regular to irregular, longitudinal, mesial striation in part directed to posterior margin and in part anteriorly divergent and posteriorly convergent, interspaces indistinguishable. Head lateral and laterodorsal regions with same striation of head dorsum, striae converging to eye margin; anterior part of laterodorsal region with few striae connecting the region posterior to frontal carina with eye margin; gena striate, irregular and semicircular striae circumscribe the torulus, not reaching eye margin, interspaces indistinguishable. Interspaces between striae on head ventral surface distinguishable. Mesosoma covered with regular to irregular striae, thinner striae (microsculpture) of variable thickness superimposed on thicker striae (macrosculpture), interspaces indistinguishable. Striation assumes multiple directions; transverse striae on pronotum anterior region (DV) continuing on lateral surface in part towards propodeal spine and in part transversely on propodeum; longitudinal striae on posterior region of pronotum and mesonotum. Promesonotal junction indistinct. Metanotal groove distinguishable by a slight depression (LV). Transverse carina inconspicuous. Dorsal margin of mesosoma slightly discontinuous, mesosoma somewhat straight (mesosoma flattened dorsally). Propodeal lobe bidentate; both teeth blunt, dorsal tooth slightly longer than ventral tooth. Propodeal spine short, straight (LV), divergent (DV), sculptured, and shorter than dorsal tooth of propodeal lobe. Procoxa striae of uniform thickness; regular and transverse striae on C 2 and C 3. Profemur and protibia mostly covered with regular striae, transverse on profemur, longitudinal on protibia. Dorsal margin of petiole discontinuous. Convex node; regular striae on anterior, lateral and dorsal surfaces, interspaces indistinguishable; interspaces distinguishable on node ventral surface; transverse striae on anterior surface continuing longitudinally on lateral surface; longitudinal striae of variable thickness on dorsal surface. Postpetiole and subpostpetiolar process with regular and longitudinal striae; subpostpetiolar process weak, convex. First gastral segment striation similar to postpetiole striae; short striae on tergum, shorter than 1 / 2 of postpetiole length; laterobasal region of sternite covered with striation. QUEEN (first description) (n = 4) (Fig. 31 A – C): HL (0.92 – 0.95); HW (0.82 – 0.88); ML (0.54 – 0.58); SL (0.58 – 0.62); MOD (0.24 – 0.26); PNW (0.75 – 0.78); WL (1.32 – 1.38); PSL (0.22 – 0.24); PL (0.60 – 0.63); PW (0.27 – 0.30); PPL (0.32 – 0.34); PPW (0.39 – 0.42); GL (1.08 – 1.12); TL (4.79 – 4.98); CI (89.13 – 93.61); SI (68.18 – 70.73); OI (27.90 – 30.48). Midsized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 12 – 13 ommatidia. Striae of variable thickness, longitudinal, regular to irregular on scutum, going from an anterior central point towards transcutal suture; thinner striae (microsculpture) between thicker striae (macrosculpture). Longitudinal and mostly regular striae of variable thickness on anepisternum and katepisternum, interspaces indistinguishable. Axilla and scutellum with the same sculpturing pattern as scutum. Irregular striae of variable thickness on propodeum, transverse in DV, directed in part to propodeal dorsum and in part to propodeal spine of lateral surface of mesosoma, interspaces indistinguishable. Wings as in Fig. 81 G. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF8FFF82F0A5F494FD74FBF4.taxon	etymology	Etymology. Dens, - entis (Latin) = tooth; lobus (Latin) and lobos (Greek) = rounded projection or protuberance.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF8FFF82F0A5F494FD74FBF4.taxon	discussion	Comments. Hylomyrma dentiloba is similar to H. blandiens, H. jeronimae, and H. macielae. Hylomyrma dentiloba and H. jeronimae are restricted to Central America (Figs. 84, 86), co-occuring in Barro Colorado Island, Panama, whereas H. blandiens shows a broad distribution in northwestern South America, from Bolivia to French Guiana (Fig. 84), overlapping in part with the area of distribution of H. macielae (south Colombia and the centernorth of Ecuador) (Fig. 86). Hylomyrma dentiloba can be distinguished from H. blandiens (characteristic in parentheses) in the indistinguishable interspaces on head dorsum striation (vs. distinguishable), the straight dorsal margin of mesonotum (vs. convex), the propodeal spine shorter than the dorsal tooth of propodeal lobe (vs. slightly longer), and the striae length on the first gastral segment shorter to the postpetiole length (vs. similar than the postpetiole length). Hylomyrma dentiloba can be distinguished from H. jeronimae in the regular striae on the mesial region of the head dorsum in part directed towards the posterior margin and in part anteriorly divergent and posteriorly convergent (vs. regular to irregular striae and divergent), the drop-shaped eye (vs. reniform), the longitudinal striation on the mesosomal dorsum (vs. striae assuming multiple directions), and the discontinuous dorsal margin of petiole (vs. continuous). Hylomyrma dentiloba can be distinguished from H. macielae in the drop-shaped eye (vs. oval), and the straight dorsal margin of the mesonotum (vs. convex). In molecular analysis, the specimens of H. dentiloba from Barro Colorado Island were separated into two clusters: (H. cf. dentiloba sp. 1 (H. plumosa (H. cf. dentiloba sp. 2, H. versuta ))) (Pierce et al. 2017). After observing the images of H. cf. dentiloba sp. 1 (CASENT 0636001) from Costa Rica, we understand that this morphospecies is H. jeronimae n. sp. and that H. cf. dentiloba sp. 2 is the true H. dentiloba. The voucher specimens of sp. 2 still need to be examined. Also, Pierce et al. (2017) stated that H. dentiloba and H. versuta are morphologically identical, being differentiated only in their geographical distribution. The authors use H. dentiloba to refer to H. dentiloba - like material from the Osa Peninsula of Costa Rica south through Panama, and H. versuta for material from the Atlantic slope of Costa Rica northward. However, we found that H. dentiloba (the H. cf. dentiloba sp. 2) can be distinguished from H. versuta in the striation with indistinguishable interspaces on the head dorsum (vs. striation with distinguishable interspaces), the mesosoma and petiolar node covered by thinner striae with indistinguishable interspaces superimposed on irregular and thicker striae (vs. thinner striae with indistinguishable interspaces between irregular and thicker striae), the profemur covered with regular and transverse striae weakly marked (vs. predominantly smooth), the protibia mainly covered with regular and weakly marked striae (vs. predominantly smooth), and the shorter striae on tergum of the first gastral segment (vs. longer striae). Hylomyrma dentiloba can be distinguished from H. jeronimae (the H. cf. dentiloba sp. 1) as mentioned in the previous paragraph.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF8FFF82F0A5F494FD74FBF4.taxon	distribution	Distribution. Hylomyrma dentiloba distribution is restricted to Panama Canal Zone (Fig. 86). Natural history. The species inhabits sites at elevations from 90 to 100 m. Some specimens were sampled in leaf-litter, which suggests that this species nests between leaves, in fallen logs, rotten wood, or inside natural cavities of the superficial soil layers.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF8FFF82F0A5F494FD74FBF4.taxon	materials_examined	Additional material examined (48 workers, 8 queens): PANAMA: C [anal] Z [one], Barro Colorado I., 27. ii. 1976, A. Newton, leaf litter, forest floor (2 W) [MCZC]; same locality, June-Oct- 1943, Zetek - 5105, Lot 43 - 16534 (9 W 3 Q) [USNM]; same data (2 W, one worker covered with gold) (MZSP 67320) [MZSP]; (1 W) [IHVL]; (2 W 1 Q) [IFML]; same locality, i. 1960, W. L. Brown, E. S. McCluskey, B- 43 (2 W) [MCZC]; (1 W) [MZSP]; same locality, Apr-May- 42, J. Zetek n ° 4953 (3 W) [USNM]; (1 W) [CASC]; same locality, vii. 1941, JasZetek n ° 4852, Lot n ° 41 - 20631 (1 Q) (MZSP 67321) [MZSP]; Colón: San Lorenzo, IBISCA, W 13115, F 3 - May 05, T 146 - 19 (2 W) [CEPLAC]; same data (1 W) (MZHY 201) [MZSP]; same except W 13057, F 2 - May 05, T 145 - 12 (3 W) [CEPLAC]; W 13044, F 1 - May 05, T 154 - 50 (3 W) (MZSP 67473, MZHY 216) [MZSP]; W 13090, F 2 - May 05, T 145 - 12 (3 W) [CEPLAC]; San Lorenzo Forest, 30. xii. 2004, 9 ° 17 ′ N, 79 ° 58 ′ W, Dejean, Orivel, Corbara, Aberlenc, Leponce, B 2, W 12585, Winkler (1 W) [CEPLAC]; same except B 1, W 12521 (1 W 1 Q) [DZSP]; F 3, W 12305 (1 Q) [MZSP]; Punta de Los Chivos, W. side Gatun Lake, 3 km SW Gatun, 100 m, 3 – 9. july. 1979, W. L. Brown col., forest (10 W 1 Q) [MCZC]; Gigante Peninsula, 90 m, 9.11363 - 79.85430 + 1 km, june. 2010 - july. 2011, T. P. Sumnicht col., Wet forest, ex leaf litter (1 W) (CASENT 0643420) [MZSP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF82FF85F0A5F061FB86FBD0.taxon	description	Figures 32, 33, 80 G, 87 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF82FF85F0A5F061FB86FBD0.taxon	diagnosis	Diagnosis. Anterior half of head dorsum with vermicular and divergent striae; posterior half with irregular and parallel striae; large and reniform eye; striae of variable thickness on mesosoma, interspaces indistinguishable; semicircular striae on mesonotum and metanotal groove region; dorsal margin of petiole discontinuous; profemur posterior surface mostly smooth; striae on protibia extensor surface weakly marked; long striae on tergum of first gastral segment; body with branched setae with multiple branches of equal size and lobed at tip.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF82FF85F0A5F061FB86FBD0.taxon	description	Redescription. WORKER (n = 4) (Fig. 32 A – C): HL (0.92 – 1.04); HW 0.98 (0.94 – 1); ML 0.62 (0.62 – 0.68); SL 0.68 (0.62 – 0.72); MOD 0.32 (0.28 – 0.33); PNW 0.72 (0.70 – 0.78); WL 1.38 (1.30 – 1.44); PSL 0.42 (0.36 – 0.46); PL 0.69 (0.66 – 0.78); PW 0.31 (0.27 – 0.31); PPL 0.42 (0.40 – 0.46); PPW 0.40 (0.37 – 0.42); GL 1.32 (1 – 1.32); TL 5.43 (4.90 – 5.43); CI 98 (98 – 102.17); SI 69.38 (65.95 – 70); OI 32.65 (29.78 – 33). Medium to large-sized. Subopaque to shiny integument. Brownish to light brown body, darker head and gaster. Thin, unbranched and branched setae, long to short, suberect to subdecumbent; branched setae with multiple short branches of equal size and lobed at tip (Fig. 80 G); unbranched setae mostly on head. Head subquadrate; posterior margin straight. Mandible masticatory margin with 6 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with 6 – 8 regular, longitudinal and thick striae, interval between thicker striae filled with thinner striae. Frontal triangle with 1 stria subdivided towards the posterior margin. Short scape, not reaching head posterior margin; apical antennomere with equal length to previous 3 antennomeres together, or slightly shorter. Frontal carina straight. Eye reniform, large-sized, larger diameter with 17 ommatidia. Longitudinal striae on head dorsum, vermicular and divergent striae on anterior half, irregular and parallel on posterior half; interspaces between thicker striae filled with thinner striae. Striation on lateral and laterodorsal regions of the head converging to eye margin; regular striae on head lateral and lateroventral regions, irregular to vermicular on laterodorsal region; gena striate, 6 – 9 regular and semicircular striae circumscribe the torulus, almost reaching eye margin. Interspaces between striae on head ventral surface striation indistinguishable. Irregular to regular striae of variable thickness on mesosoma dorsum assuming multiple directions, striae forming swollen regions, interspaces indistinguishable; pronotum mostly covered with transverse striae continuing on lateral surface; semicircular striae on dorsum of mesonotum, metanotal groove region (immediate anterior to transverse carina) and propodeum. Irregular striae on lateral of pronotum and mesepisternum continuing on propodeum lateral and reaching the propodeal spine. Promesonotal junction and metanotal groove indistinct. Transverse carina inconspicuous. Dorsal margin of mesosoma continuous, convex. Propodeal lobe bidentate, acute dorsal tooth with similar length of blunt ventral tooth; dorsal tooth length shorter than 1 / 2 of propodeal spine length. Propodeal spine long, straight (LV), divergent (DV), sculptured on base. Thin striae of uniform thickness on procoxa; irregular and transverse striae on C 2 and C 3. Profemur covered with irregular to regular transverse striae weakly marked. Protibia extensor surface entirely covered with regular and longitudinal striae. Dorsal margin of petiole discontinuous. Node elongated, convex; adjacent striae of variable thickness; regular and transverse striae on anterior surface continuing on lateral surface; irregular to vermicular longitudinal striae on lateral and dorsal surfaces; transverse striae on ventral surface weakly marked. Postpetiole and subpostpetiolar process with regular and longitudinal striae, interspaces indistinguishable; subpostpetiolar process prominent, convex. First gastral segment striation similar to postpetiole striae; long striae on tergum, shorter than postpetiole length; sternite striation covering the laterobasal region. QUEEN (n = 2, paratypes) (Fig. 33 A – C): HL 1.02 – 1.06; HW 1.02; ML 0.56 – 0.66; SL 0.72; MOD 0.34; PNW 0.86 – 0.88; WL 1.50 – 1.62; PSL 0.40; PL 0.72 – 0.74; PW 0.34; PPL 0.43 – 0.48; PPW 0.44 – 0.46; GL 1.36 – 1.42; TL 5.59 – 5.98; CI 96.22 – 100; SI 70.58; OI 33.33. Large-sized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 18 – 20 ommatidia. Longitudinal, regular to irregular striae on scutum going from an anterior central point towards transcutal suture, interspaces indistinguishable. Longitudinal striae of uniform thickness on anepisternum and katepisternum, mostly regular on anepisternum and irregular on katepisternum, interspaces indistinguishable. Axilla and scutellum with same sculpture of scutum. Scutoscutellar sulcus inconspicuous. Transverse and regular striae on propodeum (DV), interspaces indistinguishable. Lateral surface of mesosoma with mostly irregular striae directed to propodeal spine, interspaces indistinguishable. Wings unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF82FF85F0A5F061FB86FBD0.taxon	etymology	Etymology. Dolichops (Greek, dolichos = kidney-bean) is certainly a reference to the reniform eye of this species.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF82FF85F0A5F061FB86FBD0.taxon	discussion	Comments. Hylomyrma dolichops is similar to H. columbica regarding the sculpture of the mesosomal dorsum, but the former is easily distinguished from the latter by the branched setae, whose branches are lobed at the tip (Fig. 80 G). Hylomyrma columbica has unbranched setae. Also, Hylomyrma dolichops differs from H. columbica (characteristic in parentheses) in the presence of vermicular and divergent striae on anterior two-third of mesial area on head dorsum, irregular and parallel on the upper third (vs. regular striae on the mesial region in part directed towards posterior margin and in part anteriorly divergent and posteriorly convergent / parallel), and the reniform eye (vs. drop-shaped). The two species are allopatric; Hylomyrma dolichops shows a broader distribution (Brazil, Colombia, Ecuador, and Venezuela) (Fig. 87), whereas H. columbica occurs in Colombia and in western Venezuela (Fig. 85). Regarding morphological variation, three specimens from Orito, Colombia, are considerably larger than those described in the type series, although all other diagnostic characteristics are invariant.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF82FF85F0A5F061FB86FBD0.taxon	distribution	Distribution. This species is known from Brazil, Colombia, Ecuador, and Venezuela (Fig. 87). Natural history. All specimens examined were collected in leaf-litter of tropical rainforest, at elevations ranging from 260 to 700 m, which suggests that nests are located between leaves, in fallen logs, rotten wood, or inside natural cavities of the superficial soil layers.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF82FF85F0A5F061FB86FBD0.taxon	materials_examined	Additional material examined (7 workers): BR [BRAZIL]: AC [Acre]: Mâncio Lima, P. N. da Serra do Divisor, 260 m, 7 ° 27 ′ 9.22 ″ S, 73 ° 39 ′ 58.24 ″ W, 15 – 18. xi. 2016, R. M. Feitosa, T. S. Silva & A. C. Ferreira cols. (2 W) [DZUP]. COLOMBIA: Nariño: Orito, Territorio Kofan, 700 m, 00 ° 30 ′ N, 77 ° 13 ′ W, 28. ix. 1998, E. L. González leg., Bosque, Winkler 7 (1 W) [IHVL]; same data (1 W) (MZHY 197) [MZSP]; same except 29. ix. 1998, caída, T 2 T 6 (1 W covered with gold) [IHVL]. ECUADOR: Orellana: Tiputini Biodiversity Station, Jan 2004 – Jan 2006, A. L. Mertl col., dissertation 2009, litter nest, tropical rainforest, terra firme (1 W) [MCZC]. VEN [VENEZUELA]: Amazonas: San Carlos de R. Negro, 11. oct. 1981, L. Carling col., SCRN HR / JV (1 W) [MCZC].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF85FF9BF0A5F085FE14FE6C.taxon	description	Figures 34, 35, 36, 80 B, 82 E, 88 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF85FF9BF0A5F085FE14FE6C.taxon	diagnosis	Diagnosis. Large-sized; subopaque integument; body with thin striae (microsculpture) superimposed on vermicular and longitudinal striae (macrosculpture), macrosculpture absent on postpetiole and gaster, interspaces between thin striae indistinguishable; propodeal spine long; dorsal margin of petiole continuous, convex; dorsal margin of postpetiole subtriangular; subpostpetiolar process prominent, subtriangular; profemur posterior surface and protibia extensor surface striae; very long striae on tergum of first gastral segment.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF85FF9BF0A5F085FE14FE6C.taxon	description	Redescription. WORKER (n = 9) (Fig. 34 A – C): HL 1.20 (1.10 – 1.28); HW 1.15 (1.10 – 1.28); ML 0.78 (0.74 – 0.88); SL 0.84 (0.82 – 0.98); MOD 0.28 (0.24 – 0.30); PNW 0.80 (0.77 – 0.90); WL 1.56 (1.51 – 1.75); PSL 0.44 (0.36 – 0.58); PL 0.73 (0.68 – 0.82); PW 0.28 (0.27 – 0.30); PPL 0.36 (0.36 – 0.45); PPW 0.38 (0.36 – 0.42); GL 1.34 (1.24 – 1.44); TL 5.97 (5.67 – 6.47); CI 95.83 (95.76 – 101.61); SI 73.04 (70.50 – 79.64); OI 24.34 (21.81 – 24.77). Large-sized. Subopaque integument, except for the shiny gaster. Dark brown head and mesosoma, other body parts lighter. Many thin and unbranched setae (Fig. 80 B), long to midsized, erect to subdecumbent. Head subquadrate; posterior margin slightly concave at middle. Mandible masticatory margin with 6 teeth. Anterior margin of clypeus straight medially, with a pair of medium teeth laterally; median area of clypeus with clypeus centrally striate, 2 – 4 regular, longitudinal and thicker striae (macrosculpture) covered with thinner striae (microsculpture), interspaces between thin striae indistinguishable. Frontal triangle with 1 sagittal stria. Short scape, not reaching head posterior margin; apical antennomere shorter than previous 3 antennomeres together. Frontal carina straight. Eye drop-shaped, midsized, larger diameter with 13 ommatidia. Head dorsum with thin striae (microsculpture) superimposed on vermicular and longitudinal striae (macrosculpture), interspaces between thin striae indistinguishable. Head lateral and laterodorsal regions with same striation of head dorsum, striae converging to eye margin; gena striate, irregular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma sculpture similar to head dorsum. Promesonotal junction indistinct. Transverse carina inconspicuous (LV). Metanotal groove discernible by a slight depression (LV). Dorsal margin of mesosoma slightly discontinuous, convex. Propodeal lobe bidentate, dorsal tooth slightly longer and acute than the shorter and blunt ventral tooth; dorsal tooth length 1 / 2 shorter than propodeal spine length. Propodeal spine long, straight (LV), divergent (DV), sculptured. Procoxa striae of uniform thickness; irregular and transverse striae on C 2 and C 3. Profemur covered with irregular to regular striae, mostly transverse on anterior and dorsal surfaces, longitudinal on apical region. Protibia entirely covered with regular and longitudinal striae. Dorsal margin of petiole continuous, convex; sculpture similar to mesosoma; ventral surface regular and smooth. Longitudinal and adjacent striae on postpetiole and subpostpetiolar process; dorsal margin of postpetiole subtriangular; subpostpetiolar process prominent, subtriangular. First gastral segment striation similar to postpetiole striae; long striae on tergum, 11 / 2 × longer than postpetiole length; sternite striation covering entirely the basal region. QUEEN (n = 7) (Fig. 35 A – C): HL 1.13 (1.13 – 1.36); HW 1.06 (1.06 – 1.36); ML 0.73 (0.73 – 0.92); SL 0.82 (0.82 – 1); MOD 0.29 (0.28 – 0.36); PNW 0.90 (0.90 – 1.08); WL 1.62 (1.62 – 2.08); PSL 0.40 (0.39 – 0.50); PL 0.71 (0.71 – 0.94); PW 0.28 (0.28 – 0.35); PPL 0.42 (0.42 – 0.52); PPW 0.39 (0.39 – 0.51); GL 1.46 (1.46 – 1.74); TL 6.07 (6.07 – 7.54); CI 93.80 (93.80 – 101.50); SI 77.35 (69.23 – 73.77); OI 27.35 (23.73 – 27.05). Large-sized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 14 – 15 ommatidia. Vermicular striae on scutum going from an anterior central point towards transcutal suture, interspaces indistinguishable. Longitudinal striae of uniform thickness on anepisternum and katepisternum, mostly regular, interspaces indistinguishable. Axilla and scutellum with same sculpture of scutum. Scutoscutellar sulcus conspicuous. Transverse striae on propodeum (DV). Lateral surface of mesosoma with vermicular striae directed in part to propodeal dorsum and in part to propodeal spine, interspaces indistinguishable. Wings unknown. MALE (first description) (n = 4) (Fig. 36 A – C): HL (0.98 – 1.04); HW (0.82 – 90); ML (0.58 – 0.63); SL (0.30 – 0.34); MOD (0.35 – 0.36); PNW (0.90 – 0.98); WL (1.70 – 1.84); PL (0.78 – 0.85); PW (0.29 – 0.30); PPL (0.44 – 0.52); PPW (0.42 – 0.43); GL (1.34 – 1.48); TL (5.96 – 6.26); CI (83.67 – 90.81); SI (35.95 – 37.77); OI (40 – 42.68). Brownish body, yellowish appendices, darker gaster. Mandible with 6 teeth. Scutum covered with striation, striae mostly longitudinal, regular to irregular, interspaces distinguishable. Anepisternum and katepisternum mostly covered with longitudinal, regular and thinner striae, interspaces distinguishable. Scutellum with irregular to vermiculated-areolated striae, thicker than scutum striae, interspaces distinguishable. Propodeum with irregular striae assuming multiple directions, interspaces distinguishable. Propodeal lobe subquadrate. Petiole mostly covered with longitudinal and irregular striae, interspaces indistinguishable. Postpetiole with longitudinal and anastomosed striae. First gastral tergum striate on basal region. Interspaces between striae smooth. Wings as in Fig. 82 E.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF85FF9BF0A5F085FE14FE6C.taxon	etymology	Etymology. The name refers to the species size (Latin, immanis = large, huge). Hylomyrma immanis is the largest species of the genus.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF85FF9BF0A5F085FE14FE6C.taxon	discussion	Comments. Hylomyrma immanis and H. praepotens are the largest species in the genus (TL 5.67 – 6.47 mm, WL 1.51 – 1.58 mm). Both have a long propodeal spine, and an elongated and continuous dorsal margin of the petiole, but H. immanis is easily distinguished by the subopaque integument (shiny in H. praepotens), and the body covered by thin striae with interspaces indistinguishable (thick striae with interspaces distinguishable in H. praepotens). Hylomyrma immanis is more easily sampled than H. praepotens, the former has a broad distribution in the center-north of South America (Fig. 88), whereas the latter is only known from two localities in Colombia, two in Ecuador, and one in Brazil (Fig. 90). Both species co-occur in a locality 7 km NW of Letícia, Amazonas, Colombia.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF85FF9BF0A5F085FE14FE6C.taxon	distribution	Distribution. This species is widely distributed in northwest South America (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, and Venezuela), Trinidad (Trinidad and Tobago), and in eastern portions of Brazil (Fig. 88). Natural history. This species inhabits forested areas in Atlantic and other tropical rainforests of South America. Most specimens were sampled in leaf-litter with winkler extractors, which suggests that H. immanis nests between leaves, in fallen logs, rotten wood, or inside natural cavities of the superficial soil layers. Additional material examined (194 workers, 30 queens, 4 males): BOLIVIA: Cochabamba: Villa Tunari, 67.5 k E Valle Sajta, 17 ° 06 ′ 19 ″ S, 64 ° 46 ′ 57 ″ W, 6. ii. 99, R. Anderson, Lowland rain forest, litter, # 18525 (2 W) [MZSP]; same except 07. ii. 99, # 18528 (1 W) [MZSP]; La Paz: Caranavi, Saraana [Sara Ana], 410 m, 1. xi. 2011, col. A. Aduviri, Bosque, Berlese (1 W) [CBF]. BRAZIL: AC [Acre]: Mâncio Lima, P. N. da Serra do Divisor, 245 m, 7 ° 26 ′ 17.19 ″ S, 73 ° 39 ′ 27.39 ″ W, 15 – 18. xi. 2016, R. M. Feitosa, T. S. Silva & A. C. Ferreira cols. (3 W) [DZUP]; same except 260 m, 7 ° 27 ′ 9.22 ″ S, 73 ° 39 ′ 58.24 ″ W (1 W) [DZUP]; AL [Alagoas]: Murici, ESEC Murici, ICMBio, 23. ix. 2014, 09 ° 14 ′ 57.04 ″ S, 35 ° 47 ′ 28.88 ″ W, Lorenzo E. P. col. (2 W) [UFGD, MZSP]; AM [Amazonas]: PNJ Rio Carabinani, mar. esq., Cachoeira Paredão, 2 ° 10 ′ 20 ″ S, 61 ° 32 ′ 22 ″ W, 29. vii – 1. viii. 1995, A. Santos col., F. Primária 2, Arm. Pitfall (4) (1 W) [INPA]; Manaus, BR 174 km 46 - EEST, 8. v. 1991, Eq. A. Y. Harada, isca sardinha, mata, 115 (1 W) [INPA]; Pres [idente] Figueiredo, Terra-firme, Ig. Poraque, 1 ° 54 ′ 57 ″ S, 59 ° 26 ′ 94 ″ W, 23 / 04 / 94, Queiroz col., Mata primária, Arm. de solo (1 W) [MPEG]; BA [Bahia]: Aritaguá, 14 ° 39 ′ 39 ″ S, 39 ° 04 ′ 33 ″ W, 05 / x / 1998, Col. J. C. S. Carmo & J. R. M. Santos, 477 (3 W) [CEPLAC]; Jaguaripe, 9 - 03 - [19] 94, Nascimento I. C., 4817 (4 W) [CEPLAC]; Ilhéus, Mata da Boa Esperança, 14 ° 47 ′ 47 ″ S, 09 ° 03 ′ 56 ″ W, 09. ix. 2000, Santos J. R. M. col., Winkler 30 (1 W) [MZSP]; same except Winkler 7 (1 W) [MZSP]; Winkler 8 (1 W) [MZSP]; Winkler 11 (1 W) [MZSP]; Winkler 16 (1 W) [MZSP]; Winkler 12 (1 W) [MZSP]; Winkler 20 (1 W) [MZSP]; Winkler 3 (1 W) [MZSP]; Winkler 48 (3 W) [MZSP]; Ilhéus-Banco do Pedro, 12. i. 98, 144051 S, 0391524 W, Santos J. R. M. dos, Carmo J. C. S. do, Mata W-A 23 (3 W) [CEPLAC]; Mata São João, Reserva Sapiranga, 12 ° 33 ′ 29.3 ″ S, 33 ° 02 ′ 35.2 ″ W, 21 - 28. vii. 2001, Silva R. R., Brandão C. R. F. cols., Winkler 40 (2 W) [MZSP]; same except Winkler 38 (1 W) [MZSP]; Winkler 27 (1 W) [UFGD]; Winkler 44 (1 W covered with gold) [MZSP]; Winkler 45 (1 W) (MZSP 67469) [MZSP]; Winkler 41 (2 W) [MZSP]; Winkler 41 (1 W 1 Q) (MZSP 67335) [MZSP]; Porto Seguro, E. E. Pau Brasil, 16 ° 23 ′ 33 ″ S, 39 ° 10 ′ 99 ″ W, 16. vi. 2000, Santos J. R. M., Soares J. C. cols., Winkler 31 (1 W) [MZSP]; same except Winkler 27 (3 W) [MZSP]; Winkler 12 (1 W) [MZSP]; Winkler 44 (1 W) [MZSP]; Travessão, 14 ° 08 ′ 13 ″ S, 39 ° 16 ′ 39 ″ W, 28 / 04 / 1997, Santos J. R. M. (3 W) [CEPLAC]; Ubaitaba, 14 ° 15 ′ 01 ″ S, 39 ° 19 ′ 27 ″ W, 10 - 11 / 04 / 1997, Col. J. C. S. Carmo (2 W 1 Q) [CEPLAC]; Una, 15 ° 16 ′ 45 ″ S, 39 ° 05 ′ 29 ″ W, 29 / 09 - 22 / 11 / 1996, Santos J. R. M. (1 W) [CEPLAC]; same data (1 Q 1 M) (MZSP 67334) [MZSP]; MA [Maranhão]: Açailândia, Horto Faz. Pompéia, 04 ° 52 ′ 30 ″ S, 47 ° 17 ′ 40 ″ W, 12 - 22. vi. 2006, Silva R. R. & Feitosa R. M. cols., Winkler 4 (1 W) [MZSP]; same except 13 - 22. vi. 2006, Winkler 04 (1 W) [MZSP]; Winkler 01 (1 W) [MZSP]; MT [Mato Grosso]: Sinop, 55 ° 37 ′ W, 12 ° 31 ′ S, x. 1974, Alv. & Roppa, 12568 (1 W) [MZSP]; same except 12567 (2 M) (MZSP 67333) [MZSP]; Alta Floresta, Frag. 67 — C 8, 58 ° 82 ′ 36 ″ E, 89 ° 19 ′ 35 ″ N, 8. v. 2008, Castuera L. O. (3 W) [MZSP]; Novo Mundo, P. E. Cristalino, xi. 2012, R. E. Vicente col., (PPBIO), parc. 9, pitfall 7 (1 W) [MZSP]; parc. 2, pitfall 4 (1 W) [MZSP]; PA [Pará]: Belém, Utinga Forest Preserve, 45 m, 48.42876 ° ′ W, 1.41739 ° ′ S, 3. ii. 2009, J. Sosa-Calvo, rainforest, Winkler sample, JSC 090203 - LS 03 (2 W) [USNM]; same except JSC 090203 - LS 05 (1 Q) [USNM]; Monte Dourado, Área 75, 0 ° 42.38 - 146 ″ S, 52 ° 42.10102 ″ W, 27 - 29. iv. 2011, Marsh C. J. col., Floresta primária, Pt. PS-P 04,? (1 W) [MZSP]; Jari, Amazônia, corte-seletivo, 0 ° 53 ′ S 52 ° 36 ′ W, 2011, Silva E. A. col., estrato EPI, parcela 21, pto B (1 W) [DZUP]; same except parcela 7, pto F (1 W) [DZUP]; PE [Pernambuco]: Recife, Horto Dois Irmãos, 08 ° 00 ′ 32 ″ S, 34 ° 56 ′ 40 ″ W, 15 - 24. vii. 2002, Silva R. R. & Eberhardt F. cols., Winkler 27 (1 W) [MZSP]; same except Winkler 38 (1 W) [UTLP]; Winkler 40 (2 W) [MZSP]; Winkler 44 (1 W) [MZSP]; Winkler 50 (1 W) [MZSP]; Winkler 45 (1 W) [MZSP]; Winkler 18 (3 W) [MZSP]; Winkler 15 (3 W 1 Q) [MZSP]; Winkler 14 (2 W) [MZSP]; Winkler 12 (2 W) [MZSP]; Winkler 4 (2 W) [MZSP]; Winkler 26 (1 W) [MZSP]; Winkler 16 (2 W 1 Q) [MZSP]; Winkler 2 (1 Q) [MZSP]; Winkler 19 (1 Q) [MZSP]; Winkler 46 (1 Q) [MZSP]; Timbaúba, Engenho Água Azul, 07 ° 36 ′ 05.56 ″ S, 35 ° 22 ′ 30.51 ″ W, 15. vi. 2013, Lorenzo E. P. col. (3 W) [UFGD]; RO [Rondônia]: Porto Velho, UHE SAE, Módulo Jiráu, 09.43726 ° S, 064.68475 ° W, 09. i. 2011, C. E. D. Sanhudo col. (1 W) [MZSP]; same locality, Área Abunã, A 10 P 1, 09 ° 35 ′ 48 ″ S, 64 ° 21 ′ 56 ″ W, 2 - 16. x. 2013, Mazão G. R. e Mendonça R. T. T. cols. (2 W 1 Q) [DZUP]; same except A 10 P 3, 09 ° 35 ′ 53.1 ″ S, 65 ° 22 ′ 00.1 ″ W, 04 - 18. ix. 2012, Ulysséa, M. A. & Prado, L. P. cols. (1 W) [MZSP]; A 11 P 2, 09 ° 38 ′ 05.6 ″ S, 65 ° 27 ′ 11.2 ″ W, 04 - 18. ix. 2012, Ulysséa, M. A. & Prado, L. P. cols. (1 Q 3 W) [MZSP]; A 12 P 2, 09 ° 36 ′ 38 ″ S, 65 ° 22 ′ 05 ″ W, 2 - 16. x. 2013, Mazão G. R. e Mendonça R. T. T. cols. (1 W) [DZUP]; Porto Velho, Área Caiçara, C 1 P 4, 09 ° 26 ′ 31 ″ S, 64 ° 51 ′ 40 ″ W, 2 - 16. x. 2013, Mazão G. R. e Mendonça R. T. T. cols. (1 W) [DZUP]; same except C 3 P 2, 09 ° 26 ′ 46.8 ″ S, 64 ° 49 ′ 31.1 ″ W, 4 - 18. ix. 2012, Vicente R. E. e Oliveira J. cols. (2 W 1 Q) [DZUP]; 04 - 18. ix. 2012, Ulysséa, M. A. & Prado, L. P. cols. (3 W) [MZSP]; C 3 P 4, 09 ° 26 ′ 46.8 ″ S, 64 ° 49 ′ 31.1 ″ W, 17 - 30. vi. 2012, Sanhudo, C. E. D. & Andriolli, F. S. (2 W) [MZSP]; 04 - 18. ix. 2012, M. A. Ulysséa & L. P. Prado (5 W) [MZSP]; Porto Velho, Área Mutum, M 7 P 2, 09 º 35 ′ 25 ″ S, 65 º 04 ′ 08 ″ W, 11 - 19. v. 2010, R. R. Silva & R. M. Feitosa cols. (1 W) [MZSP]; same except M 7 P 3, 09 ° 35 ′ 41.6 ″ S, 65 ° 03 ′ 54.2 ″ W, 4 - 18. ix. 2012, Vicente R. E. e Oliveira J. cols. (1 W) [DZUP]; M 7 P 4, 09 ° 35 ′ 41.6 ″ S, 65 ° 03 ′ 54.2 ″ W, 17 - 27. vii. 2013, Mazão G. R. e Probst R. S. cols. (1 W) [DZUP]; 04 - 18. ix. 2012, Ulysséa, M. A. & Prado, L. P. cols. (1 W) [MZSP]; SE [Sergipe]: Sta. Luiza do Itanhy, Crasto, 11 ° 22 ′ 39.3 ″ S, 37 ° 25 ′ 07.4 ″ W, 29. vii- 03. viii. 2001, Silva R. R., Brandão C. R. F., Winkler 15 (1 W) [MZSP]; same except Winkler 2 (1 W) [MZSP]; Winkler 34 (2 W) [MZSP]; Winkler 38 (1 W) [MZSP]; Winkler 46 (2 W) [MZSP]; Winkler 20 (2 W) [MZSP]; Winkler 47 (3 W) [MZSP]; Winkler 48 (1 W) [MZSP]; Winkler 17 (2 W) [MZSP]; Winkler 28 (1 W) [MZSP]; Winkler 34 (3 W) [MZSP]; Winkler 1 (1 W 1 Q) [MZSP]. COLOMBIA: Caquetá: [Puerto] Solano, PNN [Serranía de] Chiribiquete, R. Sararamano, B. Verde military, 300 m, 0 ° 10 ′ 48 ″ N, 72 ° 37 ′ 24 ″ W, 9. iv. 2000, E. González leg., Winkler 14 (3 W) [IHVL]; Amazonas: Araracuara, 300 m, G. Ganghi (1 W) [IHVL]; Guaviare: RN Nukak Maku, Cñ Cucuy Cr Moyano, Rebalse, 02 ° 10 ′ 40 N, 71 ° 11 ′ 25 W, feb. 96, F. Fernandez F. E. (1 W) [IHVL]; Nariño: Orito, Territorio Kofan, 700 m, 00 ° 30 ′ N, 77 ° 13 ′ W, 29. ix. 1998, E. L. González leg., Bosque caída, T 2 T 4 (1 W) [IHVL]. ECUADOR: Napo: 20 km S of Tena, 600 m, 11 Jul 1976, S. & J. Peck, B 360 (2 W) [MCZC]; [Sucumbíos]: Cuyabeno, 12 / 10 - 05 / 11 / 94, J. P. Caldwell, # 10427 (1 W) [CEPLAC]. FRENCH GUIANA: [Cayenne]: Nouragues Field Station, H Trail, 108 m, 52 ° 40.856 ′ W, 4 ° 5.108 ′ N, 1. viii. 2005, T. R. Schultz, J. S. LaPolla, D. Price, 1 ° forest, litter sample, Winkler sample, TRS 050801 - 03 - LS 12 (1 W 1 Q) [USNM]; Nouragues Natural Reserve Station, 102 km SW of Cayenne, lat 4.08799, lon - 52.67978, 145 m, 21. viii- 1. ix. 2018, Rainforest, Ant Course 2018 (1 W) [MZSP]; Nouragues, 4 ° 04 ′ 58 ″ N, 52 ° 40 ′ 28 ″ O, 120 - 160 m, 29. iii. 2006, Jérôme Orivel (+ autres), Fourmis de litière, FLWTr 45 (1 W 1 Q) [MZSP]; [Saint Laurent du Maroni]: Maripassoula, 21 / 23. vi. 2000, Delabie, Durouh, Dejean, Corbara & Giberneau [leg.], Winkler (1 Q) [CEPLAC]; Mitaraka, Pente, 2.238226 N - 54.45203 W, 03. iv. 2015, J. Orivel & F. Petitclerc cols., Winkler 48 h, Transect A, Plot P 2, Point 2 MI 15 - 0374 - 27 (1 W) [ECOFOG]. GUYANA: [Cuyuni-Mazaruni]: Calm Water Creek, along Essequibo River nr. Bartica, 58 ° 37.16 ′ W, 6 ° 28.06 ′ N, 24. ix. 2002, J. S. LaPolla, 1 ° forest, litter sample, = 020923 - 7, JSL 020923 - 01 - LS 03 (1 W) [USNM]; same except = 020923 - 6, JSL 020923 - 01 - LS 02 (1 W 1 Q) [USNM]; = 020923 - 5, JSL 020923 - 01 - LS 01 (1 W) [USNM]; = 020923 - 10, JSL 020923 - 01 - LS 06 (1 W) [USNM]; = 020923 - 9, JSL 020923 - 01 - LS 05 (1 W) [USNM]; [Potaro-Siparuni]: Iwokrama For. Res., Whitewater Camp, 60 m, 58 ° 50.992 ′ W, 4 ° 43.89 ′ N, 5. xi. 2002, J. S. LaPolla et al., 1 ° forest, litter sample, = 021105 - 1 - LS 13, JSL 021105 - 01 - LS 13 (1 W) [USNM]; same except = 021105 - 1 - LS 17, JSL 021105 - 01 - LS 17 (2 W) [USNM]; = 021105 - 1 - LS 18, JSL 021105 - 01 - LS 18 (1 W) [USNM]; = 021105 - 1 - LS 20, JSL 021105 - 01 - LS 20 (2 W) [USNM]; = 021105 - 1 - LS 9, JSL 021105 - 01 - LS 09 (1 W) [USNM]; Dicymbe Forest, 717 m, 59 ° 54.63 ′ W, 5 ° 17.76 ′ N, 6. x. 2002, T. R. Schultz, J. LaPolla, C. Marshall, R. Williams, Dicymbe forest, litter sample, TRS 021006 - 01 - LS 18 (1 Q) [USNM]; [Upper Demerara-Berbice]: Mabura Hill, camp at end of rd. From Georgetown to Letham Rd., 64 m, 58 ° 41.982 ′ W, 5 ° 9.313 ′ N, 29. x. 2002, J. S. LaPolla et al., 1 ° forest, litter sample, JSL 021029 - 01 - LS 10 (1 W) [USNM]; same except JSL 021029 - 01 - LS 15 (1 W 1 Q) [USNM]; JSL 021029 - 01 - LS 17 (1 W) [USNM]; JSL 021029 - 01 - LS 14 (1 W) [USNM]; JSL 021029 - 01 - LS 14 (1 Q) [USNM]; Upper Takutu-Upper Essequibo: Acarai Mts., nr. Romeo’s Camp, 293 m, 58 ° 56.804 ′ W, 1 ° 23.171 ′ N, 10. x. 2006, T. R. Schultz & J. Sosa-Calvo, 1 ° forest, litter sample, Winkler sample, JSC 061010 - LS 04 (2 W 2 Q 1 M) [USNM]; same except 314 m, 58 ° 56.787 ′ W, 1 ° 23.137 ′ N, JSC 061010 - LS 06 (1 Q) [USNM]. PERU: Loreto: Campamento San Jacinto, 11. july. 1993, R. Leschen, # 86, rainforest, flower fall, berlasete (4 W) [MCZC]; Madre de Dios: Los Amigos Field Station, Trail 6, Huangana, 277 m, 70 ° 6 ′ 3.1 W, 12 ° 34 ′ 8.4 S, 6. x. 2004, T. R. Schultz, C. Marshall, J. Sosa-Calvo, 1 ° forest, litter sample, = JSC 041006 - 20, TRS 041006 - 01 - LS- 20 (2 W 1 Q) [USNM]; same except = JSC 041006 - 16, TRS 041006 - 01 - LS- 16 (1 W) [USNM]; Trail 3, = JSC 041009 - 10, TRS 041009 - 01 - LS- 10 (1 W) [USNM]; Trail 3, = JSC 041009 - 18, TRS 041009 - 01 - LS- 18 (1 W) [USNM]; Trail 3, = JSC 041009 - 05, TRS 041009 - 01 - LS- 05 (1 W) [USNM]; Trail 3, = JSC 041009 - 04, TRS 041009 - 01 - LS- 04 (2 W 1 Q) [USNM]; across River from Cocha Cashu, 19.90 ° S, 71.36 ° W, 23 Sept. 1999, D. W. Davidson, Ex sifted litter, FAS — 000948 (1 W) [UNMSM]; Pto [Puerto] Maldonado, nr. Lake Sandoval, 260 m, 13 - 16. vi. 81, C. Kugler & R. R. Lambert cols., 1 ° (?) for., in sandy soil, berl. lf. lit. (4 W 1 Q) [MCZC]; Cusco: Est. Biol. Villa Carmen, 590 m, 5 - 15. viii. 2013, - 12.902437 ° - 71.407672 ° + 30 m, Ant Course 2013, bamboo forest, secondary vegetation, KAM # 422 (1 W) [MZSP]. [TRINIDAD AND TOBAGO]: TRINIDAD: [Rio Claro-Mayaro / Sangre Gandre]: Nariva Swamp, 22. iv. 35, N. A. Weber, 139 (1 W) [MCZC]; [Princes Town]: 13.5 km E Preau, 27 July 2004, J. K. Wetterer, # 528 (1 W) [MCZC]; [Sangre Grande]: 2 km NW Howson, 12 July 2004, J. K. Wetterer, # 506 (1 Q) [MCZC]; [Couva-Tabaquite-Talaro]: Arena Forest, Barker Trace, 20 May 2004, J. K. Wetterer, # 416 (1 W) [MCZC]; N Balata Branch, 31 Oct 2003, Wetterer, # 218 (1 Q) [MCZC]; [Tunapuna-Piarco]: Mt. Tucuche, 25. vi. 76, J. Noyes, QCAZ I 57120 (1 W) [MZSP]; Blanchisseuse Rd., 2 km N Asa Wright, 30 Oct 2003, Wetterer, # 212 (1 W) [MCZC]. VENEZUELA: Amazonas: Alto Rio Siapa, 495 m, 01 ° 42 ′ 50 ″ N, 64 ° 33 ′ 40 ″ O, 8. ii. 89, J. Lattke (1 Q) [MIZA]; same except 4. ii. 89 (1 Q) [MIZA].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9BFF9EF0A5F5D9FBC3FC08.taxon	description	Figures 37, 38, 80 F, 84 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9BFF9EF0A5F5D9FBC3FC08.taxon	materials_examined	Holotype: PANAMA: [Panamá Oeste]: Cerro Campana, 950 m, [ca. 8.73 ° N, 79.97 ° W (Branstetter, 2013)], 5. vi. 95, R. Anderson [leg.], wet mountain, forest litter, # 17833 (1 W) [MCZC]. Paratypes: same data as holotype, # 17833 (7 W) (MZSP 67337, MZSP 67338) [MZSP]; # 17833 (1 Q) (MZSP 67336) [MZSP]; # 17754 (1 W) [UTEP]; # 17754 (2 W) [USNM]; # 17754 (2 W, one covered with gold, without gaster) [DZUP]; 14 - 23. ii. 1976, leaf litter, forest floor, A. Newton [leg.] (10 W) (MCZENT 00525548, MCZENT 00525550, MCZENT 00525555, MCZENT 00525547, MCZENT 00525493, MCZENT 00525551, MCZENT 00525552, MCZENT 00525549, MCZENT 00525494, MCZENT 00525546) [MCZC]. COSTA RICA: Puntarenas: 10 k W Rincón, 22. vi. 1997, 180 m, 8 ° 42 ′ N, 83 ° 31 ′ W, R. Anderson [leg.], ridge for., litter extr., 18689 B (2 W) (MZSP 67339, MZSP 67340) [MZSP]; 17 k NE Rincón, 21. vi. 1997, 250 m, 8 ° 45 ′ N, 83 ° 25 ′ W, R. Anderson [leg.], lowland for., litter extr., 18685 D (1 W) [CASC]; same except 18685 D (1 W covered with gold) (MZSP 67341) [MZSP]; 18685 A (1 W) (MZSP 67342) [MZSP]; 24. vi. 1997, 8 ° 45 ′ 30 ″ N, 83 ° 25 ′ 00 ″ W, lowland for., litter, 18689 (1 W) (MZHY 204) [MZSP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9BFF9EF0A5F5D9FBC3FC08.taxon	diagnosis	Diagnosis. Regular to irregular striae on head dorsum diverge towards posterior margin; irregular striae of variable thickness on mesosoma assuming multiple directions, interspaces indistinguishable; reniform eye; metanotal groove distinguishable by a slight depression (LV) and altered sculpture (DV); dorsal margin of petiole continuous, convex, dorsum and lateral with same striation of mesosoma; subpostpetiolar process weak, convex; profemur posterior surface and protibia extensor surface weakly striate; striation on tergum of first gastral segment restricted to base.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9BFF9EF0A5F5D9FBC3FC08.taxon	description	Description. WORKER (n = 3) (Fig. 37 A – C): HL (0.88 – 0.90); HW (0.80 – 0.82); ML (0.52 – 0.53); SL (0.56 – 0.58); MOD (0.25 – 0.26); PNW (0.57 – 0.58); WL (1.12); PSL (0.21 – 0.24); PL (0.56 – 0.57); PW (0.20); PPL (0.28 – 0.29); PPW (0.29 – 0.30); GL (0.93 – 1.06); TL (4.31 – 4.47); CI (90.90 – 92.13); SI (68.29 – 72.50); OI (30.48 – 32.50). Smallsized. Subopaque integument, except for the shiny gaster. Light brown body, darker gaster, yellowish leg. Many thin and branched setae, long to midsized, suberect to subdecumbent, 2 – 4 short branches of equal size arising from the main axis (Fig. 80 F). Head subquadrate; posterior margin straight. Mandible masticatory margin with 5 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with regular and longitudinal striae, being 6 – 9 thicker striae, interspaces indistinguishable. Frontal triangle with 3 striae. Short scape, not reaching head posterior margin; apical antennomere shorter than previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye reniform, midsized, larger diameter with 11 ommatidia. Regular to irregular and longitudinal striae on head dorsum, divergent towards posterior margin, interspaces between thicker striae filled with thinner striae. Head lateral and laterodorsal regions with same striation of head dorsum, striae converging to eye margin; gena striate, irregular and semicircular striae circumscribe the torulus, not reaching eye margin, being 4 – 6 thicker striae, interspaces indistinguishable. Interspaces between striae on head ventral surface distinguishable. Mesosoma covered with regular to irregular striae, thinner striae (microsculpture) superimposed on thicker striae (macrosculpture), interspaces between thinner striae indistinguishable. Striation assumes multiple directions; transverse striae on pronotum anterior region (DV) continuing on lateral surface in part towards propodeal spine and in part transversely on propodeum; pronotum posterior region, mesonotum and metanotal groove with semi-elliptical to V-shaped striae. Promesonotal junction indistinct. Metanotal groove discernible by a slight depression (LV) and altered sculpture (DV). Transverse carina well-marked. Dorsal margin of mesosoma slightly discontinuous, convex. Propodeal lobe bidentate, acute dorsal tooth slightly longer than blunt ventral tooth; dorsal tooth length slightly shorter than propodeal spine length. Propodeal spine long, straight (LV), divergent (DV), sculptured. Procoxa striae of uniform thickness; regular and transverse striae on C 2 and C 3. Profemur and protibia covered with irregular to regular striae, transverse on profemur, longitudinal on protibia. Dorsal margin of petiole continuous, convex; transverse striae on entire ventral surface, interspaces distinguishable; first third of dorsal surface smooth; striation of its second third continuing on lateral surface; regular to irregular striae of variable thickness on last third of dorsal surface and lateral surface, last third of dorsal surface with semicircular striae, interspaces indistinguishable. Postpetiole and subpostpetiolar process with regular and longitudinal striae; subpostpetiolar process weak, convex, striation weakly marked. First gastral segment striation similar to postpetiole striae; short striae on tergum, shorter than postpetiole length; sternite striation covering the laterobasal region. QUEEN (n = 3) (Fig. 38 A – C): HL (0.94); HW (0.88 – 0.89); ML (0.55 – 0.62); SL (0.60 – 0.62); MOD (0.28 – 0.29); PNW (0.78 – 0.82); WL (1.38 – 1.40); PSL (0.28); PL (0.64 – 0.66); PW (0.26); PPL (0.33 – 0.34); PPW (0.37 – 0.38); GL (1.20 – 1.28); TL (5.08 – 5.18); CI (93.61 – 94.68); SI (68.18 – 70.45); OI (31.46 – 32.95). Midsized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 14 – 15 ommatidia. Longitudinal, regular to irregular striae on scutum going from an anterior central point towards transcutal suture, interspaces indistinguishable. Longitudinal and mostly regular striae on anepisternum and katepisternum, interspaces indistinguishable. Axilla and scutellum with same sculpture of scutum. Scutoscutellar sulcus conspicuous. Transverse and regular striae on propodeum (DV). Lateral surface of mesosoma with irregular to vermicular striae directed in part to propodeal dorsum and in part to propodeal spine. Wings unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9BFF9EF0A5F5D9FBC3FC08.taxon	etymology	Etymology. The epithet jeronimae is a Latin noun in the genitive case created by adding the singular Latin genitive case suffix - e to the first name of a female person. The specific epithet is named after Jerônima Mesquita (1880 – 1972), a Brazilian feminist, pioneer of the women’s suffrage in Brazil. She also advocated for the equality of rights and opportunity of women, and co-founder, along with Berta Lutz (1894 – 1976) and Stella Guerra Duval (1879 – 1971), of the League for the Intellectual Emancipation of Women in 1918 (which was subsequently named Brazilian Federation for Women’s Progress).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9BFF9EF0A5F5D9FBC3FC08.taxon	discussion	Comments. Hylomyrma jeronimae is similar to H. blandiens, H. dentiloba and H. macielae. Hylomyrma jeronimae and H. dentiloba are restricted to Central America (Figs. 84, 86), co-occuring in Barro Colorado Island, Panama, whereas H. blandiens has a broad distribution in northwestern South America, from Bolivia to French Guiana (Fig. 84), overlapping in part with the area of distribution of H. macielae (south Colombia and the center-north of Ecuador) (Fig. 86). Hylomyrma jeronimae can be distinguished from H. blandiens and H. dentiloba (characteristic of both in parentheses) by the regular to irregular, and divergent striae on the mesial region of head dorsum (vs. regular striae in part directed to posterior margin and in part anteriorly divergent and posteriorly convergent), the reniform eye (vs. drop-shaped), the striation on the mesosoml dorsum assuming multiple directions (vs. longitudinal striation), the continuous dorsal margin of petiole (vs. discontinuous), and the striae on tergum of the first gastral segment restrict to its basal region (vs. extending up to its first third). Hylomyrma jeronimae and H. macielae share the adjacent striae on the mesosoma, but the former differs from the latter in the reniform eye (vs. oval), and the continuous dorsal margin of petiole (vs. discontinuous). After observing the images of H. cf. dentiloba sp. 1 (CASENT 0636001) from Costa Rica (Pierce et al. 2017), we understand that this species is Hylomyrma jeronimae n. sp. The authors stated that H. dentiloba and H. versuta are morphologically identical, being differentiated only in their geographical distribution. However, H. jeronimae (the H. cf. dentiloba sp. 1) can be distinguished from H. versuta in the striation on the mesosomal dorsum assuming multiple directions (vs. logitudinal striation), the striae interspaces on the mesosoma indistinguishable (vs. distinguishable), and the continuous dorsal margin of petiole (vs. discontinuous). Also, we indicate five morphological differences between H. jeronimae and the true H. dentiloba (the H. cf. dentiloba sp. 2) in the previous paragraph.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9BFF9EF0A5F5D9FBC3FC08.taxon	distribution	Distribution. This species occurs in Costa Rica and Panama (Fig. 84). Natural history. Hylomyrma jeronimae inhabits forested areas at elevations ranging from 180 to 950 m. Most known specimens were collected in the leaf-litter, which suggests that nests are located in fallen logs, rotten wood, between leaves, or inside natural cavities of the superficial soil layers.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9BFF9EF0A5F5D9FBC3FC08.taxon	materials_examined	Additional material examined (42 workers, 3 queens): CR [COSTA RICA]: Punt. [Puntarenas]: Osa Penn [Peninsula] C. Helado, 17 k NE Rincón, 24. vi. 97, R. Anderson, # 18689 (2 W) (MZHY 204) [MZSP]; San Jose: 68 km PanamHw 3 de Jun. Boq., 10. ii. 96, R. Anderson, litter ex. Forest Adjacent to Sphagnum bug [bog], Sample C, # 17750 (3 W) [MZSP]. PANAMA: [Panamá Oeste]: Cerro Campana, 950 m, 5. vi. 95, R. Anderson, # 17834 (4 W) [MZSP]; same except # 17835 (3 W) [MZSP]; # 17836 (2 W) [MZSP]; # 17836 (3 W) [IHVL]; # 17836 (1 W) [USNM]; # 17836 (1 W) [DZUP]; # 17753 (7 W 1 Q) [MZSP]; # 17753 (2 W) [UTEP]; 900 m, # 17755 (2 W, one worker just petiole, postpetiole and gaster) (MYZH 219) [MZSP]; # 17756 (12 W 2 Q) [MZSP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9EFF91F0A5F03DFA53FA24.taxon	description	Figures 39, 40, 80 D, 81 E, 84 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9EFF91F0A5F03DFA53FA24.taxon	materials_examined	Holotype: ECUADOR: Pichincha: 4 km E Santo Domingo de los Colorados, 8. vii. 1976, J. & J. Peck [leg.] (1 W) (MCZENT 00524651) [MCZC]. Paratypes: same locality, [no date and collector], (1 Q) (MCZENT 00524649) [MCZC]; same data as holotype (1 W) (MCZENT 00524652 MZSP 67343) [MZSP]; (1 Q) (MCZENT 00524647 MZSP 67345) [MZSP]; (3 W 3 Q) (MCZENT 00524641, MCZENT 00524642, MCZENT 00524643, MCZENT 00524646, MCZENT 00524648, MCZENT 00524650) [MCZC]; same except 520 m, 22 June 1975, S. & J. Peck [leg.], rainfor., B- 304 (1 W) (MCZENT 00524670) [CASC]; (1 W) (MCZENT 00524667) [DZUP]; (1 W) (MCZENT 00524669) [IHVL]; (7 W 1 Q) (MCZENT 00524654, MCZENT 00524655, MCZENT 00524656, MCZENT 00524658, MCZENT 00524660, MCZENT 00524661, MCZENT 00524663, MCZENT 00524668) [MCZC]; (1 W) (MCZENT 00524653 MZSP 67346) [MZSP]; (1 W, one covered with gold) (MCZENT 00524659 MZSP 67347) [MZSP]; (1 Q) (MCZENT 00524662 MZSP 67344) [MZSP]; (1 W) (MCZENT 00524666) [USNM]; 4 km SE Santo Domingo, 1976, for., litter, berl. 42, S. & J. Peck [leg.] (9 W) (MCZENT 00524649) [MCZC]; same data (1 W) (MZCENT 00525534 MZSP 67349) [MZSP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9EFF91F0A5F03DFA53FA24.taxon	diagnosis	Diagnosis. Vermicular striae on head dorsum diverge towards posterior margin, interspaces between thicker striae filled with thinner striae; mesosoma covered with irregular to vermicular striae of variable thickness, interspaces indistinguishable; dorsal and ventral teeth of propodeal lobe blunt; mesosoma notably convex, metanotal groove discernible by a slight depression (LV) and altered sculpture (DV); dorsal margin of petiole continuous, convex, with irregular, longitudinal and anastomosed striae on dorsum; postpetiole smooth; subpostpetiolar process smooth, weak, convex; profemur posterior surface and protibia extensor surface smooth; striation on tergum of first gastral segment restricted to base; striae very short.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9EFF91F0A5F03DFA53FA24.taxon	description	Description. WORKER (n = 6) (Fig. 39 A – C): HL (0.94 – 1.04); HW (0.88 – 0.96); ML (0.60 – 0.68); SL (0.66 – 0.72); MOD (0.20 – 0.23); PNW (0.61 – 0.68); WL (1.20 – 1.36); PSL (0.30 – 0.34); PL (0.62 – 0.73); PW (0.24 – 0.26); PPL (0.32 – 0.36); PPW (0.33 – 0.36); GL (1.04 – 1.08); TL (4.77 – 5.21); CI (88.46 – 93.61); SI (70.83 – 78.26); OI (22.72 – 24.44). Midsized. Shiny integument. Brownish body, yellowish leg (except for coxa) and antenna. Thin and branched setae, midsized to short, suberect to subdecumbent, 2 short branches of equal size diverging from the main axis (Fig. 80 D). Head subquadrate; posterior margin slightly concave at middle. Mandible masticatory margin with 6 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with 7 – 8 irregular, longitudinal and thick striae converging to a point on the anterior margin, interspaces between thicker striae filled with thinner striae. Frontal triangle with 1 sagittal stria. Short scape, not reaching head posterior margin; apical antennomere slightly shorter than previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye drop-shaped, midsized, larger diameter with 11 ommatidia. Vermicular and longitudinal striae on head dorsum, divergent towards posterior margin, interspaces between thicker striae filled with thinner striae. Head lateral and laterodorsal regions with vermicular striae converging to eye margin; gena striate, 3 irregular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma covered with vermicular striae of variable thickness, interspaces indistinguishable; mostly transverse striae on pronotum anterior region (DV), striation continues on lateral surface; semicircular striae on mesonotum; striae on propodeum lateral directed mainly to propodeum dorsum. Dorsal margin of mesosoma slightly discontinuous, notably convex. Promesonotal junction indistinct. Transverse carina inconspicuous. Metanotal groove discernible by a slight depression (LV) and altered sculpture (DV). Propodeal lobe bidentate, dorsal tooth slightly longer than ventral, both blunt; dorsal tooth length 1 / 3 shorter than propodeal spine length. Propodeal spine long, straight (LV), divergent (DV), sculptured on base. Procoxa striae of variable thickness; irregular and transverse striae on C 2 and C 3. Profemur mostly smooth. Protibia smooth. Dorsal margin of petiole continuous, convex; transverse striae on ventral surface from petiole spiracle until posterior end of petiole; first third of dorsal surface smooth; lateral and posterodorsal surfaces mostly with irregular longitudinal and anastomosed striae. Postpetiole smooth; subpostpetiolar process smooth, weak, convex. Longitudinal striae on tergum first gastral segment, striation short, weakly marked, restricted to base, striae length equivalent to 1 / 4 of postpetiole length; sternite smooth. QUEEN (n = 3) (Fig. 40 A – C): HL (1 – 1.06); HW (0.94 – 0.98); ML (0.63 – 0.70); SL (0.72 – 0.80); MOD (0.24 – 0.26); PNW (0.84 – 0.88); WL (1.44 – 1.54); PSL (0.32 – 0.36); PL (0.70 – 0.76); PW (0.29 – 0.30); PPL (0.34 – 0.40); PPW (0.40 – 0.42); GL (1.20 – 1.32); TL (5.34 – 5.78); CI (92.45 – 94.11); SI (75 – 81.63); OI (25.53 – 26.53). Largesized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 13 – 14 ommatidia. Longitudinal, regular to irregular striae on scutum going from an anterior central point towards transcutal suture, interspaces indistinguishable. Longitudinal, regular to irregular striae on anepisternum and katepisternum, some vermicular striae on katepisternum, thinner striae between thicker striae. Axilla and scutellum with same sculpture of scutum. Scutoscutellar sulcus inconspicuous. Transverse striae on propodeum (DV). Lateral surface of mesosoma with vermicular striae directed to propodeal spine. Wings as in Fig. 81 E. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9EFF91F0A5F03DFA53FA24.taxon	etymology	Etymology. The epithet lispectorae is a Latin noun in the genitive case created by adding the singular Latin genitive case suffix - ae to the last name of a female person. The specific epithet is named after Clarice Lispector (1920 – 1977), born Chaya Pinkhasovna Lispector, a Ukrainian-Brazilian novelist, poetess, and short story writer — “ Liberdade é pouco. O que eu desejo ainda não tem nome ”.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9EFF91F0A5F03DFA53FA24.taxon	discussion	Comments. This species is easily distinguished from most of its congeners, but it is very similar to H. montana regarding the shape of mesosoma, and the sculpture and shape of the procoxa, petiole, postpetiole, and gaster. Also, the two species are sympatric, co-occuring in two Ecuadorian localities: Centro Científico Río Palenque, and 4 km E of Santo Domingo de los Colorados (Figs. 83, 84). Hylomyrma lispectorae can be distinguished from H. montana (characteristic in parentheses) in the dorsum of head and mesosoma by the vermicular striae of variable thickness with indistinguishable interspaces (vs. irregular striae of uniform thickness with distinguishable interspaces), and the notably convex dorsal margin of mesosoma (vs. somewhat straight).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9EFF91F0A5F03DFA53FA24.taxon	distribution	Distribution. Hylomyrma lispectorae is known from Colombia and Ecuador (Fig. 84). Natural history. This species occurs in tropical rainforest areas at altitudes below 600 m. Specimens were collected in leaf-litter with winkler extractors and Berlese-Tüllgren funnels, which suggests that nests are located in fallen logs, rotten wood, between leaves, or inside natural cavities of the superficial soil layers.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF9EFF91F0A5F03DFA53FA24.taxon	materials_examined	Additional material examined (15 workers, 4 queens): COLOMBIA: Valle del Cauca [Cauca]: PNN Gorgona, Playa Palmera, W 78 ° 12 ′ 21.4 ″, N 2 ° 56 ′ 7.9 ″, 28 msnm, 24 - 26. ii. 2011, Stephany Valdes col., winkler (2 W) (MZSP 67437, MZHY 217) [MZSP]. ECUADOR: Los Rios: C. C. R. Palenque, 79 ° 45 ′ 10 ″ W, 01 ° 25 ′ 56 ″ S, 10 JUN 1980, S. Sandoval, QCAZ I 57121 (1 W) [MZSP]; same except 01 JUN 1980, QCAZ I 114014 (1 Q) [UTPL]; 29 MAY 1980, T. DeVries, QCAZ I 114005 (1 Q) [QCAZ]; 29 m, 05 MAR 1979, T. DeVries, QCAZ I 57123 (1 W) [QCAZ]; QCAZ I 114006 (1 W) [QCAZ]; QCAZ I 114007 (1 W) [MZSP]; QCAZ I 114002 (1 W) [QCAZ]; QCAZ I 114004 (1 W) [QCAZ]; 02 MAR 1979, QCAZ I 114009 (1 W) [MZSP]; QCAZ I 114013 (1 Q) [MZSP]; 29 MAY 1980, QCAZ I 114003 (1 W) [QCAZ]; 220 m, 79 ° 00 ′ W, 00 ° 54 ′ S, 24 FEB 1977, QCAZ I 57124 (1 W) [MZSP]; 79 ° 45 ′ 10 ″ W, 01 ° 25 ′ 56 ″ S, 29 m, 24 FEB 1977, QCAZ I 114010 (1 W) [MZSP]; 01 MAR 1979, QCAZ I 57122 (1 W) [MZSP]; 2 FEB 1977, QCAZ I 114011 (1 W) [QCAZ]; 47 km S Sto. Domingo, Rio Palenque Sta. (1 Q) (MCZENT 00525485) [MCZC]; Namabí: 78 km NE Chone, 450 m, 9. vi. 1976, S. & J. Peck, B- 345 (1 W) (MCZENT 00525491) [MCZC].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF91FF96F0A5F211FD03FC30.taxon	description	Figures 41, 42, 43, 80 I, 81 D, 82 D, 83 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF91FF96F0A5F211FD03FC30.taxon	diagnosis	Diagnosis. Regular and longitudinal striae on head dorsum, parallel towards posterior margin; long scape; vermicular striae on mesosoma; dorsal margin of petiole discontinuous; ventral surface entirely covered with transverse striae; postpetiole with regular and longitudinal striae slightly anastomosed and thick; subpostpetiolar process sculptured on laterobasal region, midbasal region smooth; long striae on tergum of first gastral segment; body with many branched setae; branches of equal size; unbranched setae predominate in the gaster.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF91FF96F0A5F211FD03FC30.taxon	description	Redescription. WORKER (n = 6) (Fig. 41 A – C): HL 0.92 (0.85 – 0.94); HW 0.80 (0.77 – 0.82); ML 0.58 (0.54 – 0.60); SL 0.81 (0.74 – 0.85); MOD 0.25 (0.22 – 0.28); PNW 0.58 (0.54 – 0.60); WL 1.20 (1.10 – 1.22); PSL 0.28 (0.25 – 0.32); PL 0.54 (0.50 – 0.58); PW 0.18 (0.18 – 0.20); PPL 0.29 (0.28 – 0.32); PPW 0.29 (0.26 – 0.30); GL 1.12 (1.04 – 1.12); TL 4.65 (4.33 – 4.78); CI 86.95 (86.95 – 90.58); SI 101.25 (95 – 103.66); OI 31.25 (28.20 – 34.14). Small to midsized. Shiny integument. Bicolored body, brownish with lighter leg. Many branched setae, midsized to short, branches of equal size (Fig. 80 I), suberect to subdecumbent; unbranched setae predominate in the gaster. Head subquadrate; posterior margin straight. Mandible masticatory margin with 5 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with 8 – 10 regular and longitudinal striae converging to a point on the anterior margin, interspaces distinguishable. Frontal triangle with 2 striae. Long scape, surpassing head posterior margin; apical antennomere slightly shorter than previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye drop-shaped, midsized, larger diameter with 11 ommatidia. Regular and longitudinal striae on head dorsum, parallel towards posterior margin, interspaces between thicker striae filled with thinner striae, anastomosed striae on posterior region. Sculpture on head lateral and laterodorsal regions converge to eye margin; laterodorsal region predominantly with longitudinal striae; lateral and lateroventral region mainly with anastomosed striae; gena striate, 6 – 8 regular and semicircular striae circumscribe the torulus, almost reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma covered with vermicular striae of uniform thickness, interspaces smooth. Promesonotal junction distinguishable by a slight depression (LV); metanotal groove indistinct. Transverse carina well-marked. Dorsal margin of mesosoma discontinuous, convex. Propodeal lobe bidentate, dorsal tooth slightly longer and sharper than shorter and blunt ventral tooth; dorsal tooth shorter than 1 / 3 of propodeal spine length. Propodeal spine long, straight (LV), divergent (DV), sculptured on base. Procoxa striae of uniform thickness; irregular and transverse striae on C 2 and C 3. Profemur and protibia smooth. Dorsal margin of petiole discontinuous; ventral surface entirely covered with transverse striae. Node with regular and transverse striae on anterior surface, striation on anterior surface continuing transversely on lateral surface to ventral surface, weakly marked; vermicular striae on dorsal surface. Postpetiole covered with regular and longitudinal striae slightly anastomosed and thick; subpostpetiolar process with same sculpture of postpetiole on laterobasal region, but weakly marked, midbasal region smooth; subpostpetiolar process weak, straight. First gastral segment striation similar to postpetiole striae, longitudinal striae on tergum 11 / 2 × the postpetiole length; sternite striation restricted to laterobasal region, weakly marked. QUEEN (first description) (n = 3) (Fig. 42 A – C): HL (0.86 – 0.94); HW (0.80 – 0.84); ML (0.59 – 0.60); SL (0.76 – 0.78); MOD (0.24 – 0.27); PNW (0.62 – 0.67); WL (1.28 – 1.34); PSL (0.28 – 0.30); PL (0.57 – 0.62); PW (0.19 – 0.22); PPL (0.32 – 0.33); PPW (0.30 – 0.32); GL (1.17 – 1.20); TL (4.76 – 5); CI (89.13 – 93.02); SI (92.68 – 97.50); OI (30 – 32.92). Medium to large-sized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 14 – 15 ommatidia. Striae on scutum anastomosed anteriorly and irregular posteriorly, going from an anterior central point towards transcutal suture. Longitudinal and regular striae on anepisternum and irregular to vermicular on katepisternum, interspaces distinguishable. Axilla and scutellum with same sculpture of scutum. Scutoscutellar sulcus inconspicuous. Transverse striae on propodeum (DV). Lateral surface of mesosoma with irregular to vermicular striae directed in part to propodeal dorsum and in part to propodeal spine. Wings as in Fig. 81 D. MALE (first description) (n = 1) (Fig. 43 A – C): HL 0.66; HW 0.79; ML 0.46; SL 0.28; MOD 0.32; PNW 0.72: CMS 1.33; PSL 0.08; PL 0.60; PW 0.22; PPL 0.34; PPW 0.32; GL 1.34; TL 4.73; CI 119.70; SI 35.44; OI 40.50. Light brown body, yellowish appendices. Mandible with 6 teeth. Scutum anterior region smooth, posterior region covered with mostly longitudinal, regular to irregular striae, interspaces distinguishable. Anepisternum and katepisternum covered with longitudinal and regular striae, thinner than those on scutum, interspaces distinguishable. Scutellum with mostly irregular striae, thicker than scutum striae, interspaces distinguishable. Propodeum with irregular striae assuming multiple directions, interspaces distinguishable. Propodeal lobe rounded. Petiole mostly covered with longitudinal and irregular striae, interspaces distinguishable. Postpetiole with longitudinal and regular striae. Gaster smooth. Wings as in Fig. 82 D.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF91FF96F0A5F211FD03FC30.taxon	etymology	Etymology. The specific epithet longiscapa (Latin, longus = long and scapus = stem) refers to the relatively long antennal scape of this species.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF91FF96F0A5F211FD03FC30.taxon	discussion	Comments. Hylomyrma longiscapa is similar to H. marielleae regarding the body size and the sculpture pattern. However, H. longiscapa can be easily distinguished from H. marielleae (characteristic in parentheses) in the thin and light setae (vs. thick and dark), and the transverse striae on the ventral surface of petiole well-marked (vs. weakly marked). Both species have broad distribution in the center-north of South America, however H. longiscapa occurs in many localities in eastern portions of Brazil (from northeast to southeast regions) (Fig. 83) where H. marielleae does not occur (Fig. 85). The type specimens (Surinam) have one central ocellus. This character, together with the larger body size [HL (0.92 – 0.98); HW (0.80 – 0.86); ML (0.58 – 0.66); SL (0.76 – 0.90); MOD (0.23 – 0.28); PNW (0.58 – 0.64): WL (1.16 – 1.30); PSL (0.28 – 0.32); PL (0.56 – 0.60); PW (0.18 – 0.23); PPL (0.33 – 0.36); PPW (0.29 – 0.32); GL (1.06 – 1.20); TL (4.65 – 5.08); CI (86.95 – 89.58); SI (95 – 104.65); OI (28.75 – 32.55)], and the promesonotal junction discernible by a more pronounced depression, likely indicate the specimens are intercastes. Intercastes were also sampled in Brazil, Ecuador, Peru, and Venezuela. Specimens from Brazil (Novo Jardim, TO) have a spine on the mesoventral surface of petiole.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF91FF96F0A5F211FD03FC30.taxon	distribution	Distribution. This species occurs from central to northern South America (Brazil, Ecuador, French Guiana, Peru, and Venezuela) (Fig. 83). The distribution of H. longiscapa is broader than that of H. marielleae, especially because the distribution of the former spans the Atlantic forest in Brazil, while the latter is more restricted to the Amazonian region. Both species occur in Madre de Dios, Peru; Napo, Ecuador; and Mato Grosso, Pará, and Tocantins, all three in Brazil. Natural history. This species inhabits forested areas in the Atlantic forest (Northeast Brazil), tropical rainforest (Amazon), “ Cerrado ” (Brazilian savanna), and semideciduous forest enclaves in the Atlantic forest biome (Minas Gerais, Brazil). Specimens are frequently found in leaf-litter samples, which suggests that this species nests in the leaf-litter, fallen logs, rotten wood, or inside natural cavities of the superficial soil layers. Except for the presence of intercastes, there is no further information available regarding the biology of H. longiscapa.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF91FF96F0A5F211FD03FC30.taxon	materials_examined	Additional material examined (104 workers, 12 queens, 1 male, 13 intercastes): BRAZIL: BA [Bahia]: Aurelino Leal, L. do Banco, Mata, 26 / 05 / 1997, Col. J. C. S. Carmo & J. R. M. Santos, 637 (1 W 1 Q 1 M) (MZSP 67353) [MZSP]; Andaraí, Mata Carrasco (Castanha), 13 – 14. xii. 1990, Brandão, Diniz & Oliveira, folhiço (8 W, one covered with gold) (MZSP 67436) [MZSP]; same except 105 (1 Q) [MZSP]; 104 (1 W 3 Q 2 I) [MZSP]; Ilhéus-Castelo Novo, 26.04. [20] 00, 143912 S, 391139 W, Santos, J. R. M. dos, Mata-A 24 (3 W) [CEPLAC]; Ilhéus-Banco do Pedro, 12.01. [19] 98, 144051 S, 0391524 W, Santos J. R. M. dos, Carmo J. C. S. do, Mata W-A 23, 61 (3 W) [CEPLAC]; Ilhéus, CEPEC, 10. [19] 96, Santos, J. R. M., # 5156 (5 W) [CEPLAC]; same except 27.01. [19] 95, col. D. Agosti, MZSP 67352 (1 W 1 Q) [MZSP]; km 22 Ilhéus-Itabuna, CEPED-Área Zoolog., x. [19] 86, J. Delabie leg., 61 (1 W) [MZSP]; Mata da Boa Esperança, 14 ° 47 ′ 47 ″ S, 09 ° 03 ′ 56 ″ W, 09. ix. 2000, Santos J. R. M. col., Winkler 28 (1 W) [MZSP]; Itapebí, M. Rio Pardo, 16.07. [19] 97, Santos J. R. M. (3 W) [CEPLAC]; Ibicaraí, km 41, 145375 S, 0392901 W, 21.11. [19] 98, Santos J. R. M. dos, 61 (1 W 1 Q) [CEPLAC]; Itacaré-Taboquinha, 20 / 12 / [19] 96, Santos J. R. M. dos, 61 (3 W 1 Q) [CEPLAC]; Itapitanga, 30.05.97, 14 ° 25 ′ 22 ″ S, 39 ° 33 ′ 54 ″ W, Santos J. R. M dos, 61 (1 W) [CEPLAC]; Maraú-Tremenbé, 07.06. [19] 97, Santos J. R. M dos, MATA-W-A 4, 61 (3 W) [CEPLAC]; Arataca, 25.05. [19] 99, 151649 S, 0392331 W, Santos J. R. M dos, Mata A 47, 61 (1 W) [CEPLAC]; Mata São João, Reserva Sapiranga, 12 ° 33 ′ 29.3 ″ S, 33 ° 02 ′ 35.2 ″ W, 21 - 28. vii. 2001, Silva R. R., Brandão C. R. F. cols., Winkler 49 (1 W) [MZSP]; same except Winkler 43 (3 W) [MZSP]; Winkler 46 (1 W) [MZSP]; Winkler 34 (1 Q) [MZSP]; Winkler 51 (1 W) [MZSP]; Serra Grande, 14 ° 27 ′ 05 ″ S, 39 ° 02 ′ 34 ″ W, 28 / 04 / 1997, Carmo J. C. S., 61 (3 W) [CEPLAC]; Una, 151104 S, 0390056 W, 24.08. [19] 98, Santos J. R. M. dos, A 45, 61 (2 W) [CEPLAC]; Unacau, 150521 S, 39 ° 1742 W, 11.02.2000, Santos J. R. dos, A 43, 61 (3 W) [CEPLAC]; Lençóis, Chap [ada] Diamantina, 25. iii. 2001, Santos J. R. M., 5, 61 (2 W) [CEPLAC]; Coaraci, 14 ° 38 ′ S, 39 ° 33 ′ W, 30 / 09 / 2002, Santos J. R. M. Dos, Mata, Winkler (1 W 2 I) [CEPLAC]; GO [Goiás]: Serranópolis, Faz. São Cristovão, 817 m, 18 ° 5 ′ 32.87 ″ S, 52 ° 2 ′ 23.85 ″ W, 10. i. 2009, Gilmar G. Santos col., Frag. 03, M. W. Ponto 23 (9 W) [DZUP]; same except Ponto 06 (6 W) [DZUP]; Jataí, Faz. Sta. Gerturdes, 876 m, 17 ° 50 ′ 07 ″ S, 51 ° 43 ′ 04 ″ W, 02. ii. 2009, G. G. Santos col., Frag. 01, M. W. Ponto 13 (1 W) [DZUP]; same except Ponto 19 (2 W) [DZUP]; MA [Maranhão]: Açailândia, Horto Faz. Pompéia, 04 ° 52 ′ 30 ″ S, 47 ° 17 ′ 40 ″ W, 13 - 22. ii. 2006, Silva R. R. & Feitosa R. M. cols., Winkler 17 (1 W) [MZSP]; [MG] Minas Gerais: Parque Estadual do Rio Doce, Trilha da Garapa Torta, Elev. 280 m, 19 ° 47 ′ 49 ″ S, 42 ° 34 ′ 38 ″ W, 23 - 24. viii. 2005, TEAM exped. Col., Floresta Atlântica Etacional Semidecidual, TGT 4.7 Mini-winkler (1 W) [MZSP]; Marlieria, Parque Estadual do Rio Doce, Trilha da Lagoa Preta, 280 m, - 19.796944, - 42.577222, 30. xi. 2005, TEAM exped. Col., Floresta Atlântica Estacional Semidecidual, LP 2 - 6 (1 W) [MZSP]; same except 25. viii. 2005, LP 1 - 6 (1 W) [MZSP]; MT [Mato Grosso]: Claudia, Acampamento módulo 1, iv. 2014, Vicente, R. E., coleta manual (1 W) [MZSP]; PA [Pará]: Novo Repartimento, Faz. Aratau, 25 - 26. vi. 2003, A. M. Elizabeth (1 W 2 I) [CEPLAC]; PE [Pernambuco]: Recife, Horto Dois Irmãos, 08 ° 00 ′ 32 ″ S, 34 ° 56 ′ 40 ″ W, 15 - 24. vii. 2002, Silva R. R. & Eberhardt F. cols., Winkler 11 (1 W) [MZSP]; same except Winkler 8 (1 W) [MZSP]; Winkler 49 (1 W) [MZSP]; Winkler 46 (1 W) [MZSP]; RO [Rondônia]: Porto Velho, Área Mutum, M 5 P 4, 09 ° 35 ′ 29.5 S, 65 ° 02 ′ 57.6 ″ W, 04 - 18. ix. 2012, Ulysséa, M. A. & Prado, L. P. cols. (2 W) [MZSP]; SE [Sergipe]: Sta. Luiza do Itanhy, Crasto, 11 ° 22 ′ 39.3 ″ S, 37 ° 25 ′ 07.4 ″ W, 29. vii- 03. viii. 2001, Silva R. R., Brandão C. R. F., Winkler 30 (5 W) [MZSP]; same except Winkler grande 18 a (2 W) [MZSP]; Areia Branca, E. E. da Serra de Itabaiana, 10 ° 45 ′ 54 ″ S, 37 ° 19 ′ 57.4 ″ W, 19 - 25. v. 2003, Silva R. R., Dietz B. H. & Ferreira L. S. cols., Winkler 29 (2 W) [MZSP]; same except Winkler 46 (1 W) [MZSP]; Winkler 24 (2 W 1 Q) (MZSP 67351) [MZSP]; Winkler 29 (1 W 1 Q) [MZSP]; Winkler 36 (1 Q) [MZSP]; TO [Tocantins]: Goiatins, 07 ° 56 ′ 28.9 ″ S, 47 ° 09 ′ 31.3 ″ W, 03 - 08. v. 05, Silva R. R. & Dietz B. H., Winkler 12 (1 W) [MZSP]; same except Winkler 11 (1 W 1 I) [MZSP]; Novo Jardim, 10 ° 50 ′ 26.9 ″ S, 46 ° 41 ′ 11.7 ″ W, 09. x. 2004, Vereda, Silva R. R. & Dietz B. H., Winkler 26 (2 I) [MZSP]. ECUADOR: Napo: Limoncocha, 280 m, 00 ° 24 ′ S, 76 ° 36 ′ W, 4. viii. 1973, Lois Morales, 237, Colony F- 563 (1 W 1 I) [MZSP]. GUYANE FRANÇAISE: [Saint-Laurent-du-Maroni: Maripasoula], Mitaraka Mts, Pente, 2.238226 N - 54.45203 W, 04. iii. 2015, J. Orivel & F. Petitclerc cols., winkler 48 h, A-P 2 - 8, MI 15 - 0380 - 26 (1 W) [ECOFOG]. PERU: Madre de Dios: Los Amigos Field Station, Trail 9 at 725 m mark, 291 m, 70 ° 6 ′ 17.39 W, 12 ° 34 ′ 26.69 S, 2. x. 2004, T. R. Schultz, C. Marshall, J. Sosa-Calvo, 1 ° forest, litter sample, = CJM 041002 - 04, TRS 041002 - 01 - LS 04 (1 W 1 I) [USNM]; same except = CJM 041002 - 06, TRS 041002 - 01 - LS 06 (1 I) [USNM]. VENEZUELA: Zulia: Dto. Perija, El Tucuco, 6. ix. 1984, cols. J. L attke, E. Rubio (1 W) [MIZA]; El Tucuco, 51 km SO de Machiques, 24. vi. 1979, R. W. Brooks, A. A. Grigarick, J. McLaughlin, R. O. Schuster (1 I) [MIZA].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF96FFEBF0A5F025FDECF958.taxon	description	Figures 2 B, 44, 45, 80 E, 80 J, 89 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF96FFEBF0A5F025FDECF958.taxon	materials_examined	Holotype: BRAZIL: MA [Maranhão]: Estreito, Fazenda Itaueiras, 06 ° 31 ′ 54.4 ″ S, 47 ° 22 ′ 16 ″ W, 07 - 13. i. 2005, Silva R. R. & Silvestre R. cols., Winkler 25, MZSP 67354 (1 W) [MZSP]. Paratypes: same data as holotype (1 W) (MZSP 67355) [MZSP]; same except Winkler 36 (1 Q) (MZSP 67356) [MZSP]; Winkler 34 (1 W) (MZSP 67357) [MZSP]; 01 - 09. vi. 2005, Silva R. R. & Feitosa R. M. cols., Winkler 23 (1 W) (MZSP 67358) [MZSP]; Winkler 14 (1 W) (MZSP 67359) [MZSP]; Winkler 1 (1 W) (MZSP 67360) [MZSP]; Winkler 8 (1 W) (MZSP 67361) [MZSP]; Winkler 5 (1 W) (MZSP 67362) [MZSP]; Winkler 2 (1 W) (MZSP 67363) [MZSP]; Winkler 4 (1 W) (MZSP 67364) [MZSP]; Winkler 24 (1 W) (MZSP 67365) [MZSP]; 12 - 22. vi. 2006, Silva R. R. & Feitosa R. M. cols., Winkler (2 W) [USNM]; Winkler 11 (1 W) [DZUP]; Winkler 15 (1 W) [DZUP]; Winkler 10 (1 W) [MCZC]; Winkler 07 (1 W) [MCZC]; Winkler 06 (2 W) [IHVL]; Winkler 14 (1 W 1 Q) [CASC]; 12 - 22. vi. 2006, Winkler, Resíduo 2 (1 W 1 Q) (MZSP 67366) [MZSP]; 13 - 22. ii. 2006, Winkler, Resíduo (1 W 1 Q) (MZSP 67367) [MZSP]; Estreito, Fazenda Planalto, 06 ° 35 ′ 59.3 ″ S, 47 ° 24 ′ 50.4 ″ W, 07 - 13. i. 2005, Silva R. R. & Silvestre R. cols., Winkler 8 (1 W) (MZSP 67368) [MZSP]; same except Winkler 10 (1 W) (MZSP 67369) [MZSP]; Winkler 5 (1 W) (MZSP 67370) [MZSP]; Winkler 4 (1 W) (MZSP 67371) [MZSP]; Winkler 2 (1 W 1 Q) (MZSP 67372) [MZSP]; 01 - 09. vi. 2005, Silva R. R. & Feitosa R. M. cols., Winkler 1 (1 W) (MZSP 67373) [MZSP]; Winkler 2 (1 W) (MZSP 67374) [MZSP]; Winkler 5 (1 W) (MZSP 67375) [MZSP]; Winkler 8 (2 W) (MZSP 67376) [MZSP]; Winkler 9 (1 W) (MZSP 67377) [MZSP]; Winkler 3 (1 W) (MZSP 67378) [MZSP]; 12 - 22. vi. 2006, Silva R. R. & Feitosa R. M. cols., Winkler (2 W 1 Q) (MZSP 67379) [MZSP]; 13 - 22. ii. 2006, Silva R. R. & Feitosa R. M. cols., Winkler 8 (1 W) (MZSP 67380) [MZSP]; Winkler 11 (2 W 1 Q) (MZSP 67381) [MZSP]; Winkler 06 (1 W 1 Q) (MZSP 67382) [MZSP]; Winkler 12 (1 W) (MZSP 67383) [MZSP]; Winkler 10 (1 W) (MZSP 67384) [MZSP]; SP [São Paulo]: Matão, Fazenda Cambuhy, Mata da Virgínia, Winkler, 21 ° 37 ′ 35.5 ″ S, 48 ° 33 ′ 24.8 ″ W, 17 - 22. iv. 2015, M. A. Ulysséa & L. P. Prado cols., A 1 P 3 W 3 (1 W covered with gold) (MZSP 67385) [MZSP]; 21 ° 37 ′ 33.56 ″ S, 48 ° 32 ′ 13.11 ″ W, 30. iii- 03. iv. 2016, M. A. Ulysséa col., A 1 P 2 W 4 (1 Q) (MZSP 67386) [MZSP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF96FFEBF0A5F025FDECF958.taxon	diagnosis	Diagnosis. Regular and longitudinal striae on head dorsum diverge towards posterior margin; mesosoma covered with regular, longitudinal and thick striae, interspaces distinguishable; striae on lateral of propodeum directed to propodeal spine; dorsal margin of petiole discontinuous; anterior and ventral surfaces of node transversely striate; longitudinal, regular to irregular striae on node dorsum and lateral surfaces, striae more straight on dorsum and sinusoid on lateral surface, interspaces distinguishable; subpostpetiolar process striate; profemur posterior surface mostly smooth; striae on protibiae extensor surface weakly marked; striation on tergum of first gastral segment restricted to base; branched setae of two types: 1) 2 – 3 short, thin and equal-sized branches diverging from the main axis; 2) thick setae flattened in its final half, with several branches.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF96FFEBF0A5F025FDECF958.taxon	description	Description. WORKER (n = 8) (Fig. 44 A – C): HL (0.80 – 0.90); HW (0.79 – 0.88); ML (0.52 – 0.55); SL (0.57 – 0.64); MOD (0.22 – 0.26); PNW (0.54 – 0.63); WL (1.08 – 1.20); PSL (0.16 – 0.19); PL (0.52 – 0.58); PW (0.24 – 0.28); PPL (0.30 – 0.34); PPW (0.33 – 0.38); GL (0.96 – 1.14); TL (4.21 – 4.53); CI (93.18 – 98.75); SI (65.90 – 78.04); OI (25.58 – 32.91). Small-sized. Shiny integument. Brownish body, lighter appendices; petiole, postpetiole and gaster darker. Mostly thin setae, unbranched or branched, midsized to short, erect to decumbent; branched setae of two types: 1) 2 – 3 short, thin and equal-sized branches diverging from the main axis (Fig. 80 E); 2) thick setae flattened in its final half, with several branches, mainly in the postpetiole and gaster (Fig. 80 J). Head subquadrate; posterior margin slightly concave at middle. Mandible masticatory margin with 5 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with 7 – 9 regular to irregular, longitudinal and thick striae converging to a point on the anterior margin, interspaces smooth or thinner striae between thicker striae. Frontal triangle with 1 sagittal stria. Short scape, not reaching head posterior margin; apical antennomere as long as previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye reniform, midsized, larger diameter with 12 ommatidia. Regular, longitudinal striae on head dorsum, diverging towards posterior margin, interspaces between striae filled with thinner regular to irregular striae. Head lateral and laterodorsal regions with same striation of head dorsum, striae converging to eye margin; striation connecting the region posterior to frontal carina with eye margin, striae irregular, not uniform in thickness; gena striate, 4 regular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma covered with regular and longitudinal striae, thicker than head striation, interspaces distinguishable; striae transverse on pronotum anterior region (DV) and longitudinal in other regions (DV); lateral of pronotum, mesepisternum and propodeum with longitudinal striae directed to propodeal spine. Promesonotal junction and metanotal groove indistinct. Transverse carina inconspicuous (DV). Dorsal margin of mesosoma continuous, convex. Propodeal lobe bidentate, dorsal tooth longer and acute than shorter and blunt ventral tooth; dorsal tooth length similar to propodeal spine length. Propodeal spine short, straight (LV), divergent (DV), sculptured on base. Transverse striae of uniform thickness on procoxa; irregular and transverse striae on C 2 and C 3. Profemur mostly smooth. Protibia extensor surface entirely striate, striae weakly marked. Dorsal margin of petiole discontinuous. Node with anterior and ventral surfaces striate transversely, striae regular to irregular, interspaces distinguishable; longitudinal, regular to irregular striae on dorsal and lateral surfaces, straight on dorsum, sinusoid on lateral. Postpetiole and subpostpetiolar process with regular, longitudinal striae; subpostpetiolar process prominent, convex. First gastral segment striation longitudinal and thinner than postpetiole striae; striae length on tergum shorter than 1 / 3 of postpetiole length; sternite striation weakly marked, covering the laterobasal region. QUEEN (n = 3) (Fig. 45 A – C): HL (0.85 – 0.88); HW (0.82 – 0.84); ML (0.52 – 0.55); SL (0.54 – 0.57); MOD (0.26 – 0.28); PNW (0.64 – 0.68); WL (1.22 – 1.26); PSL (0.20 – 0.22); PL (0.58 – 0.64); PW (0.26 – 0.28); PPL (0.35 – 0.36); PPW (0.37 – 0.39); GL (1.12 – 1.20); TL (4.71 – 4.80); CI (94.32 – 96.47); SI (64.28 – 69.47); OI (31.70 – 33.73). Midsized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 14 ommatidia. Longitudinal and regular to irregular striae on scutum going from an anterior central point towards transcutal suture, interspaces distinguishable. Longitudinal and regular striae on anepisternum and katepisternum, interspaces distinguishable. Axilla with the same sculpture of scutum. Scutoscutellar sulcus inconspicuous. Scutellum with the same sculpture of scutum. Transverse striae on propodeum (DV). Lateral surface of mesosoma with longitudinal and regular striae directed to propodeal spine, interspaces distinguishable. Wings unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF96FFEBF0A5F025FDECF958.taxon	etymology	Etymology. The epithet lopesi is a Latin noun in the genitive case created by adding the singular Latin genitive case suffix - i to the last name of a male person. This species is named in honor of Dr. Benedito Cortês Lopes, M. A. Ulysséa’s first ‘ ant’ professor, for his many years of work on ants, as a Zoology and Entomology professor, and as a scientific supervisor at the Universidade Federal de Santa Catarina.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF96FFEBF0A5F025FDECF958.taxon	discussion	Comments. This species is similar to H. balzani, H. reitteri, and H. margaridae. Hylomyrma lopesi can be misidentified as H. reitteri because of the sculpture on the mesosomal dorsum, and the shape of the dorsal margin of petiole. However, H. lopesi differs from H. reitteri (characteristic in parentheses) in the smaller body size (vs. larger), the thick striae on the mesosomal dorsum (vs. thin striae), the transverse striae on the ventral surface of petiole (vs. smooth surface), the laterals of the mesosoma covered mainly with regular and thick striae that are directed to the propodeal spine (vs. irregular and thin striae directed to the propodeal dorsum), and the striate subpostpetiolar process (vs. smooth). Both species occur broadly in Brazil, co-occurring in Botucatu (SP), Niquelândia (GO), and São José de Buerarema (BA). The distribution of H. reitteri is more circumscribed to eastern portions of Brazil from southern and southeast regions (Fig. 85), whereas H. lopesi is distributed mainly in the center of Brazil from northern (state of Pará) to southern (state of São Paulo) regions (Fig. 89). Specimens of H. lopesi have been identified as H. balzani, due to the similarity of the sculpture and shape of petiole, and based on the sculpture of gena. But both species can be easily distinguished by the rugose striae on the mesosoma of H. balzani and the regular striae on the mesosoma of H. lopesi. The distribution of these species cover a broad area in Brazil, both co-occurring in Botucatu (SP) and Milagres (BA). Hylomyrma lopesi is distributed mainly in the center of Brazil from northern (state of Pará) to southern (state of São Paulo) regions (Fig. 89), whereas H. balzani is more restricted to the eastern portions of Brazil from northeast to southern regions (Fig. 84). Hylomyrma lopesi and H. margaridae have similar shape and sculpture of mesosoma. Nevertheless, H. lopesi has reniform eye (vs. oval eye of H. margaridae), the striae on the head dorsum with smooth crests (vs. with punctuated crests), and the discontinuous dorsal margin of petiole (vs. continuous). The two species are allopatric (Fig. 89); H. lopesi occurs in Brazil, and H. margaridae in Guyana and French Guiana. There is considerable variation among Hylomyrma lopesi specimens. The striae on the petiolar node dorsum in specimens from several localities in Brazil (BA, GO, MA, MT, SP, and TO) are not exclusively longitudinal, but can also be transverse. Also, in specimens from Bahia, the striae on the laterals of the mesosoma are not entirely directed to the propodeal spine. In these specimens part of the striae on the laterals of the mesosoma are directed to propodeal dorsum. Besides, the 14 intercaste specimens present a larger phenotypic plasticity, sharing morphological traits with worker and winged queens. The modifications observed in these specimens are: a) the central ocellus and the strongly convex mesosoma [(n = 1), HL 0.85; HW 0.80; ML 0.52; PNW 0.60; WL 1.125; GL 0.87]; b) the central ocellus, the strongly convex mesosoma, and the presence of pre-transcutal suture [(n = 5), HL (0.82 – 0.95); HW (0.82 – 0.92); ML (0.52); PNW (0.55 – 0.65); WL (1.15 – 1.30); GL (0.95 – 1.15)]; c) the central ocellus, the presence of promesonotal suture, the strongly convex mesosoma, the presence of pre-transcutal suture [(n = 1), HL 0.85; HW 0.82; ML 0.55; PNW 0.60; WL 1.15; GL 1]; d) the three ocelli, the strongly convex mesosoma, the promesonotal junction discernible by a depression, and the presence of pre-transcutal suture [(n = 3), HL (0.87 – 0.92); HW (0.85 – 0.87); ML (0.55 – 0.57); PNW (0.60 – 0.65); WL (1.22); GL (0.85)]; e) the three ocelli, the strongly convex mesosoma, the distinguishable promesonotal junction and metanotal groove by a depression, and the presence of pre-transcutal suture [(n = 2), HL (0.90 – 0.92); HW (0.90); ML (0.55 – 0.57); PNW (0.65 – 0.67); WL (1.22 – 1.25); GL (1.07 – 1.10)]; f) the three ocelli, the enlarged mesosoma, the presence of promesonotal and pre-transcutal sutures, and the metanotal groove discernible by a depression [(n = 1), HL 0.90; HW 0.87; ML 0.55; PNW 0.57; WL 1.17; GL 1]; and g) the three ocelli, the enlarged mesosoma, the presence of promesonotal and transcutal sutures, and the metanotal groove discernible by a depression [(n = 1), HL 0.87; HW 0.87; ML 0.55; PNW 0.62; WL 1.17; GL 1].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF96FFEBF0A5F025FDECF958.taxon	distribution	Distribution. This species is widely distributed in Brazil, occurring in north, central-west, northeast, and southeast regions (Fig. 89). Natural history. This species occurs in the “ Cerrado ” (Brazilian savanna), “ Caatinga ” (Brazilian dry forest), and Atlantic and Amazon forests in Brazil. It is mostly collected in leaf-litter samples with winkler extractors, which suggests that it nests in the leaf-litter, fallen logs, rotten wood, or inside natural cavities of the superficial soil layers. Specimens collected with pitfall traps and with honey or sardine baits are rare. Except for the presence of intercastes, and possibly ergatoids (a few intercastes have more queen-like traits), there is no further information available regarding the biology of H. lopesi.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FF96FFEBF0A5F025FDECF958.taxon	materials_examined	Additional material examined (177 workers, 5 queens, 17 intercastes): BRAZIL: AC [Acre]: Rio Branco- Res. Catuaba, 10 ° 04 ′ S, 67 ° 37 ′ W, Dez. 2001 - Dez. 2005, Oliveira M. A. (2 W) [CEPLAC]; BA [Bahia]: Milagres, Caatinga Arbórea, área 1, S 12 ° 54.542 ′, W 39 ° 51.279 ′, 13. vii. 2010, Ulysséa, M. A., Medina, A. M. e Campos, E. M. leg., serapilheira, winkler, W 4 (1 W) [MZSP]; same except 10. i. 2011, W 110 (1 W) [MZSP]; W 113 (1 W covered with gold) [MZSP]; 23. x. 2010, W 58 (1 W) [MZSP]; área 2, S 12 ° 54.294 ′, W 39 ° 52.083 ′, Ulysséa, M. A., Medina, A. M. e Campos, E. M. leg., 13. vii. 2010, W 22 (2 W) [MZSP]; Gongogi, 17 a 18. iv. 1997, 14 ° 16 ′ 27 ″ S, 39 ° 29 ′ 03 ″ W, Santos, J. R. M. dos (3 W 1 Q) [CEPLAC]; Jussari, 15 ° 08 ′ 26 ″ S, 39 ° 31 ′ 29 ″ W, 26. v. 1999, col. J. C. S. Carmo & J. R. M. Santos, 128 (5 W) [CEPLAC]; A. de Olivença, mata-w, 23. iii. 1998, 14.58.85 S, 39.01.48 W, Santos, J. R. M. dos, 637 (1 Q 2 W) [CEPLAC]; Maraú, 14 ° 12 ′ 65 ″ S, 39 ° 03 ′ 48 ″ W, 23. x. 1998, J. C. S. Carmo & J. R. M. Santos, 637 (3 W) [CEPLAC]; Una, colônia, 12. vi. 1997, 15 ° 15 ′ 42 ″ S, 39 ° 09 ′ 12 ″ W, Santos, J. R. M. dos, 637 (2 W) [CEPLAC]; same except A 45, 24. viii. 1998, 151104 S, 0390056 W, Santos, J. R. M. dos, 637 (3 W, one covered with gold) [CEPLAC]; Res. Biológica, 13. ii. 1997, col. Jardim, J., # 5079 (4 W) [CEPLAC]; Arataca, 23. xi. 1999, 151590 S, 0391601 W, Santos, J. R. M., A 48, 637 (3 W) [CEPLAC]; Monte Pascoal, 16 ° 52 ′ 03 ″ S, 39 ° 55 ′ 03 ″ W, 18. vii. 1997, Santos, J. R. M. (3 W) [CEPLAC]; Ramal de Barreiras, 25. x. 1999, 15 ° 35 ″ 54 ″ S, 39 ° 09 ′ 38 ″ W, Santos, J. R. M. dos 637 (2 W) [CEPLAC]; Itajuipe, 18. vi. 1997, 14.42.12 S, 039.29.53 W, Santos, J. R. M. dos, 637 (3 W) [CEPLAC]; Camanan, 27. viii. 1999, 15.36.04 S, 039.31.16 W, Santos, J. R. M. dos, (3 W) [CEPLAC]; Ibicaraí, Km 41, 21. xi. 1998, 145375 S, 0392901 W, Santos, J. R. M. dos, 637 (1 Q 2 W) [CEPLAC]; Itabuna, Ferradas, A 27, 21. ix. 2000, 144933 S, 392416 W, Santos, J. R. M. dos, 637 (2 W) [CEPLAC]; Ilhéus, Pimenteira, Mata, W-A 17, 06. x. 1997, 14327 S, 392539 W, Santos, J. R. M. dos, Carmo, J. C. S. do, 637 (2 W) [CEPLAC]; Madisa, A 54, 04. x. 1999, 152323 S, 0391151 W, Santos, J. R. M. dos, 637 (1 W) [CEPLAC]; Unacau, A 43, 11. ii. 2000, 150521 S, 09 ° 1742 W, Santos, J. R. M. dos, 321 (2 W) [CEPLAC]; [São José de] Buerarema, A 39, mata, 28. x. 2002, J. R. M. dos Santos (3 W) [CEPLAC]; CE [Ceará]: Ubajara, Dist. Jaburuna, + 840 m, 03 ° 52 ′ S, 40 ° 57 ′ W, vii. 2004, Y. Quinet, W 74 (2 W) [MZSP]; same except W 6 (1 W) [MZSP]; W 76 (1 W) [MZSP]; W 33 (1 W) [MZSP]; W 56 (5 W) [MZSP]; W 39 (1 I) [MZSP]; Crateús, São Luis, 05 ° 08 ′ S, 40 ° 51 ′ W, 20 - 30. iv. 2003, Y. Quinet, Mata seca, Pitfall traps (1 W) [MZSP]; same except RPPN Serra das Almas, 5 ° 08 ′ S, 40 ° 55 ′ O, Winkler, Carrasco Preservado, 05. vi. 2011, F. A. Nunes col., CRP. 05 / VII / 2011, LUK. 06 MF. 62 (1 I) [MZSP]; CRP, WK. 10 MF. 62 (3 W) (MZSP 67438) [MZSP]; CRP, WK. 06 MF. 62 (2 W) [MZSP]; CRD. 03 / VI / 2011, PFS. 13 MF. 62 (1 W) [MZSP]; SP. 28 IV-PF (1 W) [MZSP]; SP. 28 / III (3 W) [MZSP]; MSD, PFS. 19 MF. 62 (2 W) [MZSP]; Pitfall, Carrasco, + 700 m, 27. iv. 2003, Y. Quinet col., SP. 28 / III (1 I) [MZSP]; Mata Seca Preservada, 15. iv. 2012, F. A. Nunes col., MSP, WK. 01 MF. 62 (1 I) (MZSP 67442) [MZSP]; 16. iv. 2012, MSP, WK. 17 MF. 62 (1 I) [MZSP]; Mata Seca Degradada, 15. iv. 2012, F. A. Nunes col., MSD, WK. 16 MF. 62 (1 I) [MZSP]; WK. 13 MF. 62 (1 I) [MZSP]; DF [Distrito Federal]: Brasília, Reserva Ecolo. IBGE, Projeto fogo, Cerrado sensu stricto, Parcela controle, pt.: 132, 08. ii. 2008, J. Maravalhas col. (1 W) [MZSP]; E. Ecol. Águas Emendadas, 25. v. 1992, A. Reis col., isca - mel, noite - solo, 30 (1 W) [MZSP]; same except isca - mel, dia - solo, 111 (1 W) [MZSP]; isca - sardinha, dia - solo, 85 (1 W) [MZSP]; Tabatinga, F. Z. Cooperbrás, Pitfall, 2003 - 2004, Schmidt F. G. V. (5 W) [CEPLAC]; same data (1 I) (MZSP 67441) [MZSP]; GO [Goiás]: Campo Limpo, Faz. Conceição, 16 ° 19 ′ 51 ″ S, 49 ° 09 ′ 49.2 ″ W, 20 - 24. i. 2005, Silva R. R. col., Winkler 2 (1 W) [MZSP]; Niquelândia, 18 - 30. v. 1996, 14 ° 17 ′ 05 ″ S, 48 ° 55 ′ 01 ″ W, Silvestre, Silva & Brandão col., Winkler, R. do Peixe (1 W) [MZSP]; same except Riacho Pindaíba (1 W) [MZSP]; MA [Maranhão]: Estreito, Fazenda Itaueiras, 06 ° 31 ′ 54.4 ″ S, 47 ° 22 ′ 16 ″ W, 07 - 13. i. 2005, Silva R. R. & Silvestre R. cols., Winkler 29 (1 W) [MZSP]; João Lisboa, 05 ° 19 ′ 46.3 ″ S, 47 ° 19 ′ 13.0 ″ W, 13 - 22. ii. 2006, Silva R. R. & Feitosa R. M. cols., Winkler 18 (2 W) [MZSP]; same data (2 W) [UTEP]; same except Winkler 10 (1 Q) [MZSP]; 01 - 09. vi. 2005, Silva R. R. & Feitosa R. M. cols., Winkler 28 (1 W) [MZSP]; Açailândia, Horto Faz. Pompéia, 04 ° 52 ′ 30 ″ S, 47 ° 17 ′ 40 ″ W, 12 - 22. ii. 2006, Silva R. R. & Feitosa R. M. cols., Winkler (1 W) [MZSP]; same except 13 - 22. ii. 2006, Winkler 08 (3 W) [CEPLAC]; Winkler 10 (1 W) [MZSP]; Winkler, Res [íduo] (3 W) [UFSC]; São Francisco do Brejão, 05 ° 17 ′ 19 ″ S, 47 ° 15 ′ 01.7 ″ W, 07 - 13. i. 2005, Silva R. R. & Silvestre R. cols., Winkler 22 (1 W) [MZSP]; MG [Minas Gerais]: Matias Cardoso, ii. 2012, Pitfall, CJ 2 ITPS 1 - 1032, Coletor L. F. Silva (1 W) [UFMG]; Parque Estadual Lagoa do Cajueiro, Estágio Intermediário, CJ 1 ITW 4, 16. ii. 2012, 14 ° 55 ’ 16.32 ” S, 43 ° 55 ’ 10.12 ” O, L. F. Silva col. (1 W) (MZHY 61) [MZSP]; same except CJ 2 ITW 2, 20. ii. 2012, 14 ° 56 ’ 9.01 ” S, 43 ° 55 ’ 27.51 ” O, L. F. Silva col. (1 W) (MZHY 62) [MZSP]; Pitfall JITPS 2 - 2526, Coletor L. F. Silva (1 I) (MZSP 67440) [MZSP]; Pandeiros, Efeito antropização no cerrado, APA REVISE, elev. 511 m, 15 ° 29 ′ 58.15 ″ S, 44 ° 45 ′ 39.94 ″ O, 07. i. 2016, Santiago et al., área 3, pitfall epigéico, Ponto 2, Repetição C, Q 2 (1 I) [DZUP]; same except Ponto 3, Repetição B, Quadrante 2 (1 I) [DZUP]; Uberlândia, Reserva do Panga, 19 ° 10 ′ 20.5 ″ S, 48 ° 23 ′ 46.5 ″ W, 24. i. 2012, T. Frizzo, J. Maravalhas & H. Vasconcelos col., T 36 15 S; MT [Mato Grosso]: Cotriguaçu, Faz. São Nicolau — ONF Brasil, 09 ° 48 ′ S, 58 ° 16 ′ W, col. manual, x. 2011, Vicente, R. E. (1 W) [MZSP]; PA [Pará]: Paragominas, 2 ° 59 ′ S, 47 ° 21 ′ W, i-vii. 2011, baited pitfall, B 324, T 1, P 125, R. Solar col., UFV LABECOL n ° 000144 (1 W) [MZSP]; Primavera, Projeto BIOMAS, área D, 00 ° 55 ′ 26 ″ S, 47 ° 06 ′ 27 ″ W, 25. v. 2013, Fernandes F. col., winkler D 07 (1 W) [MPEG]; PB [Paraíba]: Rio Tinto, APA Rio Mamanguape, 07. vi. 2014, Ramos E. F. et al. col., 06 ° 46 ′ 20.15 ″ S, 34 ° 55 ′ 12.90 ″ W (1 I) (MZSP 67439) [MZSP]; SP [São Paulo]: Agudos, C. Gilbert, iii. 1960, 3525 (1 I) [MZSP]; Mirassol, Faz. Campo, Reserva Lukezi, 15. x. 1971, J. L. M. Diniz, # 366, 11028 (2 W) [MZSP]; same except grota, 10. i. 1983, J. L. M. Diniz, 2068 (1 W) [DZUP]; Botucatu, 18 - 02 - 1988, Forti L. C. & Rinaldi I. M. P. col., Armadilha de solo, # 12 (1 W) [MZSP]; same except 18 - 01 - 1988, # 28 (1 W) [MZSP]; 13 - 07 - 1987, # 54 (1 W) [MZSP]; 09 - 11 - 1987, # 15 (1 W) [MZSP]; Cerrado, 17. ii. 1987, L. C. Forti col., Armadilha de solo, # 05 (1 W) [MZSP]; Luiz Antônio, Reserva Jataí, 02 / 97, Silvestre R. col. (1 W) [MZSP]; same except 05 / 97, Winkler, Cerrado (2 W) [MZSP]; Reserva Ecológica Jataí, 23 - 25. v. 1997, Cerrado, R. Silvestre & R. R. Silva col., isca sardinha I, dia solo A 49 (1 W) [MZSP]; same except Winkler, Cerrado (1 W) [MZSP]; TO [Tocantins]: Palmeiras do Tocantins, 06 ° 40 ′ 12.1 ″ S, 47 ° 31 ′ 48.6 ″ W, 14 - 19. i. 2005, Silva R. R. & Silvestre R. cols., Winkler 29 (1 W) [MZSP]; same except Winkler 45 (1 W) [MZSP]; Recursolândia, Mata Ciliar Rio Mateiros, 08 ° 45 ′ 28.6 ″ S, 47 ° 02 ′ 20.7 ″ W, 09 - 12. v. 05, Silva R. R. & Dietz B. H. cols., Winkler 15 (1 W) [MZSP]; Goiatins, 07 ° 56 ′ 28.9 ″ S, 47 ° 09 ′ 31.3 ″ W, 03 - 08. v. 05, Silva R. R. & Dietz B. H., Winkler 6 (1 W) [MZSP]; same except Winkler 2 (2 W) [MZSP]; Mata Ciliar do Ribeirão Grande, 07 ° 69 ′ 57 ″ S, 47 ° 22 ′ 42 ″ W, 28. iv- 03. v. 05, Silva R. R. & Dietz B. H., Winkler 11 (3 W) [MZSP]; same except Winkler 8 (2 W) [UTLP]; Campos Lindo, 08 ° 06 ′ 12.8 ″ S, 46 ° 50 ′ 44.9 ″ W, 03 - 08. v. 05, Mata Ciliar, Silva R. R. & Dietz B. H., Isca solo 38 (1 W) [MZSP]; same except Winkler 18 (1 W) [MZSP]; Winkler 16 (2 W) [MZSP]; Palmeirante, 07 ° 52 ′ 25.3 ″ S, 47 ° 57 ′ 07.4 ″ W, 10 - 15. xii. 2001, Albuquerque & Silva cols., Mata Ciliar / Cerradão, Transecto I, Winkler 14 (1 W) [MZSP]; same except Winkler 09 (1 W) [MZSP]; Winkler 04 (2 W) [JTLC]; Winkler 05 (1 W) [MZSP]; Isca solo 04 (1 W) [MZSP]; Isca solo 34 (1 W) [MZSP]; Porto Nacional, Fazenda Alto Paraíso, 05 - 06. x. 2001, 10 ° 43 ′ 32 ″ S, 48 ° 28 ′ 05 ″ W, Albuquerque & Silva cols., Cerradão, Transecto I, Winkler 07 (6 W) [MZSP]; same except Winkler 09 (1 W 1 I) [MZSP]; Aguiarnópolis, 06 ° 36 ′ 49.4 ″ S, 47 ° 28 ′ 53.2 ″ W, 01 - 09. vi. 2005, Silva R. R. & Feitosa R. M. cols., Winkler 3 (1 W 1 Q) [MZSP]; same except Winkler 12 (1 W) [MZSP]; Winkler 4 (1 W) [MZSP]; Winkler 9 (2 W) [MZSP]; 13 - 22. ii. 2006, # 16 (1 W) [MZSP]; 1 - 17. i. 2005, Silva R. R. & Silvestre R. cols., Winkler 9 (1 W) [MZSP]; Winkler 10 (2 W) [MZSP]; Winkler 5 (1 W) [MZSP]; Winkler 8 (2 W) [MZSP]; 14 - 19. i. 2005, Silva R. R. & Silvestre R. cols. (1 W) [MZSP]; Babaçulândia, 07 ° 05 ′ 16.3 ″ S, 47 ° 49 ′ 43.1 ″ W, 10 - 15. xii. 2001, Albuquerque & Silva cols., Mata Ciliar / Cerradão, Transecto I, Winkler 06 (1 W 1 I) [MZSP]; same except Winkler 18 (1 W) [UFGD]; Winkler 05 (1 W) [MZSP]; Winkler 20 (1 W) [MZSP]; Winkler 09 (3 W) [MZSP]; Winkler 11 (1 W 1 I) [MZSP]; Winkler 07 (2 W) [MZSP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFEBFFEEF0A5F300FAFBF85C.taxon	description	Figures 46, 47, 86 (map) Holotype: COLOMBIA: Nariño: Orito, Territorio Kofan, 00 ° 30 ′ N, 77 ° 13 ′ W, 1000 m, 25. ix. 1998, E. L. González leg., winkler 3 (1 W) [IHVL]. Paratypes: same data as holotype (1 Q covered with gold) [IHVL]; same except winkler 6 (1 W) (MZHY 208) [MZSP]; 1430 m, 23. ix. 1998, winkler 1 (1 Q) (MZSP 67387) [MZSP]. ECUADOR: Pastaza: 22 km SW Puyo, 15 July 1976, B- 362, S. & J. Peck (1 W) (MCZENT 00525515) [CASC]; same data (1 W) (MCZENT 00525513) [DZUP]; (1 W) (MCZENT 00525509) [IHVL]; (1 W) (MCZENT 00524684 MZSP 67390) [MZSP]; (7 W 1 Q) (MCZENT 00524682, MCZENT 00524683, MCZENT 00525508, MCZENT 00525511, MCZENT 00525512, MCZENT 00525514, MCZENT 00525516, MCZENT 00525507) [MCZC]; (1 W) (MCZENT 00525510) [USNM].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFEBFFEEF0A5F300FAFBF85C.taxon	diagnosis	Diagnosis. Head dorsum with regular and longitudinal striae, mesial striation in part directed to posterior margin and in part anteriorly divergent and posteriorly convergent, interspaces indistinguishable; mesosoma covered with irregular striae of variable thickness, assuming multiple directions, interspaces indistinguishable; metanotal groove discernible by a slight depression (LV) and altered sculpture (DV); propodeal spine long; dorsal margin of petiole discontinuous, dorsum with the same striation of mesosoma lateral; subtriangular projection on mesoventral surface of petiole; subpostpetiolar process weak, convex; profemur and protibia mostly smooth, few irregular to regular striae on basal and apical regions; short striae on tergum of first gastral segment, shorter than postpetiole length.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFEBFFEEF0A5F300FAFBF85C.taxon	description	Description. WORKER (n = 3) (Fig. 46 A – C): HL (0.88 – 0.91); HW (0.82 – 0.84); ML (0.54 – 0.56); SL (0.58 – 0.60); MOD (0.18 – 0.22); PNW (0.58 – 0.60); WL (1.10 – 1.14); PSL (0.28 – 0.34); PL (0.52 – 0.54); PW (0.20 – 0.23); PPL (0.32); PPW (0.31 – 0.32); GL (0.88 – 1); TL (4.28 – 4.42); CI (90.11 – 95.45); SI (70.73 – 73.17); OI (21.95 – 26.83). Small-sized. Subopaque integument, except for the shiny gaster. Brown body, lighter appendices. Many thin and unbranched setae, long to midsized, suberect to subdecumbent. Head subquadrate; posterior margin slightly concave at middle. Mandible masticatory margin with 5 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with regular and longitudinal striae, interspaces indistinguishable. Frontal triangle with 1 sagittal stria. Short scape, not reaching head posterior margin; apical antennomere shorter than previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye oval, small-sized, larger diameter with 10 ommatidia. Head dorsum, lateral and laterodorsal regions with longitudinal, regular to irregular striae, mesial striation in part directed to posterior margin and in part anteriorly divergent and posteriorly convergent, interspaces indistinguishable. Head lateral and laterodorsal striae converging to eye margin; gena striate, irregular and semicircular striae circumscribe the torulus, approaching eye margin, being 5 – 7 thicker striae, interspaces indistinguishable. Interspaces between striae on head ventral surface indistinguishable. Mesosoma covered with irregular striae of variable thickness, interspaces indistinguishable; striation assumes multiple directions, transverse on pronotum anterior region, longitudinal on pronotum posterior region and mesonotum, semicircular on metanotal groove and propodeum dorsum. Promesonotal junction indistinct. Metanotal groove discernible by a slight depression (LV) and altered sculpture (DV). Transverse carina inconspicuous. Dorsal margin of mesosoma slightly discontinuous, convex. Propodeal lobe bidentate, acute dorsal tooth longer than blunt ventral tooth; dorsal tooth length 1 / 2 of propodeal spine length. Propodeal spine long, straight (LV), divergent (DV), sculptured on base. Procoxa striae of uniform thickness; regular and transverse striae on C 2 and C 3. Profemur and protibia mostly smooth, few irregular to regular striae on basal and apical regions, transversely on profemur posterior surface, longitudinally on protibia extensor surface. Dorsal margin of petiole discontinuous; ventral surface smooth. Convex node; regular and transverse striae on ventral surface, interspaces indistinguishable; irregular and transverse striae of variable thickness on other surfaces, interspaces indistinguishable; striae on anterior surface and on anterior region of dorsal surface continuing on lateral surface; semicircular-sinusoid striae on middle and posterior region of dorsal surface. Postpetiole and subpostpetiolar process with regular and longitudinal striae; subpostpetiolar process weak, convex, striation weakly marked. First gastral segment striation similar to postpetiole striae; short striae on tergum, shorter than postpetiole length; sternite smooth. QUEEN (n = 3) (Fig. 47 A – C): HL (0.90 – 0.95); HW (0.88 – 0.92); ML (0.58 – 0.62); SL (0.62 – 0.70); MOD (0.26 – 0.28); PNW (0.68 – 0.76); WL (1.32 – 1.42); PSL (0.34 – 0.38); PL (0.54 – 0.64); PW (0.25 – 0.28); PPL (0.34 – 0.40); PPW (0.38 – 0.40); GL (1.10 – 1.26); TL (4.78 – 5.29); CI (94.44 – 96.84); SI (71.59 – 76.09); OI (29.54 – 30.59). Medium to large-sized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 13 ommatidia. Longitudinal striae on scutum, irregular to vermicular anteriorly, regular to irregular posteriorly, striae going from an anterior central point towards transcutal suture, interspaces distinguishable. Longitudinal, regular to irregular striae on anepisternum and katepisternum, interspaces indistinguishable. Axilla and scutellum with same sculpture of scutum. Scutoscutellar sulcus inconspicuous. Transverse striae on propodeum (DV). Lateral of mesosoma with irregular to vermicular striae directed partly to propodeal dorsum and partly to propodeal spine, interspaces distinguishable. Wings unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFEBFFEEF0A5F300FAFBF85C.taxon	etymology	Etymology. The epithet macielae is a Latin noun in the genitive case created by adding the singular Latin genitive case suffix - ae to the middle name of a female person. The specific epithet is named after Miraildes Maciel Mota, a black Brazilian woman and footballer, popularly known as “ Formiga ” (= ant in Portuguese). Formiga holds many international records; she is the only player present in all female football editions in the Olympic Games (the first edition happened in 1996), and has participated in seven different FIFA Women’s World Cup.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFEBFFEEF0A5F300FAFBF85C.taxon	discussion	Comments. This species is similar to H. blandiens, H. dentiloba, and H. jeronimae, sharing the adjacent striation on the mesosoma, but can be distinguished from these three species in the long propodeal spine (vs. short). Also, H. macielae can be distinguished from H. blandiens (characteristic in parentheses) in the striae on the mesosomal dorsum that assumes multiple directions (vs. longitudinal); from H. dentiloba in the oval eye (vs. drop-shaped), and the convex dorsal margin of mesonotum (vs. straight); and from H. jeronimae in the oval eye (vs. reniform), and the discontinuous dorsal margin of petiole (vs. continuous). Hylomyrma jeronimae and H. dentiloba are restricted to Central America (Figs. 84, 86), co-occuring in Panamá, whereas H. blandiens has a broad distribution in northwestern South America, from Bolivia to French Guiana (Fig. 84), overlaping in part with the area of distribution of H. macielae (south Colombia and the center-north of Ecuador) (Fig. 86).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFEBFFEEF0A5F300FAFBF85C.taxon	distribution	Distribution. This species occurs in Northern Andes, from northern Peru to southern Venezuela (Fig. 86). Natural history. This species was collected at elevations between 96 and 1430 m (Letícia and Territorio Kofán, both in Colombia) with winkler extractors or Berlese-Tüllgren funnels, which suggests that nests are located in leaflitter, fallen logs, rotten wood, or inside natural cavities of the superficial soil layers. Additional material examined (10 workers): COLOMBIA: Amaz. [Amazonas]: 7 km NW Letícia, 20 - 25 Feb 1972, # 230, forest litter, berlesate (6 W, one covered with gold) (MZSP 67388, MZSP 67389) [MZSP]; same data (2 W) [MCZC]. ECUADOR: Pastaza: 25 km N of Puyo, 1000 m, 13. vii. 1976, B 361, S. & J. Peck (1 W) [MCZC]; Pichincha: Tinalandia, 16 km SE of S. Domingo de los Colorados, 4. vi. 1976, S. & J. Peck (1 W) [MCZC].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFEFFFE2F0A5F494FD6BFE48.taxon	description	Figures 48, 49, 89 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFEFFFE2F0A5F494FD6BFE48.taxon	materials_examined	Holotype: GUYANA: Upper Takutu-Upper Essequibo: Acarai Mts., nr. New Romeo Camp, 1038 m, 58 ° 57.876 ′ W, 1 ° 20.066 ′ N, 14. x. 2006, T. R. Schultz, J. Sosa-Calvo, C. J. Marshall, R. Williams, 1 ° forest, leaf litter, Winkler sample, JSC 061014 - LS 04 (1 W) (USNMENT 00688842) [USNM]. Paratypes: same data as holotype (1 W) (USNMENT 00688836 MZHY 195) [MZSP]; (2 W 1 Q, one worker covered with gold) (USNMENT 00688820, USNMENT 00688844, USNMENT 00688842) [USNM]; same except 1050 m, 58 ° 57.876 ′ W, 1 ° 20.048 ′ N, JSC 061014 - LS 03 (1 W) (USNMENT 00688740) [MCZC]. FRENCH GUIANA: [Cayenne: Saint-Élie], Nouragues Station, FT 2 - Transition For., 4 ° 09 ′ N, 52 ° 68 ′ O, ix. 2009, Sarah Groc & al, # 5635, FT 2 Tr 5 W 29 (1 Q) (MZSP 67391) [MZSP]; [Saint-Laurent-du-Maroni: Maripasoula], Mitaraka Mts, Plateau, 2.233158 N - 54.44381 W, 27. ii. 2015, J. Orivel & F. Petitclerc cols., winkler 48 h, C-P 7 - 3, MI 15 - 0079 - 15 (1 W) (MZSP 67392) [MZSP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFEFFFE2F0A5F494FD6BFE48.taxon	diagnosis	Diagnosis. Regular and longitudinal striae on head dorsum diverge towards posterior margin, interspaces between striae smooth, striae crest punctuated; mesosoma covered with regular, longitudinal, parallel and thick striae; longitudinal striae on lateral of pronotum, mesepisternum and propodeum directed to propodeal spine; dorsal margin of petiole continuous, strongly convex; spiniform projection on mesoventral surface of petiole; node rounded with transverse striae anteriorly and longitudinal striae posteriorly on dorsum; subpostpetiolar process smooth, prominent, subtriangular; profemur posterior surface and protibia extensor surface mostly smooth, with few striae on basal and apical regions; striation on tergum of first gastral segment restricted to base.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFEFFFE2F0A5F494FD6BFE48.taxon	description	Description. WORKER (n = 3) (Fig. 48 A – C): HL (0.85 – 0.88); HW (0.79 – 0.82); ML (0.53 – 0.55); SL (0.58 – 0.59); MOD (0.20 – 0.21); PNW (0.58); WL (1.12 – 1.18); PSL (0.14 – 0.16); PL (0.53 – 0.57); PW (0.21 – 0.22); PPL (0.32 – 0.34); PPW (0.29 – 0.30); GL (0.86 – 0.98); TL (4.21 – 4.50); CI (92.94 – 94.25); SI (71.95 – 73.41); OI (25.31 – 25.61). Small-sized. Shiny integument. Brownish body, darker gaster, yellowish appendices. Few thin and unbranched setae, long to midsized, suberect to subdecumbent. Head subquadrate; posterior margin straight. Mandible masticatory margin with 5 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with 8 – 9 striae, regular to irregular, longitudinal, converging to a point on the anterior margin, interspaces distinguishable. Frontal triangle with 1 stria subdivided towards the posterior margin. Short scape, not reaching head posterior margin; apical antennomere of similar size or slightly shorter than previous 3 antennomeres together. Frontal carina straight. Eye dropshaped, small-sized, larger diameter with 9 ommatidia. Regular and longitudinal striae on head dorsum, divergent towards posterior margin, interspaces smooth, striae with punctuated crests. Head laterodorsal sculpture converges to mandible insertion region; head lateral sculpture converges to eye margin; gena striate, 2 – 3 regular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma covered with regular and longitudinal striae, thicker than head striation, transverse on pronotum anterior region (DV), longitudinal in the other regions (DV); longitudinal striae on lateral of pronotum, mesepisternum and propodeum directed to propodeal spine. Promesonotal junction and metanotal groove indistinct. Transverse carina well-marked. Dorsal margin of mesosoma continuous, convex. Propodeal lobe bidentate, dorsal tooth longer and with rounded tip; ventral tooth shorter and blunt; dorsal tooth length similar to propodeal spine length. Propodeal spine short, straight (LV), divergent (DV), sculptured on base. Procoxa striae of uniform thickness; irregular and transverse striae on C 2 and C 3. Profemur and protibia mostly smooth, few striae on basal and apical regions. Dorsal margin of petiole continuous, strongly convex; smooth spiniform projection on mesoventral surface; ventral surface smooth; first third of dorsal surface and parts of lateral surface smooth, other regions mostly striate, transverse and longitudinal striae. Longitudinal striae on postpetiole; subpostpetiolar process well-developed, subtriangular, smooth. Tergum first gastral segment striation similar to that of postpetiole; striae short, restricted to base, length shorter than 1 / 4 of postpetiole length; sternum smooth. QUEEN (n = 2) (Fig. 49 A – C): HL (0.82 – 0.90); HW (0.80 – 0.86); ML (0.52 – 0.58); SL (0.54 – 0.60); MOD (0.24 – 0.26); PNW (0.65 – 0.70); WL (1.20 – 1.36); PSL (0.20 – 0.22); PL (0.58 – 0.63); PW (0.24 – 0.26); PPL (0.36 – 0.38); PPW (0.33 – 0.34); GL (1.04 – 1.16); TL (4.52 – 5.01); CI (95.55 – 97.56); SI (67.50 – 69.76); OI (30 – 30.23). Small to midsized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 12 ommatidia. Anepisternum, katepisternum, axilla and scutellum with the same sculpture of scutum. Scutoscutellar sulcus inconspicuous. Transverse striae on propodeum (DV). Lateral of mesosoma with longitudinal and regular striae directed to propodeal spine, interspaces distinguishable. Wings unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFEFFFE2F0A5F494FD6BFE48.taxon	etymology	Etymology. The epithet margaridae is a Latin noun in the genitive case created by adding the singular Latin genitive case suffix - e to the first name of a female person. This species is named in honor of Margarida Maria Alves (1933 – 1983), a Brazilian trade unionist and precursor in defending the rights of rural workers in Paraíba, northeast region of Brazil — “ Da luta eu não fujo. É melhor morrer na luta do que morrer de fome ”. Margarida was murdered for her activism on the 12 of August. Since the year 2000, thousands of women from all of the regions of the country gather every two years in a march for the equal rights for rural women that bears her name — “ Marcha das Margaridas ”.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFEFFFE2F0A5F494FD6BFE48.taxon	discussion	Comments. This species can be misidentified mainly as H. peetersi and H. villemantae. The three species are allopatric; H. villemantae occurs in one locality in northeastern Brazil (BA) (Fig. 86), H. margaridae occurs in Guyana and French Guiana (Fig. 89), and H. peetersi in areas near the Guyana - Venezuela border (Fig. 89). Hylomyrma margaridae can be distinguished from H. peetersi (characteristic in parentheses) in the striae on the laterals of mesosoma that are directed transversely towards the propodeal spine (vs. directed to the propodeal dorsum), the mesoventral surface of petiole armed with a conspicuous spine (vs. a subtriangular projection), the petiolar dorsum covered with both transverse and logitudinal striae (vs. entirely covered with transverse striae), and the prominent subpostpetiolar process (vs. weak). Hylomyrma margaridae can be distinguished from H. villemantae in the striae on the head dorsum with punctuated crests (vs. with smooth crests), the mesosoma longitudinally striate (vs. with few elliptical and concentric striae), the shorter propodeal spine (vs. longer), the armed mesoventral surface of petiole (vs. unarmed), and the smooth dorsum of postpetiole and gaster (vs. striate). Hylomyrma margaridae and H. lopesi have similar shape and sculpture of the mesosoma. Nevertheless, H. margaridae has a reniform eye (vs. oval eye of H. lopesi), the striae on the head dorsum with punctuated crests (vs. with smooth crests), and the continuous dorsal margin of petiole (vs. discontinuous). The two species are allopatric (Fig. 89); H. margaridae occurs in Guyana and French Guiana, and H. lopesi in Brazil.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFEFFFE2F0A5F494FD6BFE48.taxon	description	Regarding morphological variation, the queen collected in French Guiana has the petiolar node covered with concentric striae that circumscribe its anterodorsal, lateral and posterior regions.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFEFFFE2F0A5F494FD6BFE48.taxon	distribution	Distribution. This species is known from the southern part of the Upper Takutu-Upper Essequibo region in Guyana, and from the Réserve Naturelle Nationale des Nouragues, in the central region of French Guiana (Fig. 89). Natural history. This species has been recorded with winkler extractors in forested areas at high elevations ranging from 1038 to 1050 m in Guyana and at lower elevations in French Guiana (Nouragues Station is located at about 124 m and Mitaraka Mountain at 554 m).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE2FFE4F0A5F5FDFC77FD28.taxon	description	Figures 50, 88 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE2FFE4F0A5F5FDFC77FD28.taxon	materials_examined	Holotype: COLOMBIA: [Magdalena]: Cincinnati, Feby 1924, W. M. Mann (1 W) [USNM]. Paratypes: same data as holotype (3 W) [CASC, DZUP, IHVL]; (2 W, one covered with gold) (MZSP 67393, MZHY 205) [MZSP]; (4 W) [USNM]; San Pedro de La Sierra — Pico Yerbabuena, C. Kugler col., 2200 m, berlesate rot. wood & humus, 18. feb. 77, vial 13 (1 W) [MCZC]; same except 1730 m, berlesate humus & lf. Litter, 18. feb. 77 (1 Q) [MCZC].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE2FFE4F0A5F5FDFC77FD28.taxon	diagnosis	Diagnosis. Irregular and longitudinal striae on head dorsum diverge towards posterior margin; rugose striae on mesosoma and petiole; promesonotal junction and metanotal groove distinguishable by a slight depression (LV) and altered sculpture (DV); dorsal margin of petiole discontinuous; subtriangular projection on mesoventral surface of petiole; node ventral surface striate transversely; profemur posterior surface entirely striate; striae transverse and well-marked; protibia extensor surface entirely striate, striae weakly marked; long striae on first gastral tergite.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE2FFE4F0A5F5FDFC77FD28.taxon	description	Description. WORKER (n = 6) (Fig. 50 A – C): HL (0.99 – 1.04); HW (0.92 – 0.96); ML (0.60 – 0.65); SL (0.65 – 0.68); MOD (0.20 – 0.22); PNW (0.64 – 0.68); WL (1.26 – 1.32); PSL (0.32 – 0.37); PL (0.62 – 0.64); PW (0.25 – 0.27); PPL (0.30 – 0.32); PPW (0.34 – 0.36); GL (1.04 – 1.30); TL (4.85 – 5.23); CI (91.34 – 94.11); SI (69.47 – 70.83); OI (20.83 – 23.91). Midsized. Shiny integument. Light brown body, darker gaster, yellowish leg. Thin and unbranched setae, long to midsized, suberect to subdecumbent. Head subquadrate; posterior margin slightly concave at middle. Mandible masticatory margin with 6 teeth. Anterior margin of clypeus slightly convex medially, with a pair of medium teeth laterally; median area of clypeus with 8 – 10 irregular and longitudinal striae converging to a point on the anterior margin, interspaces distinguishable. Frontal triangle with 2 – 3 striae. Short scape, not reaching head posterior margin; apical antennomere slightly shorter than previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye dropshaped, small-sized, larger diameter with 10 ommatidia. Irregular and longitudinal striae on head dorsum, divergent towards posterior margin, interspaces between thicker striae filled with thinner striae. Head lateral and laterodorsal regions with irregular striae converging to eye margin; rugose striae on anterior part of laterodorsal region; gena striate, 3 – 4 regular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma covered with rugose striae of uniform thickness, mostly transverse on pronotum anterior region (DV), longitudinal in other regions (DV)., interspaces between rugose striae smooth. Promesonotal junction and metanotal groove distinguishable by a slight depression (LV) and altered sculpture (DV). Transverse carina inconspicuous. Dorsal margin of mesosoma slightly discontinuous, convex. Propodeal lobe bidentate, dorsal tooth slightly longer and sharper than shorter and blunt ventral tooth; dorsal tooth length around 1 / 2 of propodeal spine length. Propodeal spine long, straight (LV), divergent (DV), sculptured on base. Procoxa striae of uniform thickness; irregular and transverse striae on C 2 and C 3. Profemur ventral surface smooth, transverse striae well-marked on posterior surface; apical region of anterior and dorsal surfaces with irregular striae. Protibia extensor surface entirely striate. Dorsal margin of petiole discontinuous. Subtriangular projection on mesoventral surface of petiole. Convex node; transverse striae on ventral and anterior surfaces weakly marked; rugose striae on lateral and dorsal surfaces. Postpetiole and subpostpetiolar process with regular and longitudinal striae; subpostpetiolar process weak, convex. First gastral segment striation similar to postpetiole striae; longitudinal striae on tergum slightly longer than postpetiole length; sternite entirely striate on basal region. QUEEN (n = 1): HL 1.06; HW 1.01; ML 0.66; SL 0.70; MOD 0.24; PNW 0.80; WL 1.50; PSL 0.42; PL 0.70; PW 0.30; PPL 0.36; PPW 0.42; GL 1.26; TL 5.54; CI 95.28; SI 69.30; OI 23.76. Large-sized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 13 ommatidia. Longitudinal and irregular striae on scutum going from an anterior central point towards transcutal suture. Longitudinal and regular to irregular striae on anepisternum and katepisternum. Axilla and scutellum with same sculpture of scutum. Scutoscutellar sulcus inconspicuous. Transverse and regular striae on propodeum (DV). Lateral of mesosoma with irregular to vermicular striae directed partly to propodeal dorsum and partly to propodeal spine. Wings unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE2FFE4F0A5F5FDFC77FD28.taxon	etymology	Etymology. The epithet mariae is a Latin noun in the genitive case created by adding the singular Latin genitive case suffix - e to the first name of a female person. The specific epithet is named after Maria do Espírito Santo da Silva (1956 – 2011), a Brazilian conservationist and environmentalist who campaigned against logging in the Amazon rainforest. Maria and her husband, José Cláudio Ribeiro da Silva (1957 – 2011), extracted sustainable rainforest products, such as oils and nuts. Both were murdered for their activism in name of the Brazilian Amazon conservation.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE2FFE4F0A5F5FDFC77FD28.taxon	discussion	Comments. Hylomyrma mariae is similar to H. adelae and H. dandarae. Hylomyrma mariae differs from H. adelae (characteristics in parentheses) in the irregular divergent-longitudinal striae on the head dorsum (vs. rugose striae), the distinct metanotal groove (vs. indistinct), the longer propodeal spine (vs. shorter), the profemur posterior surface with striae weakly marked (vs. well-marked), and the dorsum of petiole with rugose striae (vs. irregular and transverse striae). The two species are allopatric; Hylomyrma mariae occurs in northern Colombia (Fig. 88), and H. adelae in Bolivia (Fig. 83). Hylomyrma mariae differs from H. dandarae in the drop-shaped eye (vs. oval), the shorter petiolar node (vs. longer), and the long striae on tergum of the first gastral segment that extends up to its first third (vs. short striae). The two species occur in Colombia (Figs. 83, 88).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE2FFE4F0A5F5FDFC77FD28.taxon	distribution	Distribution. Hylomyrma mariae is known from northern Colombia (Fig. 88). Natural history. This species is only known from its type locality, and inhabits areas at elevations between 1340 and 2200 m. Most specimens were collected from rotten wood, humus, and leaf-litter submitted to Berlese-Tüllgren funnels.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE2FFE4F0A5F5FDFC77FD28.taxon	materials_examined	Additional material examined (19 workers): COLOMBIA: [Magdalena]: Cincinnati [pueblo, 11 ° 09 ′ N 74 ° 08 ′ W], Feby 1924, W. M. Mann (3 W, two without head, one without postpetiole and gaster) [USNM]; Cuchilla S. [San] Lorenzo, vic. El Campano, 1340 m, rainf., C. Kugler & W. L. Brown cols. (15 W) [MCZC]; El Campano, 1740 m, 16. june. 1976, Brown & Kugler cols., berlesate (1 W) [MCZC].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE4FFE7F0A5F71DFB95FA6E.taxon	description	Figures 51, 52, 80 C, 85 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE4FFE7F0A5F71DFB95FA6E.taxon	materials_examined	Holotype: BRAZIL: PA [Pará]: Primavera, PBA-Projeto Primavera, 1 ° 02 ′ 48 ″ S, 47 ° 9 ′ 25 ″ W, 16 – 26. xi. 2013, M. Souza col., Área / Winkler A / 01 (1 W) (MZSP 67394) [MZSP]. Paratypes: AC [Acre]: Mâncio Lima, P. N. da Serra do Divisor, 245 m, 7 ° 26 ′ 58.81 ″ S, 73 ° 39 ′ 37.83 ″ W, 15 – 18. xi. 2016, R. M. Feitosa, T. S. Silva & A. C. Ferreira cols. (1 W) [DZUP]; TO [Tocantins]: Araguacema, Senhor do Bonfim, 08 ° 40 ′ 20 ″ S, 49 ° 25 ′ 53 ″ W, 16 – 30. xi. 2005, Semidecídua, Silva R. R. & Feitosa R. M., Winkler (1 Q) (MZSP 67395) [MZSP]; same except Winkler 7 (1 W) (MZSP 67396) [MZSP]; Winkler 6 (1 W) (MZSP 67397) [MZSP]; Babaçulândia, 07 ° 02 ′ 19 ″ S, 47 ° 52 ′ 03.4 ″ W, 01 – 09. vi. 2005, Silva R. R. & Feitosa R. M., Winkler 13 (1 W) (MZSP 67398) [MZSP]; same except Winkler 16 (1 W covered with gold) (MZSP 67399) [MZSP]; Winkler 12 (1 Q) (MZSP 67400) [MZSP]; Winkler 4 (1 W) [USNM]; RO [Rondônia]: Porto Velho, Área Abunã, A 9 P 4, 09 ° 38 ′ 03.3 ″ S, 65 ° 26 ′ 23.9 ″ W, 17 – 27. vii. 2013, Mazão G. R. & Probst R. S. cols. (2 W) [DZUP]; same except Área Mutum, M 7 P 2, 09 ° 35 ′ 41.6 ″ S, 65 ° 03 ′ 54.2 ″ W, 03. iii. 2015, winkler 1 (1 W) (MZHY 106) [MZSP]; MT [Mato Grosso]: Diamantino, Faz. Junqueira Vilela, 11 km NE casa, 17 – 18. july. 1973, W. L. Brown col. (1 W) [MCZC].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE4FFE7F0A5F71DFB95FA6E.taxon	diagnosis	Diagnosis. Regular and longitudinal striae on head dorsum, parallel towards posterior margin; long scape; vermicular striae on mesosoma; dorsal margin of petiole discontinuous; ventral surface entirely striate; transverse striae; postpetiole covered with thick, regular and longitudinal striae, slightly anastomosed; subpostpetiolar process sculptured on laterobasal region, midbasal region smooth; long striae on tergum of first gastral segment; body with many thick and unbranched setae.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE4FFE7F0A5F71DFB95FA6E.taxon	description	Description. WORKER (n = 6) (Fig. 51 A – C): HL (0.90 – 0.96); HW (0.80 – 0.88); ML (0.60 – 0.64); SL (0.75 – 0.86); MOD (0.24 – 0.27); PNW (0.60 – 0.65); WL (1.20 – 1.30); PSL (0.31 – 0.34); PL (0.52 – 0.54); PW (0.20 – 0.22); PPL (0.30 – 0.32); PPW (0.29 – 0.32); GL (1.06 – 1.20); TL (4.58 – 4.93); CI (88.89 – 92.63); SI (88.37 – 100); OI (27.90 – 30.68). Small to midsized. Shiny integument. Bicolored body, brownish, lighter head and gaster, yellowish appendices. Many thick and unbranched setae (Fig. 80 C), long to midsized, erect to decumbent. Head subquadrate; posterior margin straight. Mandible masticatory margin with 6 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with 8 – 10 striae, regular and longitudinal, converging to a point on the anterior margin, interspaces distinguishable. Frontal triangle with 1 sagittal stria. Long scape, surpassing head posterior margin; apical antennomere slightly shorter than previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye drop-shaped, midsized, larger diameter with 11 ommatidia. Regular and longitudinal striae on head dorsum, parallel towards posterior margin, interspaces between thicker striae filled with thinner striae; anastomosed striae on posterior region. Sculpture on head lateral and laterodorsal regions converge to eye margin; longitudinal striae predominate on laterodorsal region; anastomosed striae predominate on lateral and lateroventral regions; gena striate, 6 – 9 regular and semicircular striae circumscribe the torulus, almost reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma covered with vermicular striae of uniform thickness, interspaces smooth. Promesonotal junction distinguishable by a slight depression (LV); metanotal groove indistinct. Transverse carina well-marked. Dorsal margin of mesosoma discontinuous, convex. Propodeal lobe bidentate, dorsal tooth slightly longer and sharper than the shorter and blunt ventral tooth; dorsal tooth shorter than 1 / 3 of propodeal spine length. Propodeal spine long, straight (LV), divergent (DV), sculptured on base. Procoxa striae of uniform thickness; irregular and transverse striae on C 2 and C 3. Profemur with transverse to inclined striae weakly marked. Protibia smooth. Dorsal margin of petiole discontinuous; transverse striae on its entire ventral surface. Node with regular and transverse striae on anterior surface, striation on anterior surface continuing transversely on lateral surface to ventral surface, striae weakly marked; vermicular striae on dorsal surface. Postpetiole covered with thicker, regular and longitudinal striae, slightly anastomosed; subpostpetiolar process weak, straight, with the same sculpture of laterobasal region of postpetiole, sculpture weakly marked, midbasal region smooth. First gastral segment striation similar to postpetiole striae; longitudinal striae on tergum 1 / 3 longer than postpetiole length; sternite striation weakly marked, restricted to laterobasal region. QUEEN (n = 3) (Fig. 52 A – C): HL (0.94 – 1); HW (0.85 – 0.88); ML (0.62 – 0.64); SL (0.78 – 0.84); MOD (0.30); PNW (0.72 – 0.74); WL (1.42 – 1.48); PSL (0.34 – 0.36); PL (0.62 – 0.64); PW (0.23 – 0.24); PPL (0.37 – 0.38); PPW (0.33 – 0.35); GL (1.30 – 1.44); TL (5.29 – 5.52); CI (88 – 91.66); SI (88.63 – 95.45); OI (34.09 – 35.29). Large-sized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 15 – 16 ommatidia. Striae on scutum anastomosed anteriorly and irregular posteriorly, going from an anterior central point towards transcutal suture. Longitudinal and regular striae on anepisternum, logitudinal and irregular to vermicular striae on katepisternum, interspaces distinguishable in both. Axilla and scutellum with the same sculpture of scutum. Scutoscutellar sulcus inconspicuous. Transverse striae on propodeum (DV). Lateral of mesosoma with irregular to vermicular striae directed in part to propodeal dorsum and in part to propodeal spine. Wings unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE4FFE7F0A5F71DFB95FA6E.taxon	etymology	Etymology. The epithet marielleae is a Latin noun in the genitive case created by adding the singular Latin genitive case suffix - ae to the first name of a female person. This species is named in honor of Marielle Franco (1979 – 2018), born Marielle Francisco da Silva, a black Brazilian woman, politician, feminist, and human rights activist, whose assassination, motivated by her positions and actions against all forms of discrimination, happened during her mandate as a representative in Rio de Janeiro local Assembly.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE4FFE7F0A5F71DFB95FA6E.taxon	discussion	Comments. Hylomyrma marielleae is similar to H. longiscapa regarding the body size and the sculpture pattern. However, H. marielleae can be easily distinguished from H. longiscapa (characteristic in parentheses) in the thick and dark setae (vs. thin and light), and the transverse striae on the ventral surface of petiole weakly marked (vs. well-marked). Both species have broad distribution in the center-north of South America, however H. longiscapa occurs in many localities in eastern portions of Brazil (from northeast to southeast regions) (Fig. 83) where H. marielleae does not occur (Fig. 85). One queen collected in French Guiana [USNMENT 00536047] has lighter setae when compared to the other specimens evaluated here.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE4FFE7F0A5F71DFB95FA6E.taxon	distribution	Distribution. Hylomyrma marielleae is known from Brazil, Bolivia, Ecuador, French Guiana, and Peru (Fig. 85). Natural history. This species inhabits sites at elevations between 120 and 590 m in tropical rainforest, semideciduous forest, bamboo forests, crops, and pastures. There is no further information available regarding the H. marielleae biology, besides nesting in the leaf-litter or inside natural cavities of the superficial soil layers, given that specimens were collected with winkler extractors.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE4FFE7F0A5F71DFB95FA6E.taxon	materials_examined	Additional material examined (31 workers, 3 queens): BOLIVIA: Cochabamba: Villa Tunari, 67.5 k E Valle Sajta, 6. ii. 99, R. Anderson, # 18525, 17 ° 06 ′ 19 ″ S, 64 ° 46 ′ 57 ″ W, Lowland rain forest, litter, # 18525 (3 W) [UTEP, CASC, MZSP]. ECUADOR: [Sucumbíos]: Cuyabeno, 12 / 10 - 05 / 11 / 94, J. P. Caldwell, # 10349 (1 W) [CEPLAC]; Napo: Limoncocha, 250 m, 18. vi. 1976, B- 348, S. & J. Peck (6 W 1 Q) [MCZC]; same except 25. vi. 1976, B- 355 (2 W) [MCZC]. FRENCH GUIANA: [Cayenne]: Nouragues Field Station, H Trail, 153 m, 52 ° 40.834 ′ W, 4 ° 5.196 ′ N, 1. viii. 2005, T. R. Schultz, J. S. LaPolla, D. Price, 1 ° forest, litter sample, Winkler sample, TRS 050801 - 03 - LS 07 (1 W) (USNMENT 00536053) [USNM]; same data (2 Q) (USNMENT 00536047, USNMENT 00536048) [USNM]; Nouragues, Fourmis de litière, 4 ° 04 ′ 58 ″ N, 52 ° 40 ′ 28 ″ O, 120 - 160 m, 29. iii. 2006, Jérôme Orivel (+ autres), FL-WTr 25 (1 W) [MZSP]. PERU: Cusco: Estación Biológica Villa Carmen, 590 m, - 12.902437 ° - 71.407672 ° + 300 m, bamboo forest, 2 ° vegetation, 5 - 15. viii. 2013, Ant Course cols. (2 W) [DZUP]; same except w 03 (3 W) (MZHY 68, MZHY 103) [MZSP]; successional vegetation, crops and pasture (2 W) [DZUP]; 525 m, - 12.888005 ° - 71.401526 ° + 300 m, riparian successional vegetation, 8. viii. 2013, Lattke 3385 (6 W) [MZSP]; Madre de Dios: Los Amigos Field Station, Trail 3, Huangana, 277 m, 70 ° 6 ′ 3.1 W, 12 ° 34 ′ 8.4 S, 9. x. 2004, T. R. Schultz, C. Marshall, J. Sosa-Calvo, 1 ° forest, litter sample, = JSC 041009 - 02, TRS 041009 - 01 - LS 02 (1 W) [USNM]; same except Trail 6, = JSC 041006 - 12, TRS 041006 - 01 - LS 12 (1 W) [USNM]; 180 m, Sachavacayoc Center, 34.39 k SW Puerto Maldonado, 21. vii. 2012, W. & E. Mackay, # 24994, 12 ° 51 ′ 10.9 ″ S, 69 ° 22 ′ 2.5 ″ W, litter extraction, Tropical rain forest, seasonally flooded reddishbrn clay (2 W) [MZSP, IHVL]; same except AW 10 S (1 W) [UNMSM]; Pasco: nr. Pozuzo, steep 1 ° for. above farms, 1000 m, 4 - 6. vi. 81, berl. lf. lit., C. Kugler & R. R. Lambert cols. (1 W) [MCZC].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE7FFFAF0A5F1DFFB07FA00.taxon	description	Figures 53, 54, 80 D, 81 H, 87 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE7FFFAF0A5F1DFFB07FA00.taxon	materials_examined	Holotype: GUYANE FRANÇAISE: [Saint-Laurent-du-Maroni: Maripasoula], Mitaraka Mts, Pente, 2.238226 N - 54.45203, 04. iii. 2015, J. Orivel & F. Petitclerc cols., winkler 48 h, A-P 2 - 8, MI 15 - 0380 - 25 (1 W) (MZSP 67401) [MZSP]. Paratypes: same data as holotype (3 W) [CASC, DZUP, IHVL]; (7 W) (MZHY 223) [MZSP]; same except Plateau, 2.243615 N - 54.45882, 03. iii. 2015, J. Orivel & F. Petitclerc cols., winkler 48 h, A-P 1 - 12, MI 15 - 0344 - 62 (1 W 2 Q, one worker and one queen covered with gold) (MZSP 67403, MZSP 67402, MZSP 67404) [MZSP]; (2 W) [MCZC, USNM]; Maripasoula, vii. 1999, S. Durou (1 W) [CEPLAC]; [Cayenne]: Nouragues Natural Reserve Station, 102 km SW of Cayenne, lat 4.08799, lon - 52.67978, 145 m, 21. viii- 1. ix. 2018, Rainforest, Ant Course 2018 (2 W) (MZHY 224) [MZSP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE7FFFAF0A5F1DFFB07FA00.taxon	diagnosis	Diagnosis. Vermicular to vermiculated-areolated striae on head dorsum and mesosoma; dorsal margin of petiole continuous, ventral surface smooth; longitudinal and anastomosed striae on postpetiole and tergum of first gastral segment; subpostpetiolar process prominent, subtriangular; profemur posterior surface and protibia extensor surface entirely striate; striae on tergum of first gastral segment shorter than postpetiole length; 2 short branches of equal size diverging from the main axis.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE7FFFAF0A5F1DFFB07FA00.taxon	description	Description. WORKER (n = 3) (Fig. 53 A – C): HL (0.95 – 1.06); HW (0.98 – 1.01); ML (0.67 – 0.75); SL (0.75 – 0.83); MOD (0.27 – 0.29); PNW (0.73 – 0.76); WL (1.41 – 1.52); PSL (0.32 – 0.356); PL (0.69 – 0.72); PW (0.27 – 0.30); PPL (0.30 – 0.32); PPW (0.33 – 0.36); GL (1.13 – 1.21); TL (5.27 – 5.52); CI (94.11 – 106.45); SI (74.24 – 81.81); OI (27.27 – 28.78). Large-sized. Shiny integument. Dark brown body, lighter appendices. Thin and branched setae, long to midsized, erect to subdecumbent; 2 short branches of equal size diverging from the main axis (Fig. 80 D). Head subquadrate; posterior margin straight. Mandible masticatory margin with 5 teeth. Anterior margin of clypeus concave medially, with a pair of medium teeth laterally; median area of clypeus with irregular and longitudinal striae converging to a point on the anterior margin, interspaces indistinguishable. Frontal triangle with 1 sagittal stria. Short scape, not reaching head posterior margin; apical antennomere slightly shorter than previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye drop-shaped, midsized, larger diameter with 12 ommatidia. Head dorsum with vermicular to vermiculated-areolated striae, divergent towards posterior margin, interspaces smooth. Striae on head lateral and laterodorsal regions converge to eye margin; very thin striae (microsculpture) between vermicular to vermiculated-areolated striae (macrosculpture), interspaces between microesculpture indistinguishable; gena striate, with the same striation of head lateral, 2 – 3 regular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Thick, vermicular to vermiculated-areolated striae on mesosoma, interspaces between thick striae on mesosoma dorsum smooth, but filled with thinner striae on mesosoma lateral. Promesonotal junction indistinct. Metanotal groove discernible by a slight depression. Transverse carina well-marked. Dorsal margin of mesosoma discontinuous, slightly convex. Propodeal lobe bidentate, dorsal tooth slightly longer and more acute than the shorter and blunt ventral tooth; dorsal tooth length 1 / of propodeal spine length. Propodeal spine long, straight (LV), divergent (DV), 2 sculptured on base. Thin and transverse striae of uniform thickness on procoxa; irregular and transverse striae on C 2 and C 3. Irregular to regular transverse striae on profemur dorsal and posterior surfaces; anterior and ventral surfaces smooth. Protibia extensor surface entirely striate. Dorsal margin of petiole continuous, convex; ventral surface smooth; first third of dorsal surface smooth; irregular and transverse striae on second third of dorsal surface (aligned to petiolar spiracle) continuing on lateral surface; vermiculated-areolated striae on lateral and last third of dorsal surface. Longitudinal, regular to irregular striae on postpetiole, few anastomosed; subpostpetiolar process striae restricted to lateral region; subpostpetiolar process prominent, subtriangular. First gastral segment striation similar to postpetiole striae; short striae on tergum, shorter than postpetiole length; sternite striation weakly marked, covering the laterobasal region. QUEEN (n = 2) (Fig. 54 A – C): HL (1.04); HW (1.06); ML (0.75 – 0.76); SL (0.81 – 0.86); MOD (0.30); PNW (0.89 – 0.90); WL (1.67 – 1.69); PSL (0.36); PL (0.73 – 0.76); PW (0.30 – 0.32); PPL (0.33 – 0.36); PPW (0.36 – 0.40); GL (1.40 – 1.43); TL (5.96 – 6.06); CI (95.74 – 100.97); SI (101.47); OI (28.98). Large-sized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 16 – 17 ommatidia. Longitudinal, irregular to vermicular striae on scutum going from an anterior central point towards transcutal suture, interspaces distinguishable. Longitudinal and irregular striae on anepisternum and katepisternum, interspaces distinguishable. Axilla and scutellum with the same sculpture of scutum. Scutoscutellar sulcus inconspicuous. Transverse and regular striae on propodeum (DV). Thick striae (macrosculpture) on mesosoma lateral, irregular to vermiculated-areolated, directed to propodeal spine, interspaces between thicker striae filled with thinner striae (microsculpture). Wings as in Fig. 81 H. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE7FFFAF0A5F1DFFB07FA00.taxon	etymology	Etymology. The epithet mitiae is a Latin noun in the genitive case created by adding the singular Latin genitive case suffix - e to the first name of a female person. This species is named in honor of M. A. Ulysséa’s beloved friend Mítia Heusi Silveira (1984 – 2010), who had her life interrupted by femicide. As an undergraduate student, she developed projects with Fungi and Coleoptera. After university, she worked at FUNAI, a Brazilian governmental agency for the protection of indigenous interests and culture.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE7FFFAF0A5F1DFFB07FA00.taxon	discussion	Comments. Hylomyrma mitiae is similar to H. virginiae and H. sagax. All three are allopatric, but nevertheless occur in northwestern South America. Hylomyrma mitiae has been recorded only in French Guiana (Fig. 87), H. sagax is restricted to southeast Colombia (Fig. 83), whereas H. virginiae has been recorded in Ecuador (both sides of The Andes) and in western Colombia (Fig. 89). Hylomyrma mitiae can be distinguished from H. virginiae (characteristic in parentheses) by the comparatively larger body, TL 5.27 – 5.52 mm, WL 1.41 – 1.52 mm (vs. smaller, TL 4.94 – 5.20 mm, WL 1.24 – 1.40 mm), the gena and laterodorsal region of head are covered with very thin striae between the vermicular to vermiculated-areolated striae (vs. very thin striae superimposed on vermicular to vermiculated-areolated striae), the metanotal groove distinguished by a slight depression (vs. indistinct), the continuous dorsal margin of petiole (vs. discontinuous), and the very prominent and subtriangular subpostpetiolar process (vs. weak and slightly convex). Hylomyrma mitiae is distinguished from H. sagax by the pair of medium sized teeth laterally at the clypeus anterior margin (vs. large teeth), the presence of smooth interspaces between the vermicular striae only on the mesosomal dorsum (vs. on the entire mesosoma), and the subtriangular subpostpetiolar process (vs. convex).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFE7FFFAF0A5F1DFFB07FA00.taxon	distribution	Distribution. This species is only known from French Guiana (Fig. 87). Natural history. Most specimens were found in leaf-litter samples, which suggests that nests are located in fallen logs, rotten wood, between leaves, or inside natural cavities of the superficial soil layers.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFAFFFDF0A5F235FD57F9B4.taxon	description	Figures 55, 56, 83 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFAFFFDF0A5F235FD57F9B4.taxon	materials_examined	Hylomyrma montana Pierce, Branstetter & Longino, 2017: 138 (W). Holotype: COSTA RICA: Limón: Cerro Platano, 9.86732 – 83.24131 ± 20 m, 1130 m [a. s. l.], 18 June 2015, [Project] ADMAC [collection code] # Wm-E- 03 - 1 - 01, cloud forest, ex sifted leaf litter (1 W) (CASENT 0637306) [CASC] [examined by image]. Paratypes: same data as holotype (1 W) (CASENT 0638686) [MCZC] [not examined]; (1 W) (CASENT 0638689) [UCD] [not examined]; (1 W) (CASENT 0638690) [UCR] [not examined]; (1 W) (CASENT 0638691) [USNM] [not examined]; (7 W) (CASENT 0638721, CASENT 0638702, CASENT 0638694, CASENT 0638692, CASENT 0638699, CASENT 0638696, CASENT 0638695) [JTLC] [not examined]; (1 W) (CASENT 0638719 MZSP 67405) [MZSP] [examined].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFAFFFDF0A5F235FD57F9B4.taxon	diagnosis	Worker (Fig. 55 A – C) Diagnosis. Regular and longitudinal striae on head dorsum diverge towards posterior margin, interspaces between thicker striae filled with thinner striae; regular and irregular striae on mesosoma assuming multiple directions, interspaces smooth; dorsal tooth of propodeal lobe acute, ventral tooth blunt; dorsal margin of mesosoma slightly discontinuous, somewhat straight (mesosoma slightly flattened dorsally), promesonotal junction and metanotal groove discernible by a slight depression (LV) and altered sculpture (DV); propodeal spine long; dorsal margin of petiole continuous, convex; ventral surface with transverse striae from petiole spiracle to posterior end of petiole; irregular, longitudinal and anastomosed striae node dorsum; postpetiole smooth; subpostpetiolar process predominantly smooth, weak, convex; profemur posterior surface and protibia extensor surface smooth; striation on tergum of first gastral segment restricted to base, very short striae.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFAFFFDF0A5F235FD57F9B4.taxon	description	QUEEN (first description) (n = 3) (Fig. 56 A – C): HL (1 – 1.02); HW (0.96 – 0.97); ML (0.64 – 0.68); SL (0.72 – 0.74); MOD (0.24 – 0.26); PNW (0.78 – 0.80); WL (1.48 – 1.51); PSL (0.31 – 0.34); PL (0.71 – 0.74); PW (0.28 – 0.30); PPL (0.36 – 0.38); PPW (0.42); GL (1.30 – 1.36); TL (5.49 – 5.62); CI (95.10 – 97); SI (74.22 – 77.08); OI (24.74 – 26.80). Large-sized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 12 – 13 ommatidia. Longitudinal, regular to irregular striae on scutum going from an anterior central point towards transcutal suture, interspaces distinguishable. Longitudinal and mostly irregular striae on anepisternum and katepisternum, interspaces distinguishable. Axilla and scutellum with same sculpture of scutum. Scutoscutellar sulcus inconspicuous. Transverse striae on propodeum (DV). Irregular striae on mesosoma lateral directed in part to propodeal dorsum and in part to propodeal spine, interspaces distinguishable. Wings unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFAFFFDF0A5F235FD57F9B4.taxon	etymology	Etymology. The specific epithet is in reference to the montane distribution of this species.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFAFFFDF0A5F235FD57F9B4.taxon	discussion	Comments. This species is easily distinguished from almost all of its congeners, but it is very similar to H. lispectorae regarding the shape of mesosoma, and the sculpture and shape of the procoxa, petiole, postpetiole, and gaster. The two species are sympatric, co-occuring in two Ecuadorian localities: Centro Científico Río Palenque, and 4 km E of Santo Domingo de los Colorados (Figs. 83, 84). Hylomyrma montana can be distinguished from H. lispectorae (characteristic in parentheses) by the dorsum of head and mesosoma with irregular striae of uniform thickness with interspaces distinguishable (vs. vermicular striae of variable thickness with interspaces indistinguishable), and the somewhat straight dorsal margin of mesosoma (vs. notably convex). The morphological variation displayed by H. montana is associated with the presence of intercastes. Three specimens have an inconspicuous central ocellus, and a more pronounced depression on the metanotal groove. Two specimens (# 17842, # 17840) from Chiriqui-Bocas del Toro, Panama, have more queen-like traits, as larger bodies [HL (1); HW (0.94 – 0.98); ML (0.58 – 0.62); SL (0.68 – 0.70); MOD (0.22 – 0.23); PNW (0.25 – 0.26); WL (1.32 – 1.34); PSL (0.30 – 0.31); PL (0.64 – 0.65); PW (0.25 – 0.26); PPL (0.36); PPW (0.35 – 0.38); GL (1.18 – 1.30); TL (5.09 – 5.26)], the presence of three ocelli, the well-marked promesonotal suture, the differentiated mesonotum (scutum, transcutal suture, prescutellum, and scutellum) and metanotum.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFAFFFDF0A5F235FD57F9B4.taxon	distribution	Distribution. Costa Rica, Ecuador, and Panama (Fig. 83). Natural history. This species inhabits wet forest sites at elevations between 29 and 1800 m. Specimens are frequently found in leaf-litter samples, which suggests that nests are located in leaf-litter, fallen logs, rotten wood, or inside natural cavities of the superficial soil layers. Except for the presence of intercastes, there is no further information available regarding the biology of H. montana.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFAFFFDF0A5F235FD57F9B4.taxon	materials_examined	Additional material examined (40 workers, 15 queens, 5 intercastes): COSTA RICA: Heredia: 16 km SSE La Virgem, 10.26877 - 84.08531 + 200 m, 1110 m, 1. iv. 2001, ALAS # 11 / WF / 01 / all montane wet forest, ex sifted leaf litter (2 I) [MZSP]. ECUADOR: Cotopaxi: Otonga, 00 ° 25 ′ S, 79 ° 00 ′ W, 2 AUG 1977, L. Tapia & P. Ponce, ex monte, B 5, QCAZ I 26128 (1 Q) [UTLP]; same except MN 008 I, LL 4 - P 2 WI (1 Q) [UTLP]; [Los Rios]: Rio Palenque, Univ. Miami Res. Sta., 29 July. 1978, G. J. Umphrey (1 W) (MCZENT 00525538) [MCZC]; Agosto / Septiembre, 1977, 55235, Bosque primario, cuad. cerr, QCAZ I 114015 (1 Q) [UTLP]; Pichincha: Centr. Cient. R. Palenque, 4. i. 1981, Sonia Sandoval col., Bosque primario cerrado, 196 (1 Q) [IHVL]; same except 28. ii. 1981, 279 (1 Q) [IHVL]; 4 km E Santo Domingo de los Colorados, 8. vii. 1976, S. & J. Peck (2 Q) (MCZENT 00524645, MCZENT 00524644) [MCZC]; 4 km E Santo Domingo de los Colorados, 520 m, 22 June 1975, rainfor., B- 304, S. & J. Peck (2 W) (MCZENT 00524657, MCZENT 00524665) [MCZC]; same data (1 W) (MZCENT 00524664) [MZSP]; 20 - 30 km ENE Alluriquin on Chiriboga Rd., 1400 - 1800 m, moss for., B 301, 1975, S. & J. Peck (1 W 1 Q) (MCZENT 00524672, MCZENT 00524671) [MCZC]; Otongachi, 850 m, 0 ° 18 ′ 49 ″ S, 78 ° 57 ′ 15 ″ W, 04 SEP 2009, G. Ramón, KT- 601, LL 8 W 14 (2 W) [UTLP]; same except KT- 1668, LL 8 W 7 (2 W) [UTLP]; KT- 691, LL 8 W 1 (1 W) (MZHY 213) [MZSP]; KT- 691, LL 8 W 1 (1 W) [IHVL]; 04 - AUG- 2008, D. Donoso, KT- 1270, LL 6 - W 2 (2 W 1 Q) [MZSP]; 03 AUG 2011, G. Ramón, VD- 604, LL 12 - PF 18 (1 Q) [MZSP]; 06 AUG 2009, G. Ramón, KT- 1055, GR- 018 (1 Q) [UTLP]; Tinalandia, 16 km SE S. Domingo de los Colorados, 5. vi. 1976, S. & J. Peck (5 W) (MCZENT 00524687, MZCENT 00525525, MZCENT 00524685, MZCENT 00525526, MZCENT 00524686) [MCZC]; same data (1 W) (MZCENT 00525527) [MZSP]; same except 4. vi. 1976 (1 W) (MCZENT 00525483) [MCZC]; 680 m, B- 300, (1 W 1 Q) (MIZA 0021697, MIZA 0021698) [MIZA]; 680 m (1 Q) (MIZA 0021700) [MIZA]; Río Toachi, 900 m, 0.3304033 ° S, 78.9413833 W, 05. Sep. 2003, D. Donoso, KT- 786, N. O W LL- 1, 19 / 26 (2 W, one covered with gold) (MZSP 67433) [MZSP]; same except LL 1 P 11 W 48 h, xc 275, QCAZ I 59348 (1 W 1 Q) [MZSP]. PANAMA: Chiriqui-Bocas del Toro: Cont. Div., 9. vi. 95, R. Anderson, # 17845 (2 W) MZSP]; same data (1 W) [UTEP]; (2 W) [DZUP]; same except # 17842 (8 W 1 I) [MZSP]; # 17840 (2 I) [MZSP]; La Fortuna area, 9. v. 95, R. Anderson, # 17846 (1 W 1 Q) (MZSP 67406) [MZSP]; Veraguas: 6.1 k N Santa Fe, Cerro Tute, 1220 m, 13. vi. 1996, litter extr., wet cloud for., R. Anderson, # 17893 (1 W) [MZSP]; same except # 17891 (1 W) [UTEP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFDFFFFF0A5F294FCF9F894.taxon	description	Figures 57, 80 D, 89 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFDFFFFF0A5F294FCF9F894.taxon	materials_examined	Holotype: GUYANA: [Cuyuni-Mazaruni]: Mt. Ayanganna Falls Camp, 1134 m, 59 ° 57.563 ′ W, 5 ° 22.332 ′ N, 11. x. 2002, T. R. Schultz, J. LaPolla, C. J. Marshall, R. Williams, 1 ° forest, litter sample, JSL 021011 - 01 - LS 06 (1 W) (USNMENT 00413223) [USNM]. Paratypes: same data as holotype except JSL 021011 - 01 - LS 13 (1 W) (USNMENT 00413369) [USNM]; JSL 021011 - 01 - LS 09 (1 W) (USNMENT 00413299) [MZCZ]; JSL 021011 - 01 - LS 07 (1 W) (USNMENT 00413243) [DZUP]; JSL 021011 - 01 - LS 18 (1 W) (USNMENT 00413433) [MZSP 67407]; JSL 021011 - 01 - LS 14 (1 W) (USNMENT 00413373 MZHY 194) [MZSP]; Mt. Ayanganna Base Camp, 732 m, 59 ° 55.486 ′ W, 5 ° 20.063 ′ N, 9. x. 2002, J. S. LaPolla, forest, litter sample, JSL 021009 - 03 - LS 07 (1 W) (USNMENT 00441070) [USNM]; Mt. Ayanganna Cloud Forest, 1300 m, 59 ° 57.969 ′ W, 5 ° 22.483 ′ N, 13. x. 2002, T. R. Schultz, J. LaPolla, C. J. Marshall, R. Williams, litter sample, JSL 021013 - 01 - LS 10 (1 W) (USNMENT 00413807) [USNM]; JSL 021013 - 01 - LS 18 (1 W covered with gold, without postpetiole and gaster) (USNMENT 00413896) [USNM].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFDFFFFF0A5F294FCF9F894.taxon	diagnosis	Diagnosis. Regular and longitudinal striae on head dorsum diverge towards posterior margin, interspaces between striae smooth, striae crest punctuated; mesosoma covered with regular, longitudinal, parallel and thick striae; longitudinal striae on lateral of pronotum and mesepisternum continuing transversely on propodeum; dorsal margin of petiole continuous, strongly convex; subtriangular projection on mesoventral surface of petiole; node rounded with transverse striae on dorsum; subpostpetiolar process smooth, weak, convex; profemur posterior surface and protibia extensor surface mostly smooth, few striae on basal and apical regions; striation on tergum of first gastral segment restricted to base.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFDFFFFF0A5F294FCF9F894.taxon	description	Description. WORKER (n = 4) (Fig. 57 A – C): HL (0.88 – 0.94); HW (0.84 – 0.90); ML (0.57 – 0.60); SL (0.62 – 0.66); MOD (0.20 – 0.24); PNW (0.56 – 0.64); WL (1.14 – 1.24); PSL (0.16 – 0.19); PL (0.52 – 0.59); PW (0.20 – 0.22); PPL (0.32 – 0.34); PPW (0.30 – 0.31); GL (0.98 – 1.02); TL (4.41 – 4.71); CI (94.44 – 96.62); SI (73.33 – 76.47); OI (23.81 – 26.66). Small-sized. Shiny integument. Brownish body, darker gaster, lighter appendices. Thin and branched setae (Fig. 80 D), long to midsized, suberect to subdecumbent; 2 short branches of unequal size from the main axis. Head subquadrate; posterior margin straight. Mandible masticatory margin with 5 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with 8 – 9 striae, longitudinal, regular to irregular, converging to a point on the anterior margin, interspaces distinguishable. Frontal triangle with 3 striae. Short scape, not reaching head posterior margin; apical antennomere of similar size or slightly shorter than previous 3 antennomeres together. Frontal carina straight. Eye drop-shaped, small-sized, larger diameter with 10 ommatidia. Regular and longitudinal striae on head dorsum, divergent towards posterior margin, striae with punctuated crests, interspaces smooth. Head laterodorsal sculpture converges to mandible insertion region; head lateral sculpture converges to eye margin; gena striate, 2 – 3 regular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma covered with regular and longitudinal striae, thicker than head striation, transverse on pronotum anterior region (DV), longitudinal in other regions (DV); longitudinal striae on lateral of pronotum and mesepisternum continuing transversely on propodeum. Promesonotal junction and metanotal groove indistinct. Transverse carina well-marked. Dorsal margin of mesosoma continuous, convex. Propodeal lobe bidentate, dorsal tooth longer and rounded tip; ventral tooth shorter and blunt; dorsal tooth length similar to propodeal spine length. Propodeal spine short, straight (LV), divergent (DV), sculptured on base. Procoxa striae of uniform thickness; irregular and transverse striae on C 2 and C 3. Profemur and protibia mostly smooth, few striae on basal and apical regions. Dorsal margin of petiole continuous, strongly convex; subtriangular projection on mesoventral surface of petiole; ventral surface smooth; first third of dorsal surface smooth, transverse striae on posterodorsal surface continuing on lateral surface forming arches. Longitudinal striae on postpetiole; subpostpetiolar process smooth, weak, convex. Tergum first gastral segment striation similar to postpetiole striae, short, restricted to base, striae length equivalent to 1 / 2 of postpetiole length; sternum smooth. QUEEN Unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFDFFFFF0A5F294FCF9F894.taxon	etymology	Etymology. The epithet peetersi is a Latin noun in the genitive case created by adding the singular Latin genitive case suffix - i to the last name of a male person. This species is named in honor of our friend and colleague Dr. Christian Paul Peeters (1956 – 2020), a Belgian born researcher that worked at the Université Pierre et Marie Curie in Paris, and who contributed enormously to Myrmecology in general and, specifically to the present work, in understanding the Hylomyrma intercastes. Peeters generously received the first author in his lab for some very profitable months in 2015.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFDFFFFF0A5F294FCF9F894.taxon	discussion	Comments. The diagnostic characters of this species are sufficient to distinguish it from most of its congeners. Hylomyrma peetersi is, however, more similar to H. margaridae and H. villemantae. All three species display a strongly convex petiole, with indistinguishable petiolar peduncle and node, and the body mainly covered with regular and longitudinal striae. Hylomyrma peetersi and H. margaridae can be easily distinguished from H. villemantae (characteristic in parenthesis) by the striae on the head dorsum with punctuated crests (vs. with smooth crests), the mesosoma longitudinally striate (vs. with few elliptic and concentric striae), the shorter propodeal spine (vs. longer), the armed mesoventral surface of petiole (vs. unarmed), and the longitudinal striae on the dorsum of postpetiole and gaster (vs. smooth). Hylomyrma peetersi can be distinguished from H. margaridae in the striae on the laterals of mesosoma that are directed transversely to the propodeal dorsum (vs. directed to the propodeal spine), the mesoventral surface of petiole armed with a subtriangular projection (vs. a conspicuous spine), the petiolar dorsum entirely covered with transverse striae (vs. with both transverse and longitudinal striae), and the weak subpostpetiolar process (vs. prominent). All three species are allopatric; H. villemantae is recorded in Brazil (BA) (Fig. 86), whereas H. peetersi and H. margaridae occur in relatively close areas in northern South America (French Guiana, Guyana, and Venezuela) (Fig. 89).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFDFFFFF0A5F294FCF9F894.taxon	distribution	Distribution. The specimens were collected in areas near the Guyana - Venezuela border (Fig. 89). Natural history. This species inhabits the leaf-litter of forested areas at elevations between 732 and 1200 m.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFFDFFFFF0A5F294FCF9F894.taxon	materials_examined	Additional material examined (2 workers): VZLA [VENEZUELA]: Bolívar: km 114 El Dorado to Santa Elena, 12. viii. 1986, 1000 m, 6 ° 01 ′ N, 61 ° 24 ′ W, P. S. Ward [leg.], # 8548 - 16, sifted litter (leaf mold, rotten wood), rainforest (1 W) [MIZA]; same except 1200 m (1 W) [MIZA].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF0FFF1F0A5F494FA85FAE8.taxon	description	Figures 58, 80 F, 80 K, 85 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF0FFF1F0A5F494FA85FAE8.taxon	materials_examined	Hylomyrma plumosa Pierce, Branstetter & Longino, 2017: 138 (W). Holotype: COSTA RICA: Limón: Res. Biol. Hitoy-Cerere, 9.66480 - 83.02346 ± 10 m, 250 m [a. s. l.], 10 June 2015, [Project] ADMAC [collection code] # Wa-E- 02 - 2 - 38, tropical rainforest, ex sifted leaf litter (1 W) (CASENT 0638700) [CASC] [examined by image]. Paratypes: same data as holotype (1 W) (CASENT 0638687) [MCZC] [not examined]; (1 W) (CASENT 0638688) [UCD] [not examined]; (1 W) (CASENT 0638693) [UCR] [not examined]; (1 W) (CASENT 0638697) [USNM] [not examined]; (1 W) (CASENT 0638703 MZSP 67408) [MZSP] [examined]; (8 W) (CASENT 0638720, CASENT 0638701, CASENT 0638725, CASENT 0638698, CASENT 0638700, CASENT 0638722, CASENT 0638724, CASENT 0638735) [JTLC] [not examined].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF0FFF1F0A5F494FA85FAE8.taxon	diagnosis	Worker (Fig. 58 A-C) Diagnosis. Mesosoma and petiole covered with thin striae (microsculpture) superimposed on irregular and thick striae (macrosculpture), interspaces between thinner striae indistinguishable; conspicuous and trifid setae.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF0FFF1F0A5F494FA85FAE8.taxon	description	QUEEN Unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF0FFF1F0A5F494FA85FAE8.taxon	etymology	Etymology. The specific epithet is a reference to the distinctive branched setae of this species.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF0FFF1F0A5F494FA85FAE8.taxon	discussion	Comments. Hylomyrma plumosa is identified based on the conspicuous and trifid setae (Fig. 80 K). This species resembles H. versuta, described by Kempf (1973) as “ long hairs, pointed at tip, gently curved on petiole and postpetiole ”. However, the types of H. versuta and additional examined material comprise specimens with unbranched and branched setae. The branched setae of H. versuta, mostly present on the petiole, postpetiole, and gaster, are composed of a variable number of short branches with similar size (Fig. 80 F). These branched setae are not conspicuous as in H. longiscapa, H. plumosa, and H. transversa, being better observed in SEM images. Hylomyrma plumosa and H. versuta are locally sympatric (Figs. 85, 88), having been collected in the same sample (ALAS # 03 / WF / 02 /) near La Virgen (300 m), and in different samples (H. plumosa # AMI- 1 - W- 006 - 01, and H. versuta # AMI- 1 - W- 006 - 05) from the La Selva Biological Station (50 m); both locations in Heredia Province, Costa Rica. The two species have very similar body sculpture. The difference between them is very subtle: the propodeum is laterally covered by thin striae (microsculpture) with indistinguishable interspaces superimposed on irregular and thick striae (macrosculpture) in Hylomyrma plumosa, whereas H. versuta presents thin striae (microsculpture) with indistinguishable interspaces between irregular and thick striae (macrosculpture). Initially, we understood that H. plumosa and H. versuta belonged to a continuum of variation due to their body sculpture similarity, variable setae, and co-occurrence. But molecular data show H. plumosa as sister to the H. cf. dentiloba sp. 2 (the true H. dentiloba) – H. versuta clade (Pierce et al. 2017). In this scenario, we have decided to maintain all three species as valid until additional evidence is gathered. Further investigative approaches using UCEs should include representatives of all the variation found in these taxa (or nominal species).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF0FFF1F0A5F494FA85FAE8.taxon	distribution	Distribution. Hylomyrma plumosa occurs in Costa Rica (Fig. 85). Natural history. This species occurs in wet forests at elevations between 50 and 300 m. Seventeen specimens were collected in the leaf-litter, which suggests that this species nests in fallen logs, rotten wood, between leaves, or inside natural cavities of the superficial soil layers.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF0FFF1F0A5F494FA85FAE8.taxon	materials_examined	Additional material examined (4 workers): COSTA RICA: Heredia: 11 km ESE La Virgem, 10.35, - 84.05 + 2 km, 300 m, 10. iv. 2004, ALAS # 03 / WF / 02 / all montane wet forest, ex sifted leaf litter (1 W) [MZSP]; La Selva, E. O Wilson col., 19. iii. 85 (1 W) [MCZC]; La Selva Biological Station, 10.41639, - 84.02 + 500 m, 50 m, 16. iii. 2004, TEAM # AMI- 1 - W- 006 - 01, mature wet forest, ex sifted leaf litter (2 W) (MZSP 67470, MZHY 84) [MZSP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF1FFF5F0A5F15DFD41FE80.taxon	description	Figures 59, 60, 61, 90 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF1FFF5F0A5F15DFD41FE80.taxon	diagnosis	Diagnosis. Large-sized; regular to irregular, longitudinal and thick striae on head dorsum diverge towards posterior margin, interspaces between striae smooth; propodeal spine long; a pair of large and blunt teeth laterally on anterior margin of clypeus; dorsal margin of petiole continuous, convex; subpostpetiolar process weak, convex; profemur mostly covered with irregular to regular transverse striae; protibia entirely covered with regular striae; very long striae on tergum of first gastral segment; semicircular striae on sternite of first gastral segment covering the midbasal and lateral regions.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF1FFF5F0A5F15DFD41FE80.taxon	description	Redescription. WORKER (n = 4) (Fig. 59 A – C): HL 1.22 (1.20 – 1.24); HW 1.20 (1.18 – 1.22); ML 0.82 (0.82); SL 0.90 (0.90 – 0.94); MOD 0.32 (0.30 – 0.34); PNW 0.82 (0.82 – 0.84); WL 1.54 (1.54 – 1.58); PSL 0.42 (0.36 – 0.43); PL 0.75 (0.75 – 0.77); PW 0.28 (0.26 – 0.28); PPL 0.42 (0.40 – 0.42); PPW 0.42 (0.40 – 0.42); GL 1.58 (1.36 – 1.59); TL 6.33 (6.10 – 6.40); CI 98.36 (96.72 – 100); SI 75 (75 – 78.81); OI 26.66 (25.42 – 27.87). Large-sized. Shiny integument. Brownish body, lighter petiole, postpetiole and leg. Thin and unbranched setae, long to midsized, erect to subdecumbent. Head subquadrate; posterior margin slightly concave at middle. Mandible masticatory margin with 6 teeth. Anterior margin of clypeus slightly concave medially, with a pair of large and blunt teeth laterally; median area of clypeus centrally striate, with 10 regular and longitudinal striae converging to a point on the anterior margin, interspaces distinguishable. Frontal triangle with 1 sagittal stria. Short scape, not reaching head posterior margin; apical antennomere slightly shorter than previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye drop-shaped, large-sized, larger diameter with 16 ommatidia. Head dorsum with regular to irregular, longitudinal and thick striae, divergent towards posterior margin, interspaces smooth. Longitudinal striae on head lateral and laterodorsal regions converge to mandible insertion; gena striate, around 5 – 7 regular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma sculpture similar to head dorsum, striae of uniform thickness assuming multiple directions; transverse striae on pronotum anterior region (DV) continuing on lateral surface, longitudinal striae on other regions of pronotum and metanotal groove region (anteriorly to transverse carina); irregular striae on mesonotum assuming more than one direction. Promesonotal junction and metanotal groove indistinct. Transverse carina inconspicuous. Dorsal margin of mesosoma continuous, convex. Propodeal lobe bidentate, dorsal tooth slightly longer and acute than the shorter and blunt ventral tooth; dorsal tooth length 1 / 2 of propodeal spine length. Propodeal spine long, straight (LV), divergent (DV), sculptured. Thin striae of uniform thickness on procoxa; irregular and transverse striae on C 2 and C 3. Profemur mostly covered with irregular to regular transverse striae. Protibia entirely covered with regular striae. Dorsal margin of petiole continuous, convex; sculpture similar to mesosoma; mostly covered with transverse striae; ventral surface smooth. Regular and longitudinal striae on postpetiole; subpostpetiolar process striae restricted to lateral region, midbasal region smooth; subpostpetiolar process weak, convex. First gastral segment striation similar to postpetiole striae; length of long striae on tergum similar to postpetiole length; semicircular striae on sternite covering the midbasal and lateral regions. QUEEN (first description) (n = 1) (Fig. 60 A – C): HL 1.20; HW 1.24; ML 0.84; SL 0.90; MOD 0.34; PNW 1; WL 1.78; PSL 0.38; PL 0.83; PW 0.32; PPL 0.44; PPW 0.47; GL 1.66; TL 6.75; CI 103.33; SI 72.58; OI 27.41. Largesized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 18 ommatidia. Longitudinal, regular to irregular striae on scutum going from an anterior central point towards transcutal suture, interspaces distinguishable. Longitudinal and mostly regular striae on anepisternum and katepisternum, interspaces distinguishable. Axilla and scutellum with same sculpture of scutum. Scutoscutellar sulcus inconspicuous. Transverse striae on propodeum (DV). Lateral of mesosoma with mostly irregular striae directed to propodeal spine, interspaces distinguishable. Wings unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF1FFF5F0A5F15DFD41FE80.taxon	etymology	Etymology. The specific epithet praepotens (Latin, prae = pre-, before, very and potens = powerful) is possibly a reference to the head sculpture or to the long clipeal teeth of this species.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF1FFF5F0A5F15DFD41FE80.taxon	discussion	Comments. Hylomyrma praepotens, the largest species of the genus, is similar to H. immanis, both have a long propodeal spine, and an elongated and continuous dorsal margin of petiole, but H. praepotens is easily distinguished by the shiny integument (subopaque in H. immanis), and the body covered by thick striae with distinguishable interspaces (thin striae with indistinguishable interspaces in H. immanis). Hylomyrma immanis is more easily sampled than H. praepotens, the former has a broad distribution in the center-north of South America (Fig. 88), whereas the latter is only known from two localities in Colombia, two in Ecuador, and one in Brazil (Fig. 90). Both species cooccur in a locality 7 km NW of Letícia, Amazonas, Colombia. Four type specimens, identified as workers by Kempf (1973), have an inconspicuous central ocellus, and a darker area on the head dorsum coincident with the ocelli location in winged queens (Fig. 61). They also have the mesosoma larger and more convex than in workers. Due to these characters, we rcognized these specimens as intercastes.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF1FFF5F0A5F15DFD41FE80.taxon	distribution	Distribution. The few known specimens were collected in Brazil, Colombia, and Ecuador (Fig. 90). Natural history. The type specimens were collected with Berlese-Tüllgren funnels in Colombia, and the unique specimen from Brazil was collected with winkler extractors, suggesting that colonies are located in leaf-litter or in superficial soil layers. Except for the presence of intercastes (4 paratypes), there is no further information available regarding H. praepotens biology. Additional material examined (12 workers, 1 queen): BRAZIL: MA [Maranhão]: São Francisco do Brejão, 05 ° 17 ′ 19 ″ S, 47 ° 15 ′ 01.7 ″ W, 01 - 09. vi. 2005, Silva R. R. & Feitosa R. M. cols., Winkler 10 (1 W) (MZHY 202) [MZSP]. ECUADOR: Napo: Limoncocha, 250 m, 25. vi. 1976, B- 355, S. & J. Peck (11 W) [MCZC]; same except 18. vi. 1976, B- 348 (1 Q) (MCZENT 00524675) [MCZC].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF5FFF7F0A5F5A8FE77FB1C.taxon	description	Figures 62, 80 J, 88 (map) Holotype: BRAZIL: MG [Minas Gerais]: Santana do Riacho, Serra do Cipó, APA [Environmental Protection Area] Morro da Pedreira, 19 ° 17 ′ 50.11 ″ S, 43 ° 35 ′ 39.69 ″ W, 1300 m, 16. i. 2012, campo rupestre, pitfall-solo, T. C. Lana col., Q 16 - X-V- 4 [Area code - Xeric environment - Transect number - Point number] (1 W) (MZSP 67412) [MZSP]. Paratypes: same data as holotype except abril / 2011, Q 16 - X-VI- 2 (1 W) (MZSP 67413) [MZSP]; Q 16 - X-VI- 4 (1 I) (MZSP 67414) [MZSP]; abril / 2013, Q 16 - X-V- 4 (1 I) (MZSP 67415) [MZSP]; 19 ° 13 ′ 48.89 ″ S, 43 ° 34 ′ 35.19 ″ W, 1100 m, x. 2011, C 11 - X-I- 2 (1 I) (MZSP 67416) [MZSP]; C 11 - X-I- 4 (1 I) (MZSP) [MZSP 67417]; PARNA [National Park] Serra do Cipó, 19 ° 15 ′ 49.92 ″ S, 43 ° 32 ′ 04.35 ″ W, 1400 m, 25 - 30. iv. 2011, AP-X-III- 1, pitfall-solo (1 W covered with gold) (MZSP 67418) [MZSP]; same except AP-X-III- 2 (1 I) (MZSP 67419) [MZSP]; AP-X-III- 3 (1 I) (MZSP 67420) [MZSP]; jul / 2011, AP-X-III- 2 (1 I) (MZSP 67421) [MZSP]; AP-X-II- 3 (1 I) (MZSP 67422) [MZSP]; x. 2011, AP-X-III- 4 (2 I) (MZSP 67423, MZHY 60) [MZSP]; i. 2012, AP-X-III- 1 (1 I) (MZSP 67424) [MZSP]; AP-X-III- 3 (1 I) (MZSP 67425) [MZSP]; AP-X-III- 4 (1 I) (MZSP 67426) [MZSP]; iv. 2012, AP-X-III- 1 (1 W) [DZUP]; AP-X-III- 2 (1 W) [USNM]; AP-X-III- 3 (1 I) (MZSP 67427) [MZSP]; julho / 2012, AP-X-III- 2 (1 W) [MCZC]; AP-X-III- 4 (1 I) (MZSP 67428) [MZSP]; x. 2012, AP-X-III- 2 (1 I) [MCZC]; janeiro / 2013, AP-X-III- 1 (1 I) [CASC]; AP-X-III- 2 (2 I) [DZUP]; AP-X-III- 3 (2 I) [IHVL, JTCL]; abril / 2013, AP-X-III- 3 (1 I) [USNM]; julho / 2013, AP-X-III- 4 (1 I) [CASC].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF5FFF7F0A5F5A8FE77FB1C.taxon	diagnosis	Diagnosis. Body black; regular striae on head dorsum diverge towards posterior margin; rugose striae on mesosoma and petiole; promesonotal junction and metanotal groove discernible by a slight depression (LV) and altered sculpture (VD); propodeal spine length similar to dorsal tooth of propodeal lobe; dorsal margin of petiole discontinuous; transverse striae on node ventral surface; profemur posterior surface smooth; striae on protibia extensor surface weakly marked; striation on tergum of first gastral segment restricted to base; thick branched setae flattened in its final half, with several branches.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF5FFF7F0A5F5A8FE77FB1C.taxon	description	Description. WORKER (n = 3) (Fig. 62 A – C): HL (1.16 – 1.24); HW (1.06 – 1.18); ML (0.64 – 0.72); SL (0.80 – 0.86); MOD (0.28 – 0.31); PNW (0.72 – 0.82); WL (1.34 – 1.48); PSL (0.18 – 0.20); PL (0.62 – 0.68); PW (0.28 – 0.34); PPL (0.34 – 0.40); PPW (0.36 – 0.44); GL (1.22 – 1.42); TL (5.32 – 5.78); CI (91.38 – 97.45); SI (71.30 – 75.47); OI (26.08 – 26.41). Large-sized. Shiny integument. Black body, brownish mandibles, trochanter and tarsus. Mostly thin and unbranched setae, midsized to short, suberect to subdecumbent; thick branched setae flattened in its final half, with several branches, mainly in the postpetiole and gaster (Fig. 80 J). Head subquadrate; posterior margin slightly concave at middle. Mandible masticatory margin with 5 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with 7 – 8 regular, longitudinal and thick striae converging to a point on the anterior margin, interspaces between striae smooth or thinner striae between thicker striae. Frontal triangle with 1 sagittal stria. Short scape, not reaching head posterior margin; apical antennomere length 1 / 4 shorter than previous 3 antennomeres together. Frontal carina straight. Eye drop-shaped, midsized, larger diameter with 13 ommatidia. Regular, longitudinal striae on head dorsum, divergent towards posterior margin, interspaces smooth. Thin, regular to irregular striae on head lateral and laterodorsal region near eye, interspaces indistinguishable; striae on laterodorsal region near eye converging to mandible insertion region; striae on lateral region converging to eye margin; laterodorsal region near frontal carina rugose; gena striate, 2 – 3 regular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma mostly covered with rugose striae of uniform thickness; interval between striae smooth; transverse striae on propodeum dorsum and on slope surface. Promesonotal junction and metanotal groove discernible by a slight depression (LV) and altered sculpture (DV). Transverse carina indistinct to inconspicuous. Dorsal margin of mesosoma slightly discontinuous, convex. Propodeal lobe bidentate, dorsal tooth slightly longer and acute than shorter and blunt ventral tooth; dorsal tooth length similar to propodeal spine length. Propodeal spine short, straight (LV), divergent (DV), sculptured. Transverse striae of uniform thickness on procoxa; irregular and transverse striae on C 2 and C 3. Transverse striae on profemur anterior, dorsal and posterior surfaces; ventral surface smooth. Protibia extensor surface entirely striate. Dorsal margin of petiole discontinuous. Subtriangular projection on petiole mesoventral surface. Node with transverse striae on ventral and anterior surfaces, with interspaces distinguishable; rugose striae on dorsal and latero-dorsal surfaces, interspaces sculptured; irregular and longitudinal striae on latero-ventral surface. Postpetiole and subpostpetiolar process with regular and longitudinal striae; subpostpetiolar process weak, convex. First gastral segment striation similar to postpetiole striae; striae on tergum as long as postpetiole length; sternite striation restricted to laterobasal region. QUEEN Unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF5FFF7F0A5F5A8FE77FB1C.taxon	etymology	Etymology. The epithet primavesi is a non-Latin proper noun used in apposition. This species is named in honor of Ana Maria Primavesi (1920 – 2020), an agronomist, pioneer in soil sciences and especially in the ecological management of tropical soils in Brazil. She provided the scientific basis for the development of tropical organic agriculture.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF5FFF7F0A5F5A8FE77FB1C.taxon	discussion	Comments. Hylomyrma primavesi is unique among its congeners in its blackish body. Nonetheless, its body sculpture resembles that of H. balzani. Hylomyrma primavesi can be distinguished from H. balzani (characteristic in parentheses) by the larger body, TL 5.32 – 5.78 mm, WL 1.34 – 1.48 mm (vs. smaller, TL 4.27 – 5.23 mm, WL 1.10 – 1.39 mm), the distinct promesonotal junction and metanotal groove (vs. both indistinct). Both species occur in Brazil, whereas H. primavesi is known only from higher elevations in Serra do Cipó, MG (Fig. 88), H. balzani has a broad distribution, occurring mainly in eastern portions of Brazil from north to south (Fig. 84). Half of all specimens examined have a central ocellus (n = 6) or a more or less pronounced depression on the region where winged queens present the central ocellus (n = 11). Moreover, most of the specimens have the promesonotal junction and metanotal groove distinguishable by a slight depression or altered sculpture. This morphological variation is associated with intercastes at least in other Hylomyrma species with intercastes. Winged queens are not known in this species, thus these intercastes may perform reproduction. Specimens identified as workers do not present ocelli or ocellar depression, nor the conspicuous promesonotal junction and metanotal groove. One of the workers studied presents the laterals of the mesosoma flattened. This was interpreted as a deformity.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF5FFF7F0A5F5A8FE77FB1C.taxon	distribution	Distribution. This species is only known from Santana do Riacho (Serra do Cipó National Park and APA Morro da Pedreira), Minas Gerais, Brazil (Fig. 88). Natural history. All specimens were collected with winkler extractors and pitfall traps at elevations between 1100 and 1400 m in areas of “ Campos rupestres ”. Those relatively high-elevation areas are characterized by savannas and grasslands, with plants and outcrops. Except for the presence of worker-like intercastes, there is no further information available regarding the biology of H. primavesi. The winged queen of this species is unknown, then perhaps the specimens determined here as intercastes are responsible for reproduction. Additional material examined (2 workers, 4 intercastes, 2 indeterminates): BRAZIL: MG [Minas Gerais]: Santana do Riacho, Serra do Cipó, APA Morro da Pedreira, 19 ° 17 ′ 50.11 ″ S, 43 ° 35 ′ 39.69 ″ W, 1300 m, 16. i. 2012, campo rupestre, pitfall-solo, T. C. Lana col., Q 16 - X-III- 2 (1 W, bad mesosoma formation; 1 indet., without head) [MZSP]; same except jul / 2011, Q 16 - X-VI- 4 (1 W, without postpetiole and gaster) [MZSP]; out / 2011, Q 16 - X-III- 4 (1 indet.) [MZSP], Q 16 - X-VI- 2 (1 I) [MZSP]; PARNA Serra do Cipó, 19 ° 15 ′ 49.92 ″ S, 43 ° 32 ′ 04.35 ″ W, 1400 m, abril / 2013, AP-X-III- 3, pitfall-solo (1 I) [MZSP]; same except janeiro / 2013, AP-X-III- 3 (1 I) [JTLC]; julho / 2012, AP-X-III- 1 (1 I) [MZSP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF7FFCBF0A5F149FD5EFEF8.taxon	description	Figures 63, 64, 86 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF7FFCBF0A5F149FD5EFEF8.taxon	diagnosis	Diagnosis. Regular to slightly irregular and longitudinal striae on head dorsum diverge towards posterior margin, interspaces between striae smooth; striae crest punctuated; striae on mesosoma assuming multiple directions, interspaces indistinguishable; pronotum posterior region, mesonotum, and metanotal groove region with semielliptical to V-shaped striae; dorsal margin of petiole continuous, convex; subtriangular projection on mesoventral surface of petiole; subpostpetiolar process well-developed, subtriangular; profemur and protibia entirely covered with striation; striation on tergum of first gastral segment restricted to base; sternite striation of first gastral segment covering the laterobasal region.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF7FFCBF0A5F149FD5EFEF8.taxon	description	Redescription. WORKER (n = 5, exceptionally, for this species, the measures presented outside the parentheses refer to the paratype examined [NHMB]) (Fig. 63 A – C): HL 0.94 (0.94 – 1.09); HW 0.87 (0.87 – 1.04); ML 0.60 (0.60 – 0.70); SL 0.67 (0.67 – 0.76); MOD 0.17 (0.17 – 0.26); PNW (0.64 – 0.72); WL 1.30 (1.30 – 1.52); PSL 0.25 (0.25 – 0.32); PL 0.62 (0.62 – 0.75); PW 0.29 (0.29 – 0.32); PPL 0.27 (0.27 – 0.34); PPW 0.30 (0.30 – 0.38); GL 0.97 (0.97 – 1.20); TL 4.71 (4.71 – 5.60); CI 93.33 (93.33 – 101.96); SI 77.14 (73.07 – 76.09); OI 20 (20 – 25). Medium to large-sized. Shiny integument. Brownish body, darker gaster, lighter appendices. Thin and unbranched setae, long to short, suberect to subdecumbent. Head subquadrate; posterior margin straight to slightly concave at middle. Mandible masticatory margin with 5 teeth. Labial palp 3 - articulated; maxillary palp 4 - articulated. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with 10 – 12 regular to irregular and longitudinal striae. Fontal triangle with 2 striae. Short scape, not reaching head posterior margin; apical antennomere slightly shorter than previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye drop-shaped, midsized, larger diameter with 11 ommatidia. Longitudinal, regular to slightly irregular striae on head dorsum diverge towards posterior margin, striae with punctuated crests, interspaces smooth, striae. Striation on head lateral and laterodorsal regions converge to mandible insertion; gena striate, 3 – 4 regular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Irregular to regular striae on mesosoma dorsum assuming multiple directions, interspaces indistinguishable; transverse striae on pronotum anterior region (DV) continuing on lateral surface; pronotum posterior region, mesonotum, and metanotal groove region (immediate anterior to transverse carina) with semi-elliptical to V-shaped striae; striae on lateral of pronotum continuing sinuously on mesepisternum and propodeum lateral, and transversely on propodeum. Promesonotal junction and metanotal groove indistinct. Transverse carina inconspicuous. Dorsal margin of mesosoma continuous, convex. Propodeal lobe bidentate, dorsal tooth longer and more acute than the shorter and blunt ventral tooth; dorsal tooth slightly shorter than propodeal spine. Propodeal spine midsized, straight (LV), divergent (DV), sculptured on base. Thin striae of uniform thickness on procoxa; irregular and transverse striae on C 2 and C 3. Profemur covered with regular and transverse striae. Protibia extensor surface entirely covered with regular and longitudinal striae. Dorsal margin of petiole continuous, convex; subtriangular projection on mesoventral surface; dorsal and lateral surfaces covered with vermicular striae of variable thickness, interspaces indistinguishable; ventral surface smooth. Regular and longitudinal striae on postpetiole and subpostpetiolar process, interspaces indistinguishable; subpostpetiolar process well-developed, subtriangular. First gastral segment striation similar to postpetiole striae; short striae on tergum, shorter than postpetiole length; sternite striation covering the laterobasal region. QUEEN (first description) (n = 3) (Fig. 64 A – C): HL (1.02 – 1.20); HW (0.98 – 1.22); ML (0.68 – 0.80); SL (0.72 – 0.88); MOD (0.26 – 0.32); PNW (0.84 – 0.98); WL (1.56 – 1.88); PSL (0.30 – 0.36); PL (0.75 – 0.90); PW (0.34 – 0.40); PPL (0.39 – 0.40); PPW (0.40 – 0.46); GL (1.20 – 1.46); TL (5.63 – 6.64); CI (94.23 – 101.66); SI (72.13 – 74); OI (26 – 26.53). Large-sized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 15 ommatidia. Longitudinal, regular to irregular striae on scutum going from an anterior central point towards transcutal suture, interspaces distinguishable. Longitudinal and mostly regular striae with mostly uniform thickness on anepisternum and katepisternum, interspaces distinguishable. Axilla and scutellum with same sculpture of scutum. Scutoscutellar sulcus inconspicuous. Transverse striae on propodeum (DV). Lateral of mesosoma with irregular striae of variable thickness directed to propodeal spine, interspaces distinguishable. Wings unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF7FFCBF0A5F149FD5EFEF8.taxon	etymology	Etymology. The specific epithet was named after Regina Pestalozzi, Kutter’s daughter.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF7FFCBF0A5F149FD5EFEF8.taxon	discussion	Comments. Hylomyrma reginae is unlikely to be confused with any other congener because of its very prominent and subtriangular subpostpetiolar process. Two specimens collected in Upper Takutu-Upper Essequibo, Guyana (USNMENT 00688778, USNMENT 00689009) were designated as intercaste because of the larger body measures [HL (1.16); HW (1.16 – 1.17); ML (0.76 – 0.78); SL (0.82 – 0.84); MOD (0.27 – 0.28); PNW (0.83 – 0.84); WL (1.64); PSL (0.34 – 0.35); PL (0.82 – 0.85); PW (0.36 – 0.38); PPL (0.36); PPW (0.40); GL (1.30); TL (6.04 – 6.09)], and for having one central and inconspicuous ocellus.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFF7FFCBF0A5F149FD5EFEF8.taxon	distribution	Distribution. All known specimens were collected in northern South America (Brazil, French Guiana, Guyana, and Suriname) (Fig. 86). Natural history. This species is recorded from tropical rainforests (in the Amazon and in Inselberg forests; primary and secondary forests) at elevations between 60 and 1051 m. Specimens are frequently found in leaf-litter samples, which suggests that nests are located in fallen logs, rotten wood, between leaves, or inside natural cavities of the superficial soil layers. Except for the presence of intercastes, there is no further information available regarding the biology of H. reginae. Additional material examined (23 workers, 3 queens, 2 intercastes): BRAZIL: AM [Amazonas]: Manaus, Dimona Station, INPA, 100 ha Plot, 130 m, 60.09414 ° ′ W, 2.34121 ° ′ S, 16. i. 2009, J. Sosa-Calvo, rainforest, leaf litter, Winkler sample, JSC 090116 - LS 05 (1 W) (USNMENT 00447161) [USNM]; same except JSC 090116 - LS 06 (1 W) (USNMENT 00754291) [USNM]; Pres [idente] Figueiredo, Terra-firme, Ig. Poraque, 1 ° 54 ′ 57 ″ S, 59 ° 26 ′ 94 ″ W, 23 / 04 / 94, Mata primária, Arm. De solo, Queiroz col. (1 W) [MPEG]; Terra Firme, ZF- 02, km 10, capoeira, 02 ° 34 ′ S, 60 ° 06 ′ W, M. O. de A. Ribeiro coll., 01. viii. 1990, capoeira, ZF- 2 / solo, Extração Kempsom, Camada 1, Reg 18, Prof 0 - 3.5 cm (1 W) [INPA]; MA [Maranhão]: São Francisco do Brejão, 05 ° 17 ′ 19 ″ S, 47 ° 15 ′ 01.7 ″ W, 01 - 09. vi. 2005, Silva R. R. & Feitosa R. M. cols., Winkler 1 (2 W) (MZHY 200) [MZSP]; PA [Pará]: Marituba, Mata-Winkler, 1 ° 22 ′ S, 48 ° 20 ′ W, 22. x. 2004, Santos J. R. M., 17 (1 W) [CEPLAC]; Utinga, tract. nr. Belém, BF- 19, Aug. 14.1962, P. F. Darlington (1 W) [MZSP]; same data (1 W) [MCZC]; RO [Rondônia]: Porto Velho, Área Caiçara, C 3 P 3, 09 ° 26 ′ 46.8 ″ S, 64 ° 49 ′ 31.1 ″ W, 04 - 18. ix. 2012, Vicente, R. E. & Oliveira, J. cols. (2 W) [DZUP]. FRENCH GUIANA: [Cayenne]: Kaw. Mt., Amazon Nature Lodge, 950 ′, 52 ° 12.349 ′ W, 4 ° 33.426 ′ N, T. Schultz, U. Mueller, J. LaPolla, 2 ° forest, leaf litter, Winkler sample, TRS 050719 - 01 - LS 04 (2 W) (USNMENT 00537919, USNMENT 00537922) [USNM]; Nouragues Station, IN 2 - Inselberg For., 04 ° 05 ′ N, 52 ° 40 ′ W, ix. 2009, winkler, Sarah Groc & al, # 5630, IN 2 Tr 2 W 10 (1 W) [MZSP]. GUYANA: [Cuyuni-Mazaruni]: Calm Water Creek, along Essequibo River nr. Bartica, 58 ° 37.16 ′ W, 6 ° 28.06 ′ N, 24. ix. 2002, J. S. LaPolla, 1 ° forest, litter sample, = 020923 - 8, JSL 020923 - 01 - LS 04 (2 W) (USNMENT 00442164, USNMENT 00442165) [USNM]; same except = 020923 - 9, JSL 020923 - 01 - LS 05 (1 W) [USNM USNMENT 00442185]; [Potaro-Siparuni]: Iwokrama For. Res., Whitewater Camp, 60 m, 58 ° 50.992 ′ W, 4 ° 43.89 ′ N, 5. xi. 2002, J. S. LaPolla et al., 1 ° forest, litter sample, = 021105 - 1 - LS 20, JSL 021105 - 01 - LS 20 (1 W 1 Q) (USNMENT 00413113, USNMENT 00413112) [MZSP]; same except = 021105 - 1 - LS 11, JSL 021105 - 01 - LS 11 (1 Q) (USNMENT 00412448) [USNM]; Upper Takutu-Upper Essequibo: Acarai Mts., nr. New Romeo Camp, 1050 m, 58 ° 57.867 ′ W, 1 ° 20.048 ′ N, 14. x. 2006, T. R. Schultz, J. Sosa-Calvo, C. J. Marshall, R. Williams, 1 ° forest, leaf litter, Winkler sample, JSC 061014 - LS 03 (1 Q 1 I, intercaste covered with gold) (USNMENT 00688795, USNMENT 00688778) [USNM]; same except 1051 m, 58 ° 57.885 ′ W, 1 ° 20.095 ′ N, JSC 061014 - LS 05 (1 I) (USNMENT 00689009) [USNM]. SURINAME: Sipaliwini: Lely Mts., 560 m, 55 ° 13 ′ 48.65 W, 4 ° 27 ′ 2.45 N, 29. x. 2005, J. Sosa-Calvo & R. Badal, 1 ° forest, litter sample, nr. creek, JSC 051029 - 01 - LS 09 (1 W) (USNMENT 00446454) [USNM]; same except JSC 051029 - 01 - LS 08 (1 W) (USNMENT 00446434) [IHVL]; JSC 051029 - 01 - LS 05 (1 W) (USNMENT 00446224) [USNM]; same data (1 W) (USNMENT 00446225) [MCZC]; 619 m, 54 ° 45 ′ 21.96 W, 4 ° 15 ′ 10.62 N, 26. x. 2005, J. Sosa-Calvo & R. Badal, 1 ° forest, litter sample, tall forest, rocky soil, JSC 051026 - 01 - LS 04 (1 W) (USNMENT 00447161) [USNM].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFCBFFC4F0A5F56DFB6AFBBC.taxon	description	Figures 2 A, 65, 66, 67, 68, 80 E, 80 J, 81 B, 82 B, 85 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFCBFFC4F0A5F56DFB6AFBBC.taxon	diagnosis	Diagnosis. Regular and longitudinal striae on head dorsum diverge towards posterior margin; mesosoma with longitudinal, regular to irregular striae of uniform thickness slightly divergent, interspaces between striae smooth; lateral of pronotum and mesepisternum with longitudinal striae which in part continuing transversely on propodeum dorsum, and in part continuing on propodeal spine; ventral surface of node smooth; regular to irregular striae on node dorsum; subpostpetiolar process smooth; profemur mostly smooth; protibia extensor surface entirely striate, striae weakly marked; striation on tergum of first gastral segment restricted to base; branched setae of two types: 1) 2 – 3 thin, short and equal-sized branches diverging from the main axis; 2) thick setae flattened in its final half, with several branches.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFCBFFC4F0A5F56DFB6AFBBC.taxon	description	Redescription. WORKER (n = 7) (Fig. 65 A – C): HL 1 (0.86 – 1.10); HW 0.95 (0.81 – 1.04); ML 0.62 (0.52 – 0.70); SL 0.70 (0.58 – 0.70); MOD 0.22 (0.20 – 0.24); PNW 0.70 (0.58 – 0.72); WL 1.32 (1.08 – 1.34); PSL 0.22 (0.16 – 0.28); PL 0.57 (0.50 – 0.60); PW 0.22 (0.20 – 0.26); PPL 0.30 (0.28 – 0.36); PPW 0.32 (0.28 – 0.35); GL 1.10 (0.96 – 1.26); TL 4.92 (4.26 – 5.28); CI 95 (93.20 – 96.19); SI 73.68 (66.34 – 72.91); OI 23.68 (23.07 – 27.50). Small to midsized. Shiny integument. Brownish body, yellowish appendices. Mostly thin setae, unbranched or branched, midsized to short, suberect to subdecumbent; branched setae of two types: 1) 2 – 3 thin, short and equal-sized branches diverging from the main axis (Fig. 80 E); 2) thick setae flattened in its final half, with several branches, mainly in the gaster (Fig. 80 J). Head subquadrate; posterior margin slightly concave at middle. Mandible masticatory margin with 5 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with 8 – 9 regular to irregular, longitudinal, thick striae converging to a point on the anterior margin, interspaces smooth or thinner striae between thicker striae. Frontal triangle with 1 – 2 striae. Short scape, not reaching head posterior margin; apical antennomere as long as previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye reniform, small-sized, larger diameter with 10 ommatidia. Regular and longitudinal striae on head dorsum, divergent towards posterior margin, interspaces between thicker striae filled with thinner and regular to irregular striae. Head lateral and laterodorsal regions with the same striation of head dorsum, striae converging to eye margin; anterior laterodorsal region rugose; striae connecting the region posterior to frontal carina with eye margin; gena striate, 2 – 3 regular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma striation similar to head dorsum; longitudinal, regular to irregular striae of uniform thickness slightly divergent, interspaces smooth; transverse striae on pronotum anterior region (DV) directed to lateral of pronotum, longitudinal in other regions (DV); lateral of pronotum and mesepisternum with longitudinal striae in part continuing transversely on propodeum dorsum and in part continuing on propodeal spine. Promesonotal junction and metanotal groove indistinct. Transverse carina well-marked. Dorsal margin of mesosoma continuous, convex. Propodeal lobe bidentate, dorsal tooth slightly longer and acute than shorter and blunt ventral tooth; dorsal tooth length slightly shorter than 1 / 3 of propodeal spine length. Propodeal spine short, straight (LV), divergent (DV), sculptured on base. Transverse striae of uniform thickness on procoxa; irregular and transverse striae on C 2 and C 3. Profemur mostly smooth. Protibia extensor surface entirely striate, striae weakly marked. Dorsal margin of petiole discontinuous. Node with transverse striae on anterior surface continuing on lateral surface, interspaces distinguishable; regular to irregular striae on lateral and dorsal surfaces, interspaces smooth; ventral surface smooth. Postpetiole with regular, longitudinal striae; subpostpetiolar process smooth, weak, convex. First gastral segment striation similar to postpetiole striae; striae length on tergum similar to 1 / 3 of postpetiole length; sternite smooth. QUEEN (n = 6, 2 syntypes) (Fig. 66 A – C): HL 1.05 – 1.08 (0.94 – 1.12); HW 1 – 1.05 (0.88 – 1.06); ML 0.62 – 0.67 (0.60 – 0.74); SL 0.67 – 0.72 (0.63 – 0.74); MOD 0.27 – 0.28 (0.24 – 0.28); PNW 0.75 – 0.83 (0.75 – 0.90); WL 1.45 – 1.50 (1.30 – 1.52); PSL 0.25 – 0.32 (0.22 – 0.32); PL 0.62 – 0.65 (0.58 – 0.65); PW 0.25 – 0.28 (0.25 – 0.28); PPL 0.32 – 0.37 (0.32 – 0.40); PPW 0.35 – 0.39 (0.35 – 0.39); GL 1.26 – 1.37 (1.24 – 1.38); TL 5.5 (5.01 – 5.74); CI 95.24 – 97 (93.61 – 98.11); SI 63.91 – 72.50 (63.91 – 72.50); OI 26.80 – 27.50 (25 – 27.27). Midsized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 12 – 13 ommatidia. Longitudinal and mostly regular striae on scutum going from an anterior central point towards transcutal suture, interspaces distinguishable. Longitudinal and regular striae on anepisternum, katepisternum, axilla, and scutellum. Scutoscutellar sulcus inconspicuous. Transverse striae on propodeum (DV), interspaces between thicker striae filled with thinner striae. Lateral of mesosoma with longitudinal and mostly irregular striae directed to propodeal spine, interspaces distinguishable. Wings as in Fig. 81 B. MALE (first description) (n = 3) (Fig. 67 A – C): HL (0.80 – 0.84); HW (0.76 – 0.78); ML (0.44 – 0.48); SL (0.22 – 0.24); MOD (0.36 – 0.38); PNW (0.78 – 0.90); WL (1.36 – 1.50); PL (0.61 – 0.72); PW (0.22 – 0.26); PPL (0.34 – 0.36); PPW (0.31 – 0.34); GL (1.20 – 1.40); TL (4.85 – 5.12); CI (92.85 – 95.12); SI (28.20 – 30.77); OI (46.15 – 48.71). Brownish body, lighter appendices. Mandible with 4 teeth. Scutum mostly smooth and with few regular to irregular and thinner striae. Anepisternum and katepisternum mostly smooth. Scutellum with regular to irregular striae, thicker than scutum striae, interspaces distinguishable. Propodeum with irregular striae assuming multiple directions, interspaces distinguishable. Dorsal region of propodeal lobe forming an acute angle (= small tooth) and ventral region rounded. Petiole dorsum partly smooth and partly covered with irregular striae assuming multiple directions, interspaces distinguishable. Postpetiole mostly smooth. Gaster smooth. Wings as in Fig. 82 B.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFCBFFC4F0A5F56DFB6AFBBC.taxon	etymology	Etymology. This species was named in honor of Edmund Reitter (1845 – 1920), an Austrian Entomologist specialist in Coleoptera who collected this species’s type specimens.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFCBFFC4F0A5F56DFB6AFBBC.taxon	discussion	Comments. Kempf (1960) described the queen of H. reitteri and labelled it as a paratype. However, this queen is not part of the type specimens. The first queen description was made by Forel (1912) in the original description of Pogonomyrmex goeldi. Also, Kempf (1960) indicated that five H. reitteri syntypes from “ an unknown locality in São Paulo State ” were housed at Naturhistorisches Museum Wien, Austria. While visiting this collection at the museum, we recognized two workers and one queen with the label data described by Mayr (1887), and later labelled as lectotypes (workers) and paratype (queen) by Kempf. At the same museum there is also one worker labelled “ Brazil ” and “ lectotype ”, and two workers from Santa Catarina, Brazil, labelled “ paratypes ” by Kempf. Nevertheless, as Mayr (1887) described only the worker, providing a range of body measures and stated that the type locality was São Paulo, Brazil, we understand that only the two workers with the label data similar to what was described by Mayr are part of this type series. Possibly the other three “ syntype ” workers seen by Kempf have been misplaced. Kempf did not publish the later type designations he had apparently intended, thus we are designating the lectotype and consequently the paralectotype. Therefore, any other specimen with a type label should not be considered as such. Hylomyrma reitteri is similar to H. balzani, H. lopesi, H. transversa, and H. wachiperi. The first four species show broad distributions (Figs. 84, 85, 89), while H. wachiperi is known only in one locality in Peru (Fig. 86). Hylomyrma reitteri, H. transversa, and H. wachiperi are allopatric, but present the discontinuous dorsal margin of petiole, and a similar pattern of sculpturation on head. However, H. reitteri can be easily distinguished from H. transversa (characteristic in parentheses) in the reniform eye (vs. drop-shaped), the longitudinal striae on the mesonotal dorsum (vs. transverse striae), and the smooth ventral surface of petiole (vs. striate). Hylomyrma reitteri differs from H. wachiperi in the reniform eye (vs. oval), the smooth ventral surface of petiole (vs. striate), and the tergum of the first gastral segment with short striae (vs. long striae). Hylomyrma balzani and H. reitteri occur in sympatry in Paraguay (Alto Paraná department) and in 19 localities at Brazil (Santa Teresa, ES; Lavras, and Viçosa, MG; Estrada Velha Guaricana, Irati, Laranjeiras, Morretes, Rondon, and Tuneiras do Oeste, PR; Rio de Janeiro, and Teresópolis, RJ; Frederico Westphalen, Nova Petrópolis, and 5 km N of Progresso, RS; Chapecó, Nova Teutônia, and Seara, SC; Botucatu, and Jundiaí, SP). Hylomyrma reitteri can be distinguished from H. balzani by the regular to irregular striae on the mesosoma (vs. rugose striae), the irregular striae on the petiole (vs. rugose striae), and the smooth ventral surface of petiole (vs. striate surface). The winged queens of both species are very similar, differing only in the laterodorsal sculpture of the propodeum. In Hylomyrma balzani this region presents rugose striae, whereas H. reitteri has regular to irregular striae in this region. Hylomyrma reitteri differs from H. lopesi in the smaller body size (vs. larger), the thick striae on the mesosomal dorsum (vs. thin striae), the transverse striae on the ventral surface of petiole (vs. smooth surface), the laterals of the mesosoma covered mainly with regular and thick striae that are directed to the propodeal spine (vs. irregular and thin striae directed to the propodeal dorsum), and the striate subpostpetiolar process (vs. smooth). Both species occur broadlly in Brazil, the distribution of H. reitteri is more restricted to eastern portions of Brazil from southern and southeast regions, whereas H. lopesi is distributed mainly in the center of Brazil from northern (state of Pará) to southern (state of São Paulo) regions. Hylomyrma reitteri and H. lopesi co-occur in Botucatu (SP), Niquelândia (GO), and São José de Buerarema (BA), Brazil. The most striking intraspecific morphological variation found in H. reitteri concerns the pilosity and the sculpture of the petiolar dorsum. The striae on the petiolar dorsum, usually longitudinal, can be transverse (Brigadeiro, MG, and Teresópolis, RJ, Brazil) or can be in part longitudinal and in part transverse (Niquelândia, GO; Brigadeiro, MG; Santa Teresa, ES; and Morretes, PR, Brazil). Hylomyrma reitteri setae are mostly branched, with branches of unequal length. The exception to this setae pattern is observed in some specimens from Florianópolis, SC and Viçosa, MG, Brazil, which present mostly unbranched setae, and the few branched setae have branches of equal size (found in the propodeum, petiole, postpetiole and / or gaster). It is impressive that out of the 1,037 H. reitteri female specimens analysed, 730 are intercastes. As in H. balzani and H. lopesi, H. reitteri presents considerable morphological variation among intercastes. We recognized eight morphotypes that essentially have larger body sizes, and mostly are worker-like, just two are queen-like (Fig. 2 A).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFCBFFC4F0A5F56DFB6AFBBC.taxon	distribution	Distribution. This species is widely distributed in eastern Brazil, from southern Bahia to Rio Grande do Sul. It also occurs in Paraguay (Salto Tembey and Ciudad del Este) (Fig. 85). Natural history. Hylomyrma reitteri inhabits mainly areas in the Atlantic forest, but there are also records in the “ Cerrado ” (Brazilian savanna) in Goiás, and in coffee plantations (Minas Gerais), both in Brazil. Specimens are frequently found in leaf-litter samples (Fig. 68), which suggests that nests are located in fallen logs, rotten wood, between leaves, or inside natural cavities of the superficial soil layers. Most of the intercastes are worker-like and just few have more queen-like traits, those may in the future be understood as ergatoids. One winged queen and one queen-like intercaste were collected together in two different winkler samples (winkler 49 from São Bonifácio, SC, and winkler 31 from Santa Teresa, ES, Brazil).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFCBFFC4F0A5F56DFB6AFBBC.taxon	materials_examined	Additional material examined (209 workers, 98 queens, 107 males, 730 intercastes, 6 indeterminates): BRAZIL: BA [Bahia]: São José de Buerarema, 14.09.00, 15 ° 03 ′ 06 ″ S, 39 ° 18 ′ 48 ″ W, Santos J. R. M. dos, 637 (2 W) [CEPLAC]; ES [Espírito Santo]: Santa Teresa, Estação Biológica Santa Lucia, 19 ° 58 ′ 09 ″ S, 40 ° 32 ′ 15 ″ W, 20 - 24. i. 2002, Schoereder J. H. & Ribas C. R. cols., Winkler 2 (2 I) [MZSP]; same except Winkler 4 (1 I 1 M) [MZSP]; Winkler 9 (1 W) [MZSP]; Winkler 12 (1 I) [MZSP]; Winkler 19 (1 I) [MZSP]; Winkler 21 (1 I) [MZSP]; Winkler 23 (2 I) [MZSP]; Winkler 24 (1 W) [MZSP]; Winkler 27 (1 W) [MZSP]; Winkler 31 (1 I 1 Q) [MZSP]; Winkler 44 (1 W) [MZSP]; Winkler 45 (1 W) [MZSP]; Winkler 47 (1 I 1 M) [MZSP]; Winkler 14 (1 I) [MZSP]; Winkler 36 (1 I covered with gold) [MZSP]; Winkler 46 (1 I) [MZSP]; Winkler 48 (1 I) [MZSP]; GO [Goiás]: Niquelândia, 18 - 30. v. 1996, 14 ° 17 ′ 05 ″ S, 48 ° 55 ′ 01 ″ W, Silvestre, Silva & Brandão col., peneira (1 I) [MZSP]; Campinaçu, Serra da Mesa, 13 ° 52 ′ S, 48 ° 23 ′ W, 18. ii- 2. iii. 1996, Silvestre, Brandão & Yamamoto col., Cerrado (1 I covered with gold) (MZSP 67443) [MZSP]; MG [Minas Gerais]: Lavras, 6. xii. 1978, W. D. Fronk col., berlese funnel (2 W 1 Q) [MCZC]; same locality Fragmento, 06 - 12.2002, Santos M. S. & Dias N. S. (3 I) [CEPLAC]; Viçosa, Cafezal, 7. viii. 1987, M. V. B. Queiroz col., A 1 3 (1 Q) [MZSP]; same locality Floresta secundária, ii. 1994, Sperber, Louzada & Lopes, UFV LABECOL n ° 000143 (1 I) [MZSP]; Parque Estadual da Serra do Brigadeiro, 1500 m, winkler sample, Jan. 2007, R. Solar. col. (5 I 1 W 1 Q) [MZSP]; [PR / Paraná]: Rondon, vii. 1952, 24 ° 38 ′ B, 54 ° 07 ′ L, 500 m, Fritz Plaumann (1 I) [MZSP]; Rio Azul, 1000 m, x. 1959, F. Plaumann, 3173 (2 W) [MZSP]; Bocaiuva, v. 1963, F. Plaumann, 4002 (1 Q 1 W 3 I) [MZSP]; Guaragi, 25 ° 16 ′, 54 ° 14 ′, v. 1964, F. Plaumann, 4583 (2 Q 1 W) [MZSP]; Laranjeiras, iv. 1965, F. Plaumann, 4550 (2 Q 1 I) [MZSP]; Antonina, Reserva Natural Rio Cachoeira, 25 ° 18 ′ 21.85 ″ S, 48 ° 40 ′ 26.58 ″ W, 02 - 05. iii. 2014, R. Feitosa, J. Calixto, W. Franco & A. Oliveira cols., P 1 DQ 4 DIA 1 (1 I) [DZUP]; same except P 1 EQ 4 DIA 1 (1 I) [DZUP]; P 16 EQ 1 DIA 2 (2 I) [DZUP]; P 5 EQ 2 NOITE 1 (4 W 1 I) [DZUP]; P 14 EQ 1 DIA 2 (1 W) [DZUP]; P 12 DQ 2 DIA 2 (1 W) [DZUP]; P 15 EQ 2 DIA 2 (1 W) [DZUP]; P 3 EQ 4 NOITE 1 (2 W) [DZUP]; P 14 DQ 1 DIA 2 (2 W 1 I) [DZUP]; P 15 EQ 2 DIA 2 (1 W 1 I) [DZUP]; P 19 EQ 4 NOITE 2 (1 I) [DZUP]; P 8 DQ 1 NOITE 2 (1 I) [DZUP]; P 14 DQ 4 NOITE 2 (2 I) [DZUP]; P 3 EQ 4 NOITE 1 (2 I) [DZUP]; P 1 D 5 Q 2 DIA 2 (2 I) [DZUP]; P 10 DQ 4 NOITE 2 (2 I) [DZUP]; São José dos Pinhas, pitfall, 972 m, 25 ° 33 ′ 28.89 ″ S, 48 ° 59 ′ 38.28 ″ W, 15 - 22. x. 2011, J. S. Vieira e Klemann-Júnior col. (2 W 1 I) [DZUP]; Piraquara, Parque Estadual Pico do Marumbi, 25 ° 29 ’ 32.59 ” S, 48 ° 59 ’ 33.59 ” W, 15 - 19. xii. 2014, R. Feitosa et al. cols., Floresta Ombrófila Densa, Winkler, P 10 EQ 3 NOITE 2 (1 I) [DZUP]; same except P 8 EQ 2 NOITE 2 (7 W 1 Q 1 I) [DZUP]; P 19 DQ 4 DIA 2 (2 W 3 I) [DZUP]; Ponta Grossa, 24 - 28. xi. 2014, Feitosa et al. cols, winkler, arenitos 4 (1 W) [DZUP]; same except arenitos 1 (1 I) [DZUP]; arenitos 7 (1 Q 1 I) [DZUP]; 24 - 29. xi. 2014, Feitosa et al. cols., winkler, furnas 1 (2 W) [DZUP]; furnas 12 (1 W) [DZUP]; furnas 4 (1 Q) [DZUP]; furnas 9 (1 I) [DZUP]; Morretes, Parque Estadual do pau-Ôco, 25 ° 34 ′ 33.5 ″ S, 48 ° 53 ′ 19.5 ″ W, 6 - 11. v. 2002, Silva R. R. & Dietz B. H. cols., Winkler 3 (1 I) [MZSP]; same except Winkler 8 (1 I) [MZSP]; Winkler 9 (1 W 2 I) [MZSP]; Winkler 18 (1 W) [MZSP]; Winkler 11 (2 W) [MZSP]; Winkler 44 (1 W 1 I) [MZSP]; Winkler 49 (1 W 2 I) [MZSP]; Tunas, Parque das Lauráceas, 21 - 29. ii. 2001, 24 ° 51 ′ 16 ″ S, 48 ° 43 ′ 00.4 ″ W, Silva & Eberhardt cols., Transecto I, Winkler 01 (1 W 1 I) [MZSP]; same data (1 W) (MZSP 67444) [MZSP]; same except Winkler 02 (1 Q) [MZSP]; Winkler 03 (1 W 1 I) [MZSP]; Winkler 05 (1 W 1 I) [MZSP]; Winkler 06 (1 I) [MZSP]; Winkler 07 (1 I) [MZSP]; Winkler 08 (1 W 1 I) [MZSP]; Winkler 09 (1 W) [MZSP]; Winkler 10 (1 I) [MZSP]; Winkler 11 (1 I) [MZSP]; Winkler 12 (1 Q 2 W) [MZSP]; Winkler 13 (1 I) [MZSP]; Winkler 14 (1 I) [MZSP]; Winkler 15 (1 I) [MZSP]; Winkler 16 (1 I) [MZSP]; Winkler 17 (1 I) [MZSP]; Winkler 18 (1 I) (MZSP 67448) [MZSP]; Winkler 19 (1 I) [MZSP]; Winkler 22 (1 I) [MZSP]; Winkler 23 (1 W) [MZSP]; Winkler 26 (1 W 1 I) [MZSP]; Winkler 28 (1 W) [MZSP]; Winkler 29 (1 I) [MZSP]; Winkler 31 (1 I) [MZSP]; Winkler 32 (1 indet.) [MZSP]; Winkler 34 (1 W) (MZSP 67429) [MZSP]; Winkler 36 (1 W) [MZSP]; Winkler 37 (2 W) [MZSP]; Winkler 38 (1 W 1 I) [MZSP]; Winkler 39 (1 I) [MZSP]; Winkler 40 (1 I) [MZSP]; Winkler 41 (1 W) [MZSP]; Winkler 42 (1 W 2 Q 1 I) [MZSP]; Winkler 45 (1 W) [MZSP]; Winkler 46 (1 I) [MZSP]; Winkler 47 (1 I) [MZSP]; Winkler 48 (1 W 1 Q) [MZSP]; Winkler 49 (1 I) [MZSP]; Winkler 50 (1 W) [MZSP]; Estrada Velha Guaricana, ca Cachoeira das Mulas, - 25.3423 ° 48.9226 °, 13. iv. 2017, R. López, J. Lattke, 3856 (1 I) [MZSP]; same except - 25.7303 ° 49.0911 °, 10. iv. 2017, R. López, J. Lattke, 3814 (1 W) [DZUP]; 3827 (1 W) [DZUP]; 3823 (2 W) [DZUP]; 3828 (1 W) [DZUP]; 3831 (1 W) [DZUP]; 3826 (1 I) [DZUP]; Irati, Guamirim, Fz. Arroio Grande, 09. x. 2014, 25 ° 35 ′ 36.11 ″ S, 50 ° 49 ′ 12.06 ″ W, Serapilheira, Funil de Berlese, Marques, C. G. P., Falbot, L. col., Talhão 16 - AII (2 W 1 Q) [DZUP]; Talhão 15 - AIII (1 W) [DZUP]; Talhão 11 - AII (1 W) [DZUP]; Talhão 19 - AII (1 W) [DZUP]; Tuneiras do Oeste, Reserva Biológica de Perobas, 23 ° 50 ′ 5.64 ″ S, 52 ° 45 ′ 37.52 ″ W, 18. ix. 2015, Busanello, D. & Caron, E. cols. (11 W) [DZUP]; Reserva Biológica das Perobas, 23 ° 50 ′ 39 ″ S, 52 ° 44 ′ 43.26 ″ W (2 W) [DZUP]; [RJ, Rio de Janeiro: Santa Maria] Madalena, [P. E. do Desengano, 21 ° 58 ′ 41 ″ S, 41 ° 57 ′ 00 ″ W, 30. xi. 2002, Mayhé A. & Veiga-Ferreira S. cols.], 30 (1 I) [MZSP]; same except 38 (1 I) (MZSP 67446) [MZSP]; 36 (1 W) [MZSP]; 50 (1 I) (MZSP 67445) [MZSP]; 48 (1 I) [MZSP]; 47 (1 I 1 Q) (MZSP 67449) [MZSP]; Rio de Janeiro, Floresta da Tijuca, D. Federal, ii. 1960, C. A. C Seabra (1 W) [MZSP]; Teresópolis, P. N. Serra dos órgãos, 23 - 28. xi. 1999, Dietz, Silva & Rocha cols., Winkler 10 (2 I) [MZSP]; Winkler 18 (1 W) [MZSP]; RS [Rio Grande do Sul]: N. [Nova] Petropolis, Lat 29 ° 20 ′, Lo. 51 ° 12 ′, xi. 1959, F. Plaumann, 3223 (1 Q 2 W) [MZSP]; Frederico Westphalen, UF Santa Maria, 04. xi. 2008, Granzotto F., # 5546 (2 W 1 I) [CEPLAC]; 5 km north of Progresso, Linha Araçá, near Rio Fão, 52 ° 20 ′ W, 29 ° 10 ′ S, 300 - 400 m, Secondary Forest, soil sifting, Winkler 1 qm, 23. iv. 1999, Leg. J. Bihn (2 I) [MZSP]; same except 17. iii. 1999 (1 W 1 I) [MZSP]; Tainhas, iv. 1959, F. Plaumann, 3077 (2 I) [MZSP]; B [arão de] Cotejipe, vii. 1960, F. Plaumann, 3771 (2 Q 2 I) [MZSP]; SC [Santa Catarina]: Abelardo Luz, 26 ° 33 ′ 53 ″ S, 52 ° 19 ′ 42 ″ W, 20. ix. 1999, Silva R. R. col., Transecto I, Winkler (2 I) [MZSP]; Blumenau, P. E. Nascentes, 27 ° 06 ′ 15 ″ S, 49 ° 09 ′ 14 ″ W, 20 - 27. x. 2000, Silva R. R. & Eberhardt F. cols., Transecto I, Winkler 01 (1 I) [MZSP]; same except Winkler 03 (2 I) [UTEP]; Winkler 04 (1 I) [MZSP]; Winkler 05 (1 W 1 I) [MZSP]; Winkler 07 (3 I) [MZSP]; Winkler 08 (1 Q) [MZSP]; Winkler 10 (1 I) [MZSP]; Winkler 12 (2 I) [MZSP]; Winkler 13 (1 I) [MZSP]; Winkler 17 (1 I) [MZSP]; Winkler 20 (1 I) [MZSP]; Winkler 21 (1 W 1 I) [MZSP]; Winkler 24 (1 Q) [MZSP]; Winkler 26 (1 I) [MZSP]; Winkler 29 (1 I) [MZSP]; Winkler 30 (1 W) [MZSP]; Winkler 31 (1 W) [MZSP]; Winkler 32 (1 I) [MZSP]; Winkler 34 (1 I) [MZSP]; Winkler 35 (1 I) [MZSP]; Winkler 36 (1 I) [MZSP]; Winkler 37 (1 I) [MZSP]; Winkler 38 (1 I) [MZSP]; Winkler 39 (1 I) [MZSP]; Winkler 40 (1 I) [MZSP]; Winkler 41 (1 W) [MZSP]; Winkler 42 (1 I) [MZSP]; Winkler 44 (1 I) [MZSP]; Winkler 45 (1 I) [MZSP]; Winkler 46 (1 W) [MZSP]; Winkler 47 (1 I) [MZSP]; Winkler 48 (1 I) [MZSP]; Winkler 49 (1 I) [MZSP]; Winkler 50 (1 I) [MZSP]; Chapecó, v. 1957, Plaumann col., Col. Kempf n ° 2655 (5 I 1 Q) [MZSP]; same locality xii. 1957, F. Plaumann col., Coll. Borgm. Nr. 6501 (5 I 1 Q) [MZSP]; Concórdia, ix. 1959, F. Plaumann, 8294 (1 W) [MZSP]; Florianópolis, Morro da Lagoa, 19. vi. 1993, R. R. da Silva & B. C. Lopes, funil- 56 (1 I) [MZSP]; same locality, Canto da Lagoa, - 27.533657, - 48.458410, 15. ii. 2016, JCwinkler # 003, J. Chaul col. (1 Q) [MZSP]; UCAD, 18. iii. 2005, H. Mozerle (1 W) [UFSC]; Praia dos Naufragados, 27 ° 49 ′ 58.2 ″ S, 48 ° 33 ′ 37.8 ″ W, Jan. 2013, sifted litter, J. Chaul, UFV LABE-COL n ° 000141 (1 I) [MZSP]; same except início da descida da trilha para a praia, - 27.823412, - 48.561569, 19. ii. 2016, JCwinkler # 008, J. Chaul col. (1 W 5 I) [MZSP]; Nova Teutonia, 27 ° 11 ′ B, 52 ° 23 ′ L, Fritz Plaumann, 300 - 500 m, viii. 1952 (5 I 1 Q) [USNM]; same except x. 1953 (1 W 1 Q) [MCZC]; x. 1954 (2 W) [MZSP]; i. 1957 (1 I) [MZSP]; v. 1957 (1 W) [MZSP]; vi. 1957 (1 W) [MZSP]; vii. 1957 (1 W 4 I 3 Q) [MZSP]; viii. 1957 (1 W 2 I 1 Q) [MZSP]; (1 Q) [MCZC]; vi. 1959 (2 I) [IFML]; ix. 1959, 3490 (1 W 6 I 2 Q) [MZSP]; vi. 1960 (1 Q) [MZSP]; 1961, leg. F. Plaumann (1 Q) [MHNG]; iv. 1976, 13628 (4 I 1 Q) [MZSP]; N. Teutônia, vi. 1960, 8172 (1 W 3 I 2 Q) [MZSP]; vi. 1961, 6869 (2 I 3 Q) [MZSP]; viii. 1961, 7003 (5 I 1 Q) [MZSP]; xii. 1970, 8135 (1 M) [MZSP]; iii. 1971, F. Plaumann, 7099 (1 I) [MZSP]; Palhoça, P. E. Serra do Tabuleiro, 27 ° 44 ′ 28 ″ S, 48 ° 41 ′ 50 ″ W, 02 - 10. vi. 2003, Silva R. R., Dietz B. H. & Tavares A. cols., Winkler 2 (2 I) [MZSP]; same except Winkler 3 (2 I) [MZSP]; Winkler 4 (1 I) [MZSP]; Winkler 6 (1 I 1 Q) [MZSP]; Winkler 6 (1 I 1 Q) [MCZC]; Winkler 7 (2 I) [MZSP]; Winkler 8 (2 I) [MZSP]; Winkler 9 (2 I) [MZSP]; Winkler 10 (2 I) [MZSP]; Winkler 13 (1 I) [MZSP]; Winkler 14 (2 I) [MZSP]; Winkler 20 (1 Q) [CASC]; Winkler 29 (2 I) [MZSP]; Winkler 31 (2 I) [UTLP]; Winkler 33 (2 I) [MZSP]; Winkler 35 (2 I) [MZSP]; Winkler 37 (2 I) [USNM]; Winkler 38 (1 W 1 I) [MZSP]; Winkler 39 (1 I) [MZSP]; Winkler 41 (1 I) [MZSP]; Winkler 44 (2 I) [MZSP]; Winkler 45 (1 W) [MZSP]; Winkler 47 (1 I) [MZSP]; Winkler 49 (1 I) [MZSP]; P. [Passos] Bormann, xii. 1957, Plaumann (1 W) [MZSP]; Sto. [Santo] Amaro, 17. xi. 2009, PEST, F. O. D., Romualdo Begnini, NP 1 (1 W 1 I) [UFSC]; São Bento do Sul, APA Rio Vermelho, 26 ° 21 ′ 51 ″ S, 49 ° 16 ′ 16 ″ W, 30. iii- 04. iv. 2001, Silva R. R. & Eberhardt F. cols., Transecto II, Winkler 26 (1 I) [MZSP]; same except Winkler 27 (1 I) [MZSP]; Winkler 28 (1 I) [MZSP]; Winkler 29 (1 I) [MZSP]; Winkler 30 (1 I) [MZSP]; Winkler 31 (1 I) [MZSP]; Winkler 35 (1 I) [MZSP]; Winkler 38 (1 I) [MZSP]; Winkler 40 (1 I) [MZSP]; Winkler 41 (1 Q) [MZSP]; Winkler 42 (1 I) [MZSP]; Winkler 44 (1 I) [MZSP]; Winkler 45 (1 I) [MZSP]; Winkler 46 (1 I) [MZSP]; Winkler 47 (1 I) [MZSP]; Winkler 50 (1 I) [MZSP]; Transecto I, Winkler 01 (1 I) [MZSP]; Winkler 03 (1 I) [MZSP]; Winkler 04 (1 I) [MZSP]; Winkler 05 (1 I) [MZSP]; Winkler 06 (1 I) [MZSP]; Winkler 08 (1 I) [MZSP]; Winkler 09 (2 I) [IHVL]; Winkler 10 (1 I) [MZSP]; Winkler 11 (1 I) [MZSP]; Winkler 12 (1 I) [MZSP]; Winkler 13 (1 I) [MZSP]; Winkler 14 (1 I) [MZSP]; Winkler 15 (1 I) [MZSP]; Winkler 17 (1 I) [MZSP]; Winkler 18 (1 I) [MZSP]; Winkler 19 (1 I) [MZSP]; Winkler 21 (1 I) [MZSP]; Winkler 22 (1 I) [MZSP]; Winkler 23 (1 I) [MZSP]; Winkler 24 (1 I) [MZSP]; Winkler 29 (1 I) [MZSP]; São Bonifácio, 27.8183 S, 48.9114 W, 850 m, 2 - 10. vii. 2011, Winkler, F. A. Esteves & B. L. Fisher cols. (4 W 1 I) [MZSP]; same locality, - 27.8183 - 48.9114, 850 m, winkler, 02 - 10. vii. 2011, F. A. Esteves & B. L. Fisher cols. (3 I 3 W) [MZSP]; same except BLF 27476 (1 I) [MZSP]; P. E. Serra do Tabuleiro, 27 ° 49 ′ 06 ″ S, 48 ° 54 ′ 41 ″ W, 08 - 13. iii. 2004, Silva R. R., Dietz B. H. & Albuquerque N. (5 I 1 Q) [MZSP]; same except Winkler 1 (1 I) [MZSP]; Winkler 2 (1 I) [MZSP]; Winkler 4 (2 I) [MZSP]; Winkler 5 (1 I) [MZSP]; Winkler 6 (1 Q) [MZSP]; Winkler 8 (1 I) [MZSP]; Winkler 10 (2 I) [MZSP]; Winkler 12 (2 I) [MZSP]; Winkler 13 (1 I) [MZSP]; Winkler 14 (1 I) [MZSP]; Winkler 15 (3 I) [MZSP]; Winkler 16 (1 I) [MZSP]; Winkler 18 (1 W 1 Q) [MZSP]; Winkler 19 (1 I) [MZSP]; Winkler 21 (2 I) [MZSP]; Winkler 22 (3 I) [MZSP]; Winkler 23 (2 I) [MZSP]; Winkler 24 (1 Q) [MZSP]; Winkler 27 (2 I 1 Q) [MZSP]; Winkler 28 (2 I) [MZSP]; Winkler 31 (2 I) [MZSP]; Winkler 33 (2 I) [MZSP]; Winkler 34 (2 I) [MZSP]; Winkler 35 (1 I) [MZSP]; Winkler 37 (1 I 1 Q) [MZSP]; Winkler 38 (1 Q) [MZSP]; Winkler 39 (2 I) [MZSP]; Winkler 40 (1 I) [MZSP]; Winkler 41 (1 Q) [MZSP]; Winkler 42 (2 I) [MZSP]; Winkler 43 (2 I 1 Q) [MZSP]; Winkler 45 (3 I) [MZSP]; Winkler 46 (1 I) [MZSP]; Winkler 47 (3 I) (MZSP 67447) [MZSP]; Winkler 48 (1 I) [MZSP]; Winkler 49 (2 I 2 Q) [MZSP]; Winkler 50 (1 I) [MZSP]; Seara, xii. 1958, Plaumann, 2709 (1 I) [MZSP]; same except vii. 1958, Plaumann, 2721 (2 I) [MZSP]; 24 ° 07 ′ S, 52 ° 18 ′ W, v-xii. 1998, Rogério R. Silva col., Transecto I, Winkler (2 I) [MZSP]; Xanxerê, xii. 1957, F. Plaumann (1 I) [MZSP]; SP [São Paulo]: Bertioga, RPPN SESC Bertioga, 23 ° 49 ′ 18.9 ″ S, 46 ° 07 ′ 23.7 ″ W, Restinga [arbórea], Winkler, 18 - 20. x. 2014, M. A. Ulysséa & L. P. Prado cols., PT 2 (1 W) (MZSP 67432) [MZSP]; Botucatu, 9 - 11 - 1987, Forti L. C. & Rinaldi I. M. P. col., Forti # 14 (1 I) [MZSP]; same except 07 - 07 - 1987, Forti # 34 (1 I) [MZSP]; 15 - 19 - 1987, Forti # 38 (3 I) [MZSP]; 18 - 02 - 1988, Forti # 10 (1 I) [MZSP]; 07 - 10 - 1986, Forti L. C. col. (1 I) [MZSP]; 14 - 04 - 1987, Forti L. C. col., Forti # 51 (1 I) [MZSP]; 03 - 09 - 1986, Forti # 42 (1 Q 1 I) [MZSP]; Forti # 41 (2 Q 1 I) [MZSP]; Mata Ciliar, Botignoli, 9. viii. 1988, L. C. Forti col., Armadilha de solo, Forti # 04 (1 W) [MZSP]; Cananéia, P. E. Ilha do Cardoso, 25 ° 05 ′ 48.7 ″ S, 47 ° 55 ′ 47.3 ″ W, 24 - 28. xi. 2002, Silva R. R., Brandão C. R. F., Scott C. cols., Winkler 1 (2 I) [MZSP]; same except Winkler 2 (1 I) [MZSP]; Winkler 9 (1 I) [MZSP]; Winkler 11 (2 I) [MZSP]; Winkler 12 (1 I) [MZSP]; Winkler 13 (2 I) [MZSP]; Winkler 15 (2 I) [MZSP]; Winkler 32 (1 I) [MZSP]; Winkler 35 (1 I) [MZSP]; Winkler 39 (2 W 1 I) [IHVL]; Winkler 42 (1 I) [MZSP]; Winkler 47 (1 I) [MZSP]; Winkler 48 (1 W) [MZSP]; Winkler 49 (1 I) [MZSP]; Cunha, P. E. Serra do Mar, Núcleo Cunha-Indaiá, 23 ° 15 ′ 03 ″ S, 45 ° 00 ′ 26 ″ W, 21 - 22. iv. 2001, A. A. Tavares, R. R. Silva, cols., Winkler 5 (1 W) [MZSP]; same except Winkler 10 (1 I) [MZSP]; Winkler 11 (1 W 1 Q 1 I) [MZSP]; Winkler 13 (1 W) [MZSP]; Winkler 14 (1 Q) [MZSP]; Winkler 15 (1 Q) [MZSP]; Winkler 17 (3 I) [MZSP]; Winkler 18 (1 W) [MZSP]; Winkler 19 (1 Q 3 I) [MZSP]; Winkler 20 (2 I 1 indet.) [MZSP]; Winkler 22 (1 W 1 I) [MZSP]; Winkler 23 (3 W 2 I) [MZSP]; Winkler 24 (1 W 1 I) [MZSP]; Winkler 26 (1 W 1 I 1 M) [MZSP]; Winkler 27 (3 W 1 I 1 Q) [MZSP]; Winkler 28 (1 I) [MZSP]; Winkler 29 (1 W 1 Q 1 I) [MZSP]; Winkler 31 (1 W) [MZSP]; Winkler 32 (1 Q 1 I) [MZSP]; Winkler 33 (1 I) [MZSP]; Winkler 37 (1 W 1 I) [MZSP]; Winkler 39 (2 I) [MZSP]; Winkler 42 (3 W 1 I) [MZSP]; Winkler 43 (1 I) [MZSP]; Winkler 45 (1 Q 1 I) [MZSP]; Winkler 46 (1 W 2 I) [MZSP]; Winkler 47 (1 W 1 Q) [MZSP]; Winkler 50 (3 I) [MZSP]; Igaratá, Cachoeira Pedra Bonita, 23 ° 12 ′ 28 ″ S 46 ° 10 ′ 39 ″ W, 14. iv. 2015, T. Fernandes col., nidificando em galho na serapilheira, galho compartilhado com Brachymyrmex admotus, B. heeri, Pheidole sigillata e duas espécies de Solenopsis, P 6 G- 1098, CPB 3 - 1499 (1 Q) [MZSP]; same except CPB 3 - 1500 (1 Q) [MZSP]; Iguape, E. E. Juréia-Itatins, Núcleo Rio Verde, 24 ° 32 ′ 39 ″ S, 47 ° 14 ′ 08 ″ W, 5 - 14. iii. 2001, A. A. Tavares col., Winkler 1 (1 I) [MZSP]; same except Winkler 10 (1 W) [MZSP]; Winkler 11 (1 I) [MZSP]; Winkler 22 (1 I) [MZSP]; Winkler 34 (1 I) [MZSP]; Winkler 50 (1 I) [MZSP]; Itatinga, Mata, litter, 19. x. 1991, B. H. Dietz col., Forti # 06 (1 I) [MZSP]; Jundiaí, Serra do Japi, 23 ° 13 ’ S, 46 ° 58 ’ W, pitfall, iii. 2011, T. Postali col., # 1644 (1 I) [MZSP]; Nazaré Pta. [Paulista], tronco podre, 25. i. 1992, B. H. Dietz col. (3 Q) [MZSP]; Picinguaba, P. E. Serra do Mar, 23 ° 20 ′ 10 ″ S, 44 ° 50 ′ 15.3 ″ W, 30. iii- 04. iv. 2001, Brandão C. R. F. & Eq. cols, Winkler 2 (1 I) [MZSP]; same except Winkler 3 (1 I) [MZSP]; Winkler 4 (1 I) [MZSP]; Winkler 6 (1 W) [MZSP]; Winkler 7 (1 I) [MZSP]; Winkler 15 (1 W 1 I) [MZSP]; Winkler 16 (1 W 2 I) [MZSP]; Winkler 17 (2 I) [MZSP]; Winkler 18 (1 I) [MZSP]; Winkler 19 (1 I) [MZSP]; Winkler 22 (2 I) [MZSP]; Winkler 26 (1 W 2 I) [MZSP]; Winkler 28 (1 I) [MZSP]; Winkler 35 (2 I) [MZSP]; Winkler 41 (1 W 2 I) [USNM]; Winkler 43 (1 indet.) [MZSP]; Winkler 45 (1 W 2 I) [MZSP]; Winkler 47 (2 I) [MZSP]; Winkler 49 (2 I) [MZSP]; Praia Grande, P. E. Serra do Mar, Núcleo Pilões-Cubatão, 23 ° 58 ′ 31 ″ S, 46 ° 32 ′ 24 ″ W, 26 - 27. v. 2001, A. A. Tavares, R. R. Silva, cols., Winkler 1 (1 I) [MZSP]; same except Winkler 12 (2 I) [MZSP]; Winkler 37 (1 I) [MZSP]; Ribeirão Grande, P. E. Intervales, Base Barra Grande, 02. ii. 1999, A. A. Tavares, Transecto 1, Winkler 1 (1 W 1 I) [MZSP]; same except Winkler 2 (3 I) [MZSP]; Winkler 3 (1 W 3 I) [MZSP]; Winkler 5 (1 W) [MZSP]; Winkler 6 (1 I) [MZSP]; Winkler 7 (3 I) [MZSP]; Winkler 8 (1 I) [MZSP]; Winkler 11 (1 I) [MZSP]; Winkler 12 (1 I) [MZSP]; Winkler 13 (2 I) [MZSP]; Winkler 14 (2 I) [MZSP]; Winkler 15 (3 I) [MZSP]; Winkler 16 (1 I) [JTLC]; Winkler 17 (1 W 1 I) [JTLC]; Winkler 21 (2 I) [MZSP]; Winkler 22 (3 I) [MZSP]; Winkler 24 (2 I 1 indet.) [MZSP]; Winkler 25 (1 I) [MZSP]; 05. ii. 1999, Transecto 2, Winkler 1 (1 I) [MZSP]; Winkler 6 (3 I) [MZSP]; Winkler 7 (2 W 1 I) [MZSP]; Winkler 8 (2 I) [MZSP]; Winkler 11 (3 I) [MZSP]; Winkler 12 (1 I) [MZSP]; Winkler 15 (2 I) [MZSP]; Winkler 17 (1 W) [MZSP]; Winkler 18 (1 I) [MZSP]; Winkler 21 (1 W 1 I) [MZSP]; Winkler 22 (1 I) [MZSP]; Winkler 22 (1 W 2 I) [MZSP]; Salesópolis, E. B. B. [Estação Biológica de Boracéia], 20 - 26. x. 1997, C. Klingenberg & C. I. Yamamoto col., Winkler 2 a (1 I) [MZSP]; same except Winkler 3 b (1 I) [MZSP]; Winkler 8 a (1 I) [MZSP]; Winkler 9 a (1 I) [MZSP]; Winkler 9 b (1 I) [MZSP]; Winkler 10 a (1 Q 1 I) [MZSP]; Winkler 10 b (1 I) [MZSP]; Winkler 11 a (1 W 1 I) [MZSP]; Winkler 11 b (1 I) [MZSP]; Winkler 12 a (1 I) [MZSP]; Winkler 14 a (1 I) [MZSP]; Winkler 14 b (1 I) [MZSP]; Winkler 15 a (1 I) [MZSP]; Winkler 16 a (1 I) [MZSP]; Winkler 22 a (1 I) [MZSP]; Winkler 22 b (1 I) [MZSP]; Winkler 25 a (1 Q) [MZSP]; Pitfall 19 (a) (1 I) [MZSP]; Pitfall 20 (a) (1 Q) [MZSP]; 2 - 6. v. 1997, D. Agosti, C. R. F. Brandão, C. I. Yamamoto col., Transect I, Winkler 5 a (1 W) [MZSP]; same except Winkler 5 b (4 I) [MZSP]; Winkler 7 (1 W 1 Q 2 I) [MZSP]; Winkler 12 a (2 I) [MZSP]; Winkler 12 b (1 I) [MZSP]; Winkler 13 a (1 W 1 Q) [MZSP]; Winkler 16 a (4 I) [MZSP]; Winkler 16 b (4 I) [MZSP]; Winkler 16 - 2 (1 W 5 I) [MZSP]; 29. iv. 1992, S. Ide (1 I) [MZSP]; 12 - 17. vi. 1997, B. H. Dietz & C. I. Yamamoto col., Transect I, Winkler 4 (1 Q 1 I) [MZSP]; same except Winkler 14 a (1 Q) [MZSP]; Winkler 16 a (1 I) [MZSP]; Winkler 16 b (1 I) [MZSP]; Winkler 17 a (1 I) [MZSP]; Winkler 17 b (1 I) [MZSP]; Winkler 24 a (2 I) [MZSP]; Winkler 24 b (1 I) [MZSP]; Winkler 25 a (1 I) [MZSP]; 5 - 7. vii. 1997, C. I. Yamamoto col., Transect I, Winkler 2 (1 I) [MZSP]; same except Winkler 4 (1 Q 1 I) [MZSP]; Winkler 5 (1 I) [MZSP]; Winkler 6 (1 Q) [MZSP]; Winkler 10 (1 I) [MZSP]; Winkler 11 (2 I) [MZSP]; Winkler 12 (1 I) [MZSP]; Winkler 15 (1 I) [MZSP]; Winkler 19 (2 I) [MZSP]; Winkler 25 (1 I) [MZSP]; Estação Biológica de Boracéia, 21. v- 28. vi. 2008, Malaise, 23 ° 38 ′ 28 ″ S, 45 ° 51 ′ 22 ″ O, Nihei, S. Figueiredo, R. & Andrade, M. cols. (1 Q 7 M) [MZSP]; Boraceia Biological Sta [Station], 850 m, wet mt. forest, feb. 67, W. L. Brown & R. Croizer cols. (1 W 2 I) [MCZC]; same except 24 - 28. may. 1971, 850 m, wet mt. forest (1 I) [MCZC]; 3 - 5. v. 1996, Brandão, Agosti, Diniz, Silvestre & Yamamoto (1 I) [MZSP]; 23 ° 31 ′ 56 ″ S, 45 ° 50 ′ 47 ″ W, em tronco caído (1 I) [MZSP]; em tronco caído 36, imaturos em álcool [immatures in alcohol] [these are not immatures of Hylomyrma] (1 I) [MZSP]; Winkler, folhiço (4 I) [MZSP]; Trilha do Divisor, Malaise, 23 ° 39 ’ 18.2 ″ S, 45 ° 53 ’ 18 ″ W, 23. ii. 2005, 06: 00 - 18: 00 hs, J. C. de Souza & A. P. Aguiar cols., HYMPAR-FAPESP (18 M) [MZSP]; same except R. S. M. Feitosa & A. P. Aguiar cols., HYMPAR-FAPESP (9 M) [MZSP]; same data (1 Q 1 M) (MZSP 67431) [MZSP]; same except 25. ii. 2005, 09: 00 - 10: 00 hs, J. C. de Souza & A. P. Aguiar cols., HYMPAR-FAPESP (1 M) [MZSP]; 10: 00 - 11: 00 hs, J. C. de Souza & A. P. Aguiar cols., HYMPAR-FAPESP (7 M) [MZSP]; 11: 00 - 12: 00 hs, R. S. M. Feitosa & A. P. Aguiar cols., HYMPAR-FAPESP (2 M) [MZSP]; 14: 00 - 16: 00 hs, R. S. M. Feitosa & A. P. Aguiar cols., HYMPAR-FAPESP (13 M) [MZSP]; 25 - 26. ii. 2005, 18: 00 - 06: 00 hs, J. C. de Souza & A. P. Aguiar cols., HYMPAR-FAPESP (1 M) [MZSP]; 27. ii. 2005, 16: 00 - 17: 00 hs, R. S. M. Feitosa & A. P. Aguiar cols., HYMPAR-FAPESP (2 M) [MZSP]; 17: 00 - 18: 00 hs, R. S. M. Feitosa & A. P. Aguiar cols., HYMPAR-FAPESP (1 M) [MZSP]; 23 ° 39 ’ 25.7 ″ S, 45 ° 53 ’ 23 ″ W, 18 - 22. iv. 2003, A. P. Aguiar & F. M. Rodriguez cols., pt 9, BIOTA-FAPESP (1 M) [MZSP]; 23 ° 39 ’ 18.1 ″ S, 45 ° 53 ’ 18.2 ″ W, pt 8, BIOTA-FAPESP (5 M) [MZSP]; 23 ° 39 ’ 22.9 ″ S, 45 ° 53 ’ 48.1 ″ W, 18 - 28. iv. 2003, pt 10, BIOTA-FAPESP (1 M) [MZSP]; Trilha dos Pilões, Malaise, 23 ° 39 ’ S, 45 ° 53 ’ W, 18 - 28. iv. 2003, pt 3, BIOTA-FAPESP (1 M) [MZSP]; same except 23 ° 39 ’ 05.1 ″ S, 45 ° 53 ’ 51.8 ″ W, 23. ii. 2005, 06: 00 - 18: 00 hs, A. P. Aguiar & L. K. Nogueira cols., HYMPAR-FAPESP (16 M) [MZSP]; 24. ii. 2005, 11: 00 - 12: 00 hs, L. K. Nogueira & A. P. Aguiar cols., HYMPAR-FAPESP (1 M) [MZSP]; 25. ii. 2005, 09: 00 - 10: 00 hs, L. K. Nogueira & A. P. Aguiar cols., HYMPAR-FAPESP (1 M) [MZSP]; 10: 00 - 11: 00 hs, L. K. Nogueira & A. P. Aguiar cols., HYMPAR-FAPESP (6 M) [MZSP]; 10: 00 - 11: 00 hs, A. P. Aguiar & L. K. Nogueira cols., HYMPAR-FAPESP (2 M) [MZSP]; 11: 00 - 12: 00 hs, A. P. Aguiar & L. K. Nogueira cols., HYMPAR-FAPESP (1 M) [MZSP]; 26. ii. 2005, 08: 00 - 09: 00 hs, A. P. Aguiar & L. K. Nogueira cols., HYMPAR-FAPESP (1 M) [MZSP]; 27. ii. 2005, 07: 00 - 08: 00 hs, L. K. Nogueira & A. P. Aguiar cols., HYMPAR-FAPESP (2 M) [MZSP]; 16: 00 - 17: 00 hs, A. P. Aguiar & L. K. Nogueira cols., HYMPAR-FAPESP (1 M) [MZSP]; São Paulo, P. E. da Cantareira, Núcleo Engordador, 23 ° 21 ′ 27 ″ S, 46 ° 29 ′ 42 ″ W, 12 - 22. v. 2003, R. Feitosa & A. Soliva cols., Winkler 3 (2 I) [MZSP]; same except Winkler 6 (1 Q 1 I) [MCZC]; Winkler 18 (3 I) [MZSP]; Winkler 23 (1 Q 1 I) [MZSP]; Winkler 28 (2 I) [MZSP]; Winkler 33 (2 I) [MZSP]; São Vicente, Parque Estadual Xixová-Japuí, 23 ° 59 ′ S, 46 ° 23 ’ W, 01. vii. 2011, transecto 1, pitfall 9 (1 I) [MZSP]; same except 15. v. 2011, mata secundária, transecto 2, winkler 9 (1 I) [MZSP]; 23. iv. 2011, mata sede, transecto 2, winkler 3 (2 I) [CASC]; transecto 1, winkler 9 (1 I) [MZSP]; 17 - 18. vii. 2010 (1 I) [MZSP]; Tapiraí, 08 - 14. i. 2001, 24 ° 01 ′ 55 ″ S, 47 ° 27 ′ 56 ″ W, Silva & Eberhardt cols., Transecto I, Winkler 01 (1 I) [MZSP]; same except Winkler 03 (1 I) [MZSP]; Winkler 06 (1 Q) [MZSP]; Winkler 09 (2 I) [MZSP]; [MZSP]; Winkler 14 (1 I) [MZSP]; Winkler 23 (1 I) [MZSP]; Winkler 24 (1 W) [MZSP]; Winkler 26 (1 I) [MZSP]; Winkler 27 (1 I) [MZSP]; Winkler 34 (1 I) [MZSP]; Winkler 37 (1 I) [MZSP]; Winkler 43 (1 I 1 M) [MZSP]; Winkler 45 (1 I) [MZSP]; Ubatuba, P. E. S. M., N. Picinguaba, alt. 1000 m, 23 ° 17 ′ 56.4 ″ S, 44 ° 47 ′ 13.2 ″ W, 13. iii. 2006, Winkler 19 (2 I) [MZSP]; same except Winkler 20 (2 I) [MZSP]; Winkler 21 (2 I) [MZSP]; Winkler 23 (2 I) [MZSP]; Winkler 24 (1 W 1 I) [MZSP]; Winkler 25 (1 I) [MZSP]; 10. iii. 2006, Winkler 11 (2 I) [MZSP]; Winkler 12 (2 I) [MZSP]; Winkler 13 (2 I) [MZSP]; Winkler 14 (1 W) [MZSP]; Winkler 16 (1 W) [MZSP]; Winkler 17 (1 W) [MZSP]; Winkler 18 (2 I) [MZSP]; 03. iii. 2006, Winkler 2 (2 I) [MZSP]; Winkler 3 (1 I) [MZSP]; Winkler 4 (1 I) [MZSP]; Winkler 5 (3 I) [MZSP]; 07. iii. 2006, Winkler 8 (1 W) [MZSP]; Winkler 9 (1 W 1 I) [MZSP]; 800 m, 23 ° 17 ′ 49.2 ″ S, 44 ° 47 ′ 31.2 ″ W, 10. iii. 2006, Scott-Santos C. P. & Santos E. F. cols., Winkler 22 (1 W 1 I) [MZSP]; Winkler 23 (1 W 1 I) [MZSP]; Winkler 24 (2 W) [MZSP]; 23. i. 2006, Winkler 2 (1 I) [MZSP]; Winkler 3 (1 W 1 I) [MZSP]; Winkler 4 (2 I 1 M) [CASC]; Winkler 5 (2 I) [MZSP]; Winkler 6 (2 I) [MZSP]; 03. iii. 2006, Winkler 15 (2 I) [MZSP]; Winkler 16 (2 I) [MZSP]; 13. iii. 2006, Winkler 25 (1 W 1 I) [MZSP]; Winkler 26 (2 I) [MZSP]; 07. iii. 2006, Winkler 18 (2 I) [MZSP]; Winkler 19 (2 I) [MZSP]; Winkler 20 (2 I) [MZSP]; 26. i. 2006, Winkler 7 (2 I) [MZSP]; Winkler 8 (2 I) [MZSP]; Winkler 9 (1 W 1 I) [MZSP]; Winkler 10 (2 I) [MZSP]; Winkler 11 (2 I) [MZSP]; 600 m, 23 ° 17 ′ 54.4 ″ S, 44 ° 47 ′ 49.2 ″ W, 05. ii. 2006, Winkler 19 (1 I) [MZSP]; Winkler 20 (2 I) [MZSP]; Winkler 22 (1 W 1 I) [MZSP]; 23. i. 2006, Winkler 2 (2 I) [MZSP]; Winkler 3 (2 W) [MZSP]; Winkler 6 (1 W 1 I) [MZSP]; Winkler 7 (1 Q 2 I) [MZSP]; Winkler 8 (1 I) [MZSP]; 26. i. 2006, Winkler 9 (2 I) [MZSP]; Winkler 10 (2 I) [MZSP]; Winkler 15 (2 I) [MZSP]; Winkler 17 (2 I) [MZSP]; 16. iii. 2006, Winkler 21 (1 W 1 I) [MZSP]; Winkler 23 (2 I) [MZSP]; Winkler 24 (1 W 1 I) [MZSP]; Winkler 25 (1 W 1 I) [MZSP]; 400 m, 23 ° 18 ′ 21.6 ″ S, 44 ° 48 ′ 25.2 ″ W, 24. iii. 2006, Winkler 21 (2 I) [MZSP]; Winkler 22 (1 W) [MZSP]; Winkler 24 (1 W) [MZSP]; 19. iii. 2006, Winkler 11 (1 W 1 I) [MZSP]; Winkler 14 (1 I) [MZSP]; Winkler 15 (1 I) [MZSP]; Winkler 16 (2 I) [MZSP]; 16. iii. 2006, Winkler 8 (3 I) [MZSP]; Winkler 13 (1 W) [MZSP]; 05. i. 2006, Winkler 1 (2 I) [MZSP]; Winkler 3 (1 I) [MZSP]; 200 m, 23 ° 19 ′ 08.4 ″ S, 44 ° 49 ′ 4.8 ″ W, 24. iii. 2006, Winkler 21 (1 I) [MZSP]; Winkler 22 (1 I) [MZSP]; Winkler 23 (2 I) [MZSP]; Winkler 24 (2 I) [MZSP]; Winkler 25 (1 Q 1 I) [MZSP]; 18. iii. 2006, Winkler 14 (2 W 1 I) [MZSP]; Winkler 18 (2 I) [MZSP]; 18. i. 2006, Winkler 1 (1 I) [MZSP]; Winkler 4 (2 I) [MZSP]; Winkler 5 (3 I) [UTEP]; Winkler 7 (3 I) [MZSP]; Winkler 8 (1 W 1 Q) (MZSP 67430) [MZSP]; Winkler 9 (2 I) [MZSP]; 21. i. 2006, Winkler 11 (1 I) [MZSP]; Winkler 12 (1 Q 2 I) [MZSP]; Winkler 12 (4 I) [UTLP]; Winkler 13 (3 I) [MZSP]; 50 m, 23 ° 20 ′ 24 ″ S, 44 ° 49 ′ 26.4 ″ W, 21. i. 2006, Winkler 13 (1 W 1 I) [MZSP]; 18. i. 2006, Winkler 1 (1 Q 1 I 1 indet.) [MZSP]; Winkler 5 (2 I) [MZSP]; Winkler 7 (2 I) [MZSP]; Winkler 8 (1 I 1 indet.) [MZSP]; Winkler 9 (1 I) [MZSP]; 18. iii. 2006, Winkler 15 (2 I) [MZSP]; Winkler 16 (2 I) [MZSP]; Winkler 19 (1 W 1 I) [MZSP]; Winkler 20 (1 I) [MZSP]; 24. iii. 2006, Winkler 22 (2 I) [MZSP]; Winkler 23 (1 I) [MZSP]; Winkler 25 (1 Q 1 I) [MZSP]; Winkler 26 (2 I) [MZSP]; P. E. Serra do mar, N. Picinguaba, 23 ° 17 ′ 49.2 ″ S, 44 ° 47 ′ 31.20 ″ W, 03 - 14. iii. 2008, Esteves F. A. & Feitosa R. M. cols., altitude 800 m, armadilha subterrânea - # 14 (1 I) [MZSP]; same except 23 ° 17 ′ 56.40 ″ S, 44 ° 47 ′ 49.20 ″ W, altitude 600 m, isca de sardinha sobre o solo - # 25 (1 I) [MZSP]. PARAGUAY: Alto Parana: Pto [Puerto] Pte [Presidente] Stroessner [today Ciudad del Este], 6. xi. 79, V. Mahnert, 52 (1 W 1 I) [MHNG]; Itapua: Salto Tembey, 1. xi. 82, F. Baud et al. of MHN-Geneva, forest, lf. litt. & wood, winkler apparat (1 Q 3 I) [MHNG]; same except C. Dlouhy, 82 / 20 (1 I) [MHNG].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC4FFC7F0A5F0A9FDF0FCB8.taxon	description	Figures 69, 70, 83 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC4FFC7F0A5F0A9FDF0FCB8.taxon	diagnosis	Diagnosis. Vermicular striae on head dorsum and mesosoma; propodeal spine long; dorsal margin of petiole continuous, convex; petiole ventral surface smooth; longitudinal and anastomosed striae on postpetiole and tergum of first gastral segment; subpostpetiolar process prominent, convex; profemur almost entirely striate, apical region of posterior surface smooth; protibia entirely striate; long striae on tergum of first gastral segment.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC4FFC7F0A5F0A9FDF0FCB8.taxon	description	Redescription. WORKER (n = 4) (Fig. 69 A – C): HL 1.02 (1.02 – 1.10); HW 0.98 (0.98 – 1.03); ML 0.68 (0.68 – 0.72); SL 0.74 (0.74 – 0.80); MOD 0.27 (0.26 – 0.28); PNW 0.72 (0.72 – 0.78); WL 1.38 (1.38 – 1.50); PSL 0.42 (0.42 – 0.46); PL 0.66 (0.64 – 0.68); PW 0.26 (0.26 – 0.28); PPL 0.40 (0.36 – 0.42); PPW 0.37 (0.37 – 0.40); GL 1.10 (1.10 – 1.28); TL 5.24 (5.24 – 5.70); CI 96.08 (93.63 – 97.11); SI 75.51 (75.51 – 79.20); OI 27.55 (26.53 – 27.72). Large-sized. Shiny integument. Dark brown body, lighter leg. Thin and unbranched setae, long to midsized, erect to subdecumbent. Head subquadrate; posterior margin slightly concave at middle. Mandible masticatory margin with 5 teeth. Anterior margin of clypeus slightly concave medially, with a pair of large and blunt teeth laterally; median area of clypeus with 10 irregular to regular, longitudinal striae converging to a point on the anterior margin, interspaces distinguishable. Frontal triangle with 1 sagittal stria. Short scape, not reaching head posterior margin; apical antennomere shorter than previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye drop-shaped, midsized, larger diameter with 13 ommatidia. Head dorsum with vermicular to vermiculatedareolated striae, divergent and anastomosed towards posterior margin, interspaces between striae smooth. Striae on head lateral and laterodorsal regions converge to mandible insertion; longitudinal, irregular to vermicular striae on head lateral; longitudinal striae posteriorly anastomosed on lateral and lateroventral regions; gena striate, around 4 – 6 regular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Vermicular striae of uniform thickness on mesosoma dorsum, transverse on pronotum anterior region and propodeum (DV), longitudinal on other parts, interspaces smooth. Promesonotal junction and metanotal groove indistinct. Transverse carina well-marked. Dorsal margin of mesosoma continuous, convex. Propodeal lobe bidentate, dorsal tooth slightly longer and more acute than the shorter and blunt ventral tooth; dorsal tooth length 1 / 2 of propodeal spine length. Propodeal spine long, straight (LV), divergent (DV), sculptured on base. Thin striae of uniform thickness on procoxa; irregular and transverse striae on C 2 and C 3. Irregular to regular transverse striae on profemur anterior and posterior surfaces, apical region smooth; ventral surface smooth. Protibia entirely striate. Dorsal margin of petiole continuous, convex; mostly covered with irregular to vermicular striae; ventral surface smooth. Longitudinal and anastomosed striae on postpetiole; subpostpetiolar process striae restricted to lateral region, midbasal region smooth; subpostpetiolar process prominent, convex. First gastral segment striation similar to postpetiole striae; long striae on tergum, length similar to postpetiole length; sternite striation covering the laterobasal region.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC4FFC7F0A5F0A9FDF0FCB8.taxon	materials_examined	QUEEN (n = 1, paratype) (Fig. 70 A – C): HL 1.08; HW 1; ML 0.68; SL 0.78; MOD 0.31; PNW 0.90; WL 1.64; PSL 0.44; PL 0.74; PW 0.30; PPL 0.46; PPW 0.44; GL 1.32; TL 5.92; CI 92.60; SI 78; OI 31. Large-sized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 16 ommatidia. Longitudinal, irregular, slightly anastomosed striae on scutum going from an anterior central point towards transcutal suture, interspaces distinguishable. Longitudinal striae with interspaces distinguishable on anepisternum and katepisternum, mostly irregular and anastomosed on anepisternum and regular on katepisternum. Axilla with mostly anastomosed striae, striation inclinated to anteroposterior body axis. Scutoscutellar sulcus inconspicuous. Mostly vermicular striae on scutellum. Transverse striae on propodeum (DV). Lateral of mesosoma and laterally with irregular to vermicular striae directed to propodeal spine, interspaces distinguishable. Wings unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC4FFC7F0A5F0A9FDF0FCB8.taxon	etymology	Etymology. Sagax (Latin) = wise, sagacious.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC4FFC7F0A5F0A9FDF0FCB8.taxon	discussion	Comments. This species is similar to H. virginiae and H. mitiae in the vermicular sculpture on the head dorsum and mesosoma, and the anastomosed striae on the postpetiole and first gastral tergite. Hylomyrma sagax is distinguished from H. virginiae (characteristic in parentheses) based on the following characters: a pair of large teeth laterally at the clypeus anterior margin (vs. a pair of medium teeth), the propodeal spine comparatively longer and needle-like (vs. shorter and thicker), and the continuous dorsal margin of petiole (vs. discontinuous). Hylomyrma sagax is distinguished from H. mitiae in the pair of large teeth laterally at the clypeus anterior margin (vs. medium teeth), the presence of smooth interspaces between the vermicular striae on the entire mesosoma (vs. smooth interspaces only on the mesosomal dorsum; the lateral of mesosoma is covered with very thin striae with indistinguishable interspaces), and the convex subpostpetiolar process (vs. subtriangular). All three species are allopatric; Hylomyrma sagax is restricted to southeast Colombia (Fig. 83), whereas H. mitiae occurs in French Guiana (Fig. 87), and H. virginiae is recorded in Ecuador (both sides of The Andes) and in western Colombia (Fig. 89).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC4FFC7F0A5F0A9FDF0FCB8.taxon	distribution	Distribution. This species is only known from southeast Colombia (Fig. 83).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC4FFC7F0A5F0A9FDF0FCB8.taxon	materials_examined	Natural history. Only one worker is known, apart from the type specimens. All material examined was collected from leaf-litter of tropical rainforest, which suggests that nests are located in fallen logs, rotten wood, between leaves, or inside natural cavities of the superficial soil layers. Additional material examined (1 worker): COLOMBIA: Caquetá: [Puerto] Solano, PNN [Serranía de] Chiribiquete, R. Sararamano, B. Verde military, 300 m, 0 ° 10 ′ 48 ″ N, 72 ° 37 ′ 24 ″ W, Winkler 14, 9. iv. 2000, E. González leg. (1 W covered with gold) [IHVL].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC7FFDAF0A5F7ADFB81F9B4.taxon	description	Figures 71, 72, 80 H, 89 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC7FFDAF0A5F7ADFB81F9B4.taxon	diagnosis	Diagnosis. Regular and longitudinal striae on head dorsum, parallel towards posterior margin, interspaces between thicker striae filled with thinner striae; mesosoma covered with striae of variable thickness assuming multiple directions, interspaces indistinguishable; transverse on mesonotum; dorsal margin of mesonotum straight; lateral of pronotum and mesepisternum with longitudinal striae in part continuing transversely on propodeum dorsum and in part continuing on propodeal spine; dorsal margin of petiole discontinuous; petiole ventral surface entirely covered with transverse striae; regular to irregular and mostly transverse striae on dorsal surface of petiole, interspaces indistinguishable; subpostpetiolar process striate; profemur posterior surface and protibia extensor surface entirely striate; long striae on tergum of first gastral segment; sternite striation weakly marked, covering all basal region; branched setae with multiple branches arising from the main axis; branches relatively long, subequal-sized; branched setae mainly in posterior and lateral regions of head, mesosoma, petiole and postpetiole.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC7FFDAF0A5F7ADFB81F9B4.taxon	description	Redescription. WORKER (n = 4) (Fig. 71 A – C): HL 0.98 (0.88 – 0.98); HW 0.90 (0.82 – 0.90); ML 0.58 (0.54 – 0.58); SL 0.68 (0.64 – 0.70); MOD 0.27 (0.24 – 0.28); PNW 0.64 (0.60 – 0.64); WL 1.26 (1.15 – 1.26); PSL 0.28 (0.22 – 0.28); PL 0.56 (0.48 – 0.56); PW 0.24 (0.22 – 0.24); PPL 0.31 (0.30 – 0.32); PPW 0.36 (0.33 – 0.36); GL 1.06 (1.04 – 1.06); TL 4.75 (4.40 – 4.80); CI 91.83 (90.81 – 93.18); SI 75.55 (75.55 – 78.65); OI 30 (29.26 – 31.46). Small-sized. Integument shiny to subopaque. Body light brown, head and gaster darker. Many long to short setae, appressed to decumbent; unbranched setae mostly on head dorsum and leg; branched setae with multiple branches arising from the main axis; branches relatively long, subequal in size (Fig. 80 H); branched setae mainly in posterior and lateral regions of head, mesosoma, petiole and postpetiole. Head subquadrate; posterior margin straight to slightly concave at middle. Mandible masticatory margin with 6 teeth. Anterior margin of clypeus straight medially, with a pair of medium teeth laterally; median area of clypeus with regular and longitudinal striae converging to a point on the anterior margin, interspaces indistinguishable. Frontal triangle with 1 – 3 striae. Short scape, not reaching head posterior margin; apical antennomere length 1 / 4 shorter than previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye drop-shaped, midsized, larger diameter with 14 ommatidia. Regular and longitudinal striae on head dorsum, parallels towards posterior margin, interspaces between thicker striae filled with thinner and regular to irregular striae. Head lateral and laterodorsal regions with the same striation of head dorsum, striae converging to eye margin; gena striate, regular and semicircular striae circumscribe the torulus almost reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma covered with striae of variable thickness assuming multiple directions, interspaces indistinguishable; transverse striae on pronotum anterior region (DV) continuing towards lateral of pronotum; longitudinal striae on promesonotal junction and metanotal groove; transverse striae on mesonotum; lateral of pronotum and mesepisternum with longitudinal striae in part continuing transversely on propodeum dorsum and in part continuing on propodeal spine. Dorsal margin of mesonotum straight. Promesonotal junction and metanotal groove discernible by a depression and altered sculpture. Transverse carina inconspicuous. Dorsal margin of mesosoma discontinuous, dorsal margin of mesonotum straight. Propodeal lobe bidentate, dorsal tooth slightly longer and more acute than shorter and blunt ventral tooth; dorsal tooth length is 1 / 2 of propodeal spine length. Propodeal spine midsized, straight (LV), divergent (DV), sculptured on base. Procoxa with transverse striae of uniform thickness; irregular and transverse striae on C 2 and C 3. Transverse striae on profemur. Protibia extensor surface entirely striate. Dorsal margin of petiole discontinuous; ventral surface entirely covered with transverse striae. Node with transverse striae on anterior surface continuing on lateral surface, interspaces distinguishable; regular to irregular and mostly transverse striae on dorsal surface, interspaces indistinguishable. Postpetiole and subpostpetiolar process with regular, longitudinal striae; subpostpetiolar process weak, straight to convex. First gastral segment striation similar to postpetiole striae; striae length on tergum slightly longer than postpetiole length; sternite striation weakly marked, covering all basal region.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC7FFDAF0A5F7ADFB81F9B4.taxon	materials_examined	QUEEN (n = 1, paratype) (Fig. 72 A – C): HL 0.98; HW 0.90; ML 0.60; SL 0.70; MOD 0.28; PNW 0.70; WL 1.40; PSL 0.26; PL 0.59; PW 0.27; PPL 0.36; PPW 0.36; GL 1.22; TL 5.15; CI 91.83; SI 77.77; OI 31.11. Midsized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 15 ommatidia. Longitudinal and regular striae on scutum going from an anterior central point towards transcutal suture, interspaces indistinguishable. Longitudinal striae on anepisternum and katepisternum with interspaces indistinguishable, striae regular on anepisternum and mostly irregular on katepisternum. Axilla with the same sculpture of scutum. Scutoscutellar sulcus conspicuous. Scutellum with the same sculpture of scutum, striae slightly irregular to vermicular on lateral regions. Transverse striae on propodeum (DV). Lateral of mesosoma with longitudinal and mostly regular striae directed to propodeal dorsum, interspaces indistinguishable. Wings unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC7FFDAF0A5F7ADFB81F9B4.taxon	description	MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC7FFDAF0A5F7ADFB81F9B4.taxon	etymology	Etymology. The name transversa (Latin, transversus = transverse) refers to the conspicuous transverse striae on mesonotal dorsum.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC7FFDAF0A5F7ADFB81F9B4.taxon	discussion	Comments. This species is easily distinguished from all congeners by the transverse striae on the mesonotum, and the conspicuous branched setae. Still, H. transversa resembles H. reitteri, regarding the discontinuous dorsal margin of petiole, and the head pattern of sculpturation. The two species are allopatric and have broad distributions; H. transversa occurs in the center-northwest of South America (Fig. 89), and H. reitteri in the center-southeast of South America (Fig. 85). Hylomyrma transversa can be easily distinguished from H. reitteri (characteristic in parentheses) in the drop-shaped eye (vs. reniform), the transverse striae on the mesonotal dorsum (vs. longitudinal striae), and the striate ventral surface of petiole (vs. smooth surface). Specimens of H. transversa collected at Colombia’s Parque Nacional Natural Serranía de Chiribiquete have slightly thicker setae, lack branched setae, and the transverse striae on the mesonotum are more weakly marked. We found ten intercastes among specimens examined (four from Arquipélago de Anavilhanas, AM, Brazil; two collected at Rio Negro, and four sampled at Lower Rio Madidi, both in Bolivia), identified as such for being slightly larger than workers [(n = 2), HL (0.98); HW (0.90 – 0.98); ML (0.60 – 0.64); SL (0.68 – 0.72); MOD (0.26 – 0.28); PNW (0.62 – 0.72); WL (1.24 – 1.30); PSL (0.24 – 0.28); PL (0.52 – 0.58); PW (0.22 – 0.28); PPL (0.30 – 0.32); PPW (0.32 – 0.37); GL (0.94 – 1.13); TL (4.64 – 5.02)]; for having one central ocellus, and a discernible promesonotal junction and metanotal groove by a conspicuous depression. Moreover, the specimens from Lower Rio Madidi, Bolivia, have small protuberances on the mesonotum, in the region where wings would be inserted in queens.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC7FFDAF0A5F7ADFB81F9B4.taxon	distribution	Distribution. All known specimens were collected in northwestern South America (Brazilian Amazon, Bolivia, Colombia, and Peru) (Fig. 89). Natural history. This species inhabits areas of tropical rainforest. Five specimens were collected with pitfall traps, and another was allured by an attractive bait, which suggests that workers forage on forest floor and that nests are located in fallen logs, rotten wood, between leaves, or inside natural cavities of the superficial soil layers.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFC7FFDAF0A5F7ADFB81F9B4.taxon	materials_examined	Additional material examined (27 workers, 8 intercastes): BOLIVIA: [La Paz]: Lower Rio Madidi, W. M. Mann, Mulford BioExpt 1921 – 22, January (7 W 2 I) [USNM]; same except Jan (4 W 2 I) [USNM]; Rio Negro, W. M. Mann collector, Mulford Biological Exploration 1921 – 22, Jan (2 I) [MZSP]; same data (2 W) [CASC, DZUP]. [BRAZIL]: AM [Amazonas]: Itacoatiara, 3.319 ° S, 58.723 ° W, PLOT 63 — isca # 15. J. M. S. Vilhena, 6 / 114 / 03 (1 W) (MZHY 193) [MZSP]; Novo Airão, PNJ R. Unini, mg. esq., com Lago Pedras, 1 ° 38 ′ 25 ″ S, 61 ° 39 ′ 19 ″ W, Mata Primária, 20 – 23. xi. 1995, Bindá + Alencar, Pitfall trap # 5 (1 W) [INPA]; same except Pitfall trap # 7 (1 W) [INPA]; Tarumã-Mirim, 31. iii. 1976, Joaquim Dias (1 W) [MZSP]; Rio Negro, Arquipélago d. Anavilhanas, iii. 76, R. Negre, # 13186 (2 W 2 I, one worker covered with gold) (MZSP 67452) [MZSP]; PA [Pará]: Porto Trombetas, 01.08.92, Majer J. D., 4552, Jam 76 (2 W) [CEPLAC]. COLOMBIA: Caquetá: [Puerto] Solano, PNN [Serranía de] Chiribiquete, R. Mesay, B. Caki, 300 m, 0 ° 14 ′ 24 ″ N, 72 ° 56 ′ 02 ″ W, T. caída, T 1. T 3, 27. i. 2000, F. Quebedo leg. (2 W, one covered with gold) [IHVL]; same except T 1. T 2 (1 W) [MZSP]; Cauca: Isla Gorgona, 14. xi. 1987, M. Baena (1 W) [IHVL]; Guaviare: RN Nukak Maku, 02 ° 10 ′ 40 N, 71 ° 11 ′ 25 W, Cñ Cucuy Cr Moyano, Rebalse, F. Fernandez F. E., feb. 96 (1 W without postpetiole and gaster) [IHVL]; same data (1 W) (MZHY 221) [MZSP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFDAFFD0F0A5F2A1FC35FAE8.taxon	description	Figures 73, 74, 75, 80 F, 80 J, 81 C, 82 C, 88 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFDAFFD0F0A5F2A1FC35FAE8.taxon	diagnosis	Diagnosis. Irregular and longitudinal striae on head dorsum slightly diverge towards posterior margin, interspaces between thicker striae mostly smooth or filled with thinner striae; irregular striae on mesosoma mostly longitudinal, interspaces between thick and thin striae distinguishable by smooth areas; dorsal margin of petiole discontinuous; petiole ventral surface almost entirely covered with transverse striae, anterior region smooth; node dorsal surface mostly covered with irregular striae; subpostpetiolar process smooth at middle and laterally striate; profemur and protibia mostly smooth; striae length on tergum slightly shorter than postpetiole length; branched setae of two types: 1) thin setae with multiple small branches of subequal size arising from the main axis; 2) thick setae flattened at its final half, with several branches of subequal size.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFDAFFD0F0A5F2A1FC35FAE8.taxon	description	Redescription. WORKER (n = 6) (Fig. 73 A – C): HL 0.94 (0.88 – 0.97); HW 0.86 (0.84 – 0.92); ML 0.56 (0.54 – 0.60); SL 0.60 (0.60 – 0.66); MOD 0.22 (0.22 – 0.25); PNW 0.62 (0.60 – 0.65); WL 1.20 (1.16 – 1.26); PSL 0.26 (0.22 – 0.26); PL 0.56 (0.52 – 0.60); PW 0.28 (0.25 – 0.28); PPL 0.32 (0.28 – 0.34); PPW 0.33 (0.33 – 0.35); GL 1 (1 – 1.14); TL 4.58 (4.40 – 4.88); CI 91.49 (91.30 – 95.83); SI 69.76 (69.56 – 75); OI 25.58 (23.80 – 27.17). Small-sized. Shiny integument. Brownish body, lighter appendices, darker gaster. Mostly thin setae, unbranched or branched, long to midsized, suberect to subdecumbent; branched setae of two types: 1) thin setae with multiple small branches of subequal size arising from the main axis, mostly in mesosoma (Fig. 80 F); 2) thick setae flattened in its final half, with several branches of subequal size, mainly in postpetiole and gaster (Fig. 80 J). Head subquadrate; posterior margin straight to slightly concave at middle. Mandible masticatory margin with 6 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with 7 – 9 regular to irregular, longitudinal and thick striae converging to a point on the anterior margin, interspaces between thick striae mostly smooth. Frontal triangle with 1 – 3 striae. Short scape, not reaching head posterior margin; apical antennomere as long as previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye reniform, midsized, larger diameter with 11 ommatidia. Irregular and longitudinal striae on head dorsum, slightly divergent towards posterior margin, interspaces between thicker striae mostly smooth or filled with thinner and regular to irregular striae. Head lateral and laterodorsal regions with irregular to vermicular striae converging to eye margin; anterior laterodorsal region rugose; gena striate, 2 – 5 irregular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma covered with irregular striae mostly longitudinal, interspaces between thick and thin striae distinguishable by smooth areas; transverse striae on pronotum anterior region (DV) directed towards lateral of pronotum, striae mostly longitudinal in other regions (DV). Promesonotal junction and metanotal groove discernible by a slight depression. Transverse carina inconspicuous. Dorsal margin of mesosoma discontinuous, convex. Propodeal lobe bidentate, dorsal tooth slightly longer and more acute than the shorter and blunt ventral tooth; dorsal tooth length is slightly shorter than propodeal spine length. Propodeal spine short, straight (LV), divergent (DV), sculptured. Procoxa with transverse striae of uniform thickness; irregular and transverse striae on C 2 and C 3. Profemur and protibia mostly smooth. Dorsal margin of petiole discontinuous; ventral surface almost entirely covered with transverse striae, anterior region smooth. Convex node; transverse, regular to irregular, striae on anterior surface continuing on lateral surface, interspaces distinguishable; mostly longitudinal, regular to irregular striae of variable thickness on dorsal surface, interspaces distinguishable. Postpetiole and lateral regions of subpostpetiolar process with regular, longitudinal, adjacent striae; subpostpetiolar process weak, straight to convex, smooth at middle. First gastral segment striation thinner than postpetiole striae; striae length on tergum slightly shorter than postpetiole length; sternite striation weakly marked, covering all the basal region. QUEEN (n = 4) (Fig. 74 A – C): HL 0.98 (0.94 – 1); HW 0.90 (0.90 – 0.95); ML 0.61 (0.58 – 0.62); SL 0.66 (0.64 – 0.70); MOD 0.26 (0.26 – 0.28); PNW 0.78 (0.74 – 0.82) CMS 1.44 (1.40 – 1.50); PSL 0.30 (0.26 – 0.30); PL 0.63 (0.63 – 0.70); PW 0.30 (0.28 – 0.30); PPL 0.34 (0.34 – 0.40); PPW 0.40 (0.38 – 0.42); GL 1.30 (1.24 – 1.36); TL 5.30 (5.18 – 5.58); CI 91.83 (91.83 – 95.92); SI 73.33 (71.11 – 73.68); OI 28.88 (28.88 – 29.78). Large-sized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 15 ommatidia. Longitudinal, regular to irregular and thick striae of variable thickness on scutum going from an anterior central point towards transcutal suture, interspaces between thicker striae filled with thinner striae. Longitudinal striae on anepisternum and katepisternum with interspaces distinguishable, striae mostly regular on anepisternum, irregular to vermicular on katepisternum. Axilla and scutellum with longitudinal, regular to irregular and slightly vermicular, with interspaces distinguishable. Scutoscutellar sulcus inconspicuous. Transverse and irregular striae on propodeum (DV). Lateral of mesosoma with longitudinal, mostly irregular to vermicular striae directed in part to propodeal dorsum and in part to propodeal spine, interspaces between thicker striae filled with thinner striae. Wings as in Fig. 81 C. MALE (first description) (n = 1) (Fig. 75 A – C): HL 0.75; HW 0.68; ML 0.44; SL 0.25; MOD 0.30; PNW 0.62; WL 1.30; PL 0.54; PW 0.26; PPL 0.30; PPW 0.30; GL 1.06; TL 4.39; CI 90.66; SI 36.76; OI 44.11. Head, mesosoma and appendices light brown, other parts dark brown. Mandible with 5 teeth. Interspaces between thicker striae on head dorsum partly smooth and partly filled with thinner striae. Scutum mostly smooth on anterior region and with few thinner, regular to irregular striae on posterior region, interspaces distinguishable. Anepisternum mostly covered with longitudinal, regular and thinner striae, interspaces distinguishable; katepisternum mostly smooth. Scutellum with regular to irregular striae, thicker than scutum striae, interspaces distinguishable. Propodeum with irregular striae assuming multiple directions, interspaces distinguishable. Dorsal region of propodeal lobe forming a blunt tooth, ventral region rounded. Petiole dorsum covered with irregular striae assuming multiple directions, interspaces distinguishable. Postpetiole and gaster missing. Wings as in Fig. 82 C.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFDAFFD0F0A5F2A1FC35FAE8.taxon	etymology	Etymology. Versutus (Latin) = crafty, sly. Probably the name versuta refers to the difficulty in identifying this species, given its similarity with H. balzani, according to comments made by Kempf (1973).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFDAFFD0F0A5F2A1FC35FAE8.taxon	discussion	Comments. The study of the type specimens, in addition to specimens from Guatemala, Honduras, Nicaragua, Costa Rica, and Panama, have allowed us to understand that this species has both unbranched and branched setae. The observation of the branched setae is difficult, because in some specimens the lateral branches are small, and also because sometimes both short and long branches collapse to the seta main axis, giving the wrong impression that the setae is unbranched and thick. Hylomyrma versuta is very similar to H. plumosa. There are not great differences in body sculpture. In H. versuta, the propodeum is laterally covered by thinner striae (microsculpture) with indistinguishable interspaces between irregular and thicker striae (macrosculpture), whereas H. plumosa has thinner striae (microsculpture) with indistinguishable interspaces superimposed on irregular and thicker striae (macrosculpture). The conspicuous and trifid setae of H. plumosa are easily observed, but H. versuta has unbranched and branched setae. Hylomyrma versuta and H. plumosa are restricted to Central America, co-occurring near La Virgen and in La Selva Biological Station, Heredia, Costa Rica. Hylomyrma plumosa is only known from these two localities (Fig. 85), but H. versuta has a broader distribution, from southern Mexico to western Colombia (Fig. 88). Molecular analysis using UCE and COI also confirms the proximity of these two species, with H. plumosa sister to the H. cf. dentiloba sp. 2 – H. versuta clade (Pierce et al. 2017). We emphasize that H. cf. dentiloba sp. 2 is understood here as the true H. dentiloba, but the voucher specimens of sp. 2 still need to be examined. Pierce et al. (2017) stated that there are no known morphological differences between H. versuta and H. dentiloba, with both species being differentiated only in their geographical distribution. However, we found that H. versuta can be distinguished from H. dentiloba (the H. cf. dentiloba sp. 2) in the striation with distinguishable interspaces on the head dorsum (vs. striation with indistinguishable interspaces), the mesosoma and petiolar node covered by thinner striae with indistinguishable interspaces between on irregular and thicker striae (vs. thinner striae with indistinguishable interspaces superimposed irregular and thicker striae), the profemur predominantly smooth (vs. with regular and transverse striae weakly marked), the protibia predominantly smooth (vs. mainly covered with regular and weakly marked striae), and the longer striae on tergum of the first gastral segment (vs. shorter striae). Also, Hylomyrma versuta can be distinguished from H. jeronimae (the H. cf. dentiloba sp. 1) in the longitudinal striation on the mesosomal dorsum (vs. striae assuming multiple directions), the striae interspaces on the mesosoma distinguishable (vs. indistinguishable), and the discontinuous dorsal margin of petiole (vs. continuous). Hylomyrma versuta also resembles H. reitteri (characteristic in parentheses), from which it can be distinguished in the irregular striae on the dorsum of head and mesosoma (vs. regular striae), the transverse striae on the ventral surface of petiolar node (vs. smooth surface), and the long striae on tergum of the first gastral segment (vs. short striae, restrict to its basal region). The two species are allopatric; Hylomyrma versuta occurs in Central America and Colombia, and H. reitteri in eastern Brazil and Paraguay. There is morphological variation across the range of H. versuta. Striae on the petiolar dorsum vary from being irregular and longitudinal or vermicular or transversal. The first two conditions are present in the type specimens of H. versuta, and all three can be observed in the material from Chiquiri, Panama, and Costa Rica. Additionally, the striae on the mesosomal dorsum of the specimens from Puntarenas Province, Costa Rica, vary from being more regular to irregular, and mostly longitudinal to forming semi-ellipses. The specimens from Chocó, Colombia, have an anteriorly divergent and posteriorly convergent striation on the most lateral regions of the mesial area of the head dorsum, and a drop-shaped eyes. Notwithstanding these two extremes of variation, and even being the first record to South America, we refer to the specimens from Colombia as H. versuta until we have the opportunity to re-evaluate this material deposited at MCZC.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFDAFFD0F0A5F2A1FC35FAE8.taxon	description	Intercastes are also present in this species, having the combination of the following characters: one inconspicuous central ocellus, and the metanotal groove discernible by a depression. In some specimens, darker areas on the head dorsum coincident with the ocelli location in winged queens are also observed.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFDAFFD0F0A5F2A1FC35FAE8.taxon	distribution	Distribution. Hylomyrma versuta is recorded in southern Mexico, Belize, Costa Rica, Guatemala, Honduras, Nicaragua, Panama, and western Colombia (Fig. 88). Natural history. This species inhabits sites at elevations between 50 and 1520 m. Most specimens were collected in the leaf-litter in wet montane forests, and a few were sampled in dry forest, bamboo, coffee and cardamom plantations. One specimen was collected from the stomach contents of Incilius coniferus (Cope, 1862) in Nicaragua. Wilson observed a H. versuta nest (firstly identified as H. columbica and later as H. versuta by Kempf) in captivity; workers captured flies, springtails, and other selected small invertebrates offered in the feeding chamber, where they fed directly the larvae (Wheeler & Wheeler 1960).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFDAFFD0F0A5F2A1FC35FAE8.taxon	materials_examined	Additional material examined (178 workers, 1 male, 16 queens, 21 intercastes): BRIT. HONDURAS [BELIZE: Cayo]: Caves Branch, 4 – 14. aug. 1972, B- 248, hi-canopy for., S. & J. Peck, berlese (1 Q) [MCZC]; Belmopan, 1 - 15. aug. 1972, B- 243, sift under termite nest, Berlese, S. & J. Peck (1 Q) [MCZC]; Pine Mtn. Ridge Rubber Camp, Macal River, N 1651 ′ 57 ″, W 88 58 ′ 3 ″, 415 m, 21 Feb 1992, leg Gary D. Alpert (1 W, lost specimen) [MCZC]. COLOMBIA: Chocó: 10 km SW S. [San] José del Palmar, Rio Torito, 1 - 4. june. 1978, C. Kugler col., Finca Los Guaduales, 800 m, in Guadua sp. litter (17 W) [MCZC]. COSTA RICA: Alajuela: 27 k N & 8 k W San Ramón, 14. vi. 1997, 950 m, R. Anderson, # 18698 B, 10 ° 13 ′ 30 ″ N, 84 ° 35 ′ 30 ″ W, wet premontane forest, litter, 97 - 014 A, E. B. San Ramón, # 97 - 014 B (1 W) [MZSP]; Guanacaste: Parque Rinco de la Vieja, F. Fernández, oct. 96 (1 W) [IHVL]; Maritza Field Station, 850 m, 13. ii. 1996, R. Anderson, # 17663, Dry and Wet montane, Forest litter (6 W) [MZSP]; same except 875 m, # 17736, Forest transition litter, sample B (9 W 2 I) [MZSP]; # 17736 (2 W) [UFSC, UFGD]; 875 m, # 96 - 009, # 17678, Dry tropical, wet mountain forest trans. litter (1 W) [MZSP]; 17. ii. 96, # 17737, Dry tropical, wet mountain forest trans., sample C (1 W) [MZSP]; # 17737 (2 W) [DZSP]; # 17737 (2 W) [MCZC]; # 17735, sample B (3 W 1 Q 1 I) [MZSP]; # 17734, sample A (1 Q 1 I) [MZSP]; # 96 - 010, 13. ii. 96, 950 m, # 17667, Dry tropical, wet mountain forest trans. litter (5 W 1 Q 1 I) [MZSP]; 13. xi. 1996, # 17668 (4 W, one covered with gold) (MZSP 67461) [MZSP]; # 96 - 019, 17. ii. 96, 875 m, # 17666 (9 W) [MZSP]; Pitilla Field Station, 600 m, 2. v. 1995, R. Anderson, # 17723, Berlese leaf litter, old growth Dry tropical forest, sample C, litter (4 W) [MZSP]; same except # 17720 (2 W) [USNM]; # 17722 (1 W) [MZSP]; Cacao Field Station, 96 - 008, 850 m, 13. ii. 96, R. Anderson, # 17681, Dry tropical wet montane forest trans. litter (5 W) [MZSP]; # 17681 (3 W) [UTLP, IHVL, CASC]; Heredia: P. Viejo, La Selva Biol. Station, 3. june. 1996, M. E. Kaspari col., MEK 45148.00 b, H. dentiloba M. Kaspari comp. type MCZ (2 W) [MCZC]; 11 km ESE La Virgem, 10.35, - 84.05 + 2 km, 300 m, 10. iv. 2004, ALAS # 03 / WF / 02 / all montane wet forest, ex sifted leaf litter (1 Q 2 W) (MZSP 67471, MZSP 67672, MZHY 87) [MZSP]; La Selva Biological Station, 10.41639, - 84.02 + 500 m, 50 m, 16. iii. 2004, TEAM # AMI- 1 - W- 006 - 05, mature wet forest, ex sifted leaf litter (1 Q 2 W) [JTLC]; Puntarenas: Est. Biol. Las Cruces, 4 k S San Vito, 1150 m, 19. vi. 98, R. Anderson, 8 ° 47 ′ 3 ″ N, 82 ° 59 ′ 36 ″ W, Upper wet montane forest, litter extraction, 18662 B (2 W) [MZSP]; same except 18662 A (2 W) [MZSP]; 18662 D (2 W) [MZSP]; 18662 G (2 W) [MZSP]; 18662 H (3 W) [MZSP]; 5 k SW Est. Biol. Las Cruces, 1400 m, 22. vi. 98, R. Anderson, 18665 C, 8 ° 47 ′ 13 ″ N, 82 ° 59 ′ 13 ″ W, Wet cloud forest, litter extraction (2 W) [MZSP]; same except 18665 E (2 W) [MZSP]; 18665 D (1 W) [MZSP]; 18665 I (1 W) [MZSP]; 18665 H (1 W) [MZSP]; 1425 m, 18666 A (2 W) [MZSP]; 18666 B (2 W 1 Q 1 I) [MZSP]; 1100 m, 8 ° 47 ′ N, 82 ° 59 ′ W, Wet cloud for, litter, 18660 (2 W) [MZSP]; 11 k SW Est. Biol. Las Cruces, 9. vii. 1999, 1450 m, R. Anderson, # 19908, 8 ° 46 ′ 43 ″ N, 83 ° 01 ′ 50 ″ W, Wet cloud forest, litter, 99 - 124 A (1 W) [MZSP]; same except 99 - 124 C, # 19910 (1 Q) [MZSP]; Estacion Biol. Las Alturas, 10. vii. 1999, 1520 m, R. Anderson, 8 ° 56 ′ 56 ″ N, 82 ° 50 ′ 01 ″ W, Upper montane / cloud forest transitional, litter, 99.126 B, # 19905 (1 W) [MZSP]; same except 99.126 D, # 19907 (1 W) (MZHY 211) [MZSP]; 2 k NE Alturas, 1520 m, 20. vi. 1998, R. Anderson, 8 ° 56 ′ 56 ″ N, 82 ° 50 ′ 01 ″ W, Upper montane / cloud forest transitional, litter extraction, 18663 Q (1 W) [MZSP]. HONDURAS: Cortés: PN Cusuco, 15.48713, - 88.23472 + 20 m, 1330 m, 30. v. 2010, LLAMA, # Wa-C- 06 - 1 - 11, mesophyll forest, ex sifted leaf litter (1 W 1 Q) [MZSP]. GUATEMALA: Alta Verapaz: 22.5 km SO Panzós, Finca Pablo Juc, Cancoy, 10 - 12. iii. 2013, 15.23036, - 89.71507, 811 m, bosque, winkler, F. Pacay col., 303 - 5 (1 W) [MCZC]; same except 122 - 9 (2 W) [MZSP]; Finca Miguel Putul Tut, Cancoy, 4 - 6. iv. 2013, 15.23252, - 89.71961, 905 m, bosque, winkler, F. Pacay col., 133 - 4 (1 W), 289 - 1 (1 Q) [MZSP]; 24 km SO Panzós, Finca Edgar Caal, San Vicente I, 24 - 26. viii. 2012, 15.2424942, - 89.774883, 1116 m, café, winkler, F. Pacay col., 209 - 4 (1 W) [UTEP]; same except 20 - 22. iii. 2013, 15.24317, - 89.77438, 1135 m, cardamomo, winkler, E. Sierra col., 331 - 8 (1 W) [USNM]; Petén: 13 km NW Machaquilá, 16.44522, - 89.55024 + 50 m, 400 m, 27. v. 2009, LLAMA, # Wa-B- 06 - 1 - 11, tropical forest, ex sifted leaf litter (1 W 1 Q) [MZSP]. MEXICO: Chiapas: 12 mi NW Ocozocoautla, 3200 ft, 4 - 5. ix. 73, A. Newton (1 Q 2 W, one worker covered with gold) (MZSP 67453) [MZSP]; same except forest litter (2 W) [MZSP]; berl. log & leaf litter (2 W) [MIZA]; (1 W 1 Q) [MCZC]; hojarasca, madera podrida, A. Newton (3 W) [MIZA]; Ocozingo, 2. june. 1969, J. M. Campbell col. (2 W) [MCZC]; Vera Cr. [Cruz]: Pueblo Nuevo, nr. Tetzonapa, Aug. 14 - 53, E. O. Wilson, # 221, rain forest (1 W 1 Q 1 M) [MCZC]. NICARAGUA: Masilena Creek: near Bluefields, stomach of Bufo coniferus, W. M. Wheeler (1 W) [MZSP]; RAAN: PN Cerro Saslaya, 13.76810, - 84.98546 + 10 m, 360 m, 7. v. 2011, LLAMA, # Wa-D- 02 - 1 - 50, mature wet forest, ex sifted leaf litter (1 W 1 Q) [MZSP]. PANAMA: Chiriqui: La Fortuna, area Finca la Suisse, 11. vi. 95, R. Anderson, # 17838, Oak ridge, forest litter (5 W 2 I) [MZSP]; same data (1 W 1 I) [DZUP]; (2 W) [IHVL]; (2 W) [UFSC]; (2 W) [CASC]; same except 10. vi. 95, # 17787, sample C (1 W 2 I) [MZSP]; # 17787 (1 W 1 I) [USNM]; # 17787 (1 W 1 I) [MCZC]; # 17788, Oak ridge, bamboo forest litter, sample E (4 W 1 I) [MZSP]; # 17789, sample D (6 W 1 I) [MZSP]; # 17789 (2 W) [UTLP]; # 17790, Raparina oak, forest litter, sample F (5 W 1 I) [MZSP]; # 17839, Oak forest litter, 1200 m (6 W 2 I) [MZSP]; # 17839 (3 W) [UFGD]; 20.4 km North San Felix, 950 m, 08. vi. 95, R. Anderson, # 17768, Wet mountain forest litter, sample B (1 W 3 I) [MZSP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD0FFD2F0A5F15DFCEBFDD8.taxon	description	Figures 76, 86 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD0FFD2F0A5F15DFCEBFDD8.taxon	diagnosis	Worker (Fig. 76 A – C) Diagnosis. Regular and longitudinal striae on head dorsum diverge towards posterior margin, interspaces between striae smooth, striae crest smooth; mesosoma covered with concentric and elliptical, regular and thick striae; longitudinal striae on lateral of pronotum and mesepisternum in part continuing transversely on propodeum and in part continuing on propodeal spine; transverse carina inconspicuous; propodeal spine midsized; dorsal margin of petiole continuous, strongly convex, mesoventral surface unarmed; petiole, postpetiole, subpostpetiolar process, profemur posterior surface, protibia extensor surface and tergum of first gastral segment smooth; subpostpetiolar process weak, convex.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD0FFD2F0A5F15DFCEBFDD8.taxon	description	QUEEN Unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD0FFD2F0A5F15DFCEBFDD8.taxon	etymology	Etymology. This species was named in honor of Dr. Claire Villemant, a French entomologist, curator of the Hymenoptera Collection at Muséum National d’Histoire Naturelle de Paris.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD0FFD2F0A5F15DFCEBFDD8.taxon	discussion	Comments. The diagnostic characters of this species are sufficient to easily distinguish it from most of its congeners. Still, Hylomyrma villemantae is very similar to H. peetersi and H. margaridae. All three species have a strongly convex petiole, with indistinguishable petiolar peduncle and node, and the body mainly covered with regular and longitudinal striae. Even so, H. villemantae can be easily distinguished from both (characteristic in parenthesis) in the striae on the head dorsum with smooth crests (vs. with punctuated crests), the mesosoma with few elliptical and concentric striae (vs. longitudinally striate), the longer propodeal spine (vs. shorter), the unarmed mesoventral surface of petiole (vs. armed), and the smooth dorsum of postpetiole and gaster (vs. with longitudinal striae). All three species are allopatric; H. villemantae is recorded from Brazil (BA) (Fig. 86), whereas H. peetersi and H. margaridae occur in relatively close areas in northern South America (French Guiana, Guyana, and Venezuela) (Fig. 89).	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD0FFD2F0A5F15DFCEBFDD8.taxon	distribution	Distribution. This species occurs in Ibicuí and Itororó, Bahia, Brazil (Fig. 86). Natural history. The biology of this species remains unknown. Type specimens were collected in leaf-litter with winkler extractors, at elevations between 951 and 1070 m in the region known as “ Serra das Piabas ”, a fragment of Atlantic forest in the Brazilian state of Bahia.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD0FFD2F0A5F15DFCEBFDD8.taxon	materials_examined	Additional material examined (1 worker): BRAZIL: BA [Bahia]: Itororó, [área] C, 08.08.00, 14.57.31 S, 40.02.33 W, Santos J. R. M. dos (1 W) (MZHY 199) [MZSP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD2FFD5F0A5F68DFC51FC1A.taxon	description	Figures 77, 78, 80 D, 89 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD2FFD5F0A5F68DFC51FC1A.taxon	materials_examined	Holotype: ECUADOR: Napo: Limoncocha, 250 m, 25. vi. 1976, B- 355, S. & J. Peck [leg.] (1 W) (MCZENT 00524688) [MCZC]. Paratypes: same data as holotype (5 W) (MCZENT 00525482, MCZENT 00524690, MCZENT 00525503, MCZENT 00525523, MCZENT 00524689) [MCZC]; (1 W) (MCZENT 00525522 MZSP 67462) [MZSP]; (1 W) (MCZENT 00525518 MZHY 196) [MZSP]; (1 W) (MCZENT 00525524) [USNM]; (1 W) (MCZENT 00525517) [DZUP]; (1 W) (MCZENT 00525521) [IHVL]; (1 W) (MCZENT 00525519) [CASC]; same except 18. vi. 1976, B- 348 (4 W) (MCZENT 00525501, MCZENT 00524677, MCZENT 00524676, MCZENT 00525499) [MCZC]; (1 W) (MCZENT 00525506 MZSP 67463) [MZSP]; (1 Q) (MCZENT 00524678 MZSP 67464) [MZSP]; 20 km S of Tena, 600 m, 11 Jul 1976, B 360, S. & J. Peck [leg.] (1 W) (MCZENT 00525490) [MCZC]; Pichincha: Centr. Cient. R. Palenque, 20. xii. 1980, Sonia Sandoval col., Bosque primario cerrado, 584 (1 W covered with gold) [IHVL]; Los Rios: C. C. R. Palenque, 79 ° 45 ′ 10 ″ W, 01 ° 25 ′ 56 ″ S, 02 MAR 1979, S. Sandoval (1 Q) (QCAZ I 114012) [QCAZ]; Pastaza: 22 km SW Puyo, 15 July 1976, B- 362, S. & J. Peck [leg.] (1 Q) (MCZENT 00524681) [MCZC].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD2FFD5F0A5F68DFC51FC1A.taxon	diagnosis	Diagnosis. Vermicular to vermiculated-areolated striae on head dorsum and mesosoma; petiole anterior surface well-marked; transverse striae on node ventral surface; longitudinal and anastomosed striae on postpetiole and tergum of first gastral segment; subpostpetiolar process weak, slightly convex; profemur posterior surface mostly smooth; protibia extensor surface entirely striate; long striae on tergum of first gastral segment; setae with 2 short branches of equal size arising from the main axis.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD2FFD5F0A5F68DFC51FC1A.taxon	description	Description. WORKER (n = 3) (Fig. 77 A – C): HL (0.94 – 1.04); HW (0.90 – 1.04); ML (0.66 – 0.70); SL (0.68 – 0.80); MOD (0.26 – 0.27); PNW (0.64 – 0.74); WL (1.24 – 1.40); PSL (0.28 – 0.38); PL (0.59 – 0.64); PW (0.24 – 0.27); PPL (0.36 – 0.40); PPW (0.36 – 0.38); GL (1.02 – 1.12); TL (4.94 – 5.20); CI (95.74 – 100.97); SI (73.07 – 77.67); OI (25 – 28.90). Medium to large-sized. Shiny integument. Head, postpetiole and gaster dark brown, lighter mesosoma, leg and petiole. Thin and branched setae, long to midsized, erect to subdecumbent; 2 short branches of equal size arising from the main axis (Fig. 80 D). Head subquadrate; posterior margin concave at middle. Mandible masticatory margin with 6 teeth. Anterior margin of clypeus concave medially, with a pair of medium teeth laterally; median area of clypeus with 5 irregular and longitudinal striae converging to a point on the anterior margin, interspaces distinguishable. Frontal triangle with 1 – 2 striae. Short scape, not reaching head posterior margin; apical antennomere slightly shorter than previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye drop-shaped, small-sized, larger diameter with 10 ommatidia. Head dorsum with vermicular to vermiculated-areolated striae, divergent towards posterior margin, interspaces smooth. Striae on head lateral and laterodorsal regions converge to eye margin; very thin striae with interspaces indistinguishable (microsculpture) superimposed on vermicular to vermiculatedareolated striae (macrosculpture); gena striate, with the same striation of head lateral, 2 – 3 regular and semicircular striae circumscribe the torulus, not reaching eye margin. Interspaces between striae on head ventral surface distinguishable. Vermicular to vermiculated-areolated striae on mesosoma, interspaces between thicker striae smooth on mesosoma dorsum and filled with thinner striae on mesosoma lateral. Promesonotal junction and metanotal groove indistinct. Transverse carina well-marked. Dorsal margin of mesosoma continuous, slightly convex. Propodeal lobe bidentate, dorsal tooth longer and more acute than the shorter and blunt ventral tooth; dorsal tooth length 1 / 2 of propodeal spine length. Propodeal spine midsized, straight (LV), divergent (DV), sculptured on base. Procoxa with thin and transverse striae of uniform thickness; irregular and transverse striae on C 2 and C 3. Irregular to regular transverse striae on profemur dorsal surface; posterior surface mostly smooth; anterior and ventral surface completely smooth. Protibia extensor surface entirely striate. Dorsal margin of petiole discontinuous, smooth dorsum. Node with irregular and transverse striae on anterior surface continuing on lateral surface; vermiculated-areolated striae on lateral and dorsal surfaces; irregular and transverse striae on ventral surface, interspaces distinguishable. Longitudinal and anastomosed striae on postpetiole; subpostpetiolar process striae restricted to lateral region; subpostpetiolar process weak, slightly convex. First gastral segment striation similar to postpetiole striae; long striae on tergum, 1 / 4 longer than postpetiole length; sternite striation weakly marked, covering the laterobasal region. QUEEN (n = 3) (Fig. 78 A – C): HL (0.98 – 1.12); HW (1.01 – 1.10); ML (0.66 – 0.74); SL (0.76 – 0.90); MOD (0.28 – 0.33); PNW (0.80 – 0.96); WL (1.48 – 1.72); PSL (0.35 – 0.43); PL (0.66 – 0.74); PW (0.26 – 0.30); PPL (0.40 – 0.44); PPW (0.40 – 0.49); GL (1.40 – 1.64); TL (5.58 – 6.40); CI (98.09 – 103.06); SI (75.24 – 81.81); OI (27.20 – 30). Largesized. Color, pilosity, and some sculpture characters shared with conspecific workers, only slightly larger. Larger diameter of eye with 13 – 14 ommatidia. Longitudinal, irregular to vermicular striae on scutum going from an anterior central point towards transcutal suture, interspaces distinguishable. Anepisternum, katepisternum, axilla and scutellum with the same sculpture of scutum. Scutoscutellar sulcus inconspicuous. Transverse and regular striae on propodeum (DV). Lateral of mesosoma with irregular to vermiculated-areolated, thicker striae directed mostly to propodeal dorsum, interspaces between striae smooth. Wings unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD2FFD5F0A5F68DFC51FC1A.taxon	etymology	Etymology. The epithet virginiae is a Latin noun in the genitive case created by adding the singular Latin genitive case suffix - e to the first name of a female person. This species is named in honor of Virginia Leone Bicudo (1910 – 2003), a sociologist and psychoanalyst born in São Paulo, daughter of an Italian immigrant and a black Brazilian. She was the first non-medical practitioner to be recognized as a psychoanalyst, therefore essential for the development and institutionalization of psychoanalysis in Brazil. She was a pioneer in the study of relations between races, which was the subject of her dissertation in 1945.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD2FFD5F0A5F68DFC51FC1A.taxon	discussion	Comments. Hylomyrma virginiae is not known to co-occur with H. mitiae or H. sagax, but all three occur in northwestern South America. Hylomyrma virginiae has been recorded in Ecuador (both sides of The Andes) and in western Colombia (Fig. 89), whereas H. sagax is restricted to southeast Colombia (Fig. 83), and H. mitiae only occurs in French Guiana (Fig. 87). Hylomyrma virginiae is typically smaller (TL 4.94 – 5.20 mm, WL 1.24 – 1.40 mm) than H. mitiae (TL 5.27 – 5.52 mm, WL 1.41 – 1.52 mm). Also, the gena and laterodorsal region of the head are covered by very thin striae with indistinguishable interspaces superimposed on vermicular to vermiculated-areolated striae (vs. very thin striae with indistinguishable interspaces between the vermicular to vermiculated-areolated striae in H. mitiae) (not seen in SEM images of H. mitiae due to coating artifacts), the indistinct metanotal groove (vs. distinguished by slight depression), the discontinuous dorsal margin of petiole (vs. continuous), and the weak and slightly convex subpostpetiolar process (vs. very prominent and subtriangular). Hylomyrma virginiae can be distinguished from H. sagax (characteristic in parentheses) in the medium lateral teeth at clypeus anterior margin (vs. well-developed teeth), the propodeal spine comparatively shorter and thicker (vs. longer and needle-like), and the discontinuous dorsal margin of petiole (vs. continuous). There are relatively few specimens collected of this species; most were sampled in Limoncocha, Ecuador. Specimens from Chocó, Colombia, have a more developed propodeal spine and subpostpetiolar process, and the anterior surface of petiolar node is slightly marked. These characteristics may lead to the misidentification of these specimens as H. sagax, but their body size and sculpture are more similar to those found in H. virginiae specimens recorded in Ecuador.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD2FFD5F0A5F68DFC51FC1A.taxon	distribution	Distribution. Hylomyrma virginiae is known from Colombia and Ecuador (Fig. 89). Natural history. This species occurs in tropical rainforests and areas of bamboo (Guadua sp.) plantations, at elevations between 250 and 850 m. Specimens were collected in the leaf-litter, which suggests that nests are located in fallen logs, rotten wood, between leaves, or inside natural cavities of the superficial soil layers.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD2FFD5F0A5F68DFC51FC1A.taxon	materials_examined	Additional material examined (8 workers): COLOMBIA: Chocó: 10 km SW S. [San] José del Palmar, Rio Torito, 1 – 4. june. 1978, C. Kugler col., Finca Los Guaduales, 800 m, in Guadua sp. litter (6 W) [MCZC]; same except 610 m, by river (1 W) [MCZC]; 850 m, on ridge, litter (1 W) [MCZC].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD5FF2EF0A5F053FD74FE10.taxon	description	Figures 79, 86 (map)	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD5FF2EF0A5F053FD74FE10.taxon	materials_examined	Holotype: PERU: Cusco: Kcosñipata District, Estac. Climatologica Ordinaria Rocotal, 1828 m, 71 ° 34.059 ′ W, 13 ° 6.735 ′ S, 25. ix. 2004, T. R. Schultz, J. Sosa-Calvo, C. J. Marshall, litter sample, nr. river, = JSC 040925 - 04, JSC 040925 - 01 - LS 04 (1 W) (USNMENT 00534691) [USNM]. Paratypes: same data as holotype (1 W) (USNMENT 00534693) [CASC]; same except = JSC 040925 - 05, JSC 040925 - 01 - LS 05 (1 W) (USNMENT 00534702) [USNM]; (1 W) (USNMENT 00534699) [MCZC]; = JSC 040925 - 03, JSC 040925 - 01 - LS 03 (2 W) (USNMENT 00534671, USNMENT 00534669) [USNM]; = JSC 040925 - 16, JSC 040925 - 01 - LS 16 (1 W) (USNMENT 00534760) [USNM]; = JSC 040925 - 14, JSC 040925 - 01 - LS 14 (1 W) (USNMENT 00534750) [USNM]; (1 W) (USNMENT 00534751) [IHVL]; = JSC 040925 - 13, JSC 040925 - 01 - LS 13 (1 W) (USNMENT 00534748) [USNM]; (1 W) (USNMENT 00534742) [DZUP]; = JSC 040925 - 11, JSC 040925 - 01 - LS 11 (1 W 1 I) (USNMENT 00534729 MZSP 67465, USNMENT 00534730 MZSP 67466) [MZSP]; = JSC 040925 - 12, JSC 040925 - 01 - LS 12 (1 W) (USNMENT 00534736 MZHY 206) [MZSP]; = JSC 040925 - 02, JSC 040925 - 01 - LS 02 (2 I, one covered with gold) (USNMENT 00534648 MZSP 67467, USNMENT 00534653 MZSP 67468) [MZSP].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD5FF2EF0A5F053FD74FE10.taxon	diagnosis	Diagnosis. Midsized. Propodeal lobe unidentate, dorsal region rounded, ventral region with a small and blunt tooth; apex of propodeal spine curved upwards.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD5FF2EF0A5F053FD74FE10.taxon	description	Description. WORKER (n = 3) (Fig. 79 A – C): HL (0.96 – 0.98); HW (0.92 – 0.94); ML (0.60 – 0.62); SL (0.70 – 0.72); MOD (0.20 – 0.28); PNW (0.62 – 0.64); WL (1.22 – 1.26); PSL (0.22 – 0.24); PL (0.58 – 0.60); PW (0.24 – 0.26); PPL (0.28 – 0.34); PPW (0.34 – 0.37); GL (1.22 – 1.28); TL (4.95 – 5); CI (93.87 – 97.91); SI (74.46 – 78.26); OI (21.27 – 23.40). Midsized. Shiny integument. Dark brown body, lighter trochanter, tarsi and antenna. Many thin and unbranched setae, long to midsized, erect to subdecumbent. Head subquadrate; posterior margin slightly concave at middle. Mandible masticatory margin with 5 teeth. Anterior margin of clypeus straight medially, with a pair of small teeth laterally; median area of clypeus with 8 – 9 regular, longitudinal and thicker striae (macrosculpture) covered by thinner striae with indistinguishable interspaces (microsculpture). Frontal triangle with 1 – 3 striae. Short scape, not reaching head posterior margin; apical antennomere 1 / 4 shorter than previous 3 antennomeres together. Frontal carina slightly concave posterior to antennal socket. Eye oval, small-sized, larger diameter with 9 ommatidia. Regular and longitudinal striae on head dorsum, slightly divergent towards posterior margin, interspaces between thicker striae filled with thinner striae. Head lateral and laterodorsal regions with the same striation of head dorsum; striae converging to eye margin, anterior part of laterodorsal region with few striae connecting the region posterior to frontal carina with eye margin; gena striate, 6 thicker, regular and semicircular striae circumscribe the torulus, approaching eye margin. Interspaces between striae on head ventral surface distinguishable. Mesosoma covered with irregular and longitudinal striae, interspaces smooth; transverse striae on pronotum anterior region (DV) directed to lateral of pronotum, longitudinal in other regions of pronotum (DV); lateral of pronotum and mesepisternum with longitudinal striae in part continuing transversely on propodeum dorsum and in part continuing towards propodeal spine. Dorsal margin of mesonotum convex. Promesonotal junction discernible by a depression. Metanotal groove indistinct. Transverse carina well-marked. Dorsal margin of mesosoma discontinuous, convex. Propodeal lobe unidentate, dorsal region rounded, ventral region with a small and blunt tooth. Propodeal spine midsized, apex curved upwards (LV), divergent (DV), sculptured on base. Procoxa striae of uniform thickness; regular and transverse striae on C 2 and C 3. Profemur predominantly covered with regular and transverse striae, ventral surface smooth. Protibia extensor surface entirely covered with regular and longitudinal striae. Dorsal margin of petiole discontinuous; ventral surface with few transverse, regular and thin striae, slightly marked. Petiole mesoventral surface armed with subtriangular projection. Convex node; regular and transverse striae on anterior surface continuing inclined and irregular on lateral surface, striae weakly marked; irregular and transverse striae on dorsal surface. Postpetiole covered with regular and longitudinal striae; subpostpetiolar process with the same sculpture of postpetiole on laterobasal region, striae weakly marked, midbasal region smooth; subpostpetiolar process weak, straight. First gastral segment striation similar to postpetiole striae; longitudinal striae on tergum longer than postpetiole length; sternite striation covering the laterobasal region. QUEEN Unknown. MALE Unknown.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD5FF2EF0A5F053FD74FE10.taxon	etymology	Etymology. The epithet wachiperi is a non-Latin noun used in apposition and refers to the local ethnic group. The distribution of this species is restricted to Kcosñipata, Peru and the native ethnic group of this region identify itself as “ wachiperi ”. The species name is also a tribute to the indigenous populations for protecting biodiversity, maintaining the forest, and guarding creole seeds. The specific epithet is a noun in apposition, indeclinable in accordance to articles 31.2.1 and 31.2.3 of the International Code of Zoological Nomenclature.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD5FF2EF0A5F053FD74FE10.taxon	discussion	Comments. This species is unlikely to be confused with any other congener. The combination of the apex of propodeal spine curved upwards, the propodeal lobe unidentate (only the ventral tooth is present), and the dorsal region of propodeal lobe rounded is unique in the genus. Hylomyrma wachiperi is similar to H. reitteri and H. dandarae. All three are allopatric; H. wachiperi occurs in Peru (Fig. 86), H. dandarae occur in Bolivia and Colombia (Fig. 83), and H. reitteri in Brazil and Paraguay (Fig. 85). Hylomyrma wachiperi differs from H. reitteri (characteristic in parentheses) in the oval eye (vs. reniform), the striate ventral surface of petiole (vs. smooth surface), and the tergum of the first gastral segment with long striae (vs. short striae). Hylomyrma wachiperi can be distinguished from H. dandarae in the regular striae on the head dorsum (vs. irregular striae), the irregular striae on the mesosomal dorsum (vs. rugose striae), the indistinct metanotal groove (vs. distinct), the long propodeal spine (vs. short), and the tergum of the first gastral segment with short striae (vs. long striae). Two specimens were identified as intercastes by the presence of three ocelli, a more pronounced depression on the promesonotal junction than that found in workers, modifications on the mesonotum and metanotum, and the larger body measures [HL (1.02 – 1.05); HW (0.98 – 1); ML (0.64); SL (0.72 – 0.74); MOD (0.24); PNW (0.70); WL (1.36); PSL (0.27 – 0.28); PL (0.64 – 0.66); PW (0.25 – 0.27); PPL (0.40); PPW (0.38); GL (1.32 – 1.40); TL (5.38 – 5.51)].	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
4A4D4F09FFD5FF2EF0A5F053FD74FE10.taxon	distribution	Distribution. Hylomyrma wachiperi is only known from Kcosñipata District, Cusco, Peru (Fig. 86). Natural history. This species inhabits sites in tropical rainforest at elevations of 1,825 m. All specimens were collected in leaf-litter, which suggests that nests are located in fallen logs, rotten wood, between leaves, or inside natural cavities of the superficial soil layers. Except for the presence of intercastes, there is no further information available regarding the biology of H. wachiperi.	en	Ulysséa, Mônica Antunes (2021): Taxonomic revision of the Neotropical ant genus Hylomyrma Forel, 1912 (Hymenoptera: Formicidae: Myrmicinae), with the description of fourteen new species. Zootaxa 5055 (1): 1-137, DOI: 10.11646/zootaxa.5055.1.1
