taxonID	type	description	language	source
4A5A879BFF8E6876FDD9FC6BCDFEFA31.taxon	discussion	Composition The study of Verheye et al. (2016 b) indicates that the superfamily Eusiroidea Stebbing, 1888 is a larger taxon than previously admitted. In their appendix S 2, these authors include the following families in Eusiroidea: Acanthonotozomatidae Stebbing, 1906; Acanthonotozomellidae Coleman & J. L. Barnard, 1991; Amathillopsidae Pirlot, 1934; Bateidae Stebbing, 1906; Calliopiidae G. O. Sars, 1893 (excluding Cleippides Boeck, 1871 and Weyprechtia Stuxberg, 1880); Dikwidae Coleman & J. L. Barnard, 1991; Epimeriidae Boeck, 1871; Eusiridae Stebbing, 1888; Iphimediidae Boeck, 1871; Laphystiopsidae Stebbing, 1899; Pleustidae Buchholz, 1874; Pontogeneiidae Stebbing, 1906; Sanchoidae Lowry, 2006; Stilipedidae Holmes, 1908 (including the subfamilies Alexandrellinae Holman & Watling, 1983, Astyrinae Pirlot, 1934 and Stilipedinae Holmes, 1908); Thurstonellidae Lowry & Zeidler, 2008; Vicmusiidae Just, 1990; and the orphan genera Austroregia J. L. Barnard, 1989 and Chosroes Stebbing, 1888, which were formerly included within the Gammarellidae Bousfield, 1977. Remarks The concept of Senticaudata Lowry & Myers, 2013 proposed by Lowry & Myers (2013) and adopted in major databases like the World Register of Marine Species (WoRMS, http: // www. marinespecies. org / accessed on 8 Mar. 2016) conflicts with that of Eusiroidea sensu Verheye et al. (2016 b). A brief discussion on the Senticaudata issue has therefore to be given herein. Lowry & Myers (2013) erected the suborder Senticaudata for amphipods presenting a cluster of spines on the tip of the rami of uropods 1 and 2, postulating that this character state is a synapomorphy. Surprisingly, Lowry & Myers (2013) overlooked some ‘ senticaudate’ taxa, e. g., Idunellinae d’Udekem d’Acoz, 2010, Pleustidae, Apherusa Walker, 1891, Halirages Boeck, 1871 and Paramphithoe Bruzelius, 1859). Most senticaudate families traditionally included in the Eusiroidea (the Calliopiidae and Pontogeneiidae) were transferred by Lowry & Myers (2013) to the superfamily Calliopioidea G. O. Sars, 1895 [sic] (real date of publication: 1893), i. e., the family Calliopiidae elevated to the rank of superfamily. The remaining eusiroid families were excluded from the suborder Senticaudata. Myers & Lowry (2013) also transferred the Cheirocratidae and Hornelliidae to the Calliopioidea, because the two latter families formed a clade with the Calliopiidae and Pontogeneiidae in their morphological phylogenetic tree based on the analysis of 41 characters. The molecular phylogenetic analyses of Verheye et al. (2016 b), which focused on eusiroids, but also included a representative sample of other amphipods, did not support the validity of the suborder Senticaudata. Indeed, senticaudate taxa do not form a clade, suggesting that the distal ornamentation of uropods would be subject to homoplasy, as it is the case for many other characters in amphipods (e. g., J. L. Barnard & Drummond 1978: 7, 193; J. L. Barnard & Karaman 1984: 48; J. L. Barnard & Karaman 1991: 13, 58; Verheye et al. 2016 b). In the 28 S and 18 S rDNA trees, Eusiroidea comprises senticaudate as well as nonsenticaudate taxa, and the families Calliopiidae (excluding Cleippides) and Pontogeneiidae are included in the superfamily, as it was the case in former classifications (e. g., Bousfield 1979, 1982; J. L. Barnard & Karaman 1991). The suborder Senticaudata was erected on the assumption that the senticaudate character state is a synapomorphy. However, the morphological phylogeny of the Senticaudata of Lowry & Myers (2013) does not test this hypothesis as it only includes senticaudate taxa. Moreover, the reliability of the nodes of Lowry & Myers (2013) is not assessed by any support values, e. g., bootstrap support or Bremer’s index. Additionally, it should be noted that the Cheirocratidae and Hornelliidae exhibit several important non-eusiroid character states. For example, they have a well developed accessory flagellum on antenna 1 as well as dorsal spines / setae on the urosome, which is never the case in Eusiroidea. Their position close to the Calliopiidae and Pontogeneiidae on the tree of Lowry & Myers (2013) might reflect poorly supported topologies and / or a subjective choice of characters used in their analysis. A direct observation of specimens indicates that the senticaudate and non-senticaudate character states are not always clear-cut and transitional dispositions are observed among dissimilar amphipods, suggesting multiple passages between the sentidaudate and non-sentidaudate character states. Finally, the polarity of the transition proposed by Lowry & Myers (2013): non senticaudate (plesiomorphic) to senticaudate (apomorphic) is another untested question. In conclusion, before its wide acceptance, the new classification system of amphipods based on the suborder Senticaudata should be evaluated by a combination of molecular and morphological investigations, without a priori assuming that the senticaudate character state is a synapomorphy.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF8F6872FF4AFA37CA60FD60.taxon	description	Family and sub-family key to Antarctic and sub-Antarctic Epimeriidae and their putative relatives This key includes the Epimeriidae s. str. and putatively related families and subfamilies of Eusiroidea present in the Southern Ocean.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF8F6872FF4AFA37CA60FD60.taxon	type_taxon	type genus Paramphithoe Bruzelius, 1859: 68; type species: Acanthosoma hystrix Ross, 1835 designated by J. L. Barnard & Karaman 1991: 398.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF8F6872FF4AFA37CA60FD60.taxon	discussion	Composition Two genera are accepted herein within the Epimeriidae: Epimeria Costa in Hope, 1851 and Uschakoviella Gurjanova, 1955. Actinacanthus Stebbing, 1888 and Paramphithoe Bruzelius, 1859 are removed from the Epimeriidae, and respectively transferred to the Acanthonotozomellidae Coleman & J. L. Barnard, 1991 and the Paramphithoidae G. O. Sars, 1883. Nine subgenera are recognized within Epimeria: Drakepimeria subgen. nov., Epimeria Costa in Hope, 1851; Epimeriella Walker, 1906; Hoplepimeria subgen. nov.; Laevepimeria subgen. nov.; Metepimeria Schellenberg, 1931; Pseudepimeria Chevreux, 1912; Subepimeria Bellan-santini, 1972 and Urepimeria subgen. nov. Systematic remarks and decisions The composition of the family Epimeriidae is in a state of flux. Coleman (2007) listed the following genera within the Epimeriidae: Actinacanthus Stebbing, 1888; Epimeria Costa in Hope, 1851; Epimeriella Walker, 1906; Metepimeria Schellenberg, 1931; Paramphithoe Bruzelius, 1859 and Uschakoviella Gurjanova, 1955. On the other hand, Lörz & Brandt (2004) treated Epimeriella as a junior synonym of Epimeria and questioned the validity of Metepimeria. Metepimeria was initially erected as a monotypic genus for the Magellanic species Epimeria acanthurus, based on a single character state: the absence of dactylus on the palp of the maxilliped. This character state is likely autapomorphic and Metepimeria is relegated herein to the status of subgenus of Epimeria, with a new definition. A recent molecular phylogenetic analysis (Verheye et al. 2016 b) demonstrated that Paramphithoe is unrelated to Epimeria but belongs to the Apherusa / Halirages clade. The senticaudate-type uropods 1 – 2 and the dissimilar palp of the left and right maxilla 1 (see G. O. Sars 1890 – 1895: pl. 130 for Paramphithoe and d’Udekem d’Acoz 2012 for Halirages) are characteristic of this clade, the second character being interpreted as a putative synapomorphy. The profound differences between Paramphithoe and Epimeria were previously pointed out by Gurjanova (1955: 188) and the affinity of Paramphithoe with the Calliopiidae s. lat. was already perceived by Bousfield (1979: 363). Paramphithoe sensu Stebbing, 1906 is therefore herein formally removed from the Epimeriidae and transferred to the Paramphithoidae G. O. Sars, 1883, which are re-established. The study of Verheye et al. (2016 b) showed that Alexandrella Chevreux, 1911, Bathypanoploea Schellenberg, 1939 (Alexandrellinae), Astyra Boeck, 1871 (Astyrinae), Dikwa Griffiths, 1974 (Dikwidae), Acanthonotozomella Schellenberg, 1926, Acanthonotozomoides Schellenberg, 1931 (Acanthonotozomellidae) and Acanthonotozomopsis Watling & Holman, 1980 (Vicmusiidae) are part of a more inclusive clade including Epimeria. All these families are accepted herein but an alternative could indeed be to merge all taxa related to Epimeria into one large family, which we do not advocate, at least for the time being. The genus Actinacanthus, which was traditionally included within the Epimeriidae (e. g., Coleman 2007; De Broyer et al. 2007), exhibits striking morphological similarities with the Acanthonotozomellidae, such as the presence of a pair of huge sword-like teeth on pereionites just above the insertion of coxae, and a non-incised telson. While convergent evolution cannot be ruled out, it seems more likely that these characters are synapomorphies. Actinacanthus is therefore transferred herein to the Acanthonotozomellidae. The genus Uschakoviella, also traditionally assigned to the Epimeriidae, is provisionally retained within this family, in the absence of evidence against its current taxonomic position. Nomenclatural remarks The nomenclatural status of the family Paramphithoidae G. O. Sars, 1883 (previously considered as a junior synonym of the Epimeriidae Boeck, 1871) and of the genus Paramphithoe Bruzelius, 1859 (previously considered as an Epimeriidae) requires discussion. The genus Paramphithoe was created by Bruzelius (1859) for nine unrelated or distantly related species. Stebbing (1895) noted that “ the nine species attributed to that genus by its author have since been distributed among half-a-dozen genera, amid which the original genus, from its vagueness, is an abiding source of confusion. It is a wandering star, of which the orbit is difficult to calculate. ” The family name Paramphithoidae was introduced, albeit without description by G. O. Sars (1883). According to ICZN (1999 arts. 12.1. and 12.2.4), Paramphithoidae G. O. Sars, 1883 is a valid name because it derives from an available generic name: Paramphithoe Bruzelius, 1859, an explicit description being not required for family names introduced before 1931. G. O. Sars (1883) did not explicitly define the composition of the Paramphithoidae and he designated no type species for Paramphithoe. He simply listed four species in it: Pleustes panoplus (Kröyer, 1838) (a species included in Paramphithoe by Bruzelius (1859) when he created that genus), Pleustes parvus (Boeck, 1871) (a probable junior synonym of Pleustes panoplus according to Stebbing (1906 )), Paramphithoe brevicornis G. O. Sars, 1883 and Paramphithoe assimilis G. O. Sars, 1883. All these species are currently considered as members of the family Pleustidae Buchholz, 1874. G. O. Sars (1883) did not include Paramphithoe hystrix (Ross, 1835) (under the name Acanthozone cuspidata (Lepechin, 1780 )) in the Paramphithoidae G. O. Sars, 1883 but put it in the Epimeriidae Boeck, 1871. Ten years later, G. O. Sars (1893) provided a detailed description of the Paramphithoidae, in explicitly restricting it to four genera: Pleustes Spence Bate, 1858, Paramphithoe Bruzelius, 1859, Stenopleustes G. O. Sars, 1893 and Parapleustes Buchholz, 1874, and dealing only with species currently treated as Pleustidae — two of them being part of the original Paramphithoe species of Bruzelius (1859): Paramphithoe pulchella (Kröyer, 1846) and Pleustes panoplus (Kröyer, 1838). Consistently with his previous study, G. O. Sars (1893) included Paramphithoe hystrix (again under the name Acanthozone cuspidata) in the Epimeriidae, this time in explicitely listing Acanthosoma hystrix Ross, 1835 (as Acanthosoma hystrix Owen) in its synonymy. Again, he designated no type species for Paramphithoe. Stebbing (1906) provided a new definition of the Paramphithoidae conflicting with the previous ones, without citing G. O. Sars (1883, 1893). The Paramphithoidae consisted of Actinacanthus, Epimeria and Paramphithoe. Stebbing (1906) designated no type species for Paramphithoe. He simply stated: “ 3 accepted species, 1 doubtful ”. The three accepted species were P. buchholzi Stebbing, 1906, P. hystrix and P. polyacantha (J. Murdoch, 1885) and the “ doubtful ” species P. cuspidata. In other words, only one of the original Paramphithoe species of Bruzelius (1859), namely P. hystrix, was accepted by him in that genus — while the species was clearly excluded from the Paramphithoidae by G. O. Sars (1883, 1893). Stebbing’s (1906) re-definition of Paramphithoe and of the Paramphithoidae was accepted in all subsequent literature, where the family Paramphithoidae was either cited without author’s name (e. g. Barnard 1969) or, more rarely, attributed to Stebbing (1906) (e. g. Bousfield 1979, Coleman & Barnard 1991 a). In other words, Stebbing’s (1906) proposal (not a nomenclatural act) was unanimously accepted by the community of amphipodologists as if it was the formal designation of the type species of the genus Paramphithoe; the statements of G. O. Sars (1883, 1893) were ignored. Finally Barnard & Karaman (1991) designated Acanthosoma hystrix Ross, 1835 as the type species of the genus Paramphithoe — hence also of the family Paramphithoidae. Just like Stebbing (1906), Coleman & J. L. Barnard (1991 a) believed that Paramphithoe sensu J. L. Barnard & Karaman (1991) was a close relative to the genus Epimeria and had to be included in the same family. They re-established the name Epimeriidae Boeck, 1871 for Epimeria, Paramphithoe (and other genera) because it is older than “ Paramphithoidae Stebbing, 1906 ”. It now appears that Paramphithoe is unrelated to Epimeria (Verheye et al. 2016 b) and is nested within the Apherusa / Halirages clade. The family Paramphithoidae has to be re-established for this internally poorly resolved clade including (at least) Apherusa Walker, 1891, Halirages Boeck, 1871 and Paramphithoe revealed by Verheye et al. (2016 b). Additional calliopiid genera (especially from the northern hemisphere) will possibly also have to be transferred to the Paramphithoidae, when more details on their phylogenetic position will be known. Currently, unless ICZN 1999, Article 35.5 on the precedence for names in use at higher rank is invoked, the Calliopiidae can at best be preserved for the genus Calliopius alone, which might be the sister clade of the Paramphithoidae (Verheye et al. 2016 b), and presents no close affinity with other sequenced genera traditionally included in the Calliopiidae. However in databases, we feel that the ‘ Calliopiidae s. lat. ’ should be preserved for all or most genera traditionally included within the Calliopiidae (excluding Apherusa, Halirages and Paramphithoe) as an ‘ adoptive’ artificial family, as long as their phylogenetic and systematic position remains in a too imperfect state of knowledge.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF8F6872FF4AFA37CA60FD60.taxon	description	Key to the genera of Epimeriidae	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF8B686DFDCBFCFACB8AFCAA.taxon	type_taxon	type species: Epimeria tricristata Costa in Hope, 1851: 46 (= Gammarus corniger Fabricius, 1779: 383).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF8B686DFDCBFCFACB8AFCAA.taxon	etymology	Etymology “ Probably so called from the epimera or side-plates ” (Stebbing 1888: 1672).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF8B686DFDCBFCFACB8AFCAA.taxon	description	Description Body smooth or covered with teeth or processes. If processes are present on pereionites just above connection with coxae, these processes are small, not sword-like, not forming large longitudinal carinae. Head high; lateral head lobes very reduced; inferior antennal sinus very shallow; ventral lobe of head well developed, oriented obliquely downwards, sharp or rounded. Development of rostrum variable, usually well developed but sometimes small or even tiny. Urosomite 1 always with dorsal process, which can be rounded or tooth like, projecting upwards or backwards. Telson usually distinctly incised or cleft, rarely weakly emarginate or entire, without spines or setae. Eyes usually present, rarely absent (in abyssal species), medium-sized to large, bulging when present, elliptic, pyriform, rounded or reniform, usually with distinct ommatidia. Antenna 1: peduncular articles short or fairly short, with or without teeth; peduncular article 2 shorter than 1 and longer than 3; accessory flagellum present, small, consisting of one article. Mouthparts projecting quadrately. Upper lip almost entire to notched, symmetrical; epistome not very broad. Mandible: incisor ordinary, toothed; lacinia mobilis present on both mandibles (left and right one different), uniplated; setal row present and well-developed; molar process present, large, triturative or not. Lower lip inner lobes absent, outer lobes relatively broad. Maxilla 1 of basic shape, identical on left and right appendage; palp 2 - articulate, article 2 with distal marginal row of conical spines and distal marginofacial row of strong setae. Maxilla 2, with plates of basic form; inner plate without facial row of setae. Maxilliped: inner and outer plates large and broad; outer plates reaching about mid of article 2 of palp, with short spines on medial border, with long spines on tip and with long stout setae on distal part of lateral border (the short spines intergrades into long spines which themselves intergrade in long stout setae); inner plates nearly reaching tip of article 1 of palp; palp article 4 usually well developed, but sometimes absent, with distal unguis. Coxae long. Coxae 1 – 4 progressively longer. Coxae 1 – 3 narrow. Coxa 4 five-sided: border connecting with coxa, anterodorsal border (running along the posterior margin of coxa 3) perpendicular to body axis or projecting obliquely forward; anteroventral border projecting posteriorly backwards (anterodorsal and anteroventral border either separated by angular discontinuity or smoothly intergrading into a regular curve, rarely produced into a tooth); posteroventral border straight or concave, separated from anteroventral border by angle or tooth (ventral angle or tooth); posterodorsal border concave, separate from posteroventral border by a tooth (posterior tooth); a carina or a groove often run on the surface separating ventral and posterior teeth / angles; when present this carina sometimes bear a tooth projecting laterally. Coxae 5 – 6 with or without tooth or process projecting laterally; when coxa 5 laterally dentate, its tooth can be very large. Gnathopods weak, usually similar but never identical; gnathopod 2 always longer than gnathopod 1 (usually just a bit longer, but sometimes distinctly longer); gnathopods usually subchelate, but sometimes simple; carpus and propodus short to very long, and robust to very narrow; palm well defined or not (sometimes absent), usually posteriorly with about 3 long spines; posterior border of dactylus dentate (sometimes strongly dentate), with terminal unguis. Pereiopods 3 – 7 with terminal unguis on dactylus, easily breaking at the merocarpal articulation. Pereiopod 6> pereiopod 5> pereiopod 7; basis of pereiopods 5 – 7 with longitudinal carina on both side, which is often very protruding; dactylus of pereiopods 5 – 7 usually short, rarely long. Coxal gill from gnathopod 2 to pereiopod 7. Oostegite large and broad, from gnathopod 2 to pereiopod 6. Uropods well developed; rami sword-like. In uropods 1 – 2, inner ramus slightly longer than outer ramus, slightly shorter than peduncle; tip of rami tapering to form a narrow point without spines (i. e., it is not senticaudate); peduncles without ventrolateral spines. In uropod 3, rami subequal much longer than peduncle, which is very short; outer ramus entire. Body length 8 to 80 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF8B686DFDCBFCFACB8AFCAA.taxon	distribution	Distribution and biology The genus Epimeria is almost cosmopolitan in distribution, reaching its highest diversity in Antarctic seas. It also comprises a fairly large number of tropical, deep-water Indo-Pacific species, for which the largest part remains undescribed (pers. obs.: samples from the Muséum national d’Histoire naturelle, Paris). Epimeria has been recorded from the intertidal (E. monodon Stephensen, 1947) down to 5695 m (E. abyssalis Shimomura & Tomikawa 2016 — see Shimomura & Tomikawa (2016 )), but it is predominantly found between 150 and 2500 m. No shallow-water records are known from tropical and warm-temperate seas. The majority of Epimeria species are slow moving, strictly benthic amphipods (Dauby et al. 2001 a) but they are able to swim quickly over short distances (Moore 1981). At least two highly modified species are truly pelagic: E. macronyx (Walker, 1906) and E. pelagica (Birstein & Vinogradov, 1958), and a few species, including E. scabrosa (K. H. Barnard, 1930) and E. atalanta sp. nov., are suspected by us to be semi-pelagic, as they are poorly calcified and exhibit a gracile morphology. Epimeria are usually associated with communities rich in epifauna (De Broyer et al. 2001). The material examined for this study shows that the diversity of Epimeria is directly proportional to the abundance of erect epifauna in trawl catches. The shallow-water species Epimeria monodon lives amidst macro-algae (Richardson 1977). Most species are carnivorous and feed on a fairly large range of organisms, albeit with individual preferences for each species (Dauby et al. 2001 a). One European species, Epimeria parasitica (M. Sars, 1859), is an ectoparasite of holothurians (M. Sars 1859). Epimeria species are themselves frequently consumed by fishes (e. g., Olaso et al. 2000; Dauby et al. 2003). Females largely predominate in the catches. Different hypotheses may be advanced as possible explanation: differences in behavioural pattern between males and females, a smaller size and / or shorter lifespan in males, a biased sex ratio, or even protandrous hermaphroditism. Epimeria seem to be iteroparous, at least in some cases. Indeed, within the maxilliped of the specimen of Epimeria (Drakepimeria) sp. 1 dissected by M. Verheye, which was a mature female with fully developed and setigerous oostegites, the outline of a new cuticle bordered with spines and setae in development could be observed. This indicates that the specimen was preparing an extra moult, after a presumably fertile intermoult. Klages (1991) observed that hatchlings of “ Epimeria georgiana ” (possibly E. xesta), E. robustoides (under the name E. robusta) and E. rubrieques climb on the back of their mother and remain there for a short period of time before leaving her. In contrast, he observed that juvenile “ E. macrodonta ” and “ E. similis ” leave their mother immediately after hatching. Cryptoniscin epicaridean isopods have been reported to parasitize Epimeria robusta (see K. H. Barnard 1930), E. robustoides (see Klages 1991, as E. robusta) and E. rubrieques (see Klages 1991). According to K. H. Barnard (1930), the parasites occupy the marsupium of the female.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF8B686DFDCBFCFACB8AFCAA.taxon	discussion	Remarks The body of many Epimeria species is adorned with protrusions, crests or teeth, which can sometimes be very high. Two main conjectural explanations have been previously proposed to explain the role of the dentiform processes in Epimeria and other amphipods: defence against predation (Moore 1981; Brandt 2000) and hydrodynamic stabilizers during swimming (Moore 1981). However the second explanation seems less applicable for most Epimeria, which are benthic and swim only for short distances (Moore 1981). We herein propose a third explanation, that ornamentation might play a role of camouflage, specifically disruptive camouflage. This idea is based on the analogous observations of the first author for a tuberculate pleustid Pleustes panoplus (Kröyer, 1838) [an eusiroid amphipod species superficially similar to Epimeria] on a rocky shore near Tromsø (Norway). Their protuberances, combined with a variegated colouration, made individuals almost perfectly blend with their habitat of rocks covered in epiflora and epifauna. The protrusions of some Epimeria species might constitute a form of disruptive camouflage on substrates rich in epifauna, which are their preferential habitat. As they live at depths, where the light intensity is very low or absent, a perfect camouflage might not be necessary to avoid detection by predators like fishes. A disruptive body form in a complex tridimensional epifaunal environment could efficiently protect them from predation. The genus Epimeria includes 84 described species (59 in the Southern Ocean). For practical reasons, Antarctic Epimeria species are distributed herein into different subgenera, presenting a homogeneous combination of morphological characters. These subgenera are clades (see Verheye et al. 2016 a) or putative clades (their monophyly was assumed from morphological data when some species could not be sequenced). However, the monophyly of the subgenus Urepimeria remains conjectural, since only one of its three species has been sequenced and the morphology of these species is not fully congruent. The name Vertumnus requires discussion. This generic name was first introduced by Otto (1823) for a trematod: Vertumnus thetidicola Otto, 1823, for which Otto (1823) gave a detailed description. The name Vertumnus was then introduced a second time by White (1847), as a nomen nudum, based on a manuscript of W. E. Leach about crustaceans. Later, White (1850 a) illustrated Leach’s specimens, describing them briefly as ‘ Vertumnus Cranchii ’. These specimens correspond with the description of Epimeria cornigera (Fabricius, 1779). However, White (1850 a) erroneously considered Vertumnus Cranchii as a junior synonym of Oniscus testudo Montagu, 1808 [currently Pereionotus testudo (Montagu, 1808)] and used the combination Acanthonotus testudo for it in his text and on the caption of his plate. Thus, it is clear that Epimeria Costa in Hope, 1851 and Vertumnus White, 1850 are synonyms and have the same type species. However, as indicated above, the name Vertumnus White, 1850 is preoccupied by Vertumnus Otto, 1823. Furthermore, it is unavailable in application of Article 11.6 of the International Code of Zoological Nomenclature (ICZN 1999), which states: “ a name which when first published in an available work was treated as a junior synonym of a name then used as valid is not thereby made available. ” The confusion between Oniscus testudo and Epimeria cornigera was repeated by Gosse (1855), who used again the name Acanthonotus testudo for Epimeria cornigera. A sub-generic classification of non-Antarctic and non sub-Antarctic Epimeria falls out of the scope of the present paper, as these extralimital species form a clade distinct from the Antarctic Epimeria clade (Verheye et al. 2017). We simply propose a provisional definition of the type subgenus Epimeria, which occurs in the Atlantic, Arctic and Indian Oceans, and in the Mediterranean Sea.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF8B686DFDCBFCFACB8AFCAA.taxon	description	Key to the Antarctic and sub-Antarctic subgenera of Epimeria	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF8B686DFDCBFCFACB8AFCAA.taxon	discussion	* Without dissection, Epimeria (Urepimeria) annabellae Coleman, 1994 and E. (U.) monodon Stephensen, 1947 can be readily separated from E. (Laevepimeria) spp. by the strong posterodorsal process of their pleonite 3. The pleonite 3 of E. (U.) extensa Andres, 1985 is much less produced and at first glance, the species looks fairly similar to Laevepimeria, if the mouthparts are not examined. However, E. (U.) extensa can easily be identified by its rostrum, which is about 1.1 × as long as article 1 of peduncle of antenna 1. It is about 0.5 × as long as article 1 in Laevepimeria spp.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF94686AFDCBFCA4CA60F9A7.taxon	description	urn: lsid: zoobank. org: act: 1 B 9 E 9 A 7 F- 2721 - 475 F- 9906 - 75 F 330 B 7 F 4 CC	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF94686AFDCBFCA4CA60F9A7.taxon	etymology	Etymology Combination of the Greek word: δράκων, dragon and Epimeria. The name, which is feminine, alludes to the highly processiform ornamentation in the species of the subgenus, which is reminiscent of mythological dragons.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF94686AFDCBFCA4CA60F9A7.taxon	type_taxon	Type species Epimeria loerzae sp. nov.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF94686AFDCBFCA4CA60F9A7.taxon	description	Description Body opaque, with teguments strongly calcified. Rostrum long. Ventral lobe of head acute. Pleonites 1 – 3 and at least some pereionites with strong mid-dorsal dentate carina and one pair of dorsolateral much smaller teeth, swelling or carina. On pleonite 2 and 3, a second pair of small dorsolateral teeth is sometimes observed between the main dorsolateral teeth and the mid-dorsal tooth (this accessory pair of small teeth is not positioned anteriorly to the main pair). Pereionites 1 – 7 without tooth or protrusion just above connection with coxa. Coxae 1 – 4 often with (but not always) with sharp tip. Coxae 1 – 3 usually sharply keeled along their axis. Coxa 4 with sharp carina starting at ventral tip of coxa and terminating at posterior tip of coxa; this arching carina sometimes bears a tooth projecting laterally; posteroventral border concave. Coxae 5 – 6 with strong sharp tooth projecting backwards. Mid of posterior border of epimeral plates 1 – 3 not produced into a tooth. Posteroventral tooth of epimeral plate 3 strong to very strong. Dorsal process of urosomite 1 produced into a sharp tooth, which can be directed upwards or backwards. Urosomite 2 with or without pair of small teeth pointing upwards. Lateral borders of urosomite 3 posteriorly terminated into a sharp tooth. Telson deeply bilobed. Eyes present, large to very large, not conical. Peduncle of antenna 1 with teeth (dentition sometimes reduced, but never totally absent). Mandible with molar process triturative. Lower lip with narrow (V-shaped) hypopharyngeal gap. Palp of maxilliped with 4 articles. Gnathopods small but not minute, with carpus and propodus of medium slenderness; propodus not strongly expanded distally, subcheliform, with palm obliquely transverse; posterior border of dactylus lined by row of small oblique slender teeth. Basis of pereiopods 5 – 6 moderately slender to broad, with posteroproximal rounded protrusion (sometimes weak) and posterodistal tooth projecting posteriorly. Posterior border of basis of pereiopod 7 proximally slightly convex and distally slightly concave, terminated into a tooth projecting posteriorly. Dactylus of pereiopods 5 – 7 short. Body length The maximum body length recorded in Drakepimeria species ranges between 19 and 47 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF94686AFDCBFCA4CA60F9A7.taxon	biology_ecology	Ecology Benthic, 33 – 2190 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF94686AFDCBFCA4CA60F9A7.taxon	distribution	Distribution Circum-Antarctic, as far north as the South Orkney Islands.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF94686AFDCBFCA4CA60F9A7.taxon	discussion	Remarks Drakepimeria species are superficially similar to those of the subgenus Epimeria. See e. g., illustrations of its type species, Epimeria (Epimeria) cornigera (Fabricius, 1779), given by G. O. Sars (1893). However several differences can be observed. In the subgenus Epimeria, the ventral lobe of the head is rounded instead of being sharp, the peduncle of antenna 1 is not dentate; the basis of pereiopod 5 is extremely narrow; the basis of pereiopods 5 – 6 has neither posteroproximal nor posterodistal process pointing backwards; the posterior margins of the epimeral plates have a median tooth, the lobes of the telson are blunt-tipped instead of being acute and the median notch of the telson is much shallower than in Drakepimeria. Molecular data support the idea that the subgenera Drakepimeria and Epimeria are not closely related (Verheye et al. 2016 b, 2017). Drakepimeria species can be subdivided in two phenotypical categories, which are useful for identification. The species of the first category have a pair of denticles pointing upwards on urosomite 2; they are: E. (D.) anguloce subgen. et sp. nov., E. (D.) corbariae subgen. et sp. nov., E. (D.) havermansiana subgen. et sp. nov., E. (D.) loerzae subgen. et sp. nov., E. (D.) macrodonta Walker, 1906, E. (D.) pandora subgen. et sp. nov., E. (D.) pyrodrakon subgen. et sp. nov., E. (D.) schiaparelli Lörz et al., 2007, and E. (D.) reoproi Lörz & Coleman, 2001. The species of the second category have no pair of denticles pointing upwards on urosomite 2; they are: E. (D.) acanthochelon subgen. et sp. nov., E. (D.) colemani subgen. et sp. nov., E. (D.) cyrano subgen. et sp. nov., E. (D.) leukhoplites subgen. et sp. nov., E. (D.) robertiana subgen. et sp. nov., E. (D.) similis Chevreux, 1912, E. (D.) vaderi Coleman, 1998, and E. (D.) sp. 1.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF936866FE44F9BECA2BFB9D.taxon	description	urn: lsid: zoobank. org: act: B 878492 E- 3294 - 4449 - A 17 D-DBD 502918370 Figs 1 – 7	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF936866FE44F9BECA2BFB9D.taxon	etymology	Etymology From the Greek ακανθα, spine; χελώνη, tortoise. The name, which is a noun in apposition, alludes to the toothed and heavily calcified body of the species, which can be compared to a spiny tortoise.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF936866FE44F9BECA2BFB9D.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: ♀, cruise CEAMARC, sample CEAMARC 1421, stn 62 EV 303, Adélie Coast, 66 ° 10 ʹ 34 ″ S, 143 ° 20 ʹ 45 ″ E, 532 – 550 m, beam trawl, 4 Jan. 2008 (coll. IPEV-AAD-MNHN, removed from MNHN-IU- 2014 - 4342, now MNHN-IU- 2014 - 7321) [extraction M 7; Genbank nr, 28 S: KU 759674]. Paratypes RV Polarstern cruises: SOUTHERN OCEAN: 4 specs, cruise PS 39, ANT-XIII / 3, EASIZ I, stn 1, GSN 1, eastern Weddell Sea, 71 ° 03.10 ʹ S, 11 ° 25.50 ʹ W to 71 ° 02.10 ʹ S, 11 ° 19.30 ʹ W, 462 – 481 m, bottom trawl, 5 Feb. 1996 (previously identified as Epimeria cf. similis and used for gut content studies), coll. C. De Broyer and G. Chapelle (RBINS, INV. 132698); 1 spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 1, GSN 1, eastern Weddell Sea, 71 ° 03.10 ʹ S, 11 ° 25.50 ʹ W to 71 ° 02.10 ʹ S, 11 ° 19.30 ʹ W, 462 – 481 m, bottom trawl, 5 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132699); 1 spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 6, AGT 1, eastern Weddell Sea, 71 ° 31.80 ʹ S, 13 ° 34.50 ʹ W to 71 ° 31.86 ʹ S, 13 ° 35.50 ʹ W, 254 – 261 m, Agassiz trawl, 8 Jan. 1996, coll. C. De Broyer and G. Chapelle, (RBINS, INV. 132697); 1 spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 6, AGT 1, eastern Weddell Sea, 71 ° 31.80 ʹ S, 13 ° 34.50 ʹ W to 71 ° 31.86 ʹ S, 13 ° 35.50 ʹ W, 254 – 261 m, Agassiz trawl, 8 Jan. 1996, coll. C. De Broyer and G. Chapelle, (RBINS, INV. 132706); 1 spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 9, AGT 10, eastern Weddell Sea, 71 ° 34.70 ʹ S, 12 ° 26.60 ʹ W to 71 ° 34.80 ʹ S, 12 ° 25.90 ʹ W, 560 – 571 m, Agassiz trawl, 26 Feb. 1996, coll. C. De Broyer (RBINS, INV. 132696); ca 30 specs of all sizes, cruise PS 39, ANT-XIII / 3, EASIZ I, stn 29, BPN 4, eastern Weddell Sea, 71 ° 31.50 ʹ S, 12 ° 25.50 ʹ W to 71 ° 30.30 ʹ S, 12 ° 27.80 ʹ W, 204 – 529 m, 28 Feb. 1996, coll. C. De Broyer (RBINS, INV. 132705); RV Aurora Australis cruises: SOUTHERN OCEAN: 1 spec., cruise CEAMARC, sample CEAMARC 300, stn 30 EV 66, Adélie Coast, 65 ° 59 ʹ 50 ″ S, 143 ° 39 ʹ 00 ″ E, 421 – 440 m, beam trawl, 24 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4340) [extraction M 5; Genbank nr, 28 S: KU 759672]; 2 specs, cruise CEAMARC, sample CEAMARC 1317, stn 51 AEV 215, Adélie Coast, 66 ° 44 ʹ 52 ″ S, 145 ° 26 ’ 40 ″ E, 525 – 553 m, beam trawl, 30 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4279); 1 spec. (possibly ♂), cruise CEAMARC, sample CEAMARC 1383, stn 50 AEV 220, Adélie Coast, 66 ° 45 ʹ 09 ″ S, 145 ° 20 ʹ 04 ″ E, 567 – 604 m, beam trawl, 30 – 31 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4271); 3 specs, cruise CEAMARC, sample CEAMARC 1384, stn 50 AEV 220, Adélie Coast, 66 ° 45 ʹ 09 ″ S, 145 ° 20 ʹ 04 ″ E, 567 – 604 m, beam trawl, 30 – 31 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4332); 1 spec., cruise CEAMARC, sample CEAMARC 1384, stn 50 AEV 220, Adélie Coast, 66 ° 45 ʹ 09 ″ S, 145 ° 20 ʹ 04 ″ E, 567 – 604 m, beam trawl, 30 – 31 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4333) [extraction M 8; Genbank nr, 28 S: KU 759675]; 2 specs, in one specimen lateral points of coxa 4 broken, cruise CEAMARC, sample CEAMARC 1421, stn 62 EV 303, Adélie Coast, 66 ° 10 ʹ 34 ″ S, 143 ° 20 ʹ 45 ″ E, 532 – 550 m, beam trawl, 4 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4342); 3 specs, 1 adult and 2 immatures, cruise CEAMARC, sample CEAMARC 1643, stn 31 EV 268, Adélie Coast, 66 ° 34 ʹ 30 ″ S, 145 ° 01 ʹ 15 ″ E, 429 – 451 m, beam trawl, 3 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4322) [extraction M 9; Genbank nr, 28 S: KU 759676]; 2 specs, cruise CEAMARC, sample CEAMARC 1951, stn 36 EV 297, Adélie Coast, 66 ° 20 ʹ 20 ″ S, 143 ° 41 ʹ 08 ″ E, 552 – 573 m, beam trawl, 1 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4280); 1 spec., cruise CEAMARC, sample CEAMARC 2724, stn 71 EV 447, Adélie Coast, 66 ° 23 ʹ 60 ″ S, 140 ° 32 ʹ 21 ″ E, 683 – 791 m, beam trawl, 14 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4337). RV L’Astrolabe cruises: SOUTHERN OCEAN: 3 specs, cruise REVOLTA III, stn none (Dumont d’Urville Sea), Collect _ ID: REVO _ 449, Field _ ID: CE- 000002101, Adélie Coast, 66 ° 38 ʹ S, 140 ° 42 ʹ E to 66 ° 38 ʹ S, 140 ° 40 ʹ E, 718 – 729 m, mud, beam trawl, 20 Jan. 2012, coll. G. Lecointre, A. Dettaï, J. Lanshere, C. Gallut and C. Ozouf (MNHN-IU- 2009 - 2539) [extraction K 7 (largest specimen); Genbank nr, 28 S: KU 759659]; 1 spec., cruise REVOLTA III, stn none (Dumont d’Urville Sea), Collect _ ID: REVO _ 449, Field _ ID: CE- 000002239, Adélie Coast, 66 ° 38 ʹ S, 140 ° 42 ʹ E to 66 ° 38 ʹ S, 140 ° 40 ʹ E, 718 – 729 m, mud, beam trawl, 20 Jan. 2012, coll. G. Lecointre, A. Dettaï, J. Lanshere, C. Gallut and C. Ozouf (MNHN-IU- 2009 - 2532) [extraction K 6; Genbank nr, 28 S: KU 759658]. Belgian and Belgian-Dutch Antarctic expedition: SOUTHERN OCEAN: 1 spec., Princess Ragnhild Coast, stn 133, Baie Léopold, 70 ° 19 ʹ 09 ″ S, 24 ° 13 ʹ 05 ″ E to 70 ° 19 ʹ 08 ″ S, 24 ° 12 ʹ 06 ″ E, 240 m, trawl, year 1961, coll. A. Capart (RBINS, INV. 132748); 1 spec., Princess Ragnhild Coast, stn 139, coordinates unavailable, year 1961, coll. A. Capart (RBINS, INV. 132749); 1 spec. (possibly ♂), Princess Ragnhild Coast, stn 150, Baie Léopold, 70 ° 20 ʹ 04 ″ S, 24 ° 13 ʹ E, 255 m, dredge, 20 Jan. 1961, coll. A. Capart (RBINS, INV. 132746); 1 spec., Princess Ragnhild Coast, stn 219, Baie du Glacier, 70 ° 18 ʹ 05 ″ S, 23 ° 58 ʹ 00 ″ E, 216 m, trawl, 31 Jan. 1965, coll. A. Capart (RBINS, INV. 132745); 1 spec., Princess Ragnhild Coast, stn 220, Baie du Glacier, exact coordinates unavailable [presumably about 70 ° 18 ʹ S, 23 ° 58 ʹ E], 414 – 450 m, 1 Feb. 1965, trawl, coll. A. Capart (RBINS, INV. 132734).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF936866FE44F9BECA2BFB9D.taxon	description	Description ROSTRUM. Long, reaching about tip of article 1 of peduncle of antenna 1 (teeth excluded), weakly curved, sharp-tipped in lateral view. EYE. Very large, elliptic. PEREION – PLEOSOME TOOTH PATTERN. Mid-dorsal tooth on all segments or from pereionite 2 onwards; pair of non-carinate posterodorsal teeth or swellings on all segments; pereionite 1 often with small but distinct mid-dorsal tooth or scarcely distinct bump, with pair of very low non-carinate dorsolateral swellings; pereionite 2 nearly as broad as pereionite 1, with mid-sized blunt-tipped mid-dorsal tooth and pair of dorsolateral swellings or very blunt teeth; pereionite 3 with well-developed blunt-tipped regularly curved broad, mid-dorsal tooth and pair of dorsolateral blunt teeth; pereionite 3 to pleonite 2 with acute-tipped broad and regularly curved mid-dorsal tooth (nearly same shape and size on all these segments) and pair of small non-carinate acute-tipped dorsolateral teeth (pleonites 1 – 2 without second pair of dorsolateral teeth); pleonite 3 with dorsal sharp carina with distinct median notch (lobe anterior to notch distinctly curved) and obliquely produced posteriorly into an a broad acute and sharp triangular tooth, and pair of dorsolateral non-carinate acute-tipped teeth. COXAE 1 – 3. Strongly carinate and distally sharp. COXA 4. Anterodorsal and anteroventral border nearly straight, joined by low and blunt angular discontinuity, anterior angle not strongly projecting forward; ventral tooth very long and styliform; lateral carina with lateral long and sharp triangular tooth pointing obliquely backwards, carina very distant from margin of coxa at its deepest point. COXA 5. With long, sharp and narrowly triangular, carinate, lateral tooth pointing obliquely backwards. COXA 6. With mid-sized, sharp and narrowly triangular, carinate, lateral tooth pointing obliquely backwards; posteroventral corner produced into a triangular tooth. COXA 7. With ventral and posterior border straight, with posteroventral angle produced into a strong tooth. EPIMERAL PLATES 1 – 3. Posteroventral angle produced into a very long styliform tooth. UROSOME TOOTH PATTERN. Urosomite 1 with sharp triangular process pointing upwards; urosomite 2 without pair of small posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.3; tips of lobes sharp, slit narrow. PEDUNCLE OF ANTENNA 1. Article 1 with long lateral and medial teeth reaching mid of article 2 (teeth excluded) and long ventral tooth reaching tip of article 2 (teeth excluded); article 2 with long lateral tooth nearly reaching tip of article 3, with medial tooth slightly overreaching article 3 (ventral tooth excluded), with ventral tooth slightly overreaching tip of article 3 (tooth excluded); article 3 with very long ventral tooth, longer than article itself. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; propodus not narrowing distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus very slender; basis of pereiopods 5 – 6 of normal width, with posteroproximal process rounded and strongly protruding, with posterodistal tooth strong; basis of pereiopod 7 very broad with posterodistal tooth sharp, followed more proximally by small concavity, directed posteriorly. Colour pattern Body and appendages white with a pinkish hue on posterior dorsal crests, peduncles of antennae and the distal half of pereiopods; eyes red (photograph of a specimen from Adélie Coast, MNHN-IU- 2009 - 2539). The specimen from the eastern shelf of the Weddell Sea illustrated by Rauschert & Arntz (2015) under the name of Epimeria similis exhibits a diffuse orange pattern on a whitish background. Body length Up to 45 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF936866FE44F9BECA2BFB9D.taxon	distribution	Distribution Eastern shelf of the Weddell Sea, Princess Ragnhild Coast, Adélie Coast; depth range 204 – 791 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF936866FE44F9BECA2BFB9D.taxon	biology_ecology	Biology Dauby et al. (2001 a) observed the following items in the gut content of Epimeria from the eastern Weddell Sea identified by them as E. similis: cnidocysts of various size and shape, some of them identified as from hydrozoans of the genera Tubularia and Campanula, and others to actiniids. Other items observed were: planktonic cells (diatoms and foraminifers), spicules of sponges and setae of polychaetes. Dauby et al. (2001 b) concluded that it is a micropredatory browser. These specimens were presumably Epimeria acanthochelon sp. nov., as that species was collected in large numbers during the cruise EASIZ I, and as the material studied by these authors was largely based on material collected during that cruise. Epimeria similis s. str. is not present in the eastern Weddell Sea.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF936866FE44F9BECA2BFB9D.taxon	discussion	Remarks Epimeria acanthochelon sp. nov. (type locality: Adélie Coast) is morphologically extremely similar to E. similis from the tip of the Antarctic Peninsula and South Shetland Islands. However, results of molecular analyses based on 28 S rDNA suggest that they would be distinct species (Verheye et al. 2016 a; this paper Fig. 342). Specimens from the eastern shelf of the Weddell Sea are also identified as E. acanthochelon sp. nov., albeit on basis of their morphology only. A genetic characterization of specimens from that area would be desirable whenever possible. Epimeria acanthochelon sp. nov. often has a small tooth or a trace of mid-dorsal tooth on pereionite 1, which is absent in E. similis. The dorsal teeth of other segments have a slightly different form, with those of the second and third segments being more developed in E. acanthochelon sp. nov. than in E. similis. The carina of pleonite 3 also has a stronger anterior lobe followed by a deeper median notch in E. acanthochelon sp. nov. than in E. similis. Coxa 7 often has a stronger posteroventral tooth in E. acanthochelon sp. nov. than in E. similis. Finally, the telson notch is also slightly deeper in E. acanthochelon sp. nov. than in E. similis. The specimen illustrated by a photograph in Coleman (2007: plate 1 fig. e), Lörz & Coleman (2009: unnumbered photograph on p. 17) and Rauschert & Arntz (2015: 62, plate 55) and identified by them as Epimeria similis is probably E. acanthochelon sp. nov. On page 125, Rauschert & Arntz (2015) indicated that it was collected on the eastern shelf of the Weddell Sea and in an early draft of their book made available to the authors, it is specified that it was collected during the cruise ANT-XIII / 3, which indeed sampled on the eastern shelf of the Weddell Sea (Arntz & Gutt 1997). Its morphology matches perfectly with the description of E. acanthochelon sp. nov.: the posteroventral tooth of its coxa 7 is stronger and the anterior lobe of the dorsal crest of pleonite 3 is more developed than in E. similis.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF9F6863FE22FBD3CCFDF95A.taxon	description	urn: lsid: zoobank. org: act: 9 F 72 B 38 E- 2322 - 43 C 3 - A 48 A-DCD 9823693 A 0 Figs 8 – 16	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF9F6863FE22FBD3CCFDF95A.taxon	etymology	Etymology Angulócë, dragon — Tolkien’s Quenya language (Faulskanger 2008). The name, which is a noun in apposition, alludes to the dragon-like facies of the species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF9F6863FE22FBD3CCFDF95A.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: 1 ♀, cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 228 - 3, Larsen A, 64 ° 54.96 ’ S, 60 ° 31.97 ʹ W to 64 ° 54.43 ʹ S, 60 ° 30.44 ʹ W, 277 – 309 m, bottom trawl, 27 Feb. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 132961) [extraction I 6; Genbank nr, COI: KU 870850, 28 S: KU 759627]. Paratypes RV Polarstern cruises: SOUTHERN OCEAN: 8 specs, cruise PS 69, ANT-XXIII / 8, stn 702 - 9, southeast of Larsen B, 65 ° 57.85 ʹ S, 60 ° 28.42 ʹ W to 65 ° 57.42 ʹ S, 60 ° 28.12 ʹ W, 215 – 221 m, Agassiz trawl, 12 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122565); 1 immature spec., cruise PS 69, ANT-XXIII / 8, stn 702 - 9, southeast of Larsen B, 65 ° 57.85 ʹ S, 60 ° 28.42 ʹ W to 65 ° 57.42 ʹ S, 60 ° 28.12 ʹ W, 215 – 221 m, Agassiz trawl, 12 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122563); 1 small juv., cruise PS 69, ANT-XXIII / 8, stn 702 - 9, southeast of Larsen B, 65 ° 57.85 ʹ S, 60 ° 28.42 ʹ W to 65 ° 57.42 ʹ S, 60 ° 28.12 ʹ W, 215 – 221 m, Agassiz trawl, 12 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122562); 4 specs, cruise PS 69, ANT-XXIII / 8, stn 725 - 10, south of Larsen A, 64 ° 55.89 ʹ S, 60 ° 40.06 ’ W to 64 ° 55.92 ʹ S, 60 ° 40.31 ʹ W, 189 – 192 m, Rauschert dredge, 22 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122561); 2 specs, cruise PS 69, ANT-XXIII / 8, stn 725 - 10, south of Larsen A, 64 ° 55.89 ʹ S, 60 ° 40.06 ’ W to 64 ° 55.92 ʹ S, 60 ° 40.31 ʹ W, 189 – 192 m, Rauschert dredge, 22 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (MNHN-IU- 2014 - 7322, removed from RBINS, INV. 122561); 7 specs, cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 248 - 2, Larsen B, 65 ° 57.51 ʹ S, 60 ° 28.15 ʹ W to 65 ° 57.69 ʹ S, 60 ° 28.30 ʹ W, 196 – 202 m, Agassiz trawl, 7 Mar. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 122893); 1 juv., cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 248 - 2, Larsen B, 65 ° 57.51 ʹ S, 60 ° 28.15 ʹ W to 65 ° 57.69 ʹ S, 60 ° 28.30 ʹ W, 196 – 202 m, Agassiz trawl, 7 Mar. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 122895); 2 specs, cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 248 - 2, Larsen B, 65 ° 57.51 ʹ S, 60 ° 28.15 ʹ W to 65 ° 57.69 ʹ S, 60 ° 28.30 ʹ W, 196 – 202 m, Agassiz trawl, 7 Mar. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 122900); 1 juv., posterior half missing, cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 248 - 2, Larsen B, 65 ° 57.51 ʹ S, 60 ° 28.15 ʹ W to 65 ° 57.69 ʹ S, 60 ° 28.30 ʹ W, 196 – 202 m, Agassiz trawl, 7 Mar. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 122902); 1 juv., cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 248 - 2, Larsen B, 65 ° 57.51 ʹ S, 60 ° 28.15 ʹ W to 65 ° 57.69 ʹ S, 60 ° 28.30 ʹ W, 196 – 202 m, Agassiz trawl, 7 Mar. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 132975) [extraction I 7; Genbank nr COI: KU 870851, 28 S: KU 759628]; 1 small spec., cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 248 - 2, Larsen B, 65 ° 57.51 ʹ S, 60 ° 28.15 ʹ W to 65 ° 57.69 ʹ S, 60 ° 28.30 ʹ W, 196 – 202 m, Agassiz trawl, 7 Mar. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 132655) [extraction A 20; Genbank nr COI: KU 870817, 28 S: KU 759589]; 1 spec., cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 257 - 2, Larsen A, 64 ° 54.75 ʹ S, 60 ° 39.01 ʹ W to 64 ° 54.62 ʹ S, 60 ° 39.50 ʹ W, 159 – 169 m, Agassiz trawl, 13 Mar. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 122904); 1 spec., cruise PS 81, ANT-XXIX / 3, stn 185 - 3, south east of Dundee Island, 63 ° 51.34 ʹ S, 55 ° 41.11 ʹ W, to 63 ° 51.52 ʹ S, 55 ° 41.43 ʹ W, 238 – 244 m, non muddy bottom with a lot of life (sponges, starfishes, ophiuroids, crinoids, Pentapora - like bryozoans), Agassiz trawl, 19 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122941) [extraction ANT 34; Genbank nr, COI: KU 870821, 28 S: KU 759593]; 1 ♀, cruise PS 81, ANT-XXIX / 3, stn 193 - 8, Bransfield Strait, 62 ° 43.73 ʹ S, 57 ° 29.04 ʹ W to 62 ° 43.80 ʹ S, 57 ° 29.40 ʹ W, 428 – 431 m, Agassiz trawl, 23 Feb. 2013, absolute alcohol, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 132962). Other material examined RV Polarstern cruises: 1 spec., cruise PS 10, ANT-V / 3, stn 527, eastern Weddell Sea, 72 ° 23.5 ʹ S, 16 ° 37.4 ʹ W, depth uncertain (it is indicated 582 – 631 m on the specimen label, but there are no depth record in the cruise report: Schnack-Schiel 1987), Agassiz trawl, 22 Oct. 1986 (RBINS, INV. 132725); 1 spec., cruise PS 39, ANT-XXIII / 3, EASIZ I, eastern Weddell Sea, no station, specimen kept in aquarium, 18 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132995); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 603 - 5, eastern Weddell Sea, 70 ° 30.99 ʹ S, 08 ° 48.08 ʹ W to 70 ° 30.40 ʹ S, 08 ° 48.13 ʹ W, 274 – 297 m, sponge bottom, 7 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 132978). RV Marion Dufresne cruises: SOUTHERN OCEAN: 2 specs, cruise MD 42 (SIBEX), stn 22 - CP 66, Prydz Bay, 66 ° 55 ʹ 45 ″ S, 74 ° 04 ʹ 11 ″ E, depth missing [Google Earth coordinates for that position: 427 m (accessed 27 Sep. 2016)], 26 Jan. 1985 (MNHN-IU- 2014 - 4264). Belgian and Belgian-Dutch expeditions: SOUTHERN OCEAN: 1 ovigerous ♀, Princess Ragnhild Coast, stn 220, Baie du Glacier, exact coordinates unavailable [presumably about 70 ° 18 ʹ S, 23 ° 58 ʹ E], 414 – 450 m, trawl, 1 Feb 1965, coll. A. Capart (RBINS, INV. 132728).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF9F6863FE22FBD3CCFDF95A.taxon	description	Description ROSTRUM. Long, reaching mid of article 2 of peduncle of antenna 1 (teeth excluded), very strongly curved, sharp-tipped in lateral view. EYE. Large, broadly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 with small broad and blunt mid-dorsal tooth pointing backwards, with pair of low and blunt, conical, dorsolateral teeth; pereionite 2 much narrower than pereionites 1 and 3, without mid-dorsal tooth and without pair of dorsolateral teeth; pereionite 3 with medium-sized broad and blunt mid-dorsal tooth pointing backwards and pair of low and blunt, conical, dorsolateral teeth; pereionite 4 to pleonite 2 with large, narrow, acute mid-dorsal tooth, of which the anterior border is regularly curved and the posterior border is slightly concave (except for pleonite 2 where the posterior border is straight), with pair of conical dorsolateral teeth, of which the size gradually increases posteriorly (these pairs of teeth are never duplicate); pleonite 3 with large nearly styliform acute-tipped symmetrical mid-dorsal tooth, and pair of large narrowly conical dorsolateral teeth (size of mid-dorsal teeth very weakly increasing from pereionite 4 to pleonite 2; mid-dorsal tooth of pleonite 3 almost as long as tooth of pleonite 2). COXAE 1 – 3. Strongly carinate and distally sharp. COXA 4. Anterodorsal border straight, anteroventral border distally straight, these two borders being joined by a long curve (anterior angle), this anterior angle is weakly projecting forward; ventral tooth long and acute; lateral carina with small tooth obliquely pointing backwards (in dorsal view this tooth form a broad U-shaped concavity with the coxa); carina very distant from margin of coxa at its deepest point. COXA 5. With long, sharp, narrowly triangular, carinate, lateral tooth pointing backwards (its anterior margin is parallel to body axis or weakly divergent and its posterior margin is oblique). COXA 6. With mid-sized, sharp, triangular, carinate, lateral tooth pointing obliquely backwards and with anterior border very weakly convex); posteroventral corner rounded, without angular discontinuity. COXA 7. With ventral border slightly curved, with posterior border straight, their convergence forming a blunt squared angle. EPIMERAL PLATES 1 – 3. Posteroventral angle produced into a very long and very sharp tooth. UROSOME TOOTH PATTERN. Urosomite 1 with large and sharp narrow tooth pointing upwards; urosomite 2 with pair of mid-sized posterior dorsolateral teeth pointing upwards; urosomite 3 with pair of mid-sized posterior dorsolateral teeth pointing obliquely upwards. TELSON. Cleft on 0.35; tips of lobes subacute, notch broadly and bluntly V-shaped. PEDUNCLE OF ANTENNA 1. Article 1 with long lateral and medial teeth reaching mid of article 2 (teeth excluded) and long ventral tooth slightly overreaching tip of article 2 (teeth excluded); article 2 with large lateral tooth of which 0.3 is overreaching tip of article 3, and huge medial tooth of which 0.6 is overreaching tip of article 3, without ventral tooth; article 3 with medium-sized ventral tooth, about 0.3 times as long as article itself. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; very slightly broadening distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus slender; basis of pereiopods 5 – 6 of normal width, with posteroproximal process rounded and strongly protruding, with posterodistal tooth very strong (as long as basis width); basis of pereiopod 7 broad with posterodistal tooth acute and very large, followed more proximally by distinct concavity, directed posteriorly. Colour pattern Whitish to yellowish with orange plain markings (pigmented parts not arranged in dots); rostrum, antennae orange; distal half of walking pereiopods orange or striped with orange; mouthparts and tip of gnathopods red; eyes red. Body length Up to 37 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF9F6863FE22FBD3CCFDF95A.taxon	distribution	Distribution Bransfield Strait, Dundee Island, Larsen A and B area, eastern shelf of the Weddell Sea, Princess Ragnhild Coast, Prydz Bay; 189 – 431 m and possibly 631 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF9F6863FE22FBD3CCFDF95A.taxon	discussion	Remarks Epimeria anguloce sp. nov. is morphologically similar to E. corbariae. See key to the subgenus and account for the latter species for differences. Two E. anguloce sp. nov. from the eastern shelf of the Weddell Sea were examined. These specimens were not adequately fixed for genetic studies and were not in very good condition. However they showed traces of an orange mottled colour pattern similar to that of specimens from the Antarctic Peninsula region. The mid-dorsal tooth of their pereionite 1 and 3 is a bit stronger and the posterodistal tooth of the basis of their pereiopod 7 a bit shorter than in Peninsular specimens. The same form was found off Princess Ragnhild Coast and Prydz Bay. The collection locality of the ‘ Epimeria macrodonta ’ illustrated by Rauschert & Arntz (2015: 61 plate 54) was given in a preliminary draft of the book made available to the present authors. The specimen was collected during cruise ANT-XV / 3, at stn 355: north west of King George Island, 61 ° 60 ʹ S, 59 ° 15 ʹ W, 128 – 130 m. On the published picture, the mid-dorsal tooth of pereionite 3 appears as almost as long as the tooth of pereionite 4, i. e., much longer than in any known species of the macrodonta complex, and acute-tipped. Coleman (2007) published a photograph, which was absolutely identical (plate 1 fig. d), except for the mid-dorsal tooth of pereionite 3, which is less than half as long as that of pereionite 4, and blunt-tipped (i. e., as in all known species of the macrodonta complex). It seems therefore that the photograph has been erroneously altered in the book of Rauschert & Arntz (2015). One of the authors presumably believed that the tooth was broken and tried to ‘ repair’ it.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF9A685FFE22F913CD8AFAAA.taxon	description	urn: lsid: zoobank. org: act: 74 A 8958 B- 6 C 00 - 4 D 92 - 89 B 6 - AA 6977 F 51 D 3 A Figs 17 – 26	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF9A685FFE22F913CD8AFAAA.taxon	etymology	Etymology The species is dedicated to Charles Oliver Coleman (Museum für Naturkunde, Berlin), as a recognition of his important contribution to the knowledge of the taxonomy of the genus Epimeria. The name is a genitive.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF9A685FFE22F913CD8AFAAA.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: ovigerous ♀, cruise PS 81, ANT-XXIX / 3, Bransfield Strait, stn 196 - 8, 62 ° 47.80 ʹ S, 57 ° 5.35 ʹ W to 62 ° 47.63 ʹ S, 57 ° 5.63 ʹ W, 542 – 580 m, Agassiz trawl, 24 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122942) [extraction ANT 37; Genbank nr, COI: KU 870823, 28 S: KU 759596]. Paratypes RV Polarstern cruises: SOUTHERN OCEAN: 2 specs, cruise PS 06, ANT-III / 3, stn 348, eastern Weddell Sea, 72 ° 50 ʹ S, 19 ° 23 ʹ W, 490 m, bottom trawl, 20 Feb. 1985, coll. J. Plötz (RBINS, INV. 132740); 1 spec., cruise PS 06, ANT-III / 3, station unknown, eastern Weddell Sea, Feb. 1985, coll. J. Plötz (RBINS, INV. 132742); 1 small spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 1, GSN 1, eastern Weddell Sea, 71 ° 03.10 ʹ S, 11 ° 25.50 ʹ W to 71 ° 02.10 ʹ S, 11 ° 19.30 ʹ W, 462 – 481 m, bottom trawl, 5 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132703); 1 small spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 1, GSN 1, eastern Weddell Sea, 71 ° 03.10 ʹ S, 11 ° 25.50 ʹ W to 71 ° 02.10 ʹ S, 11 ° 19.30 ʹ W, 462 – 481 m, bottom trawl, 5 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132709); 1 spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 12, GSN 5, eastern Weddell Sea, 73 ° 18.10 ʹ S, 21 ° 10.10 ʹ W to 73 ° 17.10 ʹ S, 21 ° 08.20 ʹ W, 457 – 459 m, bottom trawl, 14 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132695); 1 spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 17, GSN 10, eastern Weddell Sea, 73 ° 18.00 ʹ S, 21 ° 09.90 ʹ W to 73 ° 19.10 ʹ S, 21 ° 14.90 ʹ W, 465 – 468 m, bottom trawl, 16 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132693); 4 specs, cruise PS 69, ANT-XXIII / 8, stn 661 - 2, between Elephant Island and King George Island, 61 ° 39.29 ʹ S, 57 ° 02.89 ʹ W to 61 ° 39.20 ʹ S, 57 ° 04.75 ʹ W, 466 – 467 m, bottom trawl, 30 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122549); 3 alcohol-fixed specs, cruise PS 69, ANT-XXIII / 8, stn 662 - 1, between Elephant Island and King George Island, 61 ° 35.91 ʹ S, 57 ° 17.04 ʹ W to 61 ° 35.41 ʹ S, 57 ° 20.60 ʹ W, 425 – 432 m, bottom trawl, 30 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122492); 2 specs, initially fixed with formalin, cruise PS 69, ANT-XXIII / 8, stn 663 - 1, north of King George Island, 61 ° 38.18 ʹ S, 57 ° 33.17 ʹ W to 61 ° 38.02 ʹ S, 57 ° 37.16 ʹ W, 432 – 434 m, bottom trawl, 30 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122537); 1 immature spec., cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, eastern Weddell Sea, stn 48 - 1, 70 ° 23.94 ʹ S, 8 ° 19.14 ʹ W to 70 ° 23.89 ʹ S, 8 ° 18.67 ʹ W, 595 – 602 m, bryozoan bottom (exceptional diversity of bryozoans), Agassiz trawl, 12 Jan. 2008, coll. H. Robert (RBINS, INV. 122901); 1 spec., dissected and illustrated by unpublished line drawings, cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, eastern Weddell Sea, stn 48 - 1, 70 ° 23.94 ʹ S, 8 ° 19.14 ʹ W to 70 ° 23.89 ʹ S, 8 ° 18.67 ʹ W, 595 – 602 m, bryozoan bottom (exceptional diversity of bryozoans), Agassiz trawl, 12 Jan. 2008, coll. H. Robert (RBINS, INV. 132977) [extraction P 41; Genbank nr, COI: KU 870895, 28 S: KU 759684]; 8 specs, fixed in formalin, cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, eastern Weddell Sea, stn 48 - 1, 70 ° 23.94 ʹ S, 8 ° 19.14 ʹ W to 70 ° 23.89 ʹ S, 8 ° 18.67 ʹ W, 595 – 602 m, bryozoan bottom (exceptional diversity of bryozoans), Agassiz trawl, 12 Jan. 2008, coll. H. Robert (RBINS, INV. 132455); 2 specs, fixed in alcohol, cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, eastern Weddell Sea, stn 48 - 1, 70 ° 23.94 ʹ S, 8 ° 19.14 ʹ W to 70 ° 23.89 ʹ S, 8 ° 18.67 ʹ W, 595 – 602 m, bryozoan bottom (exceptional diversity of bryozoans), Agassiz trawl, 12 Jan. 2008, coll. H. Robert (RBINS, INV. 132986); 1 immature spec., cruise PS 77, ANT-XXVII / 3, CAMBIO, eastern Weddell Sea, stn 265 - 2, 70 ° 47.34 ʹ S, 10 ° 40.39 ʹ W to 70 ° 47.13 ʹ S, 10 ° 40.54 ʹ W, 500 – 600 m, Agassiz trawl, 22 Mar. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 132965) [extraction I 9; Genbank nr, COI: KU 870853, 28 S: KU 759630]; 1 ♀, cruise PS 81, ANT-XXIX / 3, Bransfield Strait, stn 193 - 8, 62 ° 43.73 ʹ S, 57 ° 29.04 ʹ W to 62 ° 43.80 ʹ S, 57 ° 29.40 ʹ W, 428 – 431 m, Agassiz trawl, 23 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 132976) [extraction I 13; Genbank nr, COI: KU 870839, 28 S: KU 759615]; 1 ovigerous ♀, cruise PS 81, ANT-XXIX / 3, Bransfield Strait, stn 193 - 9, 62 ° 43.50 ʹ S, 57 ° 27.92 ʹ W to 62 ° 43.53 ʹ S, 57 ° 28.28 ʹ W, 420 – 428 m, sponge bottom, Agassiz trawl, 23 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 132966) [extraction ANT 36; Genbank nr, COI: KU 870822, 28 S: KU 759595]; 1 large spec., cruise PS 81, ANT-XXIX / 3, Bransfield Strait, stn 193 - 9, 62 ° 43.50 ʹ S, 57 ° 27.92 ʹ W to 62 ° 43.53 ʹ S, 57 ° 28.28 ʹ W, 420 – 428 m, sponge bottom, Agassiz trawl, 23 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122962); 3 large specs, cruise PS 81, ANT-XXIX / 3, Bransfield Strait, stn 196 - 8, 62 ° 47.80 ʹ S, 57 ° 5.35 ʹ W to 62 ° 47.63 ʹ S, 57 ° 5.63 ʹW, 542 – 580 m, Agassiz trawl, 24 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122959); 3 specs, cruise PS 81, ANT-XXIX / 3, Bransfield Strait, stn 217 - 6, 62 ° 53.45 ʹ S, 58 ° 13.06 ʹ W to 62 ° 53.42 ʹ S, 58 ° 13.41 ʹ W, 461 – 483 m, rich sponge bottom, Agassiz trawl, 2 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122922 A) [extraction K 39, largest specimen; Genbank nr, COI: KU 870871, 28 S: KU 759651]. Other material examined RV Aurora Australis cruises: SOUTHERN OCEAN: 1 spec., cruise CEAMARC, sample CEAMARC 1317, stn 51 AEV 215, Adélie Coast, 66 ° 44 ʹ 52 ″ S, 145 ° 26 ʹ 40 ″ E, 525 – 553 m, beam trawl, 30 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 7323, removed from MNHN-IU- 2014 - 4279); 1 spec., cruise CEAMARC, sample CEAMARC 2724, stn 71 EV 447, Adélie Coast, 66 ° 24 ʹ 00 ″ S, 140 ° 32 ʹ 21 ″ E, 683 – 791 m, beam trawl, 14 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 7324, removed from MNHN-IU- 2014 - 4337) [extraction M 6; Genbank nr, COI: KU 870889, 28 S: KU 759673]; 3 specs, cruise CEAMARC, sample CEAMARC 3532, stn 87 EV 524, Adélie Coast, 65 ° 29 ʹ 29 ″ S, 139 ° 18 ʹ 37 ″ E, 397 – 411 m, beam trawl, 17 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4353); 5 specs, cruise CEAMARC, sample CEAMARC 3978, stn 36 EV 297, 66 ° 20 ʹ 20 ″ S, 143 ° 41 ʹ 08 ″ E, Adélie Coast, 552 – 573 m, 4 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4329). RV Marion Dufresne cruises: SOUTHERN OCEAN: 1 spec., cruise MD 42 (SIBEX), stn 22 - CP 73, Prydz Bay, 66 ° 57 ʹ 25 ″ S, 72 ° 41 ʹ 25 ″ E, depth missing [Google Earth depth for that position: 540 m (accessed 27 Sep. 2016)] 26 Jan. 1985 (MNHN-IU- 2014 - 4259) (typed label); 1 spec., cruise MD 42 (SIBEX), stn 22 - CP 66, Prydz Bay, 66 ° 55 ʹ 45 ″ S, 74 ° 4 ʹ 11 ″ E, depth missing [Google Earth depth for that position: 427 m (accessed 27 Sep. 2016)], 26 Jan. 1985 (MNHN-IU- 2014 - 4266) (typed label). Belgian and Belgian-Dutch Antarctic expedition: SOUTHERN OCEAN: 2 specs, Princess Ragnhild Coast, stn 220, Baie du Glacier, exact coordinates unavailable [presumably about 70 ° 18 ʹ S, 23 ° 58 ʹ E], 414 – 450 m, trawl, 1 Feb. 1965, coll. A. Capart (RBINS, INV. 132737); 2 specs, Princess Ragnhild Coast, stn 220, Baie du Glacier, exact coordinates unavailable [presumably about 70 ° 18 ʹ S, 23 ° 58 ʹ E], 414 – 450 m, trawl, 1 Feb. 1965, coll. A. Capart (RBINS, INV. 132738); 5 specs, Princess Ragnhild Coast, stn 220, Baie du Glacier, exact coordinates unavailable [presumably about 70 ° 18 ʹ S, 23 ° 58 ʹ E], 414 – 450 m, trawl, 1 Feb. 1965, coll. A. Capart (RBINS, INV. 132739). RV Discovery cruises (National Antarctic Expedition 1901 – 1904): SOUTHERN OCEAN: 1 adult ♀, Ross Sea, n ° 13, 914 m (500 fathoms), 22 Jan. 1902, together in the same tube with a juvenile Epimeria, which is identified herein as E. macrodonta s. str., obviously representing paralectotypes of Epimeria macrodonta designated by Lörz et al. (2007), albeit no label indicated their status of paralectotypes (BMNH 1907.6.6.259 - 262) (in part).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF9A685FFE22F913CD8AFAAA.taxon	description	Description ROSTRUM. Long, overreaching tip of article 1 of peduncle of antenna 1 (teeth excluded), weakly curved, sharp-tipped in lateral view. EYE. Very large, elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 3 to pleonite 3 with mid-dorsal tooth; pereionite 1 to pleonite 3 with pair of posterodorsal teeth or protrusions (those of the pereion are distinctly carinate); pereionites 1 – 2 without any trace of mid-dorsal tooth, with pair of carinate dorsolateral protrusions; pereionite 2 slightly narrower than pereionite 1; pereionite 3 with small blunt-tipped posterior middorsal tooth and pair of small dorsolateral carinate teeth; pereionite 4 to 6 with medium-sized blunttipped broad and regularly curved mid-dorsal tooth and pair of small dorsolateral carinate teeth; pereionite 7 with large acute-tipped and regularly curved mid-dorsal tooth and pair of small dorsolateral carinate teeth; pleonites 1 – 2 with large acute-tipped and regularly curved mid-dorsal tooth and pair of small dorsolateral non-carinate teeth; on pleonite 1 a second pair of (much smaller) dorsolateral teeth is observed between the mid-dorsal tooth and the main pair of dorsolateral teeth; pleonite 3 with large acute-tipped mid-dorsal tooth bearing a weak median notch and pair of small dorsolateral non-carinate teeth (teeth from pereionite 7 to pleonite 3 distinctly longer than more anterior teeth, the size difference being more important in immatures than in adults) COXAE 1 – 3. Strongly carinate and distally sharp. COXA 4. Anterodorsal and anteroventral border nearly straight joined by low and blunt angular discontinuity, anterior angle slightly but distinctly projecting forward; ventral tooth long and acute; lateral carina with obtuse angularity or very obtuse tooth, carina very distant from margin of coxa at its deepest point. COXA 5. With long, sharp, triangular, carinate, lateral tooth pointing obliquely backwards. COXA 6. With mid-sized, sharp, triangular, carinate, lateral tooth pointing obliquely backwards; posteroventral corner rounded, with trace of angular discontinuity. COXA 7. With ventral border slightly curved, with posterior border straight, their convergence forming a sharp squared angle. EPIMERAL PLATES 1 – 3. Posteroventral angle produced into a long and very sharp tooth. UROSOME TOOTH PATTERN. Urosomite 1 with sharp triangular process pointing upwards; urosomite 2 without pair of small posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.2; tips of lobes rounded, slit fairly narrow. PEDUNCLE OF ANTENNA 1. Article 1 with long lateral and medial teeth nearly reaching mid of article 2 (teeth excluded) and long ventral tooth reaching tip of article 2 (teeth excluded); article 2 with huge lateral tooth and very large medial tooth considerably overreaching tip of article 3, with ventral tooth slightly overreaching tip of article 3 (tooth excluded); article 3 with well developed ventral tooth, about as long as article itself. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; propodus not narrowing distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus slender; basis of pereiopods 5 – 6 of normal width, with posteroproximal process rounded and strongly protruding in pereiopod 5, weakly protruding in pereiopod 6, with posterodistal tooth strong; basis of pereiopod 7 very broad with posterodistal tooth sharp, followed more proximally by small concavity, directed posteriorly. Colour pattern Whitish, with vague reddish marks on the mid-dorsal tooth of the posterior body segments (pereionite 7 to urosomite 1), tip of rostrum, tip of pereiopods. Eyes reddish. Body length Up to 47 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF9A685FFE22F913CD8AFAAA.taxon	distribution	Distribution Between King George Island and Elephant Island, Bransfield Strait, eastern shelf of the Weddell Sea, Princess Ragnhild Coast, Prydz Bay, Adélie Coast, western Ross Sea, 397 – 914 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF9A685FFE22F913CD8AFAAA.taxon	discussion	Remarks In immature E. colemani sp. nov., the mid-dorsal tooth of pereionite 7 and pleonites 1 – 2 are much longer than in adults and much longer than more anterior mid-dorsal teeth. Epimeria colemani sp. nov. specimens from western stations (South Shetland Islands, Antarctic Peninsula, Eastern Weddell Sea) and the only sequenced specimen from the Adélie Coast (MNHN-IU- 2014 - 7324, extraction M 6) have identical COI haplotypes, whereas the divergence between their 28 S sequences is in the range expected between distinct species (from comparison with other species in this complex) (fig. 1 in Verheye et al. 2016 a as Epimeria SI 5). Additional data would be needed to explain this discrepancy. As morphologically, the specimens from the Adélie Coast cannot be distinguished from the remaining E. colemani, they are provisionally considered as conspecifics.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA6685CFDE5FAA3CB66F893.taxon	description	urn: lsid: zoobank. org: act: 74519 DD 0 - 1 C 75 - 4392 - AA 0 D- 5 FB 811 B 108 F 4 Figs 27 – 43	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA6685CFDE5FAA3CB66F893.taxon	etymology	Etymology The species is dedicated to Laure Corbari (Muséum national d’Histoire naturelle, Paris), who gave us the opportunity to study the very interesting collections of the CEAMARC and REVOLTA expeditions. The name is a genitive.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA6685CFDE5FAA3CB66F893.taxon	materials_examined	Type material Holotype REVOLTA cruises: (RV Seatruck) SOUTHERN OCEAN: 1 ♀, cruise REVOLTA I, stn REVO- 007 b, Collect _ ID: 249, Field _ ID: CE- 000004589, Adélie Coast, 66 ° 38 ʹ 25 ″ S, 139 ° 49 ʹ 43 ″ E, 127 – 133 m, 22 Jan. 2010, coll. M. Eléaume, L. Hemery and A. D’Hont, (MNHN-IU- 2009 - 2570) [extraction K 4; Genbank nr, COI: KU 870872, 28 S: KU 759652]. Paratypes RV Aurora Australis cruises: SOUTHERN OCEAN: 3 specs, cruise CEAMARC, sample CEAMARC 933, stn 40 EV 152, Adélie Coast, 66 ° 39 ʹ 38 ″ S, 143 ° 01 ʹ 16 ″ E, 471 – 637 m, beam trawl, 28 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4283); 1 ♀, cruise CEAMARC, sample CEAMARC 1643, stn 31 EV 268, Adélie Coast, 66 ° 34 ʹ 30 ″ S, 145 ° 01 ʹ 15 ″ E, 429 – 451 m, beam trawl, 3 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 7326, removed from MNHN-IU- 2014 - 4322); 5 specs, cruise CEAMARC, sample CEAMARC 2072, stn 65 EV 322, Adélie Coast, 65 ° 48 ʹ 29 ″ S, 143 ° 03 ʹ 46 ″ E, 750 – 788 m, beam trawl, 5 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4286); 1 ♀, cruise CEAMARC, sample CEAMARC 2724, stn 71 EV 447, Adélie Coast, 66 ° 24 ʹ 00 ″ S, 140 ° 32 ʹ 21 ″ E, 683 – 791 m, v 14 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 7327, removed from MNHN-IU- 2014 - 4337); 6 specs, cruise CEAMARC, sample CEAMARC 3978, stn 36 EV 297, Adélie Coast, 66 ° 20 ʹ 20 ″ S, 143 ° 41 ʹ 08 ″ E, 552 – 573 m, beam trawl, 4 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4330); 1 spec., cruise REVOLTA II, stn REVO _ 133, Collect _ ID: REVO _ 133, Adélie Coast, 66 ° 37 ʹ 04 ″ S, 140 ° 00 ʹ 13 ″ E, 103 – 107 m, 11 Jan. 2011, coll. N. Améziane, N. Bax, C. Gallut, A. C. Lautrédou and C. Robineau (MNHN-IU- 2014 - 4297); 1 ovigerous ♀, cruise REVOLTA II, stn REVO _ 091, Collect _ ID: REVO _ 162, Field _ ID: CE- 000001166, Adélie Coast, 66 ° 41 ʹ 07 ″ S, 139 ° 56 ʹ 41 ″ E, 33 – 34 m, 19 Jan. 2011, coll. N. Améziane, N. Bax, C. Gallut, A. C. Lautrédou and C. Robineau (MNHN-IU- 2014 - 7325, removed from MNHN- 2014 - 4299) [extraction M 10; Genbank nr, COI: KU 870878, 28 S: KU 759661]; 1 spec., cruise REVOLTA II, stn REVO _ 091, Collect _ ID: REVO _ 162, Field _ ID: CE- 000001166, 66 ° 41 ʹ 07 ″ S, 139 ° 56 ʹ 41 ″ E, 33 – 34 m, 19 Jan. 2011, coll. N. Améziane, (MNHN-IU- 2014 - 4299); 1 spec., cruise REVOLTA II, stn REVO _ 091, Collect _ ID: REVO _ 162, Adélie Coast, 66 ° 41 ʹ 07 ″ S, 139 ° 56 ʹ 41 ″ E, 33 – 34 m, 19 Jan. 2011, coll. N. Améziane, N. Bax, C. Gallut, A. C. Lautrédou and C. Robineau (MNHN-IU- 2014 - 4298); 1 juv., cruise REVOLTA II, stn REVO _ 085, Collect _ ID: REVO _ 191, Field _ ID: CE- 000001559, 66 ° 40 ʹ 12 ″ S, 139 ° 55 ʹ 56 ″ E, Adélie Coast, 37 – 44 m, 29 Jan. 2011, coll. N. Améziane, N. Bax, C. Gallut, A. C. Lautrédou and C. Robineau, (MNHN-IU- 2014 - 4296) [extraction M 11; Genbank nr, 28 S: KU 759662]; 1 spec., cruise REVOLTA II, stn REVO _ 037, Collect _ ID: REVO _ 209, Adélie Coast, 66 ° 39 ʹ 13 ″ S, 139 ° 52 ʹ 04 ″ E, 105 – 107 m, 1 Feb. 2011, coll. N. Améziane, N. Bax, C. Gallut, A. C. Lautrédou and C. Robineau (MNHN-IU- 2014 - 4305); 2 specs, cruise REVOLTA II, stn REVO _ 037, Collect _ ID: REVO _ 209, Adélie Coast, 66 ° 39 ʹ 13 ″ S, 139 ° 52 ʹ 04 ″ E, 105 – 107 m, 1 Feb. 2011, coll. N. Améziane, N. Bax, C. Gallut, A. C. Lautrédou and C. Robineau (RBINS 132717, formerly MNHN-IU- 2014 - 4305); 1 spec., cruise REVOLTA III, stn REVO _ 039, Collect _ ID: REVO _ 464, Adélie Coast, 66 ° 38.370 ʹ S, 139 ° 55.863 ʹ E to 66 ° 38.406 ʹ S, 139 ° 56.030 ʹ E, 100 m, beam trawl, 30 Jan. 2012, coll. G. Lecointre, A. Dettaï, J. Lanshere, C. Gallut and C. Ozouf (MNHN number-IU- 2009 - 2542); 1 juv., cruise REVOLTA III, stn REVO _ 040, Collect _ ID: REVO _ 470, Adélie Coast, 66 ° 38.50 ʹ S, 139 ° 57.02 ʹ E to 66 ° 38.41 ʹ S, 139 ° 57.14 ʹ W, 98 – 100 m, beam trawl, 31 Jan. 2012, coll. G. Lecointre, A. Dettaï, J. Lanshere, C. Gallut and C. Ozouf (MNHN-IU- 2009 - 2551); 1 small spec., cruise REVOLTA III, stn REVO _ 064, Collect _ ID: REVO _ 481, Adélie Coast, 66 ° 36.371 ʹ S, 140 ° 05.075 ʹ E to 66 ° 36.394 ʹ S, 140 ° 04.966 ʹ E, 110 – 120 m, beam trawl, 3 Jan. 2012, coll. G. Lecointre, A. Dettaï, J. Lanshere, C. Gallut and C. Ozouf (MNHN-IU- 2009 - 2562); 1 adult spec., cruise REVOLTA III, stn REVO _ 068, Collect _ ID: REVO _ 493, Field _ ID: CE- 000002621, Adélie Coast, 66 ° 35 ʹ 18 ″ S, 140 ° 03 ʹ 15 ″ E to 66 ° 35 ʹ 22 ″ S, 140003 ʹ 23 ″ E, 57 – 118 m, beam trawl, 7 Feb. 2012, coll. G. Lecointre, A. Dettaï, J. Lanshere, C. Gallut and C. Ozouf (MNHN-IU- 2009 - 2581). RV L’Astrolabe cruises: 5 specs, cruise REVOLTA III, stn none (Dumont d’Urville sea), Collect _ ID: REVO _ 449, Field _ ID: CE- 000002109 Adélie Coast, 66 ° 38 ʹ S, 140 ° 42 ʹ E to 66 ° 38 ʹ S, 140 ° 40 ʹ E, 718 – 729 m, mud, beam trawl, 20 Jan. 2012, coll. G. Lecointre, A. Dettaï, J. Lanshere, C. Gallut and C. Ozouf (MNHN-IU- 2009 - 2543); 1 ♀, cruise REVOLTA III, stn REVO _ 064, Collect _ ID: REVO _ 481, Field _ ID: CE- 000003196, Adélie Coast, 66 ° 36.371 ʹ S, 140 ° 05.075 ʹ E to 66 ° 36.394 ʹ S, 140 ° 04.966 ʹ E, 110 – 120 m, beam trawl, 3 Feb. 2012, coll. G. Lecointre, A. Dettaï, J. Lanshere, C. Gallut and C. Ozouf (MNHN-IU- 2009 - 2563) [extraction K 5; Genbank nr, COI: KU 870876, 28 S: KU 759657].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA6685CFDE5FAA3CB66F893.taxon	description	Description ROSTRUM. Long, reaching base of article 2 of peduncle of antenna 1 (teeth excluded), very strongly curved, sharp-tipped in lateral view. EYE. Large, broadly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 usually without mid-dorsal tooth or with a posterior bump, rarely with small subacute tooth pointing upwards (tooth with anterior border oblique and posterior border vertical), without pair of dorsolateral protrusions when pereionite has no mid-dorsal tooth or only a posterior bump, with pair of low protrusions when mid-dorsal tooth present; pereionite 2 much narrower than pereionites 1 and 3, without mid-dorsal tooth and without pair of dorsolateral teeth; pereionite 3 with medium-sized, broad, subacute to acute tooth pointing upwards or slightly backwards and pair of low and blunt, conical, dorsolateral teeth; pereionite 4 to pleonite 2 with large, narrow to fairly narrow, acute mid-dorsal tooth, of which the anterior border is regularly curved or curved with very faint angular discontinuity and the posterior border is slightly concave, with pair of conical dorsolateral teeth of which the size gradually increases posteriorly (these pairs of teeth are never duplicate); pleonite 3 with large nearly styliform acute-tipped symmetrical mid-dorsal tooth, and pair of large narrowly conical dorsolateral teeth (size of mid-dorsal teeth weakly increasing from pereionite 4 to pleonite 2; mid-dorsal tooth of pleonite 3 slightly but distinctly shorter than tooth of pleonite 2). COXAE 1 – 3. Strongly carinate and distally sharp. COXA 4. Anterodorsal border straight, anteroventral border distally slightly curved, these two borders being joined by a fairly short blunt angular discontinuity (anterior angle), this anterior angle is weakly projecting forward; ventral tooth long and acute; lateral carina with small tooth pointing backwards (its anterior border is parallel to body axis or nearly so; in dorsal view this tooth form a narrow V-shaped notch with the coxa); carina fairly distant from margin of coxa at its deepest point. COXA 5. With long, sharp, narrowly triangular, carinate, lateral tooth pointing backwards (its anterior margin is distinctly oblique to nearly parallel to body axis and its posterior margin is oblique. COXA 6. With mid-sized, sharp and triangular, carinate, lateral tooth pointing obliquely backwards; its anterior border is distinctly convex); posteroventral corner broadly rounded. COXA 7. With ventral border slightly curved, with posterior border straight or very weakly concave, their convergence forming a sharp squared angle. EPIMERAL PLATES 1 – 3. Posteroventral angle produced into a very long and very sharp tooth. UROSOME TOOTH PATTERN. Urosomite 1 with large and sharp narrow tooth pointing upwards; urosomite 2 with pair of mid-sized posterior dorsolateral teeth pointing upwards; urosomite 3 with pair of mid-sized posterior dorsolateral teeth pointing obliquely upwards. TELSON. Cleft on 0.45 – 0.55; tips of lobes subacute, notch narrowly U-shaped. PEDUNCLE OF ANTENNA 1. Article 1 with long lateral and medial teeth reaching mid of article 2 (teeth excluded) and long ventral tooth distinctly overreaching tip of article 2 (teeth excluded); article 2 with large lateral tooth of which 0.3 is overreaching tip of article 3, and huge medial teeth of which 0.6 is overreaching tip of article 3, without ventral tooth; article 3 with medium-sized ventral tooth, about 0.3 times as long as article itself. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; very slightly broadening distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus slender; basis of pereiopods 5 – 6 of normal width, with posteroproximal process rounded and strongly protruding, with posterodistal tooth very strong (as long as basis width); basis of pereiopod 7 broad with posterodistal tooth acute and large, followed more proximally by distinct concavity, directed posteriorly. Colour pattern Homogeneously bright red, or red marks on a whitish or pale reddish background. Body length Up to 31 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA6685CFDE5FAA3CB66F893.taxon	distribution	Distribution Adélie Coast, 33 – 827 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA6685CFDE5FAA3CB66F893.taxon	discussion	Remarks Epimeria corbariae sp. nov. exhibits variation in the dentition of the first body segment and the robustness of the mid-dorsal teeth (see figs 27, 28, 34, 38). The angle of divergence of the lateral tooth of coxa 5 with body axis is also variable. Due to these variations, the most extreme forms of E. corbariae sp. nov. were initially interpreted as two separate species. However, genetic data (COI, 28 S) suggest that they are indeed conspecific (Fig. 342). Epimeria corbariae sp. nov. (Adélie Coast) is morphologically similar to E. anguloce sp. nov. (Weddell Sea and Bransfield Strait), but molecular analyses based on the 28 S gene suggest that they are distinct species (Fig. 342). The following morphological differences were observed. In E. corbariae sp. nov., pereionite 1 is usually smooth or with a posterior bump (rarely with a small tooth); in E. anguloce sp. nov. a small tooth is always present. In E. corbariae sp. nov. the mid-dorsal tooth of pereionite 3 to pleonite 2 are a bit shorter and a bit broader than in E. anguloce sp. nov. and those of pereionite 3 to pleonite 2 have a trace of anterior angular discontinuity, which is not present in E. anguloce sp. nov. The tooth of the lateral carina of coxa 4 is more posteriorly directed in E. corbariae sp. nov. and forms a narrower notch with the coxa (when examined in dorsal view) than in E. anguloce sp. nov. The junction between the ventral and the posterior border of coxa 7 forms a sharp squared angle in E. corbariae sp. nov. vs a blunt squared angle in E. anguloce sp. nov., this difference being very clear. In E. corbariae sp. nov., the posterodistal tooth of basis of pereiopod 7 is shorter than in E. anguloce sp. nov. Epimeria corbariae sp. nov. is also very similar to E. schiaparelli, but the middorsal ornamentation of pereionite 3 is much more developed in the first species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA5685AFE14F8ABCD94FB45.taxon	description	urn: lsid: zoobank. org: act: 7 E 682282 - 1 C 82 - 4 A 2 B-BEEC- 54 B 05 FEEA 37 C Figs 44 – 50	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA5685AFE14F8ABCD94FB45.taxon	etymology	Etymology Cyrano de Bergerac is the central character of the eponymous play by Edmond Rostand. In the play, Cyrano is described as endowed with a prominent nose. The name, which is a noun in apposition, alludes to the long, straight and anteriorly directed rostrum of the species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA5685AFE14F8ABCD94FB45.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: 1 ♀, cruise PS 77, ANT XXVII / 3, CAMBIO, stn 263 - 6, eastern Weddell Sea, BENDEX area, 70 ° 38.66 ʹ S, 10 ° 28.16 ʹ W to 70 ° 39.50 ʹ S, 10 ° 32.91 ʹ W, depth not given [Google Earth depth range for the coordinates: 867 – 955 m (accessed 27 Sep. 2016)], gear: “ BPT ” (presumably benthopelagic trawl), 21 Mar. 2011, coll. Ch. Havermans and H. Robert (RBINS, INV. 132967) [extraction I 16; Genbank nr, COI: KU 870842, 28 S: KU 759618].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA5685AFE14F8ABCD94FB45.taxon	description	Description ROSTRUM. Very long, reaching about tip of article 2 of peduncle of antenna 1, straight and anteriorly directed, sharp-tipped in lateral view. EYE. Medium-sized, elliptic. PEREION – PLEOSOME TOOTH PATTERN. All pereionites and pleosomites with large styliform mid-dorsal tooth and pair of large styliform dorsolateral teeth; the mid-dorsal tooth of pereionite 1 is oriented forward, the others are directed posteriorly, the most posterior ones being the most backwards directed; the size of the mid-dorsal teeth slightly increase backwards; pereionite 2 nearly as broad as pereionite 1. COXAE 1 – 3. Strongly carinate, curving forward and laterally, distally very sharp. COXA 4. Anterodorsal border weakly concave, and anteroventral border straight, the two border are joined by obtuse but very distinct angular discontinuity; anterior angle slightly projecting forward; ventral tooth medium-sized, long and styliform, curving laterally; lateral carina with lateral long and sharp styliform tooth pointing laterally (scarcely oriented backwards). COXA 5. With sharp, broadly styliform, carinate, lateral tooth pointing laterally or rather slightly obliquely backwards. COXA 6. With large, sharp and narrowly triangular, carinate, lateral tooth pointing obliquely backwards; posteroventral corner broadly rounded. COXA 7. With ventral border rounded, and posterior border nearly straight, with posteroventral angle bluntly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle produced into a very long styliform tooth. UROSOME TOOTH PATTERN. Urosomite 1 with very long styliform tooth pointing upwards (or very slightly backwards); urosomite 2 without pair of small posterior dorsolateral teeth pointing upwards. TELSON. Cleft on half of its length; tips of lobes blunt-tipped, slit of medium width, U-shaped. PEDUNCLE OF ANTENNA 1. Article 1 – 2 with trace of medial tooth or teeth; articles 1 – 3 with tiny ventral tooth, otherwise without ornamentation. GNATHOPODS 1 – 2. Gnathopods 1 and 2 a bit different, not narrowing distally: gnathopod 1 with propodus and especially carpus rather stout, with palm oblique and large; gnathopod 2 of normal slenderness, with palm oblique and large. PEREIOPODS 5 – 7. Merus, carpus and propodus of pereiopods 5 – 7 very slender, dactylus fairly long; basis of pereiopods 5 – 6 narrow, with posteroproximal process reduced to a very low proximal protrusion merging without angular discontinuity with the posterior border, with posterodistal tooth very small, reduced to a broad acute angle scarcely projecting posteriorly; basis of pereiopod 7 fairly narrow with posterodistal tooth triangular and sharp, followed more proximally by low inconspicuous concavity, directed obliquely. Colour pattern White, with antennae and anterior part of body with a very pale pinkish hue; gnathopods and mouthparts blood red; eyes white. Body length 23 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA5685AFE14F8ABCD94FB45.taxon	distribution	Distribution Eastern shelf of the Weddell Sea at about 867 – 955 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA5685AFE14F8ABCD94FB45.taxon	discussion	Remarks The only other species with long styliform teeth is E. oxicarinata, but it belongs to a separate subgenus. The holotype and unique specimen of E. cyrano sp. nov. was apparently collected with a benthopelagic trawl. As the morphology of the species suggests a benthic life style, it is possible that the trawl operated close to the seafloor or hit the seafloor.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA36857FE58FB1ECA26FE3C.taxon	description	urn: lsid: zoobank. org: act: A 9 A 0 D 4 D 2 - 5 D 0 E- 4 E 40 - A 553 - 94013735 FC 7 F Figs 51 – 57	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA36857FE58FB1ECA26FE3C.taxon	etymology	Etymology The species is dedicated to Charlotte Havermans (formerly RBINS, currently Alfred-Wegener-Institute Helmholtz-Zentrum für Polar- und Meeresforschung), who collected the holotype of the species. Havermansiana is the feminine of havermansianus, which is a Latin adjective of the second declension derived from her name.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA36857FE58FB1ECA26FE3C.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., sex undetermined, cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 300 - 1, eastern Weddell Sea, BENDEX area, 70 ° 50.99 ʹ S, 10 ° 35.23 ʹ W to 70 ° 50.34 ʹ S, 10 ° 34.92 ʹ W, 227 – 266 m, bottom trawl, 1 Apr. 2011, (RBINS, INV. 132964) [extraction I 12; Genbank nr, 28 S: KU 759614]. Paratypes RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 1, GSN 1, eastern Weddell Sea, 71 ° 03.10 ʹ S, 11 ° 25.50 ʹ W, to 71 ° 02.10 ʹ S, 11 ° 19.30 ʹ W, 462 – 481 m, bottom trawl, 5 Feb. 1996, specimen kept in aquarium on board, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132700); 1 spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 17, GSN 10, eastern Weddell Sea, 73 ° 18.00 ʹ S, 21 ° 09.90 ʹ W, to 73 ° 19.10 ʹ S, 21 ° 14.90 ʹ W, 465 – 468 m, bottom trawl, 16 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132694). RV Aurora Australis cruises: SOUTHERN OCEAN: 3 specs, cruise CEAMARC, sample CEAMARC 3226, stn 20 EV 490, Adélie Coast, 66 ° 00 ʹ 59 ″ S, 140 ° 00 ʹ 02 ″ E, 189 – 196 m, beam trawl, 15 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4347); 1 spec., cruise CEAMARC, sample CEAMARC 3978, stn 36 EV 297, Adélie Coast, 66 ° 20 ʹ 20 ″ S, 143 ° 41 ʹ 08 ″ E, 552 – 573 m, beam trawl, 4 Jan. 2008 (MNHN-IU- 2014 - 7328 n, removed from MNHN-IU- 2014 - 4329). RV Marion Dufresne cruises: SOUTHERN OCEAN: 3 specs (2 large specs and 1 juv.) cruise MD 42 (SIBEX), stn 22 - CP 73, Prydz Bay, 66 ° 57 ʹ 25 ″ S, 72 ° 41 ʹ 25 ″ E, depth missing [Google Earth depth for that position: 540 m (accessed 27 Sep. 2016)], 26 Jan. 1985 (MNHN-IU- 2014 - 4260); 1 ♀, cruise MD 42 (SIBEX), stn 22 - CP 71, Prydz Bay, 66 ° 57 ʹ 37 ″ S, 72 ° 52 ʹ 07 ″ E, depth missing [Google Earth depth for that position: 513 m (accessed 27 Sep. 2016)], 26 Jan. 1985 (MNHN-IU- 2014 - 4268). Belgian and Belgian-Dutch Antarctic expedition: SOUTHERN OCEAN: 9 specs, Princess Ragnhild Coast, stn 219, Baie du Glacier, 70 ° 18 ʹ 05 ″ S, 23 ° 58 ʹ 00 ″ E, 216 m, trawl, 31 Jan. 1965 (RBINS, INV. 132743); 10 specs, Princess Ragnhild Coast, stn 219, Baie du Glacier, 70 ° 18 ʹ 05 ″ S, 23 ° 58 ʹ 00 ″ E, 216 m, trawl, 31 Jan. 1965, coll. A. Capart (RBINS, INV. 132744); 1 spec., Princess Ragnhild Coast, stn 219, Baie du Glacier, 70 ° 18 ʹ 05 ″ S, 23 ° 58 ʹ 00 ″ E, 216 m, trawl, 31 Jan. 1965 (RBINS, INV. 132747); 1 spec., Princess Ragnhild Coast, stn 219, Baie du Glacier, 70 ° 18 ʹ 05 ″ S, 23 ° 58 ʹ 00 ″ E, 216 m, trawl, 31 Jan. 1965 (RBINS, INV. 132750); 2 specs, Princess Ragnhild Coast, stn 220, Baie du Glacier, exact coordinates unavailable [presumably about 70 ° 18 ʹ S, 23 ° 58 ʹ E], 414 – 450 m, 1 Feb. 1965, trawl, coll. A. Capart (RBINS, INV. 132733); 1 spec., Princess Ragnhild Coast, stn 220, Baie du Glacier, exact coordinates unavailable [presumably about 70 ° 18 ʹ S, 23 ° 58 ʹ E], 414 – 450 m, 1 Feb. 1965, trawl, coll. A. Capart (RBINS, INV. 132735); 1 spec., Princess Ragnhild Coast, stn 220, Baie du Glacier, exact coordinates unavailable [presumably about 70 ° 18 ʹ S, 23 ° 58 ʹ E], 414 – 450 m, 1 Feb. 1965, trawl, coll. A. Capart (RBINS, INV. 132736); 1 spec., Princess Ragnhild Coast, stn 220, Baie du Glacier, exact coordinates unavailable [presumably about 70 ° 18 ʹ S, 23 ° 58 ʹ E], 414 – 450 m, 1 Feb. 1965, trawl, coll. A. Capart (RBINS, INV. 132751).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA36857FE58FB1ECA26FE3C.taxon	description	Description ROSTRUM. Long, reaching about tip of article 1 of peduncle of antenna 1 (teeth excluded), moderately curved, sharp-tipped in lateral view. EYE. Very large, broadly elliptic (nearly circular). PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 with indistinct posterior bump, with pair of low noncarinate dorsolateral swellings; pereionite 2 slightly narrower than pereionite 1, with small but distinct blunt mid-dorsal tooth and pair of low dorsolateral swellings; pereionite 3 with medium-sized blunttipped mid-dorsal tooth and pair of well-developed dorsolateral blunt teeth; pereionite 4 to pereionite 7 with large, broad, regularly curved, acute-tipped mid-dorsal tooth (dorsal length of teeth slightly and gradually increasing posteriorly) and pair of well developed conical dorsolateral teeth (pleonites 1 – 2 without second pair of dorsolateral teeth); pleonite 3 with dorsal sharp carina with weak median notch and produced posteriorly into an a broad acute and sharp triangular tooth, and pair of fairly large dorsolateral non-carinate acute-tipped teeth. COXAE 1 – 3. Strongly carinate and distally sharp. COXA 4. Anterodorsal + anteroventral borders forming a curve projecting forward (anterodorsal and anteroventral borders becoming straight respectively only at their upper and lower extremities); ventral tooth very sharp and very long; lateral carina without tooth, without distinct angularity, carina very distant from margin of coxa at its deepest point. COXA 5. With sharp and narrowly triangular carinate, lateral tooth pointing backwards (its lateral border is nearly parallel to body axis or weakly divergent). COXA 6. With mid-sized, sharp, triangular, carinate, lateral tooth pointing obliquely backwards; posteroventral corner bluntly angular. COXA 7. With ventral and posterior border straight, converging to form a sharp acute, nearly squared angle. EPIMERAL PLATES 1 – 3. Posteroventral angle produced into a very long sharp tooth. UROSOME TOOTH PATTERN. Urosomite 1 with strong triangular process pointing upwards; urosomite 2 with pair of small sharp posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.25; tips of lobes sharp, broadly V-shaped. PEDUNCLE OF ANTENNA 1. Article 1 with long lateral and medial teeth reaching beyond mid and nearly mid of article 2 (teeth excluded) and long ventral tooth nearly reaching tip of article 2 (teeth excluded); article 2 with very long lateral and medial teeth overreaching tip of article 3, with ventral tooth slightly overreaching tip of article 3 (tooth excluded); article 3 with long ventral tooth, about as long as article itself. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; propodus not narrowing distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus fairly slender; basis of pereiopods 5 – 6 of normal width, with posteroproximal process rounded and strongly protruding, with posterodistal tooth strong; basis of pereiopod 7 very broad with posterodistal tooth angulate, followed more proximally by small concavity, directed posteriorly. Body length Up to 33 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA36857FE58FB1ECA26FE3C.taxon	distribution	Distribution Eastern shelf of the Weddell Sea, Princess Ragnhild Coast, Prydz Bay, Adélie Coast; 189 – 573 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA36857FE58FB1ECA26FE3C.taxon	discussion	Remarks Epimeria havermansiana sp. nov. is morphologically very similar to E. pandora sp. nov., but delimitation methods based on the 28 S phylogeny indicate that they are indeed distinct species (Verheye et al. 2016 a, and this paper Fig. 342). On pereionite 1, E. havermansiana sp. nov. may have a trace of posterior middorsal bump, which is absent in E. pandora sp. nov. On pereionite 2, E. havermansiana sp. nov. has a posterior tooth, while E. pandora sp. nov. has only a slight bump. In E. havermansiana sp. nov., the dorsolateral teeth of the pereion pleosome and urosomite 2 are larger than in E. pandora sp. nov. Coxa 4 is anteriorly more curved than in E. pandora sp. nov. and its ventral tooth is longer and narrower.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFAE6855FE34FE35CCEBFADA.taxon	description	urn: lsid: zoobank. org: act: E 52 DDFA 4 - 14 DA- 4458 - 9 A 3 B- 403 A 37 E 19 A 4 B Figs 58 – 65	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFAE6855FE34FE35CCEBFADA.taxon	etymology	Etymology From the Greek, λευκος, white; οπλιτης, hoplite, citizen-soldier of the ancient Greece. The name, which is a noun in apposition, alludes to the armoured facies and the white or whitish colour of the species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFAE6855FE34FE35CCEBFADA.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: 1 ovigerous ♀, fixed in alcohol 70 %, cruise PS 69, ANT-XXIII / 8, stn 605 - 5, Elephant Island, 61 ° 20.27 ʹ S, 55 ° 30.92 ʹ W to 61 ° 20.37 ʹ S, 55 ° 28.99 ʹ W, 131 – 152 m, Agassiz trawl, 20 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122470). Paratypes RV Polarstern cruises: SOUTHERN OCEAN: 3 specs, initially fixed in formalin, cruise PS 69, ANT-XXIII / 8, stn 605 - 3, Elephant Island, 61 ° 20.33 ʹ S, 55 ° 31.53 ʹ W to 61 ° 20.35 ʹ S, 55 ° 30.18 ʹ W, 148 – 154 m, Agassiz trawl, 20 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122536); 1 ovigerous ♀, fixed in alcohol 70 %, cruise PS 69, ANT-XXIII / 8, stn 614 - 3, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122473); 1 spec., fixed in alcohol 70 %, cruise PS 69, ANT-XXIII / 8, stn 671 - 1, northwest of King George Island, 61 ° 59.98 ʹ S, 59 ° 14.78 ʹ W to 61 ° 60.00 ʹ S, 59 ° 10.74 ʹ W, 131 – 144 m, bottom trawl, 1 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122538); 1 juv., initially fixed in formalin, cruise PS 69, ANT-XXIII / 8, stn 728 - 2, northwest of Weddell Sea, south of Dundee Island, 63 ° 42.63 ʹ S, 56 ° 01.63 ʹ W to 63 ° 42.25 ʹ S, 56 ° 02.16 ʹ W, 293 – 298 m, Agassiz trawl, 24 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122539).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFAE6855FE34FE35CCEBFADA.taxon	description	Description ROBUSTNESS. Body and pereiopods more robust than in most Drakepimeria. ROSTRUM. Medium-sized, just reaching tip of article 1 of peduncle of antenna 1 (teeth excluded), weakly curved, sharp-tipped in lateral view. EYE. Large, elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 4 to pleonite 3 with mid-dorsal tooth; pereionite 1 to pleonite 3 with pair of dorsolateral teeth or protrusions (those of pereionites 1 – 2 so low that they are nearly inconspicuous); pereionites 1 – 3 without any trace of mid-dorsal tooth, with pair of extremely low, nearly inconspicuous, dorsolateral protrusions; pereionite 2 slightly but distinctly narrower than pereionite 1; pereionite 4 with very small blunt-tipped posterior mid-dorsal tooth and pair of very low blunt dorsolateral protrusions; pereionite 5 – 6 with medium-sized blunt-tipped broad mid-dorsal tooth and pair of small blunt dorsolateral teeth, which are anteriorly prolonged by a blunt carina; pereionite 7 with well-developed acute-tipped broad mid-dorsal tooth, which is anteriorly broadly angulate, and pair of small blunt dorsolateral teeth, which are anteriorly prolonged by a blunt carina; pleonites 1 – 3 with very broad and not very elevated, acute-tipped mid-dorsal tooth, which is anteriorly angulate and pair of small dorsolateral non-carinate teeth; on pleonite 1 a trace of second pair of (much smaller) dorsolateral teeth is observed between the mid-dorsal tooth and the main pair of dorsolateral teeth; pleonite 3 with large acute-tipped mid-dorsal tooth bearing an inconspicuous median concavity. COXAE 1 – 3. Strongly carinate and distally sharp. COXA 4. Anterodorsal border nearly straight (inconspicuously convex) and unusually long, anteroventral border nearly straight (inconspicuously concave) and short, joined by blunt but very distinct angular discontinuity, anterior corner not projecting forward; ventral tooth narrowly triangular, not long, apically subacute; lateral carina without tooth or angularity, not projecting laterally, carina very distant from margin of coxa at its deepest point. COXA 5. With well developed sharp and broadly triangular, carinate, lateral tooth, of which the anterior border diverges backwards and the posterior border is nearly perpendicular to body axis. COXA 6. With mid-sized, blunt, broadly triangular, carinate, lateral tooth, of which the anterior border diverges backwards and the posterior border is perpendicular to body axis; posteroventral corner broadly rounded. COXA 7. With ventral border distinctly curved, with posterior border nearly straight (inconspicuously convex), their convergence forming a very blunt angular discontinuity; surface of coxa posteriorly with a blunt and very low carina oriented in the dorsoventral axis. EPIMERAL PLATES 1 – 3. Posteroventral angle produced into a long and sharp tooth. UROSOME TOOTH PATTERN. Urosomite 1 with sharp triangular process pointing upwards; urosomite 2 without pair of small posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.33; tips of lobes subacute. PEDUNCLE OF ANTENNA 1. Article 1 with short lateral, medial and ventral teeth reaching the base of article 2; article 2 with large lateral and medial teeth reaching about tip of article 3 (ventral tooth excluded), with ventral tooth reduced to a tiny denticle; article 3 with small ventral tooth, distinctly shorter than article itself. GNATHOPODS 1 – 2. Carpus and propodus fairly robust, propodus narrowing distally, palm indistinct. PEREIOPODS 5 – 7. Merus, carpus and propodus very stout; basis of pereiopods 5 – 6 of normal width, with posteroproximal process rounded and distinctly protruding, with posterodistal tooth strong; basis of pereiopod 7 very broad with posterodistal tooth sharp, not followed more proximally by small concavity, directed posteriorly. Colour pattern Uniformly whitish, or whitish with a few extremely pale, small, brownish / yellowish spots. Appendages whitish. Eyes pale reddish. Body length Up to 43 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFAE6855FE34FE35CCEBFADA.taxon	distribution	Distribution Elephant Island and tip of Antarctic Peninsula, 131 – 298 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFAE6855FE34FE35CCEBFADA.taxon	discussion	Remark E. leukhoplites sp. nov. is superficially similar to E. vaderi but important differences are observed. In E. leukhoplites sp. nov., coxa 5 and 6 have large triangular carinae projecting laterally, which are very distinct in dorsal view, whilst E. vaderi has no such lateral projections; this is the most obvious difference. In E. leukhoplites sp. nov., the mid-dorsal tooth of pereionite 7 and pleonites 1 – 2 are anteriorly very angulate, whilst they form a regular curve in E. vaderi. In E. leukhoplites sp. nov., only the pair of dorsolateral teeth of pleonite 1 is duplicated, whilst this is also the case for pleonite 2 in E. vaderi. In E. leukhoplites sp. nov., the profile of the dorsal crest of pleonite 3 is nearly straight, with an inconspicuous trace of notch just on the middle, whilst in E. vaderi it presents a shallow but long concavity. In E. leukhoplites sp. nov., the central point of the lateral carina of coxa 4 is very distant from posterior border of coxa, whilst in E. vaderi the carina remains very close to the border of the coxa. The angle joining the anterodorsal and the anteroventral angle is also more distinct in E. leukhoplites sp. nov. than in E. vaderi. The posteroventral tooth of the epimeral plates is much stronger in E. leukhoplites sp. nov. than in E. vaderi, especially for the third one. Finally, in E. leukhoplites sp. nov., the propodus of the gnathopods narrows anteriorly and the palm is indistinct, whilst in E. vaderi the propodus is not tapering and the palm is normally developed. E. leukhoplites sp. nov. is also similar to Epimieria (Drakepimeria) subgen. nov. sp. 1 from the Ross Sea, which will be named and described in a separate paper by Verheye, Lörz & d’Udekem d’Acoz. The most obvious differences between the two species are given in the key.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFAC6851FE16FA93CEDCFE75.taxon	description	urn: lsid: zoobank. org: act: 878 EF 97 D-A 2 F 9 - 486 F-B 109 - 701 B 7 A 93840 A Figs 66 – 71	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFAC6851FE16FA93CEDCFE75.taxon	etymology	Etymology The species is dedicated to Anne-Nina Lörz (formerly NIWA), as a recognition of her important contribution to the taxonomy of the genus Epimeria. The name is a genitive.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFAC6851FE16FA93CEDCFE75.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: ♀, 36 mm, cruise PS 81, ANT-XXIX / 3, stn 116 - 9, north of Joinville Island, 62 ° 33.79 ʹ S, 56 ° 27.81 ʹ W to 62 ° 33.71 ʹ S, 56 ° 28.31 ʹ W, 248 m, muddy bottom with stones, Pentapora - like bryozoans and a lot of life, Agassiz trawl, 26 Jan. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122929 A) [extraction K 35; Genbank nr, 28 S: KU 759647]. Paratypes RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., photographed, cruise PS 14, ANT-VII / 4, EPOS leg 3, stn 230, eastern Weddell Sea, 75 ° 14.2 ʹ S, 26 ° 59.4 ʹ W to 75 ° 14.5 ʹ S, 26 ° 59.9 ʹ W, 270 – 275 m, Agassiz trawl, 30 Jan. 1989, coll. C. De Broyer (RBINS, INV. 132721); 4 specs, cruise PS 69, ANT-XXIII / 8, stn 605 - 1, Elephant Island, 61 ° 20.35 ʹ S, 55 ° 29.16 ʹ W to 61 ° 19.98 ʹ S, 55 ° 32.67 ʹ W, 146 – 151 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122571); 2 specs, cruise PS 69, ANT-XXIII / 8, stn 605 - 1, Elephant Island, 61 ° 20.35 ʹ S, 55 ° 29.16 ʹ W to 61 ° 19.98 ʹ S, 55 ° 32.67 ʹ W, 146 – 151 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert, (MNHN-IU- 2014 - 7329, removed from RBINS, INV. 122571); 7 medium-sized and small specs, cruise PS 69, ANT-XXIII / 8, stn 605 - 1, Elephant Island, 61 ° 20.35 ʹ S, 55 ° 29.16 ʹ W to 61 ° 19.98 ʹ S, 55 ° 32.67 ʹ W, 146 – 151 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122573); 3 specs, cruise PS 69, ANT-XXIII / 8, stn 605 - 5, Elephant Island, 61 ° 20.27 ʹ S, 55 ° 30.92 ʹ W to 61 ° 20.37 ʹ S, 55 ° 28.99 ʹ W, 131 – 152 m, Agassiz trawl, 20 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122567); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122568); 1 large spec., cruise PS 69, ANT-XXIII / 8, stn 686 - 1, Bransfield Strait, 62 ° 34.12 ʹ S, 55 ° 26.66 ʹ W to 62 ° 35.38 ʹ S, 55 ° 23.67 ʹ W, 149 m, bottom trawl, 4 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122474); 2 large specs, cruise PS 69, ANT-XXIII / 8, stn 689 - 1, north of Joinville Island, 62 ° 27.28 ʹ S, 55 ° 18.23 ʹ W to 62 ° 27.63 ʹ S, 55 ° 14.81 ʹ, 224 – 229 m, bottom trawl, 4 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122566); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 692 - 1, north of Joinville Island, 62 ° 21.76 ʹ S, 55 ° 36.96 ʹ W to 62 ° 23.62 ʹ S, 55 ° 36.42 ʹ W, 263 – 277 m, bottom trawl, 5 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122572); 2 very large specs, cruise PS 69, ANT-XXIII / 8, stn 693 - 1, north of Joinville Island, 62 ° 25.84 ʹ S, 55 ° 35.07 ʹ W to 62 ° 25.87 ʹ S, 55 ° 32.62 ʹ W, 243 – 291 m, bottom trawl, 5 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122471); 6 specs, cruise PS 69, ANT-XXIII / 8, stn 726 - 1, Snow Hill Island, 64 ° 30.86 ʹ S, 56 ° 40.23 ʹ W to 64 ° 31.16 ʹ S, 56 ° 40.51 ʹ W, 197 – 199 m, Rauschert dredge, 22 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122574); 2 large specs, cruise PS 69, ANT-XXIII / 8, stn 728 - 2, south of Dundee Island, 63 ° 42.63 ʹ S, 56 ° 01.63 ʹ W to 63 ° 42.25 ʹ S, 56 ° 02.16 ʹ W, 293 – 298 m, Agassiz trawl, 24 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122570); 1 ♀, 36 mm, cruise PS 81, ANT-XXIX / 3, stn 116 - 9, north of Joinville Island, 62 ° 33.79 ʹ S, 56 ° 27.81 ʹ W to 62 ° 33.71 ʹ S, 56 ° 28.31 ʹ W, 248 m, muddy bottom with stones and Pentapora - like bryozoans and a lot of life, Agassiz trawl, 26 Jan. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122929 B) [extraction K 36; Genbank nr, COI: KU 870868, 28 S: KU 759648]; 2 ♀♀, cruise PS 81, ANT-XXIX / 3, stn 116 - 9, north of Joinville Island, 62 ° 33.79 ʹ S, 56 ° 27.81 ʹ W to 62 ° 33.71 ʹ S, 56 ° 28.31 ʹ W, 248 m, muddy bottom with stones and Pentapora - like bryozoans and a lot of life, Agassiz trawl, 26 Jan. 2013 (RBINS, INV. 122951); 4 specs, cruise PS 81, ANT-XXIX / 3, stn 162 - 7, east of James Ross Island, 63 ° 58.78 ʹ S, 56 ° 46.24 ʹ W to 63 ° 59.02 ʹ S, 56 ° 46.26 ʹ W, 214 – 216 m, bottom with Glyptonotus and Echiniphimedia cf. hodgsoni, Agassiz trawl, 10 Feb. 2013 (RBINS, INV. 122940) [extraction ANT 35: largest specimen; Genbank nr, 28 S: KU 759594]; 3 specs, cruise PS 81, ANT-XXIX / 3, stn 162 - 7, east of James Ross Island, 63 ° 58.78 ʹ S, 56 ° 46.24 ʹ W to 63 ° 59.02 ʹ S, 56 ° 46.26 ʹ W, 214 – 216 m, bottom with Glyptonotus and Echiniphimedia cf. hodgsoni, Agassiz trawl, 10 Feb. 2013 (RBINS, INV. 122955); 2 specs, cruise PS 81, ANT-XXIX / 3, stn 162 - 7, east of James Ross Island, 63 ° 58.78 ʹ S, 56 ° 46.24 ʹ W to 63 ° 59.02 ʹ S, 56 ° 46.26 ʹ W, 214 – 216 m, bottom with Glyptonotus and Echiniphimedia cf. hodgsoni, Agassiz trawl, 10 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122958); 2 specs, cruise PS 81, ANT-XXIX / 3, stn 164 - 4, south of Dundee Island, 63 ° 37.28 ʹ S, 56 ° 9.11 ʹ W to 63 ° 37.29 ʹ S, 56 ° 9.58 ʹ W, 102 – 114 m, nonmuddy bottom with a lot of life, including Molgula, big red ophiuroids, and a lot of Glyptonotus, Agassiz trawl, 11 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 132974) [extraction I 19: largest specimen; Genbank nr, COI: KU 870844, 28 S: KU 759621]; 16 specs, cruise PS 81, ANT-XXIX / 3, stn 185 - 3, southeast of Dundee Island, 63 ° 51.34 ʹ S, 55 ° 41.11 ʹ W to 63 ° 51.52 ʹ S, 55 ° 41.43 ʹ W, 261 – 296 m, non muddy bottom (a lot of life: sponges, starfishes, ophiuroids, crinoids, Pentapora - like bryozoans), Agassiz trawl, 19 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122960); 4 small specs, cruise PS 81, ANT-XXIX / 3, stn 185 - 4 southeast of Dundee Island, 63 ° 51.53 ʹ S, 55 ° 40.74 ʹ W to 63 ° 51.53 ʹ S, 55 ° 40.43 ʹ W, 253 – 255 m, extremely fine sand mixed with some mud and gravel, Rauschert dredge, 19 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122971); 13 juvs, cruise PS 81, ANT-XXIX / 3, 185 - 4 southeast of Dundee Island, 63 ° 51.53 ʹ S, 55 ° 40.74 ʹ W to 63 ° 51.53 ʹ S, 55 ° 40.43 ʹ W, 253 – 255 m, extremely fine sand mixed with some mud and gravel, Rauschert dredge, 19 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122968). Expedition SIGNY 1991 / 92: SOUTHERN OCEAN: 1 adult, SIGNY 1991 / 92, AGT 24, transect 2, South Orkney Plateau: Signy Island, 60.704 ° S, 45.452 ° W, 190 – 200 m, 15 Feb. 1992, coll. Stefan Hain (RBINS, INV. 132686); 1 juv., SIGNY 1991 / 92, AGT 21, transect 2, South Orkney Plateau: Signy Island, 60.707 ° S, 45.437 ° W, 150 m, 14 Feb. 1992, coll. Stefan Hain (RBINS, INV. 132692).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFAC6851FE16FA93CEDCFE75.taxon	description	Description ROSTRUM. Long, overreaching tip of article 1 of peduncle of antenna 1 (teeth excluded), very strongly curved, sharp-tipped in lateral view. EYE. Very large, broadly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 with medium-sized broad and blunt mid-dorsal tooth pointing upwards, with pair of well-developed, conical, dorsolateral teeth; pereionite 2 much narrower than pereionites 1 and 3, without mid-dorsal tooth and without pair of dorsolateral teeth; pereionite 3 with well-developed broad and blunt mid-dorsal tooth pointing upwards, of which the anterior margin is strongly convex and the posterior margin weakly convex, and pair of fairly large, conical, dorsolateral teeth; pereionites 4 – 5 with large, fairly broad, acute to subacute mid-dorsal tooth, of which the anterior border is regularly curved and the posterior border is slightly concave, with pair of large narrowly conical dorsolateral teeth; pereionites 6 to pleonite 2 with large and fairly broad, acute mid-dorsal tooth of which the anterior border is subdivided into a proximal part pointing obliquely upwards and a posterior part nearly parallel to body axis, both parts being joined by distinct blunt angular discontinuity, with pair of large narrowly conical dorsolateral teeth (these pairs of teeth are never duplicate); pleonite 3 with large narrowly triangular acute-tipped symmetrical mid-dorsal tooth, and pair of large narrowly conical dorsolateral teeth (size of mid-dorsal teeth increasing from pereionite 1 to 5, those of pereionite 6 to pleonite 3 subequal to tooth of pereionite 5. COXAE 1 – 3. Strongly carinate and distally sharp. COXA 4. Anterodorsal border straight, anteroventral border weakly curved, these two borders being joined by obtuse, blunt but distinct angular discontinuity, anterior angle scarcely projecting forward; ventral tooth long and acute; lateral carina with large tooth, which is anteriorly convex and posteriorly concave, strongly projecting laterally and obliquely pointing backwards, carina very distant from margin of coxa at its deepest point. COXA 5. With long, sharp, narrowly triangular, carinate, lateral tooth pointing obliquely backwards; posteroventral corner broadly rounded. COXA 6. With long, sharp and narrowly triangular (nearly styliform), carinate lateral tooth pointing obliquely backwards. COXA 7. With ventral border slightly curved, with posterior border straight, their convergence forming a sharp squared angle. EPIMERAL PLATES 1 – 3. Posteroventral angle produced into a very long and very sharp tooth. UROSOME TOOTH PATTERN. Urosomite 1 with large and sharp narrow tooth pointing upwards; urosomite 2 with pair of mid-sized posterior dorsolateral teeth pointing upwards; urosomite 3 with pair of mid-sized posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.45; tips of lobes blunt and narrow, notch narrowly and bluntly V-shaped. PEDUNCLE OF ANTENNA 1. Article 1 with long lateral and medial teeth reaching mid of article 2 (teeth excluded) and long ventral tooth reaching about tip of article 2 (teeth excluded); article 2 with large lateral and medial teeth overreaching tip of article 3, without ventral tooth (ventral margin with scarcely distinct denticle); article 3 with medium-sized ventral tooth, about half as long as article itself. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; propodus not narrowing distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus slender; basis of pereiopods 5 – 6 of normal width, with posteroproximal process rounded and strongly protruding, with posterodistal tooth very strong (as long as basis width for pereiopod 5, nearly as long as basis width for pereiopod 6); basis of pereiopod 7 broad with posterodistal tooth subacute, followed more proximally by small concavity, directed posteriorly but somewhat obliquely. Colour pattern Whitish with irregular orange-red marks and dots arranged in complex patterns and in some places merging into small irregular transverse lines. Eyes reddish. This colour pattern is very constant and highly characteristic of the species. Body length Up to 30 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFAC6851FE16FA93CEDCFE75.taxon	distribution	Distribution South Orkney Islands; South Shetland Islands, including Elephant Island; Bransfield Strait; tip of Antarctic Peninsula; eastern shelf of the Weddell Sea (where it is very rare), 102 – 298 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFAC6851FE16FA93CEDCFE75.taxon	discussion	Remarks Epimeria loerzae sp. nov. is a common species off the South Shetland Islands and near the tip of the Antarctic Peninsula. It is also present off the South Orkney Islands, but it is very rare in the Eastern Weddell Sea, where only one specimen was found. Its bathymetrical range is apparently rather limited, since this species was not found in samples collected below 300 m. The colour pattern and the broad angulate mid-dorsal teeth of E. loerzae sp. nov. are unique in macrodonta - like Epimeria, making its identification easy (it is most similar to E. pyrodrakon sp. nov.). The collection locality of the specimen of Epimeria loerzae sp. nov. illustrated as Epimeria sp. n. 3 by Rauchert & Arntz (2015) was given in an early draft of their book made available to the authors: ANT-XV / 3 stn 355. Its coordinates are: 61 ° 59.8 ʹ S, 59 ° 14.8 ʹ W to 62 ° 00.1 ʹ S, 59 ° 14.8 ʹ W, 128 – 130 m, which corresponds to a position northwest of King George Island.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA8684FFE62FDCCCC4FFAC1.taxon	description	Figs 72 – 76	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA8684FFE62FDCCCC4FFAC1.taxon	materials_examined	Type material Lectotype National Antarctic Expedition 1901 – 1904, RV Discovery: SOUTHERN OCEAN: 1 spec., sex undetermined, Ross Sea, W. Q. [Winter Quarters — hole 12] 2 – 4 Sep. 1903, “ 236 ”, “ 237 ” (these two enigmatic numbers are put in a circle), J. 107, depth not stated, lectotype designated by Lörz et al. (2007), although no label by Lörz indicating its status as lectotype is present (BMNH 1907.6.6.259 - 262) (in part). Paralectotypes National Antarctic Expedition 1901 – 1904, RV Discovery: SOUTHERN OCEAN: 1 juv., mixed in the same tube with an adult ♀ of Epimeria colemani, Ross Sea, no locality but presumably McMurdo Sound, n ° 13, 914 m (500 fathoms), 22 Jan. 1902, obviously 2 of the paralectotypes of Epimeria macrodonta designated by Lörz et al. (2007), albeit no label by Lörz indicating their status as paralectotypes (BMNH 1907.6.6.259 - 262) (in part); 1 spec., sex undetermined, dissected by Walker, Ross Sea, no locality but presumably McMurdo Sound, 914 m (500 fathoms), 22 Jan. 1902, obviously 1 of the paralectotypes of Epimeria macrodonta designated by Lörz et al. (2007), albeit no label by Lörz indicating its status as paralectotype, 3 tubes (BMNH 1907.6.6.259 - 262) (in part).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA8684FFE62FDCCCC4FFAC1.taxon	description	Description Basis of description. Description based on lectotype. ROSTRUM. Long, reaching base of article 2 of peduncle of antenna 1 (teeth excluded), strongly curved, sharp-tipped in lateral view. EYE. Large, broadly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 with small and blunt posterior mid-dorsal protrusion, with pair of low dorsolateral protrusions; pereionite 2 much narrower than pereionites 1 and 3, without mid-dorsal tooth and without pair of dorsolateral teeth; pereionite 3 with medium-sized fairly broad and fairly blunt mid-dorsal tooth pointing upwards and pair of low and blunt, conical, dorsolateral teeth; pereionite 4 to pleonite 2 with large, narrow to very narrow, acute mid-dorsal tooth, of which the anterior border is almost regularly curved, with inconspicuous trace of angular discontinuity (pereionites 4 – 6) or with weak but distinct angular discontinuity (pereionite 7 and pleonites 1 – 2), and the posterior border is slightly concave, with pair of conical dorsolateral teeth of which the size gradually increases posteriorly (these pairs of teeth are never duplicate); pleonite 3 with large narrowly triangular acute-tipped subsymmetrical mid-dorsal tooth, and pair of large narrowly conical dorsolateral teeth (size of mid-dorsal teeth weakly and gradually increasing from pereionite 4 to pleonite 2; mid-dorsal tooth of pleonite 3 distinctly shorter than tooth of pleonite 2). COXAE 1 – 3. Strongly carinate and distally sharp; in dorsal view, the tip coxa 3 appears as projecting laterally. COXA 4. Anterodorsal border straight, anteroventral border distally nearly straight, these two borders being joined by a long weak curve (anterior angle), this anterior angle is weakly projecting forward; ventral tooth extremely long, very narrow and acute, weakly arching backwards; lateral carina with well developed tooth pointing obliquely backwards (in dorsal view its anterior border strongly diverges from body axis; in dorsal view this tooth form a narrow U-shaped notch with the coxa); carina very distant from margin of coxa at its deepest point. COXA 5. With long, sharp and narrowly triangular, carinate, lateral tooth pointing obliquely backwards (its anterior and posterior margins are distinctly oblique to body axis). COXA 6. With mid-sized, sharp and narrowly triangular, carinate, lateral tooth pointing obliquely backwards; its anterior border is distinctly convex); posteroventral corner broadly rounded. COXA 7. With ventral border distinctly curved, with posterior border nearly straight (inconspicuously convex), their convergence forming a curve (a very blunt squared angle). EPIMERAL PLATES 1 – 3. Posteroventral angle produced into a very long and very sharp tooth. UROSOME TOOTH PATTERN. Urosomite 1 with large and sharp narrow tooth pointing upwards; urosomite 2 with pair of mid-sized posterior dorsolateral teeth pointing upwards; urosomite 3 with pair of mid-sized posterior dorsolateral teeth pointing obliquely backwards. TELSON. Cleft on 0.33; tips of lobes subacute, notch narrowly V-shaped. PEDUNCLE OF ANTENNA 1. Article 1 with long lateral tooth and short medial tooth reaching respectively 0.9 and 0.33 of article 2 (teeth excluded) and long ventral tooth overreaching tip of article 2 (teeth excluded) by 0.4 of its length; article 2 with huge lateral tooth of which 0.6 is overreaching tip of article 3, and very huge medial teeth of which 0.7 is overreaching tip of article 3, without ventral tooth; article 3 with tiny ventral tooth, about 0.25 times as long as article itself. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus slender; basis of pereiopods 5 – 6 of normal width, with posteroproximal process rounded and strongly protruding, with posterodistal tooth very strong (nearly as long as basis width); basis of pereiopod 7 broad with posterodistal tooth acute, triangular, mid-sized, followed more proximally by distinct concavity, directed posteriorly. Body length Up to 25 mm. Variations The juvenile specimen has no posterodorsal bump on pereionite 1.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA8684FFE62FDCCCC4FFAC1.taxon	distribution	Distribution Ross Sea: Winter Quarters Bay [about 77 ° 50 ʹ S, 166 ° 39 ʹ E], no depth record [lectotype]; Ross Sea, no locality, 914 m [paralectotypes] (Walker 1906, 1907). The paralectotypes were presumably collected in McMurdo Sound because Hodgson (1907) stated that “ on the 20 th of January 1902, the ‘ Discovery’ passed across the mouth of the McMurdo Sound ” and the specimens were collected shortly afterwards, on 22 Jan. 1902.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFA8684FFE62FDCCCC4FFAC1.taxon	discussion	Remarks Epimeria macrodonta is morphologically similar to E. anguloce sp. nov. (Antarctic Peninsula, eastern Weddell Sea and Prydz Bay) and E. corbariae sp. nov. (Adélie Coast). Epimeria macrodonta can be distinguished from its relatives by the length of the teeth on article 1 of antenna 1 peduncle. In E. macrodonta, the lateral tooth is indeed considerably longer than the medial tooth, whilst in other species the teeth are subequal. It also exhibits other distinctive characters: lateral and medial tooth of article 2 of peduncle of antenna 1 especially long; ornamentation of pereionite 1 weak or absent; middorsal tooth of pereionite 4 to pleonite 2 especially long and slender (those of pereionite 6 and pleonite 1 – 2 exhibiting a slight angular discontinuity on their anterior border); ventral tooth of coxa 4 especially long; lateral tooth of coxae 4 and 5 very oblique in dorsal view; coxa 7 with posterior border nearly straight (inconspicuously convex), with posteroventral angle rounded; posterodistal tooth of basis of pereiopod 7 not very strong. The type locality of E. macrodonta is Ross Island (Ross Sea). Lörz et al. (2007) mistakenly reported Ross Island as being located off the Antarctic Peninsula. This lapsus might result from a confusion between Ross Island (Ross Sea) and James Ross Island (Antarctic Peninsula). E. macrodonta s. lat. has previously been recorded throughout the Southern Ocean, south of the Polar Front at depths shallower than 1000 m. However, all specimens except the lectotype and two of the three paralectotypes of E. macrodonta belong to other species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB6684CFE2DFA98CEF9FCDF.taxon	description	urn: lsid: zoobank. org: act: EABBCBFE- 2 ADE- 49 F 4 - B 739 - 35 CC 145 CE 6 A 9 Figs 77 – 82	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB6684CFE2DFA98CEF9FCDF.taxon	etymology	Etymology Πανδώρα is a well-known character of the Greek mythology. According to the myth, Pandora opened a jar, in modern accounts usually mistranslated as “ Pandora’s box ”, releasing all the evils of humanity, leaving only Hope inside once she had closed it again. She opened the jar out of simple curiosity and not as a malicious act. The name, which is a noun in apposition, should be considered as an allegory to the present study. The authors initially believed that the taxonomy of Antarctic Epimeria, was easy and fairly well-known, with perhaps a few attractive new species to describe; hence they had the curiosity to open glass jars filled with Epimeria specimens. In doing so, they opened the door to an inexhaustible flow of new species, often very difficult to separate from each other, and erased any hope of easy taxonomy for the genus.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB6684CFE2DFA98CEF9FCDF.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: ovigerous ♀, cruise PS 81, ANT-XXIX / 3, stn 217 - 6, Bransfield Strait, 62 ° 53.45 ʹ S, 58 ° 13.06 ʹ W to 62 ° 53.42 ʹ S, 58 ° 13.41 ʹ W, 461 – 483 m, rich sponge bottom, Agassiz trawl, 2 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122931 A) [extraction K 31; Genbank nr, COI: KU 870865, 28 S: KU 759644]. Paratypes RV Polarstern cruises: SOUTHERN OCEAN: 6 specs, cruise PS 69, ANT-XXIII / 8, stn 604 - 1, Elephant Island, 61 ° 20.52 ʹ S, 55 ° 09.72 ʹ W to 61 ° 20.11 ʹ S, 55 ° 07.26 ʹ W, 286 – 407 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122545); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 604 - 1, Elephant Island, 61 ° 20.52 ʹ S, 55 ° 09.72 ʹ W to 61 ° 20.11 ʹ S, 55 ° 07.26 ʹ W, 286 – 407 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (MNHN-IU- 2014 - 7330, removed from RBINS, INV. 122545); 1 spec., alcohol-fixed, cruise PS 69, ANT-XXIII / 8, stn 604 - 1, Elephant Island, 61 ° 20.52 ʹ S, 55 ° 09.72 ʹ W to 61 ° 20.11 ʹ S, 55 ° 07.26 ʹ W, 286 – 407 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122478); 2 specs, cruise PS 69, ANT-XXIII / 8, stn 604 - 1, Elephant Island, 61 ° 20.52 ʹ S, 55 ° 09.72 ʹ W to 61 ° 20.11 ʹ S, 55 ° 07.26 ʹ W, 286 – 407 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122547); 4 specs, cruise PS 69, ANT-XXIII / 8, stn 608 - 1, Elephant Island, 61 ° 11.34 ʹ S, 54 ° 43.17 ʹ W to 61 ° 11.80 ʹ S, 54 ° 40.05 ʹ W, 284 – 293 m, bottom trawl, 20 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122542); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 611 - 1, Elephant Island, 60 ° 58.90 ʹ S, 55 ° 11.31 ʹ W to 60 ° 58.52 ʹ S, 55 ° 07.82 ʹ W, 215 – 297 m, bottom trawl, 21 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122546); 6 small specs, cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122552); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 627 - 1, Elephant Island, 60 ° 59.00 ʹ S, 55 ° 42.36 ʹ W to 60 ° 57.62 ʹ S, 55 ° 40.19 ʹ W, 90 – 102 m, bottom trawl, 24 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122559); 2 specs, cruise PS 69, ANT-XXIII / 8, stn 642 - 1, Elephant Island, 61 ° 04.38 ʹ S, 55 ° 59.81 ʹ W to 61 ° 04.27 ʹ S, 55 ° 58.88 ʹ W, 254 m, Agassiz trawl, 27 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122548); 2 specs, cruise PS 69, ANT-XXIII / 8, stn 654 - 6, Elephant Island, 61 ° 22.80 ʹ S, 56 ° 03.84 ʹ W to 61 ° 23.35 ʹ S, 56 ° 04.89 ʹ W, 341 – 342 m, Agassiz trawl, 29 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122543); 1 large spec., cruise PS 69, ANT-XXIII / 8, stn 726 - 4, southeast of Snow Hill Island, 64 ° 37.83 ʹ S, 56 ° 42.10 ʹ W to 64 ° 38.03 ʹ S, 56 ° 42.57 ʹ W, 292 m, Agassiz trawl, 23 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122551); 1 ovigerous ♀, posterior part missing, cruise PS 81, ANT-XXIX / 3, stn 217 - 6, Bransfield Strait, 62 ° 53.45 ʹ S, 58 ° 13.06 ʹ W to 62 ° 53.42 ʹ S, 58 ° 13.41 ʹ W, 461 – 483 m, rich sponge bottom, Agassiz trawl, 2 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122931 B); 1 ovigerous ♀, cruise PS 81, ANT-XXIX / 3, stn 217 - 6, Bransfield Strait, 62 ° 53.45 ʹ S, 58 ° 13.06 ʹ W to 62 ° 53.42 ʹ S, 58 ° 13.41 ʹ W, 461 – 483 m, rich sponge bottom, Agassiz trawl, 2 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122935) [extraction K 32; Genbank nr, COI: KU 870866, 28 S: KU 759645].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB6684CFE2DFA98CEF9FCDF.taxon	description	Description ROSTRUM. Long, reaching about tip of article 1 of peduncle of antenna 1 (teeth excluded), moderately curved, sharp-tipped in lateral view. EYE. Very large, broadly elliptic (nearly circular). PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 without any trace of mid-dorsal tooth, with pair of very low non-carinate dorsolateral swellings; pereionite 2 nearly as broad as pereionite 1, with weak posterior mid-dorsal bump, and pair of indistinct dorsolateral swellings; pereionite 3 with small blunt-tipped middorsal tooth and pair of small dorsolateral blunt teeth; pereionite 4 with well-developed blunt-tipped mid-dorsal tooth and pair of small dorsolateral blunt teeth; pereionite 5 to pleonite 2 with large, acutetipped, broad, regularly curved mid-dorsal tooth (dorsal length of teeth slightly and gradually increasing posteriorly) and pair of dorsolateral non-carinate acute teeth (pleonites 1 – 2 without second pair of dorsolateral teeth); pleonite 3 with dorsal sharp carina with very weak median notch (lobe anterior to notch extremely low) and produced posteriorly into an a broad acute and sharp triangular tooth, and pair of dorsolateral non-carinate acute teeth. COXAE 1 – 3. Strongly carinate and distally sharp. COXA 4. Anterodorsal and anteroventral border nearly straight, joined by low and very blunt angular discontinuity, anterior angle not projecting forward; ventral tooth sharp and strong; lateral carina without tooth, without sharp angularity, carina very distant from margin of coxa at its deepest point. COXA 5. With sharp and narrowly triangular, carinate, lateral tooth pointing obliquely backwards. COXA 6. With mid-sized, sharp and narrowly triangular, carinate lateral tooth pointing obliquely backwards; posteroventral corner bluntly angular. COXA 7. With ventral and posterior border straight, converging to form a sharp acute angle (i. e. a tooth). EPIMERAL PLATES 1 – 3. Posteroventral angle produced into a very long sharp tooth. UROSOME TOOTH PATTERN. Urosomite 1 with strong triangular process pointing upwards; urosomite 2 with pair of small sharp posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.25; tips of lobes blunt, broadly V-shaped. PEDUNCLE OF ANTENNA 1. Article 1 with long lateral and medial teeth respectively nearly reaching beyond mid and nearly reaching mid of article 2 (teeth excluded) and long ventral tooth reaching tip of article 2 (teeth excluded); article 2 with very long lateral and medial teeth overreaching tip of article 3, with ventral tooth slightly overreaching tip of article 3 (tooth excluded); article 3 with long ventral tooth, about as long as article itself. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; propodus not narrowing distally, and palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus fairly slender; basis of pereiopods 5 – 6 of normal width, with posteroproximal process rounded and strongly protruding, with posterodistal tooth strong; basis of pereiopod 7 very broad with posterodistal tooth angulate, followed more proximally by small concavity, directed posteriorly. Colour pattern Body whitish with intense orange colour marks. Peduncle of antenna 1 orange-tinged. Rostrum, and posterior part of pereiopods vaguely orange-tinged; Eyes reddish. Body length Up to 45 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB6684CFE2DFA98CEF9FCDF.taxon	distribution	Distribution Elephant Island and tip of Antarctic Peninsula: Bransfield Strait and Snow Hill Island; 90 – 483 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB6684CFE2DFA98CEF9FCDF.taxon	discussion	Remarks Epimeria pandora sp. nov. is morphologically similar to the sympatric E. similis and E. colemani sp. nov. Epimeria pandora sp. nov. has a pair of small teeth pointing upwards on urosomite 2, which are absent in the two other species. It has no lateral tooth or angularity on the lateral carina of coxa 4, whilst E. similis has a well developed tooth and E. colemani sp. nov. a very obtuse tooth or angularity. Finally, the dorsolateral teeth of the pereion are conical in E. pandora sp. nov., whilst they are carinate in E. colemani sp. nov. The Peninsular E. pandora sp. nov. is replaced by the extremely similar E. havermansiana subgen. et sp. nov. in the eastern Weddell Sea, off Princess Ragnhild Coast, Prydz Bay and Adélie Coast. See also key to E. (Drakepimeria) subgen. et sp. nov. and account on E. havermansiana subgen. et sp. nov. for differences.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB56849FE31FC96CD7DFC95.taxon	description	urn: lsid: zoobank. org: act: BC 42 CFAC- 0277 - 4 B 3 D-B 6 FD- 06 A 9 C 2020 D 25 Figs 83 – 89	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB56849FE31FC96CD7DFC95.taxon	etymology	Etymology From the Greek Πυρρὸς, flame-coloured, yellowish red; and δράκων, dragon. The name alludes to the Great Red Dragon of the Book of Revelation, because its highly intricate ornamentation is not unlike that of a dragon and its colour accurately matches with the description given by John the Apostle. The Book of Revelation (12.3) indeed states: “ καὶ ὤφθη ἄλλο σημεῖον ἐν τῷ οὐρανῷ, καὶ ἰδοὺ δράκων πυρρὸς μέγας, ἔχων κεφαλὰς ἑπτὰ καὶ κέρατα δέκα καὶ ἐπὶ τὰς κεφαλὰς αὐτοῦ ἑπτὰ διαδήματα ”, which is translated as follows in the Holman Christian Standard Bible: “ Then another sign appeared in heaven: There was a great fiery red dragon having seven heads and ten horns, and on his heads were seven diadems ”. The name is a noun in apposition.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB56849FE31FC96CD7DFC95.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: ♀, in absolute alcohol, cruise PS 81, ANT-XXIX / 3, stn 193 - 8, Bransfield Strait, 62 ° 43.73 ʹ S, 57 ° 29.04 ʹ W to 62 ° 43.80 ʹ S, 57 ° 29.40 ʹ W, 428 – 431 m, Agassiz trawl, 23 Feb 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 132973) [extraction N 1; Genbank nr, 28 S: KU 759677]. Paratypes RV Polarstern cruises: SOUTHERN OCEAN: 2 specs, cruise PS 06, ANT-III / 3, stn 348, eastern Weddell Sea, 72 ° 50 ʹ S, 19 ° 23 ʹ W, 490 m, bottom trawl, 20 Feb. 1985, coll. J. Plötz (RBINS, INV. 132724); 1 spec., cruise PS 14, ANT-VII / 4, EPOS leg 3, stn 230, eastern Weddell Sea, 75 ° 14.2 ʹ S, 26 ° 59.4 ʹ W to 75 ° 14.5 ʹ S, 26 ° 59.9 ʹ W, 270 – 275 m, Agassiz trawl, 30 Jan. 1989, coll. C. De Broyer (RBINS, INV. 132722); 2 specs, kept on board in aquarium, cruise PS 39, ANT-XIII / 3, EASIZ I, stn 8, AGT 2, eastern Weddell Sea, 71 ° 18.70 ʹ S, 12 ° 17.10 ʹ W to 71 ° 18.45 ʹ S, 12 ° 16.30 ʹ W, 170 – 174 m, 9 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132685); 1 spec., kept on board in aquarium, cruise PS 39, ANT-XIII / 3, EASIZ I, stn 8, AGT 2, eastern Weddell Sea, 71 ° 18.70 ʹ S, 12 ° 17.10 ʹ W to 71 ° 18.45 ʹ S, 12 ° 16.30 ʹ W, 170 – 174 m, 9 Feb. 1996, coll. C. De Broyer and G. Chapelle (MNHN-IU- 2014 - 7331, removed from RBINS, INV. 132685); 1 spec. (used for gut content analysis and previously misidentified as E. macrodonta), cruise PS 39, ANT-XIII / 3, EASIZ I, stn 6, AGT 1, eastern Weddell Sea, 71 ° 31.80 ʹ S, 13 ° 34.50 ʹ W to 71 ° 31.86 ʹ S, 13 ° 35.50 ʹ W, 254 – 261 m, Agassiz trawl, 8 Jan. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132681); 4 specs, cruise PS 69, ANT-XXIII / 8, stn 603 - 5, eastern Weddell Sea, 70 ° 30.99 ʹ S, 08 ° 48.08 ʹ W to 70 ° 30.40 ʹ S, 08 ° 48.13 ʹ W, 274 – 297 m, sponge bottom, Agassiz trawl, 7 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122560); 1 spec., alcohol-fixed, cruise PS 69, ANT-XXIII / 8, stn 603 - 5, eastern Weddell Sea, 70 ° 30.99 ʹ S, 08 ° 48.08 ʹ W to 70 ° 30.40 ʹ S, 08 ° 48.13 ʹ W, 274 – 297 m, sponge bottom, Agassiz trawl, 7 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122475); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 726 - 4, southeast of Snow Hill Island, 64 ° 37.83 ʹ S, 56 ° 42.10 ʹ W to 64 ° 38.03 ʹ S, 56 ° 42.57 ʹ W, 292 m, Agassiz trawl, 23 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122564); 2 specs, cruise PS 69, ANT-XXIII / 8, stn 728 - 2, south of Dundee Island, 63 ° 42.63 ʹ S, 56 ° 01.63 ʹ W to 63 ° 42.25 ʹ S, 56 ° 02.16 ʹ W, 293 – 298 m, Agassiz trawl, 24 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122569); 1 ♀, dissected, absolute alcohol, cruise PS 81, ANT-XXIX / 3, stn 193 - 8, Bransfield Strait, 62 ° 43.73 ʹ S, 57 ° 29.04 ʹ W to 62 ° 43.80 ʹ S, 57 ° 29.40 ʹ W, 428 – 431 m, Agassiz trawl, 23 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 132960) [extraction I 17; Genbank nr, 28 S: KU 759619]. Belgian and Belgian-Dutch Antarctic expedition: SOUTHERN OCEAN: 1 spec., Princess Ragnhild Coast, stn 215, Baie Léopold, exact position missing, 234 m, trawl, 28 Jan. 1965, coll. A. Capart (RBINS, INV. 132730); 1 spec., Princess Ragnhild Coast, stn 219, Baie du Glacier, 70 ° 18 ʹ 05 ″ S, 23 ° 58 ʹ 00 ″ E, 216 m, trawl, 31 Jan. 1965, coll. A. Capart (RBINS, INV. 132727); 1 spec., Princess Ragnhild Coast, stn 219, Baie du Glacier, 70 ° 18 ʹ 05 ″ S, 23 ° 58 ʹ 00 ″ E, 216 m, trawl, 31 Jan. 1965, coll. A. Capart (RBINS, INV. 132729); 1 spec., Princess Ragnhild Coast, stn 219, Baie du Glacier, 70 ° 18 ʹ 05 ″ S, 23 ° 58 ʹ 00 ″ E, 216 m, trawl, 31 Jan. 1965, coll. A. Capart (RBINS, INV. 132727); 1 spec., Princess Ragnhild Coast, stn 234, between ‘ Baie des Pingouins’ and ‘ Baie du Polarhav’, 70 ° 19 ʹ S, 24 ° 26 ʹ E, 200 m, trawl, 2 Feb. 1967, coll. A. Capart (RBINS, INV. 132276).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB56849FE31FC96CD7DFC95.taxon	description	Description ROSTRUM. Long, reaching mid of article 2 of peduncle of antenna 1 (teeth excluded), strongly curved, subacute in lateral view. EYE. Large, elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 with medium-sized broad and blunt mid-dorsal tooth pointing backwards, with pair of small conical, dorsolateral teeth; pereionite 2 much narrower than pereionites 1 and 3, without mid-dorsal tooth and without pair of dorsolateral teeth; pereionite 3 with medium-sized broad and blunt mid-dorsal tooth pointing backwards and pair of fairly blunt, conical, dorsolateral teeth; pereionite 4 to pleonite 2 with large, moderately narrow, acute mid-dorsal tooth, of which the anterior border exhibits a slight angular discontinuity, and the posterior border is slightly concave, with pair of conical dorsolateral teeth of which the size gradually increases posteriorly (these pairs of teeth are never duplicate); pleonite 3 with large narrowly triangular, nearly symetrical, acutetipped mid-dorsal tooth, and pair of large styliform dorsolateral teeth (size of mid-dorsal tooth very weakly increasing from pereionite 4 to pleonite 2; mid-dorsal tooth of pleonite 3 as long as tooth of pleonite 2). COXAE 1 – 3. Strongly carinate and distally sharp. COXA 4. Anterodorsal border straight, anteroventral border weakly curved, these two borders being joined by a long curve (anterior angle), this anterior angle is weakly projecting forward; ventral tooth very long and acute; lateral carina with very large tooth obliquely pointing backwards; carina very distant from margin of coxa at its deepest point. COXA 5. With long, sharp, narrowly triangular, carinate, lateral tooth pointing obliquely backwards. COXA 6. With mid-sized, sharp and narrowly triangular, carinate, lateral tooth pointing obliquely backwards; posteroventral angle rounded, without angular discontinuity. COXA 7. With ventral border slightly curved, with posterior border straight, their convergence forming a blunt squared angle. EPIMERAL PLATES 1 – 3. Posteroventral angle produced into a very long and very sharp tooth. UROSOME TOOTH PATTERN. Urosomite 1 with large and sharp narrow tooth pointing upwards; urosomite 2 with pair of mid-sized posterior dorsolateral teeth pointing upwards; urosomite 3 with pair of mid-sized posterior dorsolateral teeth pointing obliquely upwards. TELSON. Cleft on 0.3; tips of lobes subacute, notch narrowly and sharply V-shaped. PEDUNCLE OF ANTENNA 1. Article 1 with medium-sized lateral, medial and ventral teeth reaching mid of article 2 (teeth excluded) or less; article 2 with medium-sized lateral, medial and ventral teeth, not reaching tip of article 3 (ventral tooth excluded); article 3 with well developed ventral tooth, 0.5 – 0.8 times as long as article itself. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; not broadening distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus slender; basis of pereiopods 5 – 6 of normal width, with posteroproximal process rounded and strongly protruding, with posterodistal tooth very strong (as long as basis width); basis of pereiopod 7 broad with posterodistal tooth acute and very large, followed more proximally by distinct concavity, directed posteriorly. Colour pattern Whitish with large orange red marks; eye reddish. Body length Up to 35 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB56849FE31FC96CD7DFC95.taxon	distribution	Distribution Tip of Antarctic Peninsula, eastern shelf of the Weddell Sea, Princess Ragnhild Coast; 170 – 490 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB56849FE31FC96CD7DFC95.taxon	biology_ecology	Biology Specimens previously identified as Epimeria macrodonta collected during the cruises ANT-VII / 4 and ANT-XIII / 3, and apparently used by Dauby et al. (2001 a, 2001 b) for their trophic studies were examined. They proved to be E. pyrodrakon sp. nov. According to Dauby et al. (2001 a), gut contents of their freshly collected “ Epimeria macrodonta ” (i. e., at least in part E. pyrodrakon sp. nov.) showed a wide variety of food items: cnidarians (hydroid perisarcs, gorgonian ossicles), crustaceans (pieces of euphausiids) and pycnogonids, sea cucumbers (ossicles), and plankton (foraminifers, diatoms, ostracods); sponge spicules and sand grains completed the diet. On the basis of these observations, Dauby et al. (2001 a, 2001 b) concluded that the species was an opportunistic feeder or an opportunistic predator, coupling microbrowsing on colonial organisms with active capture of small live prey and with microdetritivory.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB56849FE31FC96CD7DFC95.taxon	discussion	Remarks Epimeria pyrodrakon sp. nov. is superficially similar to the sympatric E. anguloce sp. nov., but exhibits a different size arrangement of teeth on the peduncle of antenna 1, a larger mid-dorsal tooth on pereionite 1 and a much longer tooth on the lateral carina of coxa 4. The ‘ Epimeria macrodonta ’ illustrated in fig. 21 by Coleman (2007) corresponds to the description of E. pyrodrakon sp. nov. based on the size and disposition of the teeth on the peduncle of antenna 1. The station of the specimen is not given in the book. It is 62 ° 59.38 ʹ S, 57 ° 4.82 ʹ W [tip of Antarctic Peninsula]; 3 Dec. 1984, leg. Wägele, 200 – 300 m, Agassiz trawl (Coleman pers. com.).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB06847FE83FCABCEC5FB75.taxon	description	Figs 90 – 98	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB06847FE83FCABCEC5FB75.taxon	materials_examined	Material examined RV Polarstern cruises: SOUTHERN OCEAN: ♀, partly dissected, stacking photographs, probably photographed on board — dotted colour morph without red patch, cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122477); 1 spec., probably photographed on board — dotted colour morph without red patch, cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122477); 2 specs, cruise PS 69, ANT-XXIII / 8, stn 608 - 1, Elephant Island, 61 ° 11.34 ʹ S, 54 ° 43.17 ʹ W to 61 ° 11.80 ʹ S, 54 ° 40.05 ʹ W, 284 – 293 m, 20 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122541); 2 specs, one of these specimens was photographed on board, colour morph with both red dots and red patch, cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5? 3 – 5?, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W, to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122481 A); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W, to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122481 B); ca 20 specs, cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, initially fixed with formalin, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122535); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, initially fixed with formalin, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (MNHN-IU- 2014 - 7332, removed from RBINS, INV. 122535); 1 spec., initially fixed with formalin, cruise PS 69, ANT-XXIII / 8, stn 622 - 1, Elephant Island, 60 ° 56.70 ʹ S, 55 ° 52.71 ʹ W to 60 ° 55.93 ʹ S, 55 ° 50.79 ʹ W, 218 – 307 m, Agassiz trawl, 23 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122540).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB06847FE83FCABCEC5FB75.taxon	description	Description ROSTRUM. Long, just reaching tip of article 1 of peduncle of antenna 1 (teeth excluded), distinctly curved on both sides, subacute in lateral view. EYE. Very large, broadly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 4 without any trace of mid-dorsal tooth, without pair of dorsolateral teeth or protrusions; pereionite 2 slightly narrower than pereionite 1; pereionite 5 with trace of dorsoventrally flattened posterior mid-dorsal tooth (very obtuse in dorsal view, subacute in lateral view), with pair of flattened and indistinct dorsolateral protrusions or small teeth; pereionites 6 – 7 with small and low subacute mid-dorsal tooth, with pair of flattened dorsolateral protrusions; pleonites 1 – 3 with low, acute-tipped mid-dorsal tooth (apically broader on pleonite 3), which are not anteriorly angulate (but more or less regularly curved) and pair of small dorsolateral non-carinate teeth (of which none is duplicated); pleonite 3 with large acute-tipped mid-dorsal tooth bearing an inconspicuous trace of median concavity. COXAE 1 – 3. Carinate; coxa 1 not tapering, distally very broad and very blunt; coxae 2 – 3 distally blunt and fairly broad. COXA 4. Anteriorly forming a strong curve (not projecting much forward), where the anterodorsal and the anteroventral borders merge into each other without angular discontinuity; ventral tooth fairly long, narrow, apically acute, arching backwards, lateral carina without tooth but forming a very low, very bluntly angular lobe projecting laterally, carina very distant from margin of coxa at its deepest point. COXA 5. With well developed, sharp and broadly triangular, lateral carinate tooth, of which the anterior border weakly diverges backwards and the posterior border is nearly perpendicular to body axis. COXA 6. With mid-sized, subacute and broadly subtriangular, carinate, lateral tooth, of which the proximal half of the anterior border diverges backwards, the distal half of the anterior border is nearly parallel with body axis (transition between anterior and posterior half with angular discontinuity), and the posterior border is nearly perpendicular to body axis; posteroventral corner produced broadly rounded. COXA 7. With ventral border distinctly curved, with posterior border weakly curved, their convergence forming a very blunt angular discontinuity. EPIMERAL PLATES 1 – 3. Posteroventral angle produced into a long and sharp tooth (especially the third one). UROSOME TOOTH PATTERN. Urosomite 1 with low and blunt triangular process pointing upwards; urosomite 2 with pair of small but sharp posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.3; tips of lobes subacute, slit forming a fairly broad V. PEDUNCLE OF ANTENNA 1. Article 1 with tiny lateral and medial teeth and fairly small ventral tooth reaching half of article 2; article 2 without lateral tooth (it is replaced by a weak lobe) and two tiny medial teeth, with small ventral tooth reaching about 0.3 – 0.5 of article 3; article 3 with well developed ventral tooth, slightly longer than article itself. GNATHOPODS 1 – 2. Carpus and propodus fairly robust, propodus not narrowing distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus slender; basis of pereiopods 5 – 6 of normal width, with posteroproximal process rounded, distinctly but not strongly protruding, with posterodistal tooth broadly triangular and strong; basis of pereiopod 7 very broad with posterodistal tooth triangular and sharp, followed more proximally by shallow concavity, directed posteriorly. Colour pattern Body, coxae and pereiopods studded with bright red or reddish small dots on a whitish background. In some specimens there is a large irregular red patch on the anterior half of the body. Eyes reddish to red. Body length Up to 40 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB06847FE83FCABCEC5FB75.taxon	distribution	Distribution RV Polarstern cruise, PS 56, ANT-XVIl / 3, Stn. 171 - 3 (Lörz & Coleman 2001). Exact position: 63 ° 00.10 ʹ S, 060 ° 30.70 ʹ W to 63 ° 00.10 ʹ S, 60 ° 31.00 ʹ W [Bransfield Strait: Deception Island, outside of the caldera, near South East Point], 45 – 48 m (Arntz & Brey 2001). Elephant Island, 218 – 307 m (present material).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB06847FE83FCABCEC5FB75.taxon	discussion	Remark All the specimens of Epimeria reoproi examined by us have a pair of small postero-dorsolateral teeth on urosomite 2. Lörz & Coleman (2001) stated that urosomite 2 was smooth in the type material. However, a tiny postero-dorsolateral tooth pointing backwards is visible on their figure 1 b, suggesting that their description is incorrect.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFBE6845FE3AFACDCB65F939.taxon	description	urn: lsid: zoobank. org: act: 8433 E 457 - 9757 - 41 B 0 - BC 19 - 2 A 95 EE 5 E 3176 Figs 99 – 105	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFBE6845FE3AFACDCB65F939.taxon	etymology	Etymology The species is dedicated to Henri Robert (Royal Belgian Institute of Natural Sciences), who collected a large part of the type material. Robertianus, - a, - um is an adjective derived from his name.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFBE6845FE3AFACDCB65F939.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: ♂, initially fixed in formalin, cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, eastern Weddell Sea, given by colleagues to H. Robert on 22 Dec. 2007, presumably stn 17 - 10, of which the coordinates are: 70 ° 4.58 ʹ S, 3 ° 19.66 ʹ W to 70 ° 4.48 ʹ S, 3 ° 19.20 ʹ W, 2163 – 2190 m, Agassiz trawl, coll. H. Robert (RBINS, INV. 132413). Paratypes RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., cruise PS 48, ANT-XV / 3, EASIZ II, stn 134, eastern Weddell Sea, 74 ° 32.2 ʹ S, 27 ° 13.8 ʹ W to 74 ° 32.5 ʹ S, 27 ° 14.5 ʹ W, 2054 – 2081 m, Agassiz trawl, 9 Feb. 1998, coll. C. De Broyer (RBINS, INV. 132991); 1 immature spec., sex undetermined, cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, stn 17 - 11, eastern Weddell Sea, 70 ° 5.13 ʹ S, 3 ° 23.50 ʹ W to 70 ° 4.66 ʹ S, 3 ° 21.37 ʹ W, 1724 – 2091 m, epibenthic sledge, 22 Dec. 2007, coll. H. Robert (RBINS, INV. 132963) [extraction K 44; Genbank nr, COI: KU 870842, 28 S: KU 759618]; 2 specs, initially fixed in formalin, cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, eastern Weddell Sea, given by colleagues to H. Robert on 22 Dec. 2007, presumably stn 17 - 10, of which the coordinates are: 70 ° 4.58 ʹ S, 3 ° 19.66 ʹ W to 70 ° 4.48 ʹ S, 3 ° 19.20 ʹ W, 2163 – 2190 m, Agassiz trawl, coll. H. Robert (RBINS, INV. 132413).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFBE6845FE3AFACDCB65F939.taxon	description	Description ROSTRUM. Long, overreaching mid of article 2 of peduncle of antenna 1, scarcely curved, sharp-tipped in lateral view. EYE. Very large, elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 4 without any trace of mid-dorsal tooth and dorsolateral protrusions or teeth; pereionite 2 nearly as broad as pereionite 1; pereionite 5 with very distinct posterodorsal bump and pair of weak dorsolateral teeth; pereionite 6 with small mid-dorsal tooth (of which the anterior border is straight) and pair of weak dorsolateral teeth; pereionite 7 with low mediumsized mid-dorsal tooth (of which the anterior border is weakly curved) and pair of small dorsolateral non-carinate teeth; pleonites 1 – 2 with low but large and very sharp, anteriorly weakly curved mid-dorsal tooth and pair of small dorsolateral non-carinate teeth; on pleonite 1 a second pair of (much smaller) dorsolateral teeth is observed between the mid-dorsal tooth and the main pair of dorsolateral teeth; pleonite 3 with low but large acute-tipped mid-dorsal tooth bearing an inconspicuous median notch and pair of small dorsolateral non-carinate teeth (mid-dorsal teeth gradually increasing in size in a backwards direction). COXAE 1 – 3. Weakly carinate and distally very sharp. COXA 4. Not broad; anterodorsal and anteroventral border straight joined by low and blunt angular discontinuity, anterior angle not projecting forward; ventral tooth medium-sized and acute; lateral carina not sharp, without tooth or angularity, carina very distant from margin of coxa at its deepest point. COXA 5. With long, sharp and broadly triangular, carinate, lateral tooth. COXA 6. With mid-sized, sharp and broadly triangular, carinate, lateral tooth; posteroventral corner broadly rounded. COXA 7. With ventral border strongly convex, with posterior border distinctly convex, their convergence forming a very rounded angular discontinuity. EPIMERAL PLATES 1 – 3. Posteroventral angle: produced into a medium-sized and very sharp tooth, especially the plate 3. UROSOME TOOTH PATTERN. Urosomite 1 with sharp tooth pointing backwards of which the anterior border is weakly convex and longer than the posterior border, and of which the posterior border is strongly curved and concave; urosomite 2 without pair of small posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.2; tips of lobes acute, separated by very broad V-shaped notch. PEDUNCLE OF ANTENNA 1. Article 1 without lateral tooth, with very short medial tooth, and medium-sized ventral tooth reaching 0.4 of article 2; article 2 without lateral tooth, with very short medial tooth, and short ventral tooth reaching 0.3 of article 2; article 3 with well developed ventral tooth, about as long as article itself. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; propodus not narrowing distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus slender; basis of pereiopods 5 – 6 narrow, with posteroproximal process rounded and very weak, not well individualized, with posterodistal tooth small but acute and obliquely directed in pereiopod 5, reduced to a squared angle in pereiopod 6; basis of pereiopod 7 broad with posterodistal tooth sharp, not followed more proximally by distinct concavity, directed obliquely. Colour pattern Body and appendages uniformly pink or pale purple, except for eyes and gnathopods, which are red (colour photograph published by Coleman 2007 and Rauschert & Arntz 2015, both as E. reoproi). Body length Up to 31 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFBE6845FE3AFACDCB65F939.taxon	distribution	Distribution Eastern shelf of the Weddell Sea, 1724 – 2190 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFBE6845FE3AFACDCB65F939.taxon	discussion	Remarks Rauschert & Arntz (2015) published a colour photograph of E. robertiana sp. nov. (as E. reoproi), without indicating the origin of the specimen. However, that information was given in an early draft of the book made available to the authors: ANT-XV / 3, stn 134. The coordinates of that station are: 74 ° 32.2 ʹ S, 27 ° 13.8 ʹ W to 74 ° 32.5 ʹ S, 27 ° 14.5 ʹ W, 2054 – 2081 m. The same photograph was previously published (also as E. reoproi) by Coleman (2007) and Lörz & Coleman (2009). The inclusion of E. robertiana sp. nov. in the subgenus Drakepimeria is supported by molecular data. However, its narrow basis of pereiopod 5 and its posteriorly directed process of urosomite 1 are character states unusual for the subgenus Drakepimeria. On the other hand, these character states are frequent in non-Antarctic Epimeria species, which are predominantly deep-sea forms, just like E. robertiana sp. nov.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFBC6844FEE7F931CA42FBA6.taxon	diagnosis	Diagnosis Diagnosis based on the figures of Lörz et al. (2007). Pereionites 1 – 2 smooth; pereionite 3 with posterior mid-dorsal bump and pair of dorsolateral protrusions; pereionites 3 – 7 with moderately slender mid-dorsal tooth of length increasing posteriorly and pair of dorsolateral smaller teeth; mid-dorsal tooth of pleonite 3 narrow and without median notch. Urosomite 2 with pair of small dorsolateral teeth pointing upwards. lateral carina of coxa 4 with well-developed tooth pointing backwards. Peduncle of antenna 1: article 1 with short lateral and medial teeth, with long ventral tooth reaching tip of article 2 (teeth excluded); article 2 with long lateral and medial teeth overreaching article 3; article 3 with short ventral tooth shorter than article itself.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFBC6844FEE7F931CA42FBA6.taxon	description	Colour pattern Usually whitish with irregular pale orange patches, more rarely whitish with red transverse stripes (one stripe per body segment). Body length 30 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFBC6844FEE7F931CA42FBA6.taxon	distribution	Distribution Western Ross Sea, 130 – 350 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFBC6844FEE7F931CA42FBA6.taxon	discussion	Remarks Epimeria schiaparelli is very similar to E. corbariae sp. nov. They can be differentiated by the dorsal ornamentation of pereionite 3: a posterior bump in E. schiaparelli vs a tooth in E. corbariae sp. nov.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFBD6841FE50FBB0CBADFAB7.taxon	description	Figs 106 – 124	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFBD6841FE50FBB0CBADFAB7.taxon	materials_examined	Type material Lectotype (designated here) RV Pourquoi Pas? Cruises: SOUTHERN OCEAN: undissected ovigerous ♀, second French Antarctic expedition 1908 – 1910, Draguage XVII, South Shetlands, King George Island, Admiralty Bay, 420 m (n ° 713) (MNHN Am. 5984 and MNHN-IU- 2013 - 17865). Paralectotype (designated here) RV Pourquoi Pas? Cruises: SOUTHERN OCEAN: 1 ♀ dissected by E. Chevreux, second French Antarctic expedition 1908 – 1910, Draguage XVII, South Shetlands King George Island, Admiralty Bay, 420 m (MNHN Am. 3095 and MNHN-IU- 2013 - 17864). Other material examined RV Polarstern cruises: SOUTHERN OCEAN: 3 specs, cruise PS 69, ANT-XXIII / 8, stn 604 - 1, Elephant Island, 61 ° 20.52 ʹ S, 55 ° 09.72 ʹ W to 61 ° 20.11 ʹ S, 55 ° 07.26 ʹ W, 286 – 407 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 132980); 1 spec. with a true posterodorsal bump on second body segment, cruise PS 69, ANT-XXIII / 8, stn 604 - 1, Elephant Island, 61 ° 20.52 ʹ S, 55 ° 09.72 ʹ W to 61 ° 20.11 ʹ S, 55 ° 07.26 ʹ W, 286 – 407 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122555); 2 specs, cruise PS 69, ANT-XXIII / 8, stn 608 - 1, Elephant Island, 61 ° 11.34 ʹ S, 54 ° 43.17 ʹ W to 61 ° 11.80 ʹ S, 54 ° 40.05 ʹ W, 284 – 293 m, bottom trawl, 20 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 132982); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 610 - 1, Elephant Island, 60 ° 58.59 ʹ S, 55 ° 08.39 ʹ W to 60 ° 58.05 ʹ S, 55 ° 05.00 ʹ W, 287 – 311 m, bottom trawl, 21 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122550); 2 specs, cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 132984); 2 juvs, cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122554); 1 spec., alcohol-fixed, cruise PS 69, ANT-XXIII / 8, stn 624 - 3, Elephant Island, Agassiz trawl, 61 ° 00.23 ʹ S, 55 ° 58.53 ʹ W to 61 ° 00.76 ʹ S, 55 ° 59.20 ʹ W, 287 – 319 m, 23 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122479); 4 specs, cruise PS 69, ANT-XXIII / 8, stn 627 - 1, Elephant Island, 60 ° 59.00 ʹ S, 55 ° 42.36 ʹ W to 60 ° 57.62 ʹ S, 55 ° 40.19 ʹ W, 90 – 102 m, bottom trawl, 24 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 132979); 5 specs, cruise PS 69, ANT-XXIII / 8, stn 642 - 1, Elephant Island, 61 ° 04.38 ʹ S, 55 ° 59.81 ʹ W to 61 ° 04.27 ʹ S, 55 ° 58.88 ʹ W, 254 m, Agassiz trawl, 27 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 132981); 1 small spec., cruise PS 69, ANT-XXIII / 8, stn 642 - 2, Elephant Island, 61 ° 04.28 ʹ S, 55 ° 58.93 ʹ W to 61 ° 04.24 ʹ S, 55 ° 59.27 ʹ W, 255 – 277 m, Rauschert dredge, 27 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122557); 2 specs, alcohol-fixed, cruise PS 69, ANT-XXIII / 8, stn 654 - 6, Elephant Island, 61 ° 22.80 ʹ S, 56 ° 03.84 ʹ W to 61 ° 23.35 ʹ S, 56 ° 04.89 ʹ W, 341 – 342 m, Agassiz trawl, 29 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122480); 1 small spec., cruise PS 69, ANT-XXIII / 8, stn 654 - 6, Elephant Island, 61 ° 22.80 ʹ S, 56 ° 03.84 ʹ W to 61 ° 23.35 ʹ S, 56 ° 04.89 ʹ W, 341 – 342 m, Agassiz trawl, 29 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122556); 2 ♀♀, cruise PS 69, ANT-XXIII / 8, stn 654 - 6, Elephant Island, 61 ° 22.80 ʹ S, 56 ° 03.84 ʹ W to 61 ° 23.35 ʹ S, 56 ° 04.89 ʹ W, 341 – 342 m, Agassiz trawl, 29 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 132983); 5 specs, cruise PS 69, ANT-XXIII / 8, stn 676 - 1, north of Livingstone Island, 62 ° 11.06 ʹ S, 60 ° 47.49 ʹ W to 62 ° 09.65 ʹ S, 60 ° 49.56 ʹ W, 418 – 472 m, bottom trawl, 2 Dec. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122544); 2 specs, cruise PS 69, ANT-XXIII / 8, stn 680 - 5, northeast of Livingstone Island, 62 ° 23.37 ʹ S, 61 ° 25.58 ʹ W 62 ° 22.75 ʹ S, 61 ° 25.97 ʹ W, 324 – 349 m, Agassiz trawl, 3 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122553); 1 ♀, specimen used for detailed illustrations, cruise PS 81, ANT-XXIX / 3, stn 193 - 8, Bransfield Strait, 62 ° 43.73 ʹ S, 57 ° 29.04 ʹ W to 62 ° 43.80 ʹ S, 57 ° 29.40 ʹ W, 426 - 431 m, 23 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122956 A) [extraction P 36; Genbank nr, 28 S: KU 759680]; 3 ♀♀, cruise PS 81, ANT-XXIX / 3, stn 193 - 8, Bransfield Strait, 62 ° 43.73 ʹ S, 57 ° 29.04 ʹ W to 62 ° 43.80 ʹ S, 57 ° 29.40 ʹ W, 426 - 431 m, 23 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122956 B); 6 specs, cruise PS 81, ANT-XXIX / 3, stn 193 - 9, Bransfield Strait, 62 ° 43.50 ʹ S, 57 ° 27.92 ʹ W to 62 ° 43.53 ʹ S, 57 ° 28.28 ʹ W, 420 – 431 m, sponge bottom, 23 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122961); 1 spec., cruise PS 81, ANT-XXIX / 3, stn 217 - 6, Bransfield Strait, 62 ° 53.45 ʹ S, 58 ° 13.06 ʹ W to 62 ° 53.42 ʹ S, 58 ° 13.41 ʹ W, 461 – 483 m, rich sponge bottom, Agassiz trawl, 2 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122922 B) [extraction P 38; Genbank nr, 28 S: KU 759682].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFBD6841FE50FBB0CBADFAB7.taxon	description	Description ROSTRUM. Long, reaching about tip of article 1 of peduncle of antenna 1 (teeth excluded), weakly curved, sharp-tipped in lateral view. EYE. Very large, elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 without any trace of mid-dorsal tooth, with pair of very low non-carinate dorsolateral swellings; pereionite 2 nearly as broad as pereionite 1, with or without very small blunt-tipped mid-dorsal tooth or bump, and with pair of dorsolateral non-carinate swellings or very blunt teeth; pereionite 3 with medium-sized blunt-tipped mid-dorsal tooth and pair of small dorsolateral non-carinate blunt teeth; pereionite 3 to pleonite 2 with acute-tipped broad regularly curved mid-dorsal tooth (nearly same shape and size on all these segments) and pair of dorsolateral non-carinate acute-tipped teeth (pleonites 1 – 2 without second pair of dorsolateral teeth); pleonite 3 with dorsal sharp carina with very weak median notch (lobe anterior to notch very low) and obliquely produced posteriorly into an a broad acute and sharp triangular tooth, and pair of dorsolateral non-carinate acute-tipped teeth. COXAE 1 – 3. Strongly carinate and distally sharp. COXA 4. Anterodorsal and anteroventral border nearly straight, joined by low and blunt angular discontinuity, anterior angle not strongly projecting forward; ventral tooth very long and styliform; lateral carina with lateral well-developed and sharp triangular tooth pointing obliquely backwards. COXA 5. With sharp and broadly triangular, carinate, lateral tooth pointing obliquely backwards. COXA 6. With mid-sized, sharp and narrowly triangular, carinate, lateral tooth pointing obliquely backwards; posteroventral corner produced into a triangular tooth. COXA 7. With ventral and posterior border straight, with posteroventral angle produced into a strong tooth. EPIMERAL PLATES 1 – 3. Posteroventral angle produced into a very long styliform tooth. UROSOME TOOTH PATTERN. Urosomite 1 with sharp triangular process pointing upwards; urosomite 2 without pair of small posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.25; tips of lobes rounded, slit fairly narrow. PEDUNCLE OF ANTENNA 1. Article 1 with long lateral and medial teeth nearly reaching mid of article 2 (teeth excluded) and long ventral tooth reaching tip of article 2 (teeth excluded); article 2 with long lateral tooth nearly reaching tip of article 3 (tooth excluded), with medial tooth slightly overreaching article 3 (tooth excluded), with ventral tooth slightly overreaching tip of article 3 (tooth excluded); article 3 with very long ventral tooth, longer than article itself. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; propodus not narrowing distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus slender; basis of pereiopods 5 – 6 of normal width, with posteroproximal process rounded and strongly protruding, with posterodistal tooth strong; basis of pereiopod 7 very broad with posterodistal tooth sharp, followed more proximally by small concavity, directed posteriorly. Colour pattern Whitish with faint orange marks on body; peduncle of antenna 1 carpus and propodus of pereiopods and uropods also tinged with orange. Eyes reddish. Body length Up to 40 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFBD6841FE50FBB0CBADFAB7.taxon	distribution	Distribution Elephant Island to Bransfield Strait, 90 – 483 m (present data).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFBD6841FE50FBB0CBADFAB7.taxon	discussion	Remarks The tip of the lateral tooth of coxa 4 is slightly damaged in the syntype of Epimeria similis illustrated by Chevreux (1913). This explains why the orientation of that tooth does not seem accurate on the drawings made by that author. Specimens of E. similis from Elephant Island look identical to specimens recorded further south, except for the absence of a mid-dorsal tooth on pereionite 2, which is either smooth or with a trace of bump. The difference is presumably size-related. Epimeria similis from Bransfield Strait and King George Island often reach a larger size than those of Elephant Island, and only the large ones have the extra tooth. So the specimens from Elephant Island are considered herein as small E. similis s. str. The collecting station of the E. similis illustrated by Coleman (2007) on his figure 30 are 61 ° 03.6 ʹ S, 54 ° 41.6 ʹ W [Elephant Island]; 15 Dec. 1987, leg. Coleman, 358 – 332 m, bottom trawl (Coleman pers. com.). This specimen appears to be a true E. similis. On the other hand, the specimen illustrated by a colour photograph on his plate 1, fig. e is presumably Epimeria acanthochelon sp. nov. (see discussion on that species). It seems that E. similis and the similar and closely related E. acanthochelon sp. nov. have non-overlapping distributions. Epimeria similis was recorded from the South Shetland Islands and the tip of the Antarctic Peninsula. Epimeria acanthochelon sp. nov. was recorded on the Adélie Coast (supported by 28 S rDNA) and on the eastern shelf of the Weddell Sea (identification based on morphology only). See key of Drakepimeria and account on E. acanthochelon sp. nov. for differences with E. similis.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB8683FFE5BFA8ECEC5FEFE.taxon	description	Description Description based on the figures of Coleman (1998 b). ROBUSTNESS. More robust than most Drakepimeria. Rostrum: medium-sized, just overreaching tip of article 1 of peduncle of antenna 1 (teeth excluded), weakly curved, sharp-tipped in lateral view. EYE. Very large, elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 3 without any trace of mid-dorsal tooth, without dorsolateral protrusions; pereionite 2 slightly narrower than pereionite 1; pereionite 4 with very faint posterodorsal bump and no dorsolateral protrusions; pereionite 5 to pleonite 2 with broad and low, regularly curved, dorsolateral tooth, and pair of small posterodorsal swelling or teeth; on pleonites 1 – 2 a second pair of (a bit smaller) dorsolateral teeth is observed between the mid-dorsal tooth and the main pair of dorsolateral teeth; pleonite 3 with large subacute mid-dorsal tooth bearing a shallow but long median concavity. COXAE 1 – 3. Carinate and distally sharp. COXA 4. Anterodorsal border proximally nearly straight (inconspicuously convex) and unusually long, of curvature gradually increasing downwards and followed by anteroventral border without any angular discontinuity or visible transition; posteroventral border weakly convex and short; coxa scarcely projecting forward; ventral tooth narrowly triangular, not long, apically subacute; lateral carina without tooth or angularity, not projecting laterally, carina very close from margin of coxa at its deepest point. COXA 5. Without laterally projecting carina (hence scarcely visible in dorsal view), with posteroventral corner produced into a blunt tooth oriented backwards. COXA 6. With mid-sized, blunt, triangular, lateral tooth pointing obliquely backwards (scarcely projecting laterally); posteroventral corner produced into a rounded lobe pointing downwards (posterior margin nearly straight). COXA 7. With ventral border distinctly curved, with posterior border, their convergence forming a blunt angular discontinuity; surface of coxa posteriorly without low carina oriented in the dorsoventral axis. EPIMERAL PLATES 1 – 3. Posteroventral angle produced into a small but sharp tooth. UROSOME TOOTH PATTERN. Urosomite 1 with very sharp narrow triangular process pointing upwards; urosomite 2 without pair of small posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.25; tips of lobes subacute, slit fairly broad and V-shaped. PEDUNCLE OF ANTENNA 1. Article 1 with short lateral medial and ventral teeth reaching the base of article 2 [some teeth apparently duplicated]; article 2 with medium-sized lateral and medial teeth reaching about tip of article 3, with ventral tooth very reduced to a denticle [some teeth apparently duplicated]; article 3 with small ventral tooth, about as long as article itself. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; propodus not narrowing distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus very stout; basis of pereiopods 5 – 6 of normal width, with posteroproximal process rounded and distinctly protruding, with posterodistal tooth strong and broad; basis of pereiopod 7 very broad with posterodistal tooth forming a squared angle, not followed more proximally by small concavity, directed posteriorly. Body length 19 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB8683FFE5BFA8ECEC5FEFE.taxon	distribution	Distribution RV Polarstern, PS 12, stn 231, Elephant Island. Coleman (1998 b) gives the following coordinates for station 231: 61 ° 03.8 ʹ S, 54 ° 37.6 ʹ W, 332 m, and Fütterer (1988) the following ones: 61 ° 03 ʹ S, 54 ° 45 ʹ W (start position), 331 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFB8683FFE5BFA8ECEC5FEFE.taxon	discussion	Remarks Superficially similar to the sympatric and larger E. leukhoplites sp. nov. See account on E. leukhoplites sp. nov. for differences.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC6683FFDE4FE77CEC8FD6A.taxon	distribution	Distribution Ross Sea, 151 – 300 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC6683FFDE4FE77CEC8FD6A.taxon	discussion	Remarks This undescribed species, which is similar to E. leukhoplites sp. nov. (see key) was discovered in the Ross Sea by A. - N. Lörz. It will be described as new in a forthcoming paper by M. Verheye, A. - N. Lörz & C. d’Udekem d’Acoz.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC6683FFE4BFCE3CE0EFB4F.taxon	distribution	Distribution South Shetland Islands: Clarence Island, 0 – 145 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC6683FFE4BFCE3CE0EFB4F.taxon	discussion	Remark The description of Epimeria sp. by Andres (1985) is based on two tiny (7 mm) male specimens. It exhibits similarities with E. leukhoplites sp. nov., E. reoproi and E. vaderi, of which the description is based on much larger specimens. It is likely that Andres’ (1985) specimens are juveniles of one of these species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC6683DFDDEFB06CCAFFC79.taxon	type_taxon	Type species Epimeria tricristata Costa in Hope, 1851 (= Gammarus corniger Fabricius, 1779). Good illustrated descriptions: G. O. Sars (1893); Chevreux & Fage (1925); Lincoln (1979); Ledoyer (1993).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC6683DFDDEFB06CCAFFC79.taxon	description	Description Body opaque, with teguments strongly calcified. Rostrum medium-sized to long. Ventral lobe of head rounded to angular. Eyes present, not conical. Pleonites 1 – 3 and at least pereionites 6 – 7 (sometimes all pereionites) with low mid-dorsal dentate carina projecting backwards and one pair of dorsolateral much smaller carinate teeth (sometimes duplicate or triplicate on pleonites) or carina. Pereionites 1 – 7 sometimes with low carina or trace of protrusion just above connection with coxa. Coxae 1 – 4 subacute to blunt-tipped. Coxae 1 – 3 sharply keeled along their axis. Coxa 4 with sharp carina starting at ventral tip of coxa and terminating at posterior tip of coxa; this never bears a tooth projecting laterally; posteroventral border concave. Coxae 5 – 6 with strong sharp tooth projecting backwards. Mid of posterior border of epimeral plates 1 – 3 or 1 – 2 produced into a tooth. Dorsal process of urosomite 1 produced into a tooth, which can be directed upwards or backwards. Urosomite 2 without pair of small teeth pointing upwards. Lateral borders of urosomite 3 posteriorly blunt or bluntly angular, not produced into a sharp tooth. Telson weekly bilobed. Eyes present, large, rounded or pyriform. Peduncle of antenna 1 without teeth. Mandible with molar process triturative. Lower lip with narrow (V-shaped) hypopharyngeal gap. Palp of maxilliped with 4 articles. Gnathopods of normal size, with carpus and propodus of medium slenderness, with palm obliquely transverse (gnathopods subcheliform); propodus not expanded distally; posterior border of dactylus lined by row of small oblique slender teeth. Basis of pereiopods 5 very slender, quadrate, without posteroproximal protrusion and with trace of posterodistal tooth projecting posteriorly (or posterodistal corner angulate). Basis of pereiopod 6 fairly slender, elliptic or narrowly ovate (then broad part proximal), without posteroproximal rounded protrusion and without posterodistal tooth projecting posteriorly, without posterodistal angle: the basis is distally tapering and at its apex it has the same width as the ischium, hence there is not projecting angularity. basis of pereiopod 7 fairly narrow to fairly broad, with posterior border proximally slightly convex and distally slightly concave, with posterodistal corner rounded or with a tooth projecting posteriorly. Dactylus of pereiopods 5 – 7 short. Benthic. Body length The maximum body length recorded in species of the subgenus Epimeria ranges between 9 and 40 mm (G. O. Sars 1893).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC6683DFDDEFB06CCAFFC79.taxon	biology_ecology	Ecology. Benthic, 50 m (Lincoln 1979) to 3028 m (Jones et al. 2003).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC6683DFDDEFB06CCAFFC79.taxon	distribution	Distribution European western and northern seas, Mediterranean and South Africa.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC6683DFDDEFB06CCAFFC79.taxon	discussion	Remarks As eight Antarctic and sub-Antarctic subgenera are erected herein for Epimeria, the extralimital (Atlantic) type subgenus Epimeria had to be redefined. It is cautiously restricted herein to the following species: Epimeria cornigera (Fabricius, 1779), Epimeria loricata G. O. Sars, 1879, Epimeria parasitica (M. Sars, 1858), Epimeria tuberculata G. O. Sars, 1893, an undescribed Northwest European species discovered by Beerman & Raupach (2015), and an undescribed Mediterranean deepsea species, of which specimens are deposited in the Museum of Verona (J. Beerman, pers. com.).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC4683CFDB2FBFECA61F854.taxon	type_taxon	Type species Epimeriella macronyx Walker, 1906.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC4683CFDB2FBFECA61F854.taxon	description	Description Body pellucid with teguments very weakly calcified. Rostrum minute (not reaching 0.2 of length of article 1 of peduncle of antenna 1), reduced to a narrow tiny blade-shaped process separating the bases of the first articles. Ventral lobe of head rounded. Eyes present, extremely large, not conical. Pereionite 7 and pleonites 1 – 3 with or without small posterodorsal tooth directed backwards; all pereionites and pleonites without dorsolateral pair of teeth, swelling or carina. Pereionites 1 – 7 without tooth or protrusion just above connection with coxa. Coxae 1 – 3 with rounded tip; coxa 4 with tip of variable sharpness. Coxae 1 – 3 not keeled or not strongly keeled along their axis. Coxa 4 narrow, not carinate; posteroventral border straight or very weakly concave. Coxae 5 – 6 without tooth. Mid of posterior border of epimeral plates 1 – 3 not produced into a tooth. Posteroventral tooth of epimeral plate 3 very small. Dorsal process of urosomite 1 produced into a low rounded process or into a carina posteriorly terminated in tooth directed backwards. Urosomite 2 without pair of small teeth pointing upwards. Lateral borders of urosomite 3 posteriorly terminated into an acute angle. Telson deeply cleft. Peduncle of antenna 1 with dentition very reduced (tiny ventral denticles may be present). Mandible with molar process drawn out, without triturative surface. Lower lip with wide hypopharyngeal gap. Palp of maxilliped with 4 articles. Gnathopods of normal size, with carpus and propodus broad to very broad, with palm obliquely transverse (gnathopods subcheliform); propodus not expanded distally; posterior border of dactylus lined by row of small oblique slender teeth. Basis of pereiopods 5 – 6 moderately broad, without posteroproximal protrusion or tooth and without posterodistal tooth projecting posteriorly. Posterior border of basis of pereiopod 7 convex along all its length, with tip angular or rounded. Dactylus of pereiopods 5 – 7 long to very long. Body length The maximum body length recorded in Epimeriella species ranges between 8 and 28 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC4683CFDB2FBFECA61F854.taxon	biology_ecology	Ecology Epimeria (Epimeriella) macronyx is obviously pelagic, as it is usually found in plankton nets. The fragile and poorly caclified morphology of other species suggest that they might also be pelagic or semipelagic. 0 – 1200 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC4683CFDB2FBFECA61F854.taxon	distribution	Distribution Circum-Antarctic, as far north as the South Orkney Islands.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC4683CFDB2FBFECA61F854.taxon	discussion	Remarks Epimeriella was initially erected as a genus (Walker 1906), on the basis of the non triturative molar process of the mandibles. This character is considered as a synapomorphy since Epimeriella sensu Walker, 1906 forms a clade in our phylogenetic trees. It is herein split into two subgenera corresponding to two sub-clades with very different morphotypes. The robust benthic forms of the group walkeri are now placed into the new subgenus Laevepimeria subgen. nov. Only the slender pelagic or semipelagic forms (Epimeria macronyx, E. scabrosa and their relatives) are retained within the subgenus Epimeriella. The extralimital species Epimeria pelagica Birstein & Vinogradov, 1958 also exhibits adaptations to a pelagic lifestyle: enlarged eyes and elongation of the posterior pereiopods (Birstein & Vinogradov 1958). However, this species exhibits many other characters not observed in Epimeriella species such as the shape of the mandible.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC2683AFDD0FEE3CEE3FB15.taxon	description	urn: lsid: zoobank. org: act: B 0214 FB 1 - ED 96 - 4 CEF-B 267 - 21 FC 4 B 356 C 39 Figs 125 – 130	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC2683AFDD0FEE3CEE3FB15.taxon	etymology	Etymology Ἀταλάντη is a character of the Greek mythology. She was the only woman, who took part in the nautical expedition of the Argonauts (the genus Epimeria being feminine). The name alludes to the presumed good swimming capacities of the species and its possibly pelagic life style. The name is a noun in apposition.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC2683AFDD0FEE3CEE3FB15.taxon	materials_examined	Type material Holotype RV Polarstern: SOUTHERN OCEAN: specimen initially fixated in formalin, cruise PS 69, ANT-XXIII / 8, stn 725 - 10, south of Larsen A, 64 ° 55.89 ʹ S, 60 ° 40.06 ʹ W to 64 ° 55.92 ʹ S, 60 ° 40.31 ʹ W, 189 – 192 m, Rauschert dredge, 22 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122527).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC2683AFDD0FEE3CEE3FB15.taxon	description	Description ROSTRUM. Very minute, reduced to a narrow tiny process reaching a bit more than 0.1 of article 1 of peduncle of antenna 1. EYE. Very large, elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 7 and pleonites 1 – 3 carinate and posterodorsally produced into a tooth; other body segments without mid-dorsal tooth; pereionites 1 – 7 and pleonites 1 – 3 without pair of lateral teeth. COXAE 1 – 3. Not carinate; coxa 1 distally broadly rounded; coxa 2 – 3 distally narrow but very blunt. COXA 4. Fairly broad; anterodorsal and anteroventral border nearly straight, joined by low and blunt angular discontinuity, anterior angle not strongly projecting forward; ventral projection short, bluntly but distinctly angular; no lateral carina; posteroventral border nearly straight. COXA 5 – 7. Toothless, posteriorly, ventrally and posteroventrally rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle produced into a tiny tooth. UROSOME TOOTH PATTERN. Urosomite 1 with elongate carinate process, forming a broad and regularly rounded process in lateral view. TELSON. Cleft on 0.3; tips of lobes sharp, notch broadly V-shaped. GNATHOPODS 1 – 2. Carpus and propodus short and broad; propodus not narrowing distally, palm very distinct, oblique. PEREIOPODS 5 – 7. Propodus and dactylus of pereiopods 5 – 6 very long and very slender; basis of pereiopod 5 of normal width, without posteroproximal process, with posterior border nearly straight on most of its length, posterodistally produced into a rounded lobe; basis of pereiopods 6 broad, without posteroproximal process, with posterior border rounded on proximal half and straight on distal half, posterodistally forming a rounded angle; basis of pereiopod 7 extremely broad with posterodistal corner forming a sharp squared angle, not followed more proximally by small concavity. Colour pattern Body and appendages (including gnathopods and mouthparts) whitish / pellucid studded with red chromatophores arranged in large pigmented zones. Eyes reddish. Body length 15 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC2683AFDD0FEE3CEE3FB15.taxon	distribution	Distribution Western side of the Weddell Sea: Larsen A, 189 – 192 m; eastern shelf of the Weddell Sea, 402 – 405 m (see remarks).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC2683AFDD0FEE3CEE3FB15.taxon	discussion	Remarks Epimeria atalanta sp. nov. is very similar to E. scabrosa, but differs by the presence of a posterodorsal tooth on pereionite 7 and its broader eyes (see illustrations by K. H. Barnard 1930 and Coleman 2007). Rauschert & Arntz (2015) give a photograph of a specimen that we identify as E. atalanta sp. nov. The station was given in an early draft of their book made available to us: ANT-XXI / 2 stn 145 [145 - 1]. Its coordinates are: ANT-XXI / 2 stn 145 - 1, 70 ° 56.99 ʹ S, 10 ° 48.26 ʹ W to 70 ° 56.97 ʹ S, 10 ° 47.71 ʹ W (eastern Weddell Sea), 402 – 405 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC36839FE05FB2CCA07FCDA.taxon	description	Fig 131	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC36839FE05FB2CCA07FCDA.taxon	materials_examined	Material examined RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., cruise PS 81, ANT-XXIX / 3, 182 - 1, south east of Snow Hill Island, 64 ° 47.21 ʹ S, 56 ° 41.90 ʹ W to 64 ° 46.02 ʹ S, 56 ° 42.50 ʹ W, 0 – 200 m, rectangular mid-water trawl, formalinfixed, 17 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 121486); 2 specs, cruise PS 81, ANT-XXIX / 3, stn 238 - 1, north of Livingstone Island, 62 ° 22.65 ʹ S, 61 ° 17.63 ʹ W to 62 ° 20.89 ʹ S, 61 ° 19.82 ʹ W, 0 – 200 m, rectangular midwater trawl, 8 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122943) [extraction ANT 38 (largest specimen); Genbank nr, COI: KU 870824, 28 S: KU 759597 and ANT 39 (smallest specimen); Genbank nr, COI: KU 870825, 28 S: KU 759598]. RV Aurora Australis cruises: SOUTHERN OCEAN: 1 spec., cruise CEAMARC, sample CEAMARC 1295, stn 51 AEV 215, Adélie Coast, 66 ° 44 ʹ 52 ″ S, 145 ° 26 ʹ 40 ″ E, 525 – 553 m, beam trawl, 30 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2016 - 6559); 1 spec., cruise CEAMARC, sample CEAMARC 1317, stn 51 AEV 215, Adélie Coast, 66 ° 44 ʹ 52 ″ S, 145 ° 26 ʹ 40 ″ E, 525 – 553 m, beam trawl, 30 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4276) [extraction M 19); Genbank nr, COI: KU 870885, 28 S: KU 759668].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC36839FE05FB2CCA07FCDA.taxon	description	Colour pattern Body and appendages whitish to colourless, semi-translucent; dorsal part of body orange; eyes orange; oral field and tip of gnathopods blood red. Body length Up to 28 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC36839FE05FB2CCA07FCDA.taxon	distribution	Distribution South Orkney Islands; South Shetland Islands; Antarctic Peninsula; eastern shelf of the Weddell Sea; Davis Sea; Adélie Coast, Ross Sea; 0 – 1200 m (Coleman 2007; present data). As many records of E. macronyx come from pelagic nets (De Broyer et al. 2007; present data), and as the species is very slender with large eyes reminiscent to those of hyperiids, it is obviously a true pelagic species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC06839FE24FC96CEB6F98A.taxon	description	Body length Up to 14 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC06839FE24FC96CEB6F98A.taxon	distribution	Distribution Oates Coast, 329 – 366 m (K. H. Barnard 1930).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC06839FE24FC96CEB6F98A.taxon	discussion	Remarks Coleman (pers. com.) informed us that the specimen illustrated in the figure 33 of Coleman (2007) is drawn from the type series. De Broyer et al. (2007) mention imprecise records of E. scabrosa from the southern and eastern shelf of the Weddell Sea, but they might refer to its very close relative E. atalanta sp. nov.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC06839FDF3F9C6CE1EF862.taxon	description	Body length 8 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC06839FDF3F9C6CE1EF862.taxon	distribution	Distribution Palmer Archipelago [type material], eastern shelf of the Weddell Sea, 100 – 622 m (De Broyer et al. 2007).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC16836FDCAFEE3CA60F976.taxon	description	urn: lsid: zoobank. org: act: A 5102 FE 5 - 2 D 1 C- 4365 - BFE 8 - D 5163 A 0748 D 1	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC16836FDCAFEE3CA60F976.taxon	etymology	Etymology Combination of the Greek word οπλα, armour and Epimeria. The name, which is feminine, alludes to the thick and heavily calcified teguments of the members of that subgenus.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC16836FDCAFEE3CA60F976.taxon	type_taxon	Type species Epimeria quasimodo sp. nov.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC16836FDCAFEE3CA60F976.taxon	description	Description Body opaque, with teguments very strongly calcified. Rostrum medium-sized to large. Eyes not conical. Pereionites and pleonites with a wide diversity of ornamentation. Most species have a low and broad mid-dorsal crest on their pleonites and often on their anterior pereionites. However all the pereionites and pleonites can have a very long non-flattened mid-dorsal tooth (E. rubrieques), or the body can be strongly sculptured (E. rimicarinata). Dorsolateral processes are absent except for E. rimicarinata, where they are part of its sculpured pattern. Pereionites 1 – 7 without ventrolateral tooth or protrusion just above connection with coxa. Coxae 1 – 4 with sharp or blunt tip. Coxae 1 – 3 usually weakly keeled along their axis. Coxa 4 with or without groove along the posteroventral border; when present, the anterior border of this groove forms a kind of blunt carina; when present this carina never bears a tooth projecting laterally; posteroventral border straight or concave. Coxa 5 with tooth or blunt protrusion projecting backwards (size and development very variable). Coxa 6 with or without blunt protrusion projecting backwards. Posteroventral tooth of epimeral plate 3 small to medium-sized. Dorsal process of urosomite 1 produced into a blunt tooth or a rounded lobe directed upwards. Urosomite 2 without pair of small teeth pointing upwards. Lateral borders of urosomite 3 posteriorly terminated into an acute angle. Peduncle of antenna 1 with dentition very reduced (a small ventral tooth can be present on article 1). Mandible with molar process triturative. Lower lip with narrow (V-shaped) hypopharyngeal gap. Palp of maxilliped with 4 articles. Gnathopods of normal size, with carpus and propodus very robust, with palm obliquely transverse (gnathopods subcheliform); propodus often expanded distally; posterior border of dactylus lined by row of small oblique slender teeth. Basis of pereiopods 5 – 6 broad, with posteroproximal process, which can be a (very low) rounded protrusion or a large (sometimes swordlike) tooth orientated in the same direction as the basis, with or without posterodistal tooth projecting posteriorly. Basis of pereiopod 7 narrow on its distal half; posterior border proximally slightly convex and distally either slightly concave, or deeply concave or deeply angularly notched; the posterior border of the basis of pereiopod 7 can be terminated into a small tooth projecting posteriorly or not. Dactylus of pereiopods 5 – 7 short. Body length The maximum body length recorded in Hoplepimeria subgen. nov. species ranges between 35 and 80 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC16836FDCAFEE3CA60F976.taxon	biology_ecology	Ecology Benthic, 33 – 2154 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC16836FDCAFEE3CA60F976.taxon	distribution	Distribution Circum-Antarctic, as far north as the South Georgia.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFC16836FDCAFEE3CA60F976.taxon	discussion	Remarks Hoplepimeria subgen. nov. includes species with diverse body shapes. Most species are globular with only low teeth or crests, whilst E. rubrieques is one of the Epimeria with the longest teeth. However, E. rubrieques has the same robust and distally expanded gnathopods as in other Hoplepimeria subgen. nov. and has the same proximally toothed basis of pereiopods 5 – 6 as in many other Hoplepimeria subgen. nov. The absence of dorsolateral teeth, and the non- or scarcely laterally flattened dorsal processes of E. rubrieques distinguishes it at first glance from the dentate Antarctic Epimeria of the subgenus Drakepimeria subgen. nov. Hoplepimeria subgen. nov. includes the largest known Epimeria species, namely E. gargantua sp. nov., which can reach up to 80 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCF6833FED8F8CFCDE4FEC5.taxon	description	Figs 132 – 137	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCF6833FED8F8CFCDE4FEC5.taxon	materials_examined	Material examined RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., exuvia, cruise PS 39, ANT-XIII / 3, EASIZ I, eastern Weddell Sea, no station, specimen kept in aquarium, 26 Mar. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132682); 2 specs, cruise PS 48, ANT-XV / 3, EASIZ II, stn 95, eastern Weddell Sea, 73 ° 33.5 ʹ S, 22 ° 15.3 ʹ W to 73 ° 34.0 ʹ S, 22 ° 12.3 ʹ W, 866 – 920 m, bottom trawl, 5 Feb. 1998, coll. C. De Broyer (RBINS, INV. 13992); 1 spec., cruise PS 48, ANT-XV / 3, EASIZ II, stn 120, eastern Weddell Sea, 73 ° 33.5 ʹ S, 22 ° 14.8 ʹ W to 73 ° 34.0 ʹ S, 22 ° 12.2 ʹ W, 812 – 928 m, bottom trawl, 7 Feb. 1998, coll. C. De Broyer (RBINS, INV. 13993). RV Aurora Australis cruises: SOUTHERN OCEAN: 4 specs including an ovigerous ♀, cruise CEAMARC, sample CEAMARC 3410, stn 86 EEV 518, Adélie Coast, 65 ° 28 ʹ 51 ″ S, 139 ° 24 ʹ 11 ″ E, 781 – 835 m, beam trawl, 16 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4278); 1 dissected ovigerous ♀, cruise CEAMARC, sample CEAMARC 3410, stn 86 EEV 518, Adélie Coast, 65 ° 28 ʹ 51 ″ S, 139 ° 24 ʹ 11 ″ E, 781 – 835 m, beam trawl, 16 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 7343) [removed from MNHN-IU- 2014 - 4278] [extraction M 17; Genbank nr, COI: KU 870883, 18 S: KU 759666]; 1 spec., cruise CEAMARC, sample CEAMARC 2268, stn 32 AEV 400, Adélie Coast, 65 ° 52 ʹ 44 ″ S, 144 ° 10 ʹ 55 ″ E, 953 – 1194 m, beam trawl, 12 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4285); 1 ovigerous ♀, cruise CEAMARC, sample CEAMARC 2271, stn 32 AEV 400, Adélie Coast, 65 ° 52 ʹ 44 ″ S, 144 ° 10 ʹ 55 ″ E, 953 – 1194 m, beam trawl, 12 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4281) [extraction M 18; Genbank nr, COI: KU 870884, 18 S: KU 759667]; 3 specs, cruise CEAMARC, sample CEAMARC 2271, stn 32 AEV 400, Adélie Coast, 65 ° 52 ʹ 44 ″ S, 144 ° 10 ʹ 55 ″ E, 953 – 1194 m, beam trawl, 12 Jan. 2008, coll. IPEV-AAD-MNHN, (MNHN-IU- 2014 - 7344, formerly MNHN-IU- 2014 - 4281).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCF6833FED8F8CFCDE4FEC5.taxon	description	Description ROSTRUM. Medium-sized, reaching tip of article 1 of peduncle of antenna 1, anteriorly distinctly curved, ventrally distinctly concave, subacute in lateral view; narrow and with nearly straight converging borders in frontal view. EYE. Very large, distinctly reniform. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 with trace of mid-dorsal carina and no posterior bump; pereionites 2 to pleonite 3 with low and fairly blunt mid-dorsal carina, which becomes more and more laterally compressed in the posterior body segments; profile of carina of pereionite 2 straight, of pereionites 3 to pleonite 3 with median notch (more distinct on pleonites than on pereionites); profile of carina of pleonite 3 straight with anterior very low protrusion followed by shallow notch, then nearly straight (slightly irregular), posteriorly bluntly angular; dorsolateral ornamentation absent. COXAE 1 – 3. Not carinate and apically forming a blunt squared angle. COXA 4. Anterodorsal border nearly straight (very slightly concave), anteroventral border straight, these two borders being joined by blunt but distinct angular discontinuity (anterior corner), which is weakly projecting forward; ventral corner forming a blunt squared angle (ventral projection well developed); lateral carina absent; posteroventral border distinctly concave. COXA 5. Broad, with surface smooth, with posteroventral corner forming a blunt-tipped nearly squared angle, weakly expanded laterally, forming a blunt obtuse angle in dorsal view. COXA 6. With posteroventral corner forming a blunt acute but nearly squared angle, with low projection arising from its surface (forming a very low indistinct lobe in dorsal view). COXA 7. Posteriorly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle: very obtusely rounded in plate 1; produced into a small tooth in plate 2 and into a medium-sized tooth in plate 3. UROSOME TOOTH PATTERN. Urosomite 1 with very low dorsal process, irregularly convex on both sides; urosomite 3 with dorsolateral borders nearly straight. TELSON. Cleft on 0.2; tips of lobes triangular and subacute, notch broadly V-shaped and rounded at its deepest point. GNATHOPODS 1 – 2. Carpus and propodus broad; propodus expanding distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus of medium width; dactylus medium-sized; basis of pereiopods 5 – 6 of normal width, with posteroproximal process present, sword-like, parallel to axis of basis, with posterodistal corner rounded, very weakly pointing in posterior direction; basis of pereiopod 7 broad with posterior border weakly convex, with distinct notch on distal 0.8, forming a blunt-tipped obtuse (nearly square) angle, with posterodistal corner bluntly angular and not pointing backwards. Colour pattern Uniformly whitish, eyes yellowish (Lörz et al. 2011), or uniformly pale orange, eyes golden (Rauschert & Arntz 2015 as ‘ Epimeria inermis 3 ’, p. 61, pl. 54), or uniformly salmon pink with reddish eyes (picture made by G. Chapelle published herein). Body length Up to 40 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCF6833FED8F8CFCDE4FEC5.taxon	distribution	Distribution Eastern shelf of the Weddell Sea; Adélie Coast; 781 – 1194 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCF6833FED8F8CFCDE4FEC5.taxon	discussion	Remarks The new records of E. angelikae off Adélie Coast considerably extend eastwards the distribution of the species. No morphological differences were detected between these specimens, the illustrations given by Lörz et al. (2011) and the specimens examined from the Weddell Sea. One of the main diagnostic characters proposed by Lörz et al. (2011) for distinguishing E. angelikae from other species of the georgiana complex is the obtusely notched posterior border of the basis of pereiopod 7. This character is valid for adults. However, the depth and the angularity of that notch is a character to be used with caution, as it is partly size-dependent: the notch is shallow in juveniles of all species of the georgiana complex and becomes gradually deeper and more angulate in some species only, as size increases. Epimeria angelikae is superficially similar to E. xesta sp. nov. The two species coexist on the eastern shelf of the Weddell Sea, but have apparently different depth optima. The ‘ Epimeria inermis 3 ’ of Rauschert & Arntz (2015) corresponds to the description of E. angelikae and not to that of E. inermis, as it has a short rostrum, reniform eyes, the posteroventral border of coxa 4 concave and a median spur on the basis of pereiopods 5 and 6. The station of this specimen was given in an early draft of their book made available to the authors: ANT-XXI / 2 stn 233 [233 - 1]. Its coordinates are: 71 ° 18.99 ʹ S, 13 ° 56.56 ʹ W to 71 ° 19.19 ʹ S, 13 ° 57.45 ʹ W (eastern Weddell Sea), 844 – 848 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCA6832FE33FE9DCC79F9F8.taxon	description	urn: lsid: zoobank. org: act: 5 A 5 D 8439 - 3793 - 4124 - A 65 B-C 6 EB 65 A 1587 E Figs 138 – 144	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCA6832FE33FE9DCC79F9F8.taxon	etymology	Etymology From the Greek, κυφός, humpbacked; ραχις, spine, spinal column. The name, which is a noun in apposition, alludes to the humpbacked silhouette of the species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCA6832FE33FE9DCC79F9F8.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: ♀, cruise PS 81, ANT-XXIX / 3, stn. 249 - 2, Drake Passage, north of Livingstone Island, 61 ° 56.05 ʹ S, 60 ° 5.56 ʹ W to 61 ° 56.21 ʹ S, 60 ° 5.80 ʹ W, 413 – 421 m, mud, Agassiz trawl, 12 Mar. 2013 (RBINS, INV. 122936) [extraction K 25; Genbank nr, COI: KU 870859, 28 S: KU 759637].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCA6832FE33FE9DCC79F9F8.taxon	description	Description ROSTRUM. Medium-sized, overreaching tip of article 1 of peduncle of antenna 1, anteriorly weakly curved, ventrally straight, acute in lateral view; broad and with weakly curved converging borders in frontal view. EYE. Very large, broadly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 3 smooth, pereionites 4 to pleonite 3 with extremely low and extremely blunt mid-dorsal carina; profile of carina of pereionite 4 inconspicuously curved; profile of carinae of pereionites 5 to pleonites 3 straight, those of pleonite 2 with inconspicuous concavity; profile of carina of pleonite 3 straight with anterior low very protrusion followed by shallow notch, then nearly straight, posteriorly blunt; dorsolateral ornamentation absent. COXAE 1 – 3. Weakly carinate and distally subacute to narrowly blunt-tipped. COXA 4. Anterodorsal border nearly straight (very slightly concave), anteroventral border very weakly concave, these two borders being joined by a very large rounded lobe (anterior corner), which is weakly projecting forward; ventral corner forming a sharp squared angle (ventral projection well developed); lateral carina very obtuse, parallel and close to posteroventral border; posteroventral border distinctly concave. COXA 5. Broad, with surface smooth, with posteroventral corner forming a blunt-tipped nearly squared angle, weakly laterally produced, forming a low very rounded obtuse angle in dorsal view (coxa 5 distinctly visible in dorsal view). COXA 6. With posteroventral corner broadly rounded, with low projection arising from its surface (forming a distinct lobe in dorsal view). COXA 7. Posteriorly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle: very obtusely rounded, with indistinct trace of tooth in plate 1; produced into a small tooth in plate 2 and into a medium-sized tooth in plate 3. UROSOME TOOTH PATTERN. Urosomite 1 with very low dorsal process: anteriorly nearly straight, apically very blunt, posteriorly regularly convex large; urosomite 3 with dorsolateral borders nearly straight. TELSON. Cleft on 0.2; tips of lobes bluntly triangular, notch narrowly V-shaped. GNATHOPODS 1 – 2. Carpus and propodus broad; propodus expanding distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus of medium width; dactylus long; basis of pereiopods 5 – 6 of normal width, with posteroproximal process present, triangular (pereiopod 5) or sword-like (pereiopod 6), parallel to axis of basis, with posterodistal corner obtuse, scarcely projecting posteriorly; basis of pereiopod 7 broad with posterior border distinctly convex, with deep notch on distal 0.8, forming a blunt-tipped squared angle, with posterodistal corner obtuse, scarcely projecting posteriorly. Colour pattern Red with a diffuse finely mottled pattern; some parts of the coxae and epimeral plates are white; eyes red. Body length Up to 42 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCA6832FE33FE9DCC79F9F8.taxon	distribution	Distribution South Shetland Islands: north of Livingstone Island, 413 – 421 m (present material); south of Elephant Island, 463 – 990 m (Lörz et al. 2011 as Epimeria georgiana Clade A).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCA6832FE33FE9DCC79F9F8.taxon	discussion	Remarks Epimeria cyphorachis sp. nov. can be easily distinguished from the similar sympatric species E. quasimodo sp. nov. by the shape of the anterior corner of coxa 4: it is broadly rounded in the first species, angulate in the second. Actually, E. cyphorachis sp. nov. (South Shetland Islands) is morphologically more similar to E. xesta sp. nov. (eastern shelf of the Weddell Sea). In E. xesta sp. nov., the rostrum is narrower, the ventral corner of coxa 4 is more blunt, the coxa 5 less projecting laterally and the notch of the basis of pereiopods 5 – 7 is narrower than in E. cyphorachis sp. nov.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCB682FFE38F970CDA2FD78.taxon	description	urn: lsid: zoobank. org: act: E 5 FDFA 56 - F 77 B- 46 D 9 - 9665 - 644728 A 7970 C Figs 145 – 151	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCB682FFE38F970CDA2FD78.taxon	etymology	Etymology Gargantua is a giant and one of the main characters in the tales of François Rabelais, such as ‘ La vie très horrifique du grand Gargantua, père de Pantagruel’. The name, which is a noun in apposition, alludes to the huge size of the species, which is the largest known Epimeria species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCB682FFE38F970CDA2FD78.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: ♀, cruise PS 81, ANT-XXIX / 3, Bransfield Strait, stn 196 - 8, 62 ° 47.80 ʹ S, 57 ° 5.35 ʹ W to 62 ° 47.63 ʹ S, 57 ° 5.63 ʹ W, 542 – 580 m, trawl haul with huge stones and a lot of life, Agassiz trawl, 24 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122937 A) [extraction ANT 33; Genbank nr, COI: KU 870820, 28 S: KU 759592]. Paratypes RV Polarstern cruises: SOUTHERN OCEAN: 1 medium-sized spec., cruise PS 69, ANT-XXIII / 8, stn 662 - 1, between Elephant Island and King George Island, 61 ° 35.91 ʹ S, 57 ° 17.04 ʹ W to 61 ° 35.41 ʹ S, 57 ° 20.60 ʹ W, 425 – 432 m, bottom trawl, 30 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122491); 2 large specs, cruise PS 69, ANT-XXIII / 8, stn 663 - 1, northeast of King George Island, 61 ° 38.18 ʹ S, 57 ° 33.17 ʹ W to 61 ° 38.02 ʹ S, 57 ° 37.16 ʹ W, bottom trawl, 432 – 434 m, 30 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122494); 5 specs, cruise PS 81, ANT-XXIX / 3, Bransfield Strait, stn 196 - 8, 62 ° 47.80 ʹ S, 57 ° 5.35 ʹ W to 62 ° 47.63 ʹ S, 57 ° 5.63 ʹ W, 542 – 580 m, trawl haul with huge stones and a lot of life, Agassiz trawl, 24 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122937 B); 1 spec., cruise PS 81, ANT-XXIX / 3, Bransfield Strait, stn 217 - 6, 62 ° 53.45 ʹ S, 58 ° 13.06 ʹ W to 62 ° 53.42 ʹ S, 58 ° 13.41 ʹ W, 461 – 483 m, rich sponge bottom, Agassiz trawl, 2 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122927) [extraction K 37; Genbank nr, COI: KU 870869, 28 S: KU 759649]; 1 spec., cruise PS 81, ANT-XXIX / 3, Bransfield Strait, stn 217 - 6, 62 ° 53.45 ʹ S, 58 ° 13.06 ʹ W to 62 ° 53.42 ʹ S, 58 ° 13.41 ʹ W, 461 – 483 m, rich sponge bottom, Agassiz trawl, 2 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122928) [extraction K 38; Genbank nr, COI: KU 870870, 28 S: KU 759650]; 7 specs, cruise PS 81, ANT-XXIX / 3, Bransfield Strait, stn 217 - 6, 62 ° 53.45 ʹ S, 58 ° 13.06 ʹ W to 62 ° 53.42 ʹ S, 58 ° 13.41 ʹ W, 461 – 483 m, rich sponge bottom, Agassiz trawl, 2 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 132957); 1 spec., cruise PS 81, ANT-XXIX / 3, Bransfield Strait, stn 217 - 6, 62 ° 53.45 ʹ S, 58 ° 13.06 ʹ W to 62 ° 53.42 ʹ S, 58 ° 13.41 ʹ W, 461 – 483 m, rich sponge bottom, Agassiz trawl, 2 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (MNHN-IU- 2014 - 7333, removed from RBINS, INV. 132957); 1 spec., cruise PS 81, ANT-XXIX / 3, Bransfield Strait, stn 227 - 2, 62 ° 55.83 ʹ S, 58 ° 41.09 ʹ W to 62 ° 55.76 ʹ S, 58 ° 41.46 ʹ W, 562 – 564 m, mud, Agassiz trawl, 5 Mar. 2013, RBINS, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122939) [extraction ANT 40; Genbank nr, COI: KU 870826, 28 S: KU 759599].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCB682FFE38F970CDA2FD78.taxon	description	Description ROSTRUM. Medium-sized, not reaching tip of article 1 of peduncle of antenna 1, anteriorly distinctly and regularly curved, ventrally straight, fairly narrow and subacute in lateral view; fairly narrow and with very weakly convex converging borders in frontal view. EYE. Very large, broadly elliptic to more or less reniform. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 7 smooth; pleonite 1 dorsally weakly carinate, very weakly convex, with distinct posterior bump; pleonite 2 dorsally distinctly carinate, with extremely low (nearly inconspicuous) proximal rounded lobe followed by extremely weak (nearly inconspicuous) concavity, posteriorly produced into a bluntly triangular tooth projecting backwards; pleonite 3 dorsally distinctly carinate with median very low rounded lobe, followed by distinct concavity, terminated by a blunt tooth directed upwards. COXAE 1 – 3. Not carinate, apically subacute. COXA 4. Anterodorsal border nearly straight (inconspicuously concave), anteroventral border straight, these two borders being joined by blunt but broad, very distinct squared angle (anterior corner), which is slightly projecting forward; ventral corner forming a very obtuse angle (ventral projection very short and very broad); lateral carina absent; posteroventral border nearly straight (inconspicuously convex). COXA 5. Broad, with surface smooth, with posterior border inconspicuously concave (nearly straight), with posteroventral corner forming a blunt tooth (shape: acute triangle) projecting backwards and not laterally (no tooth or corner visible in dorsal view). COXA 6. With posterior border very weakly concave, with posteroventral corner forming a blunt tooth (shape: narrow acute triangle) projecting backwards and not laterally (no tooth or corner visible in dorsal view). COXA 7. Posteriorly weakly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle: angulate in plate 1, produced into a medium-sized tooth in plates 2 – 3. UROSOME TOOTH PATTERN. Urosomite 1 with well developed blunt-tipped process of which both the anterior and the posterior borders have an angulate concavity (the anterior deeper); urosomite 3 with dorsolateral borders weakly concave and posteriorly produced into a sharp triangular tooth. TELSON. Cleft nearly on 0.2; tips of lobes triangular and subacute, notch very broadly V-shaped and subacute at its deepest point. GNATHOPODS 1 – 2. Carpus and propodus very broad; propodus expanding distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus fairly broad; dactylus small, very curved, with long unguis; basis of pereiopods 5 – 6 broad, with posteroproximal process present, sword-like, projecting obliquely, with posterodistal corner produced into a triangular tooth (with tip subacute), projecting backwards; basis of pereiopod 7 broad; posterior border with proximal 0.4 with weak concavity, with distal 0.6 deeply concave, with posterodistal corner forming a narrowly triangular tooth projecting backwards. Colour pattern Body and coxae pure white, gnathopods and oral field purplish; antennae and pereiopods 3 – 4 pale pink; pereiopods 5 – 7 and tailfan pure white; eyes red. This colour pattern was very consistent in all specimens examined during ANT-XXIX / 3; none had spots or marks on their immaculate body. Body length Up to 80 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCB682FFE38F970CDA2FD78.taxon	distribution	Distribution Between Elephant Island and King George Island; northeast of King George Island, Bransfield Strait; 404 – 580 m (present material; Coleman 1994: “ Epimeria robusta from Elephant Island ” (specimen actually collected between Elephant and King George Islands )).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFCB682FFE38F970CDA2FD78.taxon	discussion	Remarks Epimeria gargantua sp. nov. is the largest known Epimeria species, both in length and body volume. Epimeria gargantua sp. nov. (tip of Antarctic Peninsula) is morphologically similar to E. robustoides (eastern shelf of the Weddell Sea), but these two species were identified as separate species by methods based on COI and 28 S genes (Fig. 342). In E. gargantua sp. nov., the dorsolateral margins of urosomite 3 are less concave and the posterodistal corner of the basis of pereiopods 5 – 7 sharper than in E. robustoides. The colour pattern of E. gargantua sp. nov. is very constant (body white, without coloured marks), whilst it is more variable in E. robustoides.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD6682EFE89FCF2CD31F81C.taxon	description	Figs 152 – 158	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD6682EFE89FCF2CD31F81C.taxon	materials_examined	Type material Lectotype Swedish South Polar Expedition 1901 – 1903: SOUTHERN OCEAN: ♀, from the syntype series, stn 34, South Georgia, in front of mouth of Cumberland Bay, 54 ° 11 ʹ S, 36 ° 18 ʹ W, grey clay, some stones, 252 – 310 m, T = + 1.45 ° C, Skottsberg pinx. (SMNH - type 673).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD6682EFE89FCF2CD31F81C.taxon	description	Description ROSTRUM. Medium-sized, just overreaching tip of article 1 of peduncle of antenna 1, anteriorly distinctly curved, ventrally straight, subacute in lateral view; broad and with proximally nearly straight converging borders in frontal view. EYE. Fairly large, broadly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 smooth; pereionites 2 – 3 with weak posterior bump; pereionites 4 to pleonite 3 with low, non-toothed, very broad carina (not reduced on pereionites 2 – 3), those of pleonites 1 – 3 with trace of concavity in anterior 0.4; posterodorsal angle of pleonite 3 very bluntly angular; dorsolateral ornamentation absent. COXAE 1 – 3. Scarcely carinate and apically tapering and blunt. COXA 4. Anterodorsal border weakly sigmoid, anteroventral border distinctly concave, these two borders being joined by very distinct and not so blunt angle (anterior corner), which is strongly projecting forward; ventral corner forming a sharp acute angle (ventral projection short); lateral carina present, not sharp; broad hollow surface between carina and posteroventral border of coxa; posteroventral border distinctly concave. COXA 5. Broad, with surface smooth, with posteroventral corner forming a rounded lobe, weakly expanded laterally, forming a blunt squared angle which is very distinct in dorsal view. COXA 6. With posteroventral corner rounded, with lateral projection arising from its surface (broadly triangular in dorsal view). COXA 7. Posteriorly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle: forming an angle with a trace of tooth in plate 1, produced into a small tooth in plate 2 and into a medium-sized tooth in plate 3. UROSOME TOOTH PATTERN. Urosomite 1 with very low dorsal process, regularly convex on both sides; urosomite 3 with dorsolateral borders nearly straight (inconspicuously convex), with tip bluntly angular. TELSON. Cleft on 0.4; lobes with rate of divergence abruptly increasing on their distal half, triangular and subacute in their distal half, median notch narrowly V-shaped. GNATHOPODS 1 – 2. Carpus and propodus very broad; propodus expanding distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus of medium width; dactylus medium-sized; basis of pereiopods 5 – 6 very broad, with posteroproximal process present, tooth-like, parallel to axis of basis, with posterodistal corner bluntly angular, distinctly produced and pointing in posterior direction; basis of pereiopod 7 broad with posterior border weakly convex, with distinct notch on distal 0.8, forming a blunt-tipped acute angle, with posterodistal corner bluntly angular and not projecting backwards. Body length Up to 40 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD6682EFE89FCF2CD31F81C.taxon	distribution	Distribution South Georgia, 75 – 310 m (Schellenberg 1931).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD6682EFE89FCF2CD31F81C.taxon	discussion	Remarks Epimeria georgiana s. str. is considered endemic to South Georgia and all extralimital records are presumably based on related species. The mention ‘ Skottsberg pinx. ’ (i. e., Skottsberg pinxit) on one of the labels of the lectotype suggests that Carl Johan Skottsberg made a painting of it during the Swedish Antarctic Expedition. Nothing is known about the fate of that painting. Besides the adult female examined and designated herein as the lectotype, Schellenberg (1931) recorded a mid-sized (20 mm) specimen and three juveniles (7 – 8 mm) of Epimeria georgiana, which become paralectotypes. These specimens were not examined.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD4682CFE42FEE3CCF0FC4C.taxon	description	Description Description based on illustrations of Coleman (1998 a). ROSTRUM. Medium-sized, just overreaching tip of article 1 of peduncle of antenna 1, anteriorly weakly curved, ventrally straight, tip subacute in lateral view. EYE. Very large, narrowly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 5 smooth; pereionite 6 with weak mid-dorsal straight carina, pereionite 7 to pleonite 3 with low carina, each with median slight concavity and posterior corner bluntly angular; the carinae are strongly laterally compressed. COXAE 1 – 3. Scarcely carinate and apically broadly rounded. COXA 4. Anterodorsal border nearly straight (very slightly concave), anteroventral border weakly but distinctly concave, these two borders being joined by broad rounded angle (anterior corner), which is distinctly projecting forward; ventral corner forming a fairly narrow rounded projection; lateral carina present and projecting laterally (it appears as a low rounded lobe in dorsal view); at its deepest point the carina is very distant from the posteroventral border of the coxa; posteroventral border deeply concave. COXA 5. Broad, with rounded lateral projection pointing laterally. In dorsal view, the lateral projection of coxa 4 and 5 looks like a mirror version of each other, the unit coxa 4 + coxa 5 forming a low concave structure on each side of the body. COXA 6. Posteriorly broadly rounded, with very low projection arising from its surface (forming a very low rounded lobe in dorsal view). COXA 7. Posterior and ventral border strongly convex; transition between these borders indistinct. EPIMERAL PLATES 1 – 3. Posteroventral angle: rounded in plates 1 – 2, produced into a small tooth in plate 2 and into a medium-sized tooth in plate 3. UROSOME TOOTH PATTERN. Urosomite 1 with low rounded dorsal process; urosomite 3 with dorsolateral borders straight, distally forming a rounded lobe. TELSON. Cleft on 0.15; tips of lobes broadly rounded, notch broadly V-shaped. GNATHOPODS 1 – 2. Carpus and propodus broad; propodus not expanding distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus slender, dactylus medium-sized; basis of pereiopods 5 – 6 of normal width, with posteroproximal half forming a very low rounded projection; with posterodistal corner not produced; basis of pereiopod 7 narrow with posterior border straight, with posterodistal corner very bluntly angular, not projecting. Colour pattern Body and coxae whitish with a very faint dull purplish hue. Basis of pereiopods 5 – 7 pale dull purple. Antennae, other pereiopods, distal part of pereiopods 5 – 7 and tailfan dull brownish pink. Eyes pale yellow. Body length 52 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD4682CFE42FEE3CCF0FC4C.taxon	distribution	Distribution Northwest of Elephant Island. Said to have been collected at 60 ° 54.60 ʹ S, 55 ° 45.90 ʹ W at 235 m depth (RV Polarstern, PS 42, ANT-XIV / 2, stn 31) (Coleman 1998 a). The coordinates of station 31 given by Kattner (1998) are slightly different and deeper: 60 ° 53 ʹ S, 55 ° 47 ʹ W to 60 ° 54 ʹ S, 55 ° 57 ʹ W, 443 – 734 m. However, Coleman (pers. com.) informed us that the depth record given in his logbook is 230 – 235 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD4682CFE42FEE3CCF0FC4C.taxon	discussion	Remarks This highly characteristic species, which shares a number of character states with Epimeria georgiana and E. inermis, is presumably rare, as it was not found in the abundant Epimeria material from Elephant Island collected during the cruise ANT-XXIII / 8. Epimeria heldi is tentatively assigned to the subgenus Hoplepimeria subgen. nov. based on morphological assessment only.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD56828FE46FC05CD2CFAF7.taxon	description	Figs 159 – 184	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD56828FE46FC05CD2CFAF7.taxon	materials_examined	Type material Holotype British Antarctic Expedition 1898 – 1900, RV Southern Cross: SOUTHERN OCEAN: ♀, Cape Adare, 51 m (28 fathoms) (BMNH 1903.10.5.19). Other material examined RV Polarstern cruises: SOUTHERN OCEAN: 2 specs, previously identified as E. georgiana, cruise PS 39, ANT-XIII / 3, EASIZ I, stn 11, GSN 4, eastern Weddell Sea, 73 ° 22.60 ʹ S, 21 ° 10.60 ʹ W to 73 ° 23.00 ʹ S, 21 ° 12.90 ʹ W, 333 – 338 m, bottom trawl, 13 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132707); 1 juv., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 11, GSN 4, eastern Weddell Sea, 73 ° 22.60 ʹ S, 21 ° 10.60 ʹ W to 73 ° 23.00 ʹ S, 21 ° 12.90 ʹ W, 333 – 338 m, bottom trawl, 13 feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132710); 9 specs, previously mixed with 1 E. xesta sp. nov., misidentified as E. georgiana and used for gut content analysis, cruise PS 39, ANT-XIII / 3, EASIZ I, stn 12, GSN 5, eastern Weddell Sea, 73 ° 18.10 ʹ S, 21 ° 10.10 ʹ W to 73 ° 17.10 ʹ S, 21 ° 08.20 ʹ W, 457 – 459 m, bottom trawl, 14 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132996); 1 spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 17, GSN 10, eastern Weddell Sea, 73 ° 18.00 ʹ S, 21 ° 09.90 ʹ W to 73 ° 19.10 ʹ S, 21 ° 14.90 ʹ W, 465 – 468 m, bottom trawl, 16 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132708); 1 spec., cruise PS 39, ANT-XIII / 3, EASIZ I, eastern Weddell Sea, station lost, specimen kept in aquarium, 27 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132684); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 605 - 1, Elephant Island, 61 ° 20.35 ʹ S, 55 ° 29.16 ʹ W to 61 ° 19.98 ʹ S, 55 ° 32.67 ʹ W, 146 – 151 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122513); 1 large spec., cruise PS 69, ANT-XXIII / 8, stn 608 - 1, Elephant Island, 61 ° 11.34 ʹ S, 54 ° 43.17 ʹ W to 61 ° 11.80 ʹ S, 54 ° 40.05 ʹ W, 284 – 293 m, bottom trawl, 20 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122520); 1 small spec., cruise PS 69, ANT-XXIII / 8, stn 609 - 6, Elephant Island, 61 ° 08.58 ʹ S, 54 ° 31.86 ʹ W to 61 ° 08.58 ʹ S, 54 ° 31.86 ʹ W, 437 – 442 m, Rauschert dredge, 21 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122486); 4 ♀♀, cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122512); 5 ♀♀, cruise PS 69, ANT-XXIII / 8, stn 654 - 6, Elephant Island, 61 ° 22.80 ʹ S, 56 ° 03.84 ʹ W to 61 ° 23.35 ʹ S, 56 ° 04.89 ʹ W, 341 – 342 m, Agassiz trawl, 29 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122511); 6 ♀♀, cruise PS 69, ANT-XXIII / 8, stn 654 - 7, Elephant Island, 61 ° 23.37 ʹ S, 56 ° 04.35 ʹ W to 61 ° 23.94 ʹ S, 56 ° 03.79 ʹ W, 318 – 345 m, Rauschert dredge, 29 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122514); 1 spec., cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, stn 48 - 1, eastern Weddell Sea, 70 ° 23.94 ʹ S, 8 ° 19.14 ʹ W to 70 ° 23.89 ʹ S, 8 ° 18.67 ʹ W, 595 – 602 m, bryozoan bottom (exceptional bryozoan diversity), Rauschert dredge, 12 Jan. 2008, coll. H. Robert (RBINS, INV. 132954) [extraction I 3; Genbank nr, COI: KU 870847, 28 S: KU 759624]; 2 small specs, cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, stn 48 - 1, eastern Weddell Sea, 70 ° 23.94 ʹ S, 8 ° 19.14 ʹ W to 70 ° 23.89 ʹ S, 8 ° 18.67 ʹ W, 595 – 602 m, bryozoan bottom (exceptional bryozoan diversity), Rauschert dredge, 12 Jan. 2008, coll. H. Robert (RBINS, INV. 122897); 1 small spec., cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, stn 48 - 1, eastern Weddell Sea, 70 ° 23.94 ʹ S, 8 ° 19.14 ʹ W to 70 ° 23.89 ʹ S, 8 ° 18.67 ʹ W, 595 – 602 m, bryozoan bottom (exceptional bryozoan diversity), Agassiz trawl, 12 Jan. 2008, coll. H. Robert (RBINS, INV. 122905); 1 large spec., cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, stn 48 - 1, eastern Weddell Sea, 70 ° 23.94 ʹ S, 8 ° 19.14 ʹ W to 70 ° 23.89 ʹ S, 8 ° 18.67 ʹ W, 595 – 602 m, bryozoan bottom (exceptional bryozoan diversity), Agassiz trawl, 12 Jan. 2008, initially fixed in formalin, coll. H. Robert (RBINS, INV. 132459); 1 spec., cruise PS 77, ANT-XXVII / 3, stn 281 - 1, eastern Weddell Sea, BENDEX area, 70 ° 48.93 ʹ S, 10 ° 32.69 ʹ W to 70 ° 49.46 ʹ S, 10 ° 33.66 ʹ W, 283 m, bottom trawl, 28 Mar. 2011, bottom trawl, coll. C. Havermans and H. Robert (RBINS, INV. 132955) [extraction I 11, Genbank nr, 28 S: KU 759613]; 1 ♀, cruise PS 81, ANT-XXIX / 3, stn 199 - 4, Bransfield Strait, 62 ° 57.22 ʹ S, 58 ° 14.60 ʹ W to 62 ° 57.33 ʹ S, 58 ° 14.95 ʹ W, 325 – 339 m, non-muddy bottom (content of the net dominated by sponges; also small stones, Stylasteridae and crinoids) Agassiz trawl, 27 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 132953) [extraction I 18; Genbank nr, COI: KU 870843, 28 S: KU 759620]; 1 spec., cruise PS 81, ANT-XXIX / 3, stn 224 - 2, Bransfield Strait, 63 ° 0.50 ʹ S, 58 ° 37.53 ʹ W to 63 ° 0.54 ʹ S, 58 ° 37.50 ʹ W, 254 – 256 m, black sand, Rauschert dredge, 4 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122972); 1 ovigerous ♀, cruise PS 81, ANT-XXIX / 3, stn 224 - 3, Bransfield Strait, 63 ° 0.53 ʹ S, 58 ° 35.67 ʹ W to 63 ° 0.58 ʹ S, 58 ° 36.11 ʹ W, 257 – 261 m, mud, Agassiz trawl, 4 Mar. 2013 (RBINS, INV. 122945 A) [extraction ANT 45; Genbank nr, COI: KU 870831, 28 S: KU 759604]; 1 immature spec., cruise PS 81, ANT-XXIX / 3, stn 224 - 3, Bransfield Strait, 63 ° 0.53 ʹ S, 58 ° 35.67 ʹ W to 63 ° 0.58 ʹ S, 58 ° 36.11 ʹ W, 257 – 261 m, mud, Agassiz trawl, 4 Mar. 2013 (RBINS, INV. 122945 B); 1 spec., cruise PS 81, ANT-XXIX / 3, stn 227 - 2, Bransfield Strait, 62 ° 55.83 ʹ S, 58 ° 41.09 ʹ W to 62 ° 55.76 ʹ S, 58 ° 41.46 ʹ W, 562 – 564 m, mud, Agassiz trawl, 5 Mar. 2013 (RBINS, INV. 122948) [extraction ANT 44, Genbank nr, COI: KU 870830, 28 S: KU 759603]. RV Marion Dufresne cruises: SOUTHERN OCEAN: 3 specs, cruise MD 42 (SIBEX), stn 22 - CP 73, Prydz Bay, 66 ° 57 ʹ 25 ″ S, 72 ° 41 ʹ 25 ″ E, depth missing [Google depth for that position: 533 m], 26 Jan. 1985, (MNHN-IU- 2014 - 4261) (typed label). RV Aurora Australis cruises: SOUTHERN OCEAN: 1 ovigerous ♀, cruise CEAMARC, sample CEAMARC 971, stn 42 EV 167, Adélie Coast, 66 ° 53 ʹ 22 ″ S, 142 ° 38 ʹ 54 ″ E, 262 – 431 m, beam trawl, 28 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4272) [extraction M 1; Genbank nr, COI: KU 870877, 28 S: KU 759660]; 1 spec., cruise CEAMARC, sample CEAMARC 1317, stn 51 AEV 215, Adélie Coast, 66 ° 44 ʹ 52 ″ S, 145 ° 26 ʹ 40 ″ E, 525 – 553 m, beam trawl, 30 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4274); 1 ♀, cruise CEAMARC, sample CEAMARC 2724, stn 71 EV 447, Adélie Coast, 66 ° 24 ʹ 00 ″ S, 140 ° 32 ʹ 21 ″ E, 683 – 791 m, beam trawl, 14 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4338) [extraction M 2; Genbank nr, COI: KU 870886, 28 S: KU 759669]; 3 specs, cruise CEAMARC, sample CEAMARC 3226, stn 20 EV 490, Adélie Coast, 66 ° 00 ʹ 59 ″ S, 140 ° 00 ʹ 02 ″ E, 189 – 196 m, beam trawl, 15 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4346); 5 specs, cruise CEAMARC, sample CEAMARC 3532, stn 87 EV 524, Adélie Coast, 65 ° 29 ʹ 29 ″ S, 139 ° 18 ʹ 37 ″ E, 397 – 411 m, beam trawl, 17 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4351); 5 very large specs, cruise CEAMARC, sample CEAMARC 3978, stn 36 EV 297, Adélie Coast, 66 ° 20 ʹ 20 ″ S, 143 ° 41 ʹ 08 ″ E, 552 – 573 m, beam trawl, 4 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4328). RV Seatruck, cruise REVOLTA I: SOUTHERN OCEAN: 1 ovigerous ♀, stn REVO _ 020 b, Collect _ ID: 143, Field _ ID: CE- 000004560, Adélie Coast, 66 ° 40 ʹ 30 ″ S, 139 ° 55 ʹ 05 ″ E, 19 – 23 m, 15 Jan. 2010:, coll. M. Eléaume, L. Hemery and A. D’Hont, (MNHN-IU- 2009 - 2569) [extraction K 3; Genbank nr, 28 S: KU 759642]. RV Seatruck, cruise REVOLTA II: SOUTHERN OCEAN: 2 specs, stn REVO _ 091, Collect _ ID: REVO _ 162, Adélie Coast, 66 ° 41 ʹ 07 ″ S, 139 ° 56 ʹ 41 ″ E, 33 – 34 m, 19 Jan. 2011, coll. N. Améziane, N. Bax, C. Gallut, A. C. Lautrédou and C. Robineau (MNHN-IU- 2014 - 4308). RV L’Astrolabe, cruise REVOLTA III: SOUTHERN OCEAN: 1 spec., stn none (Dumont d’Urville Sea), Collect _ ID: REVO _ 449, Adélie Coast, 66 ° 38 ʹ S, 140 ° 42 ʹ E to 66 ° 38 ʹ S, 140 ° 40 ʹ E, 718 – 729 m, mud, beam trawl, 20 Jan. 2012, coll. G. Lecointre, A. Dettaï, J. Lanshere, C. Gallut and C. Ozouf (MNHN-IU- 2009 - 2545); 1 large spec., stn none (Dumont d’Urville Sea), Collect _ ID: REVO _ 449, Adélie Coast, 66 ° 38 ʹ S, 140 ° 42 ʹ E to 66 ° 38 ʹ S, 140 ° 40 ʹ E, 718 – 729 m, mud, beam trawl, 20 Jan. 2012, coll. G. Lecointre, A. Dettaï, J. Lanshere, C. Gallut and C. Ozouf (MNHN-IU- 2009 - 2584). RV Seatruck, cruise REVOLTA III: SOUTHERN OCEAN: 1 ovigerous ♀, stn REVO _ 032, Collect _ ID: REVO _ 509, Field _ ID: CE- 000002403, Adélie Coast, 66 ° 39.306 ʹ S, 140 ° 01.635 ʹ W to 66 ° 39.370 ʹ S, 140 ° 01.686 ʹ W, 72 – 100 m, beam trawl, 8 Feb. 2012, coll. G. Lecointre, A. Dettaï, J. Lanshere, C. Gallut and C. Ozouf, (MNHN-IU- 2009 - 2531) [extraction K 2; Genbank nr, 28 S: KU 759632].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD56828FE46FC05CD2CFAF7.taxon	description	Description Description based on holotype. ROSTRUM. Medium-sized, reaching mid of article 2 of peduncle of antenna 1, anteriorly strongly curved, ventrally straight, acute-tipped in lateral view; blunt-tipped, broad and with borders scarcely convex in frontal view. EYE. Medium-sized, elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 2 smooth; pereionite 3 with trace of carina and trace of posterodorsal bump; pereionite 4 to pleonite 3 with low and blunt mid-dorsal carina, which is strongly laterally compressed (especially the posterior ones); all these carinae with very weak median concavity, that of pleonite 3 with a second weak concavity and posteriorly forming a blunt squared angle. COXAE 1 – 3. Weakly carinate and apically subacute. COXA 4. Anterodorsal border nearly straight (very slightly concave), anteroventral border very weakly convex, these two borders being joined by blunt but distinct squared angle (anterior corner), which is distinctly projecting forward; ventral corner forming an obtuse angle (ventral projection moderately developed); lateral carina absent; posteroventral border straight. COXA 5. Broad, with surface smooth, with posteroventral corner forming a blunt-tipped projection, weakly expanded laterally, forming a blunt squared angle in dorsal view. COXA 6. With posteroventral corner forming a blunt squared angle, with very low projection arising from its surface (forming a very low indistinct lobe in dorsal view). COXA 7. Posterior and ventral border weakly convex, joined by blunt squared angle. EPIMERAL PLATES 1 – 3. Posteroventral angle: rounded in plate 1, produced into a small tooth in plate 2 and into a medium-sized tooth in plate 3. UROSOME TOOTH PATTERN. Urosomite 1 with very low rounded dorsal process; urosomite 3 with dorsolateral borders distinctly concave. TELSON. Cleft on 0.035; tips of lobes obtusely triangular and very blunt, notch very broadly V-shaped. GNATHOPODS 1 – 2. Carpus and propodus broad; propodus expanding distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus of medium width; dactylus medium-sized; basis of pereiopods 5 – 6 of normal width, with posteroproximal process present, forming a very low rounded projection in pereiopod 5, forming a well-developed asymmetrical rounded lobe in pereiopod 6; with posterodistal corner rounded, small, projecting forward; basis of pereiopod 7 broad with posterior border weakly convex, with distinct but very shallow notch on distal 0.8, forming a blunt-tipped extremely obtuse angle, with posterodistal corner rounded, small, projecting ventrally. Colour pattern Holotype red (Walker 1903). The colour of specimens from the Antarctic Peninsula, the Weddell Sea and Adélie Coast is very variable: uniformly orange, orange red with a finely mottled / doted pattern, whitish with sparse small mottling or body divided between white zones and red zones with a finely mottled / doted pattern; eyes always reddish. Body length 36 mm (holotype). The holotype is one of the largest specimens examined.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD56828FE46FC05CD2CFAF7.taxon	distribution	Distribution Elephant Island, tip of Antarctic Peninsula, eastern shelf of the Weddell Sea, Davis Sea, Adélie Coast, western Ross Sea; 33 – 791 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD56828FE46FC05CD2CFAF7.taxon	discussion	Remarks Examination of the holotype of Epimeria inermis (Ross Sea) revealed morphological differences with the specimens from the South Shetland Islands, the Antarctic Peninsula, the eastern Weddell Sea and Adélie Coast. The most striking difference is the telson, which is scarcely emarginate in the holotype of E. inermis, whilst it exhibits an acutely triangular V-shaped notch in all other specimens examined. However, the notch is also quite small in the specimen from the Adélie Coast illustrated by Bellan-Santini (1972). The rostrum is also shorter and more distinctly curved in the holotype of E. inermis. The dorsal carina of pereionite 4 is posteriorly slightly projecting upwards, which is not the case in other specimens. The ventral part of coxa 4 is less developed, forming a blunt-tipped obtuse angle instead of a sharp squared angle. The posterior notch of the basis of pereiopod 7 is a bit shallower. Finally, the process of urosomite 1 is more obtuse. Molecular data would be needed to investigate if the specimens from the Ross Sea (topotypical E. inermis) are conspecific with or distinct from specimens from other seas. On the other hand, the COI corrected mean distance (GTR + G + I) between the haplotypes from the Scotia Arc area and the Adélie Coast is 1.3 %. This distance is much higher than intraspecific distances observed in other Epimeria species (0 – 0.5 %; table 2 in Verheye et al. 2016 a). However, additional intraspecific sampling might increase these values. It is unclear from the molecular data if these two clades are distinct species or populations of the same widespread species (Verheye et al. 2016 a). As the single specimen from the eastern Weddell Sea was only sequenced for 28 S, it is possible that the more variable COI would show some differentiation with the haplotypes from the Adélie Coast and Scotia Arc area as well. But as no morphological differences were observed between the specimens from the three regions (compare Figs 167 – 172: Bransfield Strait and Figs 173 – 184: eastern Weddell Sea and Adélie Coast), they are provisionally considered as conspecifics. The record of E. inermis from the Falkland Islands by K. H. Barnard (1930) seems very dubious. It is possibly based on E. (Metepimeria) acanthurus (Schellenberg, 1931), which is the only Magellanic Epimeria species known to date.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD16826FE26FA4ECE7CFB0C.taxon	description	Figs 185 – 191	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD16826FE26FA4ECE7CFB0C.taxon	materials_examined	Material examined RV Aurora Australis cruises: SOUTHERN OCEAN: 3 small specs, cruise CEAMARC, sample CEAMARC 2173, stn 67 AEV 326, Adélie Coast, 65 ° 43 ʹ 07 ″ S, 143 ° 03 ʹ 37 ″ E, 1957 – 2154 m, beam trawl, 5 Jan. 2008 (MNHN-IU- 2014 - 4343); 1 adult ♀, cruise CEAMARC, sample CEAMARC 2173, stn 67 AEV 326, Adélie Coast, 65 ° 43 ʹ 07 ″ S, 143 ° 03 ʹ 37 ″ E, 1957 – 2154 m, beam trawl, 5 Jan. 2008 (RBINS, INV. 132718, removed from MNHN-IU- 2014 - 4344) [extraction M 16; Genbank nr, COI: KU 870882, 28 S: KU 759665]; 2 ♀♀, cruise CEAMARC, sample CEAMARC 2173, Adélie Coast, stn 67 AEV 326, 65 ° 43 ʹ 07 ″ S, 143 ° 03 ʹ 37 ″ E, beam trawl, 1957 – 2154 m, 5 Jan. 2008 (MNHN-IU- 2014 - 4344).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD16826FE26FA4ECE7CFB0C.taxon	description	Description ROSTRUM. Well-developed, reaching tip of article 2 of peduncle of antenna 1, anteriorly weakly curved, posteriorly nearly straight (inconspicuously concave), narrow and subacute in lateral view; narrow and with straight converging borders in dorsal view. EYE. Very large, distinctly reniform. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 3 smooth; pereionite 4 with trace of posterodorsal bump; pereionite 5 with small and low posterodorsal tooth; pereionite 6 to pleonite 2 with very low mid-dorsal carina posteriorly produced into a tooth pointing backwards and very shallow median notch; pleonite 3 with very low rounded protrusion on median 0.4 followed posteriorly by a straight line and terminated by a medium-sized broad bluntly triangular tooth pointing obliquely upwards; dorsolateral ornamentation absent. COXAE 1 – 3. Not carinate, apically rounded (coxa 1) and bluntly angular (coxae 2 – 3). COXA 4. Anterodorsal border straight, anteroventral border straight, these two borders being joined by blunt but distinct squared angle (anterior corner), which is distinctly projecting forward; ventral corner forming an acute (nearly squared) angle (ventral projection longer than in most Hoplepimeria); lateral carina absent; posteroventral border weakly concave. COXA 5. Broad, with surface smooth, with posteroventral corner forming a triangular tooth projecting laterally: in dorsal view the anterior border of the tooth points obliquely backwards (it forms a low concave curve with the more anterior part of the coxa), the tip of the tooth is blunt or subacute; the posterior border of the tooth is perpendicular with the body axis. COXA 6. With posteroventral corner forming a blunt acute but nearly squared angle, with low projection arising from its surface (forming blunt lobe in dorsal view). COXA 7. Posteriorly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle: very obtusely rounded in plate 1, produced into a small tooth in plate 2 and into a medium-sized tooth in plate 3. UROSOME TOOTH PATTERN. Urosomite 1 with very low dorsal process, forming a symmetrical and blunttipped obtuse triangle; urosomite 3 with dorsolateral borders nearly straight (very weakly convex) and posteriorly produced into a sharp triangular tooth. TELSON. Cleft on 0.2; tips of lobes triangular and sharp, notch broadly V-shaped and sharp at its deepest point. GNATHOPODS 1 – 2. With carpus and propodus very broad; propodus expanding distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus of medium width; dactylus medium-sized; basis of pereiopods 5 of normal width, with posteroproximal process present, acutely triangular, parallel to axis of basis, far from reaching tip of basis, with posterodistal corner produced into a bluntly quadrate lobe; basis of pereiopod 6 of normal width, with posteroproximal process present, sword-like, parallel to axis of basis, nearly reaching tip of basis, with posterodistal corner produced into a bluntly quadrate lobe; basis of pereiopod 7 broad with posterior border proximally distinctly convex, with very shallow concavity on distal 0.8; with posterodistal corner produced into a bluntly quadrate lobe. Variations Epimeria larsi exhibits variations (possibly related to size) in the shape of the lateral projection of coxa 5. It is very long and projecting backwards in the large holotype illustrated by Lörz (2009: 66, fig. 6). It is shorter and apparently projecting laterally in the smaller paratype illustrated by Lörz (2009: 62, fig. 2); the same disposition is observed in the material studied herein. In our specimens, coxa 1 is also less blunt and the spur of the basis of pereiopod 6 a bit longer than in the specimens illustrated by Lörz (2009). On the other hand, the colour pattern is the same. Colour pattern Body pale pink, with a brighter pink hue on the tip of the dorsal crests; coxae bright pink; lateral border of rostrum, antennae, pereiopods and tailfan blood red. Eyes orange. Body length Up to 58 mm (Lörz 2009). The specimen illustrated herein is 55 mm long.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD16826FE26FA4ECE7CFB0C.taxon	distribution	Distribution Western Ross Sea, 1954 – 1990 m (type material); Adélie Coast, 1957 – 2154 m (present material).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD16826FE26FA4ECE7CFB0C.taxon	discussion	Remarks The coordinates of the type station (TAN 0802 / 167) as given by Lörz et al. (2009): 71 ° 51 ʹ 21 ″ S, 71 ° 53 ʹ 24 ″ E are incorrect, this position being located on the Antarctic mainland far away from the Ross Sea area. The correct coordinates of this station are: 71.8557 ° S 174.033 ° E (Vargas et al. 2015: supplement 2).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFDF6824FE14FB46CB60FB20.taxon	description	urn: lsid: zoobank. org: act: 261 E 7087 - B 67 B- 47 ED- 905 B-FB 8 FF 8 C 636 D 8 Figs 192 – 197	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFDF6824FE14FB46CB60FB20.taxon	etymology	Etymology The species is dedicated to Katrin Linse (British Antarctic Survey, Cambridge), who played an important role in the study of the Epimeria georgiana complex. The name is a genitive.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFDF6824FE14FB46CB60FB20.taxon	materials_examined	Type material Holotype RV Polarstern Cruises: SOUTHERN OCEAN: ♀, cruise PS 77, ANT-XXVII / 3, South Orkney Islands, stn 217 - 6, 61 ° 09.62 ʹ S, 44 ° 02.37 ʹ W to 61 ° 10.52 ʹ S, 44 ° 04.91 ʹ W, 350 – 354 m, bottom trawl, 19 Feb. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 132958) [extraction I 10; Genbank nr, COI: KU 870838, 28 S: KU 759612]. Paratypes Expedition SIGNY 1991 / 92 (name of ship unavailable): SOUTHERN OCEAN: 1 large ♀, Transect 1, AGT 8, South Orkney Plateau: Signy Island, 60.679 ° S, 45.557 ° W, 150 – 160 m, 14 Jan. 1992, coll. Stefan Hain (RBINS, INV. 132688); 1 fairly small spec., Transect 2, AGT 23, South Orkney Plateau: Signy Island, 60.691 ° S, 45.443 ° W, 150 – 160 m, 14 Feb. 1992, coll. Stefan Hain (RBINS, INV. 132689); 1 large ♀, Transect 2, AGT 14, South Orkney Plateau: Signy Island, 60.705 ° S, 45.557 ° W, 150 m, 5 Feb. 1992, coll. Stefan Hain (RBINS, INV. 132690); 1 adult ♀, Transect 1, AGT 4, South Orkney Plateau: Signy Island, 60.685 ° S, 45.573 ° W, 100 – 140 m, 9 Jan. 1992, coll. Stefan Hain (RBINS, INV. 132691).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFDF6824FE14FB46CB60FB20.taxon	description	Description ROSTRUM. Large, nearly reaching tip of article 2 of peduncle of antenna 1, anteriorly slightly curved, ventrally inconspicuously concave, blunt-tipped in lateral view; narrow and with borders weakly convex in frontal view. EYE. Fairly large, nearly round. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 – 3 smooth; pereionites 4 with posterior bump and trace of carina; pereionites 5 to pleonite 3 with low, non-toothed, very broad carina, that of pleonite 2 with trace of concavity on anterior 0.4; pleonite 3 with weak dorsal protuberance followed by weak concavity followed by trace of second concavity; posterodorsal angle of pleonite 3 bluntly angular; dorsolateral ornamentation absent. COXAE 1 – 3. Scarcely carinate and apically tapering, sharp to subacute. COXA 4. Anterodorsal border nearly straight (very weakly sigmoid), anteroventral border weakly concave, these two borders being joined by blunt but very distinct angle (anterior corner), which is distinctly projecting forward; ventral corner forming a sharp or subacute squared angle (ventral projection short); lateral carina present but not sharp; broad hollow surface between carina and posteroventral border of coxa; posteroventral border distinctly concave. COXA 5. Broad, with surface smooth, with posteroventral corner forming a rounded lobe, weakly expanded laterally, forming a broadly rounded obtuse (nearly square) angle which is very distinct in dorsal view. COXA 6. With posteroventral corner rounded, with projection arising from its surface (broadly triangular in dorsal view). COXA 7. Posteriorly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle: produced into a small tooth in plates 1 – 2 and into a mediumsized tooth in plate 3. UROSOME TOOTH PATTERN. Urosomite 1 with very low dorsal process, anteriorly slightly concave, apically rounded and posteriorly oblique and straight; urosomite 3 with dorsolateral borders nearly straight (inconspicuously concave), with tip sharply angular. TELSON. Cleft on 0.25; lobes broad, forming subacute triangle; notch V-shaped, neither very narrow nor very broad. GNATHOPODS 1 – 2. Carpus and propodus very broad; propodus expanding distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus of medium width; dactylus rather long; basis of pereiopods 5 – 6 of normal width, with posteroproximal process present, tooth-like, parallel to axis of basis, with posterodistal corner bluntly angular, very weakly pointing in posterior direction; basis of pereiopod 7 broad with posterior border weakly convex, with distinct notch on distal 0.8, forming a blunt-tipped squared or acute angle, with posterodistal corner rounded and not projecting backwards. Colour pattern Body and appendages with a diffuse orange dotted / mottled pattern, more intense in some parts; eye reddish. Body length Up to 35 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFDF6824FE14FB46CB60FB20.taxon	distribution	Distribution Plateau of the South Orkney Islands. Our specimens were collected between 100 and 354 m. If we are correct in identifying the ‘ Epimeria georgiana clade B’ of Lörz et al. (2011) as E. linseae sp. nov., the species descends down to 964 – 1014 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFDF6824FE14FB46CB60FB20.taxon	discussion	Remarks E. linseae sp. nov. is morphologically similar to E. quasimodo sp. nov. The best differential character is the shape of the rostrum, which is narrower (both in lateral and dorsal view) and laterally more curved in the first than in the second species. The dorsal profile of pleonite 3 and urosomite 3 is also slightly different. Material from Coleman (2007) is here attributed to E. linseae sp. nov. based on morphology (especially for the shape of the rostrum) and collection location: 60 ° 43.12 ʹ S, 45 ° 30.86 ʹ W (South Orkney Islands), 141 – 190 m, Agassiz trawl, 3.1.1985, leg. Wägele (Coleman pers. com.).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFDD6820FE3CFB3ACE34FB78.taxon	description	urn: lsid: zoobank. org: act: DA 484 D 99 - BC 6 E- 4 E 9 D-B 06 E- 9 D 775 DCB 7 FD 1 Figs 198 – 207	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFDD6820FE3CFB3ACE34FB78.taxon	etymology	Etymology Quasimodo, the hunchback of Notre-Dame, is a well-known character from the novel ‘ Notre-Dame de Paris’ by Victor Hugo. The name, which is a noun in apposition, alludes to the humpbacked silhouette of the species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFDD6820FE3CFB3ACE34FB78.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: ♀, cruise PS 81, ANT-XXIX / 3, stn 160 - 3, East of Joinville Island, 63 ° 10.57 ʹ S, 54 ° 06.66 ʹ W to 63 ° 10.71 ʹ S, 54 ° 06.37 ʹ W, 238 – 244 m, Agassiz trawl, 8 Feb. 2013 (RBINS, INV. 122930 A) [extraction K 23; Genbank nr, COI: KU 870857, 28 S: KU 759635]. Paratypes RV Polarstern cruises: SOUTHERN OCEAN: 1 ♀, cruise PS 14, ANT-VII / 4, EPOS leg 3, stn 211, Elephant Island, 60 ° 59.8 ʹ S, 55 ° 12.1 ʹ W to 60 ° 59.3 ʹ S, 55 ° 10.5 ʹ W, 207 – 213 m, bottom trawl, 15 Jan. 1989, coll. C. De Broyer (RBINS, INV. 132950); 3 specs, cruise PS 69, ANT-XXIII / 8, stn 604 - 1, Elephant Island, 61 ° 20.52 ʹ S, 55 ° 09.72 ʹ W to 61 ° 20.11 ʹ S, 55 ° 07.26 ʹ W, 286 – 407 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122467); 3 specs, cruise PS 69, ANT-XXIII / 8, stn 604 - 1, Elephant Island, 61 ° 20.52 ʹ S, 55 ° 09.72 ʹ W to 61 ° 20.11 ʹ S, 55 ° 07.26 ʹ W, 286 – 407 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122496); 2 specs, cruise PS 69, ANT-XXIII / 8, stn 604 - 1, Elephant Island, 61 ° 20.52 ʹ S, 55 ° 09.72 ʹ W to 61 ° 20.11 ʹ S, 55 ° 07.26 ʹ W, 286 – 407 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122500); 1 juv., cruise PS 69, ANT-XXIII / 8, stn 604 - 1, Elephant Island, 61 ° 20.52 ʹ S, 55 ° 09.72 ʹ W to 61 ° 20.11 ʹ S, 55 ° 07.26 ʹ W, 286 – 407 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122508); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 605 - 1, Elephant Island, 61 ° 20.35 ʹ S, 55 ° 29.16 ʹ W to 61 ° 19.98 ʹ S, 55 ° 32.67 ʹ W, 146 – 151 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122504); 1 juv., cruise PS 69, ANT-XXIII / 8, stn 605 - 1, Elephant Island, 61 ° 20.35 ʹ S, 55 ° 29.16 ʹ W to 61 ° 19.98 ʹ S, 55 ° 32.67 ʹ W, 146 – 151 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122506); 4 specs, cruise PS 69, ANT-XXIII / 8, stn 605 - 5, Elephant Island, 61 ° 20.27 ʹ S, 55 ° 30.92 ʹ W to 61 ° 20.37 ʹ S, 55 ° 28.99 ʹ W, 131 – 152 m, Agassiz trawl, 20 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122501); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 608 - 1, Elephant Island, 61 ° 11.34 ʹ S, 54 ° 43.17 ʹ W to 61 ° 11.80 ʹ S, 54 ° 40.05 ʹ W, 284 – 293 m, bottom trawl, 20 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122515); 3 specs, cruise PS 69, ANT-XXIII / 8, stn 608 - 1, Elephant Island, 61 ° 11.34 ʹ S, 54 ° 43.17 ʹ W to 61 ° 11.80 ʹ S, 54 ° 40.05 ʹ W, 284 – 293 m, bottom trawl, 20 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122498); 2 specs, cruise PS 69, ANT-XXIII / 8, stn 608 - 1, Elephant Island, 61 ° 11.34 ʹ S, 54 ° 43.17 ʹ W to 61 ° 11.80 ʹ S, 54 ° 40.05 ʹ W, 284 – 293 m, bottom trawl, 20 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (MNHN-IU- 2014 - 7334, removed from RBINS, INV. 122498); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 611 - 1, Elephant Island, 60 ° 58.90 ʹ S, 55 ° 11.31 ʹ W to 60 ° 58.52 ʹ S, 55 ° 07.82 ʹ W, 215 – 297 m, bottom trawl, 21 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122558); 1 spec., initially fixed with formalin, cruise PS 69, ANT-XXIII / 8, stn 622 - 1, Elephant Island, 60 ° 56.70 ʹ S, 55 ° 52.71 ʹ W to 60 ° 55.93 ʹ S, 55 ° 50.79 ʹ W, 218 – 307 m, Agassiz trawl, 23 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122505); 7 specs, cruise PS 69, ANT-XXIII / 8, stn 654 - 6, Elephant Island, 61 ° 22.80 ʹ S, 56 ° 03.84 ʹ W to 61 ° 23.35 ʹ S, 56 ° 04.89 ʹ W, 341 – 342 m, Agassiz trawl, 29 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122499); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 668 - 1, north of King George Island, 61 ° 49.32 ʹ S, 58 ° 34.74 ʹ W to 61 ° 50.05 ʹ S, 58 ° 30.67 ʹ W, 152 – 193 m, bottom trawl, 31 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122503); 1 spec., cruise PS 69, ANT-XXIII / 8, stn; 688 - 1, north of Joinville Island, 62 ° 32.27 ʹ S, 54 ° 57.55 ʹ W to 62 ° 33.59 ʹ S, 55 ° 0.22 ʹ W, bottom trawl, 4 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122495); 1 very large spec., cruise PS 69, ANT-XXIII / 8, stn 692 - 1, north of Joinville Island, 62 ° 21.76 ʹ S, 55 ° 36.96 ʹ W to 62 ° 23.62 ʹ S, 55 ° 36.42 ʹ W, 263 – 277 m, bottom trawl, 5 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122497); 1 juv., cruise PS 69, ANT-XXIII / 8, stn 726 - 4, southwest of Snow Hill Island, 64 ° 37.83 ʹ S, 56 ° 42.10 ʹ W to 64 ° 38.03 ʹ S, 56 ° 42.57 ʹ W, 292 m, Agassiz trawl, 23 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122507); 2 ♀♀, cruise PS 81, ANT-XXIX / 3, stn 116 - 9, north of Joinville Island, 62 ° 33.79 ʹ S, 56 ° 27.81 ʹ W to 62 ° 33.71 ʹ S, 56 ° 28.31 ʹ W, 248 m, muddy bottom with stones, Pentapora - like bryozoans and a lot of life, Agassiz trawl, 26 Jan. 2013 (RBINS, INV. 122920) [extractions K 27 (largest specimen); Genbank nr, COI: KU 870861, 28 S: KU 759639 and K 28 (smallest specimen), Genbank nr, COI: KU 870862, 28 S: KU 759640]; 1 juv., cruise PS 81, ANT-XXIX / 3, stn 160 - 3, East of Joinville Island, 63 ° 10.57 ʹ S, 54 ° 6.66 ʹ W to 63 ° 10.71 ʹ S, 54 ° 6.37 ʹ W, 238 – 244 m, Agassiz trawl, 8 Feb. 2013 (RBINS, INV. 122930 B); 1 ♀, cruise PS 81, ANT-XXIX / 3, stn 160 - 3, East of Joinville Island, 63 ° 10.57 ʹ S, 54 ° 6.66 ʹ W to 63 ° 10.71 ʹ S, 54 ° 6.37 ʹ W, 238 – 244 m, Agassiz trawl, 8 Feb. 2013 (RBINS, INV. 122933) [extraction K 24; Genbank nr, COI: KU 870858, 28 S: KU 759636]; 1 immature spec., cruise PS 81, ANT-XXIX / 3, stn 162 - 7, east of James Ross Island, 63 ° 58.78 ʹ S, 56 ° 46.24 ʹ W to 63 ° 59.02 ʹ S, 56 ° 46.26 ʹ W, 214 – 216 m, muddy bottom with a lot of life, Agassiz trawl, 10 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122952); 1 very large spec., cruise PS 81, ANT-XXIX / 3, stn 162 - 7, east of James Ross Island, 63 ° 58.78 ʹ S, 56 ° 46.24 ʹ W to 63 ° 59.02 ʹ S, 56 ° 46.26 ʹ W, 214 – 216 m, muddy bottom with a lot of life, Agassiz trawl, 10 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122952); 1 very large spec., cruise PS 81, ANT-XXIX / 3, stn 164 - 4, south of Dundee Island, 63 ° 37.28 ʹ S, 56 ° 9.11 ʹ W to 63 ° 37.29 ʹ S, 56 ° 9.58 ʹ W, 102 – 114 m, non-muddy bottom with a lot of life, including Molgula, big red ophiuroids, and a lot of Glyptonotus, 11 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122923) [extraction K 29; Genbank nr, COI: KU 870863, 28 S: KU 759641]; 1 very large ♀, cruise PS 81, ANT-XXIX / 3, stn 164 - 4, south of Dundee Island, 63 ° 37.28 ʹ S, 56 ° 9.11 ʹ W to 63 ° 37.29 ʹ S, 56 ° 9.58 ʹ W, 102 – 114 m, non-muddy bottom with a lot of life, including Molgula, big red ophiuroids, and a lot of Glyptonotus, Agassiz trawl, 11 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122925) [extraction K 30; Genbank nr, COI: KU 870864, 28 S: KU 759643]; 1 ♀, cruise PS 81, ANT-XXIX / 3, stn 185 - 3, south east of Dundee Island, 63 ° 51.34 ʹ S, 55 ° 41.11 ʹ W to 63 ° 51.52 ʹ S, 55 ° 41.43 ʹ W, Agassiz trawl, 238 – 244 m, not muddy bottom with a lot of life (sponges, starfishes, ophiuroids, crinoids, Pentapora - like bryozoans), Agassiz trawl, 19 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122921 A) [extraction K 26; Genbank, COI nr: KU 870860, 28 S: KU 759638] [the station given by Verheye et al. (2016 a) is incorrect]; 1 adult ♀, cruise PS 81, ANT-XXIX / 3, stn 185 - 3, south east of Dundee Island, 63 ° 51.34 ʹ S, 55 ° 41.11 ʹ W to 63 ° 51.52 ʹ S, 55 ° 41.43 ʹ W, Agassiz trawl, 238 – 244 m, not muddy bottom with a lot of life (sponges, starfishes, ophiuroids, crinoids, Pentapora - like bryozoans), Agassiz trawl, 19 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye, dissected and photographed (RBINS, INV. 122921 B) [extraction P 35; Genbank nr, COI: KU 870893, 28 S: KU 759679]; 2 immature spec., cruise PS 81, ANT-XXIX / 3, stn 185 - 3, south east of Dundee Island, 63 ° 51.34 ʹ S, 55 ° 41.11 ʹ W to 63 ° 51.52 ʹ S, 55 ° 41.43 ʹ W, Agassiz trawl, 238 – 244 m, not muddy bottom with a lot of life (sponges, starfishes, ophiuroids, crinoids, Pentapora - like bryozoans), Agassiz trawl, 19 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122921 C) [extraction P 37; Genbank nr, COI: KU 870894, 28 S: KU 759681]; 1 spec., cruise PS 81, ANT-XXI / 3, stn 185 - 4, south east of Dundee Island, 63 ° 51.53 ʹ S, 55 ° 40.74 ʹ W to 63 ° 51.53 ʹ S, 55 ° 40.43 ʹ W, 253 – 255 m, extremely fine sand mixed with some mud and gravel, Rauschert dredge, 19 Feb. 2013 coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 132971) [extraction I 20; Genbank nr, COI: KU 870846, 28 S: KU 759623]; 1 ♀, cruise PS 81, ANT-XXIX / 3, stn 197 - 5, Bransfield Strait, 62 ° 44.73 ʹ S, 57 ° 26.79 ʹ W to 62 ° 45.05 ʹ S, 57 ° 26.68 ʹ W, 258 – 273 m, Agassiz trawl, 25 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122867) [extraction ANT 41; Genbank nr, COI: KU 870827, 28 S: KU 759600]; 1 ♀, cruise PS 81, ANT-XXIX / 3, stn 197 - 6, Bransfield Strait, 62 ° 45.05 ʹ S, 57 ° 26.68 ʹ W to 62 ° 45.09 ʹ S, 57 ° 26.47 ʹ W, 210 – 222 m, black gravel mixed with sand and a little bit of mud, Rauschert dredge, 25 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122954); 1 juv., cruise PS 81, ANT-XXIX / 3, stn 197 - 6, Bransfield Strait, 62 ° 45.05 ʹ S, 57 ° 26.68 ʹ W to 62 ° 45.09 ʹ S, 57 ° 26.47 ʹ W, 210 – 222 m, black gravel mixed with sand and a little bit of mud, Rauschert dredge, 25 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122954); 1 spec., cruise PS 81, ANT-XXIX / 3, stn 205 - 1, Bransfield Strait, 63 ° 2.35 ʹ S, 58 ° 3.63 ʹ W to 63 ° 2.67 ʹ S, 58 ° 6.46 ʹ W, depth unrecorded [Google Earth coordinates for these positions: 155 – 178 m (accessed 27 Sep. 2016], rectangular midwater trawl (which presumably hit the sea floor, as it was supposed to sample between 0 and 200 m depth), 28 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122970); 1 ♀, cruise PS 81, ANT-XXIX / 3, stn 224 - 3, Bransfield Strait, 63 ° 0.53 ʹ S, 58 ° 35.67 ʹ W to 63 ° 0.58 ʹ S, 58 ° 36.11 ʹ W, 257 – 261 m, mud, Agassiz trawl, 4 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122924) [extraction K 22; Genbank nr, COI: KU 870856, 28 S: KU 759634]; 1 very large ♀, cruise PS 81, ANT-XXIX / 3, stn 224 - 3, Bransfield Strait, 63 ° 0.53 ʹ S, 58 ° 35.67 ʹ W to 63 ° 0.58 ʹ S, 58 ° 36.11 ʹ W, 257 – 261 m, mud, Agassiz trawl, 4 Mar. 2013, red colour, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122926) [extraction K 21; Genbank nr, COI: KU 870855, 28 S: KU 759633]; 1 ♀, cruise PS 81, ANT-XXIX / 3, stn 224 - 3, Bransfield Strait, 63 ° 0.53 ʹ S, 58 ° 35.67 ʹ W to 63 ° 0.58 ʹ S, 58 ° 36.11 ʹ W, 257 – 261 m, mud, Agassiz trawl, 4 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122953); 3 large specs, cruise PS 81, ANT-XXIX / 3, stn 224 - 3, Bransfield Strait, 63 ° 0.53 ʹ S, 58 ° 35.67 ʹ W to 63 ° 0.58 ʹ S, 58 ° 36.11 ʹ W, 257 – 261 m, mud, Agassiz trawl, 4 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122957); 1 ♀, cruise PS 81, ANT-XXIX / 3, stn 234 - 5, north of Livingstone Island, 62 ° 17.36 ʹ S, 61 ° 12.06 ʹ W to 62 ° 17.31 ʹ S, 61 ° 12.63 ʹ W, 248 – 251 m, mud with a lot of specimens of a small reddish ophiuroid with very long and flexible legs, Agassiz trawl, 7 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 132970) [extraction I 14; Genbank nr, COI: KU 870840, 28 S: KU 759616].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFDD6820FE3CFB3ACE34FB78.taxon	description	Description ROSTRUM. Medium-sized, overreaching tip of article 1 of peduncle of antenna 1, anteriorly slightly curved, ventrally straight, blunt-tipped in lateral view; broad and with proximally nearly straight converging borders in frontal view. EYE. Fairly large, broadly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 smooth; pereionites 2 – 3 with weak posterior bump; pereionites 4 to pleonite 3 with low, non-toothed, very broad carina (very reduced in pereionites 2 – 3), those of pereionite 7 and pleonites 1 – 3 with very shallow concavity on anterior 0.4; posterodorsal angle of pleonite 3 broadly rounded; dorsolateral ornamentation absent. COXAE 1 – 3. Scarcely carinate and apically tapering and blunt (coxae 1 – 2) or subacute (coxa 3). COXA 4. Anterodorsal border nearly straight (very weakly sigmoid), anteroventral border weakly concave, these two borders being joined by blunt but very distinct angle (anterior corner), which is distinctly projecting forward; ventral corner forming a sharp squared angle (ventral projection short); lateral carina present not sharp; broad hollow surface between carina and posteroventral border of coxa; posteroventral border distinctly concave. COXA 5. Broad, with surface smooth, with posteroventral corner forming a rounded lobe, weakly expanded laterally, forming a broadly rounded obtuse (nearly squared) angle which is very distinct in dorsal view. COXA 6. With posteroventral corner rounded, with projection arising from its surface (broadly triangular in dorsal view). COXA 7. Posteriorly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle: forming an angle with a trace of tooth in plate 1, produced into a small tooth in plate 2 and into a medium-sized tooth in plate 3. UROSOME TOOTH PATTERN. Urosomite 1 with very low dorsal process, regularly convex on both sides; urosomite 3 with dorsolateral borders nearly straight (inconspicuously convex), with tip bluntly angular. TELSON. Cleft on 0.4; lobes broad becoming apically broadly and bluntly triangular, notch narrowly V-shaped. GNATHOPODS 1 – 2. Carpus and propodus very broad; propodus expanding distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus of medium width; dactylus medium-sized; basis of pereiopods 5 – 6 of normal width, with posteroproximal process present, tooth-like, parallel to axis of basis, with posterodistal corner bluntly angular, very weakly pointing in posterior direction; basis of pereiopod 7 broad with posterior border weakly convex, with distinct notch in distal 0.8, forming a blunt-tipped squared or acute angle, with posterodistal corner rounded and not projecting backwards. Colour pattern Red-coloured with the red pigmentation forming an irregular, finely mottled / dotted pattern; or whitish with very irregular and diffuse red patches, in which the red pigmentation is forming an irregular finely peppered pattern. Eyes reddish. Body length Up to 50 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFDD6820FE3CFB3ACE34FB78.taxon	distribution	Distribution Elephant Island to tip of Antarctic Peninsula, 131 – 407 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFDD6820FE3CFB3ACE34FB78.taxon	biology_ecology	Biology Examination of the stomach content of fifteen specimens indicates that the species is carnivorous (Coleman 1991, as Epimeria georgiana).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFDD6820FE3CFB3ACE34FB78.taxon	discussion	Remarks A juvenile E. quasimodo sp. nov. is illustrated herein (Fig. 207) to show the notch of the basis of pereiopod 7, which is much shallower than in adults. Epimeria quasimodo sp. nov. is a very common species, which was previously confused with E. georgiana. The latter is presumably endemic to South Georgia.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD9681FFEBBFAF3CA4DFD6F.taxon	description	Fig. 208	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD9681FFEBBFAF3CA4DFD6F.taxon	materials_examined	Material examined RV Marion Dufresne cruises: SOUTHERN OCEAN: 1 spec., specimen used for illustrations, cruise MD 42 (SIBEX), stn 22 - CP 66, Prydz Bay, 66 ° 55 ʹ 45 ″ S, 74 ° 04 ʹ 11 ″ E, depth not given [Google Earth depth for that position: 427 m (accessed 27 Sep. 2016)], 26 Jan. 1985 (MNHN-IU- 2014 - 4265) [extraction M 24; Genbank nr, COI: KU 870887]; 1 spec., previously identified as Epimeria grandirostris, cruise MD 42 (SIBEX), stn 22 - CP 73, Prydz Bay, 66 ° 57 ʹ 25 ″ S, 72 ° 41 ʹ 24 ″ E, depth not given [Google Earth depth for that position: 540 m (accessed 27 Sep. 2016)], 26 Jan. 1985 (MNHN-IU- 2014 - 4256).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD9681FFEBBFAF3CA4DFD6F.taxon	diagnosis	Diagnosis Morphology unique in the genus Epimeria. Pereionite 2 with small but broad posterodorsal tooth; pereionite 3 to pleonite 3 laterally sculptured, with tridimentionally sculptured and medially notched mid-dorsal tooth. Posteroventral border of coxa 4 very concave, surface just in front of this border forming a very deep groove. Colour pattern Red with on each side a broad white longitudinal stripe running along the upper part of the coxae (Coleman 2007: plate 2 fig. e) or blood red with white patches (Lörz & Coleman 2009: unnumbered photograph p. 17).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFD9681FFEBBFAF3CA4DFD6F.taxon	distribution	Distribution Western Ross Sea, 337 – 474 m (Watling & Holman 1980); Prydz Bay, 427 – 540 m (present material).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE6681DFE7CFCE5CD9FFEC0.taxon	description	Figs 209 – 211	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE6681DFE7CFCE5CD9FFEC0.taxon	materials_examined	Material examined RV Seatruck cruises: SOUTHERN OCEAN: 1 ♀, cruise REVOLTA I, stn REVO- 043, Collect _ ID: 436, Adélie Coast, 66 ° 38 ʹ 28 ″ S, 140 ° 01 ʹ 50 ″ E, 85 – 86 m, 26 Jan. 2010, coll. M. Eléaume, L. Hemery and A. D’Hont (MNHN-IU- 2009 - 2571) [extraction K 1; Genbank nr, COI: KU 870854, 28 S: KU 759631].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE6681DFE7CFCE5CD9FFEC0.taxon	description	Description ROSTRUM. Medium-sized, nearly reaching tip of article 1 of peduncle of antenna 1, anteriorly straight, ventrally straight, narrow and subacute in lateral view; of medium width and with weakly curved converging borders in frontal view. EYE. Very large, narrowly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 7 smooth; pereionite 7 dorsally scarcely carinate; pleonite 1 with weak, straight mid-dorsal carina; pleonite 2 with weak straight mid-dorsal carina, posteriorly simply angulate but not forming a tooth; pleonite 3 dorsally weakly carinate with median extremely low (inconspicuous) rounded lobe, followed by inconspicuous concavity, terminated by a narrow but blunt-tipped tooth directed upwards. COXAE 1 – 3. Not carinate, apically blunt. COXA 4. Anterodorsal border very weakly sigmoid, anteroventral border straight, these two borders being joined by very broad, blunt but very distinct squared angle (anterior corner), which is slightly projecting forward; ventral corner forming an obtuse sharp angle (ventral projection short and fairly broad); lateral carina absent; posteroventral border straight. COXA 5. Broad, with surface smooth, with posterior border straight, with posteroventral corner forming a very blunt tooth (shape: acute triangle) projecting backwards and not laterally (no tooth or corner visible in dorsal view). COXA 6. With posterior border straight, with posteroventral corner forming a blunt tooth (shape: narrow acute triangle) projecting backwards and not laterally (no tooth or corner visible in dorsal view). COXA 7. Posteriorly very weakly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle: angulate in plate 1, produced into a medium-sized tooth in plates 2 – 3. UROSOME TOOTH PATTERN. Urosomite 1 with well developed blunt-tipped process of which the anterior border is distinctly concave and the posterior border is nearly straight; urosomite 3 with dorsolateral borders distinctly concave and posteriorly produced into a sharp triangular tooth. TELSON. Cleft on 0.15; tips of lobes triangular and blunt, notch very broadly V-shaped and blunt at its deepest point. GNATHOPODS 1 – 2. Carpus and propodus very broad; propodus expanding distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus fairly broad; dactylus small, normally curved, with unguis normally developed; basis of pereiopods 5 – 6 broad, with posteroproximal process present, swordlike, projecting obliquely, with posterodistal corner produced into a subacute to sharp triangular tooth projecting obliquely backwards; basis of pereiopod 7 broad; posterior border with proximal 0.4 weakly rounded, with distal 0.6 deeply concave, with posterodistal corner forming a sharp triangular tooth projecting obliquely backwards. Colour pattern Body and coxae either yellowish, sometimes with two pairs of small yellowish spots transversally ordered on some body segments or alternatively pale yellowish with a few tiny dark dots; pereiopods and mouthparts often tinged with pink; eyes red (Lörz et al. 2009: 111, fig. 10 B – D). Body length Up to 43 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE6681DFE7CFCE5CD9FFEC0.taxon	distribution	Distribution Adélie Coast to western Ross Sea, 85 – 814 m (Lörz et al. 2009; present material).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE6681DFE7CFCE5CD9FFEC0.taxon	biology_ecology	Biology K. H. Barnard (1930) found epicaridean isopods in the marsupium of E. robusta.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE6681DFE7CFCE5CD9FFEC0.taxon	discussion	Remarks Epimeria robusta s. str. is known from Adélie Coast and the western Ross Sea. Records from the Weddell Sea and the Scotia Region are based on its close relatives E. robustoides and E. gargantua sp. nov. Epimeria robusta can be easily distinguished from its relatives by the dentition of its pleosome (see key of Hoplepimeria subgen. nov. species).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE4681BFEE7FE98CA11FCEA.taxon	description	Figs 212 – 219	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE4681BFEE7FE98CA11FCEA.taxon	materials_examined	Material examined RV Polarstern cruises: SOUTHERN OCEAN: 3 specs, dorsally opened for gut content analysis, cruise PS 14, ANT-VII / 4, EPOS 3, stn 290 (AGT 24), 71 ° 05.9 ʹ S, 12 ° 34.0 ʹ W to 71 ° 06.5 ʹ S, 12 ° 42.3 ʹ W, 522 – 531 m, 19 Feb. 1989, coll. C. De Broyer (RBINS, INV. 132944); 1 ♀, cruise PS 69, ANT-XXIII / 8, stn 603 - 5, eastern Weddell Sea, 70 ° 30.99 ʹ S, 08 ° 48.08 ʹ W to 70 ° 30.40 ʹ S, 08 ° 48.13 ʹ W, 274 – 297 m, sponge bottom, Agassiz trawl, 7 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 132985); 1 juv., cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, stn 48 - 1, eastern Weddell Sea, 70 ° 23.94 ʹ S, 8 ° 19.14 ʹ W to 70 ° 23.89 ʹ S, 8 ° 18.67 ʹ W, 595 – 602 m, bryozoan bottom (exceptional bryozoan diversity), Rauschert dredge, 12 Jan. 2008:, coll. H. Robert (RBINS, INV. 132969) [extraction I 8; Genbank nr, COI: KU 870852, 28 S: KU 759629]; 1 ♀, cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 265 - 2, eastern Weddell Sea, 70 ° 47.34 ʹ S, 10 ° 40.39 ʹ W to 70 ° 47.13 ʹ S, 10 ° 40.54 ʹ W, 500 – 600 m, 22 Mar. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 122894) [extraction Ex 114; Genbank nr, COI: KU 870834, 28 S: KU 759608].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE4681BFEE7FE98CA11FCEA.taxon	description	Description ROSTRUM. Medium-sized, nearly reaching tip of article 1 of peduncle of antenna 1, anteriorly straight, ventrally straight, narrow and subacute in lateral view; of medium width and with straight converging borders in frontal view. EYE. Very large, narrowly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 6 smooth; pereionite 7 with weak posterior bump; pleonite 1 dorsally distinctly carinate, straight, with distinct posterior bump or small very blunt tooth; pleonite 2 dorsally distinctly carinate, with very low proximal rounded lobe followed by weak concavity, posteriorly produced into a bluntly triangular tooth projecting obliquely upwards; pleonite 3 dorsally distinctly carinate with median very low rounded lobe, followed by distinct concavity, terminated by a blunt tooth directed upwards. COXAE 1 – 3. Not carinate, apically subacute. COXA 4. Anterodorsal border very weakly sigmoid, anteroventral border nearly straight (extremely weakly convex), these two borders being joined by very broad blunt, very distinct squared angle (anterior corner), which is slightly projecting forward; ventral corner forming an obtuse sharp angle (ventral projection short and fairly broad); lateral carina absent; posteroventral border straight. COXA 5. Broad, with surface smooth, with posterior border straight, with posteroventral corner forming a very blunt tooth (shape: acute triangle) projecting backwards and not laterally (no tooth or corner visible in dorsal view). COXA 6. With posterior border straight, with posteroventral corner forming a blunt tooth (shape: narrow acute triangle) projecting backwards and not laterally (no tooth or corner visible in dorsal view). COXA 7. Posteriorly very weakly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle: angulate with tiny tooth in plate 1, produced into a mediumsized tooth in plates 2 – 3. UROSOME TOOTH PATTERN. Urosomite 1 with well developed blunt-tipped process of which the anterior border is distinctly concave and the posterior border is straight; urosomite 3 with dorsolateral borders strongly concave and posteriorly produced into a sharp triangular tooth. TELSON. Cleft on 0.25; tips of lobes triangular and blunt, notch very broadly V-shaped and subacute at its deepest point. GNATHOPODS 1 – 2. Carpus and propodus very broad; propodus expanding distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus fairly broad; dactylus small, normally curved, with unguis normally developed; basis of pereiopods 5 – 6 broad, with posteroproximal process present, swordlike, projecting obliquely, with posterodistal corner produced into a rounded lobe projecting obliquely backwards (pereiopod 5) or a blunt triangular tooth projecting obliquely backwards (pereiopod 6); basis of pereiopod 7 broad; posterior border with proximal 0.4 with weak concavity, with distal 0.6 deeply concave, with posterodistal corner forming a blunt triangular tooth projecting obliquely backwards. Colour pattern Body and coxae either pure white, or white with two pairs of small yellow spots transversally ordered on some body segments; gnathopods and oral field deep purple; antennae yellowish; pereiopods 3 – 4 pink; pereiopods 5 – 7 with basis white and distal articles pale yellowish to orange; tailfan white or whitish; eye red. Body length Up to 40 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE4681BFEE7FE98CA11FCEA.taxon	distribution	Distribution Eastern shelf of the Weddell Sea, 274 – 605 m (Lörz et al. 2009; present data).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE4681BFEE7FE98CA11FCEA.taxon	biology_ecology	Biology According to Klages & Gutt (1990), the digestive tract of specimens identified as E. robusta (but presumably belonging to E. robustoides as they were collected in the eastern Weddell Sea) contained mainly (42 %) miscellaneous organic matter. Only in some specimens were siliceous sigma and other types of microspiculae of Demospongiae and calcareous spiculae of Calcispongiae found among sediment particles and organic material. In all the E. robusta stomachs examined by them, some 10 % of the contents consisted of tentacles and body wall sclerites of holothurians. The tentacle sclerites of Abyssocucumis cf. liouvillei [i. e. Staurocucumis cf. liouvillei (Vaney, 1914)] (Holothuroidea: Dendrochirotida) in one single specimen were identified. On the other hand, Dauby et al. (2001 a) recorded a wider range of organisms in the digestive tract of the species (also specimens from the Eastern Weddell Sea identified as E. robusta): ophiuroids, plankton remains, sponges, cnidarians, polychaetes and crustaceans. Dauby et al. (2001 b) concluded that E. robustoides would be an opportunistic predator. Klages (1991, as Epimeria robusta) indicates that the species is sometimes parasitized by cryptoniscin isopods. Klages (1991, as E. robusta) also observed that hatchlings of E. robustoides climb on the back of their mother and remain there for a short period of time.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE4681BFEE7FE98CA11FCEA.taxon	discussion	Remarks Epimeria robustoides is extremely similar to E. gargantua sp. nov., but the two species appear to have allopatric distributions. Epimeria robustoides has been recorded from the eastern shelf of the Weddell Sea, whilst E. gargantua sp. nov. has been found around the tip of the Antarctic Peninsula. Rauschert & Arntz (2015) do not indicate the coordinates of their illustrated ‘ Epimeria robustoides ’, but the origin of the specimens was listed in an early draft of their book made available to the authors. The specimen 2 of Rauschert & Arntz (2015) was collected during the Polarstern cruise ANT-XIII / 3 (EASIZ 1), which sampled on the eastern shelf of the Weddell Sea; hence it is presumably E. robustoides. The specimen 1 of Rauschert & Arntz (2015) was collected on the Bruce Ridge (a small isolated shallow bank situated midway between the South Orkney Islands and the South Sandwich Islands) and presumably belongs to an undescribed species, which is referred herein as Epimeria (Hoplepimeria) sp. subgen. nov.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE26819FEB9FC62CBA9FC9D.taxon	description	Figs 220 – 221	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE26819FEB9FC62CBA9FC9D.taxon	materials_examined	Material examined RV Polarstern cruises: SOUTHERN OCEAN: 3 specs, in 3 different tubes, cruise PS 10, ANT-V / 3, station missing, eastern Weddell Sea, fixed on board [presumably after aquarium observations] on 20 Nov. 1986, coll. C. De Broyer (RBINS, INV. 132732); 1 spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 17, GSN 10, eastern Weddell Sea, 73 ° 18.00 ʹ S, 21 ° 09.90 ʹ W to 73 ° 19.10 ʹ S, 21 ° 14.90 ʹ W, 465 – 468 m, bottom trawl, specimen kept on board in aquarium, 16 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132680); 1 spec., cruise PS 48, ANT-XV / 3, EASIZ II, stn 206, eastern Weddell Sea, 71 ° 00.4 ʹ S, 11 ° 42.6 ʹ W to 71 ° 00.7 ʹ S, 11 ° 42.5 ʹ W, 594 – 602 m, Agassiz trawl, 18 Feb. 1998, coll. C. De Broyer (RBINS, INV. 132990); 1 spec., cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, stn 48 - 1, eastern Weddell Sea, 70 ° 23.94 ʹ S, 8 ° 19.14 ʹ W to 70 ° 23.89 ʹ S, 8 ° 18.67 ʹ W, 595 – 602 m, bryozoan bottom (exceptional bryozoan diversity), Rauschert dredge, 12 Jan. 2008, coll. H. Robert (RBINS, INV. 132943) [extraction K 41; Genbank nr, COI: KU 870874, 28 S: KU 759654]; 1 spec., fixed in formalin, cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, stn 48 - 1, eastern Weddell Sea, 70 ° 23.94 ʹ S, 8 ° 19.14 ʹ W, to 70 ° 23.89 ʹ S, 8 ° 18.67 ʹ W, 595 – 602 m, bryozoan bottom (exceptional bryozoan diversity), Rauschert dredge, 12 Jan. 2008, coll. H. Robert (RBINS, INV. 132410); 1 small badly damaged spec., cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 265 - 2, eastern Weddell Sea, BENDEX area, 70 ° 47.34 ʹ S, 10 ° 40.39 ʹ W to 70 ° 47.13 ʹ S, 10 ° 40.54 ʹ W, depth not recorded [Google Earth depth range for these coordinates: 571 – 599 m (accessed 27 Sep. 2016)], Agassiz trawl, 22 Mar. 2011:, coll. Ch. Havermans and H. Robert (RBINS, INV. 132968) [extraction I 5; Genbank nr, COI: KU 870849, 28 S: KU 759626].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE26819FEB9FC62CBA9FC9D.taxon	diagnosis	Diagnosis ROSTRUM. Curved and very long, nearly reaching tip of article 2 of peduncle of antenna 1.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE26819FEB9FC62CBA9FC9D.taxon	description	BODY SEGMENTS. All pereionites and pleonites with strong mid-dorsal tooth, without pair of dorsolateral teeth or projections (however there is a very faint trace of dorsolateral protrusions on posterior segments); pereionites without pair of ventrolateral teeth or projections; mid-dorsal tooth of pereionites nearly cylindrical, very long, with tip not acute; mid-dorsal tooth of pleonites 1 – 2 very narrowly triangular in lateral view, scarcely laterally compressed, with slight anterior notch; pleonite 3 with low rounded lobe followed by notch which is itself followed by a subacute broad triangular tooth. COXA 4. Anteriorly produced into a blunt angle, strongly projecting forward, ventrally produced into a fairly narrow, subacute tooth, with posteroventral border very concave. COXA 5. Produced into very long styliform tooth projecting obliquely backwards. UROSOMITE 1. With rounded dorsal process. PEDUNCLE OF ANTENNA 1. Not toothed. GNATHOPODS 1 – 2. Carpus and propodus robust; propodus distally expanding, with palm oblique. PEREIOPOD 5. Basis with posteroproximal tooth forming a blunt squared angle with the basis, with posterodistal corner not produced into a tooth. PEREIOPOD 6. Basis with posteroproximal acutely triangular tooth pointing obliquely, with posterodistal corner not produced into a tooth. PEREIOPOD 7. Basis with distinct notch forming a blunt obtuse angle on 0.8 of posterior border, with posterodistal corner not produced into a tooth. Colour pattern Body orange red. Rostrum laterally lined with white. Coxae 1 – 4 largely whitish. Posterior coxae and epimeral plates with diffuse whitish zones. Pereiopods largely whitish. Eyes orange. Body length Up to 70 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE26819FEB9FC62CBA9FC9D.taxon	distribution	Distribution Eastern shelf of Weddell Sea (De Broyer et al. 2007), Davis Sea: Tressler Bank (Pyper 2010); 254 – 1030 m (De Broyer et al. 2007).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE26819FEB9FC62CBA9FC9D.taxon	biology_ecology	Biology Analyses of digestive tract contents revealed the following items: diatoms, sponge spicules, fragments of hydroids of the genus Staurotheca, spicules of Clavularia, crustacean remains (mainly amphipods), polychaete setae, and sclerites of holothurians (Sigmodota contorta (Ludwig, 1875) as Taeniogyrus contortus) (Dauby et al. 2001 a). These authors conclude that E. rubrieques is rather an opportunistic feeder with both scavenging and predatory behaviour. Dauby et al. (2001 b) stated that it is anopportunistic predator. Klages (1991) observed that the species is sometimes parasitized by cryptoniscin isopods. Klages (1991) also observed that hatchlings of E. rubrieques climb on the back of their mother and remain there for a short period of time.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE26819FEB9FC62CBA9FC9D.taxon	discussion	Remarks The collection data of the specimens illustrated on figure 29 by Coleman (2007) are: 73 ° 12.39 ʹ S, 20 ° 46.24 ʹ W (eastern shelf of the Weddell Sea), 650 – 710 m, 22.1.1985, leg. Wägele (Coleman pers. com.). The collection data of the “ specimen 2 ” illustrated by Rauschert & Arntz (2015) were given in an early draft of their book accessed by us: ANT-XV / 3 stn 222. Its coordinates are 70 ° 49.1 ʹ S, 10 ° 39.2 ʹ W 70 ° 50.5 ʹ S, 10 ° 41.8 ʹ W (eastern Weddell Sea) at 234 – 385 m. The photograph of Pyper (2010: cover art of the Australian Antarctic Magazine, as “ amphipod ”) shows E. rubrieques. That specimen came from the Tressler bank, which considerably extends eastwards the distribution of the species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE06817FE19FCD6CD14FB30.taxon	description	urn: lsid: zoobank. org: act: 85 ED 0 CCE- 6 EB 7 - 4010 - AFB 3 - 29894 C 55616 C Figs 222 – 228	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE06817FE19FCD6CD14FB30.taxon	etymology	Etymology Xestus, - a, - um is a Latinization of the Greek adjective ξεστός, which means smooth or polished. The name alludes to the smooth body surface of the species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE06817FE19FCD6CD14FB30.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: dissected ♀, cruise PS 14, ANT-VII / 4, EPOS leg 3, stn 226, eastern Weddell Sea, 75 ° 15.9 ʹ S, 25 ° 58.3 ʹ W to 75 ° 15.7 ʹ S, 25 ° 54.2 ʹ W, 569 – 574 m, bottom trawl, 28 Jan. 1989, coll. C. De Broyer (RBINS, INV. 132946). Paratypes RV Polarstern cruises: SOUTHERN OCEAN: 1 juv., cruise PS 14, ANT-VII / 4, EPOS leg 3, stn 226, eastern Weddell Sea, 75 ° 15.9 ʹ S, 25 ° 58.3 ʹ W to 75 ° 15.7 ʹ S, 25 ° 54.2 ʹ W, 569 – 574 m, bottom trawl, 28 Jan. 1989, coll. C. De Broyer (RBINS, INV. 132947); 2 adult specs, cruise PS 14, ANT-VII / 4, EPOS leg 3, stn 226, eastern Weddell Sea, 75 ° 15.9 ʹ S, 25 ° 58.3 ʹ W to 75 ° 15.7 ʹ S, 25 ° 54.2 ʹ W, 569 – 574 m, bottom trawl, 28 Jan. 1989, subsample, coll. C. De Broyer (RBINS, INV. 132948); 1 ovigerous ♀, cruise PS 14, ANT-VII / 4, EPOS leg 3, stn 226, eastern Weddell Sea, 75 ° 15.9 ʹ S, 25 ° 58.3 ʹ W to 75 ° 15.7 ʹ S, 25 ° 54.2 ʹ W, 569 – 574 m, bottom trawl, 28 Jan. 1989, coll. C. De Broyer (RBINS, INV. 132949); 3 very large specs, presumably including specimen photographed on board, cruise PS 39, ANT-XIII / 3, EASIZ I, no locality, aquarium, 3 Mar. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132994); 1 spec., previously mixed with 9 E. inermis, misidentified as E. georgiana and used for gut content analysis, cruise PS 39, ANT-XIII / 3, EASIZ I, stn 12, GSN 5, eastern Weddell Sea, 73 ° 18.10 ʹ S, 21 ° 10.10 ʹ W to 73 ° 17.10 ʹ S, 21 ° 08.20 ʹ W, 457 – 459 m, bottom trawl, 14 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132997); 1 large spec., misidentified as E. georgiana and used for gut content analysis, cruise PS 39, ANT-XIII / 3, EASIZ I, stn 29, BPN 4, eastern Weddell Sea, 71 ° 31.50 ʹ S, 12 ° 25.50 ʹ W to 71 ° 30.30 ʹ S, 12 ° 27.80 ʹ W, 504 – 529 m, 28 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132998).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE06817FE19FCD6CD14FB30.taxon	description	Description ROSTRUM. Medium-sized, overreaching tip of article 1 of peduncle of antenna 1, anteriorly weakly curved, ventrally straight, subacute in lateral view; of medium width and with nearly straight converging borders in frontal view. EYE. Very large, broadly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 3 smooth; pereionite 4 to pleonite 3 with very low and very blunt mid-dorsal carina; profile of carinae of pereionites 4 – 7 weakly and regularly curved; profile of carinae of pleonites 1 – 2 straight, profile of carina of pleonite 3 straight with anterior low protrusion followed by shallow notch, then nearly straight (slightly irregular), posteriorly blunt; dorsolateral ornamentation absent. COXAE 1 – 3. Weakly carinate and distally subacute. COXA 4. Anterodorsal border nearly straight (very slightly concave), anteroventral border slightly but distinctly concave, these two borders being joined by a very large rounded lobe (anterior corner), which is weakly projecting forward; ventral corner forming a blunt squared angle (ventral projection well developed); lateral carina very obtuse, parallel and close to posteroventral border; posteroventral border distinctly concave. COXA 5. Broad, with surface smooth, with posteroventral corner forming a blunt-tipped nearly squared angle, not expanded laterally, not forming an angle in dorsal view (scarcely visible in dorsal view). COXA 6. With posteroventral corner broadly rounded, with low projection arising from its surface (forming a very low indistinct lobe in dorsal view). COXA 7. Posteriorly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle very obtusely rounded in plate 1, produced into a small tooth in plate 2 and into a medium-sized tooth in plate 3. UROSOME TOOTH PATTERN. Urosomite 1 with very low dorsal process: anteriorly nearly straight, apically very blunt, posteriorly regularly convex large and sharp narrow tooth pointing upwards; urosomite 3 with dorsolateral borders nearly straight. TELSON. Cleft on 0.2; tips of lobes very broad and very rounded, notch narrowly V-shaped. GNATHOPODS 1 – 2. Carpus and propodus broad; propodus expanding distally, palm distinct. PEREIOPODS 5 – 7. Merus, carpus and propodus of pereiopods 5 – 7 of medium width; dactylus long; basis of pereiopods 5 – 6 of normal width, with posteroproximal process present, sword-like, parallel to axis of basis, with posterodistal corner bluntly angular and pointing in posterior direction; basis of pereiopod 7 broad with posterior border very convex, with deep notch on distal 0.8, forming a blunt-tipped acute angle, with posterodistal corner very bluntly angular and pointing backwards. Colour pattern Body and appendages with a homogeneous and diffuse orange dotted / mottled pattern; legs with colour more intense; eyes reddish. Body length Up to 55 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE06817FE19FCD6CD14FB30.taxon	distribution	Distribution Eastern shelf of Weddell Sea, 457 – 574 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE06817FE19FCD6CD14FB30.taxon	biology_ecology	Biology Dauby et al. (2001 a, as E. georgiana) state that “ this weakly mobile large amphipod (up to 40 mm) can be found, sometimes in abundance, on coarse sediment bottoms or at the base of animal colonies ”. According to them, “ digestive tract analyses of 31 specimens revealed a wide variety of food items identifiable by hard remnants: crustaceans (mysids and amphipods), polychaetes (setae of terebellids), holothurioid ossicles and hydrozoan perisarcs; planktonic items (diatoms, radiolarians, foraminifers) also form a significant part of the diet; finally, as for the other deposit-feeders, sponge spicules (which form a major component of bottom mats) and mineral particles complete the food, the latter providing evidence of feeding on the sediment; both these items, albeit ‘ inorganic’, are likely to be of considerable nutritional value, since they may be densely coated with bacteria, the importance of which has been shown in the Weddell Sea. ” Dauby et al. (2001 b) concluded that E. xesta (as E. georgiana) is a deposit feeder.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE06817FE19FCD6CD14FB30.taxon	discussion	Remarks Epimeria xesta is sympatric with E. angelikae, though they have different bathymetric ranges, with E. angelikae being found at greater depths: 781 – 1194 m vs 457 – 574 m. The morphologically most similar species to E. xesta is E. cyphorachis from the South Shetland Islands and from the tip of the Antarctic Peninsula. In E. xesta, the rostrum is narrower in frontal view than in E. cyphorachis, the ventral corner of coxa 4 bluntly angulate instead of being sharply angulate, and the posterior notch of pereiopods 5 – 7 is narrower.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEE6816FDE8FB08CD55FE31.taxon	diagnosis	Character states and colour pattern Similar to Epimeria gargantua but with a different colour pattern. Background colour of the body: white; all pereionites with two pairs of small red spots disposed transversally; pereionite 1 + coxa 1, pereionite 4 + coxa 4, pereionite 7 + coxa 7 with broad orange red stripe; urosomite 1 and basis of pereiopod 7 orange red; basis of pereiopods 5 – 6 white; antennae and pereiopods (except basis of pereiopods 5 – 7) pink; mouthparts purple; eyes red.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEE6816FDE8FB08CD55FE31.taxon	distribution	Distribution PS 61, ANT-XIX / 5, LAMPOS, stn 229 - 1, Bruce Ridge [a geomorphologic elevation situated midway between the South Orkney Islands and the southern tip of the archipelago of the South Sandwich Islands], 60 ° 08.77 ʹ S, 34 ° 54.83 ʹ W to 60 ° 07.81 ʹ S, 34 ° 56.17 ʹ W, 362 – 371 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEE6816FDE8FB08CD55FE31.taxon	discussion	Remarks Rauschert & Arntz (2015) do not indicate the coordinates of their ‘ Epimeria robustoides 1 ’, but this information is given in an early draft of the book made available to the authors: ANT-XIX / 5, LAMPOS, stn 229 - 1, Bruce Ridge. It is not known where the specimen is stored; in November 2015, it was not deposited at the Berlin Museum (Coleman pers. com.). As its colour pattern, with large red patches, is very different from the purely white E. gargantua sp. nov. found more southernly, this specimen possibly belongs to a distinct species. It should be noted that the colour pattern of all E. gargantua sp. nov. collected during the cruises ANT-XXIII / 8 and ANT-XXIX / 3 (20 specimens) was examined on board and exhibited no variation.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEF6815FDCAFE09CA60F81C.taxon	description	urn: lsid: zoobank. org: act: D 1 B 62023 - FFE 4 - 4 AEB-A 345 - 89459 C 1063 AC	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEF6815FDCAFE09CA60F81C.taxon	etymology	Etymology Combination of laevis, smooth in Latin, and Epimeria. That name, which is feminine, alludes to the absence of ornamentation in the species of that subgenus.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEF6815FDCAFE09CA60F81C.taxon	type_taxon	Type species Epimeria walkeri (K. H. Barnard, 1930).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEF6815FDCAFE09CA60F81C.taxon	description	Description Body opaque, with teguments moderately calcified. Rostrum small but not minute (reaching about mid of article 1 of antenna 1). Eyes not conical. Pereionites and pleonites completely smooth (only pleonite 3 can be posterodorsally produced into a low bump). Coxae 1 – 4 with sharp or blunt tip. Coxae 1 – 3 not keeled along their axis. Coxa 1 with anterior margin proximally concave and with subdistal dilatation. Coxa 4 without carina or groove; posteroventral border straight or nearly so. Coxae 5 – 6 toothless. Mid of posterior border of epimeral plates 1 – 3 not produced into a tooth. Posteroventral tooth of epimeral plate 3 small. Dorsal process of urosomite 1 produced into a large blunt tooth, of which the anterior border is shorter than the posterior border (the posterior border of the tooth extend to the posterior border of urosomite 1). Urosomite 2 without pair of small teeth pointing upwards. Lateral borders of urosomite 3 posteriorly terminated into a blunt angle. Peduncle of antenna 1 without teeth. Mandible with molar process drawn out, without triturative surface. Lower lip with wide hypopharyngeal gap. Palp of maxilliped with 4 articles. Gnathopods of normal size, with carpus and propodus very stout, with palm reduced or normal; propodus not expanded distally; dactylus very stout, with long slender posterior teeth perpendicular (or nearly perpendicular) to axis of dactylus (as in a comb). Basis of pereiopods 5 – 6 fairly broad, with indistinct trace of posteroproximal process, with or without trace posterodistal blunt process projecting posteriorly. Basis of pereiopod 7 with posterior border convex along all its length or with slight distal concavity, with large distal lobe reaching 0.3 of merus. Merus of pereiopod 7 broad. In pereiopod 7, the sum of the merus, carpus and propodus is short: only about as long as basis or a little bit more. Dactylus of pereiopods 5 – 7 short. Body length The maximum body length recorded in Laevepimeria species ranges between 15 and 32 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEF6815FDCAFE09CA60F81C.taxon	biology_ecology	Ecology Benthic, 170 – 889 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEF6815FDCAFE09CA60F81C.taxon	distribution	Distribution Circum-Antarctic, as far north as Elephant Island.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEF6815FDCAFE09CA60F81C.taxon	discussion	Remarks Laevepimeria subgen. nov. and Epimeriella form a large clade together. These subgenera share synapomorphies in the morphology of the mandible (molar process non triturative) and the lower lip (wide hypopharyngeal gap), and were formerly grouped into the genus Epimeriella s. lat. However, Laevepimeria subgen. nov. and Epimeriella s. str. exhibit striking differences in the morphology of the body, gnathopods and pereiopods. These differences are presumably related to their benthic lifestyle (Laevepimeria subgen. nov.) versus a pelagic or semi-pelagic lifestyle (Epimeriella). These subgenera can be identified without dissection, which was not possible with the former concept of Epimeriella.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFED6813FE16FEE3CE4CF862.taxon	description	urn: lsid: zoobank. org: act: 4328 C 52 C- 34 BA- 4 DDF-A 984 - 7 BDE 24 BADE 7 F Figs 229 – 234	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFED6813FE16FEE3CE4CF862.taxon	etymology	Etymology Combination of the prefix an-, which means ‘ lack of’ and of the Greek noun, όδούς, which means ‘ teeth’. The name, which is an apposition, alludes to the absence of tooth in the species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFED6813FE16FEE3CE4CF862.taxon	materials_examined	Type material Holotype RV Aurora Australis cruises: SOUTHERN OCEAN: ♀, cruise CEAMARC, sample CEAMARC 2724, stn 71 EV 447, Adélie Coast, 66 ° 24 ʹ 00 ″ S, 140 ° 32 ʹ 21 ″ E, 683 – 791 m, beam trawl, 14 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4336) [extraction M 13; Genbank nr, COI: KU 870880, 28 S: KU 759664]. Paratypes RV Aurora Australis cruises: SOUTHERN OCEAN: 1 spec., cruise CEAMARC, sample CEAMARC 1317, stn 51 AEV 215, Adélie Coast, 66 ° 44 ʹ 52 ″ S, 145 ° 26 ʹ 40 ″ E, 525 – 553 m, beam trawl, 30 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4275); 1 adult spec., cruise CEAMARC, sample CEAMARC 1384, stn 50 AEV 220, Adélie Coast, 66 ° 45 ʹ 09 ″ S 145 ° 20 ʹ 04 ″ E, 567 – 604 m, beam trawl, 30 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4331) [extraction M 12; Genbank nr, COI: KU 870879, 28 S: KU 759663]; 1 ♀, cruise CEAMARC, sample CEAMARC 2724, stn 71 EV 447, Adélie Coast, 66 ° 24 ʹ 00 ″ S, 140 ° 32 ʹ 21 ″ E, 683 – 791 m, beam trawl, 14 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 7335) [removed from MNHN-IU- 2014 - 4336].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFED6813FE16FEE3CE4CF862.taxon	description	Description HEAD + ROSTRUM. Normally curved in lateral view. ROSTRUM. In lateral view short and very broad, reaching mid of article 1 of peduncle of antenna 1, anteriorly distinctly curved, ventrally weakly convex, tip subacute; in frontal view very broad and with nearly straight converging borders, tip blunt. EYES. Medium-sized, elliptic, laterally oriented (interocular distance about 3 × as wide as eye size when seen in frontal view). PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 7 totally smooth; pleonites 1 – 2 with trace of mid-dorsal keel and with trace of posterior bump; pleonite 3 with trace of mid-dorsal carina and with posterior bump. COXAE 1 – 3. Tip sharp. COXA 4. Broad, anterodorsal border straight; anteroventral border nearly straight (inconspicuously convex), these two borders being joined by broad rounded convexity, which is distinctly projecting forward; anterodorsal border 1.35 × as long as anteroventral border; posteroventral border nearly straight (inconspicuously sinuate). COXA 5. Very broad, posteroventral corner very broadly rounded in lateral view, very obtusely rounded in dorsal view (almost not projecting laterally). COXA 6. Posterior border weakly convex; posteroventral corner very broadly rounded. COXA 7. Posterior border nearly straight (inconspicuously concave); posteroventral corner broadly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle very obtusely rounded in plate 1 (without any trace of tooth), produced into a small tooth in plates 2 – 3. UROSOME TOOTH PATTERN. Urosomite 1 with distinct asymmetrical dorsal process, anteriorly nearly straight, apically blunt, posteriorly strongly convex; urosomite 3 with dorsolateral borders weakly convex, with tip broadly rounded. TELSON. Cleft on 0.2; lobes laterally very convex, medially nearly straight, with tips subacute; notch with borders weakly convergent and end rounded. GNATHOPODS 1 – 2. Carpus and propodus short and very broad; propodus not expanding distally, palm very reduced (dactylus 4 × as long as palm; gnathopods achelate or nearly so). PEREIOPOD 4. Merus, carpus and propodus long and slender, dactylus short. PEREIOPOD 5. Basis of normal width, with posteroproximal process absent, with posterodistal corner forming a very obtuse angle (not projecting posteriorly) of which the tip is very blunt; merus, carpus and propodus long and slender, dactylus short. PEREIOPOD 6. Basis of normal width, with posteroproximal process absent, with posterodistal corner very broadly rounded (not angulate); merus, carpus and propodus long and slender, dactylus short. PEREIOPOD 7. Basis broad; posterior border weakly convex and slightly diverging in proximal 0.8, at this level forming a blunt but strong angular discontinuity and abruptly and very strongly converging (border becoming inconspicuously concave, terminated into a small, blunt but distinctly protruding lobe; merus very broad and short, carpus of medium width and short, propodus slender and short, dactylus short. Colour pattern Uniformly pale yellowish; eyes blood red. Body length Up to 32 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFED6813FE16FEE3CE4CF862.taxon	distribution	Distribution Adélie Coast, 525 – 791 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFED6813FE16FEE3CE4CF862.taxon	discussion	Remarks Epimeria anodon sp. nov. and E. walkeri are very similar in morphology. Epimeria anodon sp. nov. has smaller eyes and the posterodistal corner of the basis of its pereiopods 5 and 6 is very broadly rounded, which is not the case for the other two species. In dorsal view, pleonites 1 – 3 are also less smooth than in E. walkeri.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEB6810FE25FEE3CCB8FD82.taxon	description	urn: lsid: zoobank. org: act: A 77 C 9 AD 4 - 391 F- 4101 - 8 DFA-F 15 A 45185 A 0 E Figs 235 – 240	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEB6810FE25FEE3CCB8FD82.taxon	etymology	Etymology Cinderella, heroin of humble origin in a well-known folk tale. The name, which is a noun in apposition, alludes to the modest size and the absence of ornamentation of the species, which contrasts with the extravagant adornment and the impressive size of many ‘ rival’ Epimeria species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEB6810FE25FEE3CCB8FD82.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: ♂, cruise PS 81, ANT-XXIX / 3, stn 188 - 5, south east of Dundee Island, 63 ° 50.92 ʹ S, 55 ° 37.66 ʹ W to 63 ° 50.93 ʹ S, 55 ° 37.52 ʹ W, 402 – 407 m, Rauschert dredge, 20 Dec. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 132656) [extraction I 15; Genbank nr, COI: KU 870841, 28 S: KU 759617]. Paratypes RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., cruise PS 69, ANT-XXIII / 8, stn 605 - 1, Elephant Island, 61 ° 20.35 ʹ S, 55 ° 29.16 ʹ W to 61 ° 19.98 ʹ S, 55 ° 32.67 ʹ W, 146 – 151 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122528); 1 spec., cruise PS 81, ANT-XXIX / 3, stn 185 - 3, south east of Dundee Island, 63 ° 51.53 ʹ S, 55 ° 40.74 ʹ W to 63 ° 51.53 ʹ S, 55 ° 40.43 ʹ W, 253 – 255 m, extremely fine sand mixed with some mud and gravel, Rauschert dredge, 19 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122965); 1 spec., cruise PS 81, ANT-XXIX / 3, stn 234 - 5, north of Livingstone Island, 62 ° 17.36 ʹ S, 61 ° 12.06 ʹ W to 62 ° 17.31 ʹ S, 61 ° 12.63 ʹ W, 248 – 251 m, mud with a lot of specimens of a small reddish ophiuroid with very long and flexible legs, Agassiz trawl, 7 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122949) [extraction ANT 43; Genbank nr, COI: KU 870829, 28 S: KU 759602]. RV James Clark Ross cruises: SOUTHERN OCEAN: 2 specs, cruise JR 144, stn EI-EBS- 4 - supra, south of Elephant Island, 61 ° 33 ’ 544 ” S, 55 ° 20 ’ 379 ” W to 61 ° 33 ’ 637 ” S, 55 ° 20 ’ 901 ” W, epibenthic sledge, 270 m, 12 Mar. 2006, coll. BAS (RBINS, INV. 132959) [extraction I 4; Genbank nr, COI: KU 870848, 28 S: KU 759625]. Other material RV Seatruck, cruise REVOLTA II: SOUTHERN OCEAN: 1 spec., stn REVO _ 140, Collect _ ID: REVO _ 140, Adélie Coast, 66 ° 37 ʹ 53.4 ″ S, 139 ° 51 ʹ 18 ″ E, 106 – 172 m, 12 Jan. 2011, coll. N. Améziane, C. Gallut and A. C. Lautrédou (MNHN-IU- 2016 - 6885).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEB6810FE25FEE3CCB8FD82.taxon	description	Description HEAD + ROSTRUM. Very strongly curved in lateral view. ROSTRUM. In lateral view short and broad, reaching 0.6 of article 1 of peduncle of antenna 1, anteriorly and ventrally straight, subacute; in dorsal view narrow, with straight converging borders, with tip blunt. EYES. Huge, broadly elliptic, partly anteriorly oriented (interocular distance about 1.5 × as wide as eye size when seen in frontal view). PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 7 and pleonites 1 – 2 totally smooth; pleonite 3 with weak posterior bump. COXAE 1 – 3. Coxa 1 blunt-tipped, coxa 2 with tip subacute, coxa 3 blunt-tipped. COXA 4. Narrow; anterodorsal border straight; anteroventral border nearly straight (inconspicuously convex), these two borders being joined by very broad rounded convexity, which is distinctly projecting forward; anterodorsal border 1.75 × as long as anteroventral border; posteroventral border nearly straight. COXA 5. Broad, posteroventral corner rounded in lateral view, not projecting laterally. COXA 6. Posterior border weakly convex; posteroventral corner very broadly rounded. COXA 7. Posterior border nearly straight (inconspicuously concave); posteroventral corner broadly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle very obtusely rounded in plate 1 (without any trace of tooth), produced into a small tooth in plates 2 – 3. UROSOME TOOTH PATTERN. Urosomite 1 with distinct asymmetrical dorsal process, anteriorly nearly straight, apically bluntly angulate, posteriorly weakly convex; urosomite 3 with dorsolateral borders very weakly convex, with tip very bluntly angulate. TELSON. Cleft on 0.2; lobes laterally very convex, medially nearly straight, with tips rounded; notch with borders weakly convergent and end rounded. GNATHOPODS 1 – 2. Carpus and propodus short and very broad; propodus not expanding distally, palm not reduced. PEREIOPOD 4. Merus, carpus and propodus long and slender, dactylus short. PEREIOPOD 5. Basis of normal width, with posteroproximal process absent, with posterodistal corner forming a blunt squared angle (not projecting posteriorly); merus, carpus and propodus long and slender, dactylus short. PEREIOPOD 6. Basis of normal width, with posteroproximal process absent, with posterodistal corner forming a blunt squared angle (not projecting posteriorly); merus, carpus and propodus long and slender, dactylus short. PEREIOPOD 7. Basis broad; posterior border strongly convex along all its length, terminated in a large rounded lobe projecting ventrally; merus very broad and short, carpus of medium width and short, propodus slender and short, dactylus short. Body length Up to 15 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEB6810FE25FEE3CCB8FD82.taxon	distribution	Distribution Elephant Island, tip of Antarctic Peninsula, Adélie Coast; 106 – 270 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFEB6810FE25FEE3CCB8FD82.taxon	discussion	Remarks Epimeria cinderella sp. nov. can be easily distinguished from the related E. anodon sp. nov. and E. walkeri by its much narrower and more asymetrical coxa 4. The gnathopods and the basis of pereiopod 7 are also different. Epimeria cinderella sp. nov. seems more similar to Epimeria (Laevepimeria) sp. subgen. nov., which is only known from a very small juvenile. The tip of the coxae 1 – 2 of the young Epimeria (Laevepimeria) sp. subgen. nov. examined are, however, much more rounded than in Epimeria cinderella sp. nov., but this difference might disappear in adults.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE9680EFE74FDDCCE9BF923.taxon	description	Figs 241 – 249	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE9680EFE74FDDCCE9BF923.taxon	materials_examined	Material examined RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 8, AGT 2, eastern Weddell Sea, 71 ° 18.70 ʹ S, 12 ° 17.10 ʹ W to 71 ° 18.45 ʹ S, 12 ° 16.30 ʹ W, 170 – 174 m, 9 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132683); 1 spec., alcohol-fixed, cruise PS 69, ANT-XXIII / 8, stn 603 - 5, eastern Weddell Sea, 70 ° 30.99 ʹ S, 08 ° 48.08 ʹ W to 70 ° 30.40 ʹ S, 08 ° 48.13 ʹ W, 274 – 297 m, sponge bottom, Agassiz trawl, 7 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122485); 4 specs, cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122516); 1 spec., cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, stn 48 - 1, eastern Weddell Sea, 70 ° 23.94 ʹ S, 8 ° 19.14 ʹ W to 70 ° 23.89 ʹ S, 8 ° 18.67 ʹ W, 595 – 602 m, bryozoan bottom (exceptional diversity of bryozoans), Agassiz trawl, 12 Jan. 2008, coll. H. Robert (RBINS, INV. 132448); 1 spec., cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 222 - 5, southwest of King George Island, 62 ° 17.60 ʹ S, 58 ° 41.19 ʹ W to 62 ° 17.49 ʹ S, 58 ° 41.46 ʹ W, 445 – 889 m, Agassiz trawl, 23 Feb. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 132667) [extraction A 6; Genbank nr, COI: KU 870819, 28 S: KU 759591]; 1 spec., specimen missing, DNA extraction products only, PS 77, ANT-XXVII / 3, CAMBIO, stn 288 - 3, Eastern Weddell Sea, 70 ° 56.40 ʹ S, 10 ° 32.60 ʹ W to 70 ° 56.42 ʹ S, 10 ° 32.25 ʹW, 303 – 311 m, Rauschert Dredge, 30 Mar. 2011, coll. C. Havermans and H. Robert [extraction Ex 169; Genbank nr, COI: KU 870836, 28 S: KU 759610]; ♂, cruise PS 81, ANT-XXIX / 3, stn 217 - 7, Bransfield Strait, 62 ° 53.64 ʹ S, 58 ° 12.52 ʹ W to 62 ° 53.64 ʹ S, 58 ° 12.37 ʹ W, 387 – 395 m, heterogeneous bottom (black muddy sand and gravel; small stones with incrusting bryozoans; gorgonians; sea urchins), Rauschert dredge, 2 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122944) [extraction ANT 42; Genbank nr, COI: KU 870828, 28 S: KU 759601].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE9680EFE74FDDCCE9BF923.taxon	description	RV Aurora Australis cruises: SOUTHERN OCEAN: 1 spec., cruise CEAMARC, sample CEAMARC 631, stn 4 EV 112, Adélie Coast, 66 ° 20 ʹ 16 ″ S, 141 ° 59 ʹ 17 ″ E, 237 – 257 m, beam trawl, 26 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4324); 1 spec., cruise CEAMARC, sample CEAMARC 1643, Adélie Coast, 66 ° 34 ʹ 30 ″ S, 145 ° 01 ʹ 15 ″ E, 429 – 451 m, beam trawl, 3 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4323); 4 specs, cruise CEAMARC, sample CEAMARC 3532, Adélie Coast, 65 ° 29 ʹ 29 ″ S, 139 ° 18 ʹ 37 ″ E, 397 – 411 m, beam trawl, 17 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4352). Description HEAD + ROSTRUM. Very curved in lateral view. ROSTRUM. In lateral view short and broad, reaching mid of article 1 of peduncle of antenna 1, anteriorly weakly curved, ventrally straight, acute-tipped; in frontal view narrow and with straight converging borders, with tip acute. EYES. Huge, broadly elliptic, largely oriented forward (interocular distance about 1.3 × as wide as eye size when seen in frontal view). PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 7 and pleonites 1 – 2 totally smooth; pleonite 3 with posterior bump. COXAE 1 – 3. Tip blunt. COXA 4. Broad, anterodorsal border nearly straight (inconspicuously concave); anteroventral border weakly convex, these two borders being joined by broad rounded convexity, which is distinctly projecting forward; anterodorsal border 1.12 × as long as anteroventral border; posteroventral border nearly straight (inconspicuously sinuate). COXA 5. Very broad, posteroventral corner very broadly rounded in lateral view, very obtusely rounded in dorsal view (almost not projecting laterally). COXA 6. Posterior border weakly convex; posteroventral corner very broadly rounded. COXA 7. Posterior border nearly straight (inconspicuously concave); posteroventral corner broadly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle very obtusely rounded in plate 1 (without any trace of tooth), produced into a small tooth in plates 2 – 3. UROSOME TOOTH PATTERN. Urosomite 1 with distinct asymmetrical dorsal process, anteriorly nearly straight, apically blunt, posteriorly moderately convex; urosomite 3 with dorsolateral borders nearly straight (very weakly convex), with tip bluntly angulate. TELSON. Cleft on 0.15; lobes laterally very convex, medially nearly straight, with tips blunt; notch with borders weakly convergent and end rounded. GNATHOPODS 1 – 2. Carpus and propodus short and very broad; propodus not expanding distally, palm very reduced (dactylus 4 × as long as palm; gnathopods achelate or nearly so). PEREIOPOD 4. Merus, carpus and propodus long and slender, dactylus short. PEREIOPOD 5. Basis of normal width, with posteroproximal process absent, with posterodistal corner forming a blunt squared angle (slightly projecting posteriorly); carpus and propodus long and slender, dactylus short. PEREIOPOD 6. Basis of normal width, with posteroproximal process absent, with posterodistal corner forming a sharp squared angle; carpus and propodus long and slender, dactylus short. PEREIOPOD 7. Basis broad; posterior border weakly convex and slightly diverging in proximal 0.8, at this level the curvature slightly and gradually increases and becomes convergent with the anterior border; just before tip the curve becomes slightly concave; the posterior border of the basis is terminated into a small, sharp tooth (forming a small squared angle); merus very broad and short, carpus of medium width and short, propodus slender and short, dactylus short. Colour pattern Background whitish or very pale greyish; pereionites 3 – 7 yellowish / brownish; indistinct pale orange marks present here and there; peduncle of antenna 1 tinged with pale orange; urosome (especially urosomite 1) and tailfan tinged with orange; a proximal orange mark on the basis of pereiopod 7; eyes red. Body length Up to 30 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE9680EFE74FDDCCE9BF923.taxon	distribution	Distribution Type locality: Ross Sea, McMurdo Sound, 256 – 379 m (K. H. Barnard 1930). Other records: King George Island and Bransfield Strait, eastern shelf of the Weddell Sea, Adélie Coast; 170 – 889 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE9680EFE74FDDCCE9BF923.taxon	biology_ecology	Biology Dauby et al. (2001 a) provide some information on the gut content of Epimeria walkeri (as Epimeriella walkeri). According to them, the commonest items were ophiuroids (ossicles and parts of arms), striated muscle and diatoms. Less prevalent were sponge spicules and cnidocysts, while crustacean pieces and holothurian ossicles were infrequent. Dauby et al. (2001 b) concluded that it is a macropredator / scavenger.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFE9680EFE74FDDCCE9BF923.taxon	discussion	Remarks The illustrations of E. walkeri given by K. H. Barnard (1930) are difficult to understand. On his figures, the rostrum appears as broader than in specimens examined by us, but this might be related the orientation of the specimen. On the other hand, the shapes of coxa 4 and of the basis of pereiopods 5 – 7 are similar to that of our specimens. Our identification is only tentative. A comparison between topotypical E. walkeri (from the Ross Sea) and specimens from other Antarctic seas should be carried out whenever possible. The collection details of the specimen illustrated by Coleman (2007: 60, fig. 35 a – b) are: 72 ° 27.7 ʹ S, 17 ° 42.32 ʹ W (eastern shelf of the Weddell Sea), 240 – 254 m, Agassiz trawl, leg. Wägele, 15 Feb. 1985 (Coleman pers. com.). The station of the specimen illustrated by Rauschert & Arntz (2015) was indicated in an early draft made available to the authors. It is ANT-XVII / 3, stn 85 [85 - 1], of which the coordinates are: 71 ° 11.30 ʹ S, 12 ° 15.40 ʹ W to 71 ° 12.19 ʹ S, 12 ° 19.01 ʹ W (eastern shelf of the Weddell Sea), 309 – 318 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF7680CFDE8F93BCD54FDC7.taxon	description	Fig. 250	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF7680CFDE8F93BCD54FDC7.taxon	materials_examined	Material examined RV Polarstern cruise: SOUTHERN OCEAN: 1 tiny juv., cruise PS 81, ANT-XXIX / 3, stn 197 - 5, Bransfield Strait, 62 ° 44.73 ʹ S, 57 ° 26.79 ʹ W to 62 ° 45.05 ʹ S, 57 ° 26.68 ʹ W, 258 – 273 m, Agassiz trawl, 25 Feb. 2013 (RBINS, INV. 122932) [extraction K 40; Genbank nr, COI: KU 870873, 28 S: KU 759653].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF7680CFDE8F93BCD54FDC7.taxon	description	Description HEAD + ROSTRUM. Weakly curved in lateral view. ROSTRUM. In lateral view short and very broad, reaching about 0.6 of article 1 of peduncle of antenna 1, anteriorly weakly convex and ventrally straight, blunt-tipped. EYES. Huge, broadly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 7 and pleonites 1 – 2 totally smooth; pleonite 3 missing. COXAE 1 – 3. Tip broadly rounded. COXA 4. Fairly narrow; anterodorsal border straight; anteroventral border slightly convex, these two borders being joined by very broad rounded convexity, which is distinctly projecting forward (anterior corner and anteroventral border forming a regular curve, without discontinuity); anterodorsal border 1.4 x as long as anteroventral border; ventral corner bluntly and obtusely angulate; posteroventral border nearly straight. COXA 5. Broad, posteroventral corner very broadly rounded in lateral view. COXA 6. Posterior border weakly convex; posteroventral corner very broadly rounded. COXA 7. Posterior border nearly straight (inconspicuously concave); posteroventral corner broadly rounded. EPIMERAL PLATES 1 – 2. Posteroventral angle very obtusely rounded in plate 1 (without any trace of tooth), produced into a small tooth in plate 2. UROSOME. Missing. GNATHOPODS 1 – 2. Carpus and propodus short and very broad; propodus not expanding distally, palm not reduced. PEREIOPOD 6. Basis of normal width, with posteroproximal process absent, with posterodistal corner rounded or very bluntly angulate. PEREIOPOD 7. Basis broad; posterior border strongly convex along all its length, terminated in a large rounded lobe projecting ventrally. Body length 5 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF7680CFDE8F93BCD54FDC7.taxon	distribution	Distribution Tip of Antarctic Peninsula: Bransfield Strait, 258 – 273 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF7680CFDE8F93BCD54FDC7.taxon	discussion	Remarks A COI phylogenetic analysis (Verheye et al. 2016 a) indicates that this specimen would belong to a distinct species of the subgenus Laevepimeria subgen. nov. Since it is an incomplete and very small juvenile (about 5 mm long), it is considered inappropriate for formal description and naming. Only a photograph and a description of the morphological details visible without dissection are given herein. Within Laevepimeria subgen. nov., this specimen is more similar to E. cinderella sp. nov. regarding the shape of its coxa 4 and of the basis of its pereiopod 7. On the other hand, the tip of coxae 1 – 2 are broadly rounded, whilst they are subacute in E. cinderella sp. nov. However, these structures might differ in adults, as seen in other Epimeria.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF5680BFDEBFD9DCA60FAEE.taxon	type_taxon	Type species Metepimeria acanthurus Schellenberg, 1931.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF5680BFDEBFD9DCA60FAEE.taxon	description	Description Body opaque, with teguments strongly calcified. Rostrum medium-sized. Eyes not conical. Pleonites and at least posterior pereionites with low mid-dorsal carina, which is sometimes posteriorly produced into a tooth pointing backwards; some carinae with slight or strong median concavity. Dorsolateral processes (small teeth or low carinae) always present on pleosomites and posterior pereionite (s); some body segments have 2 or more dorsolateral processes, some in longitudinal arrangement. Pereionites 1 – 7 without tooth or protrusion just above connection with coxa. Coxae 1 – 4 with sharp to fairly sharp tip. Coxae 1 – 3 distinctly keeled along their axis. Coxa 4 with groove along the posteroventral border; this groove might be limited by a distinct carina; when present this carina never bears a tooth projecting laterally; posteroventral border concave. Coxae 5 – 6 without tooth or distinct protrusion. Mid of posterior border of epimeral plates 1 – 3 not produced into a tooth. Posteroventral tooth of epimeral plate 3 small to medium-sized. Dorsal process of urosomite 1 produced into a sharp tooth directed upwards. Urosomite 2 without pair of small teeth pointing upwards. Lateral borders of urosomite 3 posteriorly terminated into an acute angle. Peduncle of antenna 1 without teeth. Mandible with molar process triturative. Lower lip with narrow (V-shaped) hypopharyngeal gap. Palp of maxilliped with 3 or 4 articles. Gnathopods of normal size, with carpus and propodus elongate, with palm very reduced, forming a weak oblique convexity in continuity with the axis of the propodus (gnathopods nearly achelate); propodus not expanded distally; ornamentation of posterior border of dactylus weak. Basis of pereiopods 5 – 6 broad, with posteroproximal process present, of variable development (strong nearly dentiform lobe at the base of basis; weak lobe at the base of basis; or a low lobe occupying half of posterior border), with or without posterodistal tooth or lobe (if present not distinctly projecting posteriorly). Posterior border of basis of pereiopod 7 with distinct median blunt angle; the distal half is slightly to distinctly concave; distal corner rounded. Dactylus of pereiopods 5 – 7 short. Body length The body length recorded in Metepimeria species ranges between 15 and 22.6 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF5680BFDEBFD9DCA60FAEE.taxon	biology_ecology	Ecology. Benthic, 27 – 1025 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF5680BFDEBFD9DCA60FAEE.taxon	distribution	Distribution Sub-Antarctic and low Antarctic seas: Patagonia; Falkland Islands; Shag Rocks; South Georgia; Macquarie Ridge.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF5680BFDEBFD9DCA60FAEE.taxon	discussion	Remarks Metepimeria was initially erected as a monotypic genus for the Magellanic species Epimeria acanthurus, based on a single character state: the absence of dactylus on the palp of the maxilliped. This character state is most likely autapomorphic, and the recognition of a separate genus for E. acanthurus is not supported here. Epimeria acanthurus is morphologically similar to Epimeria intermedia (South Georgia endemic) and E. ashleyi from Macquarie Ridge. In the present paper, Metepimeria is redefined, as a subgenus for these three species only. However, the New Zealand species, E. sophie Lörz & Coleman, 2014 (see Lörz & Coleman 2014) exhibits morphological similarities with them and a future transfer of E. sophie to Metepimeria seems possible.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF2680AFE37FA78CD3AFD60.taxon	description	Body length 22 mm (Coleman 2007). Colour pattern White with scarce suffusions of orange; eyes reddish (photograph of Rauschert & Arntz 2015 as ‘ Epimeria inermis 1 ’).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF2680AFE37FA78CD3AFD60.taxon	distribution	Distribution Magellanic Region: Patagonia, Falkland Islands, 27 – 494 m (De Broyer et al. 2007), Burdwood Bank, 296 – 299 m (see remarks).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF2680AFE37FA78CD3AFD60.taxon	discussion	Remarks The specimen labelled ‘ Epimeria inermis 1 ’ in Rauschert & Arntz (2015) does not correspond to the description of E. inermis: it exhibits important differences in the shape of coxa 4, and the dorsal profiles of pleonite 3 and urosomite 1. The morphology of this specimen is more similar to E. acanthurus as illustrated by Coleman (2007), and it is therefore identified as such. Rauschert & Arntz (2015) do not indicate the collection details of their specimen, but this information was found in an early draft of their book made available to the authors: ANT XIX [obviously ANT-XIX / 5], LAMPOS, station 153 [i. e., 153 - 1]. The coordinates of this station are: 54 ° 32.75 ʹ S 56 ° 09.84 ʹ W to 54 ° 31.22 ʹ S 56 ° 08.93 ʹ W [Burdwood Bank], 296 – 299 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF3680AFDE2FCF9CDECFBD8.taxon	description	Body length 22.6 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF3680AFDE2FCF9CDECFBD8.taxon	distribution	Distribution Hjort Seamount, Macquarie Ridge, 676 – 1025 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF3680AFE60FB91CC7BF897.taxon	description	Body length 18 mm (Schellenberg 1931).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF3680AFE60FB91CC7BF897.taxon	distribution	Distribution South Georgia: Cumberland Bay, 75 m (Schellenberg 1931).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF3680AFE60FB91CC7BF897.taxon	discussion	Remarks See notes on Epimeria (Metepimeria) intermedia forma B.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF36809FE60F8AECB4BFB3C.taxon	description	Body length 15 mm (K. H. Barnard 1932). Colour pattern Pure white (Rauschert & Arntz 2015: 60, as Epimeria sp. n. 2).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF36809FE60F8AECB4BFB3C.taxon	distribution	Distribution South Georgia, 88 – 273 m (K. H. Barnard 1932); Shag Rocks, 284 m (see remarks).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF36809FE60F8AECB4BFB3C.taxon	discussion	Remarks There are substantial differences between the illustrations of E. intermedia given by Schellenberg (1931) and Coleman (2007) on one hand, and those of K. H. Barnard (1932). Coleman (2007) questions the conspecificity of these specimens. We agree and treat them as separate forma A and B. However we refrain to attribute them a formal taxonomic status, as no direct examination of these specimens was possible. Rauschert & Arntz (2015) gave a colour photograph of a juvenile (6 mm) named “ Epimeria sp. n. 2 ”, which corresponds to the drawings of Epimeria intermedia forma B given by K. H. Barnard (1932). The collection details of the specimen were given in an early draft of the book: ANT-XIX / 5 in stn 169 - 1, 53 ° 22.94 ʹ S, 42 ° 41.37 ʹ W to 53 ° 22.89 ʹ S, 42 ° 41.50 ʹ W (Shag Rocks), at 284 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF06807FDD5FB35CA61F8F2.taxon	type_taxon	Type species	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF06807FDD5FB35CA61F8F2.taxon	description	Description Body opaque, with teguments strongly calcified, very strongly ornamented and sculptured by teeth and carinae. Rostrum long. Eyes conical. Pleonites 1 – 3 and pereionites 1 – 7 with strong mid-dorsal tooth. Each pereionite is entirely lined by a strong and sharp transverse carina starting on the mid-dorsal tooth and terminating at the connection with coxa; these transverse carinae bear two pairs of small teeth: a dorsolateral one and a second one just above the connection with coxa. Pleonites 1 – 2 with one pair of well developed dorsolateral teeth and smaller teeth in various arrangement more ventrally; pleonite 3 with 3 pairs of dorsolateral teeth and smaller teeth in various arrangements more ventrally. Coxae 1 – 3 (and often coxa 4) with sharp tip. Coxae 1 – 3 sharply keeled along their axis. Coxa 4 with sharp carina starting at ventral tip of coxa and terminating at posterior tip of coxa; this carina sometimes bears a tooth or a lobe projecting laterally; carina very distant from posteroventral border of coxa at its deepest point; posteroventral border of coxa concave; anterior corner of coxa strongly bulging anteriorly and sometimes produced into a tooth. Coxa 5 with strong lateral carina which is laterally or posterolaterally produced into a tooth, which is sometimes very large. Coxa 6 with strong lateral carina which is laterally or posterolaterally produced into a tooth, which is much smaller than that of coxa 5. Posteroventral tooth of epimeral plate 3 strong to very strong. Dorsal process of urosomite 1 produced into a sharp tooth directed upwards. Urosomite 2 with pair of small teeth pointing upwards. Lateral borders of urosomite 3 posteriorly terminated into a sharp tooth or an acute angle. Peduncle of antenna 1 with teeth. Mandible with molar process triturative. Lower lip with narrow (V-shaped) hypopharyngeal gap. Palp of maxilliped with 4 articles. Gnathopods very small, achelate; gnathopod 2 distinctly more slender than gnathopod 1. Basis of pereiopods 5 – 6 broad, with strong posteroproximal protrusion (which can be acute but not sword-like and which is directed posteriorly) and posterodistal tooth projecting posteriorly. Posterior border of basis of pereiopod 7 with strong median angle followed by deep excavation, terminated into a tooth projecting posteriorly. Merus, carpus and propodus of pereiopods 3 – 7 very short and very stout; dactylus medium-sized, stout and strongly curved, partly folding on propodus (as if they were designed to clasp on the branches of bushy organisms). Benthic. Body length The maximum body length recorded in Pseudepimeria species ranges between 22 and 35 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF06807FDD5FB35CA61F8F2.taxon	biology_ecology	Ecology Benthic, 50 – 573 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF06807FDD5FB35CA61F8F2.taxon	distribution	Distribution Circum-Antarctic, as far north as the South Orkney Islands.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF06807FDD5FB35CA61F8F2.taxon	discussion	Remarks Of all the Epimeria subgenera, Pseudepimeria is probably the most distinctive. Their highly sculptured and processiform body, and their strong clasping pereiopods 4 – 7 presumably reflect a specialized habitat. Hydroid remains were found in the stomach of E. (Pseudepimeria) oxicarinata (Coleman 1990 a) and pictures of this species clinging on hydroids have been published (Coleman 2007). Coleman (1988) also observed E. grandirostris eating branches of hydroids in an aquarium on the RV Polarstern. Some species are morphologically very similar to each others, but the most similar forms appear to be allopatric. These complexes include the tetrad amoenitas / debroyeri / cf. debroyeri / pulchra and the triad callista / grandirostris / kharieis. Molecular data were not available for all taxa, hence some taxonomic decisions were taken based on morphology alone. Some observed morphological differences are possibly sizedependant, which did not facilitate taxonomic decisions (descriptions are based on adult females only). Immature specimens are assumed to be conspecific with adults of the most similar species found in the same region.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFF06807FDD5FB35CA61F8F2.taxon	description	This key applies to adult females only.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFF6804FDFFFEE3CB6CFCC4.taxon	description	urn: lsid: zoobank. org: act: 191 AE 45 C-F 84 D- 412 A-B 0 E 1 - CFA 3251 C 5824 Figs 251 – 256	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFF6804FDFFFEE3CB6CFCC4.taxon	etymology	Etymology From the Latin, a moenitas, - tatis: pleasantness, delightfulness, loveliness. This name, which is a noun in apposition, has been coined because the species is as elegant as its close relative Epimeria pulchra, of which the specific epithet means beautiful.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFF6804FDFFFEE3CB6CFCC4.taxon	materials_examined	Type material Holotype RV Aurora Australis cruises: SOUTHERN OCEAN: ♀ with hatchlings, cruise CEAMARC, sample CEAMARC 3978, stn 36 EV 297, Adélie Coast, 66 ° 20 ʹ 19.806 ″ S, 143 ° 41 ʹ 07.782 ″ E, 552 – 573 m, beam trawl, 4 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4327). Paratype RV Aurora Australis cruises: SOUTHERN OCEAN: 1 spec., cruise CEAMARC, sample ‘ CEAMARC V 3 stn 158 ’, stn 28 EV 53, Adélie Coast, 65 ° 59 ʹ 46.95 ″ S, 143 ° 02 ʹ 57.0948 ″ E, 461 – 483 m, beam trawl, 23 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4284) [extraction M 14; Genbank nr, COI: KU 870881].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFF6804FDFFFEE3CB6CFCC4.taxon	description	Description ROSTRUM. Long, overreaching tip of peduncle of antenna 1; in lateral view, anteriorly weakly curved in its middle and forming a blunt very obtuse angle with head, ventrally straight, tip not abruptly curving downwards; in frontal view, fairly narrow, with weakly convex borders, with subacute tip. EYE. Medium-sized, conical. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 with medium-sized fairly broad and blunt-tipped middorsal tooth pointing upwards, of which the anterior border is straight and the posterior border is strongly convex, with pair of well-developed, blunt, dorsolateral teeth and pair of small ventrolateral teeth; pereionite 2 much narrower than pereionites 1 and 3, with small narrow blunt-tipped tooth pointing upwards (a bit shorter than mid-dorsal tooth of pereionite 1 and much shorter than mid-dorsal tooth of pereionite 3), with pair of small very blunt dorsolateral teeth, and pair of small ventrolateral teeth; pereionite 3 with large, broad and blunt mid-dorsal tooth pointing upwards, of which the anterior and the posterior borders are both strongly convex, with pair of medium-sized, low and blunt dorsolateral teeth, and pair of small ventrolateral teeth; pereionites 4 – 7 with large and broad mid-dorsal tooth pointing upwards, of which the anterior border is strongly convex and the posterior border weakly convex to nearly straight, and of which the tip is bluntly angulate to subacute, and with pair of medium-sized, low and blunt dorsolateral teeth, and pair of small ventrolateral teeth; pleonites 1 – 2 with large and broad mid-dorsal tooth pointing upwards, of which the anterior border is strongly convex (regularly convex for pleonite 1, with weak median angular discontinuity in pleonite 2) and the posterior border straight, of which the tip is subacute, with pair of large subacute dorsolateral teeth (anteriorly preceded by small but sharp denticle); pleonite 3 with medium-sized mid-dorsal tooth pointing upwards, of medium width, of which the anterior border is proximally convex and distally weakly concave, the tip narrow and acute and the posterior border nearly straight, with 5 pairs of dorsolateral teeth (the most posterior one very large and blunt). COXAE 1 – 3. Strongly carinate and distally sharp. COXA 4. Anterodorsal border straight; anteroventral border with proximal very distinct notch forming an obtuse angle, with distal part straight; anterior angle rounded and projecting forward; ventral tooth long, narrow and acute, directed backwards; lateral carina with rounded lobe strongly projecting laterally, posteriorly followed by fairly deep rounded concavity; in lateral view, inner corner of carina forming a distinct squared angle; in lateral view posteroventral border of coxa deeply concave, the deepest point of this concavity forming a blunt squared angle (more anteriorly and more distally the posterior border is nearly straight). COXA 5. In dorsal view, with very long, triangular, carinate, lateral tooth, of which the anterior border points obliquely backwards and the posterior border is nearly perpendicular to body axis (very weakly pointing backwards); tip of carina sharp. COXA 6. With triangular carinate lateral tooth of which the anterior border points obliquely backwards and the posterior border is nearly perpendicular to body axis (very weakly pointing onwards) (character to be examined in dorsal view); tip of carina sharp. COXA 7. With ventral border distinctly curved, with posterior border very weakly concave, these two borders being joined by a broad curve. EPIMERAL PLATES 1 – 3. Plate 1 with posteroventral angle bluntly angulate, with distinct lateral carina and 2 lateral teeth; plate 2 with posteroventral angle sharply angulate (forming a sharp tooth in squared angle), with distinct lateral carina and 2 lateral teeth; plate 3 with posteroventral angle produced into a long tooth, with distinct lateral carina and 1 lateral tooth, with posterior margin weakly convex. UROSOME TOOTH PATTERN. Urosomite 1 with sharp narrow tooth pointing upwards, not anteriorly preceded by pair of denticles; urosomite 2 with pair of mid-sized posterior dorsolateral teeth pointing upwards; urosomite 3 with pair of mid-sized posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.4; tips of lobes subacute, notch forming a narrow slit. PEDUNCLE OF ANTENNA 1. Article 1 with anterior border concave, with strong anterior and posterior tooth; article 2 with strong lateral triangular tooth; article 3 with strong ventral tooth. GNATHOPODS 1 – 2. Very small, achelate; carpus and propodus of normal slenderness in gnathopod 1 (propodus tapering), extremely narrow in gnathopod 2; dactylus of gnathopod 2 moderately broad and posteriorly straight. PEREIOPODS 3 – 4. Merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPODS 5 – 6. Basis of normal width, with posteroproximal process sharply triangular, with posterodistal tooth strong, narrow and sharp; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPOD 7. Basis broad; posterior border parallel to axis of basis in proximal 0.75; at this level it forms a distinct squared angle, which is followed by a deep concavity; this concavity forms a rounded obtuse (nearly squared) angle; the posterodistal corner is produced into a long narrow tooth pointing obliquely backwards; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. Body length Up to 35 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFF6804FDFFFEE3CB6CFCC4.taxon	distribution	Distribution Adélie Coast, 461 – 573 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFF6804FDFFFEE3CB6CFCC4.taxon	discussion	Remarks Epimeria amoenitas sp. nov. (Adélie Coast) is very similar to E. debroyeri sp. nov. (eastern shelf of the Weddell Sea). In E. amoenitas sp. nov., pleonite 3 has a fairly broad and acute-tipped mid-dorsal tooth, whilst in E. debroyeri sp. nov., the same tooth is very broad and bluntly triangular. In E. amoenitas sp. nov., the posterior border of third epimeral plate is weakly curved, whilst in E. debroyeri sp. nov., it forms a strongly protruding curved projection. Finally, in E. amoenitas sp. nov., the tip of the lateral carina of coxa 5 is sharp in dorsal view, whilst in E. debroyeri sp. nov., it is blunt.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFD6802FDECFC9DCEADF91A.taxon	description	urn: lsid: zoobank. org: act: 8 E 717 EF 2 - 0 CC 3 - 4796 - 859 D- 90 F 60 B 661961 Figs 257 – 264	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFD6802FDECFC9DCEADF91A.taxon	etymology	Etymology From the Greek, καλλιστος, Latinized as callistus, - a, - um, adjective meaning very beautiful, the most beautiful.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFD6802FDECFC9DCEADF91A.taxon	materials_examined	Type material Holotype RV Aurora Australis cruises: SOUTHERN OCEAN: adult ♀, cruise CEAMARC, sample CEAMARC 3978, stn 36 EV 297, Adélie Coast, 66 ° 20 ʹ 20 ″ S, 143 ° 41 ʹ 08 ″ E, 552 – 573 m, beam trawl, 4 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 7336, removed from MNHN-IU- 2014 - 4327) [Extraction P 40; Genbank nr, 28 S: KU 759683]. Paratypes RV Aurora Australis cruises: SOUTHERN OCEAN: 1 dissected adult ♀, cruise CEAMARC, sample CEAMARC 3978, stn 36 EV 297, Adélie Coast, 66 ° 20 ʹ 20 ″ S, 143 ° 41 ʹ 08 ″ E, 552 – 573 m, beam trawl, 4 Jan. 2008, coll. IPEV-AAD-MNHN, (MNHN-IU- 2014 - 7337, removed from MNHN-IU- 2014 - 4327). RV Seatruck cruises: SOUTHERN OCEAN: 1 small and damaged spec., cruise REVOLTA II, stn REVO _ 029, Collect _ ID: REVO _ 183, Adélie Coast, 66 ° 39 ʹ 30 ″ S, 140 ° 01 ʹ 59 ″ E, 97 – 103 m, beam trawl, 26 Jan. 2011, coll. N. Améziane, N. Bax, C. Gallut, A. C. Lautrédou and C. Robineau (MNHN-IU- 2014 - 4300).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFD6802FDECFC9DCEADF91A.taxon	description	Description ROSTRUM. Long, overreaching tip of peduncle of antenna 1; in lateral view, regularly curved along all its length, not forming an angle with head, ventrally straight, tip not abruptly curving downwards; in frontal view, fairly narrow, with weakly convex borders, with acute tip. EYE. Medium-sized, conical. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 with low, very broad, rounded mid-dorsal tooth of which the anterior border is very oblique, with pair of well-developed, rounded, dorsolateral teeth and pair of small ventrolateral teeth; pereionite 2 much narrower than pereionites 1 and 3, with small broad rounded tooth pointing upwards (as high as mid-dorsal tooth of pereionite 1 and distinctly shorter than middorsal tooth of pereionite 3), with pair of very small dorsolateral teeth, and pair of small ventrolateral teeth; pereionite 3 with medium-sized, broad and rounded mid-dorsal tooth of which the anterior and the posterior borders are both strongly convex, with pair of medium-sized, low and blunt dorsolateral teeth, and pair of small ventrolateral teeth; pereionites 4 – 7 with fairly large and broad mid-dorsal tooth pointing upwards, of which the anterior border is strongly convex and the posterior border nearly straight, and of which the tip is bluntly angulate, and with pair of medium-sized, low and blunt dorsolateral teeth, and pair of small ventrolateral teeth; pleonites 1 – 2 with large and broad mid-dorsal tooth pointing upwards, of which the anterior border is strongly and regularly convex and the posterior border straight, and of which the tip is subacute, and with pair of large subacute dorsolateral teeth and 4 pairs of much smaller denticles; pleonite 3 with medium-sized, fairly broad, triangular mid-dorsal tooth pointing upwards, of which the anterior border is proximally convex, medially concave and distally straight, the tip subacute and the posterior border nearly straight, with 5 pairs of dorsolateral teeth. COXAE 1 – 3. Strongly carinate and distally sharp. COXA 4. Anterodorsal border nearly straight; anteroventral border with proximal very shallow concavity, with distal part distinctly convex; anterior angle very broadly rounded and slightly projecting forward; ventral tooth long, broad and subacute, directed backwards; lateral carina with broadly rounded lobe distinctly projecting laterally, posteriorly followed by fairly deep rounded concavity and small extremely low second lobe; in lateral view, inner corner of carina forming a distinct squared angle; in lateral view posteroventral border of coxa deeply concave; the concavity is rounded, not distinctly angulate. COXA 5. In dorsal view, with medium-sized, triangular, carinate, lateral tooth, of which the anterior border points weakly obliquely backwards and the posterior border is nearly perpendicular to body axis; tip of carina broadly rounded. COXA 6. With triangular carinate lateral tooth of which the anterior border points obliquely backwards and the posterior border is nearly perpendicular to body (character to be examined in dorsal view); tip of carina subacute. COXA 7. With ventral + posterior borders forming a regular curve, with small lateral tooth. EPIMERAL PLATES 1 – 3. Plate 1 with posteroventral angle angulate, with distinct lateral carina and 2 lateral teeth; plate 2 with posteroventral angle sharply angulate (forming a distinct blunt-tipped tooth in squared angle), with distinct lateral carina and 2 lateral teeth; plate 3 with posteroventral angle produced into a fairly long tooth, with distinct lateral carina and 1 lateral tooth, with posterior margin weakly convex. UROSOME TOOTH PATTERN. Urosomite 1 with sharp narrow tooth pointing upwards, not anteriorly preceded by pair of denticles; urosomite 2 with pair of tiny posterior dorsolateral teeth pointing upwards; urosomite 3 with pair of mid-sized posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.35; tips of lobes rounded, notch forming a fairly narrow slit. PEDUNCLE OF ANTENNA 1. Article 1 with anterior border weakly concave, with trace of anterior and posterior tooth; article 2 with medium-sized lateral triangular tooth; article 3 with weak ventral tooth. GNATHOPODS 1 – 2. Very small, achelate; carpus and propodus of normal slenderness in gnathopod 1 (propodus tapering), extremely narrow in gnathopod 2; dactylus of gnathopod 2 very broad and posteriorly strongly convex. PEREIOPODS 3 – 4. Merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPOD 5. Basis of normal width, with posteroproximal process rounded and strongly protruding, with posterodistal tooth fairly strong, triangular and sharp; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPOD 6. Basis of normal width, with posteroproximal process acutely triangular, with posterodistal tooth strong, narrow and sharp; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPOD 7. Basis broad; posterior border distinctly diverging from axis of basis in proximal 0.75; at this level it forms a distinct nearly squared (very slightly obtuse) angle, which is followed by a deep concavity; this concavity forms a broadly rounded curve; the posterodistal corner is produced into a long narrow tooth pointing obliquely backwards. Body length Up to 22 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFD6802FDECFC9DCEADF91A.taxon	distribution	Distribution Adélie Coast, 97 – 573 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFD6802FDECFC9DCEADF91A.taxon	discussion	Remarks Epimeria callista sp. nov. (Adélie Coast) is very similar to E. kharieis sp. nov. (eastern shelf of the Weddell Sea) and E. grandirostris (Antarctic Peninsula and surrounding islands). Molecular phylogenetic analysis of 28 S rDNA (Verheye et al. 2016 a) supports the species status of E. callista sp. nov. and E. grandirostris. Epimeria kharieis sp. nov. is herein recognized as a distinct species based on morphology alone, as the type material was not suitable for genetic studies. See key of the subgenus Pseudepimeria for differences between the three species. Allometric differences have been observed in immatures and juveniles of these species, therefore the morphological characters given in the key and descriptions refer to adult females only.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFB68FFFDFAF953CD7CFD81.taxon	description	urn: lsid: zoobank. org: act: 05 C 0 B 30 A-F 5 BD- 40 C 0 - A 3 FC- 07 DE 05 BFB 375 Figs 265 – 271	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFB68FFFDFAF953CD7CFD81.taxon	etymology	Etymology The species is dedicated to Claude De Broyer (Royal Belgian Institute of Natural Sciences), who collected the holotype of the species. The name is a genitive.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFB68FFFDFAF953CD7CFD81.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: large ♀, cruise PS 14, ANT-VII / 4, EPOS leg 3, stn 291 (GSN 14), eastern Weddell Sea, 71 ° 06.1 ʹ S, 12 ° 33.5 ʹ W to 71 ° 05.9 ʹ S, 12 ° 34.8 ʹ W, 499 – 515 m, bottom trawl, 19 Feb. 1989, coll. C. De Broyer (RBINS, INV. 132940).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFB68FFFDFAF953CD7CFD81.taxon	description	Description ROSTRUM. Long, overreaching tip of peduncle of antenna 1; in lateral view, anteriorly weakly curved in its middle and forming a blunt very obtuse angle with head, ventrally straight, tip abruptly curving downwards; in frontal view, fairly narrow, with weakly convex borders, with subacute tip. EYE. Medium-sized, conical. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 with medium-sized broad and blunt-tipped mid-dorsal tooth pointing upwards of which the anterior border is straight and the posterior border is strongly convex, with pair of well-developed, blunt, dorsolateral teeth and pair of small ventrolateral teeth; pereionite 2 much narrower than pereionites 1 and 3, with small narrow blunt-tipped tooth pointing upwards (a bit shorter than mid-dorsal tooth of pereionite 1 and much shorter than mid-dorsal tooth of pereionite 3), with pair of small very blunt dorsolateral teeth, and pair of small ventrolateral teeth; pereionites 3 – 4 with large, broad and blunt to subangulate mid-dorsal tooth pointing upwards, of which the anterior and the posterior borders are both strongly convex, with pair of medium-sized, low and blunt dorsolateral teeth, and pair of small ventrolateral teeth; pereionites 5 – 7 with large and broad mid-dorsal tooth pointing upwards, of which the anterior border is strongly convex and the posterior border weakly convex, of which the tip is bluntly angulate to subacute, with pair of medium-sized, low and blunt dorsolateral teeth, and pair of small ventrolateral teeth; pleonites 1 – 2 with large and broad mid-dorsal tooth pointing upwards, of which the anterior border is strongly convex (with weak median angular discontinuity) and the posterior border nearly straight, of which the tip is subacute, with pair of large subacute dorsolateral teeth (anteriorly preceded by small but sharp denticle); pleonite 3 with very broad medium-sized middorsal tooth pointing upwards, of which the anterior border is convex, the tip broad and subangulate and the posterior border weakly convex, with 5 pairs of dorsolateral teeth. COXAE 1 – 3. Strongly carinate and distally blunt. COXA 4. Anterodorsal border weakly but distinctly concave; anteroventral border with proximal very distinct notch forming an obtuse angle, with distal part convex; anterior angle rounded to very bluntly angulate and projecting forward; ventral tooth long, narrow and acute, directed backwards; lateral carina with rounded to very bluntly angulate lobe strongly projecting laterally, posteriorly followed by fairly shallow rounded concavity, and more backwards by a second, very low lobe projecting laterally; in lateral view, inner corner of carina forming a distinct obtuse (nearly squared) angle; in lateral view posteroventral border of coxa deeply concave, the deepest point of this concavity forming a blunt squared angle (more anteriorly and more distally the posterior border is nearly straight). COXA 5. In dorsal view, with very long, triangular, carinate, lateral tooth, of which the anterior border points obliquely strongly backwards and the posterior border is also distinctly pointing backwards); tip of carina blunt. COXA 6. With triangular carinate lateral tooth of which the anterior border points obliquely backwards and the posterior border is nearly perpendicular to body axis (very weakly pointing onwards) (character to be examined in dorsal view); tip of carina subacute. COXA 7. With ventral border distinctly curved, with posterior border straight, these two borders being joined by a broad curve. EPIMERAL PLATES 1 – 3. Plate 1 with posteroventral angle broadly rounded, with distinct lateral carina and 1 lateral tooth; plate 2 with posteroventral angle sharply angulate (forming a sharp tooth in squared angle), with distinct lateral carina and 1 lateral tooth; plate 3 with posteroventral angle produced into a long tooth, with distinct lateral carina and 1 lateral tooth, with posterior margin very strongly convex. UROSOME TOOTH PATTERN. Urosomite 1 with sharp narrow tooth pointing upwards, anteriorly preceded by pair of denticles; urosomite 2 with pair of mid-sized posterior dorsolateral teeth pointing upwards; urosomite 3 with pair of mid-sized posterior dorsolateral teeth pointing obliquely backwards. TELSON. Cleft on 0.2; tips of lobes subacute, notch forming a broad V. PEDUNCLE OF ANTENNA 1. Article 1 with anterior border concave, with strong anterior and posterior tooth; article 2 with strong lateral triangular tooth; article 3 with strong ventral tooth. GNATHOPODS 1 – 2. Very small, achelate; carpus and propodus of normal slenderness in gnathopod 1 (propodus tapering), extremely narrow in gnathopod 2; dactylus of gnathopod 2 fairly narrow and posteriorly straight. PEREIOPODS 3 – 4. Merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPODS 5 – 6. Basis of normal width, with posteroproximal process sharply triangular, with posterodistal tooth strong, narrow and sharp; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPOD 7. Basis broad; posterior border parallel to axis of basis in proximal 0.75; at this level it forms a distinct acute (nearly squared) angle, which is followed by a deep concavity; this concavity forms a rounded obtuse (nearly squared) angle; the posterodistal corner is produced into a long narrow tooth pointing obliquely backwards; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. Colour pattern Body and appendages entirely white; eye reddish (Coleman 2007: 47, colour plate 2 f; Rauschert & Arntz 2015: 62, pl. 55, both as Epimeria pulchra). Body length 33 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFB68FFFDFAF953CD7CFD81.taxon	distribution	Distribution Eastern shelf of the Weddell Sea, 499 – 515 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFFFB68FFFDFAF953CD7CFD81.taxon	discussion	Remarks Epimeria debroyeri sp. nov. (eastern shelf of the Weddell Sea) is very similar to E. amoenitas sp. nov. (Adélie Coast). See key and account on E. amoenitas sp. nov. The specimen of E. debroyeri sp. nov. illustrated by a photograph as E. pulchra in Coleman (2007) and reproduced in Rauschert & Arntz (2015) was collected on the eastern shelf of the Weddell Sea. Indeed, in an early draft of Rauschert & Arntz’s (2015) book, it was indicated that this specimen was collected during the cruise ANT-XIII / 3 dedicated to the study of that region.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0668FFFDDCFDD8CD67FA8C.taxon	description	Fig 272	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0668FFFDDCFDD8CD67FA8C.taxon	materials_examined	Material examined RV Polarstern cruises: SOUTHERN OCEAN: 1 juv., cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 132941); 1 white juv., photographed, cruise PS 69, ANT-XXIII / 8, stn 728 - 2, south of Dundee Island, 63 ° 42.63 ʹ S, 56 ° 01.63 ʹ W to 63 ° 42.25 ʹ S, 56 ° 02.16 ʹ W, 293 – 298 m, Agassiz trawl, 24 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122524).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0668FFFDDCFDD8CD67FA8C.taxon	description	Colour pattern Body and appendages entirely white; eyes reddish.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0668FFFDDCFDD8CD67FA8C.taxon	discussion	Remarks These two specimens possibly belong to Epimeria debroyeri sp. nov. However, they are much smaller (8 mm instead of 33 mm) and exhibit a number of differences, especially in the development of their mid-dorsal crests. It has to be noted that these differences might be size-related. The two specimens are therefore referred as E. cf. debroyeri.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0668FCFE4AFAC5CB9AFC8B.taxon	description	Figs 273 – 282	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0668FCFE4AFAC5CB9AFC8B.taxon	materials_examined	Material examined RV Polarstern cruises: SOUTHERN OCEAN: 1 very small juv., PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122523); 1 small spec., PS 69, ANT-XXIII / 8, stn 605 - 1, Elephant Island, 61 ° 20.35 ʹ S, 55 ° 29.16 ʹ W to 61 ° 19.98 ʹ S, 55 ° 32.67 ʹ W, 146 – 151 m, bottom trawl, 19 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122530); 1 small spec., PS 69, ANT-XXIII / 8, stn 608 - 1, Elephant Island, 61 ° 11.34 ʹ S, 54 ° 43.17 ʹ W to 61 ° 11.80 ʹ S, 54 ° 40.05 ʹ W, 284 – 293 m, bottom trawl, 20 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122626); 1 small spec., PS 69, ANT-XXIII / 8, stn 654 - 6, Elephant Island, 61 ° 22.80 ʹ S, 56 ° 03.84 ʹ W to 61 ° 23.35 ʹ S, 56 ° 04.89 ʹ W, 341 – 342 m, Agassiz trawl, 29 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122484); 1 adult ♀, PS 81, ANT-XXIX / 3, stn 185 - 3, south east of Dundee Island, 63 ° 51.34 ʹ S, 55 ° 41.11 ʹ W to 63 ° 51.52 ʹ S, 55 ° 41.43 ʹ W, bottom not muddy with a lot of life (sponges, starfishes, ophiuroids, crinoids, Pentapora - like bryozoans), 261 – 296 m, Agassiz trawl, 19 Feb. 2013:, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122946) [extraction ANT 46; Genbank nr, COI: KU 870832, 28 S: KU 759605]; 1 badly damaged small spec., PS 81, ANT-XXIX / 3, stn 197 - 6, Bransfield Strait, 62 ° 45.05 ʹ S, 57 ° 26.68 ʹ W to 62 ° 45.09 ʹ S, 57 ° 26.47 ʹ W, 210 – 222 m, black gravel mixed with sand and a little bit of mud, Rauschert dredge, 25 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122950) [extraction ANT 47; Genbank nr, COI: KU 870833, 28 S: KU 759606]	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0668FCFE4AFAC5CB9AFC8B.taxon	description	Description Description based on adult female of ANT-XXIX / 3, stn 185 - 3 (RBINS, INV. 122946). ROSTRUM. Long, overreaching tip of peduncle of antenna 1; in lateral view, proximally nearly straight, distinctly curved on second 0.3, not forming a distinct angle with head, ventrally straight, tip not abruptly curving downwards; in frontal view, fairly broad, with weakly convex borders, with acute tip. EYE. Medium-sized, conical. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 with low, very broad, rounded mid-dorsal tooth, of which the anterior border is very oblique, with pair of well-developed, rounded, dorsolateral teeth and pair of small ventrolateral teeth; pereionite 2 much narrower than pereionites 1 and 3, with small broad rounded tooth pointing upwards (as high as mid-dorsal tooth of pereionite 1 and distinctly shorter than mid-dorsal tooth of pereionite 3), with pair of very small dorsolateral teeth, and pair of small ventrolateral teeth; pereionites 3 – 7 with fairly large, broad and rounded mid-dorsal tooth of which the anterior and posterior borders are both strongly convex, with pair of medium-sized, low and blunt dorsolateral teeth, and pair of small ventrolateral teeth; pleonites 1 – 2 with large and broad mid-dorsal tooth pointing upwards, of which the anterior border is strongly and regularly convex (less convex in pleonite 2) and the posterior border straight, and of which the tip is subacute, and with pair of large subacute dorsolateral teeth and 4 pairs of much smaller denticles; pleonite 3 with medium-sized, fairly broad, triangular mid-dorsal tooth pointing upwards, of which the anterior border is straight (except for proximal tiny protrusion), the tip acute and the posterior border slightly but distinctly convex, with 5 pairs of dorsolateral teeth. COXAE 1 – 3. Strongly carinate and distally sharp. COXA 4. Anterodorsal border nearly straight; anteroventral border with inconspicuous concavity, with distal part distinctly convex; anterior angle extremely broadly rounded and not projecting forward; ventral tooth well developed, very broad and subacute directed backwards; lateral carina with low, very broadly rounded lobe distinctly projecting laterally, posteriorly not followed by concavity and not followed by second smaller lobe; in lateral view, the carina is forming a regular, strongly arching curve and is not at all angulate; in lateral view posteroventral border of coxa deeply concave; the concavity is rounded, not distinctly angulate. COXA 5. In dorsal view, with medium-sized, triangular, carinate, lateral tooth, of which the anterior border points distinctly obliquely backwards and the posterior border is nearly perpendicular to body axis; tip of carina broadly rounded. COXA 6. Triangular carinate lateral tooth of which the anterior border points obliquely backwards and the posterior border is nearly perpendicular to body (character to be examined in dorsal view); tip of carina very blunt. COXA 7. With ventral + posterior borders forming a regular curve, with small lateral tooth. EPIMERAL PLATES 1 – 3. Plate 1 with posteroventral angle sharply angulate and with small triangular tooth, with distinct lateral carina and 2 lateral teeth; plate 2 with posteroventral angle sharply angulate and produced into a small triangular tooth, with distinct lateral carina and 2 lateral teeth; plate 3 with posteroventral angle produced into a fairly long tooth, with distinct lateral carina and 1 lateral tooth, with posterior margin distinctly convex. UROSOME TOOTH PATTERN. Urosomite 1 with sharp narrow tooth pointing upwards, not anteriorly preceded by pair of denticles; urosomite 2 with pair of tiny posterior dorsolateral teeth pointing upwards; urosomite 3 with pair of mid-sized posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.25; lobes laterally very convex, blunt-tipped, notch fairly narrow. PEDUNCLE OF ANTENNA 1. Article 1 with anterior border weakly concave, with low anterior and posterior tooth; article 2 with medium-sized lateral triangular tooth; article 3 with weak ventral tooth. GNATHOPODS 1 – 2. Very small, achelate; carpus and propodus of normal slenderness in gnathopod 1 (propodus tapering), extremely narrow in gnathopod 2; dactylus of gnathopod 2 very broad and posteriorly weakly convex. PEREIOPODS 3 – 4. Merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPOD 5. Basis of normal width, with posteroproximal process triangular and sharp, with posterodistal tooth fairly strong, triangular and sharp; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPOD 6. Basis 6 of normal width, with posteroproximal process triangular and subacute, with posterodistal tooth strong, narrow and sharp; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPOD 7. Basis broad; posterior border distinctly diverging from axis of basis in proximal 0.75; at this level it forms a very blunt obtuse angle, which is followed by a distinct concavity; this concavity forms a broadly rounded curve; the posterodistal corner is produced into a long triangular tooth pointing obliquely backwards; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. Allometric variations See photographs of immatures (Figs 279 – 282). Colour pattern Orange red mottling on a whitish background. Body length Up to 25 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0668FCFE4AFAC5CB9AFC8B.taxon	distribution	Distribution Marguerite Bay, 254 m [type locality] (Chevreux 1912, 1913); Elephant Island, Bransfield Strait, northwestern Weddell Sea (Dundee Island); 146 – 342 m (present material). For an inventory of older records of Epimeria of the grandirostris complex, see De Broyer et al. (2007), as Epimeria grandirostris.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0668FCFE4AFAC5CB9AFC8B.taxon	biology_ecology	Biology Coleman (1988) observed E. grandirostris eating branches of hydroids in an aquarium on the RV Polarstern.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0668FCFE4AFAC5CB9AFC8B.taxon	discussion	Remarks The populations of the E. grandirostris complex sampled in the eastern Weddell Sea and Adélie Coast are described herein as distinct species, respectively E. kharieis sp. nov. and E. callista sp. nov. See key of the subgenus Pseudepimeria for the main diagnostic characters of adult females.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0568F9FDE2FCC5CC96FE3D.taxon	description	urn: lsid: zoobank. org: act: 1 B 3 DC 61 B-C 54 C- 4 D 63 - AAC 6 - 29 A 5 BD 8807 A 7 Figs 283 – 290	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0568F9FDE2FCC5CC96FE3D.taxon	etymology	Etymology Derived from the Greek adjective χαρίεις, meaning graceful and beautiful, and Latinized herein as kharieis, - is, - e.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0568F9FDE2FCC5CC96FE3D.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: adult ♀, cruise PS 48, ANT-XV / 3, EASIZ II, stn 77 (AGT 7), eastern Weddell Sea, 71 ° 08.6 ʹ S, 12 ° 26.6 ʹ W to 71 ° 10.2 ʹ S, 12 ° 30.7 ʹ W, 330 – 433 m, Agassiz trawl, 2 Feb. 1998, coll. C. De Broyer and Y. Scailteur (RBINS, INV. 132937). Paratypes RV Polarstern cruises: SOUTHERN OCEAN: 1 fairly large spec., cruise PS 14, ANT-VII / 4, EPOS leg 3, stn 281, eastern Weddell Sea, 71 ° 39.0 ʹ S, 12 ° 21.1 ʹ W to 71 ° 36.9 ʹ S, 12 ° 25.0 ʹ W, 389 – 450 m, Agassiz trawl, 17 Jan. 1989, coll. C. De Broyer (RBINS, INV. 132939); 1 spec., cruise PS 48, ANT-XV / 3, EASIZ II, stn 77 (AGT 7), eastern Weddell Sea, 71 ° 08.6 ʹ S, 12 ° 26.6 ʹ W to 71 ° 10.2 ʹ S, 12 ° 30.7 ʹ W, 330 – 433 m, Agassiz trawl, 2 Feb. 1998, coll. C. De Broyer and Y. Scailteur (RBINS, INV. 132938); 1 immature spec., cruise PS 65, ANT-XXI / 2, BENDEX, stn 144 - 1, eastern Weddell Sea, 70 ° 57.02 ʹ S, 10 ° 48.43 ʹ W to 70 ° 56.98 ʹ S, 10 ° 48.04 ʹ W, 401 – 407 m, 13 Dec. 2003, coll. C. De Broyer (RBINS, INV. 132988).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0568F9FDE2FCC5CC96FE3D.taxon	description	Description Description based on adult females. ROSTRUM. Long, overreaching tip of peduncle of antenna 1; in lateral view, straight on proximal half, regularly curved on second half, forming a very obtuse angle with head, ventrally straight, tip not abruptly curving downwards; in frontal view, fairly broad, with weakly convex borders, with acute tip. EYE. Medium-sized, conical. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 with low, broad, rounded mid-dorsal tooth pointing upwards of which the anterior border is distinctly oblique, with pair of well-developed, rounded, dorsolateral teeth and pair of small ventrolateral teeth; pereionite 2 much narrower than pereionites 1 and 3, with small elliptic tooth pointing upwards (as high as mid-dorsal tooth of pereionite 1 and distinctly shorter than mid-dorsal tooth of pereionite 3), with pair of very small dorsolateral teeth, and pair of small ventrolateral teeth; pereionites 3 – 7 with medium-sized, very broad and rounded mid-dorsal tooth of which the anterior and the posterior borders are both strongly convex, with pair of mediumsized, low and blunt dorsolateral teeth, and pair of small ventrolateral teeth; pleonites 1 – 2 with large and broad mid-dorsal tooth pointing upwards, of which the anterior border is strongly and regularly convex and the posterior border slightly less convex, and of which the tip is bluntly angulate, and with pair of large subacute dorsolateral teeth and 4 pairs of much smaller denticles; pleonite 3 with medium-sized, broad, triangular mid-dorsal tooth pointing upwards, of which the anterior border may have (holotype) a low obtuse angulate protrusion on proximal 0.6, of which the tip is subacute and the posterior border weakly sinuate, with 5 pairs of dorsolateral teeth. COXAE 1 – 3. Strongly carinate and distally sharp. COXA 4. Anterodorsal border nearly straight; anteroventral border with proximal very shallow concavity, with distal part distinctly convex; anterior angle very broadly rounded and slightly projecting forward; ventral tooth long, broad and subacute, directed backwards; lateral carina with very low rounded lobe slightly projecting laterally, in the holotype posteriorly followed by very shallow rounded concavity and small extremely low second lobe; in lateral view, inner corner of carina forming a distinct squared angle; in lateral view posteroventral border of coxa deeply concave; the concavity is rounded, not distinctly angulate. COXA 5. In dorsal view, with low, triangular, carinate, lateral tooth, of which the anterior border is nearly parallel to body axis and the posterior border is obliquely pointing backwards; tip of carina blunt. COXA 6. With triangular carinate lateral tooth, of which the anterior border (which is nearly straight, inconspicuously convex) points obliquely backwards; the posterior border is nearly perpendicular to body axis (character to be examined in dorsal view); tip of carina blunt. COXA 7. With ventral + posterior borders forming a regular curve, with small lateral tooth. EPIMERAL PLATES 1 – 3. Plate 1 with posteroventral angle angulate, with distinct lateral carina and 2 lateral teeth; plate 2 with posteroventral angle sharply angulate (forming a distinct sharp tooth), with distinct lateral carina and 2 lateral teeth; plate 3 with posteroventral angle produced into a fairly long tooth, with distinct lateral carina and 1 lateral tooth, with posterior margin weakly convex. UROSOME TOOTH PATTERN. Urosomite 1 with sharp narrow tooth pointing upwards, more anteriorly with or without pair of low dorsolateral denticles; urosomite 2 with 2 pairs of posterior dorsolateral teeth pointing upwards; urosomite 3 with pair of mid-sized posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.30; tips of lobes blunt, notch forming a fairly narrow slit. PEDUNCLE OF ANTENNA 1. Article 1 with anterior border weakly concave, with well-developed anterior and posterior tooth; article 2 with medium-sized lateral triangular tooth; article 3 with weak ventral tooth. GNATHOPODS 1 – 2. Very small; gnathopod 1 incomplete in dissected specimen; gnathopod 2 achelate; carpus and propodus extremely narrow; dactylus of gnathopod 2 fairly broad and posteriorly weakly convex. PEREIOPODS 3 – 4. Merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPOD 5. Basis of normal width, with posteroproximal process rounded and strongly protruding, with posterodistal tooth small, triangular; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPOD 6. Basis of normal width, with posteroproximal process very protruding (with proximal border very convex, with tip angulate or bluntly angulate, with distal border concave), with posterodistal tooth fairly small, triangular; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPOD 7. Basis broad; posterior border strongly diverging from axis of basis in proximal 0.75; at this level it form a distinct, blunt, nearly squared (very slightly obtuse) angle, which is followed by a deep concavity; this concavity form a broadly rounded curve; the posterodistal corner is produced into a long narrow tooth pointing obliquely backwards; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. Allometric variations See pictures of immature specimen. Colour pattern Red mottled on a white background (photographs of Rauschert & Arntz 2015). Body length Up to 26 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0568F9FDE2FCC5CC96FE3D.taxon	distribution	Distribution Eastern shelf of the Weddell Sea, 330 – 450 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0568F9FDE2FCC5CC96FE3D.taxon	discussion	Remarks Based on morphological characters, Epimeria of the grandirostris complex from the eastern shelf of the Weddell Sea are herein considered as the new species E. kharieis sp. nov. See key of the subgenus Pseudepimeria for characters. The ‘ Epimeria grandirostris ’ specimens illustrated by Rauschert & Arntz (2015) are presumably E. kharieis sp. nov. based on the collection location: eastern shelf of the Weddell Sea. However, as many morphological characters cannot be observed on their photographs, this identification is tentative. The precise localities are given in an early draft of their book. They are indicated as respectively coming from stations F 42 and 121 of the cruise ANT-XXI (obviously ANT-XXI / 2). Station 42 corresponds to a photo sledge. The letter “ F ” would suggest that it is not an error, as the official acronym of the photo sledge used during ANT-XXI / 2 is FTS (Arntz & Brey 2005: 120). The coordinates of station 42 are 70 ° 29.66 ʹ S, 08 ° 58.73 ʹ W to 70 ° 29.65 ʹ S, 08 ° 58.55 ʹ W, 428 – 429 m. Those of station 121 are 70 ° 50.08 ʹ S, 10 ° 35.54 ʹ W to 70 ° 50.08 ʹ S, 10 ° 34.76 ʹ W, 268 – 274 m. The ‘ Epimeria grandirostris ’ illustrated on plate 2 fig. b (colour photograph) by Coleman (2007) is also presumably E. kharieis sp. nov. A copy of the photograph used by Coleman (2007) was found in the archives of RBINS, indicating that it was made during the cruise ANT-XIII (obviously ANT-XIII / 3), which was focused on the study of the eastern shelf of the Weddell Sea.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0068F7FE23FE33CB25F862.taxon	description	Figs 291 – 292	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0068F7FE23FE33CB25F862.taxon	materials_examined	Material examined RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., cruise PS 14, ANT-VII / 4, EPOS leg 3, stn 211, Elephant Island, 60 ° 59.8 ʹ S, 55 ° 12.1 ʹ W to 60 ° 59.3 ʹ S, to 55 ° 10.5 ʹ W, 207 – 213 m, bottom trawl, 15 Jan. 1989, coll. C. De Broyer (RBINS, INV. 132945); 1 spec., cruise PS 48, ANT-XV / 3, EASIZ II, stn 353, north of Nelson Island, 61 ° 59.4 ʹ S, 59 ° 14.4 ʹ W to 61 ° 58.2 ʹ S, 59 ° 14.3 ʹ W, 129 – 132 m, Agassiz trawl, 20 Mar. 1998, coll. C. De Broyer (RBINS, INV. 132723); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 605 - 3, Elephant Island, 61 ° 20.33 ʹ S, 55 ° 31.53 ʹ W to 61 ° 20.35 ʹ S, 55 ° 30.18 ʹ W, 148 – 154 m, Agassiz trawl, 20 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122483) [extraction N 8; Genbank nr, COI: KU 870892]; 1 ♀, cruise PS 69, ANT-XXIII / 8, stn 605 - 5, Elephant Island, 61 ° 20.27 ʹ S, 55 ° 30.92 ʹ W to 61 ° 20.37 ʹ S, 55 ° 28.99 ʹ W, 131 – 152 m, 20 Dec. 2006, Agassiz trawl, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122482); 8 specs, cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122517); 1 spec., PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (MNHN-IU- 2014 - 7338, removed from RBINS, INV. 122517); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 132942); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 132972); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122468) [extraction N 7; Genbank nr, COI: KU 870891]; 1 spec., cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122472); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 614 - 3, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122521); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 639 - 1, Elephant Island, 61 ° 10.27 ʹ S, 55 ° 56.52 ʹ W to 61 ° 11.12 ʹ S, 55 ° 53.03 ʹ W, 127 – 128 m, bottom trawl, 26 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122518); 2 specs, cruise PS 69, ANT-XXIII / 8, stn 644 - 1, Elephant Island, 61 ° 03.19 ʹ S, 55 ° 54.36 ʹ W to 61 ° 01.78 ʹ S, 55 ° 51.83 ʹ W, 150 – 187 m, bottom trawl, 27 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122522).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0068F7FE23FE33CB25F862.taxon	description	Description ROSTRUM. Long, overreaching tip of peduncle of antenna 1; in lateral view, regularly curved, forming a distinct obtuse angle with head, ventrally very weakly concave, tip scarcely curving downwards; in frontal view, fairly broad, with weakly convex borders, with acute tip. EYE. Medium-sized, conical. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 with very long, narrow, sharp mid-dorsal tooth arching forward (anterior border strongly concave, posterior border strongly convex), with pair of long, sharp, dorsolateral teeth, and pair of small ventrolateral teeth; pereionite 2 much narrower than pereionites 1 and 3, with small, narrow and sharp tooth pointing upwards (5 × shorter than mid-dorsal tooth of pereionites 1 and 3), with pair of small dorsolateral teeth, and pair of small ventrolateral teeth; pereionites 3 – 7 with very long, very narrow, very sharp mid-dorsal tooth of which the anterior and the posterior borders are nearly straight, with pair of large and sharp dorsolateral teeth, and pair of small ventrolateral teeth; pleonites 1 – 2 with very long, very narrow, very sharp mid-dorsal tooth of which the anterior and the posterior borders are nearly straight, with pair of large acute dorsolateral teeth and very small denticles; pleonite 3 very long, very narrow, very sharp mid-dorsal tooth of which the anterior and the posterior borders are nearly straight, with about 5 pairs of dorsolateral teeth, of which 2 are large and sharp and the others small to very small. COXAE 1 – 3. Strongly carinate and distally sharp. COXA 4. Anterodorsal border weakly but distinctly concave; anteroventral border with deep notch, with distal part nearly straight; anterior angle produced into a large and sharp tooth pointing downwards; ventral tooth long, narrow and sharp, directed ventrally (scarcely arching backwards); lateral carina with long and sharp tooth projecting laterally; in lateral view, inner corner of carina forming a distinct obtuse angle; in lateral view posteroventral border of coxa deeply concave; the concavity is rounded. COXA 5. In dorsal view, with very large, narrowly triangular, carinate, lateral tooth, of which the anterior and the posterior borders point obliquely backwards; tip of carina sharp. COXA 6. With narrowly triangular carinate lateral tooth of which the anterior border (which is convex) and the posterior border (which is concave) points obliquely backwards (character to be examined in dorsal view); tip of carina very sharp. COXA 7. With ventral + posterior borders forming a regular curve, with small lateral tooth. EPIMERAL PLATES 1 – 3. Plate 1 with posteroventral angle angulate, with distinct lateral carina and 2 lateral teeth; plate 2 with posteroventral angle sharply angulate (forming a distinct sharp triangular tooth), with distinct lateral carina and 2 lateral teeth; plate 3 with posteroventral angle produced into a very long tooth, with distinct lateral carina and 1 lateral tooth, with posterior margin weakly convex. UROSOME TOOTH PATTERN. Urosomite 1 with sharp narrow mid-dorsal tooth pointing upwards, with pair of lateral denticles at the same level as mid-dorsal tooth; urosomite 2 with 2 pairs of small posterior dorsolateral teeth pointing upwards (a very small triangular one followed by a longer styliform one); urosomite 3 with pair of mid-sized posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.4; tips of lobes sharp, notch V-shaped. PEDUNCLE OF ANTENNA 1. Article 1 with anterior border weakly concave, with well-developed anterior and posterior tooth; article 2 with medium-sized lateral triangular tooth; article 3 with weak ventral tooth. GNATHOPODS 1 – 2. Very small, achelate; carpus and propodus of medium stoutness in gnathopod 1 (propodus tapering), of fairly narrow in gnathopod 2; dactylus of gnathopod 2 fairly broad and posteriorly slightly convex. PEREIOPODS 3 – 4. Merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPODS 5 – 6. Basis of normal width, with posteroproximal process sharply triangular and strongly protruding, with posterodistal tooth large and sharp; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPOD 7. Basis of medium-width; posterior border distinctly diverging from axis of basis in proximal 0.75; at this level it forms a triangular tooth (i. e., blunt-tipped acute, nearly straight angle), which is followed by a deep concavity; this concavity form a broadly rounded curve; the posterodistal corner is produced into a long narrow tooth pointing obliquely backwards; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. Colour pattern Transversally striped of white, red and orange. Body length Up to 26 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0068F7FE23FE33CB25F862.taxon	distribution	Distribution Elephant Island, 127 – 267 m (Coleman 2007; present data), Drake Passage: north of Nelson Island (present data).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0068F7FE23FE33CB25F862.taxon	biology_ecology	Biology Coleman (1990 a) found thecate hydrozoans (cf. Oswaldella antarctica (Jäderholm, 1904 )) in the stomachs of several E. oxicarinata specimens. A photograph of the species clinging onto a thecate hydrozoan with an long strong stem and thin lateral branches was published by Coleman (2007). So it is likely that E. oxicarinata is a hydroid-associated and hydroid-feeding species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0068F7FE23FE33CB25F862.taxon	discussion	Remarks There is an unconfirmed record of E. oxicarinata from the eastern shelf of the Weddell Sea (Klages 1991; De Broyer et al. 2007), but this might be a misidentification, possibly a confusion with E. cyrano sp. nov., which is also a very spiny, albeit very different in other morphological characters. The collection station of the specimen of Epimeria oxicarinata illustrated by Rauchert & Arntz (2015) was mentioned in an early draft of their book accessed by the authors: ANT-XV / 3 stn 355. This corresponds to 61 ° 59.8 ʹ S, 59 ° 14.8 ʹ W to 62 ° 00.1 ʹ S, 59 ° 14.8 ʹ W (north of Nelson Island), 128 – 130 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0F68F4FE1DFEE3CEA5FE1D.taxon	description	Fig. 293	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0F68F4FE1DFEE3CEA5FE1D.taxon	materials_examined	Material examined SOUTHERN OCEAN: 2 adult specs, SIGNY 1991 / 92, AGT 5, transect 1, South Orkney Plateau: Signy Island, 60.683 ° S, 45.581 ° W, 50 m, 10 Jan. 1992, coll. Stefan Hain (RBINS, INV. 132687).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0F68F4FE1DFEE3CEA5FE1D.taxon	description	Description ROSTRUM. Long, overreaching tip of peduncle of antenna 1; in lateral view, anteriorly weakly curved in its middle and forming a blunt very obtuse angle with head, ventrally weakly curved, tip sometimes abruptly curving downwards; in frontal view, fairly narrow, with weakly convex borders, with subacute tip. EYE. Medium-sized, conical. PEREION – PLEOSOME TOOTH PATTERN. Pereionite 1 with medium-sized broad and blunt-tipped mid-dorsal tooth pointing upwards of which the anterior border is straight and the posterior border is strongly convex, with pair of well-developed, blunt, dorsolateral teeth and pair of small ventrolateral teeth; pereionite 2 much narrower than pereionites 1 and 3, with small narrow blunt-tipped tooth pointing upwards (a bit shorter than mid-dorsal tooth of pereionite 1 and much shorter than mid-dorsal tooth of pereionite 3), with pair of small very blunt dorsolateral teeth, and pair of small ventrolateral teeth; pereionites 3 – 7 with very large, slender and blunt-tipped mid-dorsal tooth pointing upwards, of which the anterior and the posterior borders are similarly convex, with pair of small, low and blunt dorsolateral teeth, and pair of small ventrolateral teeth; pleonite 1 with very large and slender mid-dorsal tooth pointing upwards, of which the anterior and posterior borders are convex (anterior border with distinct subdistal angular discontinuity), and of which the tip is blunt to angulate, and with pair of large subacute dorsolateral teeth (associated with several pairs of smaller denticles); pleonite 2 with very large and slender mid-dorsal tooth pointing upwards, of which the anterior and posterior borders are convex, and of which the tip is angulate, and with pair of large subacute dorsolateral teeth (associated with several pairs of smaller denticles); pleonite 3 with very narrow and very large mid-dorsal tooth pointing upwards, of which the anterior border is not convex, and the tip sharp and narrow, with 5 pairs of dorsolateral teeth. COXAE 1 – 3. Strongly carinate and distally subacute. COXA 4. Anterodorsal border straight; anteroventral border with median very deep angulate concavity (anterior border of concavity concave; posterior border of concavity straight); anterior angle produced into a large broad tooth projecting forward; ventral tooth fairly long, narrow and subacute, pointing weakly obliquely backwards; lateral carina with bluntly angulate lobe strongly projecting laterally, posteriorly followed by fairly deep rounded concavity, and more backwards by a second, lower but distinct lobe projecting laterally; in lateral view, inner corner of carina forming a distinct squared angle; in lateral view posteroventral border of coxa deeply concave, the deepest point of this concavity forming a very rounded, obtuse (nearly squared) angle. COXA 5. In dorsal view, with extremely long, triangular, carinate, lateral tooth, of which the anterior border points obliquely strongly backwards and the posterior border is almost perpendicular to body axis; tip of carina sharp. COXA 6. With triangular carinate lateral tooth of which the anterior border points obliquely backwards and the posterior border is nearly perpendicular to body axis (very weakly pointing onwards); tip of carina subacute; posteriorly with a second, much smaller obtuse triangular tooth directed backwards. COXA 7. With ventral border strongly curved, with posterior border straight, these two borders being joined by a long broad curve. EPIMERAL PLATES 1 – 3. Plate 1 with posteroventral angle angulate and with posterior border distinctly angulate, with distinct lateral carina and 2 lateral teeth; plate 2 with posteroventral angle produced into a medium-sized sharp tooth, with distinct lateral carina and 2 lateral teeth; plate 3 with posteroventral angle produced into a long tooth, with distinct lateral carina and 1 lateral tooth, with posterior margin produced into a large triangular tooth. UROSOME TOOTH PATTERN. Urosomite 1 with sharp narrow tooth pointing upwards, with pair of denticle just in front of the main tooth; urosomite 2 with pair of mid-sized posterior dorsolateral teeth pointing upwards; urosomite 3 with pair of mid-sized posterior dorsolateral teeth pointing upwards. TELSON. Cleft on 0.3; tips of lobes acute, notch forming a narrow V. PEDUNCLE OF ANTENNA 1. Article 1 with anterior border concave, with strong anterior and posterior tooth; article 2 with strong lateral triangular tooth; article 3 with strong ventral tooth. GNATHOPODS 1 – 2. Very small, achelate; carpus and propodus broad in gnathopod 1 (propodus tapering), of medium width in gnathopod 2; dactylus of gnathopod 2 extremely broad and posteriorly straight. PEREIOPOD 3 – 4. Merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPODS 5 – 6. Basis of pereiopods 5 – 6 of normal width, with posteroproximal process produced into a sharp tooth, with posterodistal tooth strong, narrow and sharp; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. PEREIOPOD 7. Basis broad; posterior border nearly parallel to basis axis of proximal 0.75; at this level it forms a medium-sized sharp to very sharp tooth, which is followed by a deep rounded concavity; the posterodistal corner is produced into a medium to long narrow tooth pointing obliquely backwards; merus, carpus and propodus extremely broad and short; dactylus fairly large, strongly curved on both sides, clasping. Body length Up to 35 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0F68F4FE1DFEE3CEA5FE1D.taxon	distribution	Distribution Plateau of the South Orkney Islands, 50 – 190 m (Coleman 1990; present material).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0F68F4FE1DFEE3CEA5FE1D.taxon	discussion	Remarks Epimeria pulchra is superficially similar to E. amoenitas sp. nov. and E. debroyeri sp. nov., but it differs from these two species by a number of characters such as: longer mid-dorsal teeth on pereionites 3 – 7 and pleonites 1 – 2, much longer mid-dorsal tooth on pleonite 3, anterior corner of coxa 4 much more produced, posterior border of third epimeral plate produced into a tooth instead of being simply convex, gnathopods much broader. It is likely that E. pulchra s. str. is endemic to the South Orkney Islands, just as E. linseae sp. nov.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0D68F3FDE4FE55CFFAFCFB.taxon	type_taxon	Type species Subepimeria geodesiae Bellan-Santini, 1972.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0D68F3FDE4FE55CFFAFCFB.taxon	description	Description Body opaque, with teguments moderately calcified. Rostrum medium-sized. Eyes not conical. Pleonite 2 posteriorly produced into a small sharp tooth oriented backwards; pleonite 1 and pereionite 7 sometimes also posteriorly produced into a tooth or bump; dorsal border of pleonite 3 keeled; in lateral view that keel posteriorly terminated into a rounded bump or a squared angle. With the exception of the aforementioned mid-dorsal ornamentations, pereionites and pleosomites perfectly smooth. Coxae 1 – 3 with blunt tip, not sharply keeled. Coxa 4 with indistinct carina starting at ventral tip of coxa and terminating at posterior tip of coxa; this carina remains very close to the posteroventral border of coxa, the space in-between forming a very shallow groove; posteroventral border of coxa 4 very weakly concave, nearly straight. Coxae 5 – 6 without tooth or distinct protrusion. Mid of posterior border of epimeral plates 1 – 3 not produced into a tooth. Posteroventral tooth of epimeral plate 3 medium-sized to strong. Dorsal process of urosomite 1 produced into a triangular tooth directed upwards. Urosomite 2 without pair of small teeth pointing upwards. Lateral borders of urosomite 3 posteriorly terminated into a sharp tooth or bluntly angular. Peduncle of antenna 1 without teeth or with dentition vestigial. Mandible with pars molar process triturative. Lower lip with narrow (V-shaped) hypopharyngeal gap. Palp of maxilliped with 4 articles. Gnathopods of normal size, with carpus and propodus of medium slenderness, with palm obliquely transverse but poorly developed (gnathopods subcheliform); propodus not expanded distally; posterior border of dactylus lined by row of small oblique slender teeth. Basis of pereiopods 5 – 6 moderately broad, with posteroproximal rounded protrusion (sometimes very weak and scarcely distinct); a posterodistal tooth projecting posteriorly can be present on basis of pereiopod 5, otherwise basis posterodistally angulate or bluntly angulate. Posterior border of basis of pereiopod 7 slightly convex in proximal 0.8; at this level there is a slight angular discontinuity and the distal 0.2 can be slightly concave; posterior border terminated into a blunt angle or a blunt tooth directed in the axis of the basis. Dactylus of pereiopods 5 – 7 short. Benthic. Small species. Body length The maximum body length recorded in Subepimeria species ranges between 14 and 16 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0D68F3FDE4FE55CFFAFCFB.taxon	biology_ecology	Ecology. Benthic, 52 – 840 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0D68F3FDE4FE55CFFAFCFB.taxon	distribution	Distribution Circum-Antarctic, as far north as South Georgia.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0D68F3FDE4FE55CFFAFCFB.taxon	discussion	Remarks Molecular data (COI, 28 S) (Verheye et al. 2016 a) indicate that Subepimeria and Drakepimeria are sister clades (Fig. 342). The morphology of their gnathopods and pereiopods is similar. However, Subepimeria species are much smaller (<20 mm) and have only one or two (smaller) mid-dorsal teeth and no dorsolateral teeth on pereionites and pleosomites. The lateral carina of coxa 4 is also vestigial in Subepimeria, whilst it is very strongly developed in Drakepimeria. We believe that these conspicuous morphological differences justify the recognition of distinct subgenera for these clades. The morphological identification of species within the subgenus Subepimeria is very difficult. Interspecific differences are based on ill-defined characters, such as the curves of coxa 4. The species are also rare, which largely prevents the study of individual and allometric differences within species. The link between immatures and adult specimens of Epimeria (Subepimeria) iota is tentative as only immature specimens were available for sequencing. As Subepimeria species can only be distinguished by a combination of illdefined characters, we delineate their differences in a tabular format (Table 1) instead of an identification key.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0A68F2FDDFFC72CBCDF862.taxon	description	urn: lsid: zoobank. org: act: 76 F 9695 C- 487 B- 434 F-B 09 C- 52 CA 334 ACD 5 B Figs 294 – 298	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0A68F2FDDFFC72CBCDF862.taxon	etymology	Etymology The noun Adelia, - ae is a Latinization of Adèle (referring to Adèle Pépin, the wife of Jules Sébastien César Dumont d’Urville), and alludes to Adélie Coast (discovered by J. S. C. Dumont d’Urville), where the type material of the species was collected. The name is a genitive.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0A68F2FDDFFC72CBCDF862.taxon	materials_examined	Type material Holotype RV Aurora Australis cruises: SOUTHERN OCEAN: ♀, cruise CEAMARC, sample CEAMARC 2072, stn 65 EV 322, Adélie Coast, 65 ° 48 ʹ 09 ″ S, 143 ° 03 ʹ 46 ″ E, 750 – 788 m, beam trawl, 5 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 4288) [extraction M 4; Genbank nr, COI: KU 870888, 28 S: KU 759671]. Paratypes RV Aurora Australis cruises: SOUTHERN OCEAN: 1 spec., cruise CEAMARC, sample CEAMARC 2072, stn 65 EV 322, Adélie Coast, 65 ° 48 ʹ 09 ″ S, 143 ° 03 ʹ 46 ″ E, 750 – 788 m, beam trawl, 5 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 7339, removed from MNHN-IU- 2014 - 4288); 1 spec., cruise CEAMARC, sample CEAMARC 2072, stn 65 EV 322, Adélie Coast, 65 ° 48 ʹ 09 ″ S, 143 ° 03 ʹ 46 ″ E, 750 – 788 m, beam trawl, 5 Jan. 2008, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 7340, removed from MNHN-IU- 2014 - 4288) [extraction M 3; Genbank nr, 28 S: KU 759670].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0A68F2FDDFFC72CBCDF862.taxon	description	Description ROSTRUM. In lateral view short and narrow, nearly reaching tip of article 1 of peduncle of antenna 1, distinctly curved on the proximal half of its anterior border, posterior border straight, tip acute; in frontal view triangular: fairly broad and with straight converging borders, with tip blunt. EYES. Large, broadly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 6 totally smooth; pereionite 7 keeled with small blunt posterodorsal tooth; pleonites 1 – 2 keeled with well developed acute posterodorsal tooth; pleonite 3 keeled with posterodorsal tip forming a distinct squared angle. COXAE 1 – 3. Tip blunt but narrow. COXA 4. Of medium width; anterodorsal border proximally nearly straight, curving ventrally and followed by curved anteroventral border (transition between anterodorsal and anteroventral border very gradual, without distinct anterior corner); the coxa is not projecting forward; ventral corner forming a squared angle of which the tip is blunt but not rounded; posteroventral border nearly straight (very weakly concave); posterodorsal border as long as posteroventral border. COXA 5. Very broad, posteroventral corner forming a blunt but distinct angle. COXA 6. Posterior border regularly rounded. COXA 7. Posterior border nearly straight (very weakly convex); posteroventral corner broadly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle blunt and squared in plate 1, produced into a medium-sized tooth in plate 2, into a large tooth in plate 3. UROSOME TOOTH PATTERN. Urosomite 1 with distinct triangular dorsal process, anteriorly weakly concave, tip subacute, posterior border weakly convex. TELSON. Cleft on 0.4; lobes with tips bluntly angulate; notch narrowly V-shaped. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; propodus not narrowing distally, and palm distinct. PEREIOPOD 5. Basis of normal width, with posteroproximal process forming a low but distinct rounded lobe, with posterodistal corner forming a long, narrowly triangular blunt-tipped tooth; merus, carpus and propodus stout. PEREIOPOD 6. Basis of normal width, with posteroproximal process very low and rounded, posterior border nearly straight, diverging from anterior border and terminated in blunt squared angle; merus, carpus and propodus stout. PEREIOPOD 7. Basis broad; posterior border weakly convex, with inconspicuous trace of concavity in distal 0.9, terminated into a blunt, obtuse (nearly squared) angle. Body length Up to 16 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0A68F2FDDFFC72CBCDF862.taxon	distribution	Distribution Adélie Coast, 750 – 788 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0A68F2FDDFFC72CBCDF862.taxon	discussion	Remarks The pattern of dorsal teeth of Epimeria adeliae sp. nov. is reminiscent to that of E. (Subepimeria) sp. 1 from South Georgia and different from that of other species of the subgenus Subepimeria.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0868EFFE3BFEE3CE0EF851.taxon	description	Description Description based on the illustrations of Bellan-Santini (1972). ROSTRUM. Reaching tip of article 1 of peduncle of antenna 1, broad in lateral view. EYES. Large, rounded. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 7 totally smooth; pleonite 1 toothless; pleonite 2 with small, posterodorsal tooth; pleonite 3 with posterodorsal tip forming a small, distinct rounded (very broadly triangular) lobe projecting backwards. COXAE 1 – 3. Tip subacute in coxae 1 – 2, blunt in coxa 3. COXA 4. Very narrow; anterodorsal border weakly convex, directly followed by very broadly rounded ventral lobe (anteroventral border absent); posteroventral border nearly straight (weakly concave); ratio length of posterodorsal border / length of posteroventral border: 0.5. COXA 5. Very broad, posteroventral corner with broadly rounded angular discontinuity. COXA 6. Posterior border regularly rounded. COXA 7. Posterior border nearly straight (very weakly convex); posteroventral corner broadly rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle angulate in plate 1, produced into a small tooth in plate 2; produced into a medium-sized tooth in plate 3. UROSOME TOOTH PATTERN. Urosomite 1 with narrow triangular dorsal process; urosomite 3 with dorsolateral borders straight, with tip produced into a long tooth pointing obliquely. TELSON. Cleft on 0.25. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; propodus not narrowing distally, and palm distinct. PEREIOPOD 5. Basis of normal width, with posteroproximal process indistinct (reduced to very low proximal dilatation in continuity with the more distal part of the posterior border), with posterodistal corner forming a blunt-tipped broadly triangular process (acute angle) projecting backwards; merus, carpus and propodus stout. PEREIOPOD 6. Basis of normal width, with posteroproximal process indistinct (reduced to very low proximal dilatation in continuity with the more distal part of the posterior border), with posterior border nearly straight and parallel to anterior border, with posterodistal corner forming a blunt-tipped broadly triangular process (acute but nearly squared angle) scarcely projecting backwards; merus, carpus and propodus stout. PEREIOPOD 7. Basis broad; posterior border weakly convex, with shallow concavity in distal 0.8, terminated into a sharp triangular tooth (forming a squared angle). Body length 15 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0868EFFE3BFEE3CE0EF851.taxon	distribution	Distribution Adélie Coast: Cape Géodésie, 115 – 135 m, on coarse sand with abundant epifauna: hydroids, bryozoans and sponges (Bellan-Santini 1972).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF0868EFFE3BFEE3CE0EF851.taxon	discussion	Remarks Watling & Holman (1981) synonymised Subepimeria geodesiae with Epimeria puncticulata, which they considered as a widely distributed variable species. Molecular data (COI, 28 S) revealed different species within E. puncticulata s. lat. (Verheye et al. 2016 a; this paper Fig. 342), leading to the resurrection of Subepimeria, as a subgenus. The shape of coxa 4 of E. geodesiae as illustrated by Bellan-Santini (1972) is unique and does not fit with the descriptions of the holotype of E. puncticulata (K. H. Barnard 1930; Coleman 1994) or any other Subepimeria species. Epimeria geodesiae is considered here to be a valid species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1768ECFDCBFEE3CB32FD69.taxon	description	urn: lsid: zoobank. org: act: 7 A 557 F 8 F- 058 F- 413 B- 998 C-AED 9 C 9 D 23 EE 6 Figs 299 – 306	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1768ECFDCBFEE3CB32FD69.taxon	etymology	Etymology Iota is the smallest letter of the Greek alphabet. The name, which is a noun in apposition, alludes to the small size of the species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1768ECFDCBFEE3CB32FD69.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: sex undetermined, cruise PS 81, ANT-XXIX / 3, stn 197 - 6, Bransfield Strait, 62 ° 45.05 ʹ S, 57 ° 26.68 ʹ W to 62 ° 45.09 ʹ S, 57 ° 26.47 ʹ W, 210 – 222 m, black gravel mixed with sand and a little bit of mud, Rauschert dredge, 25 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122947) [extraction ANT 48; Genbank nr, 28 S: KU 759607]. Paratypes RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., cruise PS 69, ANT-XXIII / 8, stn 605 - 3, Elephant Island, 61 ° 20.33 ʹ S, 55 ° 31.53 ʹ W to 61 ° 20.35 ʹ S, 55 ° 30.18 ʹ W, 148 – 154 m, Agassiz trawl, 20 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122519); 2 specs, cruise PS 69, ANT-XXIII / 8, stn 614 - 3 / 4 / 5, Elephant Island, 60 ° 52.37 ʹ S, 55 ° 29.80 ʹ W to 60 ° 52.71 ʹ S, 55 ° 27.83 ʹ W, 248 – 265 m, a lot of epifauna, Rauschert dredge and Agassiz trawl, 22 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122525); 1 spec., sex undetermined, cruise PS 81, ANT-XXIX / 3, stn 164 - 5, south of Dundee Island, 63 ° 36.84 ʹ S, 56 ° 10.28 ʹ W to 63 ° 36.72 ʹ S, 56 ° 10.46 ʹ W, 121 – 122 m, Rauschert dredge, 11 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122934) [extraction K 42; Genbank nr, 28 S: KU 759655].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1768ECFDCBFEE3CB32FD69.taxon	description	Description Basis of description. Description based on holotype. ROSTRUM. In lateral view fairly short and fairly narrow; distinctly curved, posterior border straight, tip acute; in frontal view narrowly triangular, fairly narrow and with almost straight converging borders, with tip blunt; not reaching tip of article 1 of peduncle of antenna 1. EYES. Very large, nearly circular (very broadly elliptic). PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 7 and pleonite 1 totally smooth; pleonite 2 with small, posterodorsal tooth; pleonite 3 carinate, with posterodorsal tip forming a small, low, rounded lobe very weakly projecting backwards. COXAE 1 – 3. Tip blunt. COXA 4. Narrow; anterodorsal border nearly straight, curving ventrally and followed by weakly curved anteroventral border; anterior corner very rounded but distinct, projecting forward; ventral corner forming a broadly rounded squared angle; posteroventral border distinctly concave; ratio length of posterodorsal border / length of posteroventral border: 0.7. COXA 5. Very broad, posteroventral corner with broadly rounded angular discontinuity. COXA 6. Posterior border rounded. EPIMERAL PLATES 1 – 3. Posteroventral angle rounded in plate 1, produced into a small tooth in plate 2; produced into a medium-sized tooth in plate 3. UROSOME TOOTH PATTERN. Urosomite 1 with medium-sized blunt triangular dorsal process; urosomite 3 with dorsolateral borders straight, with tip bluntly angulate. TELSON. Broad, with lateral border strongly converging, with lobes distally broad, slit narrowly U-shaped, cleft on 0.3. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; propodus not narrowing distally, and palm weak but distinct. PEREIOPOD 5. Basis of normal width, with posteroproximal process indistinct (reduced to very low proximal dilatation in continuity with the more distal part of the posterior border), with posterodistal corner forming a blunt-tipped broadly triangular process (squared angle), not projecting backwards; merus, carpus and propodus stout. PEREIOPOD 6. Basis of normal width, with posteroproximal process indistinct (reduced to very low proximal dilatation in continuity with the more distal part of the posterior border), with posterior border parallel to anterior border, with posterodistal corner forming a rounded triangular process (very rounded squared angle) not projecting backwards; merus, carpus and propodus stout. PEREIOPOD 7. Basis broad; posterior border nearly straight, with trace of concavity in distal 0.8, terminated into an obtuse rounded lobe. Variations The sequenced holotype is obviously immature. Adult specimens preserved in formalin interpreted as adult Epimeria iota sp. nov. have broader coxa 4, a stronger posterodistal tooth on the basis of pereiopods 5 and 6, sometimes a trace of a posterodorsal tooth or bump on pleonite 1 and a more obtuse dorsal process on urosomite 1. Colour pattern Holotype: background whitish (some areas very pale pinkish or very pale yellowish); head, pereionites 3 – 5, lower part of epimeral plates, posterior border of pleonite 3 and urosomites 1 – 2 suffused with orange-red; two red dots on the anterior part of coxa 4; a large red dot on first epimeral plate; eyes reddish. Body length The holotype is 9.5 mm. Non sequenced specimens interpreted as adult E. iota sp. nov. reach 16 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1768ECFDCBFEE3CB32FD69.taxon	distribution	Distribution Tip of Antarctic Peninsula and Elephant Island, 121 – 265 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1768ECFDCBFEE3CB32FD69.taxon	discussion	Remarks The specimens from the stations 605 - 3 and 614 - 3 / 4 / 5 of ANT-XXIII / 8 are interpreted as adult Epimeria (Subepimeria) iota sp. nov. They possess the following character states of E. iota sp. nov.: (1) the anterior margin of their coxa 4 has an angular discontinuity (forming a regular curve in E. teres sp. nov.), and (2) the margins of the basis of pereiopod 6 are parallel (diverging in E. teres sp. nov.). The ‘ Epimeria puncticulata ’ illustrated by Watling & Holman (1981) and the ‘ Epimeria aff. puncticulata ’ illustrated by Rauschert & Arntz (2015: 61, pl. 54) presumably belong to the same morphospecies as well. The collection details are not given by Rauschert & Arntz (2015), but are indicated in an early draft of their book made available to the authors: ANT-XXI / 2 stn 276 [i. e., 276 - 1], 71 ° 06.44 ʹ S, 11 ° 27.76 ʹ W to 71 ° 06.64 ʹ S, 11 ° 27.28 ʹ W, 268 – 277 m, eastern shelf of the Weddell Sea.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1568EBFE37FCEFCE28F997.taxon	description	Description Basis of description. Description based on the accounts on the holotype published by K. H. Barnard (1930) and Coleman (1994); when illustrations are conflicting, description based on Coleman (1994). ROSTRUM. In lateral view very narrow. EYES. Medium-sized, narrowly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 7 totally smooth; pleonite 1 with feeble indication of a keel; pleonite 2 keeled with well developed acute posterodorsal tooth; pleonite 3 keeled with posterodorsal tip forming a distinct blunt process distinctly projecting backwards. COXAE 1 – 2 (cf. Coleman 1994). Tip subacute to rather blunt; coxa 3: tip rounded. COXA 4 (cf. Coleman 1994). Fairly narrow; anterodorsal border nearly straight (inconspicuously convex); anteroventral border weakly convex; anterior corner very broadly rounded but forming a distinct angular discontinuity; the coxa is slightly projecting forward; ventral corner forming a squared angle of which the tip is subacute; posteroventral border distinctly concave; posterodorsal border 0.8 × as long as posteroventral border. COXA 5. Very broad, posteroventral corner forming a blunt but distinct squared angle. COXA 6. Posterior border regularly rounded. COXA 7. Posterior border straight; posteroventral corner forming a distinct obtuse angle. EPIMERAL PLATES 1 – 3. Posteroventral angle with trace of tooth in plate 1, produced into a small tooth in plate 2, produced into a medium-sized tooth in plates 3. UROSOME TOOTH PATTERN. Urosomite 1 with triangular dorsal process. TELSON. Cleft on 0.2; lobes with tips rounded; notch V-shaped. GNATHOPODS 1 – 2. With carpus and propodus of normal slenderness; propodus not narrowing distally, and palm distinct but weak. PEREIOPOD 5. Basis of normal width, with posteroproximal process reduced to low proximal dilatation in continuity with the more distal part of the posterior border, with posterodistal corner forming a fairly narrow, acutely triangular tooth projecting backwards; merus, carpus and propodus stout. PEREIOPOD 6. Basis of normal width, with posteroproximal process reduced to very low proximal dilatation nearly in continuity with the more distal part of the posterior border), with posterior border parallel to anterior border, with posterodistal corner forming a triangular process (acute, nearly squared angle) very weakly projecting backwards; merus, carpus and propodus stout. PEREIOPOD 7. Basis broad; posterior border weakly convex, with inconspicuous trace of concavity in distal 0.9, terminated into a very blunt, obtuse angle. Body length Up to 16 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1568EBFE37FCEFCE28F997.taxon	distribution	Distribution Ross Sea, McMurdo Sound, 175 m (K. H. Barnard 1930).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1568EBFE37FCEFCE28F997.taxon	discussion	Remarks Watling & Holman (1981) and Coleman (2007) considered Epimeria puncticulata as a single widespread and variable species. Yet, genetic data (COI, 28 S) (Verheye et al. 2016 a) based on a limited material indicate the existence of four different species refered herein as E. adeliae sp. nov., E. iota sp. nov., E. teres sp. nov. and E. urvillei sp. nov. (Fig. 342). Examination of published illustrations suggest the existence of four additional species: E. geodesiae, E. puncticulata s. str., E. (Subepimeria) sp. 1 and E. (Subepimeria) sp. 2. Coxa 4 of the holotype of E. puncticulata is different in the drawings of K. H. Barnard (1930) and those of Coleman (1994). This might result from different orientations of the specimens.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1268E9FDC0F9A2CE9BFA0E.taxon	description	urn: lsid: zoobank. org: act: 87 DB 1 EDA- 29 A 4 - 493 E- 84 BF- 9 A 68 EFF 422 ED Figs 307 – 313	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1268E9FDC0F9A2CE9BFA0E.taxon	etymology	Etymology Teres, teres, terete, Latin adjective meaning round and smooth, which seems appropriate for a very smooth Epimeria species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1268E9FDC0F9A2CE9BFA0E.taxon	materials_examined	Type material Holotype RV Polarstern cruises: SOUTHERN OCEAN: adult, sex undetermined, cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 248 - 2, Larsen B, 65 ° 57.51 ʹ S, 60 ° 28.15 ʹ W to 65 ° 57.69 ʹ S, 60 ° 28.30 ʹ W, 196 – 202 m, Agassiz trawl and Rauschert dredge, 7 Mar. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 132951) [extraction I 2; Genbank nr, COI: KU 870845, 28 S: KU 759622]. Paratype RV Polarstern cruises: SOUTHERN OCEAN: 1 juv., sex undetermined, cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 248 - 2, Larsen B, 65 ° 57.51 ʹ S, 60 ° 28.15 ʹ W to 65 ° 57.69 ʹ S, 60 ° 28.30 ʹ W, 196 – 202 m, Agassiz trawl and Rauschert dredge, 7 Mar. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 122896).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1268E9FDC0F9A2CE9BFA0E.taxon	description	Description ROSTRUM. In lateral view fairly short and narrow, reaching tip of article 1 of peduncle of antenna 1, weakly and regularly curved on anterior border, posterior border straight, tip very acute; in frontal view triangular: narrow, with straight converging borders, with tip blunt. EYES. Large, broadly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 7 totally smooth; pleonite 1 keeled along all its length, posteriorly terminated by tiny but distinct bump; pleonite 2 keeled with well developed acute posterodorsal tooth; pleonite 3 keeled with posterodorsal tip forming a distinct blunt process (shape: acute angle) distinctly projecting backwards. COXAE 1 – 3. Tip acute. COXA 4. Narrow; anterodorsal and anteroventral border forming a continuous curve without any trace of discontinuity (there is no distinct anterior corner), not curving significantly more ventrally; the coxa is not projecting forward; ventral corner forming an acute (nearly squared) angle of which the tip is blunt but not broadly rounded; posteroventral border distinctly concave; posterodorsal border 0.9 × as long as posteroventral border. COXA 5. Very broad, posteroventral corner forming a blunt but distinct obtuse (nearly squared) angle. COXA 6. Posterior border regularly rounded. COXA 7. Posterior border straight; posteroventral corner forming a distinct obtuse angle. EPIMERAL PLATES 1 – 3. Posteroventral angle: forming a squared angle in plate 1, produced into a tiny tooth in plate 2, produced into a small tooth in plate 3. UROSOME TOOTH PATTERN. Urosomite 1 with distinct triangular dorsal process, anteriorly weakly concave, tip straight, posterior border nearly straight (inconspicuously concave); urosomite 3 with dorsolateral borders straight, with tip forming a squared angle. TELSON. Cleft on 0.20; lobes with tips broad; notch broadly V-shaped. GNATHOPODS 1 – 2. Carpus and propodus of normal slenderness; propodus not narrowing distally, and palm distinct but weak. PEREIOPOD 5. Basis of normal width, with posteroproximal process reduced to low proximal dilatation in continuity with the more distal part of the posterior border, with posterodistal corner forming a long, narrowly triangular tooth projecting backwards; merus, carpus and propodus stout. PEREIOPOD 6. Basis of normal width, with posteroproximal process reduced to low proximal dilatation in continuity with the more distal part of the posterior border, with posterior border distinctly diverging from anterior border, with posterodistal corner forming a triangular process (acute, nearly squared angle) weakly projecting backwards; merus, carpus and propodus stout. PEREIOPOD 7. Basis broad; posterior border nearly straight, with weak but distinct concavity in distal 0.8, terminated into a tooth forming a squared angle. Body length Up to 16 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1268E9FDC0F9A2CE9BFA0E.taxon	distribution	Distribution Western Weddell Sea: Larsen B, 196 – 202 m; probably eastern shelf of the Weddell Sea, 268 – 277 m (see remarks).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1268E9FDC0F9A2CE9BFA0E.taxon	discussion	Remarks Epimeria (Subepimeria) teres sp. nov. is morphologically very similar to E. (S.) urvillei sp. nov., while genetic data (COI, 28 S) Verheye et al. (2016 a) suggests that they are different species (Fig. 342). In E. (S.) teres sp. nov., the rostrum is a bit longer, the eyes slightly more rounded, coxa 4 a bit narrower, and the tip of the basis of pereiopod 7 more produced than in E. (S.) urvillei sp. nov. It is likely that the ‘ Epimeria aff. puncticulata ’ illustrated by Rauschert & Arntz (2015) is E. (S.) teres sp. nov., as it has a narrow coxa 4, a very strong posterodorsal protrusion on pleonite 3, a very strong dorsal protrusion on urosomite 1 and a posterodistal tooth on the basis of pereiopod 7. Rauschert & Arntz (2015) did not give the collection details of their specimen, but this information was present in an early draft of their book made available to the authors. It is indicated that the specimen was collected during the cruise ANT-XXI (obviously ANT-XXI / 2) at station 276 (i. e., 276 - 1 as there are no other sub-stations). The coordinates of this station are: 71 ° 06.44 ʹ S, 11 ° 27.76 ʹ W to 71 ° 06.64 ʹ S, 11 ° 27.28 ʹ W [eastern shelf of the Weddell Sea], 268 – 277 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1068E7FDDFFA47CA60FD78.taxon	description	urn: lsid: zoobank. org: act: 524 C 2 AA 7 - 3 B 7 D- 4 EDF-B 336 - 0 A 2 D 2 D 5 BE 294 Figs 314 – 319	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1068E7FDDFFA47CA60FD78.taxon	etymology	Etymology The species is dedicated to the memory of the French explorer Jules Sébastien César Dumont d’Urville (23 May 1790 – 8 May 1842), who discovered Adélie Coast, where the species was collected. The name is a genitive.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1068E7FDDFFA47CA60FD78.taxon	materials_examined	Type material Holotype RV Seatruck cruises: SOUTHERN OCEAN: ovigerous ♀, cruise REVOLTA III, stn. REVO _ 084, Collect _ ID: REVO _ 477, Field _ ID: CE- 000003093, Adélie Coast, 66 ° 39.279 ʹ S, 139 ° 55.846 ʹ E to 66 ° 39.291 ʹ S, 139 ° 56.043 ʹ E, 52 – 54 m, beam trawl, 1 Feb. 2012, Coll. G. Lecointre, A. Dettaï, J. Lanshere, C. Gallut and C. Ozouf, (MNHN-IU- 2009 - 2578) [Extraction K 33; Genbank nr, COI: KU 870867, 28 S: KU 759646].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1068E7FDDFFA47CA60FD78.taxon	description	Description ROSTRUM. In lateral view short and narrow, not reaching tip of article 1 of peduncle of antenna 1, very weakly and regularly curved on anterior border, posterior border very weakly concave, tip acute; in frontal view triangular: fairly narrow, with straight converging borders, with tip blunt. EYES. Large, narrowly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 7 totally smooth; pleonite 1 keeled all along its length, posteriorly terminated by tiny but distinct bump; pleonite 2 keeled with well developed acute posterodorsal tooth; pleonite 3 keeled with posterodorsal tip forming a distinct blunt triangular process distinctly projecting backwards. COXAE 1 – 2. Tip subacute. COXA 4. Fairly narrow; anterodorsal border nearly straight, followed by anterior angle + anteroventral border, which form a very regular curve (without discontinuity) — there is no distinct anterior corner; the coxa is not projecting forward; ventral corner forming an acute (nearly squared) angle of which the tip is subacute; posteroventral border distinctly concave; posterodorsal border 1.2 × as long as posteroventral border. COXA 5. Very broad, posteroventral corner forming a blunt but distinct obtuse (nearly squared) angle. COXA 6. Posterior border regularly rounded. COXA 7. Posterior border straight; posteroventral corner forming a distinct obtuse angle. EPIMERAL PLATES 1 – 3. Posteroventral angle broadly rounded in plate 1, produced into a medium-sized tooth in plates 2 – 3. UROSOME TOOTH PATTERN. Urosomite 1 with distinct triangular dorsal process, anteriorly weakly concave, tip subacute, posterior border nearly straight (inconspicuously concave); urosomite 3 with dorsolateral borders straight, with tip forming a squared angle. TELSON. Cleft on 0.25; lobes with tips bluntly angulate; notch narrowly V-shaped. GNATHOPODS 1 – 2. With carpus and propodus of normal slenderness; propodus not narrowing distally, and palm distinct but weak. PEREIOPOD 5. Basis of normal width, with posteroproximal process reduced to low proximal dilatation in continuity with the more distal part of the posterior border, with posterodistal corner forming a long, narrowly triangular sharp tooth projecting backwards; merus, carpus and propodus stout. PEREIOPOD 6. Basis of normal width, with posteroproximal process indistinct (reduced to very low proximal dilatation in continuity with the more distal part of the posterior border), with posterior border slightly but distinctly diverging from anterior border, with posterodistal corner forming a triangular process (acute, nearly squared angle) weakly projecting backwards; merus, carpus and propodus stout. PEREIOPOD 7. Basis broad; posterior border weakly convex, with inconspicuous trace of concavity in distal 0.9, terminated into a very blunt, obtuse angle. Colour pattern Whitish with scattered orange dots. Rostrum, coxae 1 – 4, anterior 0.2 of coxa 5, pleon suffused with pale orange. Eyes reddish. Body length 16 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1068E7FDDFFA47CA60FD78.taxon	distribution	Distribution Adélie Coast, 52 – 54 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1068E7FDDFFA47CA60FD78.taxon	discussion	Remarks Epimeria urvillei sp. nov. is morphologically very similar to E. teres sp. nov. and E. puncticulata. Further molecular and morphological studies are necessary to sort out completely the taxonomy of this complex.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1E68E5FD9FFCF1CB5CFE5E.taxon	materials_examined	Material (not examined) Discovery Investigations 1926 – 1927: R. V. Discovery cruises: SOUTHERN OCEAN: 1 ♀, 13 mm, stn 45, South Georgia, 2.7 miles S 85 ° E of Jason Light, N 4 - T (net with mesh of 4 mm attached to the back of the trawl) and NCS-T (tow-net of coarse silk, with 16 meshes to the linear inch, attached to trawl), 238 – 270 m, grey mud, 6 Mar. 1926 (NHM 1936.11.2. 1581 – 1590); 3 ♀♀, 11 – 13 mm, stn 123, South Georgia, off mouth of Cumberland Bay, N 4 - T (net with mesh of 4 mm attached to the back of the trawl) and OTL (large otter trawl), 230 – 250 m, grey mud, 15 Dec. 1926 (NHM 1936.11.2. 1581 – 1590); 8 ♀♀, 10 – 13 mm, stn 140, South Georgia, Stromness Harbour to Larsen Point, N 4 - T (net with mesh of 4 mm attached to the back of the trawl), 122 – 136 m, green mud, stn, 23 Dec. 1926, (NHM 1936.11.2. 1581 – 1590); 1 ♂, 10.5 mm, stn 148, South Georgia, off Cape Saunders, N 4 - T (net with mesh of 4 mm attached to the back of the trawl) and OTL (large otter trawl), 200 – 234 m, grey mud, stn, 10 Jan. 1927 (NHM 1936.11.2. 1581 – 1590). R. V. William Scoresby cruises: SOUTHERN OCEAN: 1 ♂, 7 mm, stn WS 33, South Georgia, 54 ° 59 ʹ S, 35 ° 24 ʹ W, N 100 H (1 m tow net), 135 m, grey mud, 21 Dec. 1926 (NHM 1936.11.2. 1581 – 1590); 1 juv., 5.5 mm, Marine Biological Station, stn MS 71, South Georgia, East Cumberland bay, BTS (small beam trawl) and NCS-T (townet of coarse silk, with 16 meshes to the linear inch, attache to trawl), 60 – 110 m, 9 Mar. 1926 (NHM 1936.11.2. 1581 – 1590).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1E68E5FD9FFCF1CB5CFE5E.taxon	description	Description Description based on the photograph of Rauschert & Arntz (105) and K. H. Barnard’s (1932) account. ROSTRUM. in lateral view short reaching about tip of article 1 of peduncle of antenna 1, distinctly curved on the proximal half of its anterior border. EYES. Large, broadly elliptic. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 6 totally smooth; pereionite 7 keeled with small blunt posterodorsal tooth; pleonites 1 – 2 keeled with well developed acute posterodorsal tooth; pleonite 3 keeled with posterodorsal tip forming a very strong blunt protrusion. COXAE 1 – 3. Tip blunt. COXA 4. Of medium width; anterodorsal border proximally nearly straight; anteroventral border straight (transition between anterodorsal and anteroventral border gradual, but with rather distinct anterior corner); the coxa is not projecting forward; ventral corner forming a squared angle, of which the tip is blunt but not rounded; posteroventral border nearly straight (very weakly concave); posterodorsal border as long as posteroventral border. EPIMERAL PLATES 2 – 3. Produced into a medium-sized tooth. UROSOME TOOTH PATTERN. Urosomite 1 with low and blunt triangular dorsal process. BASIS OF PEREIOPOD 7. Broad; posterior border weakly convex, terminated into a blunt, obtuse (nearly squared) angle. Colour pattern “ Pale yellow. Each of first 7 segments with a transverse pale brown band dorsally along both anterior and posterior margins. Large side-plate of 4 th segment largely red-brown. Posterior segments yellow mottled with brown. Caudal appendages and legs pale yellow. Eyes red ” (K. H. Barnard 1932). Whitish with scattered orange dotting and mottling; peduncle of antennae and dorsal process of urosomite 1 red; eyes red (specimen photographed by M. Rauschert). Body length 13 mm (K. H. Barnard 1932) or 14 mm (specimen photographed by M. Rauschert).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1E68E5FD9FFCF1CB5CFE5E.taxon	distribution	Distribution South Georgia, 60 – 270 m (K. H. Barnard 1932 as Epimeria puncticulata); “ Scotia Arc ”, 280 m (Rauschert & Arntz 2015 as Epimeria sp. 1).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1E68E5FD9FFCF1CB5CFE5E.taxon	discussion	Remarks From discussions with C. O. Coleman and W. Arntz, it transpires that large parts of Rauschert’s collections are not yet registered and thus currently inaccessible. Therefore, the “ Epimeria sp. n. 1 ” from Rauschert & Arntz (2015) could not be found and its precise collection locality is unknown. An early draft of the book of Rauschert & Arntz (2015) simply indicated that it was collected during the cruise ‘ ANT XIX’ in the ‘ Scotia Arc’. However, the diagnosis of ‘ Epimeria puncticulata ’ from South Georgia by K. H. Barnard (1932) is in agreement with the photograph of ‘ Epimeria sp. n. 1 ’: “ the 7 th peraeon segment ends in a short medio-dorsal point, the keels on pleon segments 1 and 2 end in slightly more prominent points, and the triangular projection on segment 4 is more sharply pointed than in the Terra Nova specimens. ” M. Rauschert took part in cruise ANT-XIX / 5, which sampled extensively around South Georgia (Arntz & Brey 2003). Therefore, there is a high probability that Rauschert’s specimens were collected off South Georgia, just as the specimens of K. H. Barnard (1932). Because of time constraints, it was impossible to borrow and to re-examine K. H. Barnard’s (1932) specimens, deposited in the Natural History Museum in London: NHM 1936.11.2. 1581 – 1590, six tubes together in the same jar, no specific number allocated to each tube (M. Lowe pers. com.). The species is treated herein as Epimeria (Subepimeria) sp. 1. When examination of specimens will be possible, it will presumably have to be described as a new species. The dorsal tooth pattern of the Epimeria (Subepimeria) sp. 1 is very different from that of other West-Antarctic Subepimeria examined. It is similar to that of Epimeria adeliae sp. nov. from Adélie Coast. However, the shape of the posterodorsal protrusion on pleonite 3 is different: blunt in Epimeria (Subepimeria) sp. 1 (see Rauschert & Arntz 2015, as Epimeria sp. 1), whereas it forms a squared angle in E. adeliae sp. nov. The anteroventral border of coxa 4 is also more straight and the protrusion on urosomite 1 lower in Epimeria (Subepimeria) sp. 1 than in E. adeliae sp. nov.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1C68E5FD9FFE17CDF4F868.taxon	description	Discription EYES. Large and narrowly elliptic. PEREIONITE 7. Posterodorsally smooth. PLEONITE 1. Posterodorsally smooth. PLEONITE 2. With small sharp posterodorsal tooth. PLEONITE 3. Posterodorsally produced into a blunt acute (nearly squared) angle. COXA 4. with ventral corner produced into a distinct tooth (Coleman 2007), or apparently blunt (Rauschert & Arntz 2015) [specimen not photographed in a flat orientation?]. UROSOME TOOTH PATTERN. Urosomite 1 with well developed triangular dorsal process. Colour pattern Eyes red. Body whitish to yellowish, with some areas more brownish. Distinct brown transverse stripes on body segment 4 and coxa 4. Body length 17 mm (Rauschert & Arntz 2015).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1C68E5FD9FFE17CDF4F868.taxon	distribution	Distribution Eastern shelf of the Weddell Sea, 765 – 840 m (see remarks).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1C68E5FD9FFE17CDF4F868.taxon	discussion	Remarks. The identity of the E. puncticulata illustrated by colour photographs in the books of Coleman (2007: plate 2 figure d) and Rauschert & Arntz (2015: plate 55) is not clear. The photograph examined are not sharp enough to reveal all relevant details. The species is similar to E. iota sp. nov., E. teres sp. nov., E. puncticulata and E. urvillei sp. nov. Unlike E. iota sp. nov., E. teres sp. nov. and E. urvillei sp. nov., E. (Subepimeria) sp. 2 exhibits a distinct brown stripe on each side of pereionite 4 and on coxa 4. The colour pattern of E. puncticulata s. str. is unknown. The collection details of the specimen of Rauschert & Arntz (2015) is given in an early draft of their book made available to the authors: ANT-XVII / 3 stn 138 [138 - 1]. The coordinates of this station are: 71 ° 08.90 ʹ S, 013 ° 12.80 ʹ W to 71 ° 08.80 ʹ S, 013 ° 13.20 ʹ W [eastern shelf of the Weddell Sea], 765 – 840 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1D68E3FDBDFEE3CA61FDAA.taxon	etymology	Etymology Combination of οὐρά, tail and Epimeria. The name, which is feminine, refers to the fact that the only ornamentation of the species is located on the posterior part of the animal.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1D68E3FDBDFEE3CA61FDAA.taxon	type_taxon	Type species Epimeria annabellae Coleman, 1994.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1D68E3FDBDFEE3CA61FDAA.taxon	description	Description Body fairly opaque with teguments moderately calcified. Rostrum small to medium-sized: 0.5 to 1.1 × as long as article 1 of peduncle of antenna 1. Eyes not conical. Pereionites 1 – 7 and pleonites 1 – 2 completely smooth; pleonite 3 posterodorsally produced into a triangular tooth or a blunt lobe directed backwards, without other ornamentation. Coxae 1 – 3 with tip blunt or sharp, keeled or not. Coxa 4 without groove or carina, with posteroventral border distinctly concave. Coxae 5 – 6 without tooth or distinct protrusion. Mid of posterior border of epimeral plates 1 – 3 not produced into a tooth. Posteroventral tooth of epimeral plate 3 small to strong. Dorsal process of urosomite 1 produced into a triangular tooth or a rounded lobe directed upwards. Urosomite 2 without pair of small teeth pointing upwards. Lateral borders of urosomite 3 posteriorly bluntly angular. Peduncle of antenna 1 without teeth or with dentition vestigial. Mandible with molar process triturative. Lower lip with narrow (V-shaped) hypopharyngeal gap. Palp of maxilliped with 4 articles. Gnathopods of normal size, with carpus and propodus fairly stout, with palm obliquely transverse (gnathopods subcheliform); propodus not expanded distally or slightly expanded distally; posterior border of dactylus lined by row of small oblique slender teeth. Basis of pereiopods 5 – 6 broad and quadrato-elliptic, without posteroproximal protrusion, without posterodistal tooth. Posterior border of basis of pereiopod 7 either completely rounded or concave on distal 0.25 and terminated into a tooth. Dactylus of pereiopods 5 – 7 short. Benthic. Body length The maximum body length recorded in Urepimeria species ranges between 11 and 29 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1D68E3FDBDFEE3CA61FDAA.taxon	biology_ecology	Ecology Benthic, 0 – 459 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1D68E3FDBDFEE3CA61FDAA.taxon	distribution	Distribution South Orkney Islands, South Shetland Islands, Palmer Archipelago, Antarctic Peninsula, eastern Weddell Sea.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1D68E3FDBDFEE3CA61FDAA.taxon	discussion	Remarks Epimeria annabellae, E. extensa, and E. monodon together constitute the subgenus Urepimeria, based on their morphological similarity.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1A68E2FE69FDA3CCD3FE46.taxon	description	Figs 320 – 321	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1A68E2FE69FDA3CCD3FE46.taxon	materials_examined	Material examined RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 2, MG 29, eastern Weddell Sea, 71 ° 18.60 ʹ S, 12 ° 25.40 ʹ W, 181 m, 22 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132701); 1 spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 2 AGT 2, eastern Weddell Sea, 71 ° 18.70 ʹ S, 12 ° 17.10 ʹ W, to 71 ° 18.45 ʹ S, 12 ° 16.30 ʹ W, 170 – 174 m, Agassiz trawl, 9 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132715); 10 specs, cruise PS 39, ANT-XIII / 3, EASIZ I, stn 6, AGT 1, eastern Weddell Sea, 71 ° 31.80 ʹ S, 13 ° 34.50 ʹ W to 71 ° 31.86 ʹ S, 13 ° 35.50 ʹ W, 254 – 261 m, Agassiz trawl, 8 Jan. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132679); 2 specs, cruise PS 39, ANT-XIII / 3, EASIZ I, stn 6, AGT 1, eastern Weddell Sea, 71 ° 31.80 ʹ S, 13 ° 34.50 ʹ W to 71 ° 31.86 ʹ S, 13 ° 35.50 ʹ W, 254 – 261 m, Agassiz trawl, 8 Jan. 1996, coll. C. De Broyer and G. Chapelle (MNHN-IU- 2014 - 7341, removed from RBINS, INV. 132679); 1 spec., cruise PS 39, ANT-XIII / 3, EASIZ I, stn 8, AGT 2, eastern Weddell Sea, 71 ° 18.70 ʹ S, 12 ° 17.10 ʹ W to 71 ° 18.45 ʹ S, 12 ° 16.30 ʹ W, 170 – 174 m, 9 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132999); 5 specs, cruise PS 39, ANT-XIII / 3, EASIZ I, stn 8, AGT 2, eastern Weddell Sea, 71 ° 18.70 ʹ S, 12 ° 17.10 ʹ W to 71 ° 18.45 ʹ S 12 ° 16.30 ʹ W, 170 – 174 m, 9 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132713); 2 large specs, cruise PS 39, ANT-XIII / 3, EASIZ I, stn 12, GSN 5, eastern Weddell Sea, 73 ° 18.10 ʹ S, 21 ° 10.10 ʹ W, to 73 ° 17.10 ʹ S, 21 ° 08.20 ʹ W, 457 – 459 m, bottom trawl, 14 Feb. 1996, coll. C. De Broyer and G. Chapelle (RBINS, INV. 132704); 1 spec., cruise PS 69, ANT-XXIII / 8, stn 603 - 5, eastern Weddell Sea, 70 ° 30.99 ʹ S, 08 ° 48.08 ʹ W to 70 ° 30.40 ʹ S, 08 ° 48.13 ʹ W, 274 – 297 m, sponge bottom, Agassiz trawl, 7 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122476) [extraction N 2; Genbank nr, COI: KU 870890, 28 S: KU 759678]; 1 spec., cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 260 - 6, eastern Weddell Sea, BENDEX area, 70 ° 50.56 ʹ S, 10 ° 36.20 ʹ W to 70 ° 50.71 ʹ S, 10 ° 36.56 ʹ W, 251 – 253 m, Agassiz trawl, 20 Mar. 2011, coll. Ch. Havermans and H. Robert (RBINS, INV. 132952) [extraction I 1; Genbank nr, COI: KU 870837, 28 S: KU 759611] [the station number given by Verheye et al. (2016 a) is incorrect]; 1 spec., cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 275 - 3, eastern Weddell Sea, 70 ° 56.01 ʹ S, 10 ° 29.28 ʹ W to 70 ° 56.01 ʹ S, 10 ° 28.72 ʹ W, 219 – 226 m, coll. Ch. Havermans and H. Robert (RBINS, INV. 122898).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1A68E2FE69FDA3CCD3FE46.taxon	description	Colour pattern Whitish, often speckled with scattered orange dots; peduncle of antenna 1 often orange; eyes reddish. Body length Up to 25 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1A68E2FE69FDA3CCD3FE46.taxon	distribution	Distribution Eastern shelf of Weddell Sea, 159 – 459 m (De Broyer et al. 2007).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1B68E2FE49FE13CC0BFC3E.taxon	description	Body length 11 mm. The species is known by a single specimen and it is possible that it can reach a larger size.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1B68E2FE49FE13CC0BFC3E.taxon	distribution	Distribution Southeast of tip of Antarctic Peninsula, 230 – 260 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1B68E1FE10FC0ACD22FDA3.taxon	materials_examined	Material examined SOUTHERN OCEAN: 1 exuvia, King George Island, Admiralty Bay, kept in aquarium, 14 Apr. 1992, coll. C. De Broyer (RBINS, INV. 132831); 1 exuvia, expedition ARCTOWSKI 1992, King George Island, Admiralty Bay, kept in aquarium, 1 Apr. 1992, coll. C. De Broyer (RBINS, INV. 132832); 1 exuvia, expedition ARCTOWSKI 1992, King George Island, Admiralty Bay, kept in aquarium, 16 Apr. 1992, coll. C. De Broyer (RBINS, INV. 132833); 1 exuvia, expedition ARCTOWSKI 1992, King George Island, Admiralty Bay, kept in aquarium, 30 Mar. 1992, coll. C. De Broyer (RBINS, INV. 132834); 2 specs, expedition ARCTOWSKI 1992, King George Island, Admiralty Bay, kept in aquarium, year 1992, coll. C. De Broyer (RBINS, INV. 132835).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1B68E1FE10FC0ACD22FDA3.taxon	description	Body length Up to 29 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1B68E1FE10FC0ACD22FDA3.taxon	distribution	Distribution South Orkney Islands; South Shetland Islands; Palmer Archipelago; 0 – 15 m (De Broyer et al. 2007). Epimeria monodon was recorded from rocky intertidal communities (Aghmich et al. 2016) and was found amidst macro-algae (Richardson 1977). Thurston (1972) stated: “ apparently associated with algae growing on rocks, boulder or gravel bottoms, 3 – 15 m ”.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1B68E1FE10FC0ACD22FDA3.taxon	discussion	Remarks The record of E. monodon from the eastern shelf of the Weddell Sea at 254 m listed in the data compilation of De Broyer et al. (2007) falls outside of the usual geographical and bathymetric range of the species. It is considered as very suspect. It might be based on E. annabellae, which is common at such depth in the eastern Weddell Sea. It seems possible that E. monodon, which is a very shallow-water species, uses algal rafting as a method of dispersal between islands and archipelagoes. Thurston (1972) recorded a slight sexual dimorphism in the species: “ the male has larger eyes and a slightly more prominent projection on pleon segment 3 ”.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1868E1FDF9FD3DCEADF9C6.taxon	distribution	Distribution Boreal North Pacific, Antarctica (?), 54 – 2550 m, possibly below 3000 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1868E1FDF9FD3DCEADF9C6.taxon	discussion	Remarks Uschakoviella echinophora has been recorded only once in the Southern Ocean: north of the South Orkney Islands, 59 ° 48 ʹ S, 45 ° 06 ʹ W (Watling & Holman 1981). These authors do not give the depth of their record. The Google Earth depth for that position is 3553 m [Google Earth accessed on 27.09.2016], suggesting a depth between 3000 and 4000 m. The specimen, which was a juvenile, is lost (Coleman 2007). The real identity of the Antarctic specimen is questionable (De Broyer et al. 2007). Klages (1991: 51) also listed a Uschakoviella sp. from the eastern Weddell Sea, without giving precise information on his material. It is usually assumed that Uschakoviella echinophora echinophora Gurjanova, 1955 and U. e. abyssalis Gurjanova, 1955 are the same species. However, examination of Gurjanova’s (1955) drawings reveals slight morphological differences in the basis of pereiopod 7 and there might therefore be two taxa involved.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1868E1FDC2FDBACA25FD26.taxon	type_taxon	type species: Uschakoviella echinophora Gurjanova, 1955.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1868E0FE57F99FCA60FB7B.taxon	discussion	In earlier literature, only Acanthonotozomella Schellenberg, 1926, Acanthonotozomoides Schellenberg, 1931 and the extralimital Amatiguakius Coleman & J. L. Barnard, 1991 were included in the Acanthonotozomellidae (Coleman 2007). Actinacanthus Stebbing, 1888, which is traditionally included in the Epimeriidae (e. g., Coleman & J. L. Barnard 1991 a; Coleman 2007; De Broyer et al. 2007) is herein transferred to the Acanthonotozomellidae. The entire telson and the large ventrolateral processes on the lower part of the pereionites are characters, which are typical of the Acanthonotozomellidae.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF1968DFFE08FAF2CA61FEF8.taxon	type_taxon	type species: Acanthonotozomella alata Schellenberg, 1926.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2668DFFE3BFCEBCB1DFBCB.taxon	distribution	Distribution Off extreme southern Patagonia, 384 – 494 m (Watling & Holman 1980).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2668DFFE3AFB83CD68F84B.taxon	distribution	Distribution Drake Passage, North of King George Island, 61 ° 26.8 ʹ S, 58 ° 06.2 ʹ W to 61 ° 27.1 ʹ S, 58 ° 05.2 ʹ W, 1047 – 1227 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2668DFFE3AFB83CD68F84B.taxon	discussion	Remarks Coleman & Jäger (2001) indicate that the locus typicus of Acanthonotozomella rauscherti is: RV Polarstern, cruise PS 48, ANT-XV / 3, station 336, Drake Passage, 62 ° 21.9528 ʹ S, 58 ° 41.1854 ʹ W, depth 1000 m, 19.03.1998. The coordinates of station 336 are actually 61 ° 26.8 ʹ S, 58 ° 06.2 ʹ W (gear on ground) to 61 ° 27.1 ʹ S, 58 ° 05.2 ʹ W (start of heaving), 1047 – 1227 m, 19.03.1998 (Arntz & Gutt 1999). Rauschert & Arntz (2015) give a photograph of an Acanthonotozomella cf. rauscherti, which almost certainly really belongs to the present species. The apparent slight differences might be size-related, as the specimen photographed is a juvenile. Rauschert & Arntz (2015) give very little information on their specimen. However, in the photographic archives of RBINS, an annotated version of the same photograph is present. The specimen is said to be 5 mm long and was collected at station 232 of ANT-XXI / 2. The coordinates of this station are: 71 ° 18.61 ʹ S, 13 ° 56.12 ʹ W to 71 ° 18.73 ʹ S, 13 ° 56.57 ʹ W (eastern shelf of the Weddell Sea), 900 – 910 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2668DFFDF0FE73CD9EFD53.taxon	distribution	Distribution Davis Sea, 385 m (Schellenberg 1926).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2768DEFE20FEE3CDF4FBF0.taxon	materials_examined	Material examined RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 248 - 2, Larsen B, 65 ° 57.51 ʹ S, 60 ° 28.15 ʹ W to 65 ° 57.69 ʹ S, 60 ° 28.30 ʹ W, 196 – 202 m, Agassiz trawl, 7 Mar. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 132673) [extraction E 16 of 23 Aug. 2012; Genbank nr, 28 S: KT 808684, 18 S: KT 808770].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2768DEFE20FEE3CDF4FBF0.taxon	distribution	Distribution Adélie Coast: Cape Géodésie; Scotia Sea; eastern shelf of Weddell Sea; 120 – 130 m (De Broyer et al. 2007); Larsen B, 196 – 202 m (present material).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2768DEFE20FEE3CDF4FBF0.taxon	discussion	Remarks Rauschert & Arntz (2015: 59, pl. 52) give a photograph of an Acanthonotozomella cf. trispinosa, which might be a species distinct from the true A. trispinosa. In the archives of RBINS, there is an annotated copy of this photograph. It indicates that the specimen is 6 mm long and was collected at the station 293 of ANT-XXI / 2. The coordinates of stn 293 - 1 are 72 ° 51.90 ʹ S, 19 ° 39.31 ʹ W to 72 ° 48.65 ʹ S, 19 ° 39.62 ʹ W (eastern shelf of the Weddell Sea), 518 – 542 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2768DEFE01FB4ACA60F975.taxon	type_taxon	type species: Acanthonotozomoides sublittoralis Schellenberg, 1931.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2768DDFE1BF8CDCB41FB4D.taxon	description	Fig. 322	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2768DDFE1BF8CDCB41FB4D.taxon	materials_examined	Material examined RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., cruise PS 69, ANT-XXIII / 8, stn 721 - 2, south-east of Larsen B, 65 ° 55.41 ʹ S, 60 ° 34.01 ʹ W to 65 ° 55.79 ʹ S, 60 ° 33.96 ʹ W, 295 – 299 m, hard bottom with hydrocorals (Stylasteridae), Agassiz trawl, 20 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122307); 1 spec., cruise PS 77, ANT-XXVII- 3, CAMBIO, stn 211 - 6, Shag Rocks, 53 ° 23.94 ʹ S, 42 ° 40.10 ʹ W to 53 ° 23.86 ʹ S, 42 ° 40.11 ʹ W, 285 m, Agassiz Trawl, 14 Feb. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 132669) [extraction EB 12 of 27 Feb. 2012; Genbank nr, 28 S: KT 808686, 18 S: KT 808782]; 2 specs, cruise PS 81, ANT-XXIX / 3, stn 197 - 6, Bransfield Strait, 62 ° 45.05 ʹ S, 57 ° 26.68 ʹ W to 62 ° 45.09 ʹ S, 57 ° 26.47 ʹ W, 210 – 222 m, black gravel mixed with sand and a little bit of mud, Rauschert dredge, 25 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122868). RV Aurora Australis cruises: SOUTHERN OCEAN: 1 spec., cruise CEAMARC, sample CEAMARC 933, stn 40 EV 152, Adélie Coast, 66 ° 39 ʹ 38 ″ S, 143 ° 01 ʹ 15.78 ″ E, 471 – 637 m, beam trawl, 28 Dec. 2007, coll. IPEV-AAD-MNHN (MNHN-IU- 2014 - 7342, removed from the sample MNHN-IU- 2014 - 4283).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2768DDFE1BF8CDCB41FB4D.taxon	distribution	Distribution Palmer Archipelago, South Georgia, South Shetland Islands, eastern shelf of Weddell Sea, Davis Sea, Adélie Coast, Ross Sea, (De Broyer et al. 2007), Shag Rocks, Bransfield Strait, Larsen B (present material), between 68 m (De Broyer et al. 2007) and 637 m (present material).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2468DDFE2FFB04CDC3F9AC.taxon	distribution	Distribution Falkland Islands, 197 m (Schellenberg 1931).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2468DDFDCBF9A7CBA2F862.taxon	type_taxon	type species: Acanthozone tricarinata Stebbing, 1883.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2568DCFDE6FEE3CA60FCE9.taxon	distribution	Distribution Heard Island, 274 m (Stebbing 1888).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2568DCFDE6FEE3CA60FCE9.taxon	discussion	Remarks Actinacanthus tricarinatus has been recorded only once since its original description, by Hibbert & Moore (2009, as ‘ Amphipoda spl (Cru 74) ’) in the same area as the type material, the archipelago of the Heard and McDonald Islands. The species is currently considered as endemic to the Kerguelen Plateau.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2568DCFD98FAF6CC4BFA56.taxon	type_taxon	type species: Dikwa acrania Griffiths, 1974.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2568DBFDC1F9EECEF6FD8E.taxon	description	Fig. 323	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2568DBFDC1F9EECEF6FD8E.taxon	materials_examined	Material examined RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 208 - 2, Burdwood Bank, 54 ° 32.27 ʹ S, 56 ° 9.78 ʹ W to 54 ° 30.84 ʹ S, 56 ° 9.06 ʹ W, 282 – 283 m, bottom trawl, 11 Feb. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 132666) [extraction EB 11; Genbank nr, 28 S: KT 808704, 18 S: KT 808771] [the lat. / long. coordinates given by Verheye et al. (2016 b) are incorrect].	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2568DBFDC1F9EECEF6FD8E.taxon	distribution	Distribution Burdwood Bank (Lörz & Coleman 2003; present material), 272 – 290 m (Lörz & Coleman 2003).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2568DBFDC1F9EECEF6FD8E.taxon	biology_ecology	Ecology Dikwa andresi was caught clinging to the stylasterid hydrocoral Errinopsis reticulum Broch, 1951 (Lörz & Coleman 2003).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2268DBFDBBFDD9CC46F862.taxon	discussion	Remarks Holman & Watling (1983) proposed a classification scheme where the Stilipedidae consisted of an assemblage of three well distinct homogeneous subfamilies: Alexandrellinae Holman & Watling, 1983, Astyrinae Pirlot, 1934 and Stilipedinae Holmes, 1908. The monophyly of this assemblage was questioned by Andres & Lott (1986) and was not supported by a recent phylogenetic analysis including two of these subfamilies: the Alexandrellinae and the Astyrinae (Verheye et al. 2016 b, 2017). The classification of Holman & Watling (1983) is retained herein for convenience only. In the past, members of the family Stilipedidae have been attributed to a wide range of families. For example Schellenberg (1931) included Alexandrella schellenbergi (as Iphimediopsis australis) in the Acanthonotozomatidae; K. H. Barnard (1932) put Eclysis similis in the Paramphithoidae, and Nicholls (1938) put Alexandrella mixta (as Parandaniexis mixtus) in the Stegocephalidae.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2368DAFE29FEE3CB4BFC5A.taxon	discussion	Remarks According to the phylogenetic analyses of Verheye et al. (2016 b, 2017), the Alexandrellinae are extremely close to the Epimeriidae, and might even be nested within Epimeria.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2368D8FDB5FC13CA60F888.taxon	type_taxon	(Alexandrella dentata designated as type species, by monotypy).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2368D8FDB5FC13CA60F888.taxon	etymology	Etymology The name Alexandrella presumably refers to Alexander Island, where the type species of the genus was collected.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2368D8FDB5FC13CA60F888.taxon	discussion	Remarks We decided to include the description of a new Alexandrella species herein because this genus belong to the same large clade as Epimeria (Verheye et al. 2016 b, 2017) and because the new species is large, very characteristic and was already mentioned and illustrated in the cruise report of ANT-XXIX / 3 (d’Udekem d’Acoz & Verheye 2013). Moreover, we felt that a quick overview of Alexandrella taxonomy would be a good test for checking whether the underestimated biodiversity found in Epimeria was unique for large Antarctic amphipods or representative of a general situation. This led us to draw a dire conclusion, that the taxonomy of Antarctic Alexandrella is even messier than for Epimeria before our present revision, and a significant number of undescribed species obviously occur in the Southern Ocean. The uniform morphology of Alexandrella species simply contributed to mask its true taxonomic diversity and to promote the convenient but spurious concept of ‘ variable widespread species’. The morphological differences between the genera Alexandrella and Bathypanoploea as proposed by Holman & Watling (1983) are ill-defined (Berge & Vader 2005 b, 2005 c) and genetic data (Verheye et al. 2016 b) confirm that Bathypanoploea pulchra (as A. schellenbergi) and Alexandrella cf. mixta (as A. aff. dentata) are very close relatives. The two genera are therefore considered herein as synonyms. Besides the species from the temperate and polar parts of the southern hemisphere treated herein, Alexandrella includes an Atlantic abyssal species: A. setosa Serejo, 2014 (see Serejo 2014) and two abyssal species from the North Pacific: Alexandrella carinata (Birstein & Vinogradova, 1960) and a second, undescribed species examined by the first author. The species described and illustrated as Astyroides carinatus by Birstein & Vinogradova (1960) exhibits almost all the characters of Alexandrella. J. L. Barnard (1969) considered Astyroides as a junior synonym of Alexandrella, and it is only reluctantly that Holman & Watling (1983) accepted Astyroides as valid. In the reprint of Birstein & Vinogradova (1960) available to us, there is a handwritten comment by Vinogradova relegating Astyroides into the synonymy of Alexandrella. This synonymization is formally adopted herein.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2368D8FDB5FC13CA60F888.taxon	description	This key to Antarctic and sub-Antarctic Alexandrella is tentative and has not the pretention to be a rigorous identification tool. Characters visible without dissection have been used whenever possible.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2168D7FDC1F8C0CA73FC9B.taxon	distribution	Distribution Eastern Weddell Sea: Scottish National Antarctic Expedition 1902 – 1904, RV Scotia, stn 417, 71 ° 22 ʹ S, 16 ° 34 ʹ W, 2578 m (Chilton 1912).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2168D7FDC1F8C0CA73FC9B.taxon	discussion	Remarks There are significant morphological differences between the Alexandrella australis specimens illustrated by Chilton (1912) and Holman & Watling (1983), as follows: ventral margin of coxa 1 concave vs convex; pereionite 6 with vs without small mid-dorsal tooth; crest of urosomite 1 with long vs small posterior tooth. It is probable that they are not conspecific and that the specimen illustrated by Holman & Watling (1983) belongs to an undescribed species. It is referred herein as Alexandrella sp. 3. The second (nonillustrated) specimen listed by these authors might be A. chione sp. nov. (see account on that species).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2E68D5FD99FCD2CFE1F81F.taxon	description	urn: lsid: zoobank. org: act: 90 F 1 FE 7 A-F 36 C- 4411 - A 7 A 8 - 833 B 300 A 0862 Figs 324 – 335	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2E68D5FD99FCD2CFE1F81F.taxon	etymology	Etymology From the Greek, χιων, snow. The name, which is a noun in apposition, alludes to the pure white colour of the species.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2E68D5FD99FCD2CFE1F81F.taxon	materials_examined	Type material Holotype RV Polarstern cruises SOUTHERN OCEAN: immature ♀, cruise PS 81, ANT-XXIX / 3, stn 227 - 2, Bransfield Strait, 62 ° 55.83 ʹ S, 58 ° 41.09 ʹ W to 62 ° 55.76 ʹ S, 58 ° 41.46 ʹ W, 562 – 564 m, muddy bottom, Agassiz Trawl, 5 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122887).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2E68D5FD99FCD2CFE1F81F.taxon	description	Description HEAD. Rostrum tiny, extremely narrow in frontal view, reaching 0.3 of article 1 of peduncle of antenna 1 (dorsomedial tooth included); lateral head lobes triangular (forming a sharp squared angle); ventral lobe forming an extremely strong hemi-elliptic protrusion; connection between this protrusion and head forming a deep notch (shape: sharp squared angle). EYES. Circular, small, pale, non-ommatidian, disappearing in alcohol. PEREION – PLEOSOME TOOTH PATTERN. Pereionites 1 – 6 totally smooth; pereionite 7 posteriorly carinate and with medium-sized tooth projecting backwards; pleonites 1 – 2 carinate, with medium-sized tooth projecting backwards; pleonite 3 carinate, with distinctly sigmoid profile (distinctly convex in anterior 0.8, distinctly concave in posterior 0.2), with medium-sized tooth projecting backwards. EPIMERAL PLATES 1 – 3. Posteroventral angle: very obtusely rounded in plate 1, produced into a mediumsized tooth in plates 2 – 3. UROSOME ORNAMENTATION. Urosomite 1 with deep notch on proximal third, followed by elevated carina, anteriorly rounded and slightly produced forward, dorsally nearly straight (posteriorly very slightly concave), posteriorly produced into a well-developed, narrowly triangular sharp tooth overreaching urosomite 1 and forming a blunt squared angle with it (tooth 0.23 × as long as carina); urosomite 2 toothless; urosomite 3 lateral borders distally produced into a well-developed lobe, of which the tip forms a squared angle. TELSON. Broadly rectangular, distally broad and emarginate; with very shallow broad V-shaped notch (cleft on 0.08); lobes very broad, medially forming a rounded very low projection, more laterally with shallow concavity, at their connection with lateral border forming a broad and blunt triangular tooth. ANTENNA 1. Articles of peduncle very broad; article one with large and broad dorsomedial tooth reaching tip of article two, with strong ventral tooth reaching mid of article two; article two with broadly rounded lateral lobe; article three with broadly rounded lateral lobe; accessory flagellum distinct, uniarticulate. ANTENNA 2. Article four with posterodistal blunt dentiform process; article five 1.3 × as long as article four. UPPER LIP. Strongly asymmetrical, right lobe well developed, bluntly triangular and very protruding, left lobe low and very rounded. MANDIBLES. Incisor very broad, straight and toothless (or with trace of tiny denticles near lateral corner); left lacinia mobilis very large (0.7 × as broad as incisor process); right lacinia mobilis small (0.25 × as broad as incisor process); molar absent; palp with article 1 very short, posterodistally produced into a tooth; article 2 and 3 long and narrow, posteriorly ciliate; article 2 slightly longer than article 3. LOWER LIP. Lower lip without inner lobes, outer lobes broad with small acute projection at outer corners. MAXILLA 1. Inner plate setose along medial margin and along a row on inner face, notched at the level of this row to allow imbrication of outer plate; palp biarticulate, expanded distally with 1 marginal row of spines + row of short setae. MAXILLA 2. Inner plate distinctly larger than outer, rounded, posterior setae conspicuously longer than those more anterior but not strongly plumose, also sparse setae on surface of plate; outer plate triangular. MAXILLIPED. Outer plate exceeding palp, setose medially; inner plate extending beyond distal margin of palp article 1, setae on distal margin of moderate length; palp 4 - articulate, article 2 longest, article 4 less than 0.5 length of article 3. GILLS. From gnathopod 2 to pereiopod 7. OOSTEGITES. From gnathopod 2 to pereiopod 5. GNATHOPOD 1. Coxa very broad, expanding distally, anteroventral corner very broadly rounded, ventral margin straight; leg achelate; carpus very broad, 1.8 × as long as propodus; propodus strongly tapering. GNATHOPOD 2. Coxa broad, anteriorly rounded, posterodistally angulate; leg achelate; carpus very broad, 1.5 × as long as propodus; propodus weakly tapering. PEREIOPOD 3. Coxa broad, with anterior border rounded, distally produced into a large blunt tooth, posterior margin very slightly concave; leg robust: propodus 4.8 × as long as wide; dactylus short: 3.6 × as long as broad, 0.4 × as long as propodus, with row of small spines on posterior border. PEREIOPOD 4. Coxa anteriorly forming a strong regular curve, with ventral corner angulate (forming a distinct acute angle), with posteroventral border strongly concave; leg robust very similar to that of pereiopod 3. PEREIOPOD 5. Coxa posteriorly produced into a triangular tooth (forming a squared angle), posteroventrally also produced into a triangular tooth (forming a blunt acute, nearly squared angle); basis fairly narrow, posterior border very weakly concave, posterodistal corner broadly rounded, posteromedial carina distally produced into a small rounded lobe; carpus, propodus and dactylus missing. PEREIOPOD 6. Coxa posteriorly forming a rounded obtuse angle, posteroventrally produced into a triangular tooth (forming a blunt acute angle); basis of medium width; posterior border very weakly concave, posterodistal corner forming a blunt squared angle (not rounded); carpus, propodus and dactylus missing. PEREIOPOD 7. Coxa posteriorly straight, posteroventrally forming an obtuse (nearly squared) angle; basis extremely broad, with posteroproximal strong rounded process, posterodistally produced into a sharp triangular tooth forming an acute angle projecting forward (that tooth is 0.17 × as long as basis, tooth included). Colour pattern White, except for mouthparts, gnathopods (red) and carpus, propodus and dactylus of pereiopods 3 – 4 (pale pink); eyes pale yellow. Body length 30 mm.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2E68D5FD99FCD2CFE1F81F.taxon	distribution	Distribution Bransfield Strait, 562 – 564 m (present data).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2E68D5FD99FCD2CFE1F81F.taxon	discussion	Remarks Alexandrella chione sp. nov. (Bransfield Strait shelf) is similar to Alexandrella australis (eastern Weddell Sea abyss) and Alexandrella sp. 3 (South Sandwich Islands abyss). The most visible difference concerns the ventral lobe of head, which is much stronger and proximally narrower in Alexandrella chione sp. nov. than in other species. In addition to the characters given in the key, it must be mentioned that the lacinia mobilis of the right mandible is more developed and the posterodistal tooth of the basis of pereiopod 7 is longer in Alexandrella chione sp. nov. than in Alexandrella sp. 3 as illustrated by Holman & Watling (1983). Very little information is given on the adult specimens of ‘ Alexandrella australis ’ recorded at 935 m in the Bransfield Strait by Holman & Watling (1983) and Berge & Vader (2005 a), but they may be A. chione sp. nov., which is described herein from a specimen collected at similar depths in the same area.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2D68D4FDCDFEE3CE91FAE0.taxon	distribution	Distribution Bellingshausen Sea: Alexander Island, 297 m (Chevreux 1913).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2D68D4FDCDFEE3CE91FAE0.taxon	biology_ecology	Biology The holotype of A. dentata was found inside a sponge (Chevreux 1913).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2D68D4FDCDFEE3CE91FAE0.taxon	discussion	Remarks Chevreux (1913) explicitly described and illustrated a tooth on the side of each segments of the pereion of the holotype of Alexandrella dentata, hence the name of the species. They were not reproduced, however, in the figures of J. L. Barnard (1969) and J. L. Barnard & Karaman (1991), which were copied from Chevreux (1913). Neither are they mentioned in the account of the holotype of A. dentata by Holman & Watling (1983), who only studied the appendages and not the body. Berge & Vader (2005 a) also did not mention these teeth on their specimens and put Alexandrella mixta, which is not supposed to have such teeth, in synonymy with A. dentata. The specimens of Berge & Vader (2005 a) are without lateral teeth, which indicates that the species is not A. dentata. The presence or absence of these lateral teeth is here considered as a species level character distinguishing A. mixta from A. dentata. A. mixta has been recorded in widely separate localities and across a wide bathymetric range. Moreover, small differences exist between the various descriptions and illustrations. Therefore, A. mixta might be a species complex.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2D68D4FE2DFA7BCE75F897.taxon	distribution	Distribution Prince Edward Islands, 46 ° 56.3 ʹ S, 37 ° 55.6 ʹ E, 120 m (Bellan-Santini & Ledoyer 1987); Prince Edward Islands, 46 ° 40 ʹ 32 ″ S, 37 ° 51 ʹ E, 460 – 560 m; off Prydz Bay, 67 ° 30 ʹ S, 77 ° 14 – 13 ʹ E, 341 – 333 m (Berge & Vader 2005 a).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2D68D3FE0BF8A0CA60FEDF.taxon	distribution	Distribution East of South Georgia, 55 ° 04 ʹ S, 33 ° 57 ʹ W to 55 ° 00 ʹ S, 33 ° 59 ʹ W, 3138 – 3239 m (Berge & Vader 2005 a).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2A68D3FDE9FE95CC14FD77.taxon	distribution	Distribution Middle of Scotia Sea, at the same longitude as South Georgia, 58 ° 53 ʹ S, 37 ° 15 ʹ W to 58 ° 54 ʹ S, 37 ° 19 ʹ W, 2901 m; Drake Passage, north of Livingstone Island, 61 ° 43.08 ʹ S, 60 ° 41.8 ʹ W, 2830 – 2862 m; Somov Sea: longitude of Cape Adare and latitude of (east of) Balleny Islands, 68 ° 05 ʹ S, 173 ° 38 ʹ E to 68 ° 06 ʹ S, 173 ° 44 ʹ E, 2608 – 3175 m (Berge & Vader 2005 a).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2A68D3FDDEFCCECCB8F977.taxon	description	Fig. 336	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2A68D3FDDEFCCECCB8F977.taxon	materials_examined	Material examined RV Polarstern cuises SOUTHERN OCEAN: 1 spec., cruise PS 69, ANT-XXIII / 8, stn 725 - 6, Larsen A, 64 ° 54.80 ʹ S, 60 ° 37.46 ʹ W to 64 ° 54.80 ʹ S, 60 ° 38.28 ʹ W, 105 – 207 m, amongst large grey hexactinellid sponges, Agassiz trawl, 22 Jan. 2007, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122305).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2A68D3FDDEFCCECCB8F977.taxon	distribution	Distribution Adélie Coast, Weddell Sea; 60 – 1543 m (De Broyer et al. 2007 as A. dentata, excluding type record).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2A68D3FDDEFCCECCB8F977.taxon	biology_ecology	Biology One specimen identified as Alexandrella mixta was found by the first author amongst large grey hexactinellid sponges and its colour pattern matched perfectly with that of these sponges (fig. 338). Some kind of symbiosis might be involved.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2A68D3FDDEFCCECCB8F977.taxon	discussion	Remarks See account on A. dentata for the taxonomic history of A. mixta.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2A68D2FE0AF8CECB10FCA4.taxon	distribution	Distribution Northwest of Elephant Island, 60 ° 54.60 ʹ S, 55 ° 45.90 ʹ W, 235 m (holotype); southwestern Atlantic, off Argentina (off the province of Río Negro), 40 ° 14 ʹ 06 ″ S, 55 ° 24 ʹ 07 ″ W, 1475 m (paratype) (Berge & Vader 2005 b, 2005 c).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2A68D2FE0AF8CECB10FCA4.taxon	discussion	Remarks The conspecificity between the holotype of Alexandrella polarsterni collected off Elephant Island at 235 m and the paratype collected off Argentina at 1475 m looks a priori suspect, based on the differences in location and depth, but cannot be refuted herein. The photograph of “ Alexandrella sp. nov. ” given by Rauschert & Arntz (2015) matches very well with the figures of Berge & Vader (2005 b, 2005 c), especially for the shape of the mid-dorsal carina of urosomite 1; hence this specimen is identified as such. Rauschert & Arntz (2015) do not indicate the coordinates of their specimen, but in an early draft of their book made available to the authors, the following information was given: ANT-XXI / 2, stn 145, 400 m, 7 mm. The coordinates of the station 145 - 1 (epibenthic sledge) are: 70 ° 56.99 ʹ S, 10 ° 48 ʹ 26 ″ W to 70 ° 56.97 ʹ S, 10 ° 47.7 ʹ W [eastern shelf of the Weddell Sea], 402 – 405 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2B68D1FE03FCBCCE5CFB8F.taxon	description	Fig. 337	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2B68D1FE03FCBCCE5CFB8F.taxon	materials_examined	Material examined RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., orange form, cruise PS 69, ANT-XXIII / 8, stn 654 - 6, Elephant Island, 61 ° 22.80 ʹ S, 56 ° 03.84 ʹ W to 61 ° 23.35 ʹ S, 56 ° 04.89 ʹ W, 341 – 342 m, Agassiz trawl, 29 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122440); 1 spec., orange form, cruise PS 69, ANT-XXIII / 8, stn 662 - 1, between Elephant Island and King George Island, 61 ° 35.91 ʹ S, 57 ° 17.04 ʹ W to 61 ° 35.41 ʹ S, 57 ° 20.60 ʹ W, 425 – 432 m, bottom trawl, 30 Dec. 2006, coll. C. d’Udekem d’Acoz and H. Robert (RBINS, INV. 122304); 1 spec., colour unrecorded, cruise PS 71, ANT-XXIV / 2, ANDEEP-SYSTCO, stn 48 - 1, east of Weddell Sea, 70 ° 23.94 ʹ S, 8 ° 19.14 ʹ W to 70 ° 23.89 ʹ S, 8 ° 18.67 ʹ W, 595 – 602 m, bryozoan bottom (exceptional bryozoan diversity), Rauschert dredge, 12 Jan. 2008, coll. H. Robert (RBINS, INV. 132623) [extraction EAS of 10 Feb. 2012; Genbank nr, 28 S: KT 808699, 18 S: KT 808768]; 1 spec., orange and white form, cruise PS 81, ANT-XXIX / 3, stn 217 - 6, Bransfield Strait, 62 ° 53.45 ʹ S, 58 ° 13.06 ʹ W to 62 ° 53.42 ʹ S, 58 ° 13.41 ʹ W, 421 - 483 m, rich sponge bottom, Agassiz trawl, 2 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122865); 1 spec., orange and white form, cruise PS 81, ANT-XXIX / 3, stn 227 - 2, Bransfield Strait, 62 ° 55.83 ʹ S, 58 ° 41.09 ʹ W to 62 ° 55.76 ʹ S, 58 ° 41.46 ʹ W, 562 – 564 m, muddy bottom, Agassiz Trawl, 5 Mar. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122864).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2B68D1FE03FCBCCE5CFB8F.taxon	description	Colour pattern and eyes description Head whitish; eyes very large, non-ommatidian and white, forming a large V pointing forwards; body entirely orange (orange form) or orange except for mid-dorsal crests of pleon, which are largely white (orange and white form); pereiopods and antennae orange (orange form) or largely white (orange and white form). The eyes disappear in alcohol.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2B68D1FE03FCBCCE5CFB8F.taxon	distribution	Distribution Reliable records are from the South Shetland Islands, Bransfield Strait and the eastern shelf of the Weddell Sea, 341 – 602 m.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2B68D1FE03FCBCCE5CFB8F.taxon	biology_ecology	Biology The analyses of gut contents by Coleman (1990 b) revealed the occurrence of ossicles of holothurians in the digestive tract of Alexandrella pulchra (as Bathypanoploea schellenbergi). Gut content analyses of the latter species by Dauby et al. (2001 a) revealed gorgonian remains (cnidocysts and ossicles), plankton and hard remains of crustaceans, bryozoans and holothurioids.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2B68D1FE03FCBCCE5CFB8F.taxon	discussion	Remarks The reasons for re-establishing Alexandrella pulchra as a distinct species are given in the remarks section on A. schellenbergi. In A. pulchra, some specimens have orange pereiopods and antennae and others nearly entirely white appendages. Two explanations are possible: Either the specimens exhibit individual colour variation or two very similar species are involved. The identity of the Alexandrella sp. of Rauschert & Arntz (2015) is not clear but it might be A. pulchra. The shape of the eye and the number of body segments with a tooth match with A. pulchra but coxa 4 seems to be apically rounded instead of subacute.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2868D0FE22FBC6CEFDFE6E.taxon	distribution	Distribution Sub-Antarctic Region: south of Argentine Basin: north east of Falkland Islands: 50 ° 19 ʹ S, 50 ° 50 ʹ W, 2675 m (Holman & Watling 1983).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2868D0FE22FBC6CEFDFE6E.taxon	discussion	Remarks It has been reported that Alexandrella schellenbergi has a very wide geographical and bathymetric distribution across the Southern Ocean (Holman & Watling 1983, Berge & Vader 2005 b, 2005 c, De Broyer et al. 2007). This widespread distribution is questioned here. The holotype of A. schellenbergi was collected north of the Antarctic Polar Front and north of the line joining the Falkland Islands to South Georgia, at 2675 m. Its telson is more deeply cleft than in similar Alexandrella specimens from the continental shelf of the South Shetland Islands and the tip of the Antarctic Peninsula. This telson character state is consistent with the specimens illustrated in literature (Holman & Watling 1983, Ren & Huang 1991) and examined by us. On the poor quality photograph given by Schellenberg (1931), the posterior pereiopods also appear more slender than in the forms of the Antarctic continental shelf; however, this might be an illusion created by the possibly not flattened orientation of the legs on the picture. The telson difference, the geographical wide separation and the bathymetric differences suggest that two species are involved. The name Alexandrella pulchra Ren in Ren & Huang, 1991 is available for the form of the South Shetland Islands and the tip of the Antarctic Peninsula, and it is herein resurrected for them. All topotypical shelf specimens of Alexandrella pulchra examined by us were devoid of posterodorsal tooth on pereionite 6. However, specimens similar to A. schellenbergi with a tooth on pereionite 6 are present in the Ross Sea (Holman & Watling 1983: 49, fig. 10 a), off Adélie Coast (present material) and in the Atlantic sector of the Southern Ocean (Rauschert & Arntz 2015). They are treated herein as a separate taxon: Alexandrella sp. 1.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2968D0FE3CFDFACBA5FB0A.taxon	distribution	Distribution For clarity, it seems necessary to give the station details in full for all known specimens identified as Alexandrella subchelata in literature: holotype: RV Galathea, stn 554, 5 December 1951, 37 ° 28 ʹ S, 138 ° 55 ʹ E, Great Australian Bight, 1320 – 1340 m (J. L. Barnard 1961 as A. dentata, Holman & Watling 1983). — RV Eltanin, cruise 26, stn 4, 41 ° 56 ʹ S, 160 ° 07 ʹ E [between Tasmania and New Zealand], 4846 – 4929 m. — RV Eltanin, cruise 27, stn 1880, 49 ° 40 ʹ S, 178 ° 53 ʹ E [Antipodes Island], 103 m. — RV Polarstern, ANT-XIX / 3 - 4 (ANDEEP II), stn 133 - 3, 65 ° 20.15 ʹ S, 54 ° 14.35 ʹ E to 65 ° 20.06 ʹ S, 54 ° 14.51 ʹ E [Antarctica: Kong Håkon VII Sea], 1119 – 1122 m (Berge & Vader 2005 a).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2968D0FD5BFB45CA71F862.taxon	description	Figs 338 – 339	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2968D0FD5BFB45CA71F862.taxon	materials_examined	Material examined RV l’Astrolabe cruises: SOUTHERN OCEAN: 1 very large spec., cruise REVOLTA III, no station (Dumont d’Urville Sea), Collect _ ID: REVO _ 449, Adélie Coast, 66 ° 38 ʹ S, 140 ° 42 ʹ E to 66 ° 38 ʹ S, 140 ° 40 ʹ E, 718 – 729 m, mud, beam trawl, 20 Jan. 2012, CE- 000002100 (MNHN-IU- 2009 - 2540).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF2968D0FD5BFB45CA71F862.taxon	discussion	Remarks The specimen examined has a mid-dorsal tooth on pereionite 6 in addition to the usual teeth on pereionite 7 and pleonites 1 – 3. Otherwise, it looks very similar to Alexandrella pulchra. Only minor differences in the form of coxae 1 and 4 were observed. The shape of the eye is similar to that of A. pulchra, but the body and leg pigmentation is less pronounced. A small tooth is also present in the specimen from the continental shelf of the Ross Sea illustrated by Holman & Watling (1983) and a specimen from the eastern Weddell Sea illustrated by Rauschert & Arntz (2015). The latter specimen has much smaller eyes than A. pulchra, and according to an early draft of Rauschert & Arntz’s book, it was collected during the cruise ANT-XXI (obviously ANT-XXI / 2), which almost exclusively sampled on the continental shelf.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3668CFFD5BFEE3CBB7FD4C.taxon	description	Fig 339	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3668CFFD5BFEE3CBB7FD4C.taxon	distribution	Distribution RV Polarstern, PS 77, ANT-XXVII / 3, stn 211 (there were several sub-stations for stn 211), Shag Rocks. The species is known from a photograph taken by H. Robert and Ch. Havermans. The specimen was unavailable for study.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3668CFFD5BFEE3CBB7FD4C.taxon	discussion	Remark This species looks similar to A. pulchra and A. schellenbergi but displays some differences. Pereionites 5 and 6 have each a conspicuous posterodorsal tooth, which is absent in A. pulchra and A. schellenbergi. The pereiopods are striped, which is not the case or not so pronounced in A. pulchra.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3668CFFD5BFD05CBEFF982.taxon	distribution	Distribution This taxon has been recorded once at the following station: RV Islas Orcadas, cruise 575, Stn 54, 57 ° 39 ʹ S, 26 ° 00.4 ʹ W [South Sandwich Islands], 2380 – 2609 m (Holman & Watling 1983, Berge & Vader 2005 a).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3668CFFD5BFD05CBEFF982.taxon	discussion	Remarks The specimen identified as Alexandrella australis by Holman & Watling (1983) exhibits striking differences with the illustration of the holotype given by Chilton (1912). The carina of urosomite 1 presents a much stronger posterior tooth and deeper anterior notch on the drawing of Chilton (1912) than in that of Holman & Watling (1983). The specimen of Chilton (1912) has a small posterodorsal tooth on pereionite 6, which is missing in the one illustrated by Holman & Watling (1983). The specimen of Chilton (1912) is illustrated as having a more truncated coxa 1 than the one of Holman & Watling (1983). Finally, the posterodistal tooth of the basis of pereiopod 7 looks longer in the specimen of Chilton (1912) than in the one of Holman & Watling (1983). As the conspecificity of the two specimens is questionable, the name Alexandrella sp. 3 is used herein for the one studied by Holman & Watling (1983).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3668CFFD5BF9DCCBDAF88C.taxon	distribution	Distribution Northeast of Elephant Island, 60 ° 39.53 ʹ S, 53 ° 56.93 ʹ W, 2889 m (Berge & Vader 2005 c).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3768CEFDA8F93BCD91F820.taxon	distribution	Distribution Elephant Island, 120 m (Andres 1997).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3768CEFD8BFC51CD6DF921.taxon	type_taxon	type species: Astyra abyssi Boeck, 1871.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3768CEFD8BFC51CD6DF921.taxon	etymology	Etymology “ Astyra, a town of Mysia, Mela, Pliny: Strabo calls it a hamlet, near which is a grove of Diana, thence named Astyrena: this Astyra is near Adramytrium, and to be distinguished from another near Abydos, which had a gold mine, Strabo. This last was a town of Troas; in ruins in Strabo’s time ” (MacBean 1773). Stebbing (1888: 1668) suggested that the name Astyra derives from Αστυρα or Astura, which is the name of a river in Asturia (Spain).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3468CDFDB9FEE3CBB0FD1A.taxon	type_taxon	type species: Eclysis similis K. H. Barnard, 1932.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3468CDFDBAFD56CD3EFBA2.taxon	distribution	Distribution South Georgia and Weddell Sea: eastern shelf, 230 – 250 m (De Broyer et al. 2007); Ross Sea, 659 – 741 m (Berge & Vader 2005 b).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3468CDFD98FA9DCA61F919.taxon	type_taxon	type species: Stilipes distinctus Holmes, 1908.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3468CDFDB0F953CDFDF853.taxon	distribution	Distribution Macquarie Island, 956 – 959 m (Berge 2003 a, b).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3568CCFE23FD25CE3AFB02.taxon	type_taxon	type species: Acanthonotozomella pushkini Bushueva, 1978.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3568CCFD94FEE3CD44FD2C.taxon	type_taxon	type genus: Vicmusia Just, 1990.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3568CBFE07FB5BCC83FCE0.taxon	description	Fig 341	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3568CBFE07FB5BCC83FCE0.taxon	materials_examined	Material examined RV Polarstern cruises: SOUTHERN OCEAN: 1 spec., cruise PS 77, ANT-XXVII / 3, CAMBIO, stn 211 - 5, Shag Rocks, 53 ° 24.21 ʹ S, 42 ° 41.65 ʹ W to 53 ° 24.06 ʹ S, 42 ° 42.58 ʹ W, 327 – 345 m, Agassiz trawl, 14 Feb. 2011, coll. C. Havermans and H. Robert (RBINS, INV. 132656) [extraction E 30 of 23.08.2012; Genbank nr, 28 S: KT 808687, 18 S: KT 808783]; 3 specs, cruise PS 81, ANT-XXIX / 3, stn 197 - 6, Bransfield Strait, 62 ° 45.05 ʹ S, 57 ° 26.68 ʹ W to 62 ° 45.09 ʹ S, 57 ° 26.47 ʹ W, 210 – 222 m, black gravel mixed with sand and a little bit of mud, Rauschert dredge, 25 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 122871); 1 spec., cruise PS 81, ANT-XXIX / 3, stn 164 - 5, south of Dundee Island, 63 ° 36.84 ʹ S, 56 ° 10.28 ʹ W to 63 ° 36.72 ʹ S, 56 ° 10.46 ʹ W, 121 – 122 m, Rauschert dredge, 11 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, 122886); 1 spec., cruise PS 81, ANT-XXIX / 3, stn 188 - 5, south-east of Dundee Island, 63 ° 50.92 ʹ S, 55 ° 37.66 ʹ W to 63 ° 50.93 ʹ S, 55 ° 37.52 ʹ W, 402 – 407 m, Rauschert dredge, 20 Feb. 2013, coll. C. d’Udekem d’Acoz and M. Verheye (RBINS, INV. 132206). RV Seatruck, cruise REVOLTA II: SOUTHERN OCEAN: 2 specs, stn REVO _ 128, Collect _ ID: REVO _ 128, Adélie Coast, 66 ° 38.7 ʹ S, 139 ° 56.8 ʹ E, 91 – 93 m, 10 Jan. 2011, coll. N. Améziane, N. Bax, C. Gallut, A. C. Lautrédou and C. Robineau (MNHN-IU- 2016 - 6586). RV Seatruck, cruise REVOLTA III: SOUTHERN OCEAN: 1 large spec., stn REVO _ 032, Collect _ ID: REVO _ 509, Field _ ID: CE- 000002396, Adélie Coast, 66 ° 39 ʹ S, 140 ° 02 ʹ W to 66 ° 39 ʹ S, 140 ° 02 ʹ W, 72 – 100 m, beam trawl, 8 Feb. 2012, coll. G. Lecointre, A. Dettaï, J. Lanshere, C. Gallut and C. Ozouf, (MNHN-IU- 2016 - 6602).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3568CBFE07FB5BCC83FCE0.taxon	distribution	Distribution Davis Sea, Enderby Land, South Shetland Islands, 24 – 45 m (De Broyer et al. 2007), Shag Rocks, South of Dundee Island, Bransfield Strait, Adélie Coast, down to 402 – 407 m (present material).	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
4A5A879BFF3568CBFE07FB5BCC83FCE0.taxon	discussion	Remarks The orange dotted colour pattern of A. pushkini (Fig. 340) is reminiscent of that of many Antarctic gorgonians but so far there is no direct evidence of symbiosis.	en	d’Acoz, Cédric d’Udekem, Verheye, Marie L. (2017): Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea). European Journal of Taxonomy 359: 1-553, DOI: 10.5852/ejt.2017.359
