taxonID	type	description	language	source
4A268793623CFFCAFF0DFC0FFBB7FDE4.taxon	type_taxon	Type species. Acanthoglyptus picollus sp. nov., here designated.	en	Alekseev, Vitalii Igorevich, Vitali, Francesco (2020): Description of the first extinct member of the tribe Anaglyptini (Coleoptera: Cerambycidae) from European Tertiary. Zootaxa 4816 (1): 135-143, DOI: 10.11646/zootaxa.4816.1.11
4A268793623CFFCAFF0DFC0FFBB7FDE4.taxon	diagnosis	Differential diagnosis. The specimen under consideration is assigned to the tribe Anaglyptini on the basis of the following combination of morphological characters: 1) eyes finely facetted, 2) procoxae rounded, 3) procoxal cavities narrowly open, 4) elytra gibbose at base and without transverse raised ivory-like ridges, 5) antennomeres 3 – 5 distinctly spinose. The new genus could be associated to the tribe Tillomorphini, which nevertheless, shows closed procoxal cavities. Exceptionally, they are narrowly open in the, possibly polyphyletic, American genus Euderces LeConte, 1850, which, in turn, shows evident transverse raised ivory-like ridges (peculiar character of Tillomorphini), a cribrose elytral disc, divided eyes, antennomere III more than twice as long as IV, etc. Acanthoglyptus gen. nov. differs from the representatives of extant genera with spined antennomeres (Anaglyptus Mulsant, 1839, Oligoenoplus Chevrolat, 1863, Paraclytus Bates, 1884, and Cyrtophorus LeConte, 1850) in the stout elytra and the longer first metatarsomere. For these characters, it resembles the Chinese Yoshiakioclytus Niisato, 2007, which nevertheless, has unarmed antennomeres (Huang & Chen 2016). Considering the species included in Baltic amber, Tillomorphites Vitali, 2011 (Tillomorphini Lacordaire, 1868) is the only genus characterised by spined antennae. It differs from Acanthoglyptus gen. nov. for the not gibbose elytra, much longer and finer antennal spines on antennomeres (in contrast to three shortly spined antennomeres in the new genus), shorter and more robust legs, and elytral pattern formed by transverse reddish band (s). From Clytus (Xylotrechus) pici Piton, the newly described fossil genus can be distinguished by the smaller body size (9.0 mm compared with 15.0 mm in C. (Xylotrechus) pici), the elytra gibbose at base, the antennae longer and apparently spined, and absence of frontal carina.	en	Alekseev, Vitalii Igorevich, Vitali, Francesco (2020): Description of the first extinct member of the tribe Anaglyptini (Coleoptera: Cerambycidae) from European Tertiary. Zootaxa 4816 (1): 135-143, DOI: 10.11646/zootaxa.4816.1.11
4A268793623CFFCAFF0DFC0FFBB7FDE4.taxon	description	Description. General habitus small, relatively stout, convex dorsally. Head oblique, slightly elongated; frons with longitudinal furrow; antennal tubercles rather elevated and widely separated; eyes shallowly emarginated, finely facetted; last maxillary palpomere elongate, truncate at apex, as long as two previous palpomeres combined; antennae 11 - segmented, hardly as long as body in female; antennomeres III – V with long spine at inner side decreasing in length to antennomere III; pedicel and antennomeres III – V with a few semierect setae. Pronotum distinctly narrower than elytral base, elongate, with maximal width in anterior third, sides without lateral tubercle or spine, narrowed posteriorly; base straight, disc moderately convex, covered with dimorphic pubescence; pronotal surface double punctured. Elytra comparatively short (about twice as long as wide, and nearly two and a half times longer than pronotum); base straight with a distinct elongate gibbosity bearing a sparser and finer punctation; humeri rounded; sides slightly constricted in middle, apex rounded; disc covered with obliterate punctation on apical half, and pattern consisting of a recumbent light pubescence. Legs long and slender, covered with recumbent fine pubescence; femora slightly clubbed; tibiae almost straight, with two apical spurs of equal length; procoxae rounded, narrowly separated by prosternal process; procoxal cavities narrowly open posteriorly; mesocoxal cavities open; metatarsi one-half as long as tibiae; metatarsomere I long, hardly twice as long as tarsomeres II and III combined; claws simple.	en	Alekseev, Vitalii Igorevich, Vitali, Francesco (2020): Description of the first extinct member of the tribe Anaglyptini (Coleoptera: Cerambycidae) from European Tertiary. Zootaxa 4816 (1): 135-143, DOI: 10.11646/zootaxa.4816.1.11
4A268793623CFFCAFF0DFC0FFBB7FDE4.taxon	etymology	Etymology. The name is a combination of the old Greek word ἄκανθα (ákantha, i. e. thorn, spine) and the Greek root γλυπτός (glyptus, i. e. carved, graved) referring to the tribal type-genus Anaglyptus. Gender masculine. Composition. The new genus is monotypic, represented by the type species only.	en	Alekseev, Vitalii Igorevich, Vitali, Francesco (2020): Description of the first extinct member of the tribe Anaglyptini (Coleoptera: Cerambycidae) from European Tertiary. Zootaxa 4816 (1): 135-143, DOI: 10.11646/zootaxa.4816.1.11
4A268793623FFFCEFF0DFD0BFF04FEC4.taxon	description	(Figs. 1 – 5)	en	Alekseev, Vitalii Igorevich, Vitali, Francesco (2020): Description of the first extinct member of the tribe Anaglyptini (Coleoptera: Cerambycidae) from European Tertiary. Zootaxa 4816 (1): 135-143, DOI: 10.11646/zootaxa.4816.1.11
4A268793623FFFCEFF0DFD0BFF04FEC4.taxon	materials_examined	Type strata. Baltic amber, Blaue Erde deposits, Cenozoic: mid-Eocene to Upper Eocene. Type locality. Yantarny settlement (formerly Palmnicken), Sambian (Samland) Peninsula, Kaliningrad Region, Russia. Type material. Holotype: MAIG 6080, adult. Sex: female. The specimen is included in narrow and long, orange-yellow amber piece with approximate dimensions of 42 × 7 × 5 mm and preserved without supplementary fixation. It lacks the abdomen (only imprint on amber are available), the middle left leg, except for the coxa and the left hind leg. Syninclusions are represented by few stellate fagacean trichomes.	en	Alekseev, Vitalii Igorevich, Vitali, Francesco (2020): Description of the first extinct member of the tribe Anaglyptini (Coleoptera: Cerambycidae) from European Tertiary. Zootaxa 4816 (1): 135-143, DOI: 10.11646/zootaxa.4816.1.11
4A268793623FFFCEFF0DFD0BFF04FEC4.taxon	description	Description. Total body length about 9.0 mm; maximum length of pronotum 2.1 mm; pronotal maximum width 1.6 mm; length of an elytron along suture 5.2 mm; maximal width of elytra 2.5 mm; length of antenna about 8.0 mm. Frons densely, finely punctate; longitudinal furrow reaching posteriorly the vertex; genae slightly emarginate anteriorly; distance between antennal tubercles wider than distance between inner margins upper eye lobes; posterior area of head, finely transversely wrinkled dorsally and laterally. Scape slightly curved, covered with recumbent sparse setae; pedicel as long as wide, less than one-third as long as scape; antennomere III slightly shorter than scape (excluding apical spine); antennomeres IV-VII with same length, as long as scape; antennomere VIII three-fourth as long as scape; antennomere IX slightly shorter than VIII; antennomeres X-XI with same length, two-thirds as long as antennomere III; antennomere proportions according to the formula: 1.6: 0.5: 1.5: 1.6: 1.6: 1.6: 1.6: 1.2: 1.1: 1.0: 1.0. Pronotum about one-third longer than wide; disc covered with long, erect, sparse dark setae, and large, shallow punctures separated by about 2 – 3 times their diameter; sides, base and middle with dense, recumbent light pubescence, and very fine, dense punctation. Scutellum elongate, densely covered with light recumbent pubescence. Elytra about 2.1 times as long as wide and about 2.5 times as long as pronotum; disc covered with moderately sparse, rounded punctures, irregular and deep on basal half (distance between punctures about 1 – 3 times diameter of each puncture), obliterate on apical half; each puncture bearing long erect or semirecumbent dark seta; dense light pubescence forming: 1) transverse, angulate macula laterally on basal quarter of each elytron; 2) inverted V-shaped median band; 3) apical spot on sutural angle. Ventral side not completely conserved; prosternum, meso- and metaventrite finely punctate; metanepisternum narrow and long (about five times as long as wide); metaventral process three times narrower than mesocoxa; metaventrite with long, fine metathoracic discrimen; abdominal ventrites finely punctate, bearing long, erect, sparse setae apically; pygidium broadly rounded at apex; intercoxal process narrowly triangular; ventrites proportions (measured laterally) according to the formula: 3.0: 2.0: 2.0: 2.0: 1.5. Metatarsomere I about 1.8 times as long as metatarsomeres II and III combined (or 1.1 times as long as tarsomeres II – IV together). FIGURE 6. Distribution of the tribe Anaglyptini in Recent world (green area), Yoshiakioclytus (yellowish green area), and Acanthoglyptus picollus gen. et sp. nov. (red dot).	en	Alekseev, Vitalii Igorevich, Vitali, Francesco (2020): Description of the first extinct member of the tribe Anaglyptini (Coleoptera: Cerambycidae) from European Tertiary. Zootaxa 4816 (1): 135-143, DOI: 10.11646/zootaxa.4816.1.11
4A268793623FFFCEFF0DFD0BFF04FEC4.taxon	etymology	Etymology. The name of the new species is a theonym derived from Picollus (Peckols, Pockols, or also sometimes Patolls, Patulas, Patollo), the name of the Old Prussian god of the underworld and the easy death. The species name is a noun in apposition.	en	Alekseev, Vitalii Igorevich, Vitali, Francesco (2020): Description of the first extinct member of the tribe Anaglyptini (Coleoptera: Cerambycidae) from European Tertiary. Zootaxa 4816 (1): 135-143, DOI: 10.11646/zootaxa.4816.1.11
4A268793623FFFCEFF0DFD0BFF04FEC4.taxon	discussion	Remarks. The body integument coloration and colors of the dorsal pattern of the new fossil species are unknown.	en	Alekseev, Vitalii Igorevich, Vitali, Francesco (2020): Description of the first extinct member of the tribe Anaglyptini (Coleoptera: Cerambycidae) from European Tertiary. Zootaxa 4816 (1): 135-143, DOI: 10.11646/zootaxa.4816.1.11
4A268793623BFFCEFF0DF976FD3EF858.taxon	description	6. Mesalocerus Vitali, 2015: M. tetropoides Vitali, 2015 Asemini Thomson, 1861 7. Nothorhina Redtenbacher, 1845: N. granulicollis Zang, 1905 8. Palaeotetropium Vitali, 2011: P. saxonicum Vitali, 2011 Saphanini Gistel, 1848 9. Saphanites Vitali, 2011: S. mirabilis Vitali, 2011	en	Alekseev, Vitalii Igorevich, Vitali, Francesco (2020): Description of the first extinct member of the tribe Anaglyptini (Coleoptera: Cerambycidae) from European Tertiary. Zootaxa 4816 (1): 135-143, DOI: 10.11646/zootaxa.4816.1.11
4A268793623AFFCFFF0DFE96FAE0FC20.taxon	description	11. Protachryson Vitali, 2011: P. pomeranicum Vitali, 2011 Anaglyptini Lacordaire, 1868 12. Acanthoglyptus gen. nov.: A. picollus sp. nov. Clytini Mulsant, 1839 13. Clytus Laicharting, 1784: C. pici Piton, 1940 Obriini Mulsant, 1839 14. Obrium Dejean, 1821: O. damgaardi Vitali, 2015 15. Stenhomalus White, 1855: S. hoffeinsorum Vitali, 2014 Opsimini LeConte, 1873 16. Dicentrus LeConte, 1880: D. mehli Vitali & Daamgard, 2016 17. Europsimus Vitali, 2011: E. germanicus Vitali, 2011 18. Japonopsimus Matsushita, 1935: J. balticus Vitali, 2014 Sestyrini Lacordaire, 1868 19. Procleomenes Gressitt & Rondon, 1970: P. gouverneuri Vitali, 2018 Tillomorphini Lacordaire, 1868 20. Tillomorphites Vitali, 2011: T. robustus Vitali, 2011; T. otiliae Vitali, 2017; T. spinipes Vitali, 2017	en	Alekseev, Vitalii Igorevich, Vitali, Francesco (2020): Description of the first extinct member of the tribe Anaglyptini (Coleoptera: Cerambycidae) from European Tertiary. Zootaxa 4816 (1): 135-143, DOI: 10.11646/zootaxa.4816.1.11
4A268793623AFFCFFF0DFA82FCBCF881.taxon	description	25. Eurapatophysis Vitali, 2016: E. groehni Vitali, 2016 The biodiversity described at the moment consists of 27 extinct species, distributed in 25 genera of at least 17 tribes. Two taxa are known from Bitterfeld amber (Palaeotetropium saxonicum and Europsimus germanicus), while 25 fossils are described from Baltic amber. There is no longhorn beetles described from Oise amber or Rovno amber Lagerstätten. Two fossils are of doubtful generic placement (Dorcaschema succineum and Clytus (Xylotrechus) pici) and one species is considered to be incertae sedis within Lamiinae (Dorcadionoides subaeneus). Almost twothirds of the taxa were described in the last decade; the extension of the provided above list in the nearest future is expected. An unusual low ratio of the subfamily Lamiinae in the Palaeogene amber assemblage of Cerambycidae (compared to the current fauna) is intriguing, but could not be considered as a reliable phenomenon due to insufficient knowledge state on Palaeogene European cerambycids.	en	Alekseev, Vitalii Igorevich, Vitali, Francesco (2020): Description of the first extinct member of the tribe Anaglyptini (Coleoptera: Cerambycidae) from European Tertiary. Zootaxa 4816 (1): 135-143, DOI: 10.11646/zootaxa.4816.1.11
4A268793623AFFCFFF0DFC6AFC3DFA88.taxon	description	21. Dorcaschema Chevrolat, 1844: D. succineum Zang, 1905 Parmenini Mulsant, 1839 22. Parmenops Schaufuss, 1891: P. longicornis Schaufuss, 1891 Pogonocherini Mulsant, 1839 23. Pogonocherus Dejean, 1821: P. jaekeli Zang, 1905 Incertae sedis: 24. Dorcadionoides Motschulsky, 1856: D. subaeneus Motschulsky, 1856	en	Alekseev, Vitalii Igorevich, Vitali, Francesco (2020): Description of the first extinct member of the tribe Anaglyptini (Coleoptera: Cerambycidae) from European Tertiary. Zootaxa 4816 (1): 135-143, DOI: 10.11646/zootaxa.4816.1.11
4A268793623AFFCFFF0DFF26FBBBFF7C.taxon	description	10. Necydalis (Necydalisca) Plavilstshikov, 1936: N. (Necydalisca) zangi Vitali, 2011	en	Alekseev, Vitalii Igorevich, Vitali, Francesco (2020): Description of the first extinct member of the tribe Anaglyptini (Coleoptera: Cerambycidae) from European Tertiary. Zootaxa 4816 (1): 135-143, DOI: 10.11646/zootaxa.4816.1.11
